identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E79D793130795A879FFE2BED7C63B180.text	E79D793130795A879FFE2BED7C63B180.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triplophysa daryoae Sheraliev & Kayumova & Peng 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Triplophysa daryoae sp. nov.</p>
            <p> Figs 1, 2, 3 English common name: Sokh stone loach Uzbek common name:  So‘x yalangbalig‘i Russian common name:  Сохский голец</p>
            <p>Holotype.</p>
            <p>SWU 20211207001, male, 78.5 mm SL; Uzbekistan, Fergana Region, Sokh District, Sokh River, near Limbur village, an exclave of Uzbekistan surrounded by Kyrgyzstan, Syr Darya basin, 40°3.1528'N, 71°5.8195'E, altitude 1054 m, December 07, 2021, collected by B. Sheraliev and Y. Kayumova.</p>
            <p>Paratypes.</p>
            <p>SWU 20211207002-011, 10, 49.0-94.0 mm SL; BSFC 0023, 4, 62.1-82.4 mm SL; Uzbekistan, Fergana Region, Sokh District, Sokh River, near the Limbur village, exclave of Uzbekistan, Syr Darya basin, 40°2.7387'N, 71°6.288'E, altitude 1054 m, April 12, 2021, collected by Y. Kayumova. BSFC 0024, 3, 74.1-81.3 mm SL, same data as holotype.</p>
            <p>Diagnosis.</p>
            <p> Triplophysa daryoae is distinguished from congeners by a combination of characters. It is distinguished from  T. ferganaensis by possessing a truncate caudal fin with 13-14 branched rays (vs emarginate, 16 rays), 9 pores in the pre-opercular mandibula (vs 7-8), and a slenderer body (body depth at dorsal-fin origin 1.4-1.8 times the HL vs 1.2-1.4). It is distinguished from  T. strauchii by absence of the posterior chamber of the air bladder (vs developed, with a long tube), possessing 9-10 inner gill rakers on the first gill arch (vs 12-16), and no obvious skin mottling (vs mottling).  Triplophysa daryoae is also distinguished from  T. dorsalis ,  T. dorsonotata , and  T. elegans by having a truncate caudal fin (vs emarginate) and lacking a posterior chamber of the air bladder (vs developed in  T. dorsalis and  T. elegans ). It is distinguished from  T. sewerzowi ,  T. tenuis , and  T. ulacholica by the dorsal-fin origin opposite to the pelvic-fin insertion (vs anterior to vertical line of pelvic fin origin). </p>
            <p>Description.</p>
            <p> Morphometric data of  T. daryoae are given in Table 2. Dorsal-fin rays iii, 6(2) or 7(16); anal-fin rays ii, 5; pectoral-fin rays i, 9(1), 10(16), or 11(1); pelvic-fin rays i, 6; caudal-fin rays, 13-14 (6+7 [5]; 7+7 [13]); vertebrae, 4+35 (N = 2); gill rakers, 9-10 in the inner row of first gill arch (N = 4). Cephalic lateral-line system, 2 supratemporal, 6 supraorbital, 4+10-11 infraorbital, and 9 pre-operculum mandibular pores. </p>
            <p>Body elongate; posterior portion gradually compressed from dorsal fin to caudal-fin origin. Dorsal profile slightly convex from the snout to the insertion of the anterior dorsal fin (Fig. 1). Deepest point of body slightly anterior to dorsal-fin origin; body depth at dorsal-fin origin 12.4-15.3% of SL. Head compression, maximum width always greater than depth; head maximum width 63.2-73.2% of HL. Snout slightly pointed, length shorter, equal, or slightly longer than postorbital length; snout length 34.9-47.3% of HL. Anterior and posterior nostrils adjacent; anterior nostril as short tube with elongated barbel-like tip; tip of nostril barbel not reaching the anterior margin of eyes. Eyes normal; diameter 12.5-17.1% of HL (Fig. 2). Mouth inferior, gape arched; mouth width 16.1-24.3% of HL. Rictus situated below the anterior nostril. Lips thick with furrows and papillae; upper lip pectinate, without medial notch; lower lip wide, interrupted in middle, with mental lobes and two highly developed ridges. Upper jaw covered by the upper lip; processus dentiformis absent. Three pairs of barbels: inner rostral barbel reaching rictus, length 19.8-30.2% of HL; outer rostral barbel reaching anterior margin of eye, length 22.7-42.0% of HL; maxillary barbel reaching posterior margin of eye, length 22.0-37.4% of HL.</p>
            <p>Dorsal fin convex, origin opposite to pelvic-fin insertion, situated slightly posterior to midpoint between snout tip and caudal-fin base; upper margin slightly convex; second branched ray longest; depth of dorsal fin always shorter than lateral head length; depth 14.9-18.5% of SL. Anal fin short-based, posterior margin convex; length 13.1-16.7% of SL. Pectoral fins developed; 46.6-61.6% of pectoral-pelvic distance. Tips of depressed pelvic fins reaching the anus and anus separated from the anal-fin origin by a short distance. Caudal peduncle compressed laterally; length 2.2-2.9 times the peduncle depth. Caudal fin truncate, tips rounded; length 86.2-119.9% of caudal-peduncle length.</p>
            <p>Body smooth and scaleless; cephalic lateral-line system well developed. Infraorbital and supraorbital canals stretching from the outer rostral barbel base and ethmoid, respectively, uniting in the posterior orbital region and extending posteriorly before converging with the supratemporal canal on the back of the head, and uniting with the lateral canal. Complete lateral line ending at caudal-fin base. Intestine moderately long, with two coils. Stomach U-shaped. Posterior chamber of the air bladder degenerated.</p>
            <p>Coloration.</p>
            <p>Dorsal profile grayish-brown to pale green without regular blotches in live individuals, and dark gray-brown in preserved specimens. Ventral side of the body ivory with gray tint. Dorsal side of head with small irregular dark melanophores; dorsal side of caudal peduncle with four or five irregular dark brown blotches. All fin membranes hyaline and light gray, without obvious mottling (Figs 1, 3).</p>
            <p>Sexual dimorphism.</p>
            <p>Mature males presenting granular tubercles on each side of the preorbital region and broadened and thickened external branched pectoral-fin rays dorsally covered by small and condensed epidermal breeding tubercles. Females without tubercles on the head and pectoral-fin rays.</p>
            <p>Distribution and habitat.</p>
            <p> Triplophysa daryoae sp. nov. is known only from its type locality, the Sokh River, which originates in the Alay mountains and Turkestan range (Fig. 4). Presently, Sokh River water is primarily used for irrigation and does not reach Syr Darya. The river is located at an altitude of 700-1500 m and is constantly flowing rapidly; the water is clear and cold (the water temperature was 7.3 °C when the holotype was caught), and the bottom consists of gravel and stone (Fig. 5).  Triplophysa daryoae cohabited with  Cottus spinulosus Kessler, 1872 and  Schizothorax eurystomus Kessler, 1872, which are high-altitude fish species. </p>
            <p>Etymology.</p>
            <p> Triplophysa daryoae is dedicated to Daryo Sheralieva, the lovely daughter of the first author. The specific name is a noun in the genitive case. </p>
            <p>Molecular analysis</p>
            <p> COI sequence data (Fig. 6) showed that  Triplopysa daryoae belongs to a group of species with a wide distribution in the Syr Darya, Tarim, and Ili-Balkhash river drainages, an endorheic basin in Central Asia. This group is defined here as the  T. dorsalis species group, and our molecular data suggest that it includes  T. chondrostoma (Herzenstein, 1888),  T. dorsalis ,  T. dorsonotata ,  T. elegans ,  T. ferganaensis ,  T. sewerzowi ,  T. strauchii ,  T. tenuis (Day, 1877), and  T. ulacholica . The minimum K2P distances between  T. daryoae and its closest relatives  T. ferganaensis and  T. tenuis were 2.8% and 4.5%, respectively (Table 3).  Triplophysa daryoae was distinguished from its most closely related congener,  T. ferganaensis , by 18 unique and diagnostic nucleotide substitution sites in the COI barcode region (652 bp) (Table 4). </p>
            <p> * The nucleotide position number was provided relative to the first nucleotide base of the complete COI gene of  T. tenuis (KT224363). </p>
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	https://treatment.plazi.org/id/E79D793130795A879FFE2BED7C63B180	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sheraliev, Bakhtiyor;Kayumova, Yorkinoy;Peng, Zuogang	Sheraliev, Bakhtiyor, Kayumova, Yorkinoy, Peng, Zuogang (2022): Triplophysa daryoae, a new nemacheilid loach species (Teleostei, Nemacheilidae) from the Syr Darya River basin, Central Asia. ZooKeys 1125: 47-67, DOI: http://dx.doi.org/10.3897/zookeys.1125.85431, URL: http://dx.doi.org/10.3897/zookeys.1125.85431
