taxonID	type	description	language	source
C41B87B7C473EE75A7E88565FCE1A945.taxon	materials_examined	Type: — BRAZIL. “ Habitat in Brasiliae prov. Rio Negro ”, Martius 2762 (M; lectotype designated by Landrum 2010: 381). (Fig. 2 A – B.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C473EE75A7E88565FCE1A945.taxon	description	Trees 1 – 12 m. Twigs glabrous or glabrescent, the indumentum whitish or reddish-brown. Leaves with petioles 1 – 7 mm long, glabrous or glabrescent, unchanneled or shallowly channeled; blades elliptic to broadly so, lanceolate, ovate or obovate, 3.3 – 14.4 × 2.4 – 5.5 cm, glabrous, membranaceous or chartaceous, discolorous; glands conspicuous; base cuneate, acute or rounded; apex acuminate, rarely attenuate or cuneate; midvein adaxially sulcate or impressed; lateral veins moderately visible, not arched, 10 to 20 pairs; intra-marginal vein 1 – 2 mm from the margin, margin flat. Inflorescence axillary or terminal, fascicle, the axis ca. 3 mm long, sometimes inconspicuous, glabrous; bracts deltate, 0.3 – 0.5 × 0.6 mm, puberulent adaxially, glabrous abaxially, persistent in the fruit; pedicels 9 – 40 mm long, glabrous; bracteoles triangular to narrowly so, or deltate, 0.5 – 2 × 0.2 – 0.7 mm, glabrous, persistent in the fruit; flower buds globose, 7 × 12 mm, not constricted above ovary; calyx-lobes free, five, narrowly triangular, deltate to broadly so, 5 – 10 × 3 – 6 mm, glabrous, not reflexed, persistent in the fruit; ovary 3 - 5 - locular. Immature fruits subglobose or globose, 0.4 – 0.7 × 0.5 – 1.1 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C473EE75A7E88565FCE1A945.taxon	discussion	Note: — Calycolpus goetheanus may be recognized by its glabrous leaves, inflorescence in fascicles; buds with five free calyx-lobes (5 – 10 × 3 – 6 mm); ovary 3 – 5 - locular; and glabrous, subglobose or globose fruits.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C473EE75A7E88565FCE1A945.taxon	distribution	Distribution and habitat: — Known from Colombia, French Guiana, Guyana, Suriname, Trinidad-Tobago, Venezuela, northeastern and northwestern Brazil (Amazonas, Maranhão, Pará and Roraima states) (Landrum 2010, Flora do Brasil 2020). In Amazonian Maranhão, C. goetheanus was found growing along a road, and in both “ terra firme’ ’ and flooded forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C473EE75A7E88565FCE1A945.taxon	biology_ecology	Phenology: — Flowering in January, February, May and October to December, and fruiting in January, February and October.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C473EE75A7E88565FCE1A945.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Alcântara, Piriuaçú, 02 ° 24 32 ” S, 44 ° 24 ’ 53 ” W, 11 April 1954 (st.), R. L. Fróes 30771 (IAN!). Cururupu, Maracassumé River Region, Campo de Boa Esperança, 02 ° 02 ’ 29 ” S, 45 ° 57 ’ 19 ” W, 22 October 1932 (fl., fr.), R. Fróes 1977 (NY!). São José de Ribamar, Sítio Aguahy, 02 ° 38 ’ 47 ” S, 44 ° 09 ’ 05 ” W, 28 October 2017 (fl.), G. S. Amorim 378 (MAR!); idem, 20 February 2018 (fr.), G. S. Amorim 467 (MAR!). São Luís, Alumar, 02 º 41 ’ 30 ” S, 44 ° 9 ’ 44 ” W, 29 November 994 (fl.), N. Figueiredo 7 (MAR!); 02 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 18 January 1995 [fl], N. Figueiredo 4 (MAR!); 02 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 13 November 1995 (fl.), K. B. Ferreira & E. C. Gimos s. n. (MAR!); Estrada que vai do Rio Anil para Maioba, 10 January 1950 (fl.), R. L. Fróes 25652 (U, image); Reserva Florestal do Sacavém, 02 º 34 ’ 04 ” S, 44 º 15 ’ 44 ” W, 03 December 1988 (fl.), R. C. Reis 57 (MAR!); 22 January 1992 [fl], F. H. Muniz 17 (RB); 26 February 1992 [fl], F. H. Muniz 196 (HUEFS, image); 02 ° 35 ’ 59 ” S, 44 ° 16 ’ 34 ” W, 27 May 1992 (fl.), F. H. Muniz 256 (HUEFS, image); 02 ° 30 ’ S, 44 ° 15 ’ W, 21 January 1993 (fr.), F. H. Muniz 287 (INPA, image); Rio Anil, 02 ° 28 ’ 59 ” S, 44 ° 12 ’ 00 ” W, 13 January 1951 (fl., fr.), B. E. Franz 26348 (ASU, image); Sede da Cemar, 02 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 27 November 1985 (fl.), G. T. Prance & R. Henriques 29886 (US, image); Sítio Santa Eulália, 02 ° 30 ’ 30 ” S, 44 ° 16 ’ 39 ” W, 29 January 2018 (fl.), G. S. Amorim 414, 419 (MAR!); 02 ° 30 ’ 52 ” S, 44 ° 16 ’ 34 ” W, 26 February 2018 (fr.), G. S. Amorim 491 (MAR!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE77A7E886E9FE6FA5CD.taxon	materials_examined	Type: — FRENCH GUIANA. “ Habitat in sylvis Caiennae & Guianae ”, Aublet s. n. (holotype: BM 953694, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE77A7E886E9FE6FA5CD.taxon	description	Shrubs 1.5 – 3.5 m. Twigs glabrous or sparsely pubescent, the indumentum whitish. Leaves with petioles 7 – 13 mm long, glabrous or sparsely pubescent, the indumentum whitish, channeled or shallowly channeled on the basal half; blades elliptic to broadly so, ovate or obovate, 5.6 – 10.8 × 2.9 – 5.1 cm, glabrous or sparsely pubescent, pubescent when young, the whitish, membranaceous or chartaceous, discolorous; glands conspicuous, dark; base cuneate, rounded, or obtuse; apex acute or acuminate; midvein adaxially impressed; lateral veins impressed and covered with indumentum whitish abaxially and prominent adaxially, not arched, 5 to 12 pairs; intra-marginal vein absent, margin flat. Inflorescence axillary, a solitary flower, the axis inconspicuous; bracts lanceolate, ovate or oblong, 5 – 7 × 1 mm, puberulent, caducous after anthesis; pedicels 12 – 42 mm long, puberulent; bracteoles linear, 1.5 – 3 × 0.1 – 0.2 mm, pubescent, caducous after anthesis; flower buds globose, 3.2 – 3.9 × 3 – 3.5 mm, not constricted above ovary; calyx-lobes partially fused in the lower third of the bud, five, obovate or obtriangular, 1 – 2 × 1 mm, sparsely pubescent or pubescent, not reflexed, persistent in the fruit; ovary 3 – 7 - locular. Immature fruits globose, 0.4 – 1.1 × 0.4 – 1 cm, sparsely pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE77A7E886E9FE6FA5CD.taxon	discussion	Note: — Campomanesia aromatica may be recognized by having blades with lateral veins covered with whitish indumentum abaxially, impressed, 5 to 12 on each side; the flowers are solitary with pedicels 12 – 42 mm long; calyxlobes partially fused in the lower third of the bud, five; and a 3 – 7 - locular ovary.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE77A7E886E9FE6FA5CD.taxon	distribution	Distribution and habitat: — Known from Bolivia, French Guiana, Guyana, Suriname, Trinidad-Tobago, Venezuela and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, C. aromatica was found growing along a road, and in “ terra firme ” and secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE77A7E886E9FE6FA5CD.taxon	biology_ecology	Phenology: — Flowering in January and fruiting in January and February.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE77A7E886E9FE6FA5CD.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Alcântara, 04 February 1984 (fr.), A. M. V. Carvalho 2078 (SPF, image). São Luís, Estrada do Tirirical, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, February-March 1939 (fr.), R. L. Fróes 11526 (NY!); Sítio Andiroba, 07 August 1980 (st.), M. G. Silva 5693 (MG!); Sítio Santa Eulália, 2 ° 30 ’ 56 ” S, 44 ° 16 ’ 37 ” W, 20 January 1988 (fl.), E. Barroso 47 (SLUI!); 2 ° 30 ’ 52 ” S, 44 ° 16 ’ 34 ” W, 29 January 2018 (fr.), G. S. Amorim 417 (MAR!); 2 ° 30 ’ 52 ” S, 44 ° 16 ’ 33 ” W, 26 February 2018 (fr.), G. S. Amorim 490 (MAR!). Additional specimen examined: — BRAZIL. Maranhão: São José de Ribamar, praia de Caúra, 13 January 2017 (fl.), G. S. Amorim 507 (MAR!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE76A7E882A1FE00A2ED.taxon	materials_examined	Type: — JAMAICA. P. Browne s. n. (LINN 637.6; lectotype designated by Fawcett & Rendle 1926: 338). (Fig. 2 C-D.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE76A7E882A1FE00A2ED.taxon	description	Trees or shrubs 1 – 4 m. Twigs densely pubescents, the indumentum whitish. Leaves with petioles 2 – 4 mm long, pubescent, channeled; blades elliptic to narrowly so, lanceolate or ovate, 3 – 7.8 × 1.2 – 4.1 cm, pubescent or densely pubescent, the indumentum whitish, chartaceous, concolorous; glands conspicuous; base rounded, cuneate or attenuate; apex acuminate, long acuminate, almost caudate, generally apiculate, rarely acute; midvein adaxially impressed or sulcate; lateral veins visible abaxially, not arched, 12 to 20 pairs; intra-marginal vein ca. 1 mm from the margin, margin revolute. Inflorescences axillary, raceme simple, the axis 1.5 – 4.9 cm long, pubescent, the indumentum whitish; bracts triangular to narrowly so, or lanceolate, 2 – 3 × 1 – 3 mm, pubescent, persisting in the fruit; pedicels 3 – 9 mm long, sparse to densely pubescent, the indumentum brownish or whitish; bracteoles deltate to broadly so, ovate or lanceolate, 1 – 2 × 1 – 5 mm, pubescent, persisting in the fruit; flower buds globose, 3 × 4 mm, not constricted above ovary; calyx-lobes free, four, ovate, oblong or deltate, orbicular, 2 × 3 mm, pubescent, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruits subglobose or globose, 0.6 – 0.8 × 0.4 – 0.7 cm, pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE76A7E882A1FE00A2ED.taxon	discussion	Note: — Eugenia biflora may be recognized by its pubescent or densely pubescent leaves with whitish indumentum; lateral veins sometimes not visible in dried material or evident only abaxially; revolute leaf margins; a pubescent simple raceme; subglobose or globose flowers buds; and densely pubescent fruits.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE76A7E882A1FE00A2ED.taxon	distribution	Distribution and habitat: — Throughout Tropical America, and widely distributed in central, northeastern and northwestern Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. biflora was found growing in gallery forest edges and secondary forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE76A7E882A1FE00A2ED.taxon	biology_ecology	Phenology: — Flowering all year and fruiting in January, March, November and December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C471EE76A7E882A1FE00A2ED.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Anajatuba, 3 ° 15 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 14 August 2008 (fl.), M. Vidigal 03 a (IAN!); 3 ° 5 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 29 September 2008 (st.), M. Ribeiro 02 (IAN!); Afoga, 3 ° 15 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 18 October 2008 (fl.), M. Vidigal 01 (IAN!); Margem da Estrada Colombo-Anajatuba, 27 January 1976 (fr.), B. G. S. Ribeiro & G. S. Pinheiro 1246 (IAN!); 21 November 1978 (fl.), L. R. Marinho & G. S. Pinheiro 608 (IAN!); São Benedito, 3 ° 15 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 28 June 2008 (fl.), A. Araújo 84 (IAN!). Bacabal, 4 ° 23 ’ 30 ” S, 44 ° 39 ’ 00 ” W, 18 February 1983 (fl.), G. T. Prance 28150 (NY!). Bom Jardim, Posto Indígena Pindaré (Funai Post Guajajara Indians), along Rio Pindaré, c. 15 km W of Santa Inês, 3 ° 30 ’ S, 45 ° 30 ’ W, 01 November 1983 (fl.), M. J. Balick, J. M. Frazão, J. G. A. Vieira & W. S. Silva 1501 (SP, image). Cândido Mendes, 1 ° 26 ’ 49 ” S, 45 ° 43 ’ 00 ” W, 02 November 2017 (fl.), G. S. Amorim 526 (MAR!). Itapecuru Mirim, Fazenda Serra, 3 ° 23 ’ 33 ” S, 44 ° 21 ’ 31 ” W, 24 September 1975 (fl.), D. P. Lima 13398 (PEUFR!). Santa Helena, Próximo do Lugarejo Queimadas, 2 ° 13 ’ 52 ” S, 45 º 17 ’ 59 ” W, 09 July 1978 (fl.), N. A. Rosa & O. Cardoso 2560 (IAN!). Santa Luzia, Alzilândia, Rio Pindaré, 3 ° 45 ’ S, 46 ° 04 ’ 59 ” W, 30 March 1979 (fl., fr.), J. Jangoux 1025 (NY!). São Luís, Alumar, 2 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 08 August 1995 (fl.), K. B. Ferreira s. n (MAR!); Anil, 2 ° 36 ’ S, 44 ° 14 ’ W, January 1940 (fl.), R. L. Fróes 11705 (NY!); Reserva Florestal do Sacavém, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 20 December 1988 (fl.), J. C. Silva Jr. 55 (MAR!); Sítio Santa Eulália, 2 ° 30 ’ 30 ” S, 44 ° 16 ’ 39 ” W, 16 September 2017 (fl., fr.), G. S. Amorim 454, 452 (MAR!); 2 ° 30 ’ 52 ” S, 44 ° 16 ’ 33 ” W, 26 February 2018 (fr.), G. S. Amorim 492 (MAR!); Vinhas, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 08 January 1998 (fr.), A. Salino 3876 (SP, image). São Luiz Gonzaga do Maranhão, 4 ° 19 ’ 00 ” S, 44 ° 40 ’ 00 ” W, 01 October 1980 (fl.), D. C. Daly 404 (INPA, image). São José de Ribamar, Estrada carroçal, 27 December 1997 (fr.), H. C. Nakakura s. n. (EAC!); Sítio Aguahy, 2 ° 38 ’ 48 ” S, 44 ° 09 ’ 06 ” W, 13 October 2015 (fl.), G. S. Amorim 259 (MAR!); 2 ° 38 ’ 48 ” S, 44 ° 09 ’ 06 ” W, 07 May 2016 (fl.), G. S. Amorim 272 (MAR!); 2 ° 39 ’ 00 ” S, 44 ° 07 ’ 55 ” W, 26 August 2017 (fl.), G. S. Amorim 369 (MAR!); 2 ° 38 ’ 47 ” S, 44 ° 09 ’ 05 ” W, 28 October 2017 (fl.), G. S. Amorim 377 (MAR!); 2 ° 38 ’ 47 ” S, 44 ° 09 ’ 05 ” W, 20 February 2018 (fl.), G. S. Amorim 467 (MAR!). Serrano do Maranhão, Porto do Pindobal, 1 ° 38 ’ 32 ” S, 44 ° 52 ’ 17 ” W, 19 August 2014 (fl.), G. P. Lima 495 (MAR!). Viana, between Vitoria do Mearim and Viana, 3 ° 15 ’ S, 44 ° 55 ’ 00 ” W, 16 January 1980 (fl.), D. C. Daly & G. G. Campbell 628 (IAN!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C470EE76A7E885C1FBB0A61D.taxon	materials_examined	Type: — BRAZIL, Pará, Almeirim, Gleba Monte Dourado, floresta secundária de terra firme, J. M. Pires & N. T. Silva 2015 (holotype: MG). (Fig. 2 E-F.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C470EE76A7E885C1FBB0A61D.taxon	description	Shrubs 2 – 17 m. Twigs glabrous, sparsely pubescent when young, the indumentum whitish. Leaves with petioles 3 – 8 mm long, glabrous or puberulent, channeled or shallowly channeled; blades elliptic to broadly so, lanceolate, ovate or oblong, 5.2 – 14 × 2.5 – 6.3 cm, glabrous or sparsely pubescent, the indumentum brownish, chartaceous, concolorous; glands conspicuous; base rounded, cuneate or acute; apex acute or attenuate, rarely acuminate; midvein adaxially sulcate; lateral veins visible, not arched, 8 to 10 pairs; intramarginal vein 1 – 3 from the margin, margin revolute. Inflorescence axillary or terminal, raceme simple, the axis 4.2 – 10.4 cm long; bracts not seen, caducous before anthesis; pedicels 2 – 9 mm long, glabrous; bracteoles ovate to broadly so, or deltate, 1 – 2 × 1 – 3 mm, glabrous, persistent in the fruit; flower buds globose, 5 × 5 mm, not constricted above ovary; calyx-lobes free, four, orbicular, 2 – 3.5 × 3 – 4 mm, glabrous, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruits oblong or ellipsoid, 1.4 – 1.7 × 1.1 – 1.3 cm, glabrous or puberulent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C470EE76A7E885C1FBB0A61D.taxon	discussion	Note: — Eugenia caducibracteata may be recognized by its adaxially sulcate leaf midvein; a simple raceme with a long axis; large flower buds (5 × 5 mm); and bracts that are caducous before anthesis.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C470EE76A7E885C1FBB0A61D.taxon	distribution	Distribution and habitat: — Known from northeastern and northwestern Brazil (Amazonas, Maranhão and Pará states) (Mazine & Souza 2009). E. caducibracteata can be found growing in unflooded tropical moist forest (Mazine & Souza 2009) and in Amazonian Maranhão was found growing in secondary forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C470EE76A7E885C1FBB0A61D.taxon	biology_ecology	Phenology: — Flowering in February and December and fruiting in August.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C470EE76A7E885C1FBB0A61D.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Açailândia, Estrada BR- 222, entre Santa Inês e Açailândia, 300 m, 4 ° 45 ’ S, 45 ° 10 ’ W, 17 December 1978 (fl.), J. Jangoux 559 (NY!). Alto Alegre do Pindaré, Alzilândia, Rio Pindaré, 0 - 100 m, 3 ° 45 ’ S, 46 ° 05 ’ W, 11 December 1978 (fl.), J. Jangoux & R. P. Bahia 304 (BHCB, image). Buriticupu, Reserva Florestal da CVRD, 46 ° 24 ’ 04 ” S, 46 ° 24 ’ 04 ” W, 08 February 1996 (fl.), F. H. Muniz B 2378 (SP, image). Santa Luzia, Alzilândia, Rio Pindaré, 3 ° 45 ’ S, 46 ° 05 ’ W, 11 December 1978 (fr.), J. Jangoux 304 (NY!). São Luís, February-March 1939 [fl], R. L. Fróes 11773 (NY!). São José de Ribamar, Sítio Aguahy, 2 ° 38 ’ 47 ” S, 44 ° 09 ’ 05 ” W, 03 February 2018 (fl.), G. S. Amorim 448 (MAR!); 03 February 2018 (fl.), G. S. Amorim 447 (MAR!); 26 August 2017 (fr.), G. S. Amorim 373 (MAR!). Turiaçu, Palmeirinha, 27 February 1983 (fl.), J. U. Santos 994 (MG!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE79A7E886E9FCA4A559.taxon	materials_examined	Type: — SURINAME. Brokopondo, Brownsberg, C. J. Zaandam 6428 (holotype: U [0005014]).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE79A7E886E9FCA4A559.taxon	description	Trees or shrubs 5.4 – 10 m. Twigs glabrous. Leaves with petioles 8 – 22 mm long, glabrous, unchanneled; blades elliptic to broadly so, lanceolate, ovate or oblong, 12.9 – 22.4 × 5 – 10 cm, glabrous adaxially and glabrous or sparsely pubescent abaxially, the indumentum brownish-red, chartaceous, discolorous, sometimes lustrous adaxially, darker abaxially; glands not seen; base cuneate, rounded or obtuse; apex acute, cuneate, obtuse or cuspidate; midvein adaxially flat; lateral veins prominent abaxially, not arched, 12 to 15 pairs; intra-marginal vein 1 – 6 mm from the margin, margin flat. Inflorescence axillary, raceme simple, the axis 2.7 – 4.8 cm, pubescent, the indumentum brownish-red, bracts not seen, caducous before anthesis; pedicels 5 – 6 mm long, pubescent; bracteoles deltate, 1.5 – 2 × 1 – 2 mm, pubescent, persistent in the fruit; flower buds globose, 4 – 7 × 5 mm, not constricted above ovary; calyx-lobes free, four, orbicular, 1.5 – 2 × 1.8 – 3 mm, pubescent, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruits globose, 0.5 – 0.8 × 0.6 – 0.9 cm, glabrous or sparsely pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE79A7E886E9FCA4A559.taxon	discussion	Note: — Eugenia cupulata may be recognized by its adaxially flat midvein; intra-marginal vein 1 – 6 mm from the margin; a simple raceme with brownish-red pubescence; globose flower buds; and free, pubescent calyx-lobes.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE79A7E886E9FCA4A559.taxon	distribution	Distribution and habitat: — Known from Bolivia, French Guiana, Suriname, Venezuela, central, northeastern and northwestern Brazil (Acre, Amazonas, Maranhão, Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. cupulata was found growing in “ terra firme ” forest on rocky soil.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE79A7E886E9FCA4A559.taxon	biology_ecology	Phenology: — Flowering in February and March and fruiting in April and June.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE79A7E886E9FCA4A559.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: São Luís, 2 ° 34 ’ 38 ” S, 44 ° 12 ’ 04 ” W, February-March 1939 (fl.), R. L. Fróes 11789 (NY!); 1940 (fr.), R. L. Froes 11864 (NY!). Arredores de Estiva, 08 August 1980 (st.), M. G. Silva 5738 (MG!). Additional specimens examined: — BRAZIL. Maranhão: Carolina, 7 ° 19 ’ 58 ” S, 47 ° 28 ’ 10 ” W, 12 March 2016 (fr.), A. C. Sevilha 5811 (CEN, image). Lorêto, “ Ilha das Balsas ”, 140 m, 7 ° 30 ’ S, 45 ° 3 ’ W, 01 June 1962 (fr.), G. Eiten 4810 (NY!). Pará: Santarém, Km 35 da estrada do Palhao, arredores do Acampamento do Igarapé Curupira, 04 September 1969 (fl.), M. B. Silva 2493 with R. Souza (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE78A7E8821DFB00A009.taxon	materials_examined	Type: — BRAZIL. Mato Grosso. Chapada dos Guimarães, Parque Nacional da Chapada dos Guimarães, Véu da Noiva, ao longo do rio Coxipozinho, trilha para a base da cachoeira, F. F. Mazine et al. 1072 (holotype: ESA; isotype: K [001018882], image). (Fig. 2 G)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE78A7E8821DFB00A009.taxon	description	Shrubs 3 – 15 m. Twigs densely pubescent, the indumentum whitish. Leaves with petioles 2 – 13 mm long, glabrous, channeled; blades oblanceolate, oblong, obovate or elliptic, 2.5 – 20 × 1.8 – 8.5 cm, glabrous, chartaceous or subcoriaceous, discolorous; glands conspicuous; base cuneate, acute or obtuse; apex acute, acuminate or cuspidate; midvein adaxially flat; lateral veins visible, not arched, 8 to 13 pairs; intra-marginal vein 1 – 3 mm from the margin, margin revolute. Inflorescence axillary, raceme simple, the axis 1.8 – 6 cm long, sparsely pubescent or densely pubescent, the indumentum whitish; bracts ovate to broadly so, or deltate, 0.1 – 0.2 × 0.3 mm, pubescent, persisting in the fruit; pedicels 2 – 10 mm long, pubescent; bracteoles ovate or deltate, 0.2 × 0.5 mm, glabrescent or densely pubescent; persisting in the fruit; flowers buds globose, 1 – 2 × 2 mm, not constricted above ovary; calyx-lobes partially fused in the lower third of the bud, four, orbicular, 0.3 – 2 × 0.5 – 1.9 mm, glabrescent or densely pubescent, not reflexed, persistent in the fruits; ovary 2 - 3 - locular. Immature fruits globose, 0.6 – 1.1 × 0.5 – 1.7 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE78A7E8821DFB00A009.taxon	discussion	Note: — Eugenia densiracemosa may be recognized by its chartaceous or subcoriaceous leaves; with intra-marginal vein 1 – 3 mm from the margin; long simple racemes (1.8 – 6 cm), generally with a long pedicel (2 – 10 mm long). This species is morphologically similar to E. florida, but can be distinguished by its 8 to 13 pairs of unarched lateral veins (vs. arched) that are visible adaxially and abaxially (5 to 8 pairs, visible abaxially).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE78A7E8821DFB00A009.taxon	distribution	Distribution and habitat: — Known from French Guiana and central, northeastern and northwestern Brazil (Acre, Ceará, Goiás, Maranhão, Mato Grosso, Pará and Tocantins states) (POWO 2020, Flora do Brasil 2020). E. densiracemosa grows in forest formations along rivers and streams borders, in Cerrado and Amazonia biomes (Mazine & Faria 2013). In Amazonian Maranhão it was found in “ terra firme ” and secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE78A7E8821DFB00A009.taxon	biology_ecology	Phenology: — Flowering in January, August and October to December and fruiting in March and August.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47FEE78A7E8821DFB00A009.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Alto Alegre do Pindaré: Mineirinho, 0 - 100 m, 3 ° 40 ’ S, 45 º 50 ’ W, 26 May 1979 (fr.), J. Jangoux & R. P. Bahia 875 (NY!). Santa Luzia, margem direita da Rod. BR- 222, a altura do km 124, que liga Santa Inês a Açailândia na Belém-Brasília, 3 ° 57 ’ 48 ” S, 45 ° 39 ’ 29 ” W, 16 December 1978 (fl.), N. A. Rosa 3014 (NY!); Mineirinho, 3 ° 40 ’ S, 45 ° 50 ’ W, 26 March 1979 (fr.), J. Jangoux 875 (NY!). São Luís, Estrada do Tirirical, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, February-March 1989 (fr.), R. L. Fróes 11534 (NY!); Granja Barreto, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 04 January 1950 (fl.), R. L. Fróes 25619 (IAN!); Estrada da Maioba, 10 January 1950 (fl.), R. L. Fróes 25654 (IAN!); Reserva Florestal do Sacavém, 2 ° 30 ’ S, 44 ° 15 ’ W, 27 February 1992 (fl.), F. H. Muniz 36 (INPA, image); 15 September 1992 (fr.), F. H. Muniz 159 (INPA, image); 07 October 1992 (fl.), F. H. Muniz 168 (RB, image); 10 February 1993 (fl.), F. H. Muniz 216 (RB, image). São José de Ribarmar, Sítio Aguahy, 2 ° 38 ’ 34 ” S, 44 ° 08 ’ 19 ” W, 25 August 2017 (fr., fl.), G. S. Amorim 359 (MAR!); 28 October 2017 (fl.), G. S. Amorim 379 (MAR!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47EEE7BA7E887ECFC1BA279.taxon	materials_examined	Type: — BRAZIL. “ Habitat in silvis ad villam Ega in prov. do alto Amazonas ”, Pöppig 3109 (syntypes: P [01902445], image, G [00223425], image, BM [000953773], image). (Fig 3 AB)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47EEE7BA7E887ECFC1BA279.taxon	description	Shrubs ca. 1.5 m. Twigs glabrous. Leaves with petioles 9 – 10 mm long, glabrous, unchanneled or shallowly channeled; blades elliptic, oblong, or oval, 10 – 12.1 × 3.6 – 5.1 cm, glabrous, chartaceous, discolorous; glands not seen; base cuneate or rounded; apex acute, cuneate, rarely rounded; midvein adaxially flat; lateral veins visible, not arched, 10 to 15 pairs; intra-marginal vein 1 – 3 mm from the margin, margin revolute. Inflorescence axillary, a fascicle, the axis 1 – 2 mm, glabrescent; bracts orbicular, 0.5 × 0.5 mm, glabrous, persistent in the fruit; pedicels 4 – 9 mm long, glabrous; bracteoles ovate, 1.5 × 1 mm, glabrous, persistent in the fruit; flower buds globose, 3 – 4 × 5 – 6 mm, not constricted above ovary; calyx-lobes free, four, orbicular, 1.4 – 2 × 1 – 2 mm, glabrous, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, ca. 1 cm diameter, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47EEE7BA7E887ECFC1BA279.taxon	discussion	Note: — E. dittocrepis may be recognized by its leaves, which are green-yellowish adaxially and reddish-brown abaxially in the dried material, with revolute margins; its inflorescence, which is a glabrescent fascicle with persistent bracteoles in fruit; and its globose, glabrous, and smooth fruit. This species is morphologically related to E. stictopetala but can be distinguished by its revolute leaf margins (vs. flat) and its discolorous leaves (vs. concolorous leaves).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47EEE7BA7E887ECFC1BA279.taxon	distribution	Distribution and habitat: — Known from Colombia, Ecuador, Peru, Venezuela and northeastern and northwestern Brazil (Acre, Amazonas, Maranhão, Mato Grosso and Pará states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. dittocrepis was found growing along waterways.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47EEE7BA7E887ECFC1BA279.taxon	biology_ecology	Phenology: — Fruiting in April.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47EEE7BA7E887ECFC1BA279.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Cândido Mendes, Maracassumé River Region, 30 April 1932 (fr.), R. L. Fróes 1709 (NY!). Viana, 1958 (st.), O. Carvalho 5 (RB, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47DEE7BA7E8853DFB65A6A5.taxon	materials_examined	Type: — BRAZIL. in Brasiliae deserto Bahiensi, C. F. P. Martius s. n. (holotype: M [0137665]; isotype: M [0137666] M [0137667]). (Fig 2 H)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47DEE7BA7E8853DFB65A6A5.taxon	description	Shrubs 0.35 – 15 m. Twigs glabrous or sparsely pubescent, the indumentum whitish. Leaves with petioles 2 – 7 mm long, glabrous, channeled; blades elliptic to broadly so, or oblong, 4.3 – 7 × 1.3 – 3.4 cm, glabrous, chartaceous or coriaceous, concolorous; glands conspicuous; base cuneate or rounded; apex acute, acuminate or long acuminate; midvein adaxially flat to prominent; lateral veins visible and slightly elevated, not arched, 6 to 12 pairs; intra-marginal vein 1 – 2 mm from the margin, margin flat. Inflorescence cauliflorous, a fascicle, the axis absent or 1 – 1.5 mm long, glabrous; bracts orbicular, 0.7 × 1.5 mm, glabrous, persistent in the fruit; pedicels 2 – 5 mm long, glabrous; bracteoles orbicular, 0.5 × 0.7 mm, glabrous, persistent in the fruit; flower buds globose, 3 × 4 mm, not constricted above ovary; calyx-lobes free, four, deltate or orbicular, 0.5 – 2 × 1 – 1.5 mm, glabrous, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit globose, 0.4 – 0.6 × 0.2 – 0.9 cm, glabrous or sparsely pubescent, surface marked with glands.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47DEE7BA7E8853DFB65A6A5.taxon	discussion	Note: — Eugenia flavescens may be recognized by its leaves, yellowish in the dry material, with an adaxially flat to prominent midvein and slightly elevated lateral veins; its inflorescence, which is a cauliflorous fascicle with an axis that is absent or up to 1 – 1.5 mm long, turning yellow in dry material; its small flowers (3 × 4 mm); and its globose fruits, which are glabrous or sparsely pubescent with marked glands.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47DEE7BA7E8853DFB65A6A5.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Suriname, Venezuela and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. flavescens was found growing in “ terra firme ” forest, on rocky soil, and in old clearings.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47DEE7BA7E8853DFB65A6A5.taxon	biology_ecology	Phenology: — Flowering from January to April and June and fruiting in July.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47DEE7BA7E8853DFB65A6A5.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Estrada BR 222, entre Santa Inês e Açailândia, 4 ° 45 ’ S, 47 ° 10 ’ W, 17 December 1978 (fr.), J. Jangoux 543 (NY!). Barão de Grajaú, 6 ° 36 ’ 29 ” S, 43 ° 24 ’ 17 ” W, 25 January 2012 (fr.), R. M. Harley 56478 (HUEFS, image). São Luís, 8 º Batalhão da Policia Militar, 2 ° 29 ’ 49 ” S, 44 ° 16 ’ 43 ” W, 15 June 2018 (fr.), G. S. Amorim 510 (MAR!); Estrada do Olho d’Agua, February – March 1939 (st.), R. L. Fróes 11624 (NY!); Estrada de S. Luís para o Rio Anil, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 07 January 1950 (fl.), R. L. Fróes 25635 (IAN!); Granja Barreto, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 04 January 1950 (fl.), R. L. Fróes 25621 (IAN!). São Bento, contagem em campo de São Bento, 02 July 1979 (fr.), N. A. Rosa & O. C. Nascimento 2581. Turiaçu, Ilha Trauira, 1 ° 39 ’ 48 ” S, 45 º 22 ’ 18 ” W, 05 April 1987 (fl.), M. R. Santos 700 (IAN!). Additional specimens examined: — BRAZIL. Maranhão: Barão de Grajaú: ca. 50 Km da cidade na BR 230, 6 ° 36 ’ 29 ” S, 43 ° 24 ’ 17 ” W, 21 January 2012 (fr.), R. M. Harley 56478 (NY!). Carolina, Fazenda Palestina, 7 ° 22 ’ 56 ” S, 47 ° 28 ’ 40 ” W, 01 March 2015 (fr.), G. Pereira-Silva 9746 (CEN, image). Estreito, margem direita do rio Tocantins / foz do rio Feio, 6 ° 44 ’ 15 ” S, 47 ° 29 ’ 26 ” W, 14 January 2008 (fr.), G. Pereira-Silva 12608 (CEN, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E886E9FA1CA523.taxon	materials_examined	Type: — BRAZIL. Amazonas, “ in Brasilia ad ripam flum. Amazonum ”, C. F. P. Martius s. n. (holotype: M [0137662]; isotype: M [0137663]).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E886E9FA1CA523.taxon	description	Shrubs 1 – 2.5 m. Twigs sparsely pubescent, the indumentum whitish. Leaves with petioles 4 – 10 mm long, pubescent, channeled; blades elliptic to broadly so, or oblanceolate, 11 – 15.3 × 4.3 – 5.1 cm, glabrous, membranaceous or subcoriaceous, concolorous; glands not seen; base cuneate or rounded; apex acute, acuminate, or attenuate; midvein adaxially sulcate; lateral veins visible abaxially, arched, 5 to 8 pairs; intra-marginal vein ca. 1 – 3 mm or less from the margin, margin flat. Inflorescence axillary, raceme simple, the axis 1.9 – 4.1 cm long, glabrescent; bracts lanceolate, 4 – 6 × 0.5 – 1 mm, persistent in the fruit, glabrous; pedicels 2 – 4 mm long, sparsely pubescent; bracteoles deltate, 2 – 6 × 0.5 – 2 mm, persistent in the fruit, glabrous; flower buds globose, 2 × 3.5 mm, not constricted above ovary; calyx-lobes free, four, ovate, ca. 2 mm, glabrous, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, 2.7 – 3 × 2 – 2.8 cm, pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E886E9FA1CA523.taxon	discussion	Note: — Eugenia florida may be recognized by its leaves with arched lateral veins and the first vein not confluent with the marginal vein; its glabrescent simple raceme; its globose fruits, which are pubescent and have a smooth surface. This species is morphologically related to E. densiracemosa but can be distinguished by its concolorous leaves (vs. discolorous), and its arched lateral veins (vs. not arched), with 5 to 8 pairs, visible abaxially (vs. 8 to 13 pairs, visible adaxially and abaxially).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E886E9FA1CA523.taxon	distribution	Distribution and habitat: — Known from Argentina, Bolivia, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Nicaragua, Panamá, Paraguay, Peru, Suriname, Venezuela, and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. florida was found growing in “ terra firme ” forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E886E9FA1CA523.taxon	biology_ecology	Phenology: — Flowering in January and November and fruiting in March.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E886E9FA1CA523.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Santa Luzia, Fazenda Cacique, 3 ° 49 ’ 59 ” S, 46 ° 04 ’ 00 ” W, 25 March 1983 (fr.), E. L. Taylor E 1058 et al. (MG!). Additional specimens examined: — BRAZIL. Maranhão: Mirador, lugarejo Boné, 6 ° 23 ’ 33 ” S, 44 ° 28 ’ 37 ” W, 16 November 1988 (fl.), F. Noberto 11 (SLUI!). São Pedro dos Crentes, margem direita do rio Farinha, próximo à cidade, 6 ° 50 ’ 03 ” S, 46 ° 32 ’ 04 ” W, 11 January 2008 (fr.), G. Pereira-Silva 12506 (CEN, image). Pará: Peixe-Boi, colônia do Salgado, arredores do igarapé do Ubim, 1 ° 11 ’ 38 ” S, 47 ° 19 ’ 03 ” W, 22 January 1977 (fl.), M. G. da Silva 2854 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E88219FE20A94C.taxon	materials_examined	Type: — “ in ins. Caribaeis, S. Vincentii ” [and the Grenadines]. Lambert s. n. (B as F negative 33490). (Fig. 3 C-D.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E88219FE20A94C.taxon	description	Trees or shrubs 4 m. Twigs glabrous. Leaves with petioles 5 – 10 mm long, glabrous, channeled; blades elliptic or lanceolate, rarely ovate, 4.1 – 12.2 × 1.3 – 5 cm, puberulent adaxially and glabrous abaxially, the indumentum whitish, membranaceous, discolorous; glands not seen; base attenuate, cuneate or rounded; apex acute or acuminate; midvein adaxially sulcate; lateral veins visible, not arched, 8 to 20 pairs; intra-marginal vein 1 – 2 mm or less from the margin, margin revolute. Inflorescence axillary, a fascicle, the axis 2 – 7 mm long, sparsely pubescent, the indumentum brownishred; bracts deltate, 0.5 × 0.4 mm, caducous after anthesis, glabrous; pedicels 2 – 5.9 mm long, glabrous; bracteoles deltate, ca. 1 mm, glabrous, persistent in the fruit; flower buds globose 2 – 3 × 3 mm, not constricted above ovary; calyx-lobes free, four, orbicular, 1.5 × 2 mm, glabrous, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, 0.8 – 1 × 0.7 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E88219FE20A94C.taxon	discussion	Note: — Eugenia lambertiana may be recognized by its elliptic or lanceolate, rarely ovate leaves, which are discolorous, brown adaxially and green abaxially; and the intra-marginal vein ca. 1 – 2 mm or less from the margin. This species is morphologically related to E. protenta, but can be distinguished by its leaves, which are puberulent adaxially and glabrous abaxially (vs. glabrous), the revolute margin (vs. flat), and the glabrous fruit (vs. pubescent).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E88219FE20A94C.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Leeward Is., Peru, Suriname, Trinidad-Tobago, Venezuela, Windward Is. and widely distributed in central, northeastern and northwestern Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. lambertiana was found growing in open field.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E88219FE20A94C.taxon	biology_ecology	Phenology: — Flowering in November and fruiting in March.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47CEE7AA7E88219FE20A94C.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Maracaçumé River Region, 08 September 1932 (fl.), R. L. Fróes 1872 (NY!). Additional specimens examined: — BRAZIL. Amapá: Serra do Navio, Rio Amapari, 0 ° 52 ’ 52 ” N, 52 ° 01 ’ 03 ” W, 23 November 1954 (fr.), R. S. Cowan 38547 (NY!). Pará: Oriximiná. Rio Trombetas, margem esquerda do lago Erepecu, ao longo do lago, 1 ° 20 ’ 31 ” S, 56 ° 34 ’ 43 ” W, 16 July 1980 (fr.), C. A. Cid Ferreira 1522 with J. Ramos, C. D. Mota & M. Rosas (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E886E9FB1FA291.taxon	materials_examined	Type: — FRENCH GUIANA. Cayenne, Leblond s. n. (P [01902639] image). (Fig. 2 I, 3 E-H)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E886E9FB1FA291.taxon	description	Trees or shrubs 4 m. Twigs sparsely pubescent, the indumentum reddish-brown. Leaves with petioles 6 – 9 mm long, pubescent, channeled; blades elliptic to broadly so, or oblong, 10.5 – 16.2 × 3.8 – 5.3 cm, glabrous or sparsely pubescent, the indumentum whitish, chartaceous, discolorous; glands conspicuous; base rounded, attenuate or cuneate; apex acute or acuminate; midvein adaxially sulcate; lateral veins visible adaxially, not arched, 19 to 22 pairs; intra-marginal vein ca. 1 mm from the margin, margin flat. Inflorescence axillary, raceme simple, the axis 9.8 – 12.6 cm long, pubescent; bracts not seen, caducous before anthesis; pedicels 5 – 15 mm long, pubescent; bracteoles deltate, 8 × 5 mm, pubescent adaxially and glabrous abaxially, caducous after anthesis; flower buds globose ca. 3 mm, not constricted above ovary; calyx-lobes free, four, orbicular or oblong, 2 – 3 × 3 – 4 mm, pubescent, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, 1 – 1.3 × 1.3 – 1.5 cm, pubescent, surface muricate.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E886E9FB1FA291.taxon	discussion	Note: — Eugenia patens may be recognized by its discolorous, elliptic to broadly elliptic, or oblong leaves, with an adaxially sulcate midvein; and the globose fruit that are pubescent and have a ridged surface. This species is morphologically similar to E. polystachya from which it is distinguished by its fruit with muricate surface (vs. smooth) and glabrous or sparsely pubescent leaves (vs. densely pubescent).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E886E9FB1FA291.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname and northeastern and northwestern Brazil (Acre, Amazonas, Amapá, Maranhão, Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. patens was found growing in “ terra firme ” forests and secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E886E9FB1FA291.taxon	biology_ecology	Phenology: — Flowering and fruiting in October.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E886E9FB1FA291.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Bequimão, Pontal, 2 ° 58 ’ 54 ” S, 45 ° 19 ’ 09 ” W, 12 January 2020 (fr.), G. S. Amorim 601 (JPB!); Monção: Ka’apor Indian Reserve, 3 ° 09 ’ 58 ” S, 45 ° 56 ’ 17 ” W, 07 October 1986 (fl.), W. Balée 2684 (NY!). Mineirinho, Rio Pindaré, 26 October 1979 (fl., fr), J. Jangoux 937 (MG!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E88255FF69A949.taxon	materials_examined	Type: — FRENCH GUIANA, s. l., J. P. B. Rohr, s. n. (holotype: C [10015707]). (Fig 3 I-M)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E88255FF69A949.taxon	description	Trees 2.5 – 8 m. Twigs pubescent, the indumentum reddish-brown. Leaves with petioles 1 – 7 mm long, pubescent, channeled or shallowly channeled; blades elliptic to broadly so, obovate or ovate, 6.2 – 14.4 × 3.1 – 6 cm, glabrous, pubescent on young blades, the indumentum reddish-brow, membranaceous or chartaceous, concolorous; glands conspicuous; base cuneate or abruptly rounded; apex acute or cuneate, rarely cuspidate; midvein adaxially flat to impressed; lateral veins slightly visible, arched, 5 to 9 pairs; intra-marginal vein 3 – 6 mm or less from the margin, margin flat. Inflorescence axillary, a auxotelic raceme, with cataphylls at base, the axis 0.6 – 1.3 cm long, pubescent, the indumentum reddish-brown; bracts deltate, 3 – 3.5 × 1 – 1.6 mm, glabrous adaxially and pubescent abaxially, caducous after anthesis; pedicels 15 – 30 mm long, glabrous or pubescent; bracteoles linear, 1.5 × 1.5 mm, pubescent, caducous after anthesis; flower buds globose 2 – 3 × 5 mm, not constricted above ovary; calyx-lobes free, four, orbicular or oblong, 2 × 1.5 mm, pubescent, not reflexed, persistent in the fruit; ovary not seen. Immature fruits globose, ca. 2 cm diameter, glabrous, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E88255FF69A949.taxon	discussion	Note: — Eugenia patrisii may be recognized by its leaves with arched lateral veins, intra-marginal vein 1 – 6 mm from the margin; its pubescent auxotelic raceme with cataphylls at the base; long-pedicellate flowers; and its globose fruit, which are glabrous and have marked glands on the surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E88255FF69A949.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela and widely distributed in northeastern and northwestern Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. patrisii was found growing in flooded forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E88255FF69A949.taxon	biology_ecology	Phenology: — Flowering in September and fruiting in November and December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47BEE7DA7E88255FF69A949.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Alzilândia, Rio Pindaré, 11 December 1978 (fr.), J. Jangoux & R. P. Bahia 368 (MG!); Bacabal, Fazenda São Francisco, 4 ° 00 ’ 00 ’’ S, 44 ° 55 ’ 48 ” W, 25 September 1980 [fl], D. C. Daly D 257 (NY!). Carutapera, Gurupiuna, Ka’apor Indian Reserve, 2 ° 43 ’ 12 ” S, 46 ° 25 ’ 48 ” W, 05 November 1986 (st.), W. L. Balée 2904 (NY!). Monção, Bacia do Rio Turiaçu, 22 September 1985 (fl.), W. L. Balée 1047 (MG!); 3 ° 07 ’ 12 ” S, 46 ° 04 ’ 48 ” W, 31 May 1987 (st.), W. L. Balée 3312 (NY!); Catutapera, Gurupiuna, Ka’apor Indian Reserve, 3 ° 30 ’ S, 45 ° 15 ’ W, 09 May 1986 (st.), W. L. Balée 2293 (NY!); 2 ° 43 ’ 12 ” S, 46 W 25 ’ 48 ” W, 05 November 1986 (fr.), W. L. Balée 2904 (MG!). São Luís, Alumar, 2 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 10 November 1995 (fr.), K. B. Ferreira & E. C. Girnos s. n (MAR!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7CA7E886E9FED0A5A9.taxon	materials_examined	Type: — FRENCH GUIANA. Cayenne, Leblond s. n. (P) (Fig 2 J, 4 A-D)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7CA7E886E9FED0A5A9.taxon	description	Trees 3 – 5.4 m. Twigs densely pubescent, the indumentum whitish. Leaves with petioles 4 – 6 mm long, pubescent, channeled; blades elliptic to broadly so, 4.9 – 7.5 × 1.5 – 5 cm, densely pubescent, the indumentum whitish, chartaceous, discolorous; glands not seen; base cuneate or rounded; apex acute or acuminate, rarely cuspidate; midvein adaxially impressed; lateral veins visible, not arched, 31 to 35 pairs; intra-marginal vein 1 – 2 mm from the margin, margin flat. Inflorescence terminal, raceme simple, the axis 10 – 16 cm long, pubescent, the indumentum whitish; bracts lanceolate, 1 – 2 × 2 mm, densely pubescent, persistent in the fruit; pedicels 6 – 13 mm long, pubescent; bracteoles deltate or orbicular, 2 – 3 × 1 mm, pubescent, persistent in the fruit; flower buds globose, 2 – 3 × 2 mm, not constricted above ovary; calyx-lobes free, four, orbicular or oblong, 2 × 3 mm, pubescent, not reflexed, persistent in the fruit; ovary 2 – locular. Immature fruits globose, 4 – 7 × 3 – 5 cm, pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7CA7E886E9FED0A5A9.taxon	discussion	Note: — Eugenia polystachya may be recognized by its densely pubescent, elliptic to broadly elliptic leaves; its globose buds; and its globose fruit, which are pubescent and have a smooth surface. This species is morphologically similar to E. patens from which it is distinguished by its fruit with a smooth surface (vs. muricate surface) and its densely pubescent leaves (vs. glabrescent or sparsely pubescent).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7CA7E886E9FED0A5A9.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Peru and northeastern and northwestern Brazil (Acre, Amazonas, Amapá, Maranhão Pará and Roraima states) (POWO 2020, Flora do Brasil). In Amazonian Maranhão, E. polystachya was found growing in “ terra firme ” forest and flooded forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7CA7E886E9FED0A5A9.taxon	biology_ecology	Phenology: — Flowering from October to December and fruiting in January, May, October and November.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7CA7E886E9FED0A5A9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Maracassumé River Region, 06 May 1932 (fr.), R. L. Fróes 1737 (NY!). Anajatuba, 3 ° 15 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 24 January 1976 (fr.), B. G. S. Ribeiro & G. S. Pinheiro 1198 (IAN!); 26 January 1976 (fr.), B. G. S. Ribeiro & G. S. Pinheiro 1215 (IAN!); Estrada de Anajatuba, 21 November 1978 (fl.), L. R. Marinho & G. S. Pinheiro 611 (IAN!). Santa Inês, Margem direita do Rio Pindaré, 3 ° 40 ’ 01 ” S, 45 ° 22 ’ 48 ” W, 13 December 1978 (fl.), N. A. Rosa 2953 (NY!). São Luís, Estrada do Sacavem, February-March- 1939 (fl.), R. L. Fróes 11565 (NY!). São Vicente Ferrer, Banderante N to Pinheiro, 3 ° 00 ’ 00 ” S, 45 ° 10 ’ 12 ” W, 17 October 1980 (fl.), D. C. Daly D 671 (NY!). Vitória do Mearim, Rio Mearim, 3 ° 27 ’ 44 ” S, 44 ° 52 ’ 14 ” W, 16 January 1976 (fr.), N. T. Silva 4201 (IAN!). Turiaçu, km 6 da BR 106 Maracaçumá-Sta. Helena, 1 ° 39 ’ 48 ” S, 45 ° 22 ’ 18 ” W, 01 December 1978 (fl.), N. A. Rosa 2803 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7FA7E8828DFD5EA0E9.taxon	materials_examined	Type: — VENEZUELA. “ Prope San Carlos, ad Rio Negro, Brasiliae borealis ”, R. Spruce 3816 (holotype: BR! [5260492], image). (Fig 2 K)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7FA7E8828DFD5EA0E9.taxon	description	Trees 4 m. Twigs glabrous. Leaves with petioles 3 – 6 mm long, glabrous or sparsely pubescent, channeled or shallowly channeled; blades elliptic to broadly so, 4.9 – 11.5 × 1.9 – 2.4 cm, glabrous, chartaceous, discolorous; glands conspicuous; base rounded or cuneate; apex acute, acuminate or long acuminate; midvein adaxially sulcate; lateral veins visible adaxially, not arched, 10 to 13 pairs; intra-marginal vein ca. 1 mm from the margin, margin flat. Inflorescence axillary, a fascicle, the axis 2 – 4 mm long, pubescent, the indumentum reddish-brown; bracts orbicular, 2 × 5 mm, pubescent adaxially and glabrous abaxially, caducous after anthesis; pedicels 5 – 10 mm long, glabrous or sparsely pubescent; bracteoles orbicular, 0.5 × 1.2 mm, pubescent adaxially and glabrous abaxially, persistent in the fruits; flower buds globose, 3 × 2 mm, not constricted above ovary; calyx-lobes free, four, obovate, 1.2 – 2.7 × 1.2 – 3 mm, glabrous or pubescent, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, 0.6 – 1 × 0.6 – 0.8 cm, pubescent, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7FA7E8828DFD5EA0E9.taxon	discussion	Note: — Eugenia protenta may be recognized by its elliptic to broadly elliptic leaves, the acute, acuminate or long acuminate leaf apex; and the globose fruit, which are pubescent and with marked glands on the surface. This species is morphologically related to E. lambertiana but can be distinguished by its glabrous leaves (vs. puberulent adaxially), flat margin (vs. revolute) and the pubescent fruit (vs. glabrous).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7FA7E8828DFD5EA0E9.taxon	distribution	Distribution and habitat: — Known from Venezuela and northeastern and northwestern Brazil (Acre, Amazonas, Maranhão, Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. protenta was found growing in “ terra firme’ ’ forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7FA7E8828DFD5EA0E9.taxon	biology_ecology	Phenology: — Fruiting was registered in March.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C47AEE7FA7E8828DFD5EA0E9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: São José de Ribamar, Sítio Aguahy, 2 ° 38 ’ 34 ” S, 44 ° 08 ’ 19 ” W, 25 August 2017 (fl.), G. S. Amorim 362 (MAR!). São Luís, Anil, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 14 March 1949 (fr.), R. L. Fróes 24270 (IAN!). Additional specimens examined: — BRAZIL. Amazonas: Cuieiras Campina, Rio Cuieiras near mouth of Rio Branchinho, 2 ° 32 ’ 44 ” S, 60 ° 19 ’ 10 ” W, 2 July 1975 (fl.), A. B. Anderson 145 (US, image); Presidente Figueiredo: Represa de Balbina on rio Uatumã, ca. 4 km NW of dam on D- 1 road, 1 ° 49 ’ 59 ” S, 59 ° 31 ’ 59 ” W, 4 July 1986 (fr.), W. W. Thomas 5342, with C. A. Cid & E. Fiuza Lima (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE7FA7E887C4FDEDA701.taxon	materials_examined	Type: — VENEZUELA. Monte Impossible, F. W. H. Humboldt & A. Bonpland 273 (holotype: P [00679193], image). (Fig 2 L-M.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE7FA7E887C4FDEDA701.taxon	description	Shrubs 0.9 – 2.5 m. Twigs pubescent, the indumentum whitish. Leaves with petioles 3 mm long, glabrous or sparsely pubescent, shallowly channeled; blades elliptic to broadly so, obovate or oblong, 4.7 – 8.9 × 2.2 – 4.5 cm, glabrous, chartaceous, concolorous; glands conspicuous; base cuneate or rounded; apex acute, acuminate or cuneate; midvein adaxially impressed; lateral veins visible, not arched, 6 to 12 pairs; intra-marginal vein 1 – 3 mm or less from the margin, margin flat. Inflorescence axillary, a fascicle, sometimes visible only two flowers per axis, the axis frequently inconspicuous, when visible ca. 1 mm long; bracts lanceolate, 0.6 × 4 mm, pubescent, persistent in the fruit; pedicels 18 – 26 mm long, glabrous; bracteoles orbicular, 0.3 × 2 mm, glabrescent, persistent in the fruit; flower buds globose, 3 – 4 × 3 – 6 mm, not constricted above ovary; calyx-lobes free, four, orbicular or oblong, 1.5 – 2 × 1 – 2 mm, glabrous, not reflexed, persistent in the fruit; ovary 2 – locular. Immature fruit subglobose or ellipsoid, 0.5 – 1 × 0.7 – 0.8 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE7FA7E887C4FDEDA701.taxon	discussion	Note: — Eugenia punicifolia may be recognized by its glabrous or sparsely pubescent leaves with an intra-marginal vein 1 – 3 mm from the margin; the inflorescence forming a fascicle with a very short axis (ca. 1 mm long), sometimes with only two flowers visible; its orbicular or oblong calyx lobes; and it’s subglobose or ellipsoid fruits, which are glabrous and have a smooth surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE7FA7E887C4FDEDA701.taxon	distribution	Distribution and habitat: — Widely distributed in South America, including Brazil, where it occurs in all states, except Santa Catarina and Rio Grande do Sul (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. punicifolia was found growing in secondary and flooded forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE7FA7E887C4FDEDA701.taxon	biology_ecology	Phenology: — Flowering in June, July and September and fruiting in January, November and December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE7FA7E887C4FDEDA701.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Maracassumé – Sta. Helena, Fazenda Maracaçumé, 1 ° 39 ’ 48 ” S, 45 ° 22 ’ 18 ” W, 01 December 1978 (fr.), N. A. Rosa 2824 (NY!). Grajaú, Estrada do Arame, 5 ° 49 ’ 10 ” S, 46 ° 08 ’ 19 ” W, 15 May 1983 (fr.), M. F. Silva & E. L. Taylor 1146 (IAN!). São Luís, 2 ° 31 ’ 47 ”, 44 ° 18 ’ 10 ” W, 08 May 1989 (fl., fr.), M. Ferreira 97 (ESA, image); Reserva Florestal da Sacavém, 09 January 1992 (fr.), F. H. Muniz 03 (RB, image); 2 ° 30 ’ S, 44 ° 15 ’ W, 05 May 1992 (fr.), F. H. Muniz 87 (INPA, image); 07 May 1992 (fl.), F. H. Muniz 99 (RB, image); Sede do CEMAR, 2 ° 30 ’ 32 ” S, 44 ° 15 ’ 49 ” W, 27 November 1985 (fl., fr.), G. T. Prance & R. Henrique 29896 (MAR!); Sítio Santa Eulália, 2 ° 30 ’ 30 ” S, 44 ° 16 ’ 39 ” W, 16 September 2017 (fr.), G. S. Amorim 401 (MAR!); G. S. Amorim 455 (MAR!); 8 º Batalhão da Polícia Militar, 2 ° 33 ’ 19 ” S, 44 ° 15 ’ 55 ” W, 07 April 2018 (st.), G. S. Amorim 487 (MAR!); 2 ° 29 ’ 49 ” S, 44 ° 16 ’ 43 ” W, 15 June 2018 (fl., fr), G. S. Amorim 508 (MAR!). Additional specimens examined: — BRAZIL. Pará: Serra do Cachimbo, BR 163, Cuiabá-Santarém Highway, Cachoeira de Curuá, 8 ° 44 ’ 12 ” S, 54 ° 57 ’ 52 ” W, 04 November 1977 (fl.), G. T. Prance P 24782 (NY!). Roraima: Ilha de Maracá, on road between SEMA Estação and Boa Vista, 14 km from Island, Alto Alegre, 3 ° 18 ’ N, 61 ° 19 ’ 59 ” W, 17 June 1986 (fl.), M. J. G. Hopkins 825 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE61A7E880FDFBC8A3E9.taxon	materials_examined	Type: — BRAZIL. Habitat in campis inter. Provinciae Minar, C. F. P. Martius s. n. (holotype: M [0171122]). (Fig 2 N-P)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE61A7E880FDFBC8A3E9.taxon	description	Shrubs 0.2 – 8 m. Twigs glabrous. Leaves with petioles 8 – 10 mm long, glabrous, channeled; blades elliptic to broadly so, oblong, oval or rounded, 6.5 – 13.1 × 4.3 – 6.4 cm, glabrous, chartaceous, concolorous; glands conspicuous and dark; base cuneate or rounded; apex acute or rounded; midvein adaxially impressed; lateral veins visible, not arched, 5 to 15 pairs; intra-marginal vein 1 – 5 mm from the margin, margin flat. Inflorescence axillary, a fascicle, the axis ca. 1 mm long or less; bracts orbicular, 0.6 × 2.4 mm, glabrous, persistent in the fruit; pedicels 5 – 19 mm long, glabrous or sparsely pubescent; bracteoles elliptic or oblong, 0.7 – 1 × 2 – 5 mm, glabrous, persistent in the fruit; flower buds globose, 4 – 6 × 2 – 5 mm, not constricted above ovary; calyx-lobes free, four, orbicular or oblong, 0.5 – 2.5 × 1.5 – 3 mm, glabrous, not reflexed, persistent in the fruit; ovary 2 – locular. Immature fruit globose, 0.9 – 1 × 0.6 – 0.8 cm, glabrous, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE61A7E880FDFBC8A3E9.taxon	discussion	Note: — Eugenia stictopetala may be recognized by its glabrous chartaceous leaves with dark glands visible; and its orbicular or oblong calyx-lobes; and globose fruit. This species is morphologically related to E. dittocrepis but can be distinguished by its flat marginal leaf vein (vs. revolute) and concolorous leaves (vs discolorous leaves).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE61A7E880FDFBC8A3E9.taxon	distribution	Distribution and habitat: — Known from Costa Rica, French Guiana, Guyana, Panamá, Peru, Suriname, TrinidadTobago, Venezuela and Brazil, where it is widely distributed (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. stictopetala was found growing in secondary forests and along roads.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE61A7E880FDFBC8A3E9.taxon	biology_ecology	Phenology: — Flowering from March to July and fruiting in April.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C479EE61A7E880FDFBC8A3E9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Funai a 40 Km da cidade de Arame, 14 March 1983 (fl.), R. Vilhena 1005 (NY!). Rio Maracaçumé, 01 July 1958 (fl.), R. L. Fróes 34408 (IAN!). Alcântara, 2 ° 24 ’ 32 ” S, 44 ° 24 ’ 53 ” W, 10 April 1954 (fr.), R. L. Fróes 30741 (IAN!). Grajaú, rodovia a 6 Km da cidade de Grajaú, 5 ° 49 ’ 10 ” S, 46 ° 08 ’ 18 ” W, 21 May 1983 (fr.), M. F. F. da Silva 1153 (NY!); Mondelandia, 5 ° 49 ’ 10 ” S, 46 ° 08 ’ 19 ” W, 23 May 1983 (fl.), E. L. Taylor E 1324 (NY!). São Luís, Granja Barreto, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 10 May 1949 (fl.), R. L. Fróes 24292 (IAN!); Reserva Florestal do Sacavém, 03 April 1992 (fr.), F. H. Muniz 50 - 423 (RB, image); Sítio São Raimundo, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 08 April 1954 (fl.), R. L. Fróes 30711 (IAN!); R. L. Fróes 30694 (IAN!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE61A7E884CDFAF1A775.taxon	materials_examined	Type: — SURINAME. Paramaribo, H. R. Wullschlägel 193 (holotype: BR [5288649] image; isotype: MICH [1109613] image). (Fig 4 E-G.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE61A7E884CDFAF1A775.taxon	description	Trees or shrubs 15.2 m. Twigs glabrous. Leaves with petioles 3 – 5 mm long, pubescent, unchanneled; blades elliptic to broadly so, or ovate, 6.2 – 11.1 × 3.1 – 4.7 cm, pubescent adaxially and densely pubescent abaxially, the indumentum whitish or brownish, membranaceous or chartaceous, discolorous; glands not seen; base cuneate or rounded; apex acute or attenuate; midvein adaxially impressed; lateral veins visible, not arched, 8 to 9 pairs; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence axillary, a fascicle, the axis ca. 3 mm long, pubescent, the indumentum yellowish; bracts deltate, 0.5 × 0.5 mm, pubescent, caducous after anthesis; pedicels 2 – 4 mm long, pubescent; bracteoles deltate, 0.5 × 0.5 mm, pubescent, caducous after anthesis; flower buds globose, 5 × 3 mm, not constricted above ovary; calyx-lobes free, four, deltate, 1 × 1.3 mm, pubescent adaxially and glabrous abaxially, not reflexed, persistent in the fruit; ovary not seen. Immature fruit ellipsoid, 1.5 – 1.9 × 0.8 cm, glabrous, surface with glands marked, no mature fruits examined.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE61A7E884CDFAF1A775.taxon	discussion	Note: — Eugenia wullschlaegeliana may be recognized by its leaves, which are ovate, elliptic, or broadly elliptic; its inflorescence forming a pubescent fascicle, with yellowish indumentum; and its glabrous, ellipsoid fruit, with marked glands on the surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE61A7E884CDFAF1A775.taxon	distribution	Distribution and habitat: — Known from French Guiana, Suriname and northeastern and northwestern Brazil (Amazonas, Maranhão and Pará states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, E. wullschlaegeliana was found growing in flooded forests and in open fields.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE61A7E884CDFAF1A775.taxon	biology_ecology	Phenology: — Flowering in October and fruiting in September.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE61A7E884CDFAF1A775.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Maracassumé River Region, Igarapé do Urubuquara, 09 September 1932 (fr.), R. L. Fróes 1885 (NY!); 22 October 1932 (fl.), R. L. Fróes 1967 (NY!). Additional specimens examined: — BRAZIL. Pará: Genipapo trail, Basin of Rio Xingu, Gleba Bacaja, lote 88, just below mouth of Rio Bacaja, 3 ° 22 ’ 20 ” S, 50 ° 47 ’ 50 ” W, 01 December 1980 (fl.), G. T. Prance P 26566 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE63A7E88049FD2AA2C9.taxon	materials_examined	Type: — BRAZIL. Amazonas: Solimoes, s. d., C. F. P. Martius s. n. (lectotype designated by McVaugh 1969 [p. 110]: M). (Fig. 5 A.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE63A7E88049FD2AA2C9.taxon	description	Tree or shrubs 2 – 10 m. Twigs glabrous or sparsely pubescent, the indumentum brownish, the branches and inflorescence reddish. Leaves with petioles 2 – 4 mm long, pubescent, channeled; blades elliptic, lanceolate or ovate, 3.7 – 6.7 × 1.9 – 3.2 cm, glabrous, chartaceous, discolorous; glands conspicuous adaxially and strongly marked; base cuneate or rounded; apex acute or acuminate, rarely cuspidate; midvein adaxially impressed of flat; lateral veins visible, not arched, 8 to 12 pairs; intra-marginal vein 1 – 2 mm from the margin, margin flat. Inflorescence terminal or subterminal, panicle, pyramidal, flowers lax, the axis 7.8 – 13.8 cm long, glabrous, the first branching 4.2 – 6.8 cm long; bracts linear, ca. 2 mm, caducous before anthesis, pubescent; pedicels 1 – 3 mm long or absent, sparsely pubescent; bracteoles lanceolate or linear, 0.7 – 2 × 0.2 mm, caducous before anthesis, glabrous or sparsely pubescent; flower buds obovate, 2 x 1.5 mm, not constricted above ovary; calyx-lobes free, five, orbicular, 0.5 × 1 mm, glabrous, not reflexed, persistent in the fruit; ovary 2 – locular. Immature fruit globose, 0.5 cm diameter, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE63A7E88049FD2AA2C9.taxon	discussion	Note: — Myrcia amazonica may be recognized by its reddish branchlets and inflorescence, and its pyramidal panicle with small lax buds. This species is morphologically similar to M. guianensis from which it is distinguished by its reddish branches and inflorescences (vs. grayish), leaves with an adaxially impressed or flat midvein (vs. prominent) and 2 - locular ovary (vs. 3 - locular).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE63A7E88049FD2AA2C9.taxon	distribution	Distribution and habitat: — Known from south Central America, Bolivia, Brazil, French Guiana, Suriname and Venezuela. In Brazil it is widely distributed (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. amazonica was found growing in “ terra firme ” and secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE63A7E88049FD2AA2C9.taxon	biology_ecology	Phenology: — Flowering from May to August and fruiting in August.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C467EE63A7E88049FD2AA2C9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: São José de Ribamar, Sítio Aguahy, 2 ° 38 ’ 51 ” S, 44 ° 08 ’ 04 ” W, 28 August 2017 (fr.), G. S. Amorim 374 (MAR!). São Luís, Granja Barreto, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 12 May 1950 (fl.), R. L. Fróes 24260 (IAN!); Reserva Florestal do Sacavem [Mata da Caema], 2 ° 34 ’ 04 ” S, 44 ° 15 ’ 44 ” W, 1958 (fl.), J. Ferreira et al. 58 (MAR!); 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 01 July 1992 (fl.), G. M. Barroso 61 (UEC, image); 27 May 1992 (fl.), F. H. Muniz 103 (RB, image); 2 ° 30 ’ S, 44 ° 15 ’ W, 14 August 1992 (fl.), F. H. Muniz 26 (INPA, image); 2 ° 30 ’ S, 44 ° 15 ’ W, 26 June 1993 (fl.), F. H. Muniz 392 (INPA, image); 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 28 June 1993 (fl.), F. H. Muniz 378 (INPA, image). Turiaçu, 1 ° 39 ’ 48 ” S, 45 ° 22 ’ 18 ” W, Fazenda Vila Palmares, 23 July 1977 (fl.), A. E. Silva 131 (PEUFR!). Additional specimens examined: — BRAZIL. Amazonas: Manaus, Igarapé do Passarinho, 2 ° 57 ’ S, 60 ° 02 ’ 50 ” W, 21 August 1956 (fl.), D. F. Coêlho INPA 4109 (INPA, image). Pará: Santarém, Comunidade São Brás, 2 ° 30 ’ 46 ” S, 54 ° 51 ’ 33 ” W, 12 October 2012 (fl.), V. Y. P. Calao s. n. (INPA, image). Rondônia: Porto Velho, Jaci-Paraná, 8 ° 45 ’ 43 ” S, 63 ° 54 ’ 14 ” W, 13 July 2008 (fl.), G. Pereira-Silva 13547 (RON, image); Fragmento de floresta próximo a margem direita do, Floresta estacional semidecidual com relevo plano e afloramento rochoso, 9 ° 16 ’ 19 ” S, 64 ° 23 ’ 37 ” W, 26 June 2012 (fl.), M. F. Simon 1557 (RON, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C465EE63A7E885ADFC90A96D.taxon	materials_examined	Type: — FRENCH GUIANA. Cayenne, Leblond s. n. (P [00297614] image) (Fig. 5 B.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C465EE63A7E885ADFC90A96D.taxon	description	Shrubs 1 – 3 m. Twigs pubescent, the indumentum yellowish. Leaves with petioles 2 – 5 mm long, pubescent, channeled; blades elliptic or lanceolate, 4 – 7.3 × 1.8 – 2.6 cm, glabrous or sparsely pubescent adaxially and pubescent abaxially, the indumentum yellowish, chartaceous, concolorous; glands conspicuous; base cuneate or rounded; apex acuminate; midvein adaxially impressed; lateral veins visible, not arched, 12 to 15 pairs; intra-marginal vein 1 – 2 mm from the margin, margin slightly revolute. Inflorescence axillary, thyrsoid, flowers congested, the axis 2.5 – 2.7 cm long, pubescent, the indumentum yellowish; the first branching ca. 1.5 cm; bracts lanceolate, 0.8 – 1.2 × 4 – 5 mm, pubescent abaxially and glabrous adaxially, persistent in the fruit; pedicels 3 – 5 mm long, pubescent; bracteoles lanceolate, 7 – 8 × 3 mm, pubescent abaxially and glabrous adaxially, persistent in the fruit; flower buds campanulate, 4 × 4 – 7 mm, not constricted above ovary; calyx-lobes free, five, lanceolate, 2 – 4 × 1.5 – 2 mm, pubescent abaxially and glabrous adaxially, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit ellipsoid, 0.6 – 0.8 × 0.8 – 1 cm, densely pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C465EE63A7E885ADFC90A96D.taxon	discussion	Note: — Myrcia bracteata may be recognized by its elliptic or lanceolate leaves, which are glabrous or sparsely pubescent adaxially and pubescent abaxially, the thyrsoid inflorescence, with yellowish indumentum, and long bracteoles (7 – 8 × 3 mm) with long yellowish indumentum, which are persistent, and the densely pubescent, ellipsoid fruit.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C465EE63A7E885ADFC90A96D.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela and central, northeastern and northwestern Brazil (Acre, Amazonas, Maranhão, Mato Grosso, Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. bracteata was found growing in “ terra firme ” forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C465EE63A7E885ADFC90A96D.taxon	biology_ecology	Phenology: — Flowering in February.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C465EE63A7E885ADFC90A96D.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Cândido Mendes, Fazenda 7 irmãos, 2 ° 26 ’ 31 ” S, 46 ° 18 ’ 07 ” W, 16 February 2020 (fl.), G. S. Amorim 618 (JPB!). Additional specimens examined: — BRAZIL. Amazonas: Manaus, Reserva Florestal Ducke, 2 ° 52 ’ 59 ” S, 59 ° 58 ’ 00 ” W, 24 January 1996 (fr.), M. A. D. Souza, J. R. Pirani, M. A. Costa & C. F. Silva 208 (SP, image). Pará: Santarém Novo, Bacuriteua, capoeira sul de Bacuriteua, 0 ° 57 ’ 54 ” S, 47 ° 23 ’ 37 ” W, 2010 (fl.), M. P. M. Menezes 567 (HBRA, image). Rondônia: Porto Velho, Estrada de acesso ao alojamento provisório, 9 ° 18 ’ 19 ” S, 64 ° 36 ’ 24 ” W, 07 January 2009 (fr.), G. Pereira-Silva 14019 (RON, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE62A7E886E9FC5FA489.taxon	materials_examined	Type: — BRAZIL. “ Flor. Amazon. Insula Colares. In littore ”, E. F. Poeppig 2937 (lectotype designated by McVaugh [1969, p. 290]: W [0033248] image). (Figs. 4 H-K, 5 C-D)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE62A7E886E9FC5FA489.taxon	description	Trees or shrubs 3 – 7 m. Twigs pubescent, the indumentum brownish-red. Leaves with petioles 5 – 7 mm long, pubescent, channeled; blades elliptic to broadly so, or oval, 3.8 – 6.7 × 3 – 5 cm, pubescent adaxially and densely pubescent abaxially, the indumentum brownish-red, chartaceous or coriaceous, concolorous; glands conspicuous; base rounded or cuneate; apex acute, rounded or obtuse; midvein adaxially impressed or flat; lateral veins visible, not arched, 8 to 12 pairs; intramarginal vein ca. 1 mm or less from the margin, margin flat. Inflorescence terminal or subterminal, panicle, pyramidal, flowers lax, the axis 3.6 – 6 cm long, pubescent, the indumentum brownish-red, the first branching 1.9 – 4 cm; bracts lanceolate, 4 × 2 mm, pubescent, caducous before anthesis; pedicels 0.5 – 1 mm long, pubescent; bracteoles lanceolate, 0.5 × 2 mm, pubescent, caducous before anthesis; flower buds campanulate, 2 – 3 × 2 mm, not constricted above ovary; calyx-lobes free, five, orbicular, 0.5 × 1 mm, pubescent, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit globose, 0.6 – 0.8 × 0.4 – 0.6 cm, pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE62A7E886E9FC5FA489.taxon	discussion	Note: — Myrcia cuprea may be recognized by its leaves, which are elliptic to broadly elliptic, or oval, with brownish-red indumentum; campanulate flower buds; and globose fruits, which are pubescent and have a smooth surface. This species is morphologically related to M. guianensis from which it is distinguished by its pubescent branches and leaves (vs. glabrous), campanulate flower buds (vs. globose or obovate) and 2 - locular ovary (vs. 3 - locular).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE62A7E886E9FC5FA489.taxon	distribution	Distribution and habitat: — Known from French Guiana, Suriname and northeastern and northwestern Brazil (Amazonas, Amapá, Maranhão and Pará states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. cuprea was found growing in secondary and “ terra firme ” forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE62A7E886E9FC5FA489.taxon	biology_ecology	Phenology: — Flowering in May, July, August and November and fruiting in January.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE62A7E886E9FC5FA489.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Ao longo do rio. Rio Gurupi, 15 August 1958 (fl.), R. L. Fróes 34555 (IAN!). Anajatuba, 3 ° 15 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 26 January 1976 (fr.), B. G. S. Ribeiro & G. S. Pinheiro 1228 (IAN!). Cândido Mendes, 1 ° 26 ’ 49 ” S, 45 ° 43 ’ 00 ” W, 02 November 2017 (fl.), G. S. Amorim 525 (MAR!); Maracassumé River region, Ilha do Trauira, 1 ° 12 ’ 52 ” S, 45 ° 33 ’ 33 ” W, 31 August 1932 (fl.), R. L. Fróes 1842 (NY!). Cururupu, Faz. Sto Antonio, 1 ° 49 ’ 42 ” S, 44 ° 52 ’ 05 ” W, August 1914 (fl.), A. Lisboa 56 (NY!); Perto do Palacete, 1 ° 46 ’ 18 ” S, 44 ° 53 ’ 23 ” W, 21 August 2014 (fl.), M. A. Machado 14 (MAR!). Maracassumé, Santa Helena. Próximo do Lugarejo Queimadas, 2 ° 13 ’ 52 ” S, 45 ° 17 ’ 59 ” W, 09 July 1978 (fl.), N. A. Rosa & O. Cardoso 2547 (IAN!). São Luís, Alumar, 2 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 13 November 1995 (fl.), K. B. Ferreira & E. C. Girnos s. c (MAR!); Anil, Arredores da estiva. Sitio Andiroba, 06 August 1980 (fl.), M. G. Silva 5676 (MG!); Granja Barreto, 03 September 1952 (fl.), R. L. Fróes 28549 (IAN!); Granja Federal, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 12 May 1949 (fl.), R. L. Fróes 24247 (IAN!); Sítio Santa Eulália, 2 ° 30 ’ 30 ” S, 44 ° 16 ’ 39 ” W, 16 September 2017 (fr.), G. S. Amorim 399 (MAR!), G. S. Amorim 400 (MAR), G. S. Amorim 451 (MAR!); 2 ° 33 ’ 57 ” S, 44 ° 18 ’ 51 ” W, 29 January 2018 (fr.), G. S. Amorim 428 (MAR!), G. S. Amorim 429 (MAR!); Parque Ambiental Alumar, 2 ° 41 ’ 31 ” S, 44 ° 19 ’ 42 ” W, 07 September 1999 (fl.), G. Richardson s. n (MAR!). Reserva Florestal do Sacavém, 2 ° 53 ’ 37 ” S, 44 ° 30 ’ 28 ” W, 17 July 1992 (fl.), F. H. Muniz 141 (INPA, image); 2 ° 34 ’ 04 ” S, 44 ° 15 ’ 44 ” W, 30 August 1995 (fl.), J. Ferreira & E. Barroso 92 (MAR!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE65A7E8806DFF5FA30D.taxon	materials_examined	Type: — BRAZIL. Minas Gerais, “ in Brasiliae deserto ad Minas Geraes ” C. F. P. Martius s. n. (holotype: M [0136862]). (Figs. 5 E-F)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE65A7E8806DFF5FA30D.taxon	description	Trees 5 m. Twigs glabrescent or sparsely pubescent, the indumentum yellowish. Leaves with petioles 4 – 7 mm long, pubescent, channeled; blades elliptic, oblong or oval, 7 – 11.1 × 2.8 – 5.2 cm, pubescent, the indumentum whitish, chartaceous, concolorous; glands conspicuous; base cuneate or rounded; apex acuminate or long acuminate; midvein adaxially sulcate; lateral veins visible and strongly marked, not arched, 12 to 16 pairs; intra-marginal vein 1 – 2 mm from the margin, margin revolute. Inflorescence terminal or subterminal, panicle, pyramidal, sometimes two per axis, flowers lax, the axis 7 – 11.1 cm long, pubescent, the indumentum whitish, the first branching 1 – 6.5 cm; bracts not seen, caducous before anthesis; pedicels ca. 0.5 mm long or absent, glabrous; bracteoles lanceolate, 1.5 × 0.5 mm, glabrous, caducous after anthesis; flower buds globose, 2 × 3 mm, not constricted above ovary; calyx-lobes free, five, orbicular 0.5 – 1 × 0.5 – 1.5 mm, glabrous, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit globose or ellipsoid, 0.8 – 1 × 0.6 – 1.2 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE65A7E8806DFF5FA30D.taxon	discussion	Note: — Myrcia eximia may be recognized by its elliptic, oblong or oval leaves, which are pubescent with whitish indumentum; the visible lateral veins, which are strongly marked; and the pubescent panicle. This species is morphologically related to M. splendens from which it is distinguished by its sulcate adaxially midvein (vs. impressed), and the lateral veins that are strongly marked adaxially and abaxially (vs. not marked).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE65A7E8806DFF5FA30D.taxon	distribution	Distribution and habitat: — Known from central and northeastern Brazil (Bahia, Goiás, Maranhão, Mato Grosso, Minas Gerais and Piauí states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. eximia was found growing in “ terra firme ” forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE65A7E8806DFF5FA30D.taxon	biology_ecology	Phenology: — Flowering in February.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C464EE65A7E8806DFF5FA30D.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Cândido Mendes, Fazenda 7 irmãos, 2 ° 26 ’ 31 ” S, 46 ° 18 ’ 07 ” W, 16 February 2020 (fl.), G. S. Amorim 619 (JPB!). Additional specimens examined: — BRAZIL. Pará. Altamira: Sete Palmeiras, 3 ° 12 ’ 12 ” S, 52 ° 12 ’ 23 ” W, 12 December 1986 (fr.), S. M. Athie de Souza, O. Nascimento, M. Santos & J. C. Santos 673 (US, image); Salvaterra, Condeixas, 0 ° 45 ’ 12 ” S, 48 ° 31 ’ 00 ” W, 10 January 1982 (fl.), C. S. Rosáro & E. Taylor 124 (MBM, image). Piaui. Uruçuí, estrada Waldemar Bertozzolo, 7 ° 13 ’ 46 ” S, 44 ° 33 ’ 22 ” W, 19 November 2005 (fl.), A. M. Miranda 5222 (HUEFS, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE65A7E884E1FBF1A7C5.taxon	materials_examined	Type: — GUYANA. “ Guiana Angelica, ad fines Guianae Batavae ”, R. H. Schomburgk 979 (lectotype designated by McVaugh [1989, p. 475]: G).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE65A7E884E1FBF1A7C5.taxon	description	Shrubs ca. 6 m. Twigs pubescent, the indumentum brownish-red. Leaves with petioles 4 – 5 mm long, sparsely pubescent, channeled or shallowly channeled; blades elliptic to broadly so, or ovate, 4.7 – 8.7 × 2 – 4 cm, glabrous adaxially and pubescent abaxially, the in indumentum dument brownish-red, membranaceous, discolorous; glands not seen; apex long acuminate or cuneate; base rounded; midvein adaxially slightly sulcate; lateral veins slightly visible, not arched, 16 to 20 pairs; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence axillary or subterminal, panicle, umbelliform, flowers congested, the axis 0.5 – 0.7 cm long, sometimes inconspicuous, pubescent, the indumentum brownish-red; bracts not seen, caducous before anthesis; pedicels 8 mm long, pubescent, the indumentum brownishred; bracteoles not seen, caducous before anthesis; flower buds globose, 4 × 5 mm, not constricted above ovary; calyxlobes completely fused, opening by calyptra, caducous in the fruit; ovary not seen. Immature fruits globose, 0.7 × 0.8 cm, pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE65A7E884E1FBF1A7C5.taxon	discussion	Note: — Myrcia fasciculata may be recognized by its leaves, which are elliptic to broadly elliptic, or ovate, with slightly visible lateral veins; an umbeliform panicle; and globose fruits which are pubescent and have a smooth surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE65A7E884E1FBF1A7C5.taxon	distribution	Distribution and habitat: — Known from Guyana and northeastern and northwestern Brazil (Amazonas, Pará, Maranhão, Mato Grosso, Rondônia and Roraima states) (Rosário et al. 2014). In Amazonian Maranhão, M. fasciculata was found growing in flooded forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE65A7E884E1FBF1A7C5.taxon	biology_ecology	Phenology: — Fruiting in December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE65A7E884E1FBF1A7C5.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Pinheiro, Rio Alto Turiaçu, Nova Esperança, 2 ° 55 ’ 12 ” S, 45 ° 45 ’ W, 04 December 1978 (fr.), J. Jangoux 213 (NY!). Additional specimens examined: — BRAZIL. Amazonas, Santa Isabel do Rio Negro, Margem direita do Rio Negro, serra do Jacamin, 150 m, 0 ° 18 ’ S, 65 ° 28 ’ 48 ” W, 11 October 1987 (fl.), C. A. C. Ferreira, P. Maas, M. V. Roosmalen, R. P. Lima & J. G. Oliveira 9347 (US, image). Rondonia, Espigão do Oeste, BR- 364, rodovia Cuiabá-Porto Velho, estrada da FUNAI, km 05, 11 ° 20 ’ 53 ” S, 60 ° 45 ’ 21 ” W, 20 June 1984 (fr.), C. A. C. Ferreira 4678 (NY!); Estrada Boa Vista-Venezuela, Rio Surumu, 01 December 1977 (fr.), W. C. Steward 161 (NY!). Roraima, Ilha de Maracá: Sema Ecological Reserve, 3 ° 21 ’ N, 61 ° 31 ’ 00 ” W, 28 April 1987 (fr.), W. Milliken M. 129 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE64A7E880B9FD08A3E9.taxon	materials_examined	Type: — VENEZUELA. Amazonas: Rio Siapa, Casiquiare, Maguire et al. 37617 (holotype: MICH [1109512]). (Fig. 4 L-N)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE64A7E880B9FD08A3E9.taxon	description	Trees to shrub 3 - 7 m. Twigs glabrous. Leaves with petioles 4 – 6 mm long, glabrous, channeled; blades elliptic or lanceolate, 4 – 6 × 1.9 – 2.4 cm, glabrous, chartaceous or coriaceous, discolorous, more lustrous adaxially; glands not seen; base subacute or cuneate; apex acute or acuminate, often long acuminate; midvein adaxially impressed; lateral veins numerous and slightly visible, not arched, 15 to 20 pairs; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence terminal or subterminal, panicle, pyramidal, flowers lax, the axis 4.2 – 6.6 cm long, pubescent, the indumentum whitish, the first branching 2.4 – 5.8 cm long; bracts lanceolate, 2 – 4 × 2 mm, pubescent, caducous before anthesis; pedicels 1 mm long, glabrous; bracteoles lanceolate, 1 – 2 × 1 mm, pubescent, caducous before anthesis; flower buds campanulate, 2 – 2.3 × 1.5 mm, not constricted above ovary; calyx-lobes free, four, orbicular, 0.5 – 1 mm, glabrous, not reflexed; ovary not seen. Immature fruits globose, 0.4 – 0.6 × 0.6 – 0.8 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE64A7E880B9FD08A3E9.taxon	discussion	Note: — Myrcia grandis may be recognized by its elliptic or lanceolate leaves, which are more lustrous adaxially; the pubescent panicle; and the campanulate and glabrous calyx-lobes. This species is morphologically related to M. sylvatica from which it is distinguished by its glabrous leaves (vs. abaxially pubescent) with an adaxially impressed midvein (vs. adaxially sulcate).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE64A7E880B9FD08A3E9.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Guyana, Panamá, Venezuela and northeastern and northwestern Brazil (Amazonas, Maranhão, Pará, Rondônia and Roraima states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. grandis was found growing on riversides.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE64A7E880B9FD08A3E9.taxon	biology_ecology	Phenology: — Flowering in December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C463EE64A7E880B9FD08A3E9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Pinheiro: Rio Alto Turiaçu, Nova Esperança, 100 m, 2 ° 55 ’ 12 ” S, 45 ° 45 ’ W, 01 December 1978 (fl.), J. Jangoux & R. P. Bahia 128 (NY!). Additional - sSpecimens examined: — BRAZIL. Amazonas: Igarapé Lobisomen, Rio Cuieras, 2 ° 31 ’ 12 ” S, 60 ° 15 ’ W, 22 July 1991 (fl.), S. A. Mori 21827 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE64A7E884CDFA14A669.taxon	materials_examined	Type: — BRITISH GUIANA. ‘ Habitat in silvis propè montem Serpent dictum’, s. d., Aublet s. n. (holotype BM [000953654], image; isotype LINN [HS 883 - 22]). (Figs. 5 G-H)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE64A7E884CDFA14A669.taxon	description	Trees or shrubs 3 – 10 m. Twigs glabrous. Leaves with petioles 2 – 5 mm long, glabrous, channeled; blades elliptic, obovate, oval or oblong, 2.2 – 7.2 × 2 – 4 cm, glabrous, chartaceous or coriaceous, concolorous; glands conspicuous; base rounded or cuneate; apex acute, rounded or obtuse; midvein adaxially prominent or flat; lateral veins visible, not arched, 13 to 17 pairs; intra-marginal vein 1 – 2 mm from the margin, margin flat. Inflorescence terminal or subterminal, panicle or thyrsoid, flowers lax, the axis 4 – 6.1 cm long, puberulent, the indumentum brownish, the first branching 2.7 – 3.1 cm long; bracts lanceolate, 0.5 × 1 mm, glabrous, caducous after anthesis; pedicels 0.5 – 2 mm long, glabrous; bracteoles lanceolate, 0.5 × 1 mm, glabrous, caducous after anthesis; flower buds globose or obovate, 2 × 1 mm, not constricted above ovary; calyx-lobes free, five, orbicular, 0.5 – 1 × 1 – 2.5 mm, glabrous, not reflexed, persistent in the fruit; ovary 3 - locular. Immature fruit globose, 0.4 – 0.7 × 0.5 – 0.6 cm, glabrous, smooth surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE64A7E884CDFA14A669.taxon	discussion	Note: — Myrcia guianensis may be recognized by its glabrous leaves, the globose or obovate flower buds, and the 3 - locular ovary. This species is morphologically related to M. amazonica from which it is distinguished by its grayish branches (vs. reddish) an adaxially prominent or flat midvein (vs. impressed). It is also morphologically related to M. cuprea from which it is distinguished by its glabrous branches and globose or obovate flower buds (vs. pubescent branches and campanulate flower buds).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE64A7E884CDFA14A669.taxon	distribution	Distribution and habitat: — Bolivia, Colombia, Ecuador, French Guiana, Guyana, Panamá, Paraguay, Peru, Suriname, Trinidad-Tobago and Venezuela and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, Myrcia guianensis was found growing in “ terra firme ” and secondary forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE64A7E884CDFA14A669.taxon	biology_ecology	Phenology: — Flowering in August to December and fruiting in October.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE64A7E884CDFA14A669.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Old capoeira on hillside W. of road, appr. 45 km. S. of Itauna on road from Pinheiro, 2 ° 27 ’ S, 44 ° 49 ’ 59 ” W, 18 October 1980 (fl., fr), D. C. Daly & D. G. Campbell 686 (IAN!). São Luis, Alumar, 2 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 13 November 1995 (fl.), K. B. Ferreira & E. Girnos s. n. (MAR!). Pinheiro, Nova Esperança, Rio Alto Turiaçu, 100 m, 2 ° 55 ’ 12 ” S, 45 ° 45 ’ W, 04 December 1978 (fl.), J. Jangoux 199 & R. P. Bahia (NY!). Additional specimens examined: — BRAZIL. Pará: Santarém, along E side of road to Punta Piedra, about 3.4 km from where the road departs to the NW from the Santarém – Alter do Chão road (PA- 457), ca. 21.5 km E and somewhat S of Santarém, 73 m, 2 ° 30 ’ 17 ” S, 54 ° 53 ’ 04 ” W, 24 November 2011 (fr.), B. M. Torke 1292 (NY!); Oriximinã, Porto Trombetas. Mineração Rio do Norte, Mina Saracá, 1 ° 42 ’ S, 56 ° 27 ’ W, 1999 (fl.), E. M. Barbosa 324 (INPA, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE67A7E8814DFC2CA225.taxon	materials_examined	Type: — FRENCH GUIANA. Maroni, E. M. Mélinon 442 (lectotype designated by McVaugh [1969, p. 1 - 2]: P). (Fig 6 A-B)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE67A7E8814DFC2CA225.taxon	description	Shrubs 3 m. Twigs pubescent, the indumentum yellowish. Leaves with petioles ca. 4 mm long, pubescent, channeled; blades elliptic or broadly so, 7 – 9.5 × 2.9 – 4 cm, glabrous adaxially and sparsely pubescent abaxially, the indumentum yellowish, membranaceous or chartaceous, discolorous, darker adaxially than abaxially, glands conspicuous; base cuneate or rounded; apex long-caudate; midvein adaxially sulcate; lateral veins visible and prominent abaxially, arched, 7 to 11 pairs; intra-marginal vein 4 – 6 mm from the margin, margin flat. Inflorescence axillary or terminal, a thyrsoid, sometimes much-reduced, a single flower, flowers congested, the axis 2.9 – 4.2 cm, sometimes inconspicuous, glabrous, the first branching 0.9 – 1.7 cm long, with two lateral branches; bracts lanceolate 1.5 × 1.5 mm, glabrous, caducous after anthesis; pedicels 4 mm long, pubescent; bracteoles not seen, caducous before anthesis; flower buds globose, 1 × 1 mm, not constricted above ovary; calyx-lobes free, four, deltate, 0.5 × 0.5 mm, glabrous adaxially and pubescent abaxially, not reflexed, persistent in the fruit; ovary not seen. Immature fruit globose, 1 cm diameter, sparsely pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE67A7E8814DFC2CA225.taxon	discussion	Note: — Myrcia minutiflora may be recognized by its discolorous, elliptic to broadly elliptic leaves with a longcaudate leaf apex; visible lateral veins and a prominent abaxially, arched, intra-marginal vein 4 – 6 mm from the margin; and a thyrsoid inflorescence, sometimes reduced to a solitary flower.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE67A7E8814DFC2CA225.taxon	distribution	Distribution and habitat: — Known from, French Guiana, Guyana, Peru, Suriname, Venezuela and central, northeastern and northwestern Brazil (Acre, Amazonas, Amapá, Maranhão, Mato Grosso and Pará states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. minutiflora was found growing in disturbed forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE67A7E8814DFC2CA225.taxon	biology_ecology	Phenology: — Fruiting in September.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C462EE67A7E8814DFC2CA225.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Monção, Basin of the Rio Turiaçu, 3 ° 09 ’ 58 ” S, 45 ° 56 ’ 17 ” W, 16 May 1985 (st.), W. L. Balée 883 (NY!). Nova Olinda do Maranhão, S. of Fazenda Guarany, Km 133 of Br 316, 2 ° 46 ’ 12 ” S, 45 ° 43 ’ 12 ” W, 21 September 1980 (fr.), D. C. Daly D 162 (NY!). Additional specimens examined: — BRAZIL. Amazonas: Manaus-Itacoatiara, Reserva Florestal Ducke, 2 ° 52 ’ 59 ” S, 59 ° 58 ’ 00 ” W, 30 December 1997 (fl.), P. A. C. L. Assunção 759 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C461EE67A7E88519FCFCA6A5.taxon	materials_examined	Lectotype: — FRENCH GUIANA. s. l., Leprieur, s. n. (syntype: P [P 00163142], image). (Fig. 5 I-J.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C461EE67A7E88519FCFCA6A5.taxon	description	Trees or shrubs 2 – 10 m. Twigs glabrescent, the young branchlets pubescent, the indumentum whitish. Leaves with petioles 2 – 4 mm long, glabrous, pubescent on young branches, channeled; blades elliptic, lanceolate or ovate, 3.8 – 6 × 1 – 3.8 cm, glabrescent, membranaceous, concolorous, glands conspicuous and abundant; base cuneate or rounded; apex acute or acuminate; midvein adaxially impressed; lateral veins dark, flat to impressed, with numerous intermediates veins, not arched, 10 to 21 pairs; intra-marginal vein 1 – 2 mm from the margin, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers lax, the axis 12 – 31 mm long, glabrescent, the first branching 1.5 cm; bracts lanceolate 1 – 2 × 1 – 2 mm, glabrous, caducous after anthesis; pedicels 1 – 3 mm long, glabrous; bracteoles lanceolate, 1 × 1 mm, glabrous, caducous after anthesis; flower buds globose, 1 – 2.5 × 1 – 2 mm, not constricted above ovary; calyx-lobes free, five, orbiculate, 0.5 – 1 × 0.5 – 1 mm, glabrous adaxially and pubescent abaxially, not reflexed; ovary 2 - locular. Immature fruits globose, 0.5 cm diameter, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C461EE67A7E88519FCFCA6A5.taxon	discussion	Note: — Myrcia multiflora may be recognized by its elliptic, lanceolate or ovate leaves, which are concolorous and with abundant glands, and the dark lateral veins with numerous intermediate veins visible in dried material. This species is morphologically related to M. selloi, from which it is distinguished by its lateral leaf veins, which are flat abaxially and not arched (vs. prominent abaxially and arched) and erect calyx-lobes (vs. calyx-lobes reflexed).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C461EE67A7E88519FCFCA6A5.taxon	distribution	Distribution and habitat: — Known from northeastern Argentina, Bolivia, French Guiana, Guyana, Paraguay, Peru, Suriname, Trinidad-Tobago, Uruguay and Venezuela and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. multiflora was found growing in “ terra firme ” and secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C461EE67A7E88519FCFCA6A5.taxon	biology_ecology	Phenology: — Flowering in May and September.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C461EE67A7E88519FCFCA6A5.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Fazenda São Francisco, 11 km. N. of km. 337 of BR- 316, 4 ° 00 ’ 00 ” S, 44 ° 55 ’ 59 ” W, 25 September 1980 (fl.), D. C. Daly & D. G. Campbell 268 (IAN!). Bacabal, Fazenda São Francisco, 4 ° 00 ’ 00 ” S, 44 ° 55 ’ 48 ” W, 25 September 1980 (fl.), D. C. Daly D 268 (NY!). Cândido Mendes, 1 ° 26 ’ 49 ” S, 45 ° 43 ’ 00 ” W, 02 November 2017 (fl.), G. S. Amorim 524 (MAR!). São José de Ribamar, Sítio Aguahy, 2 ° 38 ’ 34 ” S, 44 ° 08 ’ 19 ” W, 07 May 2016 (fl.), G. S. Amorim 271 (MAR!). Santa Helena, próximo do Lugarejo Queimadas, 2 ° 13 ’ 52 ” S, 45 ° 17 ’ 59 ” W, 09 September 1978 (fl.), N. A. Rosa & O. Cardoso 2559 (IAN!). Additional specimens examined: — BRAZIL. Pará: Marajás, Carajás, Serra Norte. Estrada H- 7,30 km do acampamento, 6 ° 04 ’ 15 ” S, 50 ° 21 ’ 04 ” W, 04 August 1982 (fl.), U. N. Maciel 742 (NY!): Serra dos Carajás, 25 - 30 km NW of Serra Norte mining camp Pará, 5 ° 55 ’ 00 ” S, 50 ° 25 ’ 59 ” W, 05 December 1981 (fr.), D. C. Daly with R. Callejas, M. G. da Silva, E. L. Taylor, C. Rosario, M R. dos Santo 1739 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E886E9FF5FA5E1.taxon	materials_examined	Type: — BRAZIL. Minas Gerais, Widgren 536 U (lectotype designated by Araújo & Lucas [2013, p. 434]). (Fig. 6 C-E.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E886E9FF5FA5E1.taxon	description	Shrubs 2 – 4 m. Twigs glabrous or sparsely pubescent, the indumentum ochraceous. Leaves with petioles 9 – 14 mm long, pubescent, unchanneled; blades elliptic or broadly so, ovate or oval, 7 – 11.5 × 3.5 – 6.5 cm, glabrous adaxially and sparsely pubescent abaxially, the indumentum ochraceous, chartaceous or coriaceous, slightly discolorous, glands not seen; base rounded, rarely acute; apex acute or rounded; midvein adaxially impressed or sulcate distally, densely covered with indumentum; lateral veins visible, not arched, 11 to 14 pairs; intra-marginal vein 1 – 3 mm from the margin, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers lax, the axis 5.4 – 9.3 cm long, pubescent, the indumentum ochraceous, with two lateral branches, opposite, 2.9 – 3 cm long; bracts not seen, caducous before anthesis; pedicels absent; bracteoles lanceolate or deltate, 2 – 4 × 2 – 4 mm, pubescent, caducous before anthesis; flower buds obovoid, 2 × 2 mm, not constricted above ovary; calyx-lobes completely fused, opening by calyptra, not reflexed, caducous in fruit; ovary not seen. Immature fruit globose, 0.5 cm diameter, densely pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E886E9FF5FA5E1.taxon	discussion	Note: — Myrcia neoclusiifolia may be recognized by its slightly discolorous elliptic to broadly elliptic, ovate or oval leaves; the pubescent panicle with two lateral branches; the ochraceous indumentum; the calyx-lobes, which are completely fused and opening by calyptra; and the globose fruits, which are densely pubescent and have a smooth surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E886E9FF5FA5E1.taxon	distribution	Distribution and habitat: — Known from Bolivia, Venezuela and central, southeastern and northeastern Brazil (Bahia, Distrito Federal, Maranhão, Minas Gerais, Pernambuco and São Paulo states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. neoclusiifolia was found growing in “ terra firme ” and secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E886E9FF5FA5E1.taxon	biology_ecology	Phenology: — Flowering and fruiting in January and December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E886E9FF5FA5E1.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: São Luís, 2 ° 34 ’ 38 ” S, 44 ° 12 ’ 04 ” W, February-March 1939 (st.), R. L. Fróes 11732 (NY!); Estrada que vai do Rio Anil para Maioba, terra baixa perto de uma enseada, 10 January 1950 (fl., fr.), R. L. Fróes 25656 (IAN!); reservatório Sacaven, 15 January 1951 (fl. Fr.), R. L. Fróes 26819 (IAN!); vizinhanças do Reservatório de Sacaven, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 27 December 1951 (fl., fr.), R. L. Fróes 27809 (IAN!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E882C5FB3CA949.taxon	materials_examined	Type: — FRENCH GUIANA. M. Mélinon 488 (type: P [00723171], image). (Fig. 6 F-G.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E882C5FB3CA949.taxon	description	Trees 6 – 8 m. Twigs glabrous. Leaves with petioles 13 – 25 mm long, pubescent, unchanneled; blades ovate-lanceolate, oblong, rarely elliptic, 14.5 – 31.4 × 6 – 10 cm, glabrous adaxially and pubescent abaxially, the indumentum brownishred, chartaceous, discolorous, more lustrous adaxially; glands not seen; base rounded; apex acute or long acuminate; midvein adaxially impressed or sulcate; lateral veins slightly visible, not arched, 30 to 38 pairs; intra-marginal vein 1 mm or less from the margin, nearly straight, very close the margin, margin revolute. Inflorescence axillary, pyramidal panicle, flowers lax, the axis 2.3 – 3 cm long, pubescent, the indumentum brownish-red, with two lateral branches, opposite, 3.7 – 6.6 cm long; bracts not seen, caducous before anthesis; pedicels 3 mm long or sessile, pubescent; bracteoles not seen, caducous before anthesis; flower buds not seen; calyx-lobes completely fused, opening by calyptra, not reflexed, caducous in fruit; ovary not seen. Immature fruit subglobose to globose, 0.6 – 0.8 × 0.7 – 0.9 cm, pubescent, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E882C5FB3CA949.taxon	discussion	Note: — Myrcia neospeciosa may be recognized by its ovate-lanceolate or oblong, rarely elliptic leaves, which are more lustrous adaxially; the nearly straight intra-marginal vein, which is 1 mm or less from the margin; the calyxlobes, which are completely fused and opening by calyptra; and the subglobose or globose fruits, which are pubescent and have a smooth surface.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E882C5FB3CA949.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Peru, Suriname, Venezuela and northeastern and northwestern Brazil (Acre, Amapá, Maranhão, Pará and Rondônia states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. neospeciosa was found growing in “ terra firme ” forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E882C5FB3CA949.taxon	biology_ecology	Phenology: — Fruiting in December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46FEE69A7E882C5FB3CA949.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Rio Maracassumé, 03 July 1958 (st.), R. L. Fróes 34439 (IAN!). Additional specimens examined: — BRAZIL. Pará: Ilha de Marajó. Rio Anajas, 1 ° 00 ’ 00 ” S, 50 ° 10 ’ 00 ” W, 28 October 1987 (fr.), H. T. Beck 209 (INPA, image). Amazonas: Manaus, Reserva Florestal Adolfo Ducke, Próx. picada da Petrobrás, 2 ° 52 ’ 59 ” S, 59 ° 58 ’ 00 ” W, 06 August 1997 (fr.), M. A. D. Souza 395 (INPA, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE68A7E886E9FE56A291.taxon	materials_examined	Type: — SURINAME. “ Raleigh Falls, in saxis Raleighvallen fluv. Coppenam ”, A. A. Pulle 341 (holotype: U [0004963]).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE68A7E886E9FE56A291.taxon	description	Trees ca. 5 m. Twigs glabrous, sometimes the branches terminal and the axis of inflorescence are reddish. Leaves with petioles 2 – 4 mm long, glabrous, unchanneled, dark; blades broadly elliptic or ovate 2 – 3.8 × 1.8 – 3.3 cm, glabrous, chartaceous, slightly discolorous; glands not seen; base obtuse, rounded or subcordate; apex rounded or obtuse, rarely acute; midvein adaxially impressed; lateral veins slightly visible, not arched, ca. 15 pairs; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers congested, the axis 3 – 6.7 cm long, glabrous, with two lateral branches, opposite, 1.9 – 2.1 cm long; bracts not seen, caducous before anthesis; pedicels 0.2 – 0.7 mm long or sessile, glabrous; bracteoles not seen, caducous before anthesis; flower buds not seen; calyx-lobes completely fused, opening by calyptra, not reflexed, caducous in fruit; ovary not seen. Immature fruit globose, 0.3 × 0.5 cm, glabrous, surface smooth.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE68A7E886E9FE56A291.taxon	discussion	Note: — Myrcia pullei may be recognized by its terminal branches and reddish axis of the inflorescence; the chartaceous, broadly elliptic or ovate leaves; the midvein of the leaf, which is adaxially slightly sulcate at the base and impressed distally; and the completely fused calyx-lobes, opening by calyptra.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE68A7E886E9FE56A291.taxon	distribution	Distribution and habitat: — Known from Colombia, Guyana, Peru, Suriname, Venezuela and from northeastern Brazil (Amazonas, Amapá, Maranhão and Pará states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. pullei was found growing along a river.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE68A7E886E9FE56A291.taxon	biology_ecology	Phenology: — Fruiting in April and February.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE68A7E886E9FE56A291.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Turiaçú, Rio Maraçumé, Cachoeira Grande, 04 April 1981 (fr.), M. R. Santos 697 (IAN!). Additional specimens examined: — BRAZIL. Amapá: Rio Oiapoque. Cachoeira Grande Roche, 11 August 1960 (fl.), H. S. Irwin 47412 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE6BA7E88255FCA6A1B9.taxon	materials_examined	Type: — BRAZIL. “ Brasilia ”, Sellow s. n. (neotype designated by Lannoy et al. [2019, p. 112]: K [000344138]). (Figs. 5 K-L.)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE6BA7E88255FCA6A1B9.taxon	description	Shrubs 1.5 – 3 m. Twigs sparsely pubescent or pubescent, the indumentum whitish. Leaves with petioles 2 mm long, pubescent, channeled; blades elliptic, lanceolate, oval or ovate, 3 – 10.2 × 1.2 – 5.1 cm, glabrescent, membranaceous, concolorous; glands conspicuous and dark abaxially; base cuneate or rounded; apex acute or acuminate; midvein adaxially flat; lateral veins visible and prominent abaxially, arched, 14 to 21 pairs; intra-marginal vein 3 – 4 mm from the margin, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers lax, the axis 3 – 4.3 cm long, puberulent, the indumentum whitish, the first branching 1.7 – 2.7 cm long; bracts lanceolate, 0.5 × 1 mm, glabrescent, caducous before anthesis; pedicels 2.5 – 3.5 mm long, glabrescent; bracteoles lanceolate, 0.5 – 1 × 1 mm, glabrescent, caducous before anthesis; flower buds globose or obovate, 1 × 3 mm, constricted above ovary; calyx-lobes free, five, triangular, 1 × 1 mm, glabrous, reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit globose, 0.3 × 0.5 cm, glabrous, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE6BA7E88255FCA6A1B9.taxon	discussion	Note: — Myrcia selloi may be recognized by its leaves with glands visible on both sides and dark abaxially; the globose or obovate flower buds constricted above ovary; and the reflexed calyx-lobes. This species is morphologically related to M. multiflora, from which it is distinguished by its abaxially prominent arched lateral veins (vs. flat and not arched), and reflexed calyx-lobes (vs. not reflexed). It is also morphologically related to M. tomentosa from which it is distinguished by its glabrescent and concolorous leaves (vs. glabrescent or sparsely pubescent adaxially and densely pubescent abaxially and discolorous), channeled petiole (vs. unchanneled), pedicellate flowers (vs. sessile flowers), and glabrous calyx-lobes (vs. pubescent).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE6BA7E88255FCA6A1B9.taxon	distribution	Distribution and habitat: — Known from northeastern Argentina, Bolivia, Paraguay and Uruguay and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, Myrcia selloi was found growing in secondary forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE6BA7E88255FCA6A1B9.taxon	biology_ecology	Phenology: — Flowering in January and October and fruiting in January.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46EEE6BA7E88255FCA6A1B9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Gururupú, December 1957 (st.), R. L. Fróes 33998 (IAN!). São José de Ribamar, Sítio Aguahy, 2 ° 38 ’ 48 ” S, 44 ° 08 ’ 47 ” W, 28 October 2017 (fl.), G. S. Amorim 385 (MAR!). São Luís, Sítio Santa Eulália, 2 ° 30 ’ 30 ” S, 44 ° 16 ’ 39 ” W, 29 January 2018 (fr.), G. S. Amorim 421 (MAR!); 2 ° 30 ’ 21 ” S, 44 ° 16 ’ 48 ” W, 11 January 1988 (fl.), R. Nina 3 (SLUI!); 2 ° 30 ’ 56 ” S, 44 ° 16 ’ 37 ” W, 20 January 1988 (fl.), E. Barroso 46 (SLUI!); Reserva da Caema, 2 ° 34 ’ 30 ” S, 44 ° 15 ’ 10 ” W, 15 January 1992 (fl.), J. Azevedo 11 (SLUI!). Palmeirândia, 2 ° 38 ’ 43 ” S, 44 ° 53 ’ 42 ” W, 07 January 2007 (fl.), C. M. Vieira 77 (IAN!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6BA7E88773FC47A473.taxon	materials_examined	Type: — HISPANIOLA, Olof Swartz s. n. (type: S [S-R- 3706], image). (Fig. 5 M-N)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6BA7E88773FC47A473.taxon	description	Trees 2 – 13 m. Twigs pubescent, the indumentum yellowish. Leaves with petioles 2 – 4 mm long, glabrous or pubescent, channeled; blades elliptic, lanceolate or ovate-lanceolate, 3 – 12.8 × 1.3 – 3.8 cm, glabrous or sparsely pubescent adaxially and pubescent abaxially, the indumentum yellowish, chartaceous, concolorous; glands inconspicuous; base cuneate or rounded; apex acute or acuminate; midvein adaxially flat; lateral veins slightly visible, not arched, 21 to 34 pairs; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers lax, the axis 4.9 – 8.5 cm long, pubescent, the indumentum yellowish, the first branching 2.5 – 3.9 cm long; bracts lanceolate, 1 × 2 mm, pubescent, caducous after anthesis; pedicels 1 – 4 mm long, pubescent; bracteoles not seen, caducous before anthesis; flower buds globose, 1 – 4 × 1 – 4 mm, not constricted above ovary; calyx-lobes free, five, orbicular, 0.5 – 1 × 2.1 – 3 mm, densely pubescent adaxially and pubescent or sparsely pubescent abaxially, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit ellipsoid, 0.5 – 1 × 0.4 – 0.7 cm, pubescent, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6BA7E88773FC47A473.taxon	discussion	Note: — Myrcia splendens may be recognized by its elliptic, lanceolate to ovate-lanceolate leaves, which are adaxially glabrous or puberulent and abaxially pubescent with yellowish indumentum, and its ellipsoid fruits, which are pubescent and have the surface marked with glands. This species is morphologically related to M. sylvatica, from which it is distinguished by its adaxially flat midvein (vs. adaxially sulcate) and channeled petiole (vs. unchanneled).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6BA7E88773FC47A473.taxon	distribution	Distribution and habitat: — Known from northeastern Tropical America, including all Brazilian states (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. splendens was found growing in secondary and lowland forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6BA7E88773FC47A473.taxon	biology_ecology	Phenology: — Flowering in May, June, August and October and fruiting in January.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6BA7E88773FC47A473.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: between Viana & Bandeirante, back road from Viana to Pinheiro, 3 ° 00 ’ 00 ” S, 45 ° 10 ’ 00 ” W, 17 October 1980 (fl.), D. C. Daly & D. G. Campbell 637 (IAN!). Rio Maracassumé, 01 January 1958 (fr.), R. L. Fróes 34418 (IAN!). Anajatuba, São Benedito, 2 ° 57 ’ 20 ” S, 44 ° 48 ’ 25 ” W, 30 May 2008 (fl.), A. Araújo 48 (IAN!). Grajaú, estrada do Arame, 5 ° 49 ’ 10 ” S, 46 ° 08 ’ 18 ” W, 15 May 1983 (fr.), M. F. F. da Silva 1145 (INPA, image). São Luís, 8 º Batalhão da Polícia Militar, 2 ° 29 ’ 49 ” S, 44 ° 16 ’ 43 ” W, 15 June 2018 (fl.), G. S. Amorim 509 (MAR!); Estação Ecológica do Rangedor, 2 ° 29 ’ 54 ” S, 44 ° 15 ’ 55 ” W, 30 October 2010 (fl.), S. M. Santos s. n (MAR!); Granja Federal, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 12 May 1949 (fl.), R. L. Fróes 24253 (IAN!). Palmeirândia, [s. l], 2 ° 38 ’ 43 ” S, 44 ° 53 ’ 42 ” W, 16 August 2011 (fl.), M. Ribeiro PM 25 (IAN!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6AA7E88349FF56A321.taxon	materials_examined	Type: — GUYANA. “ In silvis continentis, inprimis circa rivum Arowabischkreek ”, E. K. Rodschied 296 (holotype: GOET [GOET 008254]).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6AA7E88349FF56A321.taxon	description	Shrubs 2 m. Twigs glabrous or sparsely pubescent, the indumentum yellowish. Leaves with petioles 2 – 3 mm long, sparsely pubescent, unchanneled; blades elliptic or lanceolate, 1 – 5.1 × 1 – 1.6 cm, glabrous adaxially and pubescent abaxially, the indumentum yellowish, chartaceous, slightly discolorous, lustrous adaxially; glands conspicuous and marked; base cuneate or attenuate; apex attenuate or long-acuminate; midvein adaxially sulcate; lateral veins slightly visible, not arched, 17 to 22 pairs; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers lax, the axis 2.2 – 5.3 cm long, pubescent, the indumentum yellowish, the first branching 1.4 – 2.5 cm long; bracts not seen, caducous before anthesis; pedicels 1 – 2 mm long, pubescent; bracteoles not seen, caducous before anthesis; flower buds globose, 1.5 × 2 mm, not constricted above ovary; calyx-lobes free, five, deltate, 0.5 – 1 × 0.3 – 1 mm, pubescent adaxially and glabrous abaxially, not reflexed, persistent in the fruit; ovary 2 - locular. Immature fruit oblong or ellipsoid, 0.8 – 1 × 0.3 – 0.6 cm, pubescent, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6AA7E88349FF56A321.taxon	discussion	Note: — Myrcia sylvatica may be recognized by its adaxially sulcate midvein and slightly visible lateral veins, and the oblong or ellipsoid fruits which are pubescent and have the surface marked with glands. This species is morphologically related to M. splendens, from which it is distinguished by its adaxially sulcate midvein (vs. adaxially flat) and the unchanneled petiole (vs. channeled).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6AA7E88349FF56A321.taxon	distribution	Distribution and habitat: — Known from Tropical America, widely distributed in central, northeastern and northwestern Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. sylvatica was found growing in secondary forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6AA7E88349FF56A321.taxon	biology_ecology	Phenology: — Flowering in January, November and December and fruiting in January and December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46DEE6AA7E88349FF56A321.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Carutapera, Gurupiuna, Ka’apor Indian Reserve, 2 ° 43 ’ 12 ” S, 46 ° 25 ’ 48 ” W, 04 November 1986 (fl.), W. L. Balée 2839 (NY!). São Luís, Alumar, 2 º 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 18 January 1995 (fl.), N. Figueiredo s. n. (MAR!); 2 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 20 December 1994 (fl.), N. Figueiredo 89 (MAR!); Reserva Florestal do Sacavém, 2 ° 34 ’ 04 ” S, 44 ° 15 ’ 44 ” W, 03 April 1991, (fr.), J. Ferreira & E. Barroso 56 (MAR!); 2 ° 34 ’ 04 ” S, 44 ° 15 ’ 44 ” W, 03 December 1973 (fl., fr), L. F. Gomes 56 (MAR!). Alto Turiaçu, Carutapera, Gurupiuna, Ka’apor Indian Reserve, affluent of Rio Gurupi, 04 November 1986 (fl.), 2 ° 43 ’ 12 ” S, 46 ° 25 ’ 48 ” W, W. L. Balée 2839 with B. Ribeiro (NY!). Additional specimens examined: — BRAZIL. Maranhão: Carolina, Parque Nacional Chapada das Mesas, 7 ° 04 ’ 25 ” 47 ° S, 05 ’ 45 ” W, 20 October 2015 (fl.), A. C. Sevilha 5504 (CEN, image). Pará: Tucuruí, 25 km S of Tucuruí, just off old BR- 422 at junction with old railroad bed, 3 ° 58 ’ 00 ” S, 49 ° 37 ’ 59 ” W, 30 October 1981 (fl.), D. C. Daly 1071 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6AA7E88405FEAFA7A1.taxon	materials_examined	Type: — FRENCH GUIANA. “ Habitat ad ripam fluvii Sinemariensis tribus milliaribus à maris littore ”, 1775, Aublet s. n. (holotype: BM [000953642]).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6AA7E88405FEAFA7A1.taxon	description	Trees 1.5 – 8 m. Twigs glabrous or densely pubescent, the indumentum yellowish. Leaves with petioles 8 – 17 mm long, densely pubescent, unchanneled; blades elliptic to broadly so, lanceolate or oblong, 6.8 – 10.1 × 3 – 4.1 cm, glabrescent or sparsely pubescent adaxially and densely pubescent abaxially, the indumentum yellowish, membranaceous or chartaceous, discolorous; glands inconspicuous; base cuneate or attenuate; apex acute, rarely acuminate; midvein adaxially impressed; lateral veins visible and prominent abaxially, arched, 6 to 10 pairs; intra-marginal vein absent, margin flat. Inflorescence terminal or subterminal, pyramidal panicle, flowers congested, the axis 4.7 – 10 cm long, densely pubescent, the indumentum yellowish, the first branching 0.7 – 1.1 cm long; bracts lanceolate, 5 × 5 mm, pubescent, caducous after anthesis; pedicels absent; bracteoles linear 0.2 × 0.2 mm, pubescent, caducous after anthesis; flower buds ovate, 2 – 2.5 × 1.5 mm, constricted above ovary; calyx-lobes free, five, pubescent, ovate or triangular, 0.5 × 0.5 mm, reflexed; ovary 2 - locular. Immature fruit globose, 0.3 × 0.4 cm, glabrous or sparsely pubescent, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6AA7E88405FEAFA7A1.taxon	discussion	Note: — Myrcia tomentosa may be recognized by its midvein, which is abaxially densely pubescent and an adaxially impressed; the arched lateral veins; and the densely pubescent panicle with ovate flower buds. This species is morphologically related to M. selloi, from which it is distinguished by its discolorous leaves, which are glabrescent or sparsely pubescent adaxially and densely pubescent abaxially (vs. glabrescent and concolorous); the unchanneled petiole (vs. channeled); the sessile flowers (vs. pedicelate); and the pubescent calyx-lobes (vs. glabrous).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6AA7E88405FEAFA7A1.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Panamá, Peru, Suriname, Trinidad-Tobago and Venezuela and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. tomentosa was found growing in secondary forest.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6AA7E88405FEAFA7A1.taxon	biology_ecology	Phenology: — Flowering in February and November.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6AA7E88405FEAFA7A1.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: São José de Ribamar, Sítio Aguagy, 2 ° 38 ’ 47 ” S, 44 ° 09 ’ 05 ” W, 02 February 2018 (fl.), G. S. Amorim 434 (MAR!). São Luís, Alumar, 2 ° 41 ’ 30 ” S, 44 ° 19 ’ 44 ” W, 13 November 1995 (fl.), K. B. Ferreira & E. C. Girnos s. n (MAR!). Additional specimens examined: — BRAZIL. Amazonas: Southern ridge of Pico Rondon, perimetral norte highway km 211, 700 m, 1 ° 31 ’ 48 ” N, 62 ° 48 ’ W, 03 February 1984 (fr.), G. T. Prance 28776 (NY!). Maranhão: Estreito, Margem direita do rio Tocantins (área de mata úmida próximo à sua foz), 06 December 2008 (fr.), G. P. Silva 13888 (CEN, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6DA7E88085FC99A2C9.taxon	materials_examined	Type: — UNITED STATES. Virgin Islands, Ins. St. Crucis [St. Croix], M. Vahl s. n. (HAL [0089630], image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6DA7E88085FC99A2C9.taxon	description	Trees 3 – 4 m. Twigs puberulent, the indumentum whitish. Leaves with petioles 2 – 4 mm long, puberulent, unchanneled; blades elliptic or lanceolate, 4.4 – 7.3 × 2.8 – 3 cm, glabrous, membranaceous, concolorous; glands not seen; base cuneate; apex long acuminate; midvein adaxially flat, sometimes prominent; lateral veins inconspicuous; intra-marginal vein 1 mm or less from the margin, margin flat. Inflorescence glomerules clustered in axils, the axis inconspicuous; bracts not seen, caducous before anthesis; pedicels absent; bracteoles orbicular, ca. 0.5 × 0.5 mm, glabrous, caducous after anthesis; flower buds ovate, 1.5 × 1.5 mm, not constricted above ovary; calyx-lobes free, four, glabrous, ovate, 1 – 1.5 × 1 mm, not reflexed; the fruit falling together with the calyx, corolla and hypanthium; ovary not seen. Immature fruit globose, 0.5 – 0.7 × 0.5 – 0.9 cm, glabrous, surface with glands marked.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6DA7E88085FC99A2C9.taxon	discussion	Note: — Myrciaria floribunda may be recognized by its elliptic or lanceolate leaves with inconspicuous lateral veins, the long-acuminate leaf apex, the inflorescence of glomerules clustered in leaf axils, and the fruit, which falls together with the petals and hypanthium. This species is related to M. tenella, from which it is distinguished by leaves 4.4 – 7.3 × 2.8 – 3 cm (vs. 1 – 2.5 × 0.5 – 1.2 cm), the long-acuminate leaf apex (vs. acute leaf apex), and the 2 – 4 mm long petiole (vs. 1 – 2 mm).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6DA7E88085FC99A2C9.taxon	distribution	Distribution and habitat: — Widely distributed from Mexico to tropical America (POWO 2020). In Amazonian Maranhão, M. floribunda was found growing in secondary forests and along roads.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6DA7E88085FC99A2C9.taxon	biology_ecology	Phenology: — Flowering in May.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46CEE6DA7E88085FC99A2C9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Nova Esperança, Rio Alto Turiaçú, 17 May 1979 (fl.) I. J. Jangoux & R. P. Bahia 689 (MG!). Santa Inês, Colônia Três Satubas, margem da estrada de terra, a 12 km da BR- 316, cruzamento à direita a 20 km de Araguaná, 3 ° 04 ’ 48 ” S, 45 ° 45 ’ W, 13 May 1979 (st.), J. Jangoux 587 & R. P. Bahia (NY!). Santa Luzia, Faz Agripec-Varig, caminho para o Rio Mutum, margem esquerda do Rio Pindaré, 3 ° 7 ’ 48 ” S, 45 ° 39 ’ 29 ” W, 05 April 1983 (fl.), M. F. F. Silva 1008 (INPA, image); Km 20 da estrada Santa Luzia / Santa Inez, 3 ° 57 ’ 48 ” S, 45 ° 39 ’ 29 ” W, 29 July 1977 (fl.), D. Thomaz 406 (PEUFR!). São Luís, Reserva Florestal do Sacavém, 2 ° 30 ’ S, 44 ° 15 ’ W, 07 May 1992 (fl.), F. H. Muniz 95 (INPA, image); Rio Anil, 2 ° 31 ’ 14 ” S, 44 ° 17 ’ 47 ” W, 14 May 1949 (fl.), R. L. Fróes 24281 (IAN!); Granja Federal, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 12 May 1949 (fl.), R. L. Fróes 24243 (IAN!). Additional specimens examined: — BRAZIL. Amazonas: Manaus: Manaus - Porto Velho Highway, Km. 25, 4 ° 46 ’ 06 ” S, 61 ° 19 ’ 26 ” W, 19 March 1974 (fr.), G. T. Prance 20735 (NY!). Pará. Serra dos Carajás, 475 - 525 m, 5 ° 48 ’ S, 50 ° 33 ’ W, 07 June 1982 (fr.), C. R. Sperling 5952 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46BEE6DA7E885ADFBF0A6F9.taxon	materials_examined	Type: — BRAZIL. C. Martius 47 (holotype: BR; isotypes: MO [313601], P [K 000001445]). (Fig. 5 O)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46BEE6DA7E885ADFBF0A6F9.taxon	description	Trees 2 – 6 m. Twigs pubescent, the indumentum whitish. Leaves with petioles 1 – 2 mm long, pubescent, channeled; blades elliptic or lanceolate, 1 – 2.5 × 0.5 – 1.2 cm, glabrous, chartaceous, discolorous; glands inconspicuous; base cuneate; apex acute; midvein adaxially flat; lateral veins few visible, not arched, 10 to 26 pairs, intra-marginal vein 0.5 mm or less from the margin, margin flat. Inflorescence glomerules clustered in axils, the axis inconspicuous; bracts deltate, 0.5 × 0.5 mm, glabrous, caducous after anthesis; pedicels absent; bracteoles deltate, 0.5 × 0.5 mm, glabrous, caducous after anthesis; flower buds ovate, 1.5 × 1.5 mm, not constricted above ovary; calyx-lobes free, four, glabrous, ovate, 1 × 1 mm, not reflexed; the fruit falling as a unit together with the calyx, corolla and hypanthium; ovary 2 - locular. Immature fruit globose, 0.5 cm diameter, glabrous, surface with glands marked, no mature fruits examined.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46BEE6DA7E885ADFBF0A6F9.taxon	discussion	Note: — Myrciaria tenella may be recognized by its elliptic or lanceolate leaves with acute leaf apex, the few visible lateral veins, the inflorescence glomerules clustered in leaf axils, and the fruit falling as a unit together with the calyx, corolla and hypanthium. This species is related to M. floribunda, from which it is distinguished by leaves 1 – 2.5 × 0.5 – 1.2 cm (vs. 4.4 – 7.3 × 2.8 – 3 cm), leaf apex acute (vs. long-acuminate) and petiole with 1 – 2 mm (vs. 2 – 4 mm).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46BEE6DA7E885ADFBF0A6F9.taxon	distribution	Distribution and habitat: — Known from Argentina, Bolivia, Dominican Republic, French Guiana, Haiti, Paraguay, Peru, Uruguay and Venezuela and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, M. tenella was found growing in secondary and “ terra firme ” forests.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46BEE6DA7E885ADFBF0A6F9.taxon	biology_ecology	Phenology: — Flowering in May and August and fruiting in August.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46BEE6DA7E885ADFBF0A6F9.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Colônia Três Satubas, 16 May 1979 (fl.), I J. Jangoux & R. P. Bahia 639 (MG). Rio Alto Turiaçú, Nova Esperança, 07 December 1978 (fl.), I J. Jangoux & R. P. Bahia 288 (MG!). Rio Gurupi, 13 August 1958 (fl., fr), R. L. Fróes 34525 (IAN!). Monção, Guajá, Rio Turiaçu, 3 ° 07 ’ 12 ” S, 46 ° 04 ’ 48 ” W, 24 April 1887 (st.), W. L. Balée 3343 (NY!); 3 ° 07 ’ 12 ” S, 46 ° 04 ’ 48 ” W, 25 June 1987 (st.), W. L. Balée 3486 (NY!); 3 ° 07 ’ 12 ” S, 46 ° 04 ’ 48 ” W, 29 June 1988 (st.), W. L. Balée 3514 (NY!); Lago do Junco, Fazenda Bacaba, 4 ° 25 ’ 48 ” S, 44 ° 58 ’ 12 ” W, 05 October 1980 (fr.), D. C. Daly D 493 (NY!). São Luís, Base do Carbral, 06 January 1988 (st.), R. Nina 39 (SLUI!); Reserva Florestal do Sacavém, 2 S ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 07 May 1992 (fr.), F. H. Muniz 276 (UEC, image); Rio Anil, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 10 May 1949 (fl.), R. L. Fróes 24211 (IAN!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E886E9FA82A505.taxon	materials_examined	Type: — BRAZIL. “ Provinciae Rio Negro ”, C. F. P. Martius 2931 (holotype: M [0032369]). (Fig. 6 H-J)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E886E9FA82A505.taxon	description	Shrubs ca. 2 m. Twigs glabrous. Leaves with petioles 0.5 – 1 mm long, densely pubescent, unchanneled; blades elliptic to broadly so, 3.9 – 6 × 2.9 – 3 cm, glabrous, membranaceous or chartaceous, discolorous; glands conspicuous; base attenuate or rounded; apex acute, apiculate; midvein adaxially sulcate; lateral veins visible, arched, 5 to 7 pairs; intramarginal vein absent, margin flat. Inflorescence axillary, a solitary flower, dichasia, or rarely an auxotelic raceme, the axis inconspicuous, glabrous; bracts not seen, caducous before anthesis; pedicels 2.1 – 5.3 mm long; bracteoles not seen, caducous before anthesis; flower buds globose, apiculate, 4 – 9 × 6 – 7 mm, constricted above ovary; calyx-lobes completely fused, four to five, opening by irregular tearing, glabrous, ovate, 0.5 × 0.5 mm, not reflexed, persisting in the fruit; ovary not seen. Immature fruit globose, 1 – 2 × 1.5 – 2.3 cm, glabrous, surface marked with glands.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E886E9FA82A505.taxon	discussion	Note: — Psidium acutangulum may be recognized by its glabrous, elliptic to broadly elliptic leaves with an apiculate leaf apex; a solitary flower, or sometimes with an auxotelic raceme; the four to five calyx-lobes opening by irregular tearing; and the globose fruits which are glabrous and have the surface marked with glands.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E886E9FA82A505.taxon	distribution	Distribution and habitat: — Known from Bolivia, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela and central, northeastern and northwestern Brazil (Acre, Amazonas, Amapá, Maranhão, Mato Grosso, Pará, Rondônia, Roraima, and Tocantins states) (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, P. acutangulum was found growing in flooded forest and along rivers.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E886E9FA82A505.taxon	biology_ecology	Phenology: — Flowering in February and October and fruiting in November.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E886E9FA82A505.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Maracassumé River Region, 08 October 1932 (fl.), R. L. Fróes 1927 (NY!). Monção, Urutawy, Ka’ apor Indian Reserve, basin of Rio Turiaçu, 05 November 1986 (fl), W. L Balée & B. G. Ribeiro 2904 (MG!). Pinheiro, Rio Alto Turiaçu, Nova Esperança, 100 m, 2 ° 55 ’ 12 ” S, 45 ° 45 ’ W, 29 November 1978 (fr.), J. Jangoux 78 & R. P. Bahia (NY!). São Luis, Alumar, 2 ° 31 ’ 47 ” S, 44 ° 18 ’ 10 ” W, 20 February 1985 (fl.), M. Salgado & R. M. G. Cardoso 69 (IAN!). Additional specimens examined: — BRAZIL. Acre: Sena Madureira, Riozinho do Andirá, colocaçao Curitiba, 9 ° 43 ’ 45 ” S, 68 ° 08 ’ 53 ” W, 10 June 1995 (fl.), A. R. S. de Oliveira 572 (NY!). Pará. Oriximiná, Rio Paru do Oeste, Cachoeira Chuvisco, 0 ° 59 ’ 58 ” S, 56 ° 04 ’ 42 ” W, 07 November 1980 (fl.), C. A. C. Ferreira 2247 (NY!).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E882F9FF5FA96D.taxon	materials_examined	Type: — “ Culta in Hispaniola, ” “ ex Africa ”. Presumably Swartz s. n. (holotype: S-r- 5302). (Fig. 5 P)	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E882F9FF5FA96D.taxon	description	Shrubs 2 m. Twigs sparsely pubescent or pubescent, indumentum brownish-red. Leaves with petioles 0.5 – 0.7 mm long, densely or sparsely pubescent, channeled; blades elliptic, elliptic-oblong, oblong or oval, 4 – 13.5 × 1.8 – 6.8 cm, densely pubescent, the indumentum reddish-brown, chartaceous, discolorous; glands not seen; base cuneate or rounded; apex acute, mucronate or obtuse; midvein adaxially impressed; lateral veins visible, arched, 7 to 10 pairs; intra-marginal vein absent, margin flat. Inflorescence axillary, a solitary flower or dichasium, 1 – 3 flowers, the axis 0.9 – 1.2 mm long, pubescent; bracts not seen, caducous before anthesis; pedicels 0.5 – 0.7 mm long or absent; bracteoles linear, 1 × 1 mm, pubescent, caducous after anthesis; flower buds pyriform, 4 – 5 × 7 – 9 mm, constricted above ovary; calyx-lobes completely fused, four to five, opening by irregular tearing, pubescent, ovate, 5 × 3 – 5 mm, not reflexed, persisting in the fruit; ovary 3 – 5 - locular. Immature fruit globose, ca. 1.1 × 1.4 cm, glabrous, surface marked with glands.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E882F9FF5FA96D.taxon	discussion	Note: — Psidium guineense may be recognized by its densely or sparsely pubescent, elliptic, oblong or oval leaves; the acute, mucronate or obtuse leaf apex; the inflorescence, which is a solitary flower or dichasium; and the completely fused calyx lobes, opening by irregular tearing.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E882F9FF5FA96D.taxon	distribution	Distribution and habitat: — Known from Tropical America, and widely distributed in Brazil (POWO 2020, Flora do Brasil 2020). In Amazonian Maranhão, P. guineense was found growing in flooded forests and along roads.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E882F9FF5FA96D.taxon	biology_ecology	Phenology: — Flowering in January, February and December.	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
C41B87B7C46AEE6CA7E882F9FF5FA96D.taxon	materials_examined	Specimens examined: — BRAZIL. Maranhão: Anajatuba, São Bento, 3 ° 15 ’ 52 ” S, 44 ° 37 ’ 11 ” W, 09 February 2008 (fl.), A. Araujo 10 (IAN!). Pameirândia, 2 ° 38 ’ 43 ” S, 44 ° 53 ’ 42 ” W, 28 January 2007 (fl.), C. M. Vieira 87 (IAN!). Turiaçu, Km 6 da BR 106 Maracaçumé-Sta. Helena, fazenda Maracaçumé Agro Industrial Grupo Mesbla, 1 ° 39 ’ 48 ” S, 45 ° 22 ’ 18 ” W, 01 December 1978 (fl.), N. A. Rosa 2819 & H. Vila (NY!). Additional specimens examined: — BRAZIL. Pará: Vigia, 17 km southeat of Vigia along road (PA- 140) to Belém, 50 m, 0 ° 57 ’ S, 48 ° 04 ’ 59 ” W, 30 March 1980 (fr.), G. Davidse, N. A. Rosa, C. S. Rosario & M. G. Silva 17655 (US, image).	en	Amorim, Gabriela Dos Santos, Barbosa, Maria Regina De Vasconcellos, Jr, Eduardo Bezerra De Almeida, Thomas, William Wayt (2022): Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil. Phytotaxa 568 (1): 27-60, DOI: 10.11646/phytotaxa.568.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.568.1.3
