identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C74FFC13303DFF9DFF685645D8F6FA40.text	C74FFC13303DFF9DFF685645D8F6FA40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhithrops Bilton & Shepard 2022	<div><p>Rhithrops gen. nov.</p> <p>(Figs 1–11)</p> <p>Type species. Rhithrops capensis sp. nov., by monotypy.</p> <p>Diagnosis. Rhithrops is unique amongst aquatic dryopid genera in that the labrum is transversely upturned apically. In South Africa, Rhithrops can be separated from Ahaggaria and Dryops by the lack of long erect setae on the eyes and dorsum of the body. Rhithrops can be separated from Strina and Rapnus by the antennae having 10 antennomeres as opposed to nine in Strina and seven in Rapnus. Additionally, Rhithrops has shallow sublateral pronotal furrows rather than sublateral step-like carinae, with discrete gibbosities on the pronotum outside of the sublateral furrows. The dorsally serrate coxites of the ovipositor are only shared with Strina, another South African dryopid genus. Abdominal ventrite 5 lacking a plastron is a character state commonly seen in the genus Helichus Erichson, 1847.</p> </div>	https://treatment.plazi.org/id/C74FFC13303DFF9DFF685645D8F6FA40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bilton, David T.;Shepard, William D.	Bilton, David T., Shepard, William D. (2022): Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae). Zootaxa 5195 (6): 539-553, DOI: https://doi.org/10.11646/zootaxa.5195.6.3
C74FFC13303EFF9BFF68544DDD82FA37.text	C74FFC13303EFF9BFF68544DDD82FA37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhithrops capensis Bilton & Shepard 2022	<div><p>Rhithrops capensis sp. nov.</p> <p>(Figs 1–9)</p> <p>Type locality. South Africa, Western Cape, Cederberg, Matjies Rivier, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.350111&amp;materialsCitation.latitude=-32.518726" title="Search Plazi for locations around (long 19.350111/lat -32.518726)">Matjiesrivier</a>, 32°31’07.42”S 19°21’00.4”E, 725 m. (Fig. 9).</p> <p>Type material. Holotype (male): “ 22/ix/2011 South Africa WC// <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.350111&amp;materialsCitation.latitude=-32.518726" title="Search Plazi for locations around (long 19.350111/lat -32.518726)">Cederberg</a>, Matjies Rivier // Matjiesrivier 32°31’07.42”S // 19°21’00.4”E, 725 m. D T Bilton leg.” (AMG).</p> <p>Paratypes (23): 5♂, 10♀ same data as holotype; 1♀ “ 21/ix/2010 South Africa WC// Cederberg, Matjies Rivier // <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.350111&amp;materialsCitation.latitude=-32.518726" title="Search Plazi for locations around (long 19.350111/lat -32.518726)">Matjiesrivier</a> 32°31’07.42”S // 19°21’00.4”E, 725 m. D T Bilton leg.”; 2♂, 3♀ “ 22/ix/2011 South Africa WC// Cederberg, Matjies Rivier // <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.350111&amp;materialsCitation.latitude=-32.518726" title="Search Plazi for locations around (long 19.350111/lat -32.518726)">Matjiesrivier</a> 32°31’07.42”S // 19°21’00.4”E, 725 m. D T Bilton leg.”; 1♂, 1♀ “ South Africa: W. Cape // Cederberg, Maalgat // <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.280277&amp;materialsCitation.latitude=-32.494446" title="Search Plazi for locations around (long 19.280277/lat -32.494446)">Matjiesrivier</a> // 28-29 XII 2004 820 m asl // 32°29’40”S 19°16’49”E // P. S. Cranston, leg.” (AMG, CBP, DMSA, EMEC, NMW).</p> <p>Description: With all the above characters of the genus.</p> <p>Size: Holotype: BL 4.10 mm; EL 2.65 mm; EW 1.35 mm; PL 0.85 mm; PW 1.20 mm. Paratypes: Males BL 4.40–4.75 mm; EL 2.60–3.00 mm; EW 1.45–1.60 mm; PL 0.85–1.05 mm; PW 1.20–1.35 mm. Females BL 4.65– 4.80 mm; EL 2.75–2.90 mm; EW 1.40–1.60 mm; PL 0.90–1.05 mm; PW 1.25–1.35 mm.</p> <p>Head: Epicranium densely and shallowly punctate, with short decumbent setae, epicranial suture absent; frontal area twice as wide as one compound eye, antennal ridges absent, frontoclypeal suture straight between antennal bases.</p> <p>Elytra: Conjointly twice as long as wide, parallel-sided, apices moderately protruding (Figs 1 &amp; 6D–E).</p> <p>Legs: Relatively long, elongate, with large, strongly-developed tarsi and claws (Fig. 8A–C).</p> <p>Male: Last abdominal ventrite with broad, shallow apicomedial emargination (Fig. 7E). Labium with modified, spathulate setae on anteroventral margin (Fig. 4F). Anterior edge of sternite 8 with short median strut (Fig. 9F); sternite 9 as in Fig. 9C. Tergite 8 with numerous long setae close to posterior margin (Fig. 9E).</p> <p>Aedeagus: Long and tubular (Figs 8D–F &amp; 9A, B &amp; D); basal piece:paramere ratio 0.6:0.4. Basal piece hookshaped basally, open on left side. Median lobe long, narrow, acute apically. Parameres parallel-sided, apically a little longer than median lobe, apices rounded on outer side, clasping apex of median lobe (Fig. 8F).</p> <p>Female: Few external differences to males. The sublateral furrows of the pronotum are deeper and more evident (Fig. 2B). The last abdominal ventrite has a conical apex, somewhat raised centrally and lacks an apicomedian emargination (Fig. 7F). The labium has fewer, shorter modified setae on the anteroventral margin (Fig. 4E). Sternite 8 as in Fig. 10C.</p> <p>Ovipositor: Elongate (Fig. 10A–B) and strongly sclerotized; type 1 (sensu Kodada et al. 2016); apically sharp-tipped. Valvifers ventral, asymmetrical, long, thin, forming a V-shaped sclerite. Coxites asymmetrical, laterally compressed, blade-shaped, apex very acute, dorsal side serrate, with basally directed long narrow dorsal extension.</p> <p>Distribution and habitat. Known only from type locality, the Matjies River (Fig. 11), a tributary of the Olifants River in the Cederberg, Western Cape Province, South Africa.</p> <p>Etymology. The trivial name, capensis, refers to location of this species in the Western Cape Province of South Africa.</p> </div>	https://treatment.plazi.org/id/C74FFC13303EFF9BFF68544DDD82FA37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bilton, David T.;Shepard, William D.	Bilton, David T., Shepard, William D. (2022): Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae). Zootaxa 5195 (6): 539-553, DOI: https://doi.org/10.11646/zootaxa.5195.6.3
C74FFC133031FF91FF6856BDD8F8F875.text	C74FFC133031FF91FF6856BDD8F8F875.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dryopidae Billberg 1820	<div><p>Revised key to South African Dryopidae genera</p> <p>The key below is based on that of Nelson (2007), adapted to incorporate Rhithrops gen. nov. and Pedestrodryops Kodada, 2001.</p> <p>1 Almost entire dorsum and venter with long, fine, sparse erect sensillae; terrestrial, in plant litter away from water … Pedestrodryops Kodada, 2001</p> <p>- Cuticular vestiture shorter and denser; aquatic taxa.......................................................... 2</p> <p>2 Eyes conspicuously pubescent.......................................................................... 3</p> <p>- Eyes not conspicuously pubescent, at most with scattered peg-like setae between facets............................. 4</p> <p>3 First antennomere elongate; second antennomere large, transverse, more or less rectangular, distal border rounded......................................................................................... Ahaggaria Bollow, 1938</p> <p>- First antennomere not elongate; second antennomere enlarged, ear-shaped, distal border angular...... Dryops Olivier, 1791</p> <p>4 Antennae with 10 antennomeres; second antennomere enlarged, ear-shaped, covering most of rest of antennomeres; pronotum with sublateral furrows but no carinae..................................................... Rhithrops gen. nov.</p> <p>- Antennae with fewer than 10 antennomeres; second antennomere smaller; pronotum with sublateral stepped carinae...... 5</p> <p>5 Antennae with 9 antennomeres; sublateral pronotal carinae with lateral flange; prosternal process large, gradually narrowed apically; mesotibial cleaning fringe a dense tuft of golden setae; ovipositor with coxites dorsally serrate Strina Redtenbacher, 1867</p> <p>- Antennae with 7 antennomeres; lateral margin of pronotum without lateral flange; prosternal process large, abruptly narrowed apically; mesotibial cleaning fringe not a dense tuft of golden setae; ovipositor with coxites not serrate dorsally........................................................................................... Rapnus Grouvelle, 1899</p></div> 	https://treatment.plazi.org/id/C74FFC133031FF91FF6856BDD8F8F875	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bilton, David T.;Shepard, William D.	Bilton, David T., Shepard, William D. (2022): Rhithrops capensis gen. et sp. nov., a new aquatic dryopid beetle from the Western Cape of South Africa (Coleoptera: Dryopoidea: Dryopidae). Zootaxa 5195 (6): 539-553, DOI: https://doi.org/10.11646/zootaxa.5195.6.3
