taxonID	type	description	language	source
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	description	urn: lsid: ipni. org: names: 77305496 - 1 Figs 1 – 2	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	diagnosis	Diagnosis Croton sertanejus sp. nov. differs from C. echioides due to pseudomonopodial branching, with an elongated main stem with shortened secondary branches in an alternate arrangement (vs di- or trichotomous branching in C. echioides), petiole 0.4 – 1.7 (– 2) cm long (vs (1.8 –) 3 – 8.1 cm), leaf blade base with 4 – 6 subsessile nectary glands facing both surfaces (vs with 2 stipitate nectary glands facing the abaxial surface), equal pistillate sepals (vs subequal or strongly unequal), and styles 2 – 3 mm long (vs 4 – 5.5 mm).	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	etymology	Etymology ʻ Sertanejo ʼ, in Portuguese, refers to people native to the Caatinga region, also known as ʻ sertão ʼ, that extends to all the states of northeast Brazil and the northern Minas Gerais.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	materials_examined	Material examined Type BRAZIL • Bahia, Oliveira dos Brejinhos, 6 km ao Sul da cidade pela vicinal “ Corredor da Barra ”; 12 ° 22 ′ 02.7 ″ S, 42 ° 54 ′ 14.3 ″ W; 884 m a. s. l.; 20 Dec. 2017; fl.; R. C. Sodré, A. O. Souza & U. S. Amaral 3350; holotype: BOTU; isotypes: CEPEC, NY, TEPB, UFG. Paratypes BRAZIL – Bahia • Barra, Conglomerado BA- 360; 10 ° 58 ′ 48 ″ S, 44 ° 33 ′ 36 ″ W; 20 Dec. 2017; fl.; A. S. Soares 401; RB, UB • Barra do Mendes, Carretão, estrada para Canarina; 11 ° 49 ′ 33 ″ S, 42 ° 3 ′ 17 ″ W; 750 m a. s. l.; 15 Dec. 2009; fl.; E. Melo, F. França & B. M. Silva 7505; ALCB, HUEFS • ibid.; E. Melo, F. França & B. M. Silva 7507; HUEFS • ca 10 km na estrada de terra para Seabra; 11 ° 53 ′ 41 ″ S, 42 ° 2 ′ 53 ″ W; 750 m a. s. l.; 16 Dec. 2009; fl.; E. Melo, F. França & B. M. Silva 7539; HUEFS • ibid.; E. Melo, F. França & B. M. Silva 7541; ALCB, HUEFS • ca 16.5 km na estrada de terra para Seabra, Lagoa do Peixe; 11 ° 56 ′ 22 ″ S, 42 ° 2 ′ 3 ″ W; 743 m a. s. l.; 16 Dec. 2009; fl.; E. Melo et al. 7563; HUEFS • ca 27 km na estrada de terra para Seabra; 12 ° 1 ′ 13 ″ S, 41 ° 59 ′ 58 ″ W; 750 m a. s. l.; 16 Dec. 2009; fl.; E. Melo, F. França & B. M. Silva 7580; ALCB, HUEFS • Brotas de Macaúbas, estrada para a BR- 242; 12 ° 17 ′ 7 ″ S, 42 ° 34 ′ 7 ″ W; 519 m a. s. l.; 30 Jan. 2008; fl.; J. L. Ferreira, E. Melo & B. M. Silva 266; HUEFS • ibid.; J. L. Ferreira, E. Melo & B. M. Silva 268; HUEFS • Brumado; 28 Oct. 1965; fl.; A. P. Duarte 9544; HUEFS, NY, RB, SPF • Caetité; 15 Jan. 1983; fl.; M. L. Guedes et al. 2787; ALCB • ramal a 29 km na estrada Caetité / Brumado, ca 3 km ramal adentro; 19 Feb. 1992; fr.; A. M. Carvalho et al. 3784; CEPEC, NY • Campo Formoso; 21 Apr. 1981; fr.; W. N. Fonseca 386; RB • Canudos, Bedengó, Ramal a 5 km da BR- 115, sentido Euclides da Cunha; 10 ° 4 ′ 42 ″ S, 39 ° 7 ′ 46 ″ W; 560 m a. s. l.; 24 Jun. 2005; no fl. or fr.; D. S. Carneiro-Torres et al. 514; HUEFS • Carfanaum, entre Lapa Cercada e Lapa da Igreja, proximidades do povoado de Cercadinho; 11 ° 44 ′ 17.71 ″ S, 41 ° 27 ′ 20.55 ″ W; 748 m a. s. l.; 19 Mar. 2016; fl. and fr.; P. H. A. Melo & F. F. Pezzini 4885; HUEFS • ibid.; 11 ° 44 ′ 2.4 ″ S, 41 ° 27 ′ 41.7 ″ W; 731 m a. s. l.; 9 Apr. 2017; fl.; P. H. A. Melo & L. C. A. Gonçalves 6744; HUEFS • ibid.; P. H. A. Melo & L. C. A. Gonçalves 6752; HRCB • margem esquerda da BR- 122, a 22 km de Cafarnaum, sentido Morro do Chapéu; 11 ° 31 ′ 58.7 ″ S, 41 ° 23 ′ 12.2 ″ W; 737 m a. s. l.; 10 Apr. 2017; fl.; P. H. A. Melo & L. C. A. Gonçalves 6783; HRCB, HUEFS • ibid.; P. H. A. Melo & L. C. A. Gonçalves 6792; HRCB, HUEFS • ibid.; P. H. A. Melo & L. C. A. Gonçalves 6794; HRCB, HUEFS • Contendas do Sincorá, estrada entre Sussuarana e Contendas do Sincorá; 13 ° 48 ′ 56 ″ S, 41 ° 3 ′ 57 ″ W; 404 m a. s. l.; 15 Mar. 2015; fr.; A. G. Azevedo-Neto et al. 14; HUEFS • Curaçá, Comunidade de Patamuté; 9 ° 25 ′ 19 ″ S, 39 ° 28 ′ 13 ″ W; 420 m a. s. l.; 23 Mar. 2012; fl.; E. Melo et al. 11078; HUEFS • Glória, Complexo Itaparica, margem do lago; 9 ° 4 ′ 11 ″ S, 38 ° 24 ′ 13 ″ W; 333 m a. s. l.; 14 Jun. 2007; fl.; M. Oliveira et al. 2831; MAC, UFP • Iaçu, Fazenda Lapa, Lagedo Alto; Apr. 1974; fl.; G. C. P. Pinto 42272; ALCB • rodovia Itaberaba / Milagres, entre Iaçu e Milagres, a 10 – 20 km a E de Iaçu, 400 – 500 m a. s. l.; 25 Dec. 1979; fl.; S. A. Mori & F. P. Benton 13243; CEPEC, NY • BA- 046, trecho entre Milagres e Iaçu, a 25 km ao SE de Iaçu, 300 m a. s. l.; 28 Feb. 1980; fl. and fr.; S. A. Mori 13272; CEPEC, NY • Fazenda Lapa; 12 ° 42 ′ S, 39 ° 56 ′ 1 W; 26 Feb. 1983; fl. and fr.; H. P. Bautista 716; HUEFS • Fazenda Suíbra, Morro do Gado Bravo; 12 ° 43 ′ S, 40 ° 7 ′ W; 14 Mar. 1985; fl. and fr.; L. R. Noblick 3687; CEPEC, HUEFS • rodovia Iaçu / Milagres (BA- 046) km 13, coletas no início do ramal para as fazendas Suíbra e Morros; 12 ° 49 ′ S, 40 ° 10 ′ W; 240 m a. s. l.; 22 Mar. 1988; fl.; S. Ginzbarg, L. A. Mattos-Silva & H. S. Brito 803; CEPEC, NY • Morro da Garrafa; 12 ° 45 ′ 18 ″ S, 39 ° 51 ′ 0 ″ W; 250 m a. s. l.; 23 Feb. 1997; fr.; E. Melo et al. 2116; HUEFS, HUESB, SP • Ibipeba, estrada para Ibititá; 11 ° 37 ′ 6 ″ S, 42 ° 0 ′ 3 ″ W; 742 m a. s. l.; 25 Oct. 2009; fr.; E. Melo et al. 6881; ALCB, HUEFS • Irecê, Lapão, Morro Pelado; 11 ° 24 ′ 22 ″ S, 41 ° 49 ′ 22 ″ W; 787 m a. s. l.; 27 Oct. 2009; fr.; M. L. Guedes et al. 16178; ALCB, HUEFS • Área do CETEP; 11 ° 19 ′ 0 ″ S, 41 ° 49 ′ 49 ″ W; 22 Feb. 2016; fl. and fr.; G. Costa & A. C. S. Moraes 1656; ALCB, HUEFS, HURB • 11 ° 19 ′ 0 ″ S, 41 ° 49 ′ 42 ″ W; 25 Feb. 2016; fr.; G. Costa & A. C. S. Moraes 1742; ALCB, HUEFS, HURB • ibid.; G. Costa & A. C. S. Moraes 1746; ALCB, HUEFS, HURB • Itatim, Morro do Agenor; 12 ° 43 ′ S, 39 ° 46 ′ W; 310 – 430 m a. s. l.; 25 Feb. 1996; fl.; E. Melo et al. 1483; CEN, HUEFS, MAC • Morro da Quixaba; 12.7119 ° S, 39.6981 ° W; 410 m a. s. l.; 14 Dec. 1996; fl.; E. Melo et al. 1889; HUEFS, SP, VIES • Morro do Bastião; 12 ° 45 ′ 12 ″ S, 39 ° 46 ′ 59 ″ W; 282 m a. s. l.; 25 Jan. 1997; fl.; E. Melo et al. 1919; HUEFS, HUESB, UEC • ibid.; 24 May 1997; fl.; V. M. Monteiro, C. Aguiar & J. Resende 68; HUEFS • Itiúba, Serra de Itiúba, about 6 km E of Itiúba; 10 ° 41 ′ S, 39 ° 48 ′ W; 500 m a. s. l.; 19 Feb. 1974; fl.; R. M. Harley 16214; RB • Fazenda Experimental da EPABA; 10 ° 43 ′ S, 39 ° 48 ′ W; 27 May 1983; fl. and fr.; G. C. P. Pinto & H. P. Bautista 109 / 83; CEPEC, K, NY • 10 km da cidade de Itiúba; 10 ° 41 ′ 14 ″ S, 39 ° 48 ′ 17 ″ W; 24 Jun. 2005; no fl. or fr.; D. S. CarneiroTorres et al. 531; HUEFS • Ituaçu, Morro da Mangabeira; 22 Dec. 1983; fl.; E. P. Gouveia 60 / 83; ALCB • ibid.; E. P. Gouveia s. n.; ALCB [51564 [• Jacobina; 8 Apr. 1991; fr.; M. Andrade Neto s. n.; EAC [17121], K [001184431], NY [01281082] • Jaguari, Fazenda Pé de Laje, próximo ao povoado de Gameleira; 10 ° 12 ′ 52.4 ″ S, 40 ° 8 ′ 8.2 ″ W; 644 m a. s. l.; 18 Apr. 2017; fl.; E. D. S. Almeida & J. G. B. Silva 81; HVASF, HUEFS • Jeremoabo, estrada de chão sentido Canudos; 10 ° 4 ′ 7 ″ S, 38 ° 28 ′ 21 ″ W; 297 m a. s. l.; 30 Jun. 2007; no fl. or fr.; D. S. Carneiro-Torres et al. 981; HUEFS • Jequié, estrada da Barragem de Pedra; 5 Apr. 1987; fl.; V. Pereira 99; HUEFS • Juazeiro, Serra do Mulato; 9 ° 44 ′ 40 ″ S, 40 ° 40 ′ 39 ″ W; 26 Mar. 2000; fl.; N. G. Jesus et al. 880; ALCB, SPF • Lagoa Real; 565 m a. s. l.; 17 Dec. 2008; fl.; C. Novello 1; HUEFS, HUEM • ibid.; C. Novello 2; HUEFS, HUEM • Lajedinho, proximidade do povoado de Simpatia; 12 ° 27 ′ 7.36 ″ S, 41 ° 0 ′ 41.52 ″ W; 593 m a. s. l.; 17 Mar. 2016; fl.; P. H. A. Melo & F. F. Pezzini 4833 A; HRCB • 12 ° 27 ′ 18.9 ″ S, 41 ° 0 ′ 33.3 ″ W; 591 m a. s. l.; 8 Apr. 2017; fl.; P. H. A. Melo & L. C. A. Gonçalves 6691; HUEFS • ibid.; P. H. A. Melo & L. C. A. Gonçalves 6734; HRCB, HUEFS • Livramento do Brumado, 3 – 5 km da cidade, na estrada para Rio de Contas; 13 ° 38 ′ S, 41 ° 50 ′ W; 580 – 600 m a. s. l.; 12 Dec. 1988; fl.; R. M. Harley et al. 27140; CEPEC • ibid.; R. M. Harley et al. 27141; K, NY, SPF • Manoel Vitorino, estrada que liga Manoel Vitorino com Caatingal, a 5 km da BR- 116; 350 m a. s. l.; 10 Mar. 1980; fr.; S. Mori 13448; CEPEC, NY • Maracás, estrada para Marcionílio Souza, ca 12.6 km da cidade; 13 ° 14 ′ 20 ″ S, 40 ° 34 ′ 15 ″ W; 395 m a. s. l.; 3 Nov. 2011; fl.; E. Melo et al. 10603; HUEFS • Marcionílio Souza, 8 km ao S de Marcionílio Souza na estrada para Maracás; 13 ° 5 ′ S, 40 ° 30 ′ W; 22 Mar. 1988; fl.; S. Ginzbarg, L. A. Mattos-Silva & H. S. Brito 816; NY • Monte Santo; 10 ° 26 ′ S, 39 ° 19 ′ W; 11 Jan. 2006; fl.; M. L. Guedes et al. 12037; ALCB • ibid.; 22 Jan. 2016; fl.; F. A. Silva 18; HURB • Morpará; 11 ° 35 ′ 50 ″ S, 43 ° 17 ′ 4 ″ W; 16 Dec. 2007; fl.; A. A. Conceição et al. 2652; HUEFS • estrada para Morpará, beira do Rio Paramirim; 11 ° 42 ′ 11 ″ S, 43 ° 14 ′ 2 ″ W; 444 m a. s. l.; 16 Dec. 2007; fl.; A. A. Conceição et al. 2682; HUEFS • ibid.; A. A. Conceição et al. 2686; HUEFS • Morro do Chapéu, ca 2 km da Comunidade Gruta dos Brejões; 11 ° 0 ′ 53 ″ S, 40 ° 24 ′ 19 ″ W; 650 m a. s. l.; 4 May 2007; no fl. or fr.; D. Cardoso & R. M. Santos 1820; HUEFS • estrada para Cafarnaum; 11 ° 29 ′ 50 ″ S, 41 ° 21 ′ 34 ″ W; 728 m a. s. l.; 1 Feb. 2008; fl.; F. B. L. Silva & F. França 78; HUEFS • Oliveira dos Brejinhos, estrada para Água Quente; 12 ° 20 ′ 11 ″ S, 42 ° 54 ′ 29 ″ W; 673 m a. s. l.; 18 Dec. 2007; fl.; A. A. Conceição et al. 2759; HUEFS • 6 km ao Sul da cidade pela vicinal “ Corredor da Barra ”; 12 ° 22 ′ 02.7 ″ S, 42 ° 54 ′ 14.3 ″ W; 884 m a. s. l.; 20 Dec. 2017; fl.; R. C. Sodré et al. 3347; BOTU, UFG • ibid.; R. C. Sodré et al. 3348; BOTU, UFG • ibid.; R. C. Sodré et al. 3349; BOTU, UFG • Palmeiras, 12 ° 26 ′ 48 ″ S, 41 ° 31 ′ 6 ″ W; 697 m a. s. l.; 1 Dec. 2011; fl.; S. P. S. Neves 393; HUEFS • subida para a Serra Negra; 12 ° 32 ′ S, 41 ° 35 ′ W; 740 m a. s. l.; 29 Jan. 2015; fl.; D. S. Carneiro-Torres et al. 1288; HUEFS • 12 ° 31 ′ 51.6 ″ S, 41 ° 35 ′ 17.4 ″ W; 736 m a. s. l.; 19 Dec. 2017; fl.; R. C. Sodré et al. 3341; UFG • Paulo Afonso; 12 Apr. 1952; fl.; Labouriau 941; RB, SPF • Piatã, Chapada Diamantina, subida da Capela Nossa Senhora Santana; 13 ° 9 ′ 2 ″ S, 41 ° 45 ′ 0 ″ W; 15 Jun. 2014; fl.; M. L. Guedes et al. 21918; ALCB • Piemonte da Diamantina, Miguel Calmon, entorno do Parque de Sete Passagens; 11 ° 22 ′ 25 ″ S, 40 ° 35 ′ 14 ″ W; 546 m a. s. l.; 23 Dec. 2006; fl.; M. L. Guedes et al. 13215; ALCB • João Dourada, Ponto 7; 11 ° 19 ′ 51 ″ S, 41 ° 44 ′ 14 ″ W; 363 m a. s. l.; 10 Apr. 2013; fl.; M. L. Guedes et al. 20656; ALCB • Pilão Arcado, sentido Brejo de Zacarias; 9 ° 58 ′ 31 ″ S, 42 ° 49 ′ 12 ″ W; 6 Sep. 2005; no fl. or fr.; D. S. Carneiro-Torres et al. 593; HUEFS • sentido Iú Grande – Pilão Arcado Dunas, Barra do Iú; 10 ° 1 ′ 25 ″ S, 42 ° 48 ′ 15 ″ W; 18 Mar. 2006; fr.; D. S. Carneiro-Torres et al. 621; HUEFS • Quijingue, fazenda Urubu, a ca de 3 km E do povoado Quixaba do Mandacaru; 10 ° 55 ′ 1 ″ S, 39 ° 3 ′ 49 ″ W; 270 m a. s. l.; 14 May 2005; fr.; D. Cardoso 515; HUEFS, NY • Remanso, ca 18 km de Remanso, entrada a direita no sentido Remanso; 9 ° 29 ′ 20 ″ S, 42 ° 14 ′ 54 ″ W; 457 m a. s. l.; 28 Dec. 2001; fl.; T. S. Nunes et al. 690; HUEFS • Rio de Contas, 10 km do Rio de Contas na estrada para Marcolino Moura; 13 ° 36 ′ S, 41 ° 43 ′ W; 500 – 600 m a. s. l.; 15 Nov. 1988; fl.; R. M. Harley, D. J. N. Hind & T. B. Cavalcanti 26445; K, NY, RB, SPF, U • caminho para Lagoa Nova; 13 ° 47 ′ 42 ″ S, 41 ° 46 ′ 44 ″ W; 490 m a. s. l.; 5 Feb. 1997; fr.; E. Saar et al. PCD 5093; ALCB, K, SPF • ibid.; fl. and fr.; L. Passos et al. PCD 5109; ALCB, CEPEC, K, SPF • São Gabriel, fazenda Boa Sorte; 11 ° 1 ′ S, 41 ° 39 ′ W; 798 – 800 m a. s. l.; 3 Apr. 2009; fl.; R. F. Machado, F. F. Rocha & A. F. Rocha 138; HUEFS • estrada para Jussara; 11 ° 9 ′ 54 ″ S, 41 ° 56 ′ 34 ″ W; 770 m a. s. l.; 25 Oct. 2009; fl. and fr.; E. Melo et al. 6956; ALCB, HUEFS • Senhor do Bonfim, Fazenda Rancharia; 24 Mar. 2007; fl.; L. P. Felix 11562; HUEFS • ibid.; L. P. Felix 11581; HUEFS • Sento Sé, base do Morro da Mina; 9 ° 51 ′ 39 ″ S, 42 ° 3 ′ 23 ″ W; 930 m a. s. l.; 19 Dec. 2007; fl. and fr.; J. G. Carvalho-Sobrinho & C. T. Lima 1728; HST, HUEFS • Umburanas, Ourolândia, estrada para Umburanas, estrada de chão para Pedreira Araras; 10 ° 56 ′ 57 ″ S, 41 ° 4 ′ 38 ″ W; 682 m a. s. l.; 26 Jan. 2008; fl.; M. M. Silva-Castro et al. 1435; HUEFS • Parque Nacional Boqueirão da Onça, rio Murim; 10 ° 30 ′ 16.9 ″ S, 41 ° 19 ′ 52.4 ″ W; 744 m a. s. l.; 29 Jan. 2010; fr.; A. P. Fontana et al. 6426; HUEFS, HVASF • Xique-Xique, 15 km S de Xique-Xique, camino a Santo Inácio; 10 ° 58 ′ S, 42 ° 41 ′ W; 400 – 500 m a. s. l.; 19 Feb. 1997; fr.; M. M. Arbo et al. 7519; CEPEC, CTES, NY. – Ceará • Aiuaba; 6.3712 ° S, 40.0923 ° W; 478 m a. s. l.; 21 Feb. 2014; fl.; A. L. B. Sartori et al. 1153; EAC • Campos Sales, Várzea da Vaca; Feb. 1839; fl.; G. Gardner 2441; K [001184426], K [001184427], L [0146109] • Piquet Carneiro, Ibicuã, Miguel Calmon; 17 Apr. 1909; no fl. or fr.; A. Ducke s. n.; MG [2202], RB [187679] • Saboeiro, entre Saboeiro e Aiuaba, no entroncamento da estrada do Algodão; 21 Mar. 1984; fl.; E. Nunes s. n.; EAC [12357], HUEFS [111120] • Tauá, Fazenda Angico; 29 Apr. 1981; fr.; E. Nunes & M. A. Figueiredo s. n.; EAC [10110], HUEFS [106374]. – Minas Gerais • Mato Verde, São João do Bonito, estrada para as lavras na Serra Geral, 2.5 km de São João do Bonito, 12.5 km da rodovia Mato Verde – Monte Azul (BR- 122), próximo ao Córrego do Pé da Serra; 15 ° 18 ′ 21 ″ S, 42 ° 49 ′ 37 ″ W; 580 m a. s. l.; 7 Apr. 2004; fl. and fr.; J. R. Pirani et al. 5466; SPF. – Paraíba • Itaporanga, Fazenda Cafula; 7 ° 13 ′ 52 ″ S, 38 ° 9 ′ 21.6 ″ W; 540 m a. s. l.; 2 Nov. 2015; fl.; P. F. Souza 107; CSTR, EAC • São José dos Cordeiros, RPPN-Fazenda Almas; 7 ° 28 ′ 15 ″ S, 36 ° 53 ′ 51 ″ W; 645 m a. s. l.; 24 Jan. 2003; fl.; Z. G. Quirino & J. R. Lima 3; HUEFS • ibid.; Z. G. Quirino & J. R. Lima 15; HUEFS • Manga geral, Área I, Laje do Cara; 7 ° 29 ′ 21 ″ S, 36 ° 55 ′ 36 ″ W; 16 Feb. 2003; fl.; M. R. Barbosa et al. 2686; HUEFS, JPB • ibid.; M. R. Barbosa et al. 2715; HUEFS, JPB • Taperoá, 5 km após a entrada da estrada para Taperoá; 4 Mar. 1994; fl. and fr.; S. I. da Silva s. n.; PEUFR, SPF [137104]. – Pernambuco • Afogados da Ingazeira, estrada para Afogados da Ingazeira, km 35 da BR- 292; 10 Feb. 1993; fl.; S. I. da Silva 64; PEUFR, SPF • Betânia, Vale dos Navios; 20 Apr. 1955; fl.; A. Guerra 5528; SPF • Vale do Pajéu; 5 Apr. 2002; fr.; S. M. Freitas Neto et al. 1; HUEFS • Bodocó, arredores da cidade; 12 Feb. 1991; fl.; P. Lisboa & C. Silva 4515 - 34; EAC • Buíque, fazenda Laranjeiras; 8 ° 37 ′ 30 ″ S, 37 ° 9 ′ 15 ″ W; 799 m a. s. l.; 22 Feb. 1996; fl.; K. Andrade et al. 306; NY, PEUFR • Custódia, proximidades do Reservatório Copiti, Projeto de Transposição do Rio São Francisco; 8 ° 14 ′ 53.9 ″ S, 37 ° 41 ′ 33.3 ″ W; 526 m a. s. l.; 18 Feb. 2009; fl.; M. Oliveira & J. Cardoso Junior 3763; HVASF, HUEFS • estrada formada pela área de influência da linha de transmissão de alta tensão que acompanha o canal do PISF; 8 ° 12 ′ 16.48 ″ S, 37 ° 41 ′ 52.1 ″ W; 434 m a. s. l.; 14 May 2013; fr.; V. M. Cotarelli & J. Silva 2530; HVASF, HUEFS • ibid.; 8 ° 9 ′ 55.6 ″ S, 37 ° 33 ′ 20.32 ″ W; 513 m a. s. l.; 15 Mar. 2016; fl. and fr.; M. Oliveira, A. P. Fontana & A. C. P. Oliveira 6407; RB • Lagoa Grande, Sertão, Fazenda Planaltino, Rodovia PE- 574; 8 ° 59 ′ S, 40 ° 16 ′ W; 10 Jan. 2018; fl.; M. L. Guedes 30231; ALCB • Petrolândia, trecho do canal da transposição; 8 ° 41 ′ 32.6 ″ S, 38 ° 16 ′ 28.3 ″ W; 391 m a. s. l.; 29 Jan. 2009; fl. and fr.; V. D. Silva et al. 87; HUEFS, HVASF • Petrolina, limite sul do CPATSA; 9 ° 7.5 ′ S, 40 ° 31.11 ′ W; 13 Jan. 1983; fl.; G. Fotius 3304; HTSA, RB • ibid.; 24 Jan. 1983; fl.; G. Fotius 3310; HTSA, RB • área da CODEVASF; 9 ° 9.17 ′ S, 40 ° 29.44 ′ W; 4 Feb. 1983; fl.; G. Fotius 3333; HTSA, RB • CPATSA; 3 Mar. 1993; fl.; S. I. da Silva 114; PEUFR, SPF • Salgueiro; 9 Mar. 2007; fr.; E. R. Silveira s. n.; EAC [39845] • ibid.; 8 ° 7 ′ S, 39 ° 7 ′ W; 3 Mar. 2014; fr.; E. R. Silveira & F. C. L. Pinto s. n.; ALCB [122233] • Santa Maria da Boa Vista, margem da PE 4 em direção à Jutaí; 29 Apr. 1971; fr.; Academia Brasileira de Ciências 406; HUEFS, IPA • Serra Talhada, Estação Experimental do IPA, Pimenteira; 25 Mar. 1995; fl.; A. M. Miranda et al. 2192; MAC, PEUFR, SPF • ibid.; 7.59 ° S, 38.1916 ° W; 30 Jan. 1996; fl.; M. L. Gomes 129; IPA, HUEFS • 3 km SW of road from Serra Talhada to Lagoinha, around permanent plot and flux tower; 7 ° 58 ′ 5 ″ S, 38 ° 23 ′ 3 ″ W; 17 Mar. 2017; fl.; P. W. Moonlight & T. Sarkinen 419; HUEFS • ibid.; 18 Mar. 2017; fl.; P. W. Moonlight & T. Sarkinen 430; HUEFS • Sertânia, Centro da Caprino-ovinocultura do IPA, Pimenteira; 8 ° 4 ′ 25 ″ S, 37 ° 12 ′ 24 ″ W; 160 m a. s. l.; 6 Apr. 2001; fl. and fr.; R. M. Harley & A. M. Giulietti 54177; HUEFS, K • Caroalina, Sítio do Sr. Sibas; Jul. 2003; fl.; L. Melo & J. Sampaio 34; IPA, HUEFS • Verdejante, Vila Produtiva Rural de Pilões; 7 ° 52 ′ 57.23 ″ S, 39 ° 2 ′ 16.13 ″ W; 534 m a. s. l.; 21 Feb. 2013; fl.; F. F. S. Silva 751; HUEFS, HVASF. – Piauí • São João do Piauí, Porfírio; 8 ° 21 ′ S, 42 ° 14 ′ W; 222 m a. s. l.; 14 Feb. 1995; fl. and fr.; F. G. Alcoforado Filho 477; HUEFS, IPA, K, TEPB • ibid.; 14 Apr. 1995; fl. and fr.; F. G. Alcoforado Filho & J. H. Carvalho 484; IPA, K, TEPB • Embrapa Meio Norte; 8 ° 20 ′ 44 ″ S, 42 ° 19 ′ 34 ″ W; 24 Mar. 2015; fl. and fr.; K. N. C. Castro & J. B. A. Souza 471; CEN • São Raimundo Nonato, Humaitá – São Raimundo Nonato; 15 Dec. 1978; fl.; F. M. T. Freire 11; EAC • Barreirinhas – São Raimundo Nonato; 18 Dec. 1978; fl.; A. Fernandes et al. s. n.; EAC [5151] • fazenda Joá; 27 Jan. 1984; fl.; L. Emperaire 2180 a; RB • Parque Nacional Serra da Capivara; 8 ° 25 ′ S, 42 ° 20 ′ W; 8 Dec. 2017; fl.; M. Mizushima, P. Rodrigues & R. M. Santos 218; HUEFS. – Sergipe • Poço Verde, Fazenda Santa Maria da Lage; 10 ° 42 ′ 30 ″ S, 38 ° 11 ′ 0 ″ W; 16 Nov. 2010; no fl. or fr.; E. V. R. Ferreira et al. 213; ASE • Assentamento Santa Maria da Lage; 22 Jul. 2015; fr.; E. V. S. Oliveira 563; ASE, HUEFS • locality unknown [caatingas arbustivo-arbórea e arbórea]; Apr. 1989; fl. and fr.; E. L. Araújo s. n.; SPF [137105].	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	description	Description Shrubs or trees, (1 –) 1.5 – 5 (– 7) m tall, erect, pseudomonopodial branching, with main stem extended bearing lateral alternate branches, frequently with galls; latex scarce, clear; sweetish odor; stems very rigid, cylindric, straight or tortuous, striated or smooth in young branches; whitish, yellowish, or slightly ferrugineous tomentose indumentum of multiradiate or stellate-porrect trichomes with stipe 0.05 – 0.1 mm long, stellate-porrect trichomes with one-plane whorl of 8 – 12 free lateral rays and multiradiate trichomes with two or three whorls of 16 – 30 free lateral rays, in both trichomes lateral rays 0.15 – 0.7 mm long and central ray 0.15 – 0.25 (– 0.7) mm long. Leaves alternate, leaf blades (3 –) 4 – 8 times as long as petioles; stipules 0.5 – 0.8 × 0.15 – 0.3 mm, triangular, lanceolate or linear, with 4 sessile ovoid glands at base and / or one subglobose gland at apex, with stellate-porrect trichomes on both surfaces; petioles 0.4 – 1.7 (– 2) cm long; leaf blades 2.2 – 8.9 × 1.2 – 4.7 cm, oval-elliptic, elliptic, ovate or oval-lanceolate, sometimes lanceolate or narrowly elliptic, membranaceous, base obtuse or rounded, sometimes slightly cordate or truncate with 4 – 6 patelliform nectary glands 0.2 – 0.7 mm diam., subsessile with stipe 0.1 – 0.2 mm long, at least two facing adaxial surface and other two facing abaxial surface; apex acute, obtuse, mucronate or apiculate, sometimes slightly emarginate or acuminate; margin entire or less commonly serrulate with globose sessile glands (colleters); venation brochidodromous with 4 – 8 pairs of secondary veins impressed on adaxial surface and slightly prominent on abaxial surface; discolorous, adaxial surface dark green with hirtellous indumentum of simple, 2 – 4 - radiate or stellate-porrect trichomes, stellate-porrect trichomes with 6 – 11 lateral rays, sessile, lateral rays 0.2 – 0.5 mm long, porrect ray up to 1 mm long; abaxial surface light green with shortly tomentose indumentum of stellate-porrect trichomes with 6 – 8 lateral rays or multiradiate trichomes with 11 – 16 lateral rays, both sessile, lateral rays 0.2 – 0.7 mm long, central ray 0.2 – 0.8 mm long. Thyrses 2.8 – 16 cm long, peduncle 0.4 – 2.5 cm long, terminal, bisexual with 4 – 10 solitary pistillate flowers sparsely, rarely densely, distributed on ¼ – ½ of inflorescence axis, or thyrses unisexual staminate, less commonly unisexual pistillate; staminate cymules with 2 – 4 flowers, staminate flower bracts 0.4 – 1 × 0.4 – 0.7 mm, triangular, oval or oblong, margin entire, with 2 or 3 subglobose sessile glands at base, indumentum of stellate-porrect trichomes on external surface and hirsute indumentum of simple trichomes on margin, glabrous internally; 2 staminate flower bracteoles per cymule, 0.3 – 0.7 × 0.15 – 0.3 mm, triangular or ovate, with glands and indumentum similar to those of bracts; pistillate flower bracts 0.5 – 1 (– 2.2) × 0.3 – 0.8 (– 1) mm, triangular, margin entire, with ovoid or subglobose sessile glands, with sessile stellate-porrect trichomes externally, glabrous or glabrescent internally; 2 bracteoles per pistillate flower, 0.3 – 0.8 × 0.1 – 0.4 mm, linear or triangular, with subglobose glands at base. Staminate flowers 5 – 8 mm long; pedicels 1.6 – 4.7 mm long, glabrous, glabrescent, pubescent or tomentose; calyces 5 (6) - partite, greenish with whitish indumentum, lobes 1.4 – 2 × 0.9 – 1.3 mm, ovate or largely ovate, united in ⅙ – ¼ of length, apex obtuse or acute, dorsal surface with shortly tomentose indumentum of 2 stellate-porrect sessile trichomes, villous at the apex, glabrous ventrally; petals 5 (6), 2 – 2.9 × 0.6 – 1.2 mm, obovate or oblanceolate, apex rounded or obtuse, villous basally along margins at ⅓ – ½ of length; stamens 15 – 25 (– 27), 2.5 – 3.5 mm long, filaments 2 – 3 mm long, glabrous or villous only basally, anthers 0.5 – 0.7 × 0.4 – 0.7 mm, suborbicular or largely ellipsoid; nectary disks 5 - segmented, segments transversely oblong, whitish, receptacle villous. Pistillate flowers 3.5 – 5.3 mm long, pedicels 0.8 – 1.5 mm long; sepals 5, 1 – 1.7 × 0.6 – 0.8 mm, equal, ovate or triangular, apex acute, margin entire, without glands, externally with pubescent or tomentose indumentum of stellate-porrect or multiradiate sessile trichomes, internally sparsely pubescent or glabrescent; petals globose 0.1 – 0.2 mm diam. or laminar 0.5 – 1.3 × 0.05 – 0.4 mm, linear or triangular, glabrous or with stellate trichomes externally; ovaries 1.2 – 1.4 × 1.3 – 1.8 mm, subglobose, with whitish tomentose indumentum of stellate-porrect shortly stipitate trichomes; styles 3, 2 – 3 mm long, united in ⅙ – ⅓ of length, then branching into 6 terminal tips, externally with glabrescent or pubescent indumentum of stellate trichomes basally; nectary disks 5 - segmented or 5 - lobed, segments or lobes transversely oblong, whitish. Capsules 5.7 – 6 mm diam., globose, greenish, pedicel 1.2 – 2.5 (– 4) mm long, columella with three flattened or slightly ascending tips; seeds 4.8 – 4.9 × 3 – 3.1 mm, ellipsoid, brownish, with smooth surface, slightly foveolate, caruncle hat-shaped, 1.8 – 2.2 mm large, whitish.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	distribution	Distribution, habitat and phenology Croton sertanejus sp. nov. is endemic to the Caatinga, a Brazilian Seasonally Dry Tropical Forest, with records for the states of Bahia, Ceará, Minas Gerais, Paraíba, Pernambuco, Piauí and Sergipe (Fig. 3). It grows amid the shrubby or arboreal physiognomies of scrub / forest vegetation, close or not to water courses, sometimes associated with rocky outcrops and on sandy or clay soils, between 160 and 930 m of elevation. This species was found with flowers from December to April and fruits from January to May.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	vernacular_names	Vernacular names The new species is popularly known as ʻCaatinga-brancaʼ (Pernambuco), ʻCassatingaʼ (Bahia), ʻCassutingaʼ (Bahia), ʻMarmeleiroʼ (Bahia), ʻMarmeleiro-de-folha-miúdaʼ (Paraíba), ʻMarmeleiro-pretoʼ (Piauí), ʻQuebra-facaʼ (Bahia, Ceará, Pernambuco, Piauí) or ʻQuebra-facãoʼ (Bahia, Pernambuco, Piauí).	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B60BFFDF526CFCA4FBBF8FB0.taxon	conservation	Preliminary conservation status This species has an Extent of Occurrence estimated to be 494 273 km 2 and Area of Occupancy of 5696 km 2 and is known from more than 100 different locations in seven states, many of them recently collected. Therefore, its conservation status was provisionally assessed as Least Concern. Despite this, the frequent use of its wood for firewood or fences and its leaves and bark in popular medicine or as food for goats may affect the conservation of their natural populations in the near future.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	description	Figs 4 – 5	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	materials_examined	Material examined BRAZIL – Bahia • Abaíra, arredores de Catolés; 13 ° 17 ′ S, 41 ° 51 ′ W; 1000 – 1200 m a. s. l.; 22 Nov. 1991; fl.; V. C. Souza & C. M. Sakuragui H 50256; NY, SPF • ibid.; 24 Dec. 1991; fl.; R. M. Harley et al. H 50312; NY, MO, SPF • estrada de engenho entre Catolés e Abaíra; 13 ° 18 ′ S, 41 ° 49 ′ W; 1100 m a. s. l.; 31 Jan. 1992; fl. and fr.; J. R. Pirani et al. H 51381; SPF, UB • Mendonça de Daniel Abreu; 13 ° 16 ′ S, 41 ° 49 ′ W; 1050 m a. s. l.; 24 Feb. 1992; fl. and fr.; B. Stannard et al. H 51568; HUEFS, MBM, NY, RB, SPF • estrada Catolés-Abaíra, 7 km de Catolés, Mata do Criminoso; 13 ° 19 ′ S, 41 ° 48 ′ W; 1000 m a. s. l.; 26 Feb. 1992; fr.; B. Stannard et al. H 51621; SPF • Morro do Zabumba, Engenho de Baixo; 13 ° 18 ′ S, 41 ° 48 ′ W; 850 m a. s. l.; 13 Mar. 1992; fr.; B. Stannard et al. H 51937; NY, SPF • estrada Catolés-Abaíra, ca 5 km de Catolés, Mata do Engenho; 13 ° 19 ′ S, 41 ° 49 ′ W; 1000 m a. s. l.; 24 Nov. 1992; fl.; W. Ganev 1547; HUEFS, HUFU, NY, SP, SPF • 13 ° 18 ′ S, 41 ° 49 ′ W; 930 m a. s. l.; 22 Jan. 1994; fl.; W. Ganev 2856; HUEFS, NY, SPF, UESC • Serra do Atalho, Complexo Serra da Tromba, 18 Apr. 1994; fr.; E. Melo et al. 992; HUEFS • estrada de chão sentido Cascavel; 13 ° 19 ′ 0 ″ S, 41 ° 34 ′ 24 ″ W; 767 m a. s. l.; 18 Nov. 2015; fl.; B. R. L. Barbosa et al. 1404; HUEFS • estrada Abaíra-Catolés; 13 ° 19 ′ S, 41 ° 47 ′ W; 17 Dec. 2017; fl.; R. C. Sodré et al. 3314; UFG • Almadina, rodovia de Almadina para Ibitupã, ca 20 km, Fazenda São Roque, ca 10 km da entrada do ramal; 14 ° 38 ′ 27 ″ S, 39 ° 42 ′ 47 ″ W; 12 Mar. 2005; fr.; P. Fiaschi et al. 2754; CEPEC, SPF, NY • Barra, Serra do Estreito, 5 Nov. 2009; no fl. or fr.; A. P. Prata et al. 1724; ASE • 10 ° 58 ′ 48 ″ S, 43 ° 33 ′ 36 ″ W; 20 Dec. 2017; fl.; A. S. Soares 404; RB, UB • Barra do Mendes, estrada do Mendes; 11 ° 52 ′ S, 42 ° 8 ′ W; 27 Nov. 1977; fl.; C. A. Miranda 11; RB • estrada para Canarina, localidade Carretão; 11 ° 49 ′ 33 ″ S, 42 ° 3 ′ 17 ″ W; 750 m a. s. l.; 15 Dec. 2009; fl. and fr.; E. Melo et al. 7514; ALCB, HUEFS • ca 21.5 km na estrada de terra para Seabra; 11 ° 58 ′ 25 ″ S, 42 ° 0 ′ 38 ″ W; 775 m a. s. l.; 16 Dec. 2009; fl.; E. Melo et al. 7571; ALCB, HUEFS • ca 32 km na estrada de terra para Seabra; 12 ° 3 ′ 7 ″ S, 42 ° 0 ′ 33 ″ W; 828 m a. s. l.; 16 Dec. 2009; fl.; E. Melo et al. 7591; ALCB, HUEFS • ibid.; fl. and fr.; E. Melo et al. 7597; ALCB, HUEFS • estrada para Canarina; 12 ° 9 ′ 3 ″ S, 42 ° 1 ′ 3 ″ W; 1038 m a. s. l.; 16 Dec. 2009; fl.; E. Melo et al. 7621; ALCB, HUEFS • ibid.; E. Melo et al. 7634; ALCB, HUEFS • estrada para Ipupiara; 11 ° 47 ′ 16 ″ S, 42 ° 11 ′ 25 ″ W; 850 m a. s. l.; 17 Dec. 2009; fl.; E. Melo et al. 7679; ALCB, HUEFS • Bom Jesus da Lapa, ca 18 km NE de Bom Jesus da Lapa na estrada para Ibotirama; 13 ° 7 ′ 55 ″ S, 43 ° 18 ′ 59 ″ W; 326 m a. s. l.; 8 Feb. 2000; fr.; L. P. Queiroz et al. 5745; ALCB, CEPEC, HUEFS • Buritirama; 10 ° 26 ′ 23 ″ S, 43 ° 44 ′ 24 ″ W; 15 Jan. 2018; fl.; O. Neto 974; UB • Caetité, Caetité / Bom Jesus da Lapa, ca 10 km de Caetité; 13 ° 58 ′ 31 ″ S, 42 ° 32 ′ 54 ″ W; 950 m a. s. l.; 6 Dec. 1999; fl.; E. Melo & F. França 3222; HUEFS • Região de Brejinho das Ametistas, Área da Bahia Mineração; 14 ° 19 ′ 24 ″ S, 42 ° 33 ′ 25 ″ W; 980 m a. s. l.; Aug. 2008; fr.; M. S. Mendes et al. 371; BHCB, RON • Cafarnaum, Propriedade do Sr. Cícero Pereira Lima, 14 Jan. 2012; fl.; J. G. Carvalho-Sobrinho et al. 3237; HUEFS • Campo Formoso; 10 ° 28 ′ S, 41 ° 14 ′ W; 29 Apr. 1981; fl.; R. P. Orlandi 377; HUEFS, RB • Casa Nova, margem do lago do Sobradinho; 9 ° 33 ′ 43 ″ S, 41 ° 46 ′ 6 ″ W; 415 m a. s. l.; 29 Feb. 2008; fl.; C. Correia et al. 424; HST, HUEFS, UEC • Correntina, rio Correntina; 17 Mar. 1995; fl. and fr.; G. Hatschbach et al. 62009; MBM • Delfino, estrada Delfino – Mimoso de Minas, 20 km de Delfino; 10 ° 29 ′ 31 ″ S, 41 ° 20 ′ 35 ″ W; 826 m a. s. l.; 7 Mar. 1997; fr.; P. Gasson et al. PCD 6118; ALCB, CEPEC, HUEFS, SPF • Formosa do Rio Preto, Estação; 10 ° 58 ′ S, 44 ° 57 ′ W; 24 Feb. 2005; fr.; M. L. Guedes & A. B. Xavier 11631; ALCB, CEPEC • Ibiquera, Gruta da Lapinha; 12 ° 27 ′ 51 ″ S, 40 ° 58 ′ 47 ″ W; 610 – 710 m a. s. l.; 1 May 2004; fr.; F. França et al. 4960; HUEFS, HUESB • Igaporã, ca 3 km após Igaporã em direção a Caetité; 13 ° 48 ′ 1.5 ″ S, 42 ° 39 ′ 30 ″ W; 1010 m a. s. l.; 15 Dec. 2017; fl.; R. C. Sodré et al. 3284; UFG • Itaetê, estrada para Itaetê, ca 8.7 km do entroncamento com a estrada Mucugê – Andaraí; 12 ° 55 ′ 40 ″ S, 41 ° 9 ′ 52 ″ W; 440 – 460 m a. s. l.; 13 Jul. 2003; fl.; F. França et al. 4786; HUEFS • Licínio de Almeida, Rodovia Caculé / Licínio de Almeida, 38.3 km; 14 ° 38 ′ 29 ″ S, 42 ° 27 ′ 41 ″ W; 850 m a. s. l.; 30 Mar. 2001; fl. and fr.; J. G. Jardim et al. 3231; ALCB, HUEFS, SP • estrada para Urandi, 7 km de Licínio de Almeida; 14 ° 44 ′ 43 ″ S, 42 ° 31 ′ 2 ″ W; 750 m a. s. l.; 9 Dec. 2009; fl.; E. Melo et al. 7369; ALCB, HUEFS • Serra Geral, Mata do Xaxá; 14 ° 39 ′ 6 ″ S, 42 ° 32 ′ 48 ″ W; 930 m a. s. l.; 22 Jan. 2013; fl.; F. Hurbath 452; ALCB, HUEFS, MBM • Serra Geral, Saco da Onça, Trilha Fazenda Serrana; 14 ° 43 ′ 53 ″ S, 42 ° 34 ′ 10 ″ W; 1103 m a. s. l.; 5 Apr. 2013; fr.; F. Hurbath et al. 507; ALCB • Riacho Fundo; 14 ° 35 ′ 28 ″ S, 42 ° 32 ′ 18 ″ W; 840 m a. s. l.; 5 Nov. 2013; fl.; F. Hurbath et al. 600; ALCB • Morro do Chapéu, entre 25 e 26 km de Morro do Chapéu, estrada vicinal próximo à estrada do feijão, próximo ao Ventura; 11 ° 39 ′ 7 ″ S, 40 ° 57 ′ 42 ″ W; 765 – 800 m a. s. l.; 18 Nov. 1989; fl.; E. Melo et al. 3188; HUEFS • Parque Estadual de Morro do Chapéu; 11 ° 20 ′ 43 ″ S, 41 ° 18 ′ 56 ″ W; 1000 m a. s. l.; 2 Feb. 2008; fl.; F. B. L. Silva & F. França 103; HUEFS • Buraco do Possidônio; 11 ° 38 ′ 47 ″ S, 41 ° 16 ′ 11 ″ W; 960 m a. s. l.; 17 Apr. 2008; fl. and fr.; E. Melo et al. 5511; HUEFS • Buraco da Duda ou Buracão; 19 Apr. 2008; fr.; E. Melo et al. 5656; HUEFS • divisa com o município de Várzea Nova, Serra da Babilônia, Fazenda Bom Jesus, futura área de implantação do Parque Eólico da Serra da Babilônia; 11 ° 5 ′ 4 ″ S, 41 ° 18 ′ 51 ″ W; 966 m a. s. l.; 14 Apr. 2016; fl.; L. J. T. Cardoso et al. 1613; RB • Mucugê, Fazenda Pedra Grande, estrada para Boninal; 13 ° 0 ′ 20 ″ S, 41 ° 33 ′ 48 ″ W; 990 m a. s. l.; 17 Feb. 1997; fr.; B. Stannard et al. PCD 5813; ALCB, SPF • Palmeiras, caminho para Conceição dos Gatos; 12 ° 31 ′ 59 ″ S, 41 ° 33 ′ 11 ″ W; 21 Mar. 2003; fr.; E. Melo et al. 3564; HUEFS • caminho para Serra Preta; 12 ° 31 ′ 55 ″ S, 41 ° 35 ′ 14 ″ W; 28 Mar. 2003; fr.; E. Melo et al. 3633; HUEFS • ibid.; 12 ° 26 ′ 48 ″ S, 41 ° 36 ′ 6 ″ W; 697 m a. s. l.; 25 Mar. 2010; fl.; S. P. S. Neves 329; HUEFS • ibid.; fl. and fr.; S. P. S. Neves 330; HUEFS • Piatã, estrada de terra para a mineração; 13 ° 6 ′ 45 ″ S, 41 ° 41 ′ 56 ″ W; 1112 m a. s. l.; 18 Apr. 2014; fr.; E. Melo et al. 12574; HUEFS • Ruy Barbosa, Serra do Orobó, trilha para o pátio das orquídeas; 12 ° 18 ′ 9 ″ S, 40 ° 29 ′ 15 ″ W; 521 m a. s. l.; 19 Dec. 2004; fl.; L. P. Queiroz et al. 9961; HUEFS • Seabra, Serra da Água de Rega, ca 28 km N of Seabra road to Água de Rega; ca 1000 m a. s. l.; 27 Feb. 1971; fl. and fr.; H. S. Irwin et al. 31178; RB • Senhor do Bonfim, Serra de Santana; 7 Feb. 2003; fl.; V. J. Santos et al. 266; HUEFS, HUNEB • Tucano, Fazenda Boqueirão; 12 ° 1 ′ 33 ″ S, 38 ° 38 ′ 22 ″ W; 258 m a. s. l.; 5 Jan. 2006; fl.; D. Cardoso 929; HUEFS, SPF • Wanderley; 11 ° 42 ′ 0 ″ S, 43 ° 44 ′ 24 ″ W; 24 Jan. 2018; fl.; T. M. Moraes 1402; RB. – Ceará • Aiuaba; 6 ° 39 ′ 36 ″ S, 40 ° 40 ′ 48 ″ W; 8 Apr. 2014; fl.; W. Batista 353; EAC • Caucaia; 11 Mar. 2007; fl.; A. S. F. Castro 1921; EAC • Crateús, RPPN Serra das Almas, 6 Apr. 2001; fr.; M. S. Sobrinho & M. M. A. Bruno 36; EAC • ibid.; M. S. Sobrinho & M. M. A. Bruno 40; EAC, PEUFR • ibid.; 16 Feb. 2002; fl.; F. S. Araújo & L. C. Girão 1316; HUEFS • ibid.; 5 ° 8.166 ′ S, 40 ° 54.545 ′ W; 25 Feb. 2002; fl. and fr.; F. S. Araújo 1453; HUEFS • 20 May 2002; fr.; F. S. Araújo & J. R. Lima 1454; EAC • ibid.; 25 May 2005; fr.; J. R. Lima 81; EAC • Itarema, margem da rodovia para Acaraú; 21 Apr. 2003; fl.; A. S. F. Castro s. n.; EAC [32597] • Novo Oriente, Planalto de Ibiapaba, Baixa Fria; 16 Feb. 1991; fr.; F. S. Araújo s. n.; EAC [19451] • São Gonçalo do Amarante, Varjota / Pecém; 13 Mar. 2011; fl. and fr.; A. S. F. Castro 2454; EAC • Tianguá; 3 ° 48 ′ S, 41 ° 4 ′ W; 550 m a. s. l.; 15 Mar. 1978; fl.; O. A. Salgado 39; RB • Chapada da Ibiapaba; 23 Mar. 1979; fl. and fr.; A. Fernandes & F. J. A. Matos s. n.; EAC [5729], UFRN [2348] • ibid.; 16 Feb. 1981; fl.; A. Fernandes & F. J. A. Matos s. n.; EAC [9696] • Ubajara, Planalto de Ibiapaba, Sul do Distrito de Jaburuna; 21 Feb. 1995; fl.; F. S. Araújo 1119; EAC • ibid.; F. S. Araújo 1120; EAC, UFRN • ibid.; 27 Jan. 1996; fl.; F. S. Araújo 1171; EAC • ibid.; F. S. Araújo s. n.; EAC [23595]. – Espírito Santo • São Roque do Canaã, Pedra do Pionte (São Bento), propriedade do Pionte; 750 m a. s. l.; 28 Feb. 2004; fl. and fr.; A. P. Fontana et al. 762; MBM. – Minas Gerais • Dom Joaquim, São Domingos, près de Conceição, dans le bois; Nov. or Dec.; fl.; A. F. M. Glaziou 13180; P • Montezuma, ca 23 km NE da cidade, em direção a Montugaba (BA); 15 ° 3 ′ 28 ″ S, 42 ° 22 ′ 32 ″ W; 900 – 1000 m a. s. l.; 14 Mar. 1994; fl. and fr.; V. C. Souza et al. 5495; ESA, SPF • Ninheira, sudoeste de Ninheira; 17 ° 18 ′ 45 ″ S, 41 ° 34 ′ 45 ″ W; 776 m a. s. l.; 13 Jan. 2014; fl.; D. M. Neves 1613; HUEFS, RB • without further locality; s. d.; fl.; A. F. M. Glaziou 11513; P. – Paraíba • Cajazeiras, Engenheiro Ávidos, Serra de Santa Catarina; 11 May 2008; fr.; P. C. Gadelha Neto 2238; HUEFS, JPB • São José de Piranhas, topo da serra a direita da Barragem de Engenheiro Ávidos, ca de 2 km a SE da localidade Engenheiro Ávidos; 6 ° 59 ′ 17.55 ″ S, 38 ° 26 ′ 47.43 ″ W; 592 m a. s. l.; 15 Jan. 2015; fl.; J. L. Costa-Lima & L. S. Lima 2023; HUEFS. – Pernambuco • Exu, BR- 122 sentido Crato / Exu, depois da divisa CE / PE; 7 ° 17 ′ 52 ″ S, 39 ° 32 ′ 53 ″ W; 927 m a. s. l.; 13 Feb. 2007; fl.; D. S. Carneiro-Torres et al. 874; HUEFS, INPA • Ipubi, Serrolândia, Chapada do Araripe, 3 km Serra Branca; 15 Feb. 1984; fl.; G. Fotius 3761; HUEFS, IPA • Moreilândia, Sítio dos Moreiras; 22 Feb. 1994; fl. and fr.; M. Andrade-Neto s. n.; EAC [20390], HUEFS [111103] • Ponta da Serra; 4 Mar. 1995; fl.; E. Silveira s. n.; EAC [21992], HUEFS [111106] • ibid.; 11 Mar. 1997; fl.; F. S. Cavalcanti & E. Silveira s. n.; EAC [24710] • Zé de Belina; 10 May 2000; fr.; F. S. Cavalcanti 763; EAC • Serra do Catolé, Refúgio das Abelhas Professor Paulo Nogueira Neto; 7.2751 ° S, 39.28132 ° W; 880 m a. s. l.; 8 Mar. 2015; fl.; V. M. Mascena C 88; EAC • Tacaratu; 21 Jun. 2013; fl. and fr.; P. Bohrer s. n.; MBM [394421]. – Piauí • Canto do Buriti, Fazenda Itaueira; 500 m a. s. l.; 9 Jan. 1995; fl.; I. Pinto & E. Alves s. n.; EAC [22098] • ibid.; I. Pinto & E. Alves s. n.; EAC [22114], HUEFS [111108] • Caracol, área do entorno dos cálices rochosos; 9 ° 13 ′ 15 ″ S, 43 ° 28 ′ 56 ″ W; 681 m a. s. l.; 26 Feb. 2011; fr.; E. Melo 9251; HUEFS • Cocal, Jacarandá; 3 ° 32 ′ 44.3 ″ S, 41 ° 3 ′ 54.6 ″ W; 340 m a. s. l.; 22 Feb. 2003; fl.; E. M. F. Chaves et al. 125; TEPB • ibid.; 3 ° 32 ′ 43 ″ S, 41 ° 3 ′ 51 ″ W; 340 m a. s. l.; 18 Feb. 2003; fr.; E. M. F. Chaves & E. M. Sérvio Júnior 346; HUEFS • Eliseu Martins; 7 ° 55 ′ 12 ″ S, 43 ° 33 ′ 36 ″ W; 6 Aug. 2017; no fl. or fr.; M. S. Gomes 241; UB • Padre Marcos, Serra Velha, km 2 da vicinal para Alagoinha; 420 m a. s. l.; 17 Jun. 1993; no fl. or fr.; M. E. Alencar & A. J. Castro s. n.; EAC [19687] • ibid.; 24 Jan. 1994; fl.; M. E. Alencar s. n.; EAC [20717] • ibid.; 10 Mar. 1994; no fl. or fr.; M. E. Alencar s. n.; EAC [20718] • Pio IX, Cova Donga; 11 Feb. 1982; fl.; M. A. Figueiredo s. n.; EAC [11105] • São João do Piauí, BR- 420, São Raimundo Nonato sentido São João do Piauí; 8 ° 28 ′ 57 S, 42 ° 18 ′ 17 ″ W; 354 m a. s. l.; 6 Feb. 2007; fl.; D. S. Carneiro-Torres et al. 797; HUESB, INPA, HUEFS • São Raimundo Nonato, Chapada da Serra Branca; 7 Feb. 1984; fl.; L. Emperaire 2420; P, RB • without municipality, Serra Branca; Jan. 1907; fl.; E. Ule 7454; L. – Rio de Janeiro • Entre Rio Bonito et Araruama; Oct.; fl.; A. Glaziou 10039; P.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	description	Description Subshrubs or shrubs, 0.6 – 2 (– 3) m tall, erect, usually few-branched with tender, cylindric, straight, smooth (not striate) branches ramifying in di- or trichotomies, sometimes with galls throughout plant; latex abundant, orange or reddish; odoriferous (but not sweetish); whitish or ferrugineous indumentum of stellate-rotate, stellate-lepidote or stellate-porrect trichomes, all with (5 –) 8 – 18 lateral rays in singleplane whorl, each ray 0.2 – 0.6 mm long, free or united up to 10 – 30 % of their length, central ray 0.1 – 0.3 (– 0.5) mm long, stipe 0.1 – 0.2 mm long. Leaves alternate, or sometimes subopposite or subwhorled near the inflorescences, leaf blades as long as petioles or up to 3 times as long as petioles; stipules 1 – 3.5 × 0.45 – 0.6 mm, triangular or lanceolate, without conspicuous glands or with 2 very short globose glands, with hirtellous or glabrescent indumentum of simples trichomes ventrally and dense to sparse indumentum of stellate-porrect trichomes dorsally; petioles (1.8 –) 3 – 8.1 cm long; leaf blades 4.3 – 13.6 × 2.7 – 8.2 cm, ovate, elliptic, largely ovate, largely or narrowly elliptic, oblong or lanceolate, membranaceous, base slightly cordate or cordate, less commonly rounded, truncate or obtuse, with 2 (3) stipitate-patelliform, cylindric or obconic nectary glands 0.3 – 0.7 mm diam., stipe 0.5 – 1.4 mm long, with stellate trichomes, facing abaxial surface; apex acute, obtuse, acuminate, mucronate or apiculate; margin entire with globose sessile glands; venation brochidodromous with 6 – 10 pairs of secondary veins slightly prominent on both surfaces; discolorous, adaxial surface dark green, glabrous or with sparse indumentum of simple, 2 – 4 - radiate trichomes and sessile stellate-porrect (6 – 10 lateral rays) trichomes, lateral rays 0.2 – 0.6 mm long, porrect ray 0.2 – 0.8 mm long, abaxial surface light green or whitish with shortly tomentose, pubescent or glabrescent indumentum of stellate-porrect or stellate-rotate trichomes, sometimes multiradiate, all with 8 – 16 lateral rays usually in single-plane whorl, or two-plane whorls in multiradiate trichomes, lateral rays 0.4 – 1.2 mm long, central ray 0.05 – 0.8 mm long, stipe 0.1 – 0.3 mm long. Thyrses 10.4 – 34.6 cm long, peduncles 1.5 – 4.5 cm long, terminal, bisexual with 2 – 18 solitary pistillate flowers sparsely distributed on ⅛ – ½ of inflorescence axis and staminate cymules with 3 – 6 flowers, less commonly with bisexual cymules in median portion of rachis, or thyrses unisexual staminate; staminate flower bracts 0.7 – 1.2 × 0.4 – 0.7 mm, triangular or ovate, margin entire, sometimes with 2 ovoid sessile glands (colleters) at base, with pubescent, hirtellous or glabrescent indumentum of simple trichomes ventrally and hirtellous indumentum of stellate-porrect trichomes dorsally, mainly on margin, 2 or more staminate flower bracteoles per cymule 0.5 – 0.6 × 0.2 – 0.3 mm, lanceolate or triangular, with glands and indumentum similar to those of bracts; pistillate flower bracts 0.9 – 1.1 × 0.7 – 1 mm, triangular or largely ovate, margin entire, apex acute or acuminate, with two ovoid or subglobose sessile glands (colleters) at base and one at apex, with glabrescent or hirsute-tomentose indumentum of stipitate stellate-porrect or 4 - radiate trichomes on both surfaces, 2 pistillate flower bracteoles 0.2 – 0.6 × 0.15 – 0.3 mm, triangular, with glands and indumentum similar to those of bracts. Staminate flowers 3.5 – 6.5 (– 8) mm long; pedicels 0.8 – 3 (– 5.7) mm long, glabrous, glabrescent, or sometimes pubescent; calyces 5 - partite, yellow-orange or slightly reddish with hyaline indumentum, lobes 1.6 – 2.1 × 0.9 – 1.3 mm, ovate, united in ⅙ – ⅕ of length, apex obtuse, dorsal surface with densely to sparsely pubescent or hirtellous indumentum of stellate-porrect sessile trichomes, villous at apex, glabrous internally; petals 5, 1.9 – 3 × 0.75 – 0.9 mm, obovate or oboval-oblanceolate, apex obtuse, truncate or rounded, villous basally along margins at ¼ of length and shortly villous at apex; stamens 16 – 19, 2.3 – 3.8 mm long, filaments 1.6 – 3 mm long, glabrous, anthers 0.6 – 0.8 × 0.4 – 0.5 mm, ellipsoid or oblongoid; nectary disks 5 - segmented, segments transversely oblong or suborbicular, orange, receptacle villous. Pistillate flowers 3 – 5 mm long, sessile or pedicels 0.1 – 1 (– 1.2) mm long, tomentose; sepals 5 (6), 1.3 – 3 × 0.6 – 1.3 mm, subequal or strongly unequal, ovate, narrowly triangular, oval-oblong or oval-lanceolate, apex acute, margin entire, without glands, dorsally with sparsely to densely pubescent or tomentose indumentum of stellate-porrect trichomes, margin with hirsute indumentum of simple trichomes and ventrally with glabrescent, pubescent or hirtellous indumentum of simple and fasciculate trichomes; petals globose ca 0.2 mm diam. or laminar 0.6 – 1.3 × 0.25 – 0.35 (– 0.4) mm, lanceolate, glabrous or pubescent; ovary 0.8 – 1.1 × 0.9 – 1.6 mm, with subglobose, hirtellous or tomentose indumentum of stellate-porrect shortly stipitate trichomes; styles 3, (2.8 –) 4 – 5.5 mm long, united along lower ⅒ of their length, then branching into 6 terminal tips, externally with hirsute, hirtellous or pubescent indumentum of stellate-porrect or simple trichomes along less than ½ of their length; nectary disks 5 - segmented, segments transversely oblong, yellowish. Capsules 4.8 – 5.5 × 5.5 – 6 mm, compressed-globose, greenish, pedicels 0.5 – 1.5 mm long, columella with three flattened or slightly ascending tips; seeds 4.2 – 4.8 × 3.1 – 3.5 mm, largely ellipsoid, grayish, smooth surface, slightly foveolate, caruncle hat-shaped 1 – 2 mm large.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	distribution	Distribution, habitat and phenology Croton echioides has been cited by Carneiro-Torres (2009), Silva et al. (2010) and Caruzo et al. (2020) as a species endemic to Brazil distributed in areas of Caatinga and Caatinga-Cerrado transition in the states of Alagoas, Bahia, Ceará, Minas Gerais, Paraíba, Pernambuco, Piauí and Rio Grande do Norte. We confirm occurrence of the species in the states mentioned above, except for Alagoas and Rio Grande do Norte, and provid the first record for the states of Espírito Santo and Rio de Janeiro. This species was found in seasonal dry forest, humid forest, caatinga, in transition areas between caatinga and cerrado, ʻcarrascoʼ or ʻtabuleiro litorâneoʼ growing preferentially on sandy soils, sometimes with outcrops of quartzite, and stony or sandy-clay soils between 30 and 1200 meters a. s. l. It was usually found with flowers from November to April, and with fruits from December to June.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	vernacular_names	Vernacular names ʻVelameʼ (Ceará), ʻVelame-da-serraʼ (Pernambuco), ʻVelame-brancoʼ (Piauí).	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	conservation	Preliminary conservation status Croton echioides has an Extent of Occurrence estimated to be 910 000 km 2 comprising at least 80 subpopulations and has an estimated Area of Occupancy of 4544 km 2 (based on an 8 km 2 cell size) and is therefore provisionally assessed as Least Concern. However, it is worth noting that only three of these known subpopulations occur within protected areas (Experimental Station of the IPA, Serra Talhada, Pernambuco; RPPN Serra das Almas, Cratéus, Ceará; and Morro do Chapéu State Park, Bahia), all the others are in non-protected areas, which may have implications for the conservation of the species in the future.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
76215954B603FFC752F0F9B0FAD68BF1.taxon	discussion	Leaf anatomy notes In general, the anatomy of the leaf blades and petiole of Croton echioides and C. sertanejus sp. nov. is relatively similar. Both species have hipostomatic leaves, with stomata on the same level as the other cells of the epidermis, which is unistratified and composed of cells of varying sizes, rectangular or rounded, and has a thin cuticle. The mesophyll in both species is dorsoventral, with a layer of palisade parenchyma and another layer of spongy parenchyma, both with druses (Fig. 6 F, H, L). The leaf margins have rounded outlines and the mesophyll and epidermis are not differentiated from the rest (Fig. 6 I, M). The vascular bundles are collateral, surrounded by a conspicuous parenchyma sheath in the smaller vascular bundles and with annular (Fig. 6 E) or angular collenchyma cells adjacent to the phloem on the abaxial surface (Fig. 6 J). The primary veins of both species have a convex ventral side with 5 to 6 layers of angular collenchyma cells (Fig. 6 P, T). The dorsal side is rounded with 4 layers of angular collenchyma cells in subepidermal position (Fig. 6 U) followed by a variable number of layers of ground parenchyma (Fig. 6 N, R). The primary vein has a collateral vascular cylinder (Fig. 6 O, S) with druses in the phloem. The petiole of the two species has a circular dorsal side and truncated or slightly concave ventral side in cross section. The cortex is composed of angular collenchyma and ground parenchyma. The vascular cylinder is collateral with united bundles, and there are also two accessory vascular cylinders on the ventral side (Fig. 7 A, F). The pericycle is always associated with gelatinous fibers (Fig. 7 C – E, I – J), which also surround the accessory vascular bundles (Fig. 7 B, G). Druses are associated with phloematic parenchyma (Fig. 7 D – E, I – J), as in primary veins. Despite the similar anatomical characteristics mentioned above, which are to be expected from morphologically and geographically related species, important anatomical characteristics differentiate C. sertanejus sp. nov. from C. echioides. Leaf trichomes are stipitate in C. echioides, with stipe 50 – 300 µm long (Figs 6 A, 7 A), and the leaf blade is often glabrous on the adaxial face (Fig. 6 A), whereas in C. sertanejus the trichomes occur on both faces and are sessile (Fig. 6 B – C), or with a stipe up to 30 µm (Fig. 7 F) or just with a set of cells at the base forming a cushion (Fig. 6 D). The leaf blade of C. sertanejus also has subglobose unicellular idioblasts on the abaxial surface (Fig. 6 K), which were not observed in C. echioides. In the primary veins, C. echioides presents the U-shaped vascular cylinder (or open arch) and the palisade parenchyma is continuous between the two faces of the leaf blade (Fig. 6 N), as can also be seen in the smaller vascular bundles of the leaf blades (Fig. 6 E). In C. sertanejus, however, the vascular cylinder of primary veins is circular (or closed arch), and the palisade parenchyma is discontinuous between the two faces of the leaf blade (Fig. 6 R). In the petiole, the accessory vascular cylinders are semi-lunar (open arch) and the pith has approximately 10 to 12 layers of cells in C. echioides, while in C. sertanejus the accessory vascular cylinders are circular (closed arch) and the pith has only 4 to 6 layers. In addition, the cortex of the primary vein and petiole of C. echioides have evident and numerous laticifers close to the phloem; while in C. sertanejus the laticifers were not observed or are not evident.	en	Sodré, Rodolfo Carneiro, Silva, Marcos José da (2022): Croton sertanejus, a new species from Seasonally Dry Tropical Forest in Brazil, and redescription of C. echioides (Euphorbiaceae). European Journal of Taxonomy 839: 14-38, DOI: https://doi.org/10.5852/ejt.2022.839.1929
