taxonID	type	description	language	source
5F877B8DC6A85A1FB76CB46C08BADE22.taxon	description	Figs 2, 3, 4	en	Pham, Cuong The, Hoang, Chung Van, Phan, Tien Quang, Nguyen, Truong Quang, Ziegler, Thomas (2022): Hidden in the jungle of Vietnam: a new species of Quasipaa (Amphibia, Anura, Dicroglossidae) from Ngoc Linh Mountain. ZooKeys 1124: 23-42, DOI: http://dx.doi.org/10.3897/zookeys.1124.89282, URL: http://dx.doi.org/10.3897/zookeys.1124.89282
5F877B8DC6A85A1FB76CB46C08BADE22.taxon	diagnosis	Diagnosis. Both morphological characters (body very stout, skin rough with dermal ridges and tubercles, forelimbs of males strongly enlarged, with inner side of arms or fingers or chest and belly with black spines (see Fei et al. 2009) and molecular data revealed the new species to be nested within Quasipaa. Quasipaa taoi sp. nov. is distinguishable from its congeners by a combination of the following morphological characters: (1) SVL 79.6 - 84.3 mm in males, 64.6 - 69.9 mm in females; (2) head broader than long (HL / HW 0.90 in males, 0.92 in females); (3) vomerine teeth present; (4) external vocal sacs absent; (5) tympanum slightly visible; (6) dorsum with lines of thick ridges and small round tubercles; (7) flanks covered by oval and round tubercles; (8) supratympanic fold present; (9) dorsolateral fold absent; (10) ventrolateral sides, ventral surface of arms, and all fingers with spines in males; (11) the absence of spines on chest and belly in males; (12) toes fully webbed to distal end of terminal phalanx; (13) in life, dorsum dark brown, chest and belly immaculate white.	en	Pham, Cuong The, Hoang, Chung Van, Phan, Tien Quang, Nguyen, Truong Quang, Ziegler, Thomas (2022): Hidden in the jungle of Vietnam: a new species of Quasipaa (Amphibia, Anura, Dicroglossidae) from Ngoc Linh Mountain. ZooKeys 1124: 23-42, DOI: http://dx.doi.org/10.3897/zookeys.1124.89282, URL: http://dx.doi.org/10.3897/zookeys.1124.89282
5F877B8DC6A85A1FB76CB46C08BADE22.taxon	description	Description of holotype. A large frog (SVL 84.3 mm); habitus robust with enlarged head (HL / SVL 0.40, HW / SVL 0.43); head broader than long (HL 33.5 mm, HW 36.3 mm); snout round anteriorly in dorsal view, projecting beyond lower jaw; nostril lateral, closer to eye than to the tip of snout (NS 7.6 mm, EN 5.5 mm); canthus rostralis indistinct; loreal region oblique and slightly concave; rostral length greater than eye diameter (RL 13.1 mm, ED 9.7 mm); internarial distance wider than interorbital distance and upper eyelid width (IND 8.6 mm, IOD 6.2 mm, UEW 7.7 mm); tympanum slightly visible (TYD 4.1 mm) smaller than the distance from tympanum to eye (TYE 4.9 mm); vomerine teeth in two oblique ridges; tongue cordiform, notched posteriorly; external vocal sac absent. Forelimbs: arms short; upper arm length (UAL) 17.1 mm, forearm length (FAL) 41.5 mm; relative finger lengths: II <I <IV <III; fingers free of webbing; dermal ridge on sides of fingers present on fingers I, II, III; tips of fingers swollen, not expanded; subarticular tubercles prominent, round, formula 1, 1, 2, 2; inner metatarsal tubercle round; outer metatarsal tubercle elongate; finger I with nuptial pad. Hindlimbs: tibia length longer than thigh length (FeL 44.2 mm, TbL 49.7 mm), approximately 3.4 times longer than wide (TbW 14.5 mm); tips of toes swollen, slightly round; relative length of toes: I <II <V <III <IV; toes fully webbed to distal end of terminal phalanx; dermal ridge present on outer sides of toes I and V; subarticular tubercles prominent, elongate, formula 1, 1, 2, 3, 2; inner metatarsal tubercle elongate; outer metatarsal tubercle absent; tibio-tarsal articulation reaching to tip of snout. Skin texture in life: dorsal surface of head with oval and round tubercles, dorsum with six lines of thick ridges intermixed with small round tubercles; flanks covered by oval and round tubercles; supratympanic fold distinct, extending from eye to angle of jaw; dorsolateral fold absent; dorsal surface of forelimbs and hindlimbs with small tubercles; belly and ventral surface of thighs smooth. Nuptial spines: body of males with spines except for chest, belly, and ventral surface of hindlimbs; dense spines on dorsum, flanks, ventral surface of forelimbs, ventrolateral sides, and fingers I, II; spines present on throat, dorsal surface of fore- and hindlimbs, and fingers III, IV, small and scattered. Coloration in life: iris dark copper; dorsum and upper part of flanks dark brown; lower part of flanks whitish brown with white tubercles and black spines on top; supratympanic fold dark brown; dorsal surface of limbs yellowish brown with dark crossbars; ventral surface of limbs light yellow with brown markings; throat white with brown markings; chest and belly immaculate white; toe webbing dark brown. Coloration in preservative: coloration in preservative is the same in life but somewhat faded.	en	Pham, Cuong The, Hoang, Chung Van, Phan, Tien Quang, Nguyen, Truong Quang, Ziegler, Thomas (2022): Hidden in the jungle of Vietnam: a new species of Quasipaa (Amphibia, Anura, Dicroglossidae) from Ngoc Linh Mountain. ZooKeys 1124: 23-42, DOI: http://dx.doi.org/10.3897/zookeys.1124.89282, URL: http://dx.doi.org/10.3897/zookeys.1124.89282
5F877B8DC6A85A1FB76CB46C08BADE22.taxon	etymology	Etymology. The new species is named in honor of our colleague and friend, Assoc. Prof. Dr. Tao Thien Nguyen from the Institute of Genome Research, Vietnam Academy of Science and Technology, in recognition of his numerous scientific contributions towards a better understanding of the amphibians of Vietnam. We recommend " Tao's Spiny Frog " as the common English name of the new species and the common name in Vietnamese as " Ếch gai s ần t ạo ".	en	Pham, Cuong The, Hoang, Chung Van, Phan, Tien Quang, Nguyen, Truong Quang, Ziegler, Thomas (2022): Hidden in the jungle of Vietnam: a new species of Quasipaa (Amphibia, Anura, Dicroglossidae) from Ngoc Linh Mountain. ZooKeys 1124: 23-42, DOI: http://dx.doi.org/10.3897/zookeys.1124.89282, URL: http://dx.doi.org/10.3897/zookeys.1124.89282
5F877B8DC6A85A1FB76CB46C08BADE22.taxon	distribution	Distribution. Quasipaa taoi sp. nov. is currently known from Ngoc Linh Mountain of the Central Highlands in Vietnam (Fig. 6). Data obtained from GenBank show that this species was also recorded from Xekong Province, Lao PDR (Yan et al. 2021; see Discussion below).	en	Pham, Cuong The, Hoang, Chung Van, Phan, Tien Quang, Nguyen, Truong Quang, Ziegler, Thomas (2022): Hidden in the jungle of Vietnam: a new species of Quasipaa (Amphibia, Anura, Dicroglossidae) from Ngoc Linh Mountain. ZooKeys 1124: 23-42, DOI: http://dx.doi.org/10.3897/zookeys.1124.89282, URL: http://dx.doi.org/10.3897/zookeys.1124.89282
