taxonID	type	description	language	source
038B493AFF80FF87FF52FC493C63A1DA.taxon	description	Description: See Ünal (1999) and Çıplak & Heller (2001). Considering above given molecular and morphological data, Bolua consists of the type species Bolua turkiyae and at least two new species named as Bolua balikesirensis Uluar & Çıplak sp. n. and Bolua bursaensis Uluar & Çıplak sp. n .. Although the Bursa and Kütahya populations show differences in temporal characteristics of the song, molecular data do not support this difference, thus were considered both as a single species until more song data is available.	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF80FF89FF52FAA1391FA42F.taxon	description	(Figures 1 – 3, 6, 9, 12, 15, 18, 21, 24, 27, 31; Tables 1 – 5) Bolua turkiyae Ünal; 1999: 246	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF80FF89FF52FAA1391FA42F.taxon	materials_examined	Type information: Holotype male, TURKEY: Bolu, Gerede, Aktas II, 1200 m, 21.08.1996, leg. M. Ünal (coll. M. Ünal in Abant Izzet Baysal University, Bolu, Turkey). Paratypes 7 males, 11 females, same data as holotype; 8 males, 5 females, same locality as holotype, 4.08.1995, leg M. Ünal (coll. M. Ünal in Abant Izzet Baysal University, Bolu, Turkey). Material examined: 10 males, 8 females TURKEY: Çankırı, Çerkes, Isık Dağı, 1050 – 1250 m, 12.08.1996 (leg. B. Çıplak) (1 male, 1 female in The Natural History Museum, London (E- 1997 / 322), others in AUZM); 2 males, 5 females, TURKEY: Kastamonu, Araç, Sıragömü yaylası, 900 m, 11.09.1998 (leg. B. Çıplak) (AUZM); 4 males, 8 females, TURKEY, Bolu, Yedigöller National Park, 40.8804 ° N, 31.68140 ° E, 1725 m, 15.08.2010 (leg. B. Çıplak, S. Kaya & Z. Boztepe) (MEVBIL); 16 males, 11 females, TURKEY, Kastamonu, Safranbolu, Karaevler, 41.34805 ° N, 32.69805 ° E, 996 m, 17.07.2017 (leg. B. Çıplak & O. Uluar) (MEVBIL); 2 males, 2 females, TURKEY, Kastamonu, Şenpazar-Ağlı road, 41.724 ° N, 33.48833 ° E, 997 m, 18.07.2017 (( leg. B. Çıplak & O. Uluar) (AUZM); 2 males, 3 females, TURKEY, Kastamonu, Yenice, 41.23168 ° N, 33.79239 ° E, 1025 m, 13.08.2004 (leg. D. Berger, D. Şirin, M. S. Taylan) (MEVBIL); 1 male, 4 females; TURKEY, Karabük, Keltepe, 41.0555 ° N, 32.468 ° E, 1453 m, 16.07.2017 (leg. B. Çıplak & O. Uluar) (MEVBIL).	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF80FF89FF52FAA1391FA42F.taxon	description	Description, diagnosis and remarks: Description of the type species Bolua turkiyae is given in Ünal (1999) and supplemented by Çıplak & Heller (2001). Data presented here (see Figures 3, 6, 9, 12, 15, 18, 21, 24, 27) are consistent with these descriptions. This species can be distinguished from B. balikesirensis sp. n. by male cercus (compare Figures 9 and 10). However, differences between B. turkiyae with B. bursaensis sp. n. are not prominent, but they can be identified by male cercus again (compare Figures 9 and 11). Apart from the qualitative morphology, the PCA analyses applied to female measurements distinguished B. turkiyae and B. bursaensis sp. n. from each other (Figure 27). Along with morphological differences, B. turkiyae corresponding PG 1 in phylogenetic tree (Figure 2) is definable by 53 unique mutations (Figure 31). The high net pairwise distance and F ST values with the other two phylogroups also support its distinctness as a species (Table 3). Although Çıplak & Heller (2001) presented and described a male calling song for this species, that song record is from Uludağ — Bursa Province, the type locality of the new species described here, P. bursaensis sp. n. Presently song of this species is not available to study and to compare with the other two. This species occupies the eastern part of the generic range in the western part of the Anatolian Black Sea Basin (Figure 1).	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8EFF88FF52F89F3C12A7B7.taxon	description	(Figures 4, 7, 10, 13, 16, 19, 22, 25, 27, 28, 31; Tables 1 – 5) LSID urn: lsid: zoobank. org: act: CD 84 EF 90 - FE 17 - 45 C 0 - 9121 - F 1902 C 616071	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8EFF88FF52F89F3C12A7B7.taxon	materials_examined	Material examined: Holotype male, TURKEY, Balıkesir, Edremit, Kaz Dağları National Park, Sarıkız Rd., 39.68975 ° N, 26.90948 ° E, 1470 m, 10.08.2010 (leg. B. Çıplak, S. Kaya & Z. Boztepe) (MEVBIL); Paratypes 6 males, 2 females, same data as holotype.	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8EFF88FF52F89F3C12A7B7.taxon	description	Description and diagnosis: As mentioned for the previous new species, members of the genus are similar in general appearance. Thus, the generic description can be applied to all species for certain features of the head, thorax and abdomen (Figures 4, 7, 10, 13, 16, 19, 22, 25, 27, 28, 31; Tables 1 – 5). To avoid repetition, a detailed description is not presented here. The new species B. balikesirensis sp. n. well differs from the other two by typical male cercus (tooth is near to middle in the first while in the beginning of apical 1 / 3 in the second; compare Figure 9 with 8 and 10). The male calling song consists of regular echemes including 10 – 26 (mean 18) syllables is another prominent diagnostic characteristic of B. balikesirensis sp. n. (Figure 28), distinguishing it from B. bursaensis sp. n. (B. turkiyae is unknown for the song). In addition to morphology, the new species that corresponds to PG 2 in the phylogenetic tree (Figure 2) can be well distinguished from the other two by 71 unique mutations (Figure 31). The high net pairwise distances and F ST values with other phylogroups also support its distinctness as a species (Table 3). This species seems to be restricted to the Kazdagları Mts in Balıkesir Province of Turkey (Figure 1).	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8EFF88FF52F89F3C12A7B7.taxon	etymology	Etymology: The new species Bolua balikesirensis was recorded from Kazdağları Mts located in Balıkesir Province of Turkey.	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8FFF8BFF52F8D138FBA067.taxon	description	(Figures 1, 2, 5, 8, 11, 14, 17, 20, 23, 26, 27, 29, 30, 31; Tables 1 – 5) LSID urn: lsid: zoobank. org: act: 13680 AA 4 - 1996 - 44 BD-B 4 E 8 - 3 A 5 A 73 F 13094	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8FFF8BFF52F8D138FBA067.taxon	materials_examined	Material examined: Holotype male, TURKEY, Bursa, Keleş-Inegöl Road, 39.93813 ° N, 29.34448 ° E, 1359 m, 19.07.2009 (leg. B. Çıplak, S. Kaya & E. M. Korkmaz) (MEVBIL); Paratypes 1 males, 2 females, same data as holotype; 1 male, 5 females, TURKEY, Kütahya, Domaniç-Inegöl Road, 39.86321 ° N, 29.64143 ° E, 1321 m, 12.08.2010 (leg. B. Çıplak, S. Kaya & Z. Boztepe) (MEVBIL); 1 female (nymph), TURKEY, Kocaeli, İzmit, Ambarcı, 40.90281 ° N, 30.18606 ° E, 138 m, 29.06.2018 (leg. B. Çıplak & S. Kaya) (MEVBIL); 1 female (nymph), TURKEY, Kocaeli, Karayakuplu, 40.87708 ° N, 29.80807 ° E, 300 m, 20.06.2016 (leg. B. Çıplak & S. Kaya) (MEVBIL); 1 male, TURKEY, Bursa, Uludağ, Kirazlı Yayla, 1465 m, 22.07.1998 (( leg. B. Çıplak) (AUZM). Apart of the material listed here, the records of B. turkiyae from Bursa and Bilecik provinces of Turkey (see Figure 1) by Çıplak & Heller (2001) refer to this species.	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8FFF8BFF52F8D138FBA067.taxon	description	Description and diagnosis: As documented in the ‘ morphology’ part of the Results section, members of Bolua are similar in overall general appearance. Thus, the generic description can be applied to all species for certain features of the head, thorax, and abdomen (Figures 5, 8, 11, 14, 17, 20, 23, 26, 27, 29, 30, 31; Tables 1 – 5). For this reason, all species are similar in general appearance except for the diagnostic character male cercus, and describing each structure in details is considered unnecessary. The new species B. bursaensis sp. n. differs from the other new species B. balikesirensis sp. n. in the structure of the male cercus (tooth is near to middle in the first while in the beginning of apical 1 / 3 in the second; compare Figures 10 and 11). Apart from morphology, B. bursaensis sp. n. well differs from B. balikesirensis sp. n. in the male calling song. In the first species, the male calling song consists of irregular long sequences of regularly repeated syllables, while in the second the song is arranged in regularly repeated echemes consisting of 10 – 26 (mean 18) syllables. B. bursaensis sp. n. cannot be easily distinguished from the type species B. turkiyae by morphology. The only structure somewhat exhibits difference is the more robust male cercus (compare Figures 9 and 11) and relatively slender ovipositor (compare Figures 24 and 26) in the first species. Additionally, the PCA applied to female measurements distinguishes the new species from the type species. Apart from phenotypic characters, the new species corresponds to PG 3 in a phylogenetic tree, can be well distinguished by 76 unique mutations (Figure 31). The high values of net pairwise distance and F ST with the other two also support its uniqueness as a species (Table 3). Sequence based species delimitation tests placed Bursa, Kütahya and Kocaeli (possibly also Bilecik) populations in the same species. Here we present the song of the Bursa and Kütahya populations. Although paternal characteristics are similar in these two populations, they are different in durations of opening and closing hemisyllables, and the number of pulses per closing hemisyllable (Figures 29, 30). Although bPTP suggests one haplotype (but not the second) from Kütahya population as a separate species, we prefer to follow other species delimitation tests and to consider both as single species until more song data are available.	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
038B493AFF8FFF8BFF52F8D138FBA067.taxon	etymology	Etymology: The new species Bolua bursaensis sp. n. occurs along the Uludağ Mts, mainly located in Bursa Province and some associated altitudes extending into Kütahya, Bilecik, and Kocaeli provinces of Turkey (Figure 1).	en	Uluar, Onur, Yahyaoğlu, Özgül, Çiplak, Battal (2022): Anatolian endemic genus Bolua (Orthoptera: Tettigoniidae: Tettigoniinae): genetic and phenotypic data indicate inconsistent diversity and evolutionary patterns. Zootaxa 5195 (3): 224-240, DOI: https://doi.org/10.11646/zootaxa.5195.3.2
