taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
E84587EDFFA3087BE7BCFD7062DEFB9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223836/files/figure.png	https://doi.org/10.5281/zenodo.7223836	FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X.	FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA3087BE7BCFD7062DEFB9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223858/files/figure.png	https://doi.org/10.5281/zenodo.7223858	FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9).	FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA3087BE7BCFD7062DEFB9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223862/files/figure.png	https://doi.org/10.5281/zenodo.7223862	FIGURE 10. Psilotreta spp. and their habitats. 10A, P. japonica (Banks 1906), male, dorsal (the locality name in the label “Aoyagi Shin-Asahi” needs to be corrected to “Kawashima Adogawa”). 10B, P. voluta Kawase 2021, female habitus, left dorsolateral (photo by N. Shimura). 10C, mountain stream where adults of P. kisoensis Iwata 1928 and P. voluta were collected at light. 10D, small stream where many adults of P. kisoensis were collected by a Malaise trap. 10E, pools of a small lowland stream inhabited by larvae of P. japonica.	FIGURE 10. Psilotreta spp. and their habitats. 10A, P. japonica (Banks 1906), male, dorsal (the locality name in the label “Aoyagi Shin-Asahi” needs to be corrected to “Kawashima Adogawa”). 10B, P. voluta Kawase 2021, female habitus, left dorsolateral (photo by N. Shimura). 10C, mountain stream where adults of P. kisoensis Iwata 1928 and P. voluta were collected at light. 10D, small stream where many adults of P. kisoensis were collected by a Malaise trap. 10E, pools of a small lowland stream inhabited by larvae of P. japonica.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA3087BE7BCFD7062DEFB9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA1087EE7BCFB6862DEFA9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223838/files/figure.png	https://doi.org/10.5281/zenodo.7223838	FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. & pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X.	FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. & pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA1087EE7BCFB6862DEFA9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223858/files/figure.png	https://doi.org/10.5281/zenodo.7223858	FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9).	FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA1087EE7BCFB6862DEFA9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223862/files/figure.png	https://doi.org/10.5281/zenodo.7223862	FIGURE 10. Psilotreta spp. and their habitats. 10A, P. japonica (Banks 1906), male, dorsal (the locality name in the label “Aoyagi Shin-Asahi” needs to be corrected to “Kawashima Adogawa”). 10B, P. voluta Kawase 2021, female habitus, left dorsolateral (photo by N. Shimura). 10C, mountain stream where adults of P. kisoensis Iwata 1928 and P. voluta were collected at light. 10D, small stream where many adults of P. kisoensis were collected by a Malaise trap. 10E, pools of a small lowland stream inhabited by larvae of P. japonica.	FIGURE 10. Psilotreta spp. and their habitats. 10A, P. japonica (Banks 1906), male, dorsal (the locality name in the label “Aoyagi Shin-Asahi” needs to be corrected to “Kawashima Adogawa”). 10B, P. voluta Kawase 2021, female habitus, left dorsolateral (photo by N. Shimura). 10C, mountain stream where adults of P. kisoensis Iwata 1928 and P. voluta were collected at light. 10D, small stream where many adults of P. kisoensis were collected by a Malaise trap. 10E, pools of a small lowland stream inhabited by larvae of P. japonica.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA1087EE7BCFB6862DEFA9A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA40871E7BCFA6C62C0FDFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223842/files/figure.png	https://doi.org/10.5281/zenodo.7223842	FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text.	FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA40871E7BCFA6C62C0FDFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223858/files/figure.png	https://doi.org/10.5281/zenodo.7223858	FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9).	FIGURE 9. Larvae of Psilotreta spp. 9A, larvae and cases of P. voluta Kawase 2021. 9B–9C, P. japonica (Banks 1906): 9B, head and pro-, meso-, and metanota, dorsal; 9C, larva and case, left lateral. 9D–9F, head and pro-, meso-, and metanota, primary setae numbered: 9D, P. japonica; 9E, P. kisoensis Iwata 1928; 9F, P. spatulata sp. nov. 9G–9I, abdominal gill diagrams, left lateral: 9G, P. japonica; 9H, P. kisoensis; 9I, P. spatulata (sizes of dots represent the numbers of gill branches: large dots = 20 or greater; medium dots = 10–19; small dots = 1–9).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA40871E7BCFA6C62C0FDFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA40871E7BCFA6C62C0FDFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223836/files/figure.png	https://doi.org/10.5281/zenodo.7223836	FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X.	FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA40871E7BCFA6C62C0FDFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223838/files/figure.png	https://doi.org/10.5281/zenodo.7223838	FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. & pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X.	FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. & pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFA40871E7BCFA6C62C0FDFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223844/files/figure.png	https://doi.org/10.5281/zenodo.7223844	FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired).	FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0871E7BCFDCC6228FC04.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223844/files/figure.png	https://doi.org/10.5281/zenodo.7223844	FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired).	FIGURE 4. Psilotreta moritai Kawase 2021. 4A–4E, male genitalia: 4A, dorsal; 4B, left lateral; 4C1–4C2, variations of lateral process of tergum X and intermediate appendage. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); par. = paramere (paired); v.pr. = ventral projection (paired).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0871E7BCFDCC6228FC04.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0874E7BCFB9F62D7FEFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223846/files/figure.png	https://doi.org/10.5281/zenodo.7223846	FIGURE 5 Psilotreta atrocaudata sp. nov. 5A, female right wings, dorsal; 5B–5G, male genitalia: 5B, dorsal; 5C, ventral; 5D, left lateral; 5E, lateral process of tergum X and intermediate appendage, left lateral; 5F, phallus, left lateral; 5G, same, ventral. 5H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X. Arrows see text.	FIGURE 5 Psilotreta atrocaudata sp. nov. 5A, female right wings, dorsal; 5B–5G, male genitalia: 5B, dorsal; 5C, ventral; 5D, left lateral; 5E, lateral process of tergum X and intermediate appendage, left lateral; 5F, phallus, left lateral; 5G, same, ventral. 5H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X. Arrows see text.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0874E7BCFB9F62D7FEFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0874E7BCFB9F62D7FEFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223850/files/figure.png	https://doi.org/10.5281/zenodo.7223850	FIGURE 6. Psilotreta voluta Kawase 2021. 6A–6E, male genitalia: 6A, dorsal; 6B, left lateral; 6C1–6C2, variations of lateral process of tergum X and intermediate appendage, left lateral; 6D, phallus, left lateral; 6E, same, ventral. 6F, female genitalia, ventral. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 6A–6F, redrawn from the publication by Kawase (2021).	FIGURE 6. Psilotreta voluta Kawase 2021. 6A–6E, male genitalia: 6A, dorsal; 6B, left lateral; 6C1–6C2, variations of lateral process of tergum X and intermediate appendage, left lateral; 6D, phallus, left lateral; 6E, same, ventral. 6F, female genitalia, ventral. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 6A–6F, redrawn from the publication by Kawase (2021).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0874E7BCFB9F62D7FEFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223836/files/figure.png	https://doi.org/10.5281/zenodo.7223836	FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X.	FIGURE 1. Psilotreta japonica (Banks 1906). 1A, male head and pro- and mesonota, dorsal; 1B, male right wings, dorsal; 1C, female right wings, dorsal; 1D–1I, male genitalia: 1D, dorsal; 1E, ventral; 1F, left lateral; 1G, variation of lateral process of segment X, left lateral; 1H, phallus, left lateral; 1I, same, ventral. 1J–1K, female genitalia: 1J, dorsal; 1K, ventral.Abbreviations: I, II, III, V = apical forks I, II, III, V; A1+2+3 = combined anal vein A1+A2+A3; aed. = aedeagus; amw. = anteromesal warts; aw. = anterior warts; bsw. = basal setal wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a and 1b; dc = discoidal cell; end. = endotheca; fwcs on false vein = forewing coupling setae on false vein; hwcs on C = hind wing coupling setae on costa; inf.app. = inferior appendage (paired); int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); m-cu = cross vein between median and cubital veins; M1, M2, M3, M4 = median veins 1–4; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); phall. = phallotheca; plw. = posterolateral warts; pr.app. = preanal appendage (paired); pw. = posterior warts; sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0874E7BCFB9F62D7FEFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223838/files/figure.png	https://doi.org/10.5281/zenodo.7223838	FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. & pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X.	FIGURE 2. Psilotreta kisoensis Iwata 1928. 2A, male head and pro- and mesonota, dorsal; 2B, male right wings, dorsal; 2C, female right wings, dorsal; 2D–2I, male genitalia: 2D, dorsal; 2E, ventral; 2F, left lateral; 2G, lateral process of segment X, left lateral; 2H, phallus, left lateral; 2I, same, ventral. 2J–2K, female genitalia: 2J, dorsal; 2K, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; aw. & pw. = anterior wart fused with posterior wart (paired); Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; X = segment X.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAB0874E7BCFB9F62D7FEFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223842/files/figure.png	https://doi.org/10.5281/zenodo.7223842	FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text.	FIGURE 3. Psilotreta spatulata sp. nov. 3A, male head, dorsal; 3B, male right wings, dorsal; 3C, female right wings, dorsal. 3D–3K, male genitalia: 3D, dorsal; 3E, variation of segment X, dorsal; 3F1–3F2, variations of median dorsal process, dorsal; 3G, ventral; 3H, left lateral; 3I1–3I4, variations of lateral process (paired) and intermediate appendage (paired), left lateral; 3J, phallus, left lateral; 3K, same ventral. 3L–3M, female genitalia: 3L, dorsal; 3M, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; dc = discoidal cell; int.app. = intermediate appendage (paired); lat. pr. = lateral process (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; mdp. tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; v.pr. = ventral projection (paired); va. = vaginal apparatus.Arrows see text.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0874E7BCFEC8620AFBE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223850/files/figure.png	https://doi.org/10.5281/zenodo.7223850	FIGURE 6. Psilotreta voluta Kawase 2021. 6A–6E, male genitalia: 6A, dorsal; 6B, left lateral; 6C1–6C2, variations of lateral process of tergum X and intermediate appendage, left lateral; 6D, phallus, left lateral; 6E, same, ventral. 6F, female genitalia, ventral. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 6A–6F, redrawn from the publication by Kawase (2021).	FIGURE 6. Psilotreta voluta Kawase 2021. 6A–6E, male genitalia: 6A, dorsal; 6B, left lateral; 6C1–6C2, variations of lateral process of tergum X and intermediate appendage, left lateral; 6D, phallus, left lateral; 6E, same, ventral. 6F, female genitalia, ventral. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process (paired); par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 6A–6F, redrawn from the publication by Kawase (2021).	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0874E7BCFEC8620AFBE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223862/files/figure.png	https://doi.org/10.5281/zenodo.7223862	FIGURE 10. Psilotreta spp. and their habitats. 10A, P. japonica (Banks 1906), male, dorsal (the locality name in the label “Aoyagi Shin-Asahi” needs to be corrected to “Kawashima Adogawa”). 10B, P. voluta Kawase 2021, female habitus, left dorsolateral (photo by N. Shimura). 10C, mountain stream where adults of P. kisoensis Iwata 1928 and P. voluta were collected at light. 10D, small stream where many adults of P. kisoensis were collected by a Malaise trap. 10E, pools of a small lowland stream inhabited by larvae of P. japonica.	FIGURE 10. Psilotreta spp. and their habitats. 10A, P. japonica (Banks 1906), male, dorsal (the locality name in the label “Aoyagi Shin-Asahi” needs to be corrected to “Kawashima Adogawa”). 10B, P. voluta Kawase 2021, female habitus, left dorsolateral (photo by N. Shimura). 10C, mountain stream where adults of P. kisoensis Iwata 1928 and P. voluta were collected at light. 10D, small stream where many adults of P. kisoensis were collected by a Malaise trap. 10E, pools of a small lowland stream inhabited by larvae of P. japonica.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0874E7BCFEC8620AFBE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0876E7BCFBF162C5FE86.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223852/files/figure.png	https://doi.org/10.5281/zenodo.7223852	FIGURE 7. Psilotreta bitubercula sp. nov. 7A–7E, male genitalia: 7A, dorsal; 7B, ventral; 7C, left lateral; 7D, phallus, left lateral; 7E, same, ventral. 7F–7G, female genitalia: 7F, dorsal; 7G, ventral. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). Arrow see text.	FIGURE 7. Psilotreta bitubercula sp. nov. 7A–7E, male genitalia: 7A, dorsal; 7B, ventral; 7C, left lateral; 7D, phallus, left lateral; 7E, same, ventral. 7F–7G, female genitalia: 7F, dorsal; 7G, ventral. Abbreviations: int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); mdp.tX = median dorsal process of tergum X; par. = paramere (paired); sIX = sternum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). Arrow see text.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0876E7BCFBF162C5FE86.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0876E7BCFBF162C5FE86.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223856/files/figure.png	https://doi.org/10.5281/zenodo.7223856	FIGURE 8. Psilotreta flavida Kawase 2021. 8A, head, dorsal; 8B, male right wings, dorsal; 8C, female right wings, dorsal. 8D–8G, male genitalia: 8D, dorsal; 8E, left lateral; 8F, phallus, left lateral; 8G, same, ventral. 8H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 8A–8B, 8D–8H.	FIGURE 8. Psilotreta flavida Kawase 2021. 8A, head, dorsal; 8B, male right wings, dorsal; 8C, female right wings, dorsal. 8D–8G, male genitalia: 8D, dorsal; 8E, left lateral; 8F, phallus, left lateral; 8G, same, ventral. 8H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 8A–8B, 8D–8H.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAE0876E7BCFBF162C5FE86.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223846/files/figure.png	https://doi.org/10.5281/zenodo.7223846	FIGURE 5 Psilotreta atrocaudata sp. nov. 5A, female right wings, dorsal; 5B–5G, male genitalia: 5B, dorsal; 5C, ventral; 5D, left lateral; 5E, lateral process of tergum X and intermediate appendage, left lateral; 5F, phallus, left lateral; 5G, same, ventral. 5H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X. Arrows see text.	FIGURE 5 Psilotreta atrocaudata sp. nov. 5A, female right wings, dorsal; 5B–5G, male genitalia: 5B, dorsal; 5C, ventral; 5D, left lateral; 5E, lateral process of tergum X and intermediate appendage, left lateral; 5F, phallus, left lateral; 5G, same, ventral. 5H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired); X = segment X. Arrows see text.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAC0876E7BCF9B46236F832.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7223856/files/figure.png	https://doi.org/10.5281/zenodo.7223856	FIGURE 8. Psilotreta flavida Kawase 2021. 8A, head, dorsal; 8B, male right wings, dorsal; 8C, female right wings, dorsal. 8D–8G, male genitalia: 8D, dorsal; 8E, left lateral; 8F, phallus, left lateral; 8G, same, ventral. 8H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 8A–8B, 8D–8H.	FIGURE 8. Psilotreta flavida Kawase 2021. 8A, head, dorsal; 8B, male right wings, dorsal; 8C, female right wings, dorsal. 8D–8G, male genitalia: 8D, dorsal; 8E, left lateral; 8F, phallus, left lateral; 8G, same, ventral. 8H, female genitalia, ventral. Abbreviations: I, II, III, V = apical forks I, II, III, V; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; int.app. = intermediate appendage (paired); lat.pr. = lateral process of tergum X (paired); M1, M2, M3, M4 = median veins 1–4; m-cu = crossvein between median and cubital veins; par. = paramere (paired); sIX = sternum IX; tIX = tergum IX; va. = vaginal apparatus; v.pr. = ventral projection (paired). 8A–8B, 8D–8H.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
E84587EDFFAC0876E7BCF9B46236F832.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7231471/files/figure.png	https://doi.org/10.5281/zenodo.7231471	FIGURE 11. Distributions of Japanese Psilotreta species.	FIGURE 11. Distributions of Japanese Psilotreta species.	2022-10-17	Kawase, Naoki		Zenodo	biologists	Kawase, Naoki			
