taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120213/files/figure.png	https://doi.org/10.5281/zenodo.7120213	FIGURE 5. Holotype (USNM 596065/RAP0892) of Desmognathus adatsihifrom Cataloochee Balsam, Great Smoky Mountains (NC: Swain) in life, photo courtesy of T.W. Pierson (KSU). This solidly striped morph is most commonly representative of Smokies populations and is distinguishable from D. imitator by the straight and well-defined borders of the uniformly colored dorsal stripe.	FIGURE 5. Holotype (USNM 596065/RAP0892) of Desmognathus adatsihifrom Cataloochee Balsam, Great Smoky Mountains (NC: Swain) in life, photo courtesy of T.W. Pierson (KSU). This solidly striped morph is most commonly representative of Smokies populations and is distinguishable from D. imitator by the straight and well-defined borders of the uniformly colored dorsal stripe.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120215/files/figure.png	https://doi.org/10.5281/zenodo.7120215	FIGURE 6. Paratypes (USNM 596063–4/RAP0890–1; a, b) and holotype (USNM 596065/RAP0892, c) of Desmognathus adatsihi from Cataloochee Balsam, Great Smoky Mountains (NC: Swain) isolated in life. Photos courtesy of T.W. Pierson; specimens not to scale.	FIGURE 6. Paratypes (USNM 596063–4/RAP0890–1; a, b) and holotype (USNM 596065/RAP0892, c) of Desmognathus adatsihi from Cataloochee Balsam, Great Smoky Mountains (NC: Swain) isolated in life. Photos courtesy of T.W. Pierson; specimens not to scale.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120217/files/figure.png	https://doi.org/10.5281/zenodo.7120217	FIGURE 7. Paratypes (USNM 596063–4/RAP0890–1; a,b) and holotype (USNM 596065/RAP0892, c) of Desmognathus adatsihi from Cataloochee Balsam, Great Smoky Mountains (NC: Swain) in dorsal view in preservative, approximately to scale with tick marks in mm. Photos by RAP.	FIGURE 7. Paratypes (USNM 596063–4/RAP0890–1; a,b) and holotype (USNM 596065/RAP0892, c) of Desmognathus adatsihi from Cataloochee Balsam, Great Smoky Mountains (NC: Swain) in dorsal view in preservative, approximately to scale with tick marks in mm. Photos by RAP.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120219/files/figure.png	https://doi.org/10.5281/zenodo.7120219	FIGURE 8. Paratypes (USNM 596063–4/RAP0890–1; a,b) and holotype (USNM 596065/RAP0892, c) of Desmognathus adatsihi from Cataloochee Balsam, Great Smoky Mountains (NC: Swain) in ventral view in preservative, approximately to scale with tick marks in mm. Photos by RAP. Dorsal pattern of erythristic morph (a) shows significant intrusion onto ventral surfaces, compared to the relatively immaculate spotted and striped morphs (b, c).	FIGURE 8. Paratypes (USNM 596063–4/RAP0890–1; a,b) and holotype (USNM 596065/RAP0892, c) of Desmognathus adatsihi from Cataloochee Balsam, Great Smoky Mountains (NC: Swain) in ventral view in preservative, approximately to scale with tick marks in mm. Photos by RAP. Dorsal pattern of erythristic morph (a) shows significant intrusion onto ventral surfaces, compared to the relatively immaculate spotted and striped morphs (b, c).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120221/files/figure.png	https://doi.org/10.5281/zenodo.7120221	FIGURE 9. The type locality of Desmognathus adatsihi on Cataloochee Balsam, Great Smoky Mountains National Park (NC: Swain), ~1650m ASL. The habitat is classic “spruce-fir” forest. The type series were captured under rocks and logs in the foreground. Photo courtesy of T.W. Pierson.	FIGURE 9. The type locality of Desmognathus adatsihi on Cataloochee Balsam, Great Smoky Mountains National Park (NC: Swain), ~1650m ASL. The habitat is classic “spruce-fir” forest. The type series were captured under rocks and logs in the foreground. Photo courtesy of T.W. Pierson.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120227/files/figure.png	https://doi.org/10.5281/zenodo.7120227	FIGURE 10. Paratype (USNM 596063/RAP0890) of Desmognathus adatsihi from Cataloochee Balsam (NC: Swain) in life, photo courtesy of T.W. Pierson. This brilliantly colored erythristic morph and its xanthic counterpart are infrequently observed, primarily on Cataloochee Balsam near the type locality, but also very rarely on Clingmans Dome in the Smokies (Fig. 14a) and Waterrock Knob in the Plott Balsams (S. G. Tilley, pers. comm.). A similar specimen from near the type locality was mislabeled as D. imitator by Dodd (2004).	FIGURE 10. Paratype (USNM 596063/RAP0890) of Desmognathus adatsihi from Cataloochee Balsam (NC: Swain) in life, photo courtesy of T.W. Pierson. This brilliantly colored erythristic morph and its xanthic counterpart are infrequently observed, primarily on Cataloochee Balsam near the type locality, but also very rarely on Clingmans Dome in the Smokies (Fig. 14a) and Waterrock Knob in the Plott Balsams (S. G. Tilley, pers. comm.). A similar specimen from near the type locality was mislabeled as D. imitator by Dodd (2004).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120231/files/figure.png	https://doi.org/10.5281/zenodo.7120231	FIGURE 11. Paratype (USNM 596064/RAP0891) of Desmognathus adatsihi from Cataloochee Balsam (NC: Swain) in life, photo courtesy of T. W. Pierson. This partially spotted morph is the only color pattern that is likely to be confused with D. imitator, but the spots are more contiguous, particularly towards the tail, with an overall lighter ground color (see Tilley 2000).	FIGURE 11. Paratype (USNM 596064/RAP0891) of Desmognathus adatsihi from Cataloochee Balsam (NC: Swain) in life, photo courtesy of T. W. Pierson. This partially spotted morph is the only color pattern that is likely to be confused with D. imitator, but the spots are more contiguous, particularly towards the tail, with an overall lighter ground color (see Tilley 2000).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120233/files/figure.png	https://doi.org/10.5281/zenodo.7120233	FIGURE 12. Comparison of large, darkened adult males (based on elongated snouts and peramorphic jaw musculature) of Desmognathus adatsihi (a: RAP2816; Rough Ridge, NC: Haywood) and D. balsameus (b: RAP2827; Bear Pen Gap, NC: Jackson; Fig. 16a) in life, showing the similar overall melanic coloration taken on with age. Photos courtesy of M.A. Seldes (UGA).	FIGURE 12. Comparison of large, darkened adult males (based on elongated snouts and peramorphic jaw musculature) of Desmognathus adatsihi (a: RAP2816; Rough Ridge, NC: Haywood) and D. balsameus (b: RAP2827; Bear Pen Gap, NC: Jackson; Fig. 16a) in life, showing the similar overall melanic coloration taken on with age. Photos courtesy of M.A. Seldes (UGA).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120237/files/figure.png	https://doi.org/10.5281/zenodo.7120237	FIGURE 13. Photographs of two adult Desmognathus adatsihi (a: RAP2818; b: RAP2820) in life, from the paratype locality on Rough Ridge (NC: Haywood). Photos courtesy of M.A. Seldes.	FIGURE 13. Photographs of two adult Desmognathus adatsihi (a: RAP2818; b: RAP2820) in life, from the paratype locality on Rough Ridge (NC: Haywood). Photos courtesy of M.A. Seldes.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120239/files/figure.png	https://doi.org/10.5281/zenodo.7120239	FIGURE 14. Comparison of small adult Desmognathus adatsihi (a: not collected; Clingmans Dome, TN: Sevier/NC: Swain) and exceptionally large (54mm SVL) adult D. balsameus (b: RAP2831; Lost Knob, NC: Jackson) in life, showing the dramatic possible differences in body form and coloration. Photos courtesy of M.A. Seldes.	FIGURE 14. Comparison of small adult Desmognathus adatsihi (a: not collected; Clingmans Dome, TN: Sevier/NC: Swain) and exceptionally large (54mm SVL) adult D. balsameus (b: RAP2831; Lost Knob, NC: Jackson) in life, showing the dramatic possible differences in body form and coloration. Photos courtesy of M.A. Seldes.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120211/files/figure.png	https://doi.org/10.5281/zenodo.7120211	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5023F4FC4BA1F2770890916.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120243/files/figure.png	https://doi.org/10.5281/zenodo.7120243	FIGURE 15. An aberrant specimen (RAP0908) of Desmognathus adatsihi from Rough Ridge (NC: Haywood) in life on isolated (a) and natural (b) backgrounds. In contrast to the regularly encountered erythristic and xanthic morphs, we hypothesize this individual to be a singular mutant, probably leucistic, rather than a form which occurs with any frequency in the population. Photos courtesy of T.W. Pierson.	FIGURE 15. An aberrant specimen (RAP0908) of Desmognathus adatsihi from Rough Ridge (NC: Haywood) in life on isolated (a) and natural (b) backgrounds. In contrast to the regularly encountered erythristic and xanthic morphs, we hypothesize this individual to be a singular mutant, probably leucistic, rather than a form which occurs with any frequency in the population. Photos courtesy of T.W. Pierson.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120245/files/figure.png	https://doi.org/10.5281/zenodo.7120245	FIGURE 16. The holotype (AMNH A-193885/RAP0650) of Desmognathus balsameus from Bubbling Spring Branch (NC: Haywood), ~1530m ASL. This adult specimen (43mm SVL) has a relatively dull dorsal color-pattern partially obscured by ontogenetic darkening, common in this species. Photos by RAP.	FIGURE 16. The holotype (AMNH A-193885/RAP0650) of Desmognathus balsameus from Bubbling Spring Branch (NC: Haywood), ~1530m ASL. This adult specimen (43mm SVL) has a relatively dull dorsal color-pattern partially obscured by ontogenetic darkening, common in this species. Photos by RAP.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120247/files/figure.png	https://doi.org/10.5281/zenodo.7120247	FIGURE 17. Comparative specimens of Desmognathus balsameus in preservative in dorsal (a, c) and ventral (b, d) views, showing dorsal stripe with wavy or undulating edges (RAP0979; Black Balsam Knob, NC: Haywood; a, b), and the early ontogenetic stage of this pattern with more distinct paired spots (RAP0990; Burnette Cove Creek, NC: Haywood; c, d).	FIGURE 17. Comparative specimens of Desmognathus balsameus in preservative in dorsal (a, c) and ventral (b, d) views, showing dorsal stripe with wavy or undulating edges (RAP0979; Black Balsam Knob, NC: Haywood; a, b), and the early ontogenetic stage of this pattern with more distinct paired spots (RAP0990; Burnette Cove Creek, NC: Haywood; c, d).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120249/files/figure.png	https://doi.org/10.5281/zenodo.7120249	FIGURE 18. Photographs of a juvenile (a: RAP2824) and adult (b: RAP2825) Desmognathus balsameus from Bear Pen Gap (NC: Jackson) in life. Photos courtesy of M.A. Seldes.	FIGURE 18. Photographs of a juvenile (a: RAP2824) and adult (b: RAP2825) Desmognathus balsameus from Bear Pen Gap (NC: Jackson) in life. Photos courtesy of M.A. Seldes.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120233/files/figure.png	https://doi.org/10.5281/zenodo.7120233	FIGURE 12. Comparison of large, darkened adult males (based on elongated snouts and peramorphic jaw musculature) of Desmognathus adatsihi (a: RAP2816; Rough Ridge, NC: Haywood) and D. balsameus (b: RAP2827; Bear Pen Gap, NC: Jackson; Fig. 16a) in life, showing the similar overall melanic coloration taken on with age. Photos courtesy of M.A. Seldes (UGA).	FIGURE 12. Comparison of large, darkened adult males (based on elongated snouts and peramorphic jaw musculature) of Desmognathus adatsihi (a: RAP2816; Rough Ridge, NC: Haywood) and D. balsameus (b: RAP2827; Bear Pen Gap, NC: Jackson; Fig. 16a) in life, showing the similar overall melanic coloration taken on with age. Photos courtesy of M.A. Seldes (UGA).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120239/files/figure.png	https://doi.org/10.5281/zenodo.7120239	FIGURE 14. Comparison of small adult Desmognathus adatsihi (a: not collected; Clingmans Dome, TN: Sevier/NC: Swain) and exceptionally large (54mm SVL) adult D. balsameus (b: RAP2831; Lost Knob, NC: Jackson) in life, showing the dramatic possible differences in body form and coloration. Photos courtesy of M.A. Seldes.	FIGURE 14. Comparison of small adult Desmognathus adatsihi (a: not collected; Clingmans Dome, TN: Sevier/NC: Swain) and exceptionally large (54mm SVL) adult D. balsameus (b: RAP2831; Lost Knob, NC: Jackson) in life, showing the dramatic possible differences in body form and coloration. Photos courtesy of M.A. Seldes.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120255/files/figure.png	https://doi.org/10.5281/zenodo.7120255	FIGURE 19. Live photograph of two large adult male Desmognathus balsameus (RAP2826–7) in combat (a), found in situ under a log in the understory of a coniferous forest (b) near Bear Pen Gap (NC: Jackson; ~1670m ASL). No females were observed nearby, suggesting a possible territorial nature for the interaction. Both individuals show overall darkened adult coloration with few if any discernable color-pattern remnants.Adult Eurycea cf. wilderae and Plethodon metcalfi were found under similar logs, and D. wrighti is known from the site (Pyron et al. 2022c). The encounter occurred at ~1000h on 24 July 2022 and was found by M.A. Seldes. Photos by RAP.	FIGURE 19. Live photograph of two large adult male Desmognathus balsameus (RAP2826–7) in combat (a), found in situ under a log in the understory of a coniferous forest (b) near Bear Pen Gap (NC: Jackson; ~1670m ASL). No females were observed nearby, suggesting a possible territorial nature for the interaction. Both individuals show overall darkened adult coloration with few if any discernable color-pattern remnants.Adult Eurycea cf. wilderae and Plethodon metcalfi were found under similar logs, and D. wrighti is known from the site (Pyron et al. 2022c). The encounter occurred at ~1000h on 24 July 2022 and was found by M.A. Seldes. Photos by RAP.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5183F4EC4BA1EC072920E26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120211/files/figure.png	https://doi.org/10.5281/zenodo.7120211	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5193F4AC4BA191F72C40EB6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120207/files/figure.png	https://doi.org/10.5281/zenodo.7120207	FIGURE 2. Illustrated holotype (ERA-WTN 14150) of Desmognathus perlapsus Neill, 1950 from a “rocky outcropping on the western wall of Tallulah Gorge, near the town of Tallulah Falls, Rabun [actually Habersham] County, Georgia.” The specimen, an adult male 38.6mm SVL, is not currently known to exist but may be in the FLMNH. Reprinted from the original description.	FIGURE 2. Illustrated holotype (ERA-WTN 14150) of Desmognathus perlapsus Neill, 1950 from a “rocky outcropping on the western wall of Tallulah Gorge, near the town of Tallulah Falls, Rabun [actually Habersham] County, Georgia.” The specimen, an adult male 38.6mm SVL, is not currently known to exist but may be in the FLMNH. Reprinted from the original description.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C5193F4AC4BA191F72C40EB6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120211/files/figure.png	https://doi.org/10.5281/zenodo.7120211	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C51D3F45C4BA19A272EB0F56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120203/files/figure.png	https://doi.org/10.5281/zenodo.7120203	FIGURE 1. Holotype (USNM 128007, field tag J.C.N. 1001) of Desmognathus ocoee Nicholls, 1949 from “the surface and in crevices of cliffs at Ship’s Prow Rock, in Ocoee Gorge, beside U.S. Highway 64, nine miles airline west of Ducktown, in Polk County, Tennessee,” collected 14 November 1948 by J.C. Nicholls, Jr. Note prominence of offset paired larval spots on the dorsum. Photos in dorsal (a) and ventral (b) views courtesy of the NMNH.	FIGURE 1. Holotype (USNM 128007, field tag J.C.N. 1001) of Desmognathus ocoee Nicholls, 1949 from “the surface and in crevices of cliffs at Ship’s Prow Rock, in Ocoee Gorge, beside U.S. Highway 64, nine miles airline west of Ducktown, in Polk County, Tennessee,” collected 14 November 1948 by J.C. Nicholls, Jr. Note prominence of offset paired larval spots on the dorsum. Photos in dorsal (a) and ventral (b) views courtesy of the NMNH.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C51D3F45C4BA19A272EB0F56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120257/files/figure.png	https://doi.org/10.5281/zenodo.7120257	FIGURE 20. Live specimens of Desmognathus ocoee Nicholls, 1949 in lateral views (a, b) from the isolated Cumberland Plateau (TN: Marion) population (ocoee H), west of the Tennessee River close to the southernmost populations of D. abditus (see Drukker et al. 2018). Note the relatively prominent dorsal keel on the distal portion of the tail, and the indistinct or uniform, mostly brownish color-pattern. Photos courtesy of S. Drukker (Sewanee).	FIGURE 20. Live specimens of Desmognathus ocoee Nicholls, 1949 in lateral views (a, b) from the isolated Cumberland Plateau (TN: Marion) population (ocoee H), west of the Tennessee River close to the southernmost populations of D. abditus (see Drukker et al. 2018). Note the relatively prominent dorsal keel on the distal portion of the tail, and the indistinct or uniform, mostly brownish color-pattern. Photos courtesy of S. Drukker (Sewanee).	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C51D3F45C4BA19A272EB0F56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120211/files/figure.png	https://doi.org/10.5281/zenodo.7120211	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	FIGURE 4. Map of the five species considered by our taxonomy from genetically verified recent localities (RAP collection; Anderson & Tilley 2003; Beamer & Lamb 2020; Karlin & Guttman 1986; Kozak et al. 2005; Means et al. 2017; Pyron et al. 2022c; Tilley 2000; Tilley et al. 1978, 2008; Tilley & Mahoney 1996). We include apalachicolae A2 and ocoee E within our concept of D. ocoee, for which the nominotypical lineage is ocoee F/G/H. Note that some individuals from these populations are admixed between one or more species, and hybrid zones exist between phylogeographic lineages within D. ocoee and D. perlapsus, primarily at the geographic margins where each adjacent lineage contacts its neighbor. Type localities are indicated with triangles, the population of unknown affinity from Horse Cove (GA: Gilmer; Fig. 21) is indicated with a gray dot, and the potential Cohutta populations are indicated with a “?.”	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
039087A5C51D3F45C4BA19A272EB0F56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7120261/files/figure.png	https://doi.org/10.5281/zenodo.7120261	FIGURE 21. Photographs of Desmognathus cf. ocoee from Horse Cove, Rich Mountains (GA: Gilmer) in life (a: one of the specimens in the series MCZ A-143377–86 collected 29 June 2003; photo courtesy of C.D. Camp, Piedmont) and preservative (RAP2715, collected 14 July 2022) in dorsal (b) and ventral (c) views. The paired dorsal spots on the trunk and tail of the latter specimen were brighter and more yellowish overall in life. The phylogenetic and taxonomic affinity of this population is unknown.	FIGURE 21. Photographs of Desmognathus cf. ocoee from Horse Cove, Rich Mountains (GA: Gilmer) in life (a: one of the specimens in the series MCZ A-143377–86 collected 29 June 2003; photo courtesy of C.D. Camp, Piedmont) and preservative (RAP2715, collected 14 July 2022) in dorsal (b) and ventral (c) views. The paired dorsal spots on the trunk and tail of the latter specimen were brighter and more yellowish overall in life. The phylogenetic and taxonomic affinity of this population is unknown.	2022-09-28	Pyron, R. Alexander;Beamer, David A.		Zenodo	biologists	Pyron, R. Alexander;Beamer, David A.			
