taxonID	type	description	language	source
3E9A836F7D0F594AB3F0259B6D80B1CE.taxon	description	Figs 3 A-G, 4 A-D, 5 A-C, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
3E9A836F7D0F594AB3F0259B6D80B1CE.taxon	diagnosis	Diagnosis. Male genitalia are required for identification: lateral articular areas of tectum pointed and strongly projected distally (Fig. 3 E); paramera in dorsal view slightly constricted medially, with middle portion almost as wide as apical portion (Fig. 3 E); apex of paramera in caudal view obtusely triangulate, with a strong median tooth (Fig. 3 F, G).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
3E9A836F7D0F594AB3F0259B6D80B1CE.taxon	description	Description. Holotype male (Fig. 3 A, B, E-G). Length 10.6 mm, width 5.9 mm. Body rounded-oval. Coloration. Head, elytra, and scutellar shield shiny green, with brownish reflections. Pronotum shiny green, anterolateral areas with brownish reflection, and posterolateral areas with yellow maculae laterally extending to anterior margins. Pygidium and venter shiny green with strong brownish reflections. Head. Vertex sparsely punctate on disc, laterally punctostriate. Frons with slight V-shaped depression, moderately punctate, punctures moderate and deep. Interocular width 3.7 times wider than transverse eye diameter. Clypeus confluently punctate, with anterior margin subtrapezoidal, slightly raised medially. Mandible with outer teeth strongly raised, outer margin slightly curved near base. Pronotum shallowly and sparsely punctate on disc, punctures small and shallow; anterolaterally punctostriate, punctures large and deep. Scutellar shield moderately punctate, longer than elytral suture. Elytra 2 times longer than mid-width, moderately punctate, punctuations large and shallow. Posthumeral depression well developed laterally. Apical umbone wide and poorly defined. Pygidium strongly convex, with weak and concentric sculpturing, slightly effaced posteriorly. Venter glabrous, moderately punctate. Mesometaventral process anteriorly directed between procoxae, ventrally flat, with apex abruptly acute in anteroventral view. Mesepimera partially exposed in dorsal view, strongly convex and transversally ridged. Legs. Protibia externally tridentate, with proximal tooth well defined and acute. Protarsomere V longer than protarsomeres I-IV combined. Anterior protarsal claw enlarged, unequally bifid and obliquely truncated. Mesotibia with internal margin straight, with inner apex not dilated. Mesotarsomere IV with ventroapical projection well developed, thickened and ventrally swollen. Abdomen with ventrite 6 broadly and slightly emarginated posteriorly. Aedeagus (Fig. 3 E-G). Tectum abruptly narrowed towards the apical edge, with lateral articular areas pointed and strongly projected distally. Paramera in dorsal view slightly constricted medially, with middle portion almost as wide as apical portion; apex in caudal view triangulated with a strong median tooth, strongly deflected ventrally. Endophallus (Fig. 4 A-D) divided into three portions: one narrow, tube-shaped basal portion; one wide, sac-shaped medial portion; and one hairy, slender apical portion (partially lost in Fig. 4 A, B, D). Proximal portion distally hairy; V-shaped sclerite with thin, long arms; and temones large, fused into a single sclerite with a mediolongitudinal carina. Medial portion with a broad ventral raspula and a small dorsomedial raspula bearing moderately dense, thin-walled asperites; a dorsodistal raspula bearing multiple, irregular, and dense rows of thick-walled asperites; and a large, triangular lateral sclerite, with distal edge thick and slightly raised. Paratype (1 female) (Fig. 3 C, D). Length 11.2 mm. Width 6.3 mm. The female differs from male by the more robust and more convex body; interocular width 4.2 times wider than transverse eye diameter; clypeus longer, with anterior margin narrower and more raised; pygidium plano-convex; protibia with outer teeth stronger and apically rounded; Mesotarsomere IV with a short ventroapical projection straight and pointed; and abdominal ventrite 6 not emarginated. External genitalia (Fig. 5 A-C). Gonocoxites dark brown, strongly sclerotized and moderately setose apically, setae moderately long. Proximal gonocoxites rugostriate and large, as long as wide, overlapping the distal gonocoxites; inner margin abruptly deflected to apex narrow. Distal gonocoxites with inner margin curved and apex narrowly rounded.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
3E9A836F7D0F594AB3F0259B6D80B1CE.taxon	etymology	Etymology. This species is named after the Brazilian zoologist Paulo Friederich Buehrnheim (1937 - 2001), who greatly contributed to education and research in Amazonas state, Brazil. In addition, he founded the insect collection at the Universidade Federal do Amazonas (UFAM) and collected the type series of this species.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
3E9A836F7D0F594AB3F0259B6D80B1CE.taxon	distribution	Distribution (Fig. 12). Brazil (2). Amazonas: Coari.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
6209BC0395425304AE0FD2B0C1AA67E9.taxon	description	Figs 1 O, 3 H-J, 4 E-G, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
6209BC0395425304AE0FD2B0C1AA67E9.taxon	materials_examined	Additional material examined (2 males). Brazil, Roraima, Caracarai, Parque Nacional da Serra da Mocidade, 15 - 26. I. 2016, F. F. Xavier, R. Boldrini, & P. Barroso (legs.) (♂, INPA); Brazil, Amazonas, Manaus, Fazenda Dimona - PDBFF, 19. VIII. 2000, R. Andreazze (leg.) (♂, INPA).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
6209BC0395425304AE0FD2B0C1AA67E9.taxon	distribution	Distribution (Fig. 12). Colombia (1). Meta (Neita-Moreno 2014). Brazil (2). Roraima, Amazonas.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
F08D8D808CF25F469A361D04E7FB1650.taxon	description	Figs 6 A-D, 7 A-E, 10 H-J, 11 A-D, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
F08D8D808CF25F469A361D04E7FB1650.taxon	distribution	Distribution (Fig. 12). Suriname (Hielkema and Hielkema 2019). French Guiana. Guiana. Venezuela (Soula 1998, 2002, 2005). Brazil. Amazonas, Para (Ohaus 1898; present paper).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
6588A24D1B2659B9ADD46C0850DDA833.taxon	description	Figs 2 E, 8 A-D, 9 I-M, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
6588A24D1B2659B9ADD46C0850DDA833.taxon	materials_examined	Additional material examined (18 males, 13 females). Brazil, Bahia, Prado, 05. III. 1971, C. Elias (leg.) (♀, DZUP); Brazil, Espirito Santo, Conceicao da Barra, 04. X. 1969, C. T. & C. Elias (legs.) (♂, ♀, DZUP); idem, but 26. IX. 1968 (2 ♂, ♀, DZUP); idem, but 19 - 25. XI. 1968 (♂, ♀, DZUP); Brazil, Espirito Santo, Linhares, X. 1965 (♂, DZUP); idem, but XI. 1965 (♀, DZUP); idem, but no date (♂, DZUP); idem, but XI. 1966, A. Maller (leg.) (♂, ♀, DZUP); idem, but Rio Itabapoana, 26. X. 1906, J. F. Zikan Coll. (♀, CEIOC); Brazil, Rio de Janeiro, Rio de Janeiro, Corcovado, XII. 1961, Seabra & Alvarenga (legs.) (♀, DZUP); Brazil, Minas Gerais, Rio Jose Pedro, 28. XI. 1926, J. F. Zikan Coll. (♀, CEIOC); Brazil, Tris. [= Trisanga (= Irisanga)], N. 8 (3 ♂, NHMW).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
6588A24D1B2659B9ADD46C0850DDA833.taxon	distribution	Distribution (Fig. 12). Brazil. Sao Paulo (Irisanga = Oricanga; Burmeister 1844), Rio de Janeiro, Goias (Soula 1998, 2003), Bahia, Espirito Santo, Minas Gerais (new state records).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
DE103152DEDE5E5BB5852C0B156C4700.taxon	description	Figs 1 H, 2 A, G, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
DE103152DEDE5E5BB5852C0B156C4700.taxon	materials_examined	Material examined (4 females). Brazil, Amazonas, Manaus, Fazenda Esteio, 30. VIII. 1984, R. C. Klein (leg.) (♀, INPA); Brazil, Amazonas, BR- 174 Km 18, 11. VIII. 1980, C. Fonseca & E. Binda (legs.) (♀, INPA); Brazil, Amazonas, Manaus, Reserva Ducke, 14. VIII. 1969, A. Faustino (leg.) (♀, INPA); Brazil, Amazonas, Presidente Figueiredo, 21. IX. 1992, Aquino (leg.) (♀, INPA).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
DE103152DEDE5E5BB5852C0B156C4700.taxon	distribution	Distribution. Suriname (Hielkema and Hielkema 2019). French Guiana (Soula 1998). Brazil (Fig. 12). Amazonas (present paper).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
0861E88ABE06553BBFB62F743A10BB46.taxon	description	Figs 9 A-H, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
0861E88ABE06553BBFB62F743A10BB46.taxon	diagnosis	Diagnosis. Apex of pygidium smooth (Fig. 9 C); metatarsomere IV with ventroapical projection short and apex acute; apical edge of tectum wider than base of paramera, with lateral articular areas horizontally rotated (Fig. 9 E-H); paramera strongly declivous laterally, with sides inconspicuous in caudal view and strongly excavated medially (Fig. 9 F-H).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
0861E88ABE06553BBFB62F743A10BB46.taxon	description	Description. Holotype male (Fig. 9 A-G). Length 9.7 mm, width 5.6 mm. Body rounded-oval. Coloration. General color copper with green or red reflections. Head, pronotum, and scutellar shield with strong green reflections. Pronotum with yellow posterolateral maculae effaced. Elytra brownish copper, with two median yellow maculae somewhat effaced. Pygidium and venter with color more diffuse than dorsal surface. Head. Vertex sparsely punctate at disc, laterally punctostriate. Frons with slight V-shaped depression, densely punctate, punctures large. Interocular width 4.2 times wider than transverse eye diameter. Clypeus confluently punctate, with anterior margin subtrapezoidal, slightly raised medially. Pronotum shallowly and sparsely punctate at disc, with slight anterolateral depression densely punctate at lateral corner, punctures moderate and deep. Scutellar shield moderately punctate, longer than elytral suture. Elytra 1.8 times longer than mid-width, moderately punctate, punctuations moderate and shallow. Posthumeral depression weak. Apical umbone wide and well defined. Pygidium (Fig. 9 C) strongly convex and apically smooth. Venter glabrous, moderately punctate. Mesometaventral process anteriorly directed between procoxae, ventrally flat, with apex abruptly acute in anteroventral view. Mesepimera partially exposed in dorsal view, slightly convex and transversally ridged. Legs. Protibia strongly tridentate externally, with proximal tooth well defined and pointed. Protarsomere V longer than protarsomeres I-IV combined. Anterior protarsal claw enlarged, unequally cleft and obliquely truncated. Mesotibia with internal margin straight, with inner apex not dilated. Mesotarsomere IV with ventroapical projection well developed, thickened and ventrally curved. Metatarsomere IV with spine-like ventroapical projection short and pointed. Abdomen with ventrite 6 broadly and slightly emarginated posteriorly. Spiculum gastrale (Fig. 9 D) Y-shaped with proximal arms short and distal stem slender, about 2.8 times longer than arm length. Aedeagus (Fig. 9 E-G). Tectum broadly curved laterally, not narrowed towards the apical third; apical edge wider than base of paramera, with lateral articular areas horizontally rotated. Paramera strongly declivous laterally, with sides inconspicuous in caudal view and strongly excavated medially to form a laterolongitudinal carina. Female. Unknown. Paratype (1 male). The male paratype differs from holotype by the general coloration darker with stronger green reflections and elytra without yellow maculae; paramera slightly narrower, with lateral margins parallel and apex slightly more rounded (Fig. 9 H).	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
0861E88ABE06553BBFB62F743A10BB46.taxon	etymology	Etymology. The specific epithet is derived from the Latin word ' opalus ' (= precious stone) in reference to opal gemstone, alluding to the metallic, multicolor cuticular surface.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
0861E88ABE06553BBFB62F743A10BB46.taxon	distribution	Distribution (Fig. 12). Brazil (2). Para: Itaituba, Rondonia: Candeias do Jamari. The two male specimens composing the type series were collected within the Madeira-Tapajos interfluvium.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
1DFAB3780AE1562E85F89A42ED723B33.taxon	description	Figs 1 L, 5 D-G, 10 A-G, 11 E-H, 12	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
1DFAB3780AE1562E85F89A42ED723B33.taxon	diagnosis	Diagnosis. Male: lateral articular areas of tectum thickened and deflected outward (Fig. 10 E); paramera in caudal view rounded-oval, with apex rounded to slightly constricted and sides not declivous (Fig. 10 F). Female: Females of this species are diagnosed in association with male specimens.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
1DFAB3780AE1562E85F89A42ED723B33.taxon	description	Description. Male holotype (Fig. 10 A, B, E-G). Length 11.3 mm, width 6.1 mm. Body rounded-oval. Coloration. General color shiny green with brownish reflections. Pronotum with posterolateral yellow maculae laterally extending to anterior margins. Head. Vertex sparsely punctate at disc, laterally punctostriate. Frons with slight V-shaped depression, densely punctate, punctures moderate and deep. Interocular width 4 times wider than transverse eye diameter. Clypeus confluently punctate, with anterior margin subtrapezoidal, slightly raised medially. Pronotum shallowly and sparsely punctate at disc and moderately punctate anterolaterally, punctures moderate and deep. Scutellar shield moderately punctate, longer than elytral suture. Elytra 2.3 times longer than wide, moderately punctate, punctures large. Posthumeral depression weak. Apical umbone wide and poorly defined. Pygidium strongly convex, with concentric sculpturing and moderately punctate posteriorly. Venter glabrous, moderately punctate. Mesometaventral process anteriorly directed between procoxae, ventrally flat, with apex abruptly acute in anteroventral view. Mesepimera partially exposed in dorsal view, slightly convex and transversally ridged. Legs. Protibia externally tridentate, with proximal tooth well defined. Protarsomere V enlarged, longer than protarsomeres I-IV combined. Anterior protarsal claw enlarged, unequally cleft and obliquely truncated. Mesotibia with internal margin straight, with inner apex not dilated. Mesotarsomere IV with ventroapical projection well developed, thickened and ventrally curved. Abdomen with ventrite 6 broadly and slightly emarginated posteriorly. Aedeagus (Fig. 10 E-G). Tectum moderately narrowed towards the apical edge, with lateral articular areas thickened and deflected outward. Paramera in caudal view rounded-oval, with apex parabolic and sides not declivous. Endophallus (Fig. 11 E-H) divided into three portions: one narrow, tube-shaped basal portion; one wide, sac-shaped medial portion; and one hairy, slender apical portion. Medial portion with a large ventral raspula and a small dorsoproximal raspula bearing dense, thin-walled asperites; a dorsodistal raspula bearing an irregular, dense multiple rows of thick-walled asperites; and a large and cultrate lateral sclerite, with proximal and distal edges thick and slightly raised. Paratypes (3 males, 14 females): male paratypes differ from holotype in length (10.7 - 11.7 mm), width (5.8 - 6.7 mm), and form of the apex of paramera (more round to slightly narrowed). Female paratypes (Fig. 10 C, D). Length 10.8 - 11.8 mm, width 6 - 6.4 mm. The females differ from males by the clypeus longer, anteriorly narrower and more raised; pygidium plano-convex; protibia with outer teeth stronger and apically rounded; protarsomere V simple, with anterior claw unenlarged and equally cleft; inner metatibial spur apically rounded; Mesotarsomere IV with a ventroapical projection short and straight; and abdominal ventrite 6 not emarginated. External genitalia (Fig. 5 D-F). Gonocoxites light brown, slightly sclerotized and moderately setose apically, setae moderately long. Proximal gonocoxites short and semicircular, wider than long, barely overlapping the distal gonocoxites; surface moderately punctate. Distal gonocoxites broadly rounded apically, with inner margin almost straight.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
1DFAB3780AE1562E85F89A42ED723B33.taxon	etymology	Etymology. The specific epithet is Greek for ' phallos ' (= penis) and ' kardia ' (= heart), refers to the heart-shaped male paramera of this species.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
1DFAB3780AE1562E85F89A42ED723B33.taxon	distribution	Distribution (Fig. 12). Brazil (10). Rondonia: Ouro Preto d'Oeste.	en	Bento, Matheus, Jameson, Mary Liz, Seidel, Matthias (2022): New species and illustrated key of Macraspis (Scarabaeidae, Rutelinae, Rutelini) from the Amazon biome of Brazil. ZooKeys 1124: 161-189, DOI: http://dx.doi.org/10.3897/zookeys.1124.91156, URL: http://dx.doi.org/10.3897/zookeys.1124.91156
