identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
911BE60E5548FFE78CC5FEA1FEBB83C9.text	911BE60E5548FFE78CC5FEA1FEBB83C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Longitermes JOUAULT, ENGEL, LEGENDRE, NEL 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> GENUS  LONGITERMES JOUAULT, ENGEL, LEGENDRE &amp; NEL GEN. NOV.</p>
            <p>Z o o b a n k r e g i s t r a t i o n: h t t p: / / z o o b a n k. org/ urn:lsid:zoobank.org:act: AB77601A-A6D5-4CB9- 9236-DEC7FF2144B7</p>
            <p> Type species:  Longitermes pulcher Jouault, Engel, Legendre &amp; Nel sp. nov.</p>
            <p> Etymology: The generic name is a combination of the Latin longus, long, referring to the elongate head of the new species, and termes, termite, commonly used to refer to  Isoptera . Gender masculine. </p>
            <p>Diagnosis: Imago. Head elongate, with anterior corners rounded orthogonal and posterior corners broadly rounded; occiput and frons convex; epicranial ecdysial line visible; ocelli absent; postclypeus trapezoidal; anteclypeus small; left mandible apical tooth elongate, sharp, three marginal teeth, LM 1 -LM 2 interdental space small, v-shaped, LM 1 small, LM 2 elongate, forming broad cutting edge, LM 2 -LM 3 separated by a notch, LM 3 bent toward base of mandible; right mandible with long, sharp apical tooth, and at least one marginal tooth of similar shape and length; pronotum slightly wider than head, anterior margin concave, posterior margin broadly convex with a median concavity, sides broadly convex; tarsi pentamerous (fourth tarsomere cryptic); tibial spur formula 3-4-2; all tibiae with strong setae but no additional spines; arolium present; forewing with scale large, with humeral margin nearly straight, claval suture straight (PCu), basal suture faintly arched; Sc, RA, RP heavily sclerotized; R branched within scale; RP nearly encompassing wing apex, with main branch bending to wing apex, with five secondary branches; M and CuA weaker than other surrounding veins; CuA with three branches in scales; anal veins absent; hindwing with delineated scale.</p>
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	https://treatment.plazi.org/id/911BE60E5548FFE78CC5FEA1FEBB83C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jouault, Corentin;Engel, Michael S.;Fls;Legendre, Frédéric;Huang, Diying;Grandcolas, Philippe;Nel, André	Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe, Nel, André (2022): Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera). Zoological Journal of the Linnean Society 196: 608-629, DOI: 10.1093/zoolinnean/zlac064
911BE60E5548FFE08C34F9C4FB7683EB.text	911BE60E5548FFE08C34F9C4FB7683EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Longitermes JOUAULT, ENGEL, LEGENDRE & NEL 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> LONGITERMES PULCHER JOUAULT, ENGEL, LEGENDRE, &amp; NEL SP. NOV. </p>
            <p>(FIGS 1, 2)</p>
            <p>Z o o b a n k r e g i s t r a t i o n: h t t p: / / z o o b a n k. org/ urn:lsid:zoobank.org:act: 6327861A-3757-4810- BEED-9800D20146C2</p>
            <p>Material: Holotype, IGR.BU-052, preserved in an ovoid piece of amber measuring 18 × 12 × 2 mm, housed in the amber collection of the Geological Department and Museum (IGR), Rennes, France.</p>
            <p>Locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Upper Albian to Lower Cenomanian, ‘Mid’-Cretaceous.</p>
            <p> Etymology: The specific epithet is from the Latin adjective  pulcher , beautiful. </p>
            <p>Diagnosis: As for the genus (vide supra).</p>
            <p> Description: Imago specimen. Body c. 7.34 mm long (as preserved and measured from tip of labrum to abdominal apex). Head elongate, c. 1.44 mm long (measured from posterior margin to tip of labrum) and c. 0.82 mm wide excluding compound eyes (measured beyond eyes), trapezoidal in dorsal view, prognathous, sides slightly convex, postclypeus c. 0.11 mm long, anteclypeus c. 0.11 mm long, labrum lobe-shape, sides nearly parallel, slightly narrower basally, apical margin nearly straight, with few setae; mandibles triangular shaped, c. 0.20 mm high, covered by labrum; left mandible apical tooth elongate, sharp; three marginal teeth, LM 1 small, LM 1 -LM 2 interdental space small, v-shaped, LM 2 elongate, forming broad cutting edge, LM 2 -LM 3 separated by a notch, LM 3 bend toward base of mandible (general aspect similar to left imago mandible of  Hodotermopsis ); right mandible with long, sharp apical tooth, and at least one marginal tooth of similar shape and length; buccal pieces: glossa with two visible segments, paraglossa with three segments, length from base to apex (in mm) 0.05, 0.06, 0.1; lacinial incisor with two teeth; five maxillary palpomeres visible, length from base to apex (in mm): 0.06, 0.05, 0.08, 0.09, 0.12; three labial palpomeres present, total length as preserved c. 0.21 mm; compound eyes, c. 0.37 mm long, ovoid, situated laterally near head midlength, and separated from posterior head margin by more than their length; ocelli absent; fontanelle absent; antennae moniliform with 14 antennomeres. Pronotum c. 0.54 mm long and c. 1.0 mm wide, nearly flat with anterior margin concave, anterior corners arched, sides broadly convex and posterior margin convex with a medial indentation, posterior corners broadly arched. </p>
            <p>Legs robust; profemur medially swollen, c. 0.63 mm long, protibia c. 0.37 mm long, protarsus c. 0.22 mm long; protibia with three spurs f1, f2, f3; mesofemur medially swollen c. 0.63 mm long, mesotibia c. 0.42 mm long, mesotarsus c. 0.22 mm long; mesotibia with four observable spurs m1, m2, m3, m4; metafemur c. 0.51 mm long, metatibia c. 0.52 mm long, with two observable spurs h1, h2, mesotarsus c. 0.25 mm long; all tarsi pentamerous, with arolium. Forewing scale c. 0.80 mm long and 0.54 mm wide (measured distally), RP with additional tertiary branches joining costal margin; median (M) weak, closer to cubitus (Cu) than to radial sector (RP), with five branches terminating at lower margin; CuA with six primary and secondary branches extending to lower margin; anal vein absent. Hindwing (folded or damage): humeral suture weak, barely visible; wing venation apparently similar to forewing except anal vein present.</p>
            <p>Abdomen at least 4.0 mm long (damaged during the fossilization process) with at least ten observable segments; abdominal segments apparently slightly wider near midlength. Cerci multi-merous c. 0.16 mm long, five(?) cercomeres (sometimes fused so not clearly countable).</p>
            <p>Colour: Not preserved.</p>
            <p> Remarks:  Longitermes pulcher is unique among Cretaceous termites, with an unprecedented combination of diagnostic characters making it close to the family  Archotermopsidae , as treated in Jiang et al. (2021), i.e. encompassing  Archotermopsis ,  Hodotermopsis and  Zootermopsis . Here we follow the new system of Wang et al. (2022: table 1), which recently resolved paraphyly of  Archotermopsidae by elevating Hodotermopsinae to family rank. In fact,  L. pulcher shares with  Archotermopsidae and  Hodotermopsidae the lack of a fontanelle; a Y-shaped ecdysial line present; large compound eyes; ocelli absent; pronotum faintly arched, slightly wider than head; left mandible with an apical tooth and three marginal teeth. Legs: five tarsomeres (some cryptic); tibial spur formula 3-4-2; arolium present. Cerci with five(?) cercomeres. Forewing: costal margin flat to faintly arched; humeral suture well defined, almost straight; Sc, RA, RP sclerotized, RP with the main branch bending to apex of wing, five to six secondary branches and additional tertiary branches joining costal margin; M weak, with secondary branches terminating at lower margin; Cu with primary and secondary branches extending to lower margin; anal vein absent. The wing venation with the radial field encompassing most of the wing apex and the shape of the left mandibular teeth, nearly similar to that of  Hodotermopsis , strengthen the taxonomic placement. However, the phylogenetic position of the new genus is hard to ascertain and it may be a stem representative of  Hodotermopsidae or of  Archotermopsidae (sensu Wang et al., 2022) . Since the new specimen differs from the genera  Archotermopsis and  Zootermopsis (  Archotermopsidae sensu Wang et al., 2022 ), at least based on its mandibular shape with LM 2 elongate and forming a broad cutting edge (vs. sharp and not forming a cutting edge in the aforementioned genera) (Krishna et al., 2013), and resembles more the genus  Hodotermopsis in its wing venation and mandibular configuration, we presume that  Longitermes is probably a stem-Hodotermopsidae (sensu Wang et al., 2022).  Hodotermopsis differs from the new specimen in possessing small eyes, while they are large in  Longitermes , but also in having numerous antennomeres (at least 23), while the new specimen only possesses 14 (the antenna might not be complete but the number of antennomeres seems to be lower than in  Hodotermopsis ) (Krishna et al., 2013). Recently, several families were erected based on Cretaceous specimens from Kachin amber (e.g. Melqartitermitidae and Mylacrotermitidae) but our specimen cannot be placed in any of them (Jiang et al., 2021). Its broad radial field precludes affinities with the Melqartitermitidae and the presence of a welldefined forewing basal suture demarcating the scale precludes affinities with Mylacrotermitidae (Jiang et al., 2021). Similarly, the new specimen differs from representatives of the family  Krishnatermitidae , at least, because of its forewing with PCu straight (vs. arched in  Krishnatermes ), RP encompassing the wing apex (variable in  Krishnatermes ) and its hindwing with a defined scale (vs. absent in  Krishnatermes ). However, the habitus of  Longitermes and  Krishnatermes are at first sight similar, which, if not symplesiomorphic, may indicate a proximity between the two genera and a hypothetical proximity of  Krishnatermes with the Teletisoptera. </p>
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	https://treatment.plazi.org/id/911BE60E5548FFE08C34F9C4FB7683EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jouault, Corentin;Engel, Michael S.;Fls;Legendre, Frédéric;Huang, Diying;Grandcolas, Philippe;Nel, André	Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe, Nel, André (2022): Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera). Zoological Journal of the Linnean Society 196: 608-629, DOI: 10.1093/zoolinnean/zlac064
911BE60E554FFFE08F98F957F9BF85F0.text	911BE60E554FFFE08F98F957F9BF85F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisotermes ZHAO	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> GENUS  ANISOTERMES ZHAO ET AL., 2019 </p>
            <p> Included species:  Anisotermes xiai Zhao, Eggleton &amp; Ren 2019 (type species) and  A. bourguignoni Jouault, Engel, Huang &amp; Nel sp. nov.</p>
            <p> Emended diagnosis: Head approximately rounded; ocelli absent; fontanelle absent; Y-shaped ecdysial line absent; mandible not exceeding labrum, with la and ra conspicuously longer than LM1 and RM1; antenna moniliform, with about 25 antennomeres, terminal eight antennomeres tapered; compound eyes hemispheric, lying at head midlength; pronotum definitely saddle-shaped, slightly wider than head in dorsal view; lateral margin weakly upcurved; procoxal carina present; additional tibial spines present on all legs; tibial spur formula 3-4-4; tarsi wholly pentamerous; arolium variable (absent in  A. xiai but present in  A. bourguignoni ); wings membranous, densely reticulate; forewing scale large, overlapping hindwing base, basal suture convex, veins Sc, RA, RP and M more heavily pigmented than CuA. Forewing: all major veins originate within scale; Sc simple or double; RP with five to six main branches, terminating on costal margin anterior to wing apex; radial field of moderate width, parallel to costal margin; M running about midway between RP and CuA, terminating on posterior margin; medial field relatively narrow; CuA branched, terminating about same distance with Rs to wing apex; PCu (claval suture) arched, meeting basal suture on posterior margin. Hindwing without basal suture, large anal lobe present; M first inferior branch fading away, terminating in center of hindwing; cerci short, trimerous or tetramerous (with additional indented rings); abdominal styli absent in female but present in male. </p>
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	https://treatment.plazi.org/id/911BE60E554FFFE08F98F957F9BF85F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jouault, Corentin;Engel, Michael S.;Fls;Legendre, Frédéric;Huang, Diying;Grandcolas, Philippe;Nel, André	Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe, Nel, André (2022): Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera). Zoological Journal of the Linnean Society 196: 608-629, DOI: 10.1093/zoolinnean/zlac064
911BE60E554FFFEC8C02FB96FEC68424.text	911BE60E554FFFEC8C02FB96FEC68424.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisotermes bourguignoni Jouault, Engel, Huang & Nel 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> ANISOTERMES BOURGUIGNONI JOUAULT, ENGEL, HUANG &amp; NEL SP. NOV. </p>
            <p>(FIGS 3–5)</p>
            <p>Z o o b a n k r e g i s t r a t i o n: h t t p: / / z o o b a n k. org/ urn:lsid:zoobank.org:act: A406BDD6-D799-4DB8- 93D0-4C2E61C71F1A</p>
            <p>Holotype: Specimen identifier NIGP177751 (a complete dealate specimen preserved in a rounded amber piece), housed in the collection of the Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences (NIGPAS), Nanjing, China.</p>
            <p>Etymology: The specific epithet honours Dr Thomas Bourguignon for his work on termite evolution. It is to be treated as a noun in the genitive case.</p>
            <p>Locality and age: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Late Albian to Early Cenomanian, ‘Mid’-Cretaceous.</p>
            <p>Diagnosis: Imago. Head rounded, with anterior corners slightly orthogonal and posterior corners rounded; left mandible with la longer than LM 1, both sharp; right mandible with an apical and two marginal teeth, ra shorter than RM 1, RM 1 and RM 2 separated by a sharp angle; occiput and frons convex; postclypeus trapezoidal; anteclypeus small; ocelli absent; pronotum massive, trapezoidal, wider than head, anterior margin concave surrounding head, posterior margin broadly convex, with dorsal surface flatly arched in profile, sides convex; tarsi pentamerous; tibial spur formula 3-4-4; protibia with two additional spine (fa, fb); mesotibia with six additional spines (ma-mf); metatibia with at least three additional spines (ha-hc); arolium present; forewing with scale large, with humeral margin welldefined and slightly arched, claval suture strongly curved (PCu), basal suture convex; Sc, RA, RP and CuA heavily sclerotized; Sc branched within scale and going outside of scale; R branched within scale; M weaker than other surrounding veins; CuA with three branches in scales; PCu strongly curved; anal veins not discernible within scale.</p>
            <p>Description: Imago. Body c. 10 mm long (from tip of labrum to abdomen apex). Head robust, c. 2.20 mm long (measured from under the pronotum to the tip of the labrum) and c. 1.93 mm wide excluding compound eyes, square-shaped in dorsal view, hypognathous, sides slightly convex, anterolateral corner slightly orthogonal, posterolateral corners rounded, Y-shaped ecdysial scar absent; tentorial pit visible (ttp); postclypeus (pc) 0.32 mm long, trapezoidal; labrum c. 0.50 mm long, lobe-shape, narrower basally; mandibles massive, triangular shaped, at least 0.53 mm high; buccal pieces with glossa (gl), paraglossa (gl), stipe (st), mentum (mt), submentum (sm) visible but not clearly describable; five maxillary palpomeres, combined length as preserved c. 1.2 mm; four(?) labial palpomeres present, with three discernible palpomeres, combined length c. 0.75 mm; compound eyes, c. 0.45 mm long, circular, situated laterally near head midlength, and separated from posterior head margin by more than their length; fontanelle not visible and probably absent; antenna moniliform with at least 22 antennomeres. Pronotum c. 1.60 mm long marginal teeth (number corresponding); sm, submentum; st, stipe; ttp, tentorial pit. Scale bars 0.5 mm (A, C, D); 1 mm (B). and c. 2.73 mm wide, overlapping head dorsally, sides slightly upturned.</p>
            <p>Legs robust; procoxa c. 1.38 mm long, profemur c. 1.51mm long, protibia c. 1.41mm long, protarsus at least 0.76 mm long; protibia with three spurs f1, f2, f3 and additional spines; mesocoxa c. 1.13 mm long, mesofemur medially swollen c. 1.57 mm long, mesotibia c. 1.72 mm long, mesotarsus c. 0.64 mm long; mesotibia with four observable spurs m1, m2, m3, m4 and additional spines; metacoxa at least 1.10 mm long, metafemur at least 2.24mm long, metatibia at least 1.50 mm long, with three observable spurs h1, h2, h3, h4 and additional spines; all spurs flattened with serration; all tarsi pentamerous; pretarsal arolium present. Forewing scale c. 2.36 mm long and 1.41 mm wide (measured distally), sclerotized, especially in anal area, with sparse setation. Hindwing scale not visible but proximal part of hindwing clearly cut beyond basal suture.</p>
            <p>Abdomen at least.6.0mm long with at least11observable segments; abdominal segments widest at midlength. Cerci multimerous c. 0.52 mm long, five to seven cercomeres (sometimes fused so not clearly countable).</p>
            <p>Colour: Not preserved.</p>
            <p> Remarks:  Anisotermes bourguignoni differs from most other Cretaceous ‘lower’  Isoptera in having a plesiomorphic character in scale configuration (e.g. fine reticulation between veins in the scale), tarsi pentamerous (plesiomorphic character), a tibial spur formula of 3-4-4 and an overall massive trapezoidal-shaped pronotum. The wholly pentamerous tarsi and enriched tibial spur formula tend to imply an early diverged position within  Isoptera , among groups like  Archotermopsidae ,  Hodotermitidae ,  Mastotermitidae and their various extinct relatives. Additionally, its general habitus, and particularly its large pronotum, suggests that  A. bourguignoni is related to  Mastotermitidae or to taxa belonging to the ‘  Meiatermes -grade’. Interestingly, the hindwings of  A. bourguignoni were cut slightly distal from the hindwing basal scale suture, while in the forewing the cut follows the basal suture. The hindwing basal scale suture was probably not fragile enough to break alone after the mating swarm, as it normally happens in a majority of extant termites. This suggests that the suture of the hindwing is rudimentary (symplesiomorphy) as in  Mastotermes darwiniensis (Tillyard, 1931) , and that the new specimen could be related to  Mastotermitidae . Another character suggesting placement of  Anisotermes within  Mastotermitidae is the mandibular shape. In fact,  Anisotermes and  Mastotermes exhibit a similar configuration of teeth on both mandibles, and length variations (RM 1 conspicuously longer than ra vs. equal in length) are usually indicative of different genera from the same family. Additionally, this placement is strengthened by the diagnostic characters of the new species: ocelli absent, procoxal carina present, arolium present and the aforementioned scale characters. The absence of ocelli indicates affinities with  Idanotermitinae , but the new fossil differs from the genus  Idanotermes Engel, 2008 in at least its different tibial spur formula (i.e. 3-4-4 vs. 3-4-3). The genus  Anisotermes is here tentatively considered as a member of the  Idanotermitinae (Jiang et al., 2021: 379); we confirm this placement mainly based on the presence of the anal lobe (characteristics of  Mastotermitidae within the  Isoptera ) and based on the absence of ocelli. The new species differs from  A. xiai by, inter alia, the presence of the pretarsal arolium and more additional spines along the tibiae. </p>
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	https://treatment.plazi.org/id/911BE60E554FFFEC8C02FB96FEC68424	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jouault, Corentin;Engel, Michael S.;Fls;Legendre, Frédéric;Huang, Diying;Grandcolas, Philippe;Nel, André	Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe, Nel, André (2022): Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera). Zoological Journal of the Linnean Society 196: 608-629, DOI: 10.1093/zoolinnean/zlac064
911BE60E5543FFEC8C03FD49FF61817A.text	911BE60E5543FFEC8C03FD49FF61817A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Magnifitermes KRISHNAI JOUAULT, ENGEL & NEL 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> GENUS  MAGNIFITERMES JOUAULT, ENGEL &amp; NEL GEN. NOV.</p>
            <p>Zoobank registration: http://zoobank.org/urn:lsid: z o o b a n k. o r g:a c t: B D E A7C 1 1- 0 2 9 6- 4 7F 8- A7 3 5 - 817A4D58F844</p>
            <p> Etymology: The generic name is a combination of the Latin adjective magnificus, magnificent, and the noun termes, meaning ‘maggot’ or ‘termite’, and commonly used as the root for many genera of  Isoptera . Gender masculine. </p>
            <p> Type species:  Magnifitermes krishnai Jouault, Engel &amp; Nel sp. nov.</p>
            <p>Diagnosis: Imago. Head massive, square-shaped, dorsally flattened, with posterior corners broadly rounded; epicranial scar present; ocelli present, located just above compound eyes (not separated from eyes by a space greater than their length); postclypeus trapezoidal and flat; anteclypeus anteriorly convex; left mandible with an apical and two distinct marginal teeth, LM 1 -LM 2 interdental space broad, v-shaped, posterior margin of LM 1 slightly longer than anterior margin LM 2; pronotum massive, trapezoidal, wider than head, anterior margin concave surrounding head, posterior margin broadly convex, with dorsal surface flatly arched in profile, sides convex; tarsi pentamerous, with fifth tarsomere extremely elongate; tibial spur formula 3-4-4; protibia with two additional spines (fa, fb); mesotibia with four(?) additional spines (ma, mb, mc, md); metatibia with at least three additional spines (ha, hb, hc); pretarsal arolium present; forewing scale large, overlapping hindwing scale, claval suture (PCu) broadly arched, basal suture broadly arched; Sc, RA, RP, M heavily sclerotized; Sc with two main branches; R branched within scale; RP running parallel to costal margin, with at least five branches joining costal margin; M slightly closer to RP than to CuA, with at least four secondary branches radiating to apical margin; CuA less sclerotized, with several branches joining lower margin. Hindwing: scale (basal) suture absent; anal lobe (ano-jugal) developed (folded)</p>
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	https://treatment.plazi.org/id/911BE60E5543FFEC8C03FD49FF61817A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jouault, Corentin;Engel, Michael S.;Fls;Legendre, Frédéric;Huang, Diying;Grandcolas, Philippe;Nel, André	Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe, Nel, André (2022): Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera). Zoological Journal of the Linnean Society 196: 608-629, DOI: 10.1093/zoolinnean/zlac064
911BE60E5546FFEB8FF0F907FF1B8168.text	911BE60E5546FFEB8FF0F907FF1B8168.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mastotermes myanmarensis Jouault & Engel & Fls & Legendre & Huang & Grandcolas & Nel 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> MASTOTERMES MYANMARENSIS JOUAULT  SP. NOV.</p>
            <p>(FIGS 9, 10)</p>
            <p>Z o o b a n k r e g i s t r a t i o n: h t t p: / / z o o b a n k. o r g / urn:lsid:zoobank.org:act: F5CF600D-2068-424E-B255- 9F92051F952B</p>
            <p> Material:   Holotype, IGR.BU-054, preserved in an elongate, pale and ovoid piece of amber measuring 50 × 20 × 9 mm, housed in the amber collection of the  Geological Department and Museum (IGR), Rennes, France. </p>
            <p>Locality and horizon: Hkamti site, Hkamti District, Sagaing Region, Myanmar; Early Albian (c. 110 Mya), Early Cretaceous.</p>
            <p>Etymology: The specific epithet refers to the country of origin of the amber piece, Myanmar, combining the country name with the Latin toponymic adjectival suffix, – ēnsis, meaning ‘from’.</p>
            <p>Diagnosis: Imago. Wings membranous, long and broad, densely reticulate, with ‘cross-veins’ present; veins Sc, RA, RP and M more heavily sclerotized than CuA, reticulate veins obviously pigmented among radial and medial fields; forewing Sc with two main branches; R with two branches; width of radial field moderate, RA simple, RP with five main branches; medial field encompassing wing apex, M main vein closely parallel to RP main vein, with three main branches, first branch of M arising near apical-third of wing; CuA with five main branches, apical-most branch of CuA terminating into posterior margin and in apical-third of forewing length. Hindwing with large anal lobe; costal space wide; Sc simple, terminating on costal margin at basal-third of hindwing length; RA long, simple and terminating on costal margin before apical-third of hindwing length; RP with two main branches; medial field encompassing wing apex; M separating from RP near wing base, relatively closely parallel to RP, with three main branches, secondary branches present; CuA uniformly branching, terminating just posterior to wing apex.</p>
            <p>Description: Imago. Wings membranous, reticulate veins present; forewing scale part missing, forewing preserved part at least 10.70 mm long and 4.62 mm wide, apex rounded, middle section of posterior slightly convex basally then convex; veins Sc, RA, RP and M more heavily sclerotized than CuA and A; forewing Sc short, bifurcate at suture, posterior branch with a faint subbranch at margin; RA with one branch; radial field width moderate, occupying about one-sixth area on average; RP with five branches, proximal branch fading apically before anterior wing margin; M closely parallel to RP, with three main branches; CuA with numerous posterior branches, apical-most branch of CuA terminating on posterior margin well proximal to apex, more proximal branches loosely pectinate, apicalmost of these branches terminating on margin near wing midlength.</p>
            <p>Hindwing with large anal lobe, scale not preserved, 14.10 mm long and 6.0 mm wide with anal lobe and 5.16 mm wide without anal lobe; Sc longer than that of forewing; RA long, simple and terminating on costal margin before apical-third of hindwing length; RP with three branches, first branch originating just before point of contact between Sc and wing margin; M with four branches, separating from RP away from wing base, then closely parallel to RP, apical branch of M with a secondary branch; CuA with numerous posterior branches, some dichotomous, terminating posterior to wing apex.</p>
            <p>Colour: Not preserved.</p>
            <p> Remarks: Compared with the known ‘Mid’-Cretaceous fossil and the extant mastotermitids,  M. myanmarensis stands out in having broadly rounded wings, wider intervals between the longitudinal veins (shared with  M. monostichus Zhao et al., 2019 ), and a broad space above Sc on the hindwing. Additionally, the space occupied by Sc + RA + RP is extremely wide near the basal suture.  Mastotermes myanmarensis has a hindwing with vein A1 without branches, while the extant  M. darwiniensis has a vein A1 with numerous posterior branches (similar to  M. monostichus ), and the anal veins in the anal lobe are all simple (vs. with some dichotomies in  M. darwiniensis and  M. monostichus ).  Mastotermes myanmarensis can be differentiated from  M. monostichus by the forewing with RA simple, M with only a reduced number of branches and the first dichotomy far from the wing base, CuA with more dichotomous branches; hindwing with a broad space above Sc, and the anal field with more branches.  Mastotermes myanmarensis also differs from the extant  M. darwiniensis by its temporal range (‘Mid’-Cretaceous vs. extant). </p>
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	https://treatment.plazi.org/id/911BE60E5546FFEB8FF0F907FF1B8168	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jouault, Corentin;Engel, Michael S.;Fls;Legendre, Frédéric;Huang, Diying;Grandcolas, Philippe;Nel, André	Jouault, Corentin, Engel, Michael S., Fls, Legendre, Frédéric, Huang, Diying, Grandcolas, Philippe, Nel, André (2022): Incrementing and clarifying the diversity and early evolution of termites (Blattodea: Isoptera). Zoological Journal of the Linnean Society 196: 608-629, DOI: 10.1093/zoolinnean/zlac064
