taxonID	type	description	language	source
03F14B1A5B40BA33FF24E88B190749F2.taxon	materials_examined	Material examined. NEPAL: 2 ♂♂: Mahakali / Darchula, vicinity of Thaisain, around Brum (holy) Lake, 29 ° 52 ’ 16 ″ N 80 ° 41 ’ 21 ″ E. 3600 m a. s. l. 20.06.2017. A. Weigel leg. (cSh, NME).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B40BA33FF24E88B190749F2.taxon	discussion	Remarks. Amphichroum anthobioides is known from several localities in India and Nepal (Shavrin 2021).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B40BA31FF24EA481D044F2C.taxon	description	(Figs. 1 – 5)	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B40BA31FF24EA481D044F2C.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 1): ‘ CHINA: YUNNAN PROv., | Gongshan Co., | Biluo Mts. Pass, 3890 – 3910 m, | 28 ° 04.5 ′ N, 098 ° 45.6 ′ E, | D. Král & J. Růžička leg. ’ <printed>, ‘ 5. vii. 2019, sift # 06 [Y 06], | mixed forest with dominant | Abies and Rhododendron shrubs, | litter under shrubs and along | fallen trunks’ <printed>, ‘ HOLOTYPE | Amphichroum | discolor sp. n. | Shavrin A. V. des. 2022 ’ <red, printed> (NMPC). Paratypes: 24 ♂♂ (four specimens dissected), 27 ♀♀: same data as the holotype, with additional red printed label: ‘ PARATYPE | Amphichroum | discolor sp. n. | Shavrin A. V. des. 2022 ’ <red, printed> (3 ♂♂, 3 ♀♀: cSh; 21 ♂♂, 24 ♀♀: NMPC).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B40BA31FF24EA481D044F2C.taxon	description	Description. Measurements (n = 52): HW: 0.47 – 0.60; HL: 0.27 – 0.35; AL (holotype): 1.12; OL: 0.15 – 0.17; PL: 0.47 – 0.65; PW: 0.75 – 0.90; ESL: 0.88 – 1.20; EW: 0.95 – 1.02; AW: 0.94 – 1.07; MTbL (holotype): 0.57; MTrL (holotype): 0.32 (MTrL 1 – 4: 0.15; MTrL 5: 0.17); AedL: 0.45 – 0.60; TL: 2.35 – 3.35 (holotype: 2.70). Habitus as in Figs. 1 – 2. Body yellow-brown to brown; antennomeres 3 – 11 (brown specimens) or 5 – 11 (pale specimens) brownish; mouthparts, antennomeres 1 – 2 or 1 – 4 and legs yellow to yellow-brown (some specimens with yellow-brown forebody and narrow brown longitudinal spot along midline (Fig. 2), brown abdomen and yellowish paratergites; pronotum of specimens with brown body (Fig. 1) with yellow-brown lateral margins and slightly paler elytra). Body shiny; head with dense transverse microsculpture, denser and isodiametric on infraorbital ridges (some pale specimens without distinct meshes on infraorbital ridges); neck with distinct isodiametric or transverse microreticulation; pronotum with indistinct transverse microsculpure or without it; visible part of scutellum with very fine transverse meshes or without them; abdominal tergites with very dense isodiametric microsculpture. Head with several very fine and sparse punctures in middle; pronotum with moderately dense, large and deep punctation, sparser in middle and mediobasal portions; punctation of elytra distinctly larger, deeper and denser than that on pronotum, denser and coarser in mediobasal portion, finer and sparser along suture; abdominal tergites with indistinct and very sparse moderately fine punctation. Frontal portion of head with several long setae; pronotum and elytra with dense long setation; abdominal tergites with regular, short and moderately dense pubescence. Head 1.7 times as broad as long, distinctly flattened in middle and slightly elevated on infraorbital ridges; anteocellar foveae (grooves in front of ocelli) narrow and very long, diagonally stretching toward antennal bases to about level of anterior margins of eyes. Ocelli large, located at level of posterior margins of eyes near occipital furrow, distance between ocelli about as long as distance between ocellus and posterior margin of eye or slightly longer. Eyes very large, convex. Apical segment of maxillary palpi 1.1 – 1.3 times as long as preceding segment, from basal portion gradually narrowed apicad. Antenna moderately long, reaching basal third of elytral length when reclined; basal antennomere moderately long, 1.4 – 1.6 times as long as broad, antennomere 2 about as broad as and distinctly shorter than basal antennomere, 3 about as long as and narrower than 2, 4 slightly shorter and broader than 3, 5 – 9 about as long as and distinctly broader than 4, 10 slightly shorter and broader than 9, apical antennomere about twice longer than 10, from apical third strongly narrowed toward subacute apex. Pronotum convex, 1.3 – 1.5 times as broad as long, 1.5 times as broad as head, widest in middle, gradually or slightly more narrowed anteriad than posteriad; anterior angles rounded, distinctly protruded apicad; posterior angles widely rounded; lateral portions widely flattened and slightly explanate, gradually broadened posteriad. Elytra about as broad as long, distinctly broadened apicad, reaching apical margin of abdominal tergite IV or V, 1.8 times as long as pronotum, with widely rounded apical margins. Legs moderately long; metatibia 1.7 times as long as metatrasus, with long strong thorns on inner and outer surface. Abdomen about as broad as elytra, with a pair of small transverse tomentose spots in middle of tergite IV, with narrow palisade fringe on apical margin of tergite VII. Male. Protarsomeres 1 – 4 wide. Medial margin of apical half of protibia with two parallel rows of several short peg setae; mesotibia strongly curved mediad, with dense row of 17 – 21 very short thorns beginning from medial side of bend and stretching to apex of mesotibia. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII slightly concave. Aedeagus with very wide basal portion, strongly narrowed apicad toward very narrow and long apical portion of median lobe; parameres narrow, slightly exceeding apex of median lobe, with three short apical setae; internal sac long, with two fields of small thorns (Fig. 3 – 4). Lateral aspect of the aedeagus as in Fig. 5. Female. Protarsomeres 1 – 4 narrow. Medial margin of protibia without peg setae; mesotibia without modifications, slender, gradually slightly widened apicad. Abdominal tergite VIII and sternite VIII with truncate or rounded apical margin. Comparative notes. Based on the general shape of the body and apical portion of the median lobe, it is similar to Chinese A. subaequale Shavrin & Smetana, 2018 (Shaanxi), A. cuccodoroi Shavrin, 2022 (Yunnan), and the Himalayan A. altivagans Cameron, 1941 (India: Kashmir) and A. telnovi Shavrin, 2021 (Nepal). From A. subaequale it differs by the less transverse pronotum, and shorter and narrower parameres. From A. cuccodoroi, A. altivagans and A. telnovi it differs by the absence of the large sclerotized teeth in the internal sac. Additionally, it can be distinguished from: A. cuccodoroi by the less transverse pronotum, longer elytra, and significantly shorter parameres; A. altivagans by the longer antennomeres and shorter elytra; A. telnovi by the smaller body, longer antennomeres 6 – 10, slightly broader pronotum and elytra. From all these species it can be distinguished by the darker elytra, and the shape of very narrow and elongate apical portion of the median lobe.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B40BA31FF24EA481D044F2C.taxon	distribution	Distribution. The species is at present known only from the type locality in Biluo Mts., Yunnan, China. Bionomics. Specimens were collected at elevation from 3890 to 3910 m a. s. l. together with specimens of A. grandidentatum sp. n. (see below). They were taken by sifting litter under shrubs and along fallen trunks in mixed forest with Abies and Rhododendron.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B40BA31FF24EA481D044F2C.taxon	etymology	Etymology. The specific epithet (Latin adjective: of different colours) refers to the different coloration of the body of specimens.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B42BA37FF24EDF41D5A4FB8.taxon	description	(Figs. 7 – 8, 11)	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B42BA37FF24EDF41D5A4FB8.taxon	materials_examined	Type material examined. Holotype ♂ (Fig. 11; left antennomeres 10 – 11 missing): ‘ CHINA: YUNNAN PROv., | Gongshan Co., | Biluo Mts. Pass, 3890 – 3910 m, | 28 ° 04.5 ′ N, 098 ° 45.6 ′ E, | D. Král & J. Růžička leg. ’ <printed>, ‘ 5. vii. 2019, sift # 06 [Y 06], | mixed forest with dominant | Abies and Rhododendron shrubs, | litter under shrubs and along | fallen trunks’ <printed>, ‘ HOLOTYPE | Amphichroum | grandidentatum sp. n. | Shavrin A. V. des. 2022 ’ <red, printed> (NMPC). Paratypes: 3 ♂♂ (two specimens dissected; one specimen without left antennomeres 7 – 11), 5 ♀♀ (three specimens without left antennomeres 10 – 11, 7 – 11 or 11 respectively): same data as the holotype, with additional red printed label: ‘ PARATYPE | Amphichroum | grandidentatum sp. n. | Shavrin A. V. des. 2022 ’ <red, printed> (1 ♂, 1 ♀: cSh; 2 ♂♂, 4 ♀♀: NMPC).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B42BA37FF24EDF41D5A4FB8.taxon	description	Description. Measurements (n = 9): HW: 0.64 – 0.68; HL: 0.41 – 0.44; AL (holotype): 1.68; OL: 0.16 – 0.20; PL: 0.56 – 0.67; PW: 0.98 – 1.17; ESL: 1.26 – 1.30; EW: 1.31 – 1.37; AW: 1.19 – 1.29; MTbL (holotype): 0.72; MTrL (holotype): 0.61 (MTrL 1 – 4: 0.32; MTrL 5: 0.29); AedL: 0.55 – 0.58; TL: 2.95 – 3.70 (holotype: 3.32). Habitus as in Fig. 11. Head, antennomeres 4 – 11, pronotum and abdomen brown or reddish-brown (frontal portion and infraorbital ridges of head and sometimes paratergites distinctly paler); lateral and basal portions of pronotum and elytra yellow-brown; mouthparts, antennomeres 1 – 3 and legs yellow (two preapical segments of maxillary palpi darker, brown or reddish-brown). Head with very dense transverse microsculpture, sometimes finer in middle and coarser on infraorbital ridges; neck with dense isodiametric microreticulation; pronotum with dense isodiametric microsculpture, missing in mediobasal third in some paratypes; visible part of scutellum with dense isodiametric meshes. Head with sparse and moderately large punctation, denser and deeper in middle, finer, sparser and sometimes indistinct on infraorbital ridges; pronotum with moderately dense punctation, slightly finer than that in middle portion of head, sometimes sparser in middle and mediobasal portions; punctation of elytra denser, larger and deeper than that on pronotum, finer along suture. Head 1.5 times as broad as long; anteocellar foveae narrow and very deep, diagonally stretching toward supraantennal elevation. Ocelli very large, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palpi slightly longer than precending segment, from basal portion strongly narrowed toward acute apex. Antennomeres 5 – 7 about as long as and slightly broader than 4, 8 – 10 distinctly shorter and slightly broader than 7, apical antennomere 1.5 – 1.7 times as long as 10. Pronotum transverse, 1.7 times as broad as long, 1.5 – 1.7 times as broad as head, disitnctly more narrowed anteriad than posteriad; widely rounded anterior angles distinctly protruded anteriad; lateral portions very wide, flattened and explanate. Elytra wide, distinctly broader than long, reaching basal portion of abdominal tergite IV, about or more than twice as long as pronotum. Metatibia 1.1 times as long as very long metatarsus. Abdomen distinctly narrower than elytra, with a pair of indistinct and very wide transverse tomentose spots in middle of tergite IV. Male. Medial margin of apical half of protibia with two parallel rows of several moderately long peg setae; mesotibia strongly curved mediad, with dense row of 18 – 20 short thorns. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII widely rounded. Aedeagus with very wide basal portion, strongly narrowed towards rounded apex; parameres moderately wide, long and slightly curved, significantly exceeding somewhat rounded apex of median lobe, with one short apical and one preapical setae; internal sac long, with a row of large sclerotized thorns in middle and four strongly sclerotized, very large teeth in basal portions (Fig. 7). Lateral aspect of the aedeagus as in Fig. 8. Female. Medial margin of protibia and mesotibia without modifications. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded. Comparative notes. Amphichroum grandidentatum sp. n. differs from remaining species of the genus by the presence of large sclerotized teeth in basal portion of the internal sac of the aedeagus. Regarding the shape of the median lobe with rounded apex and the length of the parameres considerably exceeding apex of median lobe, it is somewhat similar to A. rotundatum Shavrin & Smetana, 2018 and A. feldmanni Shavrin, 2019 (both from Sichuan), but can be distinguished from them by the different coloration of the body and the morphology of the aedeagus. Additionally, it differs from A. feldmanni Shavrin, 2019 by the smaller body, more transverse pronotum and slightly broader elytra.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B42BA37FF24EDF41D5A4FB8.taxon	distribution	Distribution. The species is at present known only from the type locality in Biluo Mts., Yunnan, China. Bionomics. Specimens were collected in the same locality together with A. discolor sp. n. (see above).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B42BA37FF24EDF41D5A4FB8.taxon	etymology	Etymology. The specific epithet is an adjective composed of the Latin adjectives grandis, - e (large) and dentatus, - a, - um (toothed). It alludes to the presence of large sclerotized teeth in basal portion of the internal sac of the aedeagus.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B44BA37FF24ED87192C4D52.taxon	materials_examined	Material examined. NEPAL: 2 ♀♀: ‘ Kosi, Chauki, 27 ° 11 ′ – 12 ′ N 87 ° 27 ′ – 28 ′ E. 2600 – 3000 m a. s. l. 22 – 24.06. [20] 01 ’, ‘ NHMB Basel expedition to Nepal, 2001 ’ (NHMB); BHUTAN: 11 ♂♂: ‘ Dorjula 3100 m 6.6. ’, ‘ Nat. — Hist. Museum Basel — Bhutan Expedition 1972 ’ (NHMB); 1 ♂: ‘ Chimakothi 22.5. ’, ‘ Nat. — Hist. Museum Basel — Bhutan Expedition 1972 ’ (NHMB).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B44BA37FF24ED87192C4D52.taxon	discussion	Remarks. Amphichroum monticola is known from several localities of China (Tibet), Nepal and Bhutan. I restudied specimens of A. monticola reported by Coiffait (1978) from Bhutan and confirmed his identification.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B44BA37FF24EFE819034CEF.taxon	materials_examined	Material examined. NEPAL: 1 ♂: Bagmati, Sindhupalchok, Hile Kharka, 27 ° 58.607 ′ N 85 ° 48.626 ′ E. 3514 m a. s. l. 03.05.2010. E. Sprecher leg. (NHMB); 1 ♂: Mahakali / Darchula vic. Sitaula, Kopu Lekh Kulanga Khola, 29 ° 53 ′ 04 ’’ N 80 ° 44 ′ 38 ’’ E. 3500 m a. s. l., river valley, by light. A. Kopetz leg. (NME); 3 ♂♂, 2 ♀♀: Dhaulagari Mts., S slope, Bagar Khola, 28 ° 30 ′ 44 ’’ N 83 ° 32 ′ 36 ’’ E. 2250 m a. s. l. 10.05.2009. J. Schmidt leg. (cSh, NME); BHUTAN: 4 ♂♂: ‘ Dorjula 3100 m 6.6. ’, ‘ Nat. — Hist. Museum Basel — Bhutan Expedition 1972 ’ (NHMB).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B44BA37FF24EFE819034CEF.taxon	discussion	Remarks. Amphichroum pindarense is known from India, China (Tibet) and Bhutan (Shavrin 2021, 2022).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B47BA35FF24E9E21D454E14.taxon	description	(Figs. 9 – 10, 12)	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B47BA35FF24E9E21D454E14.taxon	materials_examined	Type material examined. Holotype ♂ (dissected): ‘ Sampa-Kotoka | 1400 – 2600 m’ <printed>, ‘ Nat. - Hist. Museum | Basel — Bhutan | Expedition 1972 ’ <printed>, ‘ HOLOTYPE’ <red, printed>, ‘ Amphichroum | reticulatum | H. COIFFAIT det. 19 [printed] 77 ’ <handwritten>, ‘ Amphichroum | reticulatum Coiffait, 1978 | Shavrin A. V. det. 2022 ’ (NHMB).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B47BA35FF24E9E21D454E14.taxon	description	Redescription. Measurements: HW: 0.75; HL: 0.42; AL: 2.24; OL: 0.20; PL: 0.75; PW: 1.22; ESL: 1.35; EW: 1.53; AW: 1.35; MTbL: 0.82; MTrL: 0.77 (MTrL 1 – 4: 0.45; MTrL 5: 0.32); AedL: 0.85; TL: 3.95. Habitus as in Fig. 12. Forebody, antennomeres 4 – 11 and legs yellow-brown (middle portions of elytra somewhat darker; paratergites slightly paler); mouthparts, antennomeres 1 – 3, lateral portions of pronotum and apical tarsomeres yellow. Head with dense and regular microsculpture, transverse in frontal portion and isodiametric in middle and on infraorbital ridges; neck with very dense and coarse isodiametric microreticulation; pronotum with regular, very dense, small isodiametric microsculpture; visible part of scutellum with dense transverse meshes; elytra with indistinct, fine isodiametric meshes; abdominal tergites with dense microreticulation, transverse on abdominal tergites IV – VI or isodiametric on tergites VII – VIII. Head without visible punctation; middle portion of pronotum with indistinct, very sparse and fine punctation, slightly denser and larger in basal portion; elytra with moderately large and dense punctation, finer in middle portion; abdomen without visible punctures. Body without setation except for frontal portion of head with several moderately short setae, and pronotum with very short setae along lateral margins. Head 1.7 times as broad as long, slightly flattened in middle; anteocellar foveae narrow, moderately deep and very long, almost exceeding supra-antennal elevation. Ocelli moderately large, distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Apical segment of maxillary palpi 1.3 times as long as preceding segment. Antennomere 4 distinctly shorter and slightly broader than 3, 5 – 7 slightly shorter and broader than 4, 8 – 10 slightly shorter than 7, apical antennomere about 1.3 times as long as 10. Pronotum 1.6 times as broad as long, 1.6 times as broad as head, widest in middle, gradually narrowed both apicad and posteriad; widely rounded anterior angles strongly protruded apicad. Elytra slightly broader than long, reaching apical margin of abdominal tergite IV. Metatibia slightly longer than very long metatarsus. Abdomen distinctly narrower than elytra. Male. Medial margin of apical half of protibia with two parallel rows of eight short peg setae; mesotibia strongly curved mediad, with dense row of about 20 short thorns. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII slightly concave. Aedeagus with wide basal portion, strongly narrowed apicad toward moderately narrow median lobe with subacute apex; parameres narrow, significantly exceeding apex of median lobe, with two short apical and two preapical setae; internal sac long, without additional large sclerites (Fig. 9). Lateral aspect of the aedeagus as in Fig. 10. Female unknown. Comparative notes. Based on the general shape of the body and the aedeagus, it is similar to the Himalayan A. monticola Cameron, 1928, A. nepalicum (Coiffait, 1982) and A. pindarense Champion, 1920. From A. monticola and A. nepalicum it differs by the more transverse pronotum, slightly longer elytra, and narrower apical portion of the median lobe. Additionally, from A. nepalicum it can be distinguished by the larger body. From A. pindarense it differs by the slightly broader apical portion of the median lobe and the absence of large sclerotized teeth in the internal sac.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B47BA35FF24E9E21D454E14.taxon	distribution	Distribution. Amphichroum reticulatum is at present known only from the type locality in Sampa-Kotoka, Bhutan. Bionomics. The holotype was collected at elevation between 1400 – 2600 m a. s. l. The detailed ecological data are unknown.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B46BA35FF24EC2B18764DAE.taxon	description	(Fig. 6)	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B46BA35FF24EC2B18764DAE.taxon	materials_examined	Material examined. CHINA: SICHUAN: 2 ♂♂, 5 ♀♀: Pass Zheduo Shankuo, W Kangding, E slope, 29 ° 58 N 101 ° 23 E. 3850 m a. s. l. 18. VII. 1998, J. Farkač leg. (cSh, NHMB).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B46BA35FF24EC2B18764DAE.taxon	discussion	Remarks. Amphichroum rotundatum is known from two localities in Daxue Shan and Gonnga Shan ranges in Sichuan, China (Shavrin & Smetana 2018). The present material was collected in the type locality of this species. The apical portions of the parameres is variable and can be very wide (Fig. 6).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B46BA3AFF24EE7C1C854BE0.taxon	materials_examined	Material examined. CHINA: SICHUAN: 61 ♂♂, 45 ♀♀: Pass Zheduo Shankou, W Kangding, E slope, 29 ° 59 ′ N 101 ° 50 ′ E. 3850 m a. s. l. 18.07.1998. J. Farkač leg. (cSh, NHMB); 6 ♀♀: same data, 15 km W Kangding, 29 ° 57 ′ N 101 ° 54 ′ E. 19.07.1998 (NHMB).	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
03F14B1A5B46BA3AFF24EE7C1C854BE0.taxon	discussion	Remarks. Amphichroum squamosum is known from several localities in Sichuan (Daxue Shan, Gongga Shan), China (Shavrin & Smetana 2018, Shavrin 2019). The present material was collected in the type locality of this species.	en	Shavrin, Alexey V. (2022): New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa 5190 (4): 575-583, DOI: https://doi.org/10.11646/zootaxa.5190.4.7
