taxonID	type	description	language	source
DD1006389744FF94FEDEF97F604AF8D1.taxon	description	(FIGS 4 – 10) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org: act: 5 DB 813 D 0 - FDE 9 - 45 D 8 - B 0 E 3 - 40336 A 28 DF 25 Type species: Morinoia paludosus (Cheng et al., 2011). Material examined: Holotype ♂, 8.5 mm; CHINA, Beijing, Chaoyang District, lakeside along Aosen (short name in Chinese for Olympic Forest Park); 40.01 N, 116.37 E; 67 m a. s. l.; 16 November 2017; collected by Hongguang Liu and Zhonge Hou; IZCAS-I-A 1700 – 1. Paratype. ♀, 7.6 mm; same data as for preceding; IZCASI-A 1700 – 2. Other materials, two males and two females (IZCAS-I-A 1700 – 3, 4, 5, 6), same data as holotype. All type specimens are deposited in the Institute of Zoology, Chinese Academy of Sciences (IZCAS) in Beijing, China. Etymology: The specific name refers to the type locality and is a noun in apposition. Diagnosis: Male gnathopod I carpus subtriangularshaped, with tumescent humps at posterodistal angle, female gnathopod II carpus and propodus tumescent posteriorly; uropod III peduncle with two spines on distal corner, ramus slightly shorter than peduncle; telson with one or two distal spines and rounded on distal margin. Description: Male (IZC AS-I-A 1700 – 1), 8.5 mm (Fig. 4 A). Head: Eyes rounded, inferior antennal sinus distinct. Antenna I (Fig. 6 A): short, reaching end of peduncular article 4 of antenna II; peduncular articles 1 – 3 subequal in length; flagellum with four articles. Antenna II (Fig. 6 A): peduncular article 5 slightly longer than article 4, with short spines on both margins; flagellum with 12 articles, each article with some setae. Upper lip (Fig. 5 A): convex, with minute setae. Mandible (Fig. 5 B, C): left mandible incisor five-dentate, lacinia mobilis with five weak teeth. Right mandible incisor five-dentate, lacinia mobilis bifurcate, with weak definitions, molar triturative with one long seta. Maxilla I (Fig. 5 D, E): inner plate with two plumose distal setae, outer plate with nine apical serrated spines. Maxilla II (Fig. 5 F): inner plate with one plumose seta at medial margin, outer plate with setae on apical margin and outer margin. Lower lip (Fig. 5 G): concave, with fine setae. Maxilliped (Fig. 5 H): inner plate with some plumose setae on distal margin; outer plate with apical setae; articles 1 – 2 of palp broad, article 4 distinct. Pereon: Gnathopod I (Fig. 6 C): basis with short spines on anterior and posterior margins; merus without tumescent humps, posterior margin with spines; carpus subtriangular-shaped, with tumescent humps at posterodistal angle, and three spines at base of hump; propodus about 0.7 times as long as carpus, with tumescent hump at posterodistal angle, with rows of spines, palmar margin with stiff setae; dactylus shorter than palm. Gnathopod II (Fig. 6 D): coxal plate with four spines on ventral margin; anterior margin of basis bare; propodus 1.26 times as long as wide, palmar margin convex and fringed with spines; dactylus bare on both margins. Pereopods III – VII (Fig. 7): Pereopod III (Fig. 7 A) longer than pereopod IV (Fig. 7 B); carpus of pereopod IV shorter than that of pereopod III; propodus with a distal spine at dactylar hinge; dactylus cuspidactylate. Pereopod V (Fig. 7 C): almost as long as pereopod III, coxal plate bilobed, anterior lobe slightly larger than posterior lobe, bearing setae on posterior lobe; posterior margin of basis expanded, with seven short spines, anterior margin with eight spines; merus, carpus and propodus with spines on anterior and posterior margins. Pereopod VI (Fig. 7 D): much longer than pereopod V, coxal plate bilobed; basis subrounded with eight spines on anterior margin and six setae on posterior margin; merus, carpus and propodus with spines on anterior and posterior margins; dactylus slender. Pereopod VII (Fig. 7 E): as long as pereopod VI, coxal plate small; basis strongly expanded, with 11 short spines on anterior margin and nine setae on posterior margin; merus, carpus and propodus with spines on anterior and posterior margins; dactylus 0.29 times as long as propodus. Coxal gills: present on gnathopod II and pereopods III – VI. Pleon: Epimeral plates (Fig. 6 B) with slightly pointed posterior angles, ventral margin unarmed, posterior margin with two or three minute setae. Pleopods (Fig. 10 E): similar to each other; peduncle about 1.2 times as long as rami, inner and outer rami nearly the same length, both rami fringed with plumose setae. Urosome: Uropod I (Fig. 6 E): peduncle longer than rami, with five and four spines on outer and inner margins, respectively; outer ramus a little shorter than inner ramus, outer ramus marginally bare, with four distal spines; inner ramus with one spine on outer margin, three spines on inner margin and five distal spines. Uropod II (Fig. 6 F): peduncle subequal in length with rami, with three spines on outer margin; inner ramus with two spines on outer margin and one spine on inner margin, outer ramus with one marginal spine. Uropod III (Fig. 8 A, B): peduncle with two spines on distal corner, ramus slightly shorter than peduncle, with one lateral spine, and two or four distal spines. Telson (Fig. 9 A, B): bilobed, longer than wide, apically notched, each lobe with one or two distal spines and two or three facial spines. Description of paratype female: (IZCAS-I-A 1700 – 2), 7.6 mm (Fig. 4 B). Head: Antennae I – II (Fig. 10 A) similar to those of male. Pereon: Gnathopod I (Fig. 10 B): carpus and propodus not tumescent, propodus parallel-sided, with spines ramus (ramus subequal in length to peduncle in uropod III; Fig. 8 C, D); telson rounded on distal margin (sharp on distal margin; Fig. 9 D); telson with one or two distal spines (with three spines). along anterior and posterior margins, palmar margin short, with four slender spines at posterodistal corner; cuspidactylate. Gnathopod II (Fig. 10 C, D): coxal plate with cuspidate posterior margin; basis anteriorly broad, anterior margin with many minute spines; merus with small facial spines; carpus and propodus tumescent posteriorly; propodus with spine-row on lateral surface, palm margin with stiff setae; dactylus about half of palmar margin, with a seta at hinge of nail. Pereopods III – VII: similar to those of male. Oostegites present on gnathopod II and pereopods III – V. Remarks: Morinoia aosen is similar to M. paludosus and M. japonica in antenna I shorter than antenna II, gnathopods I – II sexually dimorphic and the shape of pereopods III – VII. The new species can be distinguished from M. paludosus by the following characters (M. paludosus in parentheses): the tip of ramus in uropod III blunt (with a sharper ramus of uropod III; Fig. 8 E – G); the peduncle of uropod III longer than ramus (the peduncle of uropod III almost same length as ramus); telson rounded on distal margin (sharp on distal margin; Fig. 9 C); telson with one or two distal spines (with three or four distal spines). The new species can be distinguished from M. japonica by the following characters (M. japonica in parentheses): the peduncle of uropod III longer than	en	Liu, Hongguang, Tong, Yan, Zheng, Yami, Li, Shuqiang, Hou, Zhonge (2022): Sea-land transition drove terrestrial amphipod diversification in East Asia, with a description of a new species. Zoological Journal of the Linnean Society 196 (2): 940-958, DOI: 10.1093/zoolinnean/zlab119, URL: https://academic.oup.com/zoolinnean/article/196/2/940/6529773
