taxonID	type	description	language	source
F00D87CAB27A3730FF1A61ABFB82F924.taxon	description	(Figs 1 – 4)	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB27A3730FF1A61ABFB82F924.taxon	diagnosis	Diagnosis. Female: preapical papilla on flagellomere III present posterior teeth slanted, facing the central region of cibarium; anterior teeth arranged in a transverse row; spermathecae also transversally striated, though without the tubular aspect, clearly wider than the individual spermathecal ducts and with sessile terminal knob. Male: preapical papilla on flagellomere III present; gonocoxite without basal tuft of setae; flagellomere I ≥ 405 µm; aedeagal ducts ≤ 440 µm; epandrial lobe ≤ 157 µm; presence of one apical spine and upper external spine at the subapical level in the gonostyle. Female. Head (Fig. 1 A): 367 (363 / 381, n = 2) in length, 328 (330 / 339, n = 2) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 122 (116 / 136, n = 2) in length; eyes 177 (177 / 180, n = 3) in length, 99 (96 / 99, n = 2) in width; interocular distance 148 (133 / 151, n = 2). Pharynx with small teeth (Fig. 1 B). Cibarium (Fig. 1 B) with four posterior teeth well developed, curved towards the central region of the cibarium, eight anterior teeth distributed in a transverse row and eight lateral teeth flanking the posterior teeth; sclerotized area short and triangular; sclerotized arch incomplete. Labrum-epipharynx (LE) (Fig. 1 C): 174 (162 / 168, n = 2); mandible as shown (Fig. 1 D). Apical region of hypopharynx with 18 teeth (Fig. 1 F). Lacinia of maxilla with 11 – 12 external teeth and 16 internal teeth (Fig. 1 E). Antenna (Figs 1 G – I, 2 A – K): flagellomere length: FI 470 (470 / 493, n = 2); FII 237 (228 / 242, n = 2); FIII 209 (228 / 251, n = 2); FXIII 96 (78 / 110, n = 2) and FXIV 67 (67 / 72, n = 2). Ascoids: short posterior spur with a peduncle type of insertion and the anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 1 G); the external ascoids implanted more apical than the internal ones, in FI; presence of preapical papilla on FI – FIII (Fig. 1 G – I); papilla on FIV – FVI absent (Figs 2 A – C); presence of one papilla on FVII, FVIII and FIX (Figs 2 D – F); two papillae on FX (Fig. 2 G), presence of three, three, seven and six papillae distributed on FXI, FXII, FXIII and FXIV, respectively (Figs 2 H – K); preapical spiniform papilla observed on FIX (Fig. 2 F), FXI (Fig. 2 H), and FXIII (Fig. 2 J). Presence of simple setae on FIV – FXIV (Figs 2 A – K), Labial suture complete (Fig. 1 A). Palpi (P) (Fig. 1 J): palpal segment length: PI 35 (38 / 41, n = 2), PII 61 (55 / 61, n = 2), PIII 128 (116 / 125, n = 2), PIV 75 (70 / 72, n = 2) and PV 145 (125 / 148, n = 2) – Palpal formula: 1 - 2 - 4 - 3 - 5; PIII (Fig. 1 J) with Newstead’s sensilla scattered along the segment. Cervix: ventro-cervical sensillae absent. Cervical sclerite bearing a pair of spiniform sensilla. Thorax: Mesonotum 599 (577, n = 1) in length. Mesonotum, pronotum, paratergite, anepisternum, metanotum and postnotum light brown, pleura off-white. Two proepimeral setae; 13 upper anepisternal setae. Setae present on the anterior margin of the katepisternum. Wing (Fig. 2 L): 2,294 (2,183 / 2,294, n = 2) in length, 664 (657 / 672, n = 2) in width; veins: R 5 1,350 (1.295 / 1,350, n = 3); alfa 518 (577 / 613, n = 2); beta 372 (298 / 311, n = 2); gamma 212 (136 / 200, n = 2); delta 96 (154 / 261, n = 2); pi 95 (66 / 125, n = 2). Legs (anterior; median; posterior): Coxae: 349 (339 / 363, n = 2), 353 (325 / 339, n = 2), 330 (316 / 335, n = 2); Femur: 876 (832 / 888, n = 2), 832 (814 / 832, n = 2), 943 (925, n = 1); Tibia: 1,369 (1,239 / 1,295, n = 2), 1,480 (1,332 / 1,350, n = 2), 1,628 (1,535, n = 1); Tarsomere I: 803 (740, n = 2), 869 (795 / 851, n = 2), 906 (851, n = 1). Sum of tarsomeres II + III + IV + V, 744 (715 / 730, n = 2), 759 (744 / 766, n = 2), 773 (788, n = 1). Abdomen: 1,480 (1,424 / 1,776, n = 2) in length; tergal papillae absent. Spermathecae: 43 (41 / 43, n = 2) in length, 14 (14, n = 2) in width; with poorly defined rings (Fig. 2 M); Terminal knob: 12 (9 / 12, n = 2) in length and 12 (12, n = 2) in width, not individualized, with rounded shape and bristles in the apical region; common spermathecal duct: 70 (58 / 61, n = 3) in length and 6 (6, n = 2) in width, does not go beyond the middle of the genital fork; individual spermathecal duct: 162 (128 / 142, n = 2) in length and 6 (6, n = 2) in width, they are membranous with smooth walls and their widths are uniform throughout their length (Fig. 2 M). Cercus: 154 (154 / 159 n = 2) in length and 78 (69 / 78, n = 2) in width.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB27A3730FF1A61ABFB82F924.taxon	materials_examined	Material examined: 3 females (Nº 87550, 87016, 87549). Capture location: municipality of Lassance (17 ° 53 ′ 22.31 ″ S, 44 ° 34 ′ 53.63 ″ W), located in the state of Minas Gerais, Brazil. Collection date: 2 females (87549, 87550) 08 / ix / 2008; 1 female (87106): 12 / viii / 2008. Trap: HP trap. Collector: Mariana Campos das Neves Farah Ramos. Complementary material: 2 females from “ Parque Estadual do Rio Doce ” (19 º 30 ′ 45 ″ S, 42 º 33 ′ 10 ″ W), state of Minas Gerais, Brazil. Collection date: from June 2003 to July 2004. Trap: Malaise trap. Collector: Rogério Parentoni Martins. Material deposited at FIOCRUZ / COLFLEB.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB27A3730FF1A61ABFB82F924.taxon	description	Redescription of the Male. Head (Fig. 3 A) 335 (321 / 325, n = 2) in length, 270 (270 / 281, n = 2) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 122 (110 / 121, n = 2) in length; eyes 154 (151 / 154, n = 2) in length, 72 (72 / 84, n = 2) in width; interocular distance 125 (122 / 136, n = 2). Pharynx with streaks and no teeth (Fig. 3 B). Cibarium without teeth (Fig. 3 B). Labrum-epipharynx (LE) 136 (136 / 148, n = 2) (Fig. 3 C). Specimen with damaged (broken) antennas, description based on the other specimen (Nº 92113). Antenna (Fig. 3 D – F, 4 A – K): flagellomere length (F): FI 437 (404 / 469, n = 2), FII 227 (223 / 228, n = 2), FIII 223 (209 / 223, n = 2), FXIII 78 (81, n = 1) and FXIV 70 (67, n = 1). Ascoids not visible in specimen, description based on the other specimen (Nº 92113). Ascoids with short posterior spur and with a peduncle type of insertion, anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 3 D), but does not reach FII; external ascoids implanted at the same level as the internal; presence of preapical papilla on FI – FIII (Figs 3 D – F); papillae on FIV – FVI absent (Figs 4 A – C); presence of one papilla on FVII and FVIII (Figs 4 D and E), two papillae on FIX (Fig. 4 F), three papillae on FX and FXI (Figs 4 G and H), presence of four, six, six papillae distributed on FXII, FXIII and FXIV, respectively (Figs 4 I – K); preapical spiniform papilla on FIX (Fig. 4 F), FXI (Fig. 4 H) and FXIII (Fig. 4 J). Presence of simple setae on FIV – FXIV (Fig. 4 A – K). Palpi damaged in specimen, description based on the other specimen (Nº 92113). Palpi (P) (Fig. 3 G): palpal segment length: PI 32 (29 / 38, n = 2), PII 52 (52, n = 2), PIII 107 (107, n = 2), PIV 55 (58 / 61, n = 2), PV 96 (96 / 116, n = 2). The length of PIII and PV varied between specimens, with the PIII being longer in some specimens, while others had the largest PV. Palpal formula: 1 - 2 - 4 - (3 - 5) (n = 2); PIII with Newstead’s sensilla scattered along the segment (Fig. 3 G). Labial suture complete (Fig. 3 A). Cervix. Ventro-cervical sensilla absent. Cervical sclerites bearing a pair of spiniform sensilla. Thorax. Mesonotum 416 (416 / 423, n = 2) in length; pronotum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off-white. Two proepimeral setae; seven-eight upper anepisternal setae. Setae on the anterior margin of the katepisternum present. Wing (Fig. 4 L): 1,776 (1,813 / 1,831, n = 2) in length, 467 (496 / 504, n = 2) in width; veins: R 5 1,504 (1,504 / 1,091, n = 2); alfa 344 (381 / 396, n = 2); beta 290 (234 / 278, n = 2); gamma 180 (194 / 206, n = 2); delta 17 (17 / 38, n = 2); pi 88 (80 / 131, n = 2). Legs (anterior, median, posterior): Coxae: 307 (298 / 307, n = 2), 288 (279 / 293, n = 2), 288 (292 / 297, n = 2); Femur: 752 (694 / 740, n = 2), 679 (684, n = 1), 760 (759, n = 1); Tibia: 1,147 (1,147 / 1,202, n = 2), 1,258 (1,313, n = 1), 1,369 (1,461, n = 1); Tarsomere I: 693 (701 / 777, n = 2), 760 (832, n = 1), 803 (851, n = 1). Sum of tarsomeres II + III + IV + V (anterior, median, posterior): 657 (642 / 693, n = 2), 679 (730, n = 1), 686 (737, n = 1). Abdomen: 1,221 (1,350 / 1,535, n = 2) in length; tergal papillae absent. Terminalia (Fig. 4 M): gonocoxite 148 (145 / 148, n = 2) in length, 55 (55 / 58, n = 2) in width, without basal tuft of setae. Gonostyle 128 (130 / 142,1, n = 2) in length, without spiniform preapical seta and with four spines: one apical, one subapical (upper external), one lower external and one internal spine. The four spines are well-developed. Lower external spine implanted at a level closer to the upper external spine than the internal. Internal spine implanted in the apical third of the gonostylus. Paramere (Fig. 4 M): dorsal margin 128 (119 / 128, n = 2) and ventral 151 (148 / 151, n = 2) in length; basal level of implantation of the bristles of dorsal margin reaches the apical level implantation of the bristles in the ventral margin (Fig. 4 M). Parameral sheath sclerotized and coniform. Epandrial lobe 148 (146 / 148, n = 2) in length, 32 (29 / 32, n = 2) in width and rounded at the apex. Sperm pump (Fig. 4 N) 107 (107 / 119, n = 2); ejaculatory apodeme 78 (78 / 96, n = 2); aedeagal ducts with bevel shaped apex (Fig. 4 N), 397 (383 / 397, n = 2) in length and 3 (3, n = 2) in width; 3,7 times the length of the sperm pump. Cercus 119 (128 / 134, n = 2) in length, 49 (38 / 43, n = 2) in width.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB27A3730FF1A61ABFB82F924.taxon	materials_examined	Material examined: 3 males (Nº 89333, 89327, 89331). Capture location: municipality of Lassance (17 ° 53 ′ 22.31 ″ S, 44 ° 34 ′ 53.63 ″ W), located in the state of Minas Gerais, Brazil. Collection date: 26 / iv / 2010. Trap: CDC trap. Collector: not identified. Complementary material: 2 males (Nº 92113, 92114) from “ Parque Estadual do Rio Doce ” (19 º 30 ′ 45 ″ S, 42 º 33 ′ 10 ″ W), state of Minas Gerais, Brazil. Collection date: from June 2003 to July 2004. Trap: Malaise trap. Collector: Rogério Parentoni Martins. Material deposited in the FIOCRUZ / COLFLEB.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB27A3730FF1A61ABFB82F924.taxon	distribution	Distribution based on published articles: ARGENTINA: Resistencia, Chaco, Shannon & Del Ponte (1927); Misiones, Puerto Iguazú, Salomón et al. (2010: 69); Colonia Benítez; Margarita Belén, Szelag et al. (2017). COSTA RICA: Murilo & Zeledón (1985). COLOMBIA: Amazonas-Letícia; Boyacá-Puerto Boyacá; Caquetá-Solano; Tolima-Melga; Valle-Buenaventura, Lower Anchicaya Dam, Martins et al. (1978); La Macareña, Bejarano et al. (2006: 26). ECUADOR: Cañar-Conchacay; Guayas-Bucay, Cerro Cutuguay; Los Rios – Quevedo, La Montaña, Martins et al. (1978). FRENCH GUIANA: Approugue-Balourou, Guillaume, Saut Canori, Saut Machicou; Caux; Cayenne-Baduel, Cabossou, Crique, Anguille, Montabo, Rorota; Haute Mana-Souvenir; Iracoubo-Crique Blanche; Oyapock-Maripa, Saint Georges, Tampac, Martins et al. (1978). Montabo, Cayenne, French Guyana, Floch & Abonnenc (1943: 6). PANAMA: Darien-Altos de Quia, District de Pinogana; Panama canal area-Madden Forest Reserve, Martins et al. (1978). PERU: Cusco-Assunción, Pilcopata; Huánuco-Cachicoto; Loreto-Pucallpa, Serafin Filomeno, Zungarococha; Madre de Dios-Salvacíon, Alto Madre de Dios, Martins et al. (1978). REPUBLIC OF SURINAME: Sabajo Hills, Brokopondo, Kent et al. (2013: 6). TRINIDAD AND TOBAGO: Martins et al. (1978). VENEZUELA: Floresta Tropical úmida, Floresta Tropical Seca, Amazonas, Feliciangeli et al. (1988). BOLIVIA: Rio Yapacani, Santa Cruz, Bermudez & Young (1983: 5). BRAZIL: CEARÁ: Nova Olinda, Mangabeira (1942: 327). ACRE: Rio Branco, Martins et al. (1978). AMAPÁ: Macapá, Martins et al. (1978); Oiapoque, Santos et al. (2019: 14). AMAZONAS: Presidente Figueiredo, Alencar et al. (2011: 118); Terra Indígena Caititu, Lábrea, Silva et al. (2014). BAHIA: Parque Nacional do Pau-Brasil, Porto Seguro, Shimabukuro et al. (2016). ESPÍRITO SANTO: Vargem Alta, Pinto et al. (2009: 1). MARANHÃO: Cururupu; São Luís; Martins et al. (1978); Paço do Lumiar, Barros et al. (2000: 16); Barreirinhas, Rebêlo et al. (2010: 26); São Luís, Azevedo et al. (2011: 40); Águas do Miranda, Brilhante et al. (2015: 57); Santo Amaro, Pereira Filho et al. (2015): 8; Caxias, Guimarães & Silva et al. 2017. MATO GROSSO: Barra do Garças, Queiroz et al. (2012: 45); Sinop, Thies et al. (2018: 177). MATO GROSSO DO SUL: Corumbá, Galati et al. (1997: 31), Oliveira et al. (2016: 11), Barrios et al. (2019: 14); Campo Grande, Oliveira et al. (2003: 19), Galati et al. (2003: 47), Oliveira et al. (2006: 101), Dorval et al. (2016: 23); Bela Vista, Dorval et al. (2009; 2010). MINAS GERAIS: Timóteo, Andrade-Filho et al. (1997: 13); Pedra do Indaiá, Andrade-Filho et al. (2008: 37); Divinópolis, Margonari et al. (2010: 47), Nascimento et al. (2013: 125); Lassance, Carvalho et al. (2013; 2017); Terra Indígena Xacriabá, São João das Missões, Rêgo et al. (2015: 10), Costa et al. (2019: 52); Barra do Guaicuí, Várzea da Palma, Sanguinette et al. (2015: 8); Parque Estadual do Sumidouro, Saraiva et al. (2015: 10); Santuário do Caraça, Tonelli et al. (2017: 12); Ipatinga, Lana et al. (2018: 13); Rio Acima, Costa et al. (2021: 120). PARÁ: Belém, Martins et al. (1978); Bacarena, Oliveira et al. (2011: 106); Paraense; Santa Maria, Chagas et al. (2016: 7). PARANÁ: Itambaracá, Cruz et al. (2012: 45). PERNAMBUCO: Recife, Silva & Vasconcelos (2005: 38), Silva et al. (2020); São Vicente Férrer, Guimarães et al. (2012; 2014); Ipojuca, Miranda et al. (2015: 146); Iguaraçu, Silva & Vasconcelos (2005: 38), Silva et al. (2017: 26). PIAUÍ: Andrade-Filho et al. (2001: 96). RIO DE JANEIRO: Cantagalo, Peres Dias et al. (2016: 58). RIO GRANDE DO NORTE: Parnamirim, Pinheiro et al. (2013: 38); Nísia Floresta, Pinheiro et al. (2016: 9); Complexo Rio Doce, Silva et al. (2020). RONDÔNIA: Guajará Mirim; Porto Velho, Martins et al. (1978); Nova Mamoré, Parque Estadual Guajará-Mirim, Pereira-Júnior et al. (2019: 114); Ji-Paraná, Torchitte et al. (2020). SÃO PAULO: Botucatu, Cutolo et al. (2009: 18); Província Espeleológica do Vale do Ribeira, Parque Estadual Intervales, Galati et al. (2010: 54); Rio Claro, Cutolo et al. (2013: 19). TOCANTINS: Porto Nacional, Vilela et al. (2011: 6); Taquaraçu, Machado et al. (2012: 107). Distribution based on the analyzed slides: BRAZIL. Amapá: Pedra Branca do Amapari; Ceará: Maranguape; Goiás: Itumbiara; Mato Grosso: Alta Floresta, Chapada dos Guimarães; Mato Grosso do Sul: Camapuã; Minas Gerais: Belo Horizonte, Divinópolis, Jaboticatubas, Jequitinhonha, Lassance, Pains, Paracatu, Monsenhor Paulo, São João das Missões, Santana do Riacho, Rio Acima, Tupaciguara, Várzea da Palma; Pará: Oriximiná; Pernambuco: Recife; Piauí: Curralinho, Miguel Leão; Tocantins: Porto Nacional. Medical importance: Studies have not yet been conducted on this species regarding vector competence or capacity, or natural infection by parasites. Species feed in anurans (Costa et al. 2021).	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2753734FF1A6571FDB1FE30.taxon	description	(Figs 5 – 6)	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2753734FF1A6571FDB1FE30.taxon	diagnosis	Diagnosis. Male: preapical papilla on flagellomere III present; gonocoxite without basal tuft of setae; flagellomere I ≥ 405 µm; aedeagal ducts ≤ 440 µm; epandrial lobe ≤ 157 µm; presence of one apical spine and upper external spine at the apical level in the gonostyle. Male holotype. Head (Fig. 5 A) 330 (320 – 330, n = 6) in length, 264 (255 – 304, n = 6) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 119 (107 – 118, n = 7) in length; eyes 154 (133 – 159, n = 6) in length, 96 (70 – 93, n = 6) in width; interocular distance 122 (101 – 125, n = 5). Pharynx with streaks and no teeth (Fig. 5 B). Cibarium without teeth (Fig. 5 B). Labrum-epipharynx (LE) 136 (142 – 148, n = 6) (Fig. 5 C). Incomplete antenna on type specimen, description based on paratype (Nº 32847). Flagellomere XIII and FXIV absent in all specimens. Antenna (Figs 5 D – F, 6 A – I): flagellomere length (F): FI 442 (349 – 474, n = 6), FII 225 (172 – 228, n = 6), FIII 229 (172 – 228, n = 6), FXIII (lost), FXIV (lost). Ascoids: with short posterior spur and atrophied peduncle, anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 5 D) but does not reach FII; ascoids: implanted at the same level as the internal; flagellomere FX, FXI and FXII absent in type specimen, description based on paratype (Nº 32847). Presence of preapical papilla on FI – FIII (Figs 5 D – F); papilla absent on FIV – FVI (Figs 6 A – C); presence of one papilla on FVII, FVIII and FIX (Figs 6 D – F), two papillae on FX (Fig. 6 G), three papillae on FXI (Fig. 6 H); presence of four papillae on FXII (Fig. 6 I); spiniform preapical papilla on FIX (Fig. 6 F) and FXI (Figs 6 H). Presence of simple setae on FVII – FXII (Figs 6 A – I). Palpi (P) (Fig. 5 G), palpal segment length: PI 32 (29 – 35, n = 6), PII 58 (55 – 64, n = 6), PIII 101 (92 – 116, n = 6), PIV 61 (64 – 70, n = 6) and PV 128 (107 – 130, n = 5). Palpal formula: 1 - 2 - 4 - 3 - 5 (n = 5); PIII with Newstead’s sensilla scattered along the segment (Fig. 5 G). Labial suture complete (Fig. 5 A). Cervix. Ventro-cervical sensilla absent. Cervical sclerites bearing a pair of spiniform sensilla. Thorax. Mesonotum 409 (372 – 457, n = 6) in length; pronotoum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off – white. One proepimeral setae; 9 – 11 upper anepisternal setae. Setae present on the anterior margin of the katepisternum. Wing (Fig. 6 J): 1,794 (1,461 – 1,905, n = 5) in length, 445 (416 – 525, n = 5) in width; veins: R 5 1,091 (1,036 – 1,128, n = 5); alfa 399 (320 – 404, n = 5); beta 246 (197 – 284, n = 5); gamma 197 (194 – 255, n = 5); delta 90 (41 – 113, n = 5); pi 95 (51 – 117, n = 5). Legs (anterior, median, posterior): Coxae: 279 (288 – 316, n = 6), 279 (288 – 302, n = 6), 279 (293 – 316, n = 6); Femur: 759 (642 – 774, n = 3), 715 (598 – 737, n = 5), 723 (657 – 795, n = 4); Tibia: 1,147 (943 – 1,221, n = 3), 1,258 (1,091 – 1,406, n = 5), 1,276 (1,147 – 1,480, n = 3); Tarsomere I: 672 (591 – 723, n = 3), 715 (650 – 803, n = 5), 752 (679 – 854, n = 3). Sum of tarsomeres II + IV + V (anterior, median, posterior): 644 (584 – 679, n = 3), 693 (605 – 715, n = 5), 715 (657 – 715, n = 3). Abdomen: 1,720 (1,128 – 1,720, n = 6) in length; tergal papillae absent. Terminalia (Fig. 6 K): gonocoxite 154 (145 – 154, n = 6) in length, 58 (52 – 61, n = 6) in width, without basal tuft of setae. Gonostyle 136 (122 – 133, n = 7) in length, without preapical seta and with four spines: one apical, one upper external that migrates towards the apex, one lower external, and one internal spine. The four spines are well-developed. The lower external spine is located at a level closer to the apex spines than the internal. Internal spine located in the apical third of the gonostylus. Paramere (Fig. 6 K): dorsal margin 110 (90 – 133, n = 6) and ventral 128 (124 – 159, n = 6) in length; the basal level of the bristles in the dorsal margin reaches the apical level of the implantation of the bristles in the ventral margin (Fig. 6 K). Parameral sheath sclerotized and coniform. Epandrial lobe 154 (145 – 177, n = 6) in length, 29 (20 – 38, n = 6) in width and rounded at the apex. Sperm pump (Fig. 6 L) 130 (101 – 130, n = 6); ejaculatory apodeme 96 (81 – 101, n = 6); aedeagal ducts with bevelled apex (Fig. 6 L), 394 (375 – 414, n = 6) in length and 3 (3, n = 6) in width; 2,7 times the length of sperm pump. Cercus 96 (107 – 130, n = 6) in length and 43 (40 – 58, n = 6) in width.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2753734FF1A6571FDB1FE30.taxon	materials_examined	Material examined: Holotype (Nº 32827) and two paratypes (Nº 32832, 32847). Capture location: Fazenda São Geraldo, house of Mr. José Marques dos Santos, municipality of Monsenhor Paulo (21 ° 45 ′ 30.14 ″ S, 45 ° 32 ′ 22.15 ″ W), state of Minas Gerais, Brazil. Collection date: uninformed. Trap: uninformed, Collectors: Antônio Leôncio and Eduardo. Identified as Lu. nordestina. Paratypes from other locations: 1 male (Nº 49201). Capture location: Engenho de São Francisco da Várzea (8 º 03 ′ 27 ″ S, 34 º 57 ′ 52 ″ W), municipality of Recife, state of Pernambuco, Brazil. Collection date: 24 / ix / 1974, from 1: 00 p. m. to 4: 00 p. m. Trap: manual capture in wild animal burrows (termites) in closed florest. Collectors: João Evangelista da Silva. Determined by Amilcar Vianna Martins as L. nordestina in October 1974. 1 male (Nº 64671). Capture location: “ Sítio do Sr. Alencar ” municipality of Maranguapé (3 º 52 ′ 58.62 ″ S, 38 º 40 ′ 36.20 ″ W), state of Ceará, Brazil. Collection date: 17 / x / 1983, from 11: 00 a. m. to 11: 30 a. m. Trap: Manual capture in uninhabited shelters approximately 3 m away from the water collection, feeding on frogs. Collectors: Edelberto Santos Dias. Determined by Alda Lima Falcão and Armando Lemos Ribeiro as L. nordestina. 1 male (Nº 71674). Capture location: municipality of Monte do Carmo (Nº 10 º 47 ′ 12.71 ″ S, 48 º 00 ′ 03.70 ″ W), state of Tocantins, Brazil. Collection date: i / 1998. Trap: uninformed. Collector: W. A. Andrade. 1 male (Nº 86806). Capture location: Caverna Hall two of Caverna do Arrebenta Bomba, localized in the municipality of Lassance (17 º 53 ′ 22.31 ″ S, 44 º 34 ′ 53.63 ″ W), state of Minas Gerais, Brazil. Collection date: 15 / iv / 2008, from 2: 00 p. m. (16 / iv / 2008) to 2: 00 p. m. (17 / iv / 2008). Trap: uninformed. Collector: uninformed. Determined by Gustavo Mayr de Lima Carvalho as Lu. sordellii in May 2008. Material deposited in the FIOCRUZ / COLFLEB. Female: Unknown.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2753734FF1A6571FDB1FE30.taxon	etymology	Etymology: the species name refers to the position of the upper external spine, which is located more at the apex of the gonostyle, different from the males of Sc. sordellii.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2753734FF1A6571FDB1FE30.taxon	distribution	Distribution: BRAZIL. Ceará: Maranguapé; Minas Gerais: Lassance, Monsenhor Paulo, Nova Lima; Pernambuco: Recife; Tocantins: Monte do Carmo. Medical importance: Studies have not yet been conducted on this species regarding vector competence or capacity, or natural infection by parasites.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB271372BFF1A601DFDB1F832.taxon	description	(Figs 7 – 10)	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB271372BFF1A601DFDB1F832.taxon	diagnosis	Diagnosis. Male: preapical papilla on flagellomere III present; 5 th palpal segment larger than 3 rd and shorter than or as long as sum of 3 rd + 4 th; gonocoxite with basal tuft of setae; gonocoxite ≤ 175 µm, not arch-shaped on its dorsal margin; aedeagal ducts ≤ 600 µm. Female: preapical papilla on flagellomere III present; 5 th palpal segment larger than 3 rd and shorter than or as long as sum of 3 rd + 4 th; posterior teeth of the cibarium straight, facing the pharynx; anterior teeth arranged in a “ V ” shaped row. Male holotype. Head (Fig. 7 A) 334 (325 / 339, n = 2) in length, 299 (278, n = 2) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 113 (113, n = 2) in length; eyes 151 (142 / 157, n = 2) in length, 81 (81 / 84, n = 2) in width; interocular distance 142 (139, n = 2). Pharynx with streaks and no teeth (Fig. 7 B). Cibarium without teeth (Fig. 7 B). Labrum-epipharynx (LE) 148 (148 / 153, n = 2) (Fig. 7 C). Labial suture complete (Fig. 7 A). Damaged antennae on the type specimen, description based on paratype (Nº 92116). Antenna (Figs 7 D – F, 8 A – K): flagellomere length (F): FI 363 (386 / 404, n = 2), FII 181 (186 / 190, n = 2), FIII 190 (190 / 195, n = 2), FXIII 75 (81 / 84, n = 2) and FXIV 75 (72 / 81, n = 2). Ascoids: with short posterior spur and atrophied peduncle, anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 7 D), but does not reach FII; external ascoids located on a more apical level than the internal in FI; presence of preapical papilla on FI – FIII (Figs 7 D – F); papilla absent on FIV – FVIII (Figs 8 A – E); one papilla on FIX (Fig. 8 F), three papillae on FX and four papillae on FXI (Figs. 8 G and H); presence of four, five, six papillae distributed on FXII, FXIII and FXIV, respectively (Figs 8 I – K); spiniform preapical papilla on FIX (Fig. 8 F), FXI (Fig. 8 H) and FXIII (Fig. 8 J). Presence of simple setae on FVII – FXIV (Fig. 8 D – K). Palpi (P) (Fig. 7 G), segment palpal length: PI 35 (35 / 38, n = 2), PII 72 (69 / 75, n = 2), PIII 122 (116 / 122, n = 2), PIV 75 (78 / 84, n = 2) and PV 159 (185 / 191, n = 2). Palpal formula: 1 - 2 - 4 - 3 - 5 (n = 2); PIII with Newstead’s sensilla scattered along the segment (Fig. 7 G). Cervix. Ventro-cervical sensilla absent. Cervical sclerites bearing a pair of spiniform sensilla. Thorax. Mesonotum 489 (438 / 460, n = 2) in length; pronotum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off-white. One proepimeral setae; 11 – 15 upper anepisternal setae. Setae present on the anterior margin of the katepisternum. Wing (Fig. 8 L): 1,850 (1,831 / 1,850, n = 2) in length, 533 (511, n = 2) in width; veins: R 5 1,054 (1,091 / 1,110, n = 2); alfa 391 (418 / 423, n = 2); beta 171 (171 / 186, n = 2); gamma 275 (258 / 267, n = 2); delta 41 (55 / 64, n = 1); pi 66 (73 / 80, n = 1). Median and posterior legs lost in the holotype, description based on paratype (Nº 92116). Legs: (anterior, median, posterior): Coxae: 339 (325, n = 2), 325 (302, n = 2), 330 (302 / 316, n = 2); Femur: 781 (751, n = 1), 745, 818; Tibia: 1,091 (1,128, n = 2), 1,129, 1,461; Tarsomere I: 737 (715, n = 1), 745, 810. Sum of tarsomeres II + III + IV + V (anterior, median, posterior): 737 (708, n = 1), 708, 730. Abdomen: 1,998 (1,628 / 1,757, n = 2) in length; tergal papillae absent. Terminalia (Fig. 8 M): gonocoxite 171 (162 / 171, n = 2) in length, 58 (64, n = 2) in width, with basal tuft of setae, 9 – 10 setae (9 / 12, n = 2). Gonostyle 133 (124 / 133, n = 2) in length, without preapical seta and with four spines: one apical, one upper external, one lower external, and one internal. The four spines are well-developed. The lower external spine is located at a level closer to the upper external spine than to the internal. The internal spine is located in the apical third of the gonostyle. Paramere (Fig. 8 M): dorsal margin 157 (145 / 151, n = 2) and ventral 183 (168 / 191, n = 2) in length; the basal level of the bristles in the dorsal margin reaches the apical level of the insertion of the bristles in the ventral margin (Fig. 8 M). Parameral sheath sclerotized and coniform. Epandrial lobe 209 (194 / 206, n = 2) in length, 32 (32 / 35, n = 2) in width and rounded at the apex. Sperm pump (Fig. 8 N) 136 (122 / 125, n = 2); ejaculatory apodeme: 104 (93 / 101, n = 2); aedeagal ducts with bevelled apex (Fig. 8 N), 577 (565 / 568, n = 3) in length and 2.9 (2.9, n = 2) in width; 4.2 times the length of the sperm pump. Cercus: 116 (110 / 116, n = 2) in length, 46 (40, n = 2) in width.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB271372BFF1A601DFDB1F832.taxon	materials_examined	Material examined: Male holotype (Nº 92115). Capture location: neighborhood of Jacarepaguá (23 º 00 ′ 31 ″ S, 43 º 21 ′ 01 ″ W), located in the municipality of Rio de Janeiro, state of Rio de Janeiro, Brazil. Collection date: 14 / iv / 2015. Trap: HP. Collector: Godoy R. E. Three males paratypes (Nº 92116, 92117, 92118). Same data of the holotype. Collection date: 13 to the 16 / iv / 2015 and 19, 21 / v / 2015. Material deposited in FIOCRUZ / COLFLEB. Female. Head (Fig. 9 A): 353 (339 – 372, n = 4) in length, 296 (290 – 345, n = 3) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 119 (116 – 122, n = 4) in length; eyes 159 (145 – 168, n = 4) in length, 87 (81 – 104, n = 4) in width; interocular distance 148 (142 – 151, n = 3). Pharynx with streaks and no teeth (Fig. 9 B). Cibarium (Fig. 9 B) with four posterior teeth well-developed, slight curved towards the central region of the cibarium, 14 anterior teeth arranged in a “ V ” shaped row and eight lateral teeth flanking the posterior teeth; sclerotized area short and triangular; sclerotized arch incomplete. Labial suture complete (Fig. 9 A). Mouthparts difficult to see in the type specimen, description based on the other paratype (92120) Labrumepipharynx (LE) (Fig. 9 C): 186 (177 – 194, n = 4); mandibles as shown (Fig. 9 D). Hypopharynx with 20 – 24 teeth on apical region (Fig. 9 E). Lacinia of maxilla with 16 – 17 internal teeth and without external teeth (Fig. 9 F). Damaged antenna on paratype (Nº 92119), description based on the other paratype (Nº 92120). Antenna (Figs 9 G – I, 10 ª – K), flagellomere length: FI 358 (358 – 386, n = 4), FII 167 (167 – 181, n = 4), FIII 167 (167 – 181, n = 4), FXIII 87 (75 – 90, n = 3) and FXIV 72 (61 – 78, n = 3). Ascoids: with short posterior spur and atrophied peduncle, anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 9 G), but does not reach FII; external ascoids implanted on more apical level than the internal in FI; presence of preapical papillae on FI – FIII (Figs 9 G – I); Papillae absent on FIV – FIX (Figs 10 A – F), three and four papillae on FX and FXI (Figs 10 G and H), respectively; presence of four, five and six papillae on FXII, FXIII and FXIV, respectively (Figs 10 I – K); preapical spiniform papillae observed on FIX (Fig. 10 F), FXI (Fig. 10 H), and FXIII (Fig. 10 J). Presence of setae simple on FX – FXIV (Figs 10 G – K), Palpi (P) (Fig. 9 J), palpal segment lenght: PI 38 (38 – 41, n = 4), PII 78 (78 – 81, n = 4), PIII 125 (125 – 142, n = 4), PIV 81 (78 – 96, n = 4) and PV 191 (162 – 209, n = 4). Palpal formula: 1 - 2 - 4 - 3 - 5 (n = 4); PIII with Newstead’s sensilla scattered along the segment (Fig. 9 J). Cervix: Ventro-cervical sensilla absent. Cervical sclerites bearing a pair on of spiniform sensilla. Thorax: Mesonotum 533 (496 – 577, n = 4) in length. Mesonotum, pronotum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off-white. One proepimeral seta; 12 – 13 upper anepisternal setae. Setae presence on the anterior region of katepisternum. Wing (Fig. 10 L): 2,072 (1,905 – 2,165, n = 4) in length, 628 (599 – 679, n = 3) in width; veins: R 5 1,202 (1,184 – 1,278, n = 3); alfa 526 (460 – 545, n = 3); beta 205 (218 – 232, n = 3); gamma 261 (255 – 293, n = 3); delta 110 (78 – 122, n = 3); pi 73 (73 – 139, n = 3). Posterior legs absent on paratype (92119), description based on the other paratype (92120). Coxae (anterior, median, posterior): 358 (325 – 377, n = 4), 344 (307 – 358, n = 4), 349 (311 – 363, n = 4); Femur: 854 (752 – 876, n = 3), 818 (737, n = 2), posterior lost (818 / 920, n = 2); Tibia: 1,128 (1,073 – 1,202, n = 3), 1,313 (1,221, n = 2), posterior lost (1,369 / 1,461, n = 2); Tarsomere I: 730 (664 – 803, n = 3), 766 (701, n = 2), posterior lost (766 / 832, n = 2). Sum of tarsomeres II + III + IV + V, 693 (686 – 796, n = 2), 715 (701, n = 2), posterior lost (715 / 796, n = 2). Abdomen: 1,887 (1,554 – 1,887, n = 3 = 4) in length; tergal papillae absent. Spermathecae not visible on paratype (Nº 92119), description based on another paratype (Nº 92121). Spermathecae (Fig. 10 M): 64 (n = 1) in length, 9 (n = 9) in width; with poorly defined rings; terminal knob: 6 (n = 1) in length and 6 (n = 1) in width, individualized, with rounded shape and bristles in the apical region; common spermathecal duct: not visualized; individual spermathecal duct not fully visualized, 6 (n = 1) in width, they are membranous with smooth walls and their widths are uniform throughout their length (Fig. 10 M). Cercus: 130 (116 – 159, n = 4) in length, 70 (61 – 75, n = 4) in width. Material examined: 5 females paratypes (Nº 92119, 92120, 92121, 92122, 92123). Capture location: 4 females from neighborhood of Jacarepaguá (23 º 00 ′ 31 ″ S, 43 º 21 ′ 01 ″ W), localized in the municipality of Rio de Janeiro, state of Rio de Janeiro, Brazil. Collection date: 13, 14 / iv / 2015 and 21 / v / 2015. Trap: HP. Collector: Rodrigo Espíndola Godoy. One female (Nº 92123) from municipality of Muniz Freire (20 º 27 45 ″ S, 41 º 24 ′ 55 ″ W), state of Espírito Santo, Brazil. Collection date: iii / 2016. Trap: HP. Collector: Israel Pinto. Material deposited in the FIOCRUZ / COLFLEB.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB271372BFF1A601DFDB1F832.taxon	etymology	Etymology: The species name pays homage to Dr. Felipe Dantas Torres, “ Instituto Aggeu Magalhães ”, Fiocruz, Pernambuco, for his important contribution in the studies of leishmaniases and infectious diseases in Brazil.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB271372BFF1A601DFDB1F832.taxon	distribution	Distribution: BRAZIL. Espírito Santo: Muniz Freire. Rio de Janeiro: Jacarepaguá. Medical importance: Studies have not yet been conducted on this species regarding vector competence or capacity, or natural infection by parasites.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB26B3721FF1A62CBFD2FF994.taxon	description	(Figs 11 – 14)	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB26B3721FF1A62CBFD2FF994.taxon	diagnosis	Diagnosis. Male: preapical papilla on flagellomere III present; 5 th palpal segment larger than 3 rd and shorter than or as long as sum of 3 rd + 4 th; gonocoxite with basal tuft of setae; gonocoxite ≥ 195 µm, arch-shaped on its dorsal margin; aedeagal ducts ≥ 695 µm. Female: preapical papilla on flagellomere III present; 5 th palpal segment larger than 3 rd and shorter than or as long as sum of 3 rd + 4 th; posterior teeth of the cibarium straight, facing the pharynx; anterior teeth arranged in a “ V ” shaped row. Male holotype. Head (Fig. 11 A) 363 (349 / 377, n = 2) in length, 290 (275 / 278, n = 2) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 128 (116 / 128, n = 2) in length; eye 142 (122 / 145, n = 2) in length, 81 (61 / 96, n = 2) in width; interocular distance 139 (136 / 142, n = 2). Pharynx with streaks and no teeth (Fig. 11 B). Cibarium without teeth (Fig. 11 B). Labial suture complete (Fig. 11 A). Labrumepipharynx (LE) 159 (162 / 168, n = 2) (Fig. 11 C). Antenna (Figs 11 D – F, 12 A – K): flagellomere length (F): FI 502 (400 / 469, n = 2), FII 246 (218 / 237, n = 2), FIII 237 (214 / 242, n = 2), FXIII 90 (90 / 99, n = 2) and FXIV 84 (78 / 84, n = 2). Ascoids: with short posterior spur and atrophied peduncle, anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 11 D), but does not reach FII; external ascoids located on a more apical level than the internal in FI; presence of preapical papillae on FI – FIII (Figs 11 D – F); papillae absent on FIV – FVIII (Figs 12 A – E); one papilla on FIX (Fig. 12 F), three papillae on FX (Fig. 12 G), four papillae on FXI (Fig. 12 H), presence of five, six, six papillae distributed on FXII, FXIII and FXIV, respectively (Figs 12 I – K); spiniform preapical papilla on FIX (Fig. 12 F), FXI (Fig. 12 H) and FXIII (Fig. 12 J). Presence of simple setae on FVIII – FXIV (Figs 12 E – K). Palpi (P) (Fig. 11 G), palpal segment length: PI 41 (32 / 41, n = 2), PII 78 (75, n = 2), PIII 145 (136 / 148, n = 2), PIV 93 (93, n = 2) and PV 206 (229, n = 2). Palpal formula: 1 - 2 - 4 - 3 - 5 (n = 3); PIII with Newstead’s sensilla scattered along the segment (Fig. 11 G). Cervix. Ventro-cervical sensilla absent. Cervical sclerites bearing a pair of spiniform sensilla. Thorax. Mesonotum 496 (453 / 533, n = 2) in length; pronotum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off-white. One proepimeral seta; 16 upper anepisternal setae. Setae presence on the anterior margin of the katepisternum. Wing (Fig. 12 L): 2,275 (2,183 / 2,432, n = 2) in length, 606 (664, n = 2) in width; veins: R 5 1,387 (1,628, n = 2); alfa 586 (614, n = 2); beta 232 (273 / 278, n = 2); gamma 275 (203 / 296, n = 2); delta 159 (81 / 154, n = 2); pi 153 (80, n = 2). Legs: median and posterior legs absent in all specimens. Legs (anterior, median, posterior): Coxae: 372 (349 / 381, n = 2), 353 (311 / 358, n = 2), 363 (326 / 363, n = 2); Femur: 956, median lost, posterior lost; Tibia: 1,480, median lost, posterior lost; Tarsomere I: 956, median lost, posterior lost. Sum of tarsomeres II + III + IV + V (anterior, median, posterior): 836, median lost, posterior lost. Abdomen: 1,739 (1,498 / 1,665, n = 2) in length; tergal papillae absent. Terminalia (Fig. 12 M): gonocoxite 206 (197 / 206, n = 2) in length, 49 (46 / 58, n = 2) in width, with basal tuft of setae, 10 – 11 (10 / 12, n = 2) long setae. Gonostyle 157 (148 / 174, n = 2) in length, without preapical seta and with four spines: one apical, one upper external, one lower external, and one internal. The four spines are well-developed. The lower external spine is located at a level closer to the internal spine than the upper external. Internal spine located in the apical third of the gonostyle. Paramere (Fig. 12 M): dorsal margin 148 (171, n = 2) and ventral 183 (191 / 206, n = 2) in length; the basal level of the bristles in the dorsal margin reaches the apical level of the implantation of the bristles in the ventral margin (Fig. 12 M). Parameral sheath sclerotized and coniform. Epandrial lobe 252 (261, n = 2) in length, 29 (26 / 35, n = 2) in width and rounded at the apex. Sperm pump (Fig. 12 N) 128 (125, n = 2); ejaculatory apodeme 101 (96 / 101, n = 2); aedeagal ducts with bevelled apex (Fig. 12 N), 702 (696 / 734, n = 2) in length and 3 (3, n = 2) in width; 7 times the length of the ejaculatory apodeme. Cercus: 157 (165 / 168, n = 2) in length, 49 (46 / 52, n = 2) in width.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB26B3721FF1A62CBFD2FF994.taxon	materials_examined	Material examined: Male holotype (Nº 92124) and two males paratypes (Nº 92125, 92126). Capture location: municipality of Santa Bárbara (19 º 59 ′ 08 ″ S, 43 º 45 ′ 01 ″ W), localized in state of Minas Gerais, Brazil. Collection date: 09 and 15 / xii / 2011. Trap: CDC. Collector: Bess et al. Material deposited at FIOCRUZ / COLFLEB. Female. Head (Fig. 13 A): 367 (367 – 437, n = 4) in length, 293 (293 – 348, n = 3) in width. Arrangement of deciduous bristles in the occiput region forming an “ X ” shape; clypeus 122 (119 – 139, n = 4) in length; eyes 139 (139 – 165, n = 4) in length, 78 (72 – 97, n = 4) in width; interocular distance 145 (142 – 162, n = 4). Pharynx with streaks and short teeth (Fig. 13 B). Labrum-epipharynx (LE) (Fig. 13 C): 206 (191 – 235, n = 4). Mandible, hypopharynx and lacinia of maxilla not visible on paratype (Nº 92127), description based on the other paratype (Nº 92128): mandible as shown (Fig. 13 F). Hypopharynx with 28 – 30 on apical region (Fig. 13 D). Lacinia of maxilla with 32 internal teeth and without external teeth (Fig. 13 E). Cibarium (Fig. 13 B) with four posterior teeth well-developed, straight, and facing the pharynx, 16 anterior teeth arranged in a “ V ” shaped row; sclerotized area short and triangular; sclerotized arch incomplete. Antenna (Figs 13 G – I, 14 A – K): flagellomere length (F): FI 437 (390 – 465, n = 4), FII 200 (186 – 209, n = 4), FIII 200 (186 – 209, n = 4), FXIII 93 (96, n = 2), and FXIV 87 (90, n = 2). Ascoids: with short posterior spur and atrophied peduncle, anterior spur is long in FI and reaches the level of the preapical papilla (Fig. 13 G), but does not reach FII; external ascoids located on a more apical level than the internal in FI; presence of preapical papilla on FI – FIII (Figs 13 G – I); papillae absent on FIV – FVIII (Figs 14 A – E); presence of one papilla on FIX (not visible in the paratype (Nº 92127), present in the other paratypes), three papillae on FX and FXI (Figs 14 G and H); presence of five papillae on FXII, FXIII and FXIV (Figs 14 I – K); preapical spiniform papilla observed on FIX (Fig. 14 F), FXI (Fig. 14 H), and FXIII (Fig. 14 J). Presence of simple setae on FVIII – FXIV (Figs 14 E – K). Labial suture complete (Fig. 13 A). Palpi (P) (Fig. 13 J), palpal segment lenght: PI 41 (41 – 46, n = 4), PII 90 (72 – 93, n = 4), PIII 136 (136 – 157, n = 4), PIV 96 (84 – 104, n = 5) and PV 215 (191 – 215, n = 4). Palpal formula: 1 - 2 - 4 - 3 - 5; PIII (Fig. 13 J) with Newstead’s sensilla scattered along the segment. Cervix: Ventro-cervical sensilla absent. Cervical sclerites bearing a pair of spiniform sensilla. Thorax: Mesonotum 547 (547 – 679, n = 4) in length. Mesonotum, pronotum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off-white. Two to three proepimeral setae; 10 – 11 upper anepisternal setae. Setae presence on the anterior region of katepisternum. Wings damaged in the paratype (Nº 92127); description based on the other paratype (Nº 92128). Wing (Fig. 14 L): 2,460 (2,275 – 2,682, n = 4) in length, 803 (664 – 803, n = 4) in width; veins: R 5 1,542 (1,332 – 1,665, n = 4); alfa 730 (591 – 788, n = 4); beta 256 (209 – 284, n = 3); gamma 325 (220 – 333, n = 4); delta 130 (130 – 241, n = 4); pi 73 (51 – 73, n = 4). Legs absent in all specimens. Coxae (anterior, median, posterior): 353 (353 – 437, n = 4), 349 (349 – 433, n = 4), 353 (353 – 428, n = 4). Femur: anterior lost, median lost, posterior lost. Tibia: anterior lost, median lost, posterior lost. Tarsomere I: anterior lost, median lost, posterior lost. Sum of tarsomeres II + III + IV + V: anterior lost, median lost, posterior lost. Abdomen: 1,924 (1,887, n = 2) in length; tergal papillae absent. Spermathecae not visible in paratype (Nº 92127), description based on another paratype (Nº 92129). Spermathecae (Fig. 14 M): 49 (n = 1) in length, 12 (n = 1) in width; with poorly defined rings; terminal knob: 9 (n = 1) in length and 6 (n = 1) in width, individualized, with rounded shape and without bristles in the apical region; common spermathecal duct: 44 (n = 1) in length, 10 (n = 1) in width; individual spermathecal duct not fully visualized, 6 (n = 1) in width, they are membranous with smooth walls and their widths are uniform throughout their length (Fig. 14 M). Cercus: 159 (139 – 150, n = 4) in length and 52 (64 – 70, n = 4) in width. Material examined: 5 females paratypes (Nº 92127, 92128, 92129, 92130, 92131). Capture location: municipality of Santa Bárbara (19 º 59 ′ 08 ″ S, 43 º 45 ′ 01 ″ W), localized in state of Minas Gerais, Brazil. Collection date: 08 / vii / 2011, 09 and 13 / i / 2012. Trap: CDC Collector: Bess et al. Material deposited in the FIOCRUZ / COLFLEB.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB26B3721FF1A62CBFD2FF994.taxon	etymology	Etymology: The species name pays homage to Dr. Paloma Helena Fernandes Shimabukuro, “ Instituto René Rachou ”, Fiocruz, Minas Gerais for her studies in taxonomy of the American sand flies.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB26B3721FF1A62CBFD2FF994.taxon	distribution	Distribution: BRAZIL. Minas Gerais: Santa Bárbara; Rio Acima (Costa et al. 2021); Pains. Medical importance: Studies have not yet been conducted on this species regarding vector competence or capacity, or natural infection by parasites.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB26B3721FF1A62CBFD2FF994.taxon	discussion	Comments: Females and males identified as “ Sciopemyia aff. microps ” by Costa et al. (2021) are the same specimens proposed as Sciopemyia shimabukuroae sp. n. in the present study. It is known that the species can be found in caves feeding on anurans (Costa et al. 2021).	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2643720FF1A64FFFDB1F850.taxon	description	(Figs 15 – 16)	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2643720FF1A64FFFDB1F850.taxon	diagnosis	Diagnosis. Female: preapical papilla on flagellomere III absent; individual spermathecal ducts practically of the same width throughout their length and eight times or less longer than the spermatheca; this latter with the intermediate rings perceptibly of the same width as the apical rings. Width of individual spermathecal ducts at the spermatheca’s junction equivalent to half or more of the spermatheca’s width, measured at its central part. Width of the basal part of the individual spermathecal ducts (junction with the common duct) ca. half of the spermaheca’s width, measured at its central part. Cibarium with the posterior teeth not lateralized and the distance between the internal ones slightly greater than that between these latter and the external ones. Fifth palpal segment clearly shorter than the sum of 3 rd + 4 th. Individual spermathecal ducts at most 3.5 times longer than the spermatheca. Individual spermathecal ducts ca. two times longer than the spermatheca and increases in caliber as they get closer to the spermatheca. Presence of “ callosity ” in the individual spermathecal ducts. Female holotype. Head (Fig. 15 A): 381 (348 – 391, n = 8) in length, 339 (316 – 359, n = 8) in width. Arrangement of deciduous bristles in the occiput region in an “ X ” shape; clypeus 125 (104 – 133, n = 8) in length; eyes 171 (156 – 174, n = 8) in length, 99 (81 – 101, n = 8) in width; interocular distance 145 (142,1 – 153,7, n = 7). Pharynx with streaks and short teeth (Fig. 15 B). Labrum-epipharynx (LE) (Fig. 15 C): 188 (176 – 203, n = 8); mandible as shown (Fig. 15 D). Hypopharynx with 6 – 18 teeth on apical region (Fig. 15 E). Lacinia of maxilla with nine external teeth and 16 internal teeth (Fig. 15 F). Cibarium (Fig. 15 B) with four posterior teeth well-developed, 10 anterior teeth arranged in a “ U ” shaped row; sclerotized area short and triangular; sclerotized arch complete. Antenna (Figs 15 G – I, 16 A – K), flagellomere length (F): FI 428 (413 – 483, n = 7), FII 223 (205 – 242, n = 7), FIII 218 (209 – 223, n = 7), FXIII 93 (75 – 99, n = 7) and FXIV 75 (72 – 78, n = 7). Ascoids: with short posterior spur and atrophied peduncle, anterior spur is long in FI but does not reach the level of the preapical papilla (Fig. 15 G); external ascoids is located at a more basal level than the internal, in FI; presence of preapical papilla on FI – FII (Figs 15 G and H); papillae absent on FIII – FIX (Figs 15 I, 16 A – F); one papilla on FX and FXI (Figs 16 G – H), presence of five, six and four papillae on FXII, FXIII and FXIV, respectively (Figs 16 I – K); preapical spiniform papilla observed on FIX (Fig. 16 F), FXI (Fig. 16 H), and FXIII (Fig. 16 J). Presence of simple setae on FVIII – FXIV (Figs 16 E – K). Labial suture complete (Fig. 15 A). Palpi (P) (Figs 15 J), palpal segment lenght: PI 46 (38 – 46, n = 8), PII 82 (72 – 87, n = 8), PIII 128 (110 – 128, n = 8), PIV 82 (75 – 81, n = 8) and PV 165 (157 – 171, n = 6). Palpal formula: 1 - 4 - 2 - 3 - 5 (n = 9); PIII with Newstead’s sensilla scattered along the segment. Cervix: Ventro-cervical sensilla absent. Cervical sclerites bearing a pair of spiniform sensilla. Thorax: Mesonotum 599 (511 – 642, n = 7) in length. Mesonotum, pronotum, anepisternum, metanotum and postnotum light brown, paratergite and pleura off-white. Two proepimeral setae; 10 – 11 upper anepisternal setae. Setae presence on the anterior region of katepisternum. Wing (Fig. 16 L): 2,127 (1,850 – 2,201, n = 8) in length, 635 (577 – 679, n = 8) in width; veins: R 5 1,334 (1,184 – 1,387, n = 8); alfa 547 (460 – 569, n = 8); beta 177 (139 – 200, n = 8); gamma 406 (336 – 423, n = 8); delta 232 (162 – 232, n = 8); pi 73 (36 – 73, n = 8). Legs (anterior; median; posterior): Coxae: 390 (339 – 391, n = 8), 372 (316 – 372, n = 8), 381 (307 – 381, n = 8); Femur: 920 (912 / 934, n = 2), 869 (825 – 869, n = 3), posterior lost (1,000, n = 1; Tibia: 1,184 (1,147 / 1,202, n = 2), 1,332 (1,332 – 1,406, n = 3), posterior lost (1,572,5, n = 1); Tarsomere I: 752 (730 / 760, n = 2), 832 (803 – 854, n = 3), posterior lost (891, n = 1). Sum of tarsomeres II + III + IV + V, 723 (715 / 766, n = 2); 737 (737 – 788, n = 3), posterior lost (781, n = 1). Abdomen: 1,757 (1,702 – 2,183, n = 8) in length; tergal papillae absent. Spermathecae with poorly defined ring (Fig. 16 M); the apical ring is wider and longer than the ring that precedes it, 64 (61 – 64, n = 7) in length and 14 (12 – 17, n = 7) in width. Terminal knob: 6 (6 – 9, n = 8) in length and 6 (6, n = 8) in width; common sperm duct 38 in length and 9 in width; does not go beyond the middle of the genital fork; individual sperm duct 122 in length and 6 in width, showing a callus in the apical third. The individual and common sperm ducts are membranous with smooth walls, and the individual duct width increases as it approaches the body of the spermatheca (Fig. 16 M). Cercus 174 (162 – 177, n = 9) in length and 72 (70 – 75, n = 9) in width. Male: unknown.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2643720FF1A64FFFDB1F850.taxon	materials_examined	Material examined: 1 female (87106) holotype (Nº 92132). Capture location: Sítio dos Stur, Córrego Palmital, in the municipality of Pancas (19 ° 13 ′ 44 ″ S, 40 ° 45 ′ 31 ″ W), located at the state of Espírito Santo, Brazil. Collection date: 24 / vi / 2017. Trap: HP. Collector: Israel de Souza Pinto. 8 females paratypes (92133, 92134, 92135, 92136, 92137, 92138, 92139, 92140), same data of the female holotype. Collection date: 15 / iv / 2017 – 24 / vi / 2017. Material deposited in the FIOCRUZ / COLFLEB.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2643720FF1A64FFFDB1F850.taxon	etymology	Etymology: The species name pays homage to Claudiney Biral dos Santos, in recognition of his significant contribution to systematics of Psychodidae. The species was described in collaboration with the Dr. Israel de Souza Pinto of the “ Instituto Federal de Educação, Ciência e Tecnologia do Pará ”.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
F00D87CAB2643720FF1A64FFFDB1F850.taxon	distribution	Distribution: BRAZIL. Espírito Santo: Pancas. Medical importance: Studies have not yet been conducted on this species regarding vector competence or capacity, or natural infection by parasites.	en	Chaves Júnior, Salvador P., Shimabukuro, Paloma H. F., Andrade, Andrey J. (2022): Description of the female and redescription of the male of Sciopemyia sordellii (Shannon & Del Ponte, 1927), including the description of four new species of the genus Sciopemyia Barretto, 1962 (Diptera: Psychodidae: Phlebotominae) from Brazil. Zootaxa 5195 (4): 301-336, DOI: 10.11646/zootaxa.5195.4.1
