taxonID	type	description	language	source
03D379474868FFF6A0B1FCAD44D8021F.taxon	materials_examined	Type species: Alvania cimex (Linnaeus, 1758) by subsequent designation.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474868FFF8A0B1FF8344C2043E.taxon	description	(Fig. 3)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474868FFF8A0B1FF8344C2043E.taxon	materials_examined	Material examined. (more than five hundred shells, about half of them with soft parts, in 36 samples): SPAIN • 1 sh; 43 ° 51.265 ′ N, 008 ° 54.480 ′ W to 43 ° 51.498 ′ N, 008 ° 53.123 ′ W; 827 – 819 m; 08 – 15 SeP. 2002; DIVA-Artabria I DRN- 800 • 2 sh; 43 ° 53.457 ′ N, 008 ° 48.461 ′ W to 43 ° 54.000 ′ N, 008 ° 48.524 ′ W; 629 – 631 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 600 • 1 spc (juv) + 4 sh; 43 ° 47.188 ′ N, 008 ° 53.053 ′ W to 43 ° 55.312 ′ N, 008 ° 53.101 ′ W; 770 – 842 m; 08 – 15 SeP. 2002; DIVAArtabria I AT- 800 • 1 sPc + 6 sh; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 15 sh + 32 sPc; 43 ° 48.421 ′ N, 008 ° 51.453 ′ W to 43 ° 49.160 ′ N, 008 ° 51.091 ′ W; 599 – 607 m; 10 – 20 Sep. 2003; DIVA-Artabria I DRN- 600 • 9 sPc; 43 ° 53.575 ′ N, 008 ° 56.868 ′ W to 43 ° 54.015 ′ N, 008 ° 56.959 ′ W; 965 – 974 m; 10 – 20 Sep. 2003; DIVA-Artabria I DRN- 1000 • 7 sPc; 43 ° 48.514 ′ N, 008 ° 51.439 ′ W to 43 ° 49.163 ′ N, 008 ° 51.157 ′ W; 616 – 616 m; 10 – 20 SeP. 2003; DIVA-Artabria I AT- 600 • 1 sh; 43 ° 51.774 ′ N, 008 ° 53.640 ′ W to 43 ° 52.516 ′ N, 008 ° 53.478 ′ W; 798 – 801 m; 10 – 20 SeP. 2003; DIVA-Artabria I AT- 800 • 2 sPc + 5 sh; 43 ° 53.847 ′ N, 008 ° 57.324 ′ W to 43 ° 54.621 ′ N, 008 ° 57.261 ′ W; 993 – 1004 m; 10 – 20 SeP. 2003; DIVA-Artabria I AT- 1000 • 1 sPc; 43 ° 41.590 ′ N, 008 ° 45.328 ′ W to 43 ° 42.396 ′ N, 008 ° 44.286 ′ W; 301 – 301 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 300 •> 50 sh + 150 spc (100 juv); 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 6 sPc (4 juv); 43 ° 51.873 ′ N, 008 ° 53.683 ′ W to 43 ° 53.120 ′ N, 008 ° 53.301 ′ W; 802 – 788 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 800 • 2 sPc + 4 sh; 43 ° 38.812 ′ N, 009 ° 07.949 ′ W to 43 ° 39.841 ′ N, 009 ° 07.405 ′ W; 999 – 1001 m; 17 – 28 Sep. 2004; VERTIDOS GA-DRN- 1000. • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 720 m; SARRIDAL (2006 – 2007) SARRI- 1 • 12 sPc + 15 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 1 spc + 4 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 07) SARRI- 3 • 7 spc + 6 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 2007) SARRI- 4 • 25 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 •> 100 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 2 sh; 44 ° 08.461 ′ N, 009 ° 04.634 ′ W to 44 ° 08.607 ′ N, 009 ° 04.413 ′ W; 917 – 896 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 - 1 •> 100 sh + 10 spc; 44 ° 09.896 ′ N, 008 ° 39.581 ′ W to 44 ° 10.129 ′ N, 008 ° 39.494 ′ W; 438 – 459 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 • 1 sh; 44 ° 07.628 ′ N, 008 ° 45.871 ′ W to 44 ° 07.989 ′ N, 008 ° 45.542 ′ W; 412 – 411 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 - 4 • 2 sh; 44 ° 07.309 ′ N, 008 ° 48.289 ′ W to 44 ° 07.488 ′ N, 008 ° 48.047 ′ W; 445 – 438 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 - 5 • 7 sh; 43 ° 56.478 ′ N, 008 ° 54.199 ′ W to 43 ° 55.934 ′ N, 008 ° 54.849 ′ W; 620 – 933 m; 15 – 24 Jul. 2008; A SELVA DRN- 15 - 2 • 3 sh + 1 spc (juv); 43 ° 48.511 ′ N, 008 ° 51.393 ′ W to 43 ° 49.092 ′ N, 008 ° 50.959 ′ W; 312 – 576 m; 15 – 24 Jul. 2008; A SELVA DRN- 30 - 1 • 5 sh + 6 spc; 44 ° 07.495 ′ N, 008 ° 46.999 ′ W to 44 ° 07.675 ′ N, 008 ° 45.225 ′ W; 416 – 407 m; 15 – 24 Jul. 2008; A SELVA AT- 11 - 2 • 2 sh + 3 spc; 44 ° 06.496 ′ N, 008 ° 23.522 ′ W to 44 ° 07.16 ′ N, 008 ° 23.201 ′ W; 337 – 324 m; 15 – 24 Jul. 2008; A SELVA AT- 13 • 1 sPc; 44 ° 08.516 ′ N, 008 ° 40.788 ′ W to 44 ° 09.688 ′ N, 008 ° 39.721 ′ W; 436 – 428 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 • 1 sh + 3 spc; 44 ° 06.943 ′ N, 008 ° 47.232 ′ W to 44 ° 07.173 ′ N, 008 ° 46.031 ′ W; 440 – 429 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 - 2 • 4 spc; 44 ° 00.806 ′ N, 008 ° 33.098 ′ W to 44 ° 01.967 ′ N, 008 ° 32.57 ′ W; 346 – 344 m; 15 – 24 Jul. 2008; A SELVA EBS- 14 • 3 sh + 2 spc; 43 ° 58.84 ′ N, 008 ° 15.625 ′ W to 43 ° 59.621 ′ N, 008 ° 14.285 ′ W; 233 – 237 m; 15 – 24 Jul. 2008; A SELVA EBS- 20 • 9 sPc (juv); 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1 • 1 sh; 43 ° 24.95 ′ N, 009 ° 25.5 ′ W to 43 ° 25.6 ′ N, 009 ° 24.62 ′ W; 458 – 470 m; 15 – 30 SeP. 2008; DIVA-Artabria II DRNP- 07 • 1 sh; 43 ° 20.49 ′ N, 009 ° 34.03 ′ W to 43 ° 21.22 ′ N, 009 ° 32.96 ′ W; 846 – 770 m; 15 – 30 SeP. 2008; DIVA-Artabria II DRNP- 16 • 1 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 04 • 2 sh; 43 ° 32.5 ′ N, 009 ° 25.37 ′ W to 43 ° 33.6 ′ N, 009 ° 24.5 ′ W; 848 – 1027 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 05 • 8 sh; 43 ° 24.63 ′ N, 009 ° 31.55 ′ W to 43 ° 25.67 ′ N, 009 ° 30.73 ′ W; 880 – 827 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 08 • 4 sh + 1 sPc; 42 ° 55.76 ′ N, 009 ° 44.04 ′ W to 42 ° 55.79 ′ N, 009 ° 44.57 ′ W; 709 – 728 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 25.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474868FFF8A0B1FF8344C2043E.taxon	discussion	Remarks. It is a very common bathyal rissoid (in a wide depth range) known from high latitudes (Greenland, Iceland, and along the continental slopes from Norway) to the Cape Verde Islands (including Azores, Madeira and Canaries) and into the Mediterranean (Fretter & Graham 1978; Bouchet & Warén 1993; Hoenselaar & Goud 1998; Gofas 2007; Hoffman et al. 2011; Bitlis & Öztürk 2017). The shells studied conform to the previous descriptions of the species. One of the shells studied is shown in Figure 3 a, as well as its protoconch (Figs. 3 c – e) and details of the microsculpture of the teleoconch (Fig. 3 b) and protoconch (Figs. 3 f – g). It is characterized by its rectangular reticulate patterns with small nodules at the intersections of axial ribs and spirals cords. Multispiral protoconch of almost 3 whorls with sculpture of irregular spiral zig-zag elements, forming continuous lines near the suture. In the Mediterranean it has been regarded as a preferential characteristic element of the bathyal mud and deepsea white corals biocoenoses, usually in 100 – 1000 m (Negri & Corselli 2016). Alvania rykeli Hoenselaar & Goud, 1998, from around circalittoral and bathyal bottoms around Cape Verde Islands, strongly resembles A. cimicoides as far as the multispiral protoconch is concerned but differs in having much fewer axial (12 – 14 instead of 22 – 25) and spiral ribs (7 – 8 instead of 10 – 11). Among the samples studied, it was collected in the depth range 300 – 1200 m, but it was especially frequent in those collected at around 600 m in sandy bottoms with phosphorites.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474866FFFAA0B1F99B43850126.taxon	description	(Fig. 4)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474866FFFAA0B1F99B43850126.taxon	materials_examined	Material examined. (more than 250 shells, 154 with soft parts): SPAIN • 1 sPc; 43 ° 52.823 ′ N, 008 ° 56.151 ′ W to 43 ° 52.837 ′ N, 008 ° 55.597 ′ W; 988 – 920 m; 08 – 15 SeP. 2002; DIVA-Artabria I DRN- 1000 • 2 sh; 43 ° 53.457 ′ N, 008 ° 48.461 ′ W to 43 ° 54.000 ′ N, 008 ° 48.524 ′ W; 629 – 631 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT – 600 • 1 sh (juv); 43 ° 47.188 ′ N, 008 ° 53.053 ′ W to 43 ° 55.312 ′ N, 008 ° 53.101 ′ W; 770 – 842 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 800 • 3 sPc + 4 sh; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 10 sPc + 10 sh (1 juv); 43 ° 48.421 ′ N, 008 ° 51.453 ′ W to 43 ° 49.160 ′ N, 008 ° 51.091 ′ W; 599 – 607 m; 10 – 20 Sep. 2003; DIVA-Artabria I DRN- 600 • 1 sPc; 43 ° 53.575 ′ N, 008 ° 56.868 ′ W to 43 ° 54.015 ′ N, 008 ° 56.959 ′ W; 965 – 974 m; 10 – 20 Sep. 2003; DIVA-Artabria I DRN- 1000 • 1 sh; 43 ° 48.514 ′ N, 008 ° 51.439 ′ W to 43 ° 49.163 ′ N, 008 ° 51.157 ′ W; 616 – 616 m; 10 – 20 SeP. 2003; DIVA-Artabria I AT- 600 • 1 sh (juv); 43 ° 42.348 ′ N, 008 ° 45.889 ′ W to 43 ° 43.269 ′ N, 008 ° 45.289 ′ W; 344 – 354 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 350 • 90 sPc + 20 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 3 sh; 43 ° 51.873 ′ N, 008 ° 53.683 ′ W to 43 ° 53.120 ′ N, 008 ° 53.301 ′ W; 802 – 788 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 800 • 2 sPc + 2 sh; 43 ° 38.812 ′ N, 009 ° 07.949 ′ W to 43 ° 39.841 ′ N, 009 ° 07.405 ′ W; 999 – 1001 m; 17 – 28 SeP. 2004; VERTIDOS GA-DRN- 1000 SARRIDAL (2006 – 07) • 2 spc + 11 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 5 sPc + 5 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 07) SARRI- 3 • 5 sPc + 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 2007) SARRI- 4 • 19 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 •> 50 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 3 sh; 44 ° 08.461 ′ N, 009 ° 04.634 ′ W to 44 ° 08.607 ′ N, 009 ° 04.413 ′ W; 917 – 896 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 - 1 • 30 sPc + 70 sh; 44 ° 09.896 ′ N, 008 ° 39.581 ′ W to 44 ° 10.129 ′ N, 008 ° 39.494 ′ W; 438 – 459 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 • 5 sh; 44 ° 07.309 ′ N, 008 ° 48.289 ′ W to 44 ° 07.488 ′ N, 008 ° 48.047 ′ W; 44 ° 5 – 438 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 - 5 • 1 sPc; 43 ° 48.511 ′ N, 008 ° 51.393 ′ W to 43 ° 49.092 ′ N, 008 ° 50.959 ′ W; 312 – 576 m; 15 – 24 Jul. 2008; A SELVA DRN- 30 - 1 • 1 sPc + 1 sh; 44 ° 07.495 ′ N, 008 ° 46.999 ′ W to 44 ° 07.675 ′ N, 008 ° 45.225 ′ W; 416 – 407 m; 15 – 24 Jul. 2008; A SELVA AT- 11 - 2 • 1 sh; 44 ° 14.929 ′ N, 008 ° 30.255 ′ W to 44 ° 15.367 ′ N, 008 ° 30.438 ′ W; 2121 – 2516 m; 15 – 24 Jul. 2008; A SELVA AT- 12 • 7 sPc; 43 ° 58.84 ′ N, 008 ° 15.625 ′ W to 43 ° 59.621 ′ N, 008 ° 14.285 ′ W; 233 – 237 m; 15 – 24 Jul. 2008; A SELVA EBS- 20 • 3 sPc (juv); 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1 • 2 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ° ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 04.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474866FFFAA0B1F99B43850126.taxon	discussion	Remarks. It extends from higher latitudes (southeastern Greenland, Iceland, and Scandinavia) southwards to Portugal, usually in 50 – 500 m (Rodríguez-Babío & Thiriot-Quiévreux 1974; Fretter & Graham 1978; Bouchet & Warén 1993; Warén 1996; Hoffman et al. 2011). The shells studied show the characteristic square reticulate sculpture (Figs. 4 a – b). High magnification reveals minute spires and fuzzy growth lines (Fig. 4 e). Blunt protoconch with a marked zigzag sculpture (Figs. 4 c – d). According to Gofas & Warén (1982) records from the Mediterranean are based on Alvania sororcula GranataGrillo, 1877. Alvania jeffreysi is characterized by the zig-zag sculpture of its protoconch (Figs. 4 c – d), which it shares with some sublittoral Alvania, such as A. zylensis Gofas & Warén, 1982, A. imperspicua (Pallary, 1920) and A. vermaasi van Aartsen, 1975. In the studied samples it was present between 300 and 1000 m, mainly in the range 400 – 650 m where many shells had remains of the soft parts. It was abundant in the same samples than Alvania cimicoides.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474864FFFAA0B1FCF942C60479.taxon	description	(Figs. 5 a – d)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474864FFFAA0B1FCF942C60479.taxon	materials_examined	Material examined. (16 empty shells and 3 with soft parts): SPAIN • 1 sh; 43 ° 52.823 ′ N, 008 ° 56.151 ′ W to 43 ° 52.837 ′ N, 008 ° 55.597 ′ W; 988 – 920 m; 08 – 15 SeP. 2002; DIVA-Artabria I DRN- 1000 • 1 sh; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 1 sh: 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 150 • 2 sh; 43 ° 26.703 ′ N, 008 ° 30.669 ′ W to 43 ° 27.452 ′ N, 008 ° 29.612 ′ W; 103 – 102 m; 10 – 20 SeP. 2003; DIVAArtabria I EBS- 100 • 6 sh (juv); 43 ° 29.412 ′ N, 008 ° 28.362 ′ W to 43 ° 29.779 ′ N, 008 ° 28.240 ′ W; 110 – 111 m; 17 - 28 Sep. 2004; VERTIDOS GA-AT- 110 • 3 sPc + 8 sh; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 Sep. 2004; VERTIDOS AG-EBS- 150.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474864FFFAA0B1FCF942C60479.taxon	discussion	Remarks. This is a characteristic species of northeastern Atlantic continental shelf from the White Sea (Nekhaev 2013) to the Strait of Gibraltar (absent from Baltic and eastern basin of English Channel), south to the Alborán Sea, rare and rather local in the remainder of the Mediterranean (Gofas & Warén 1982). In the northern part of its range, it may be found intertidally, whereas it is usually found in circalittoral bottoms in the Mediterranean. Few shells were recorded on Gorringe and Ampère seamounts (Gofas 2007). Two of the shells studied, adult and juvenile, are shown in Figure 5 a – b, and a detail of the microsculpture can be seen in Figure 5 d. The protoconch in the studied shells was eroded (Fig. 5 c). Shells with soft parts were only found in one of the studied sampled collected at 150 m in muddy sand bottom.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474864FFFCA0B1F9D345FB0126.taxon	description	(Figs. 5 e – g)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474864FFFCA0B1F9D345FB0126.taxon	materials_examined	Material examined. (nearly 2000 shells, most of them with soft parts, in 33 samples): SPAIN • 1 s; 43 ° 40.165 ′ N, 008 ° 43.697 ′ W to 43 ° 40.513 ′ N, 008 ° 43.159 ′ W; 204 – 209 m; 08 – 15 SeP. 2002; DIVA-Artabria I DRN- 200 •> 80 sPc; 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 150 • 40 spc; 43 ° 40.192 ′ N, 008 ° 43.760 ′ W to 43 ° 40.943 ′ N, 008 ° 42.366 ′ W; 207 – 212 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 200 • 10 sPc; 43 ° 41.689 ′ N, 008 ° 45.195 ′ W to 43 ° 42.556 ′ N, 008 ° 44.226 ′ W; 298 – 303 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 300 • 1 spc (juv); 43 ° 53.575 ′ N, 008 ° 56.868 ′ W to 43 ° 54.015 ′ N, 008 ° 56.959 ′ W; 965 – 974 m; 10 – 20 SeP. 2003; DIVA-Artabria I DRN- 1000 •> 80 sPc; 43 ° 34.116 ′ N, 008 ° 36.535 ′ W to 43 ° 34.689 ′ N, 008 ° 35.589 ′ W; 149 – 148 m; 10 – 20 Sep. 2003; DIVA-Artabria I AT- 150 •> 200 spc; 43 ° 34.127 ′ N, 008 ° 36.562 ′ W to 43 ° 34.820 ′ N, 008 ° 35.585 ′ W; 152 – 149 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 150 •> 50 spc; 43 ° 40.250 ′ N, 008 ° 43.755 ′ W to 43 ° 40.760 ′ N, 008 ° 42.120 ′ W; 207 – 197 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 200 •> 300 spc; 43 ° 41.590 ′ N, 008 ° 45.328 ′ W to 43 ° 42.396 ′ N, 008 ° 44.286 ′ W; 301 – 301 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 300 • 1 sh; 43 ° 42.348 ′ N, 008 ° 45.889 ′ W to 43 ° 43.269 ′ N, 008 ° 45.289 ′ W; 344 – 354 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 350 • 30 spc + 200 sh; 43 ° 29.412 ′ N, 008 ° 28.362 ′ W to 43 ° 29.779 ′ N, 008 ° 28.240 ′ W; 110 – 111 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 110 • 50 sPc; 43 ° 32.026 ′ N, 008 ° 37.520 ′ W to 43 ° 32.806 ′ N, 008 ° 35.993 ′ W; 150 – 151 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 150 • 1 spc (juv); 43 ° 38.105 ′ N, 009 ° 02.006 ′ W to 43 ° 37.738 ′ N, 009 ° 00.905 ′ W; 836 – 826 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 800 • 30 sPc; 42 ° 29.455 ′ N, 009 ° 17.472 ′ W to 42 ° 31.292 ′ N, 009 ° 19.660 ′ W; 149 – 147 m; 17 – 28 SeP. 2004; VERTIDOS AG-AT- 150 •> 40 sPc; 42 ° 47.353 ′ N, 009 ° 25.161 ′ W to 42 ° 44.866 ′ N, 009 ° 25.913 ′ W; 147 – 146 m; 17 – 28 SeP. 2004; VERTIDOS CA-AT- 150 • 30 spc; 43 ° 31.970 ′ N, 008 ° 37.670 ′ W to 43 ° 32.697 ′ N, 008 ° 3.171 ′ W; 151 – 151 m; 17 – 28 SeP. 2004; VERTIDOS GA-EBS- 150 • 25 spc; 43 ° 36.222 ′ N, 008 ° 52.845 ′ W to 43 ° 36.797 ′ N, 008 ° 52.112 ′ W; 201 – 201 m; 17 – 28 Sep. 2004; VERTIDOS GA-EBS- 200 •> 800 spc; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 Sep. 2004; VERTIDOS AG-EBS- 150 •> 50 spc; 42 ° 31.176 ′ N, 009 ° 23.380 ′ W to 42 ° 32.132 ′ N, 009 ° 24.151 ′ W; 242 – 248 m; 17 – 28 Sep. 2004; VERTIDOS AG-EBS- 250 •> 200 sPc; 42 ° 50.507 ′ N, 009 ° 25.773 ′ W to 42 ° 48.810 ′ N, 009 ° 25.198 ′ W; 151 – 148 m; 17 – 28 SeP. 2004; VERTIDOS CA-EBS- 150 • 30 sPc; 42 ° 15.780 ′ N, 009 ° 10.500 ′ W to 42 ° 17.100 ′ N, 009 ° 10.630 ′ W; 149 – 150 m; 17 – 28 SeP. 2004; VERTIDOS CH-EBS- 150 • 8 sh; 43 ° 29.13 ′ N, 008 ° 43.233 ′ W to 43 ° 29.596 ′ N, 008 ° 42.838 ′ W; 149 – 151 m; 15 – 24 Jul. 2008; A SELVA DRN- 22 • 14 spc; 43 ° 31.727 ′ N, 008 ° 52.374 ′ W to 43 ° 32.678 ′ N, 008 ° 52.687 ′ W; 173 – 179 m; 15 – 24 Jul. 2008; A SELVA DRN- 23 • 12 sPc; 44 ° 06.496 ′ N, 008 ° 23.522 ′ W to 44 ° 07.16 ′ N, 008 ° 23.201 ′ W; 337 – 324 m; 15 – 24 Jul. 2008; A SELVA AT- 13 • 1 spc + 1 sh; 44 ° 08.516 ′ N, 008 ° 40.788 ′ W to 44 ° 09.688 ′ N, 008 ° 39.721 ′ W; 436 – 428 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 • 14 sPc; 44 ° 05.687 ′ N, 008 ° 24.558 ′ W to 44 ° 07.253 ′ N, 008 ° 24.302 ′ W; 337 – 330 m; 15 – 24 Jul. 2008; A SELVA EBS- 13 • 15 sPc; 44 ° 00.806 ′ N, 008 ° 33.098 ′ W to 44 ° 01.967 ′ N, 008 ° 32.57 ′ W; 346 – 344 m; 15 – 24 Jul. 2008; A SELVA EBS- 14 • 17 sPc + 10 sh; 43 ° 58.84 ′ N, 008 ° 15.625 ′ W to 43 ° 59.621 ′ N, 008 ° 14.285 ′ W; 233 – 237 m; 15 – 24 Jul. 2008; A SELVA EBS- 20 •> 50 sPc; 43 ° 31.687 ′ N, 008 ° 52.188 ′ W to 43 ° 33.149 ′ N, 008 ° 53.142 ′ W; 171 – 182 m; 15 – 24 Jul. 2008; A SELVA EBS- 23 • 5 sPc; 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1 • 4 sPc + 3 sh; 43 ° 33.20 ′ N, 008 ° 58.78 ′ W to 43 ° 34.57 ′ N, 008 ° 56.90 ′ W; 230 – 236 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 01 • 1 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 04 • 2 sPc; 43 ° 19.92 ′ N, 009 ° 38.13 ′ W to 43 ° 21.25 ′ N, 009 ° 37.11 ′ W; 1267 – 1286 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 15 • 1 sh; 42 ° 31.58 ′ N, 009 ° 40.14 ′ W to 42 ° 33.87 ′ N, 009 ° 38.75 ′ W; 2030 – 1886 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 30.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474864FFFCA0B1F9D345FB0126.taxon	discussion	Remarks. Very common species that extends from Norway to off western Morocco and throughout the Mediterranean, in a wide bathymetric range 30 – 1000 m, occasionally deeper (Oliverio et al. 1992; Bouchet & Warén 1993; Warén 1996; Cachia et al. 1996; Giribet & Peñas 1997; Gofas & Oliver 2011; Negri & Corselli 2016; Bitlis & Öztürk 2017). The shell is characterized by the opisthocline orientation of the peristome (Fig. 5 e). Protoconch sculptured by an irregular network (Figs. 5 f – g). It was the most abundant species in studied the samples along with Obtusella macilenta, predominantly between 100 and 400 m in muddy sand bottoms. Its putative planktrotrophic larval development (deduced from its protoconch) enables a large dispersal capacity.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474862FFFCA0B1FCE445C0043F.taxon	description	(Fig. 6)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474862FFFCA0B1FCE445C0043F.taxon	materials_examined	Material examined. (6 empty shells and 6 with soft parts in 4 samples): SPAIN • 1 sh; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 1 sh; 43 ° 58.84 ′ N, 008 ° 15.625 ′ W to 43 ° 59.621 ′ N, 008 ° 14.285 ′ W; 233 – 237 m; 15 – 24 Jul. 2008; A SELVA EBS- 20 • 1 sh; 43 ° 32.89 ′ N, 009 ° 30.88 ′ W to 43 ° 33.79 ′ N, 009 ° 29.60 ′ W; 1211 – 1207 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 10 • 6 sPc + 3 sh; 42 ° 45.9 ′ N, 009 ° 41.68 ′ W to 42 ° 47.00 ′ N, 009 ° 42.12 ′ W; 1499 – 1373 m; 15 – 30 SeP. 2008; DIVAArtabria II EBS- 27.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474862FFFCA0B1FCE445C0043F.taxon	discussion	Remarks. This species ranges from Norway to off Morocco, also in the Mediterranean, usually in a wide bathymetric range of 200 – 2000 m (Bouchet & Warén 1993; Warén 1996). In the Mediterranean it has been recorded from 200 down to 700 m in depth, probably as a Pleistocene subfossil (Gaglini 1991; Oliverio et al. 1992). Records from the Canaries (Bouchet & Warén 1993; Hoenselaar & Goud 1998) are extremely scanty and need confirmation. The record from shallow water in Madeira is based (Segers et al. 2009) on the misidentification of Alvania spreta (Watson, 1873). Alvania subsoluta resembles A. testae from which it mainly differs by having a paucispiral protoconch (Figs. 6 d – f). The shells studied show some variability being characterized by their square reticulate sculpture. The interspaces bear a micropattern of very thin spiral lines (Fig. 6 h). Paucispiral protoconch bearing also delicate spiral lines (Figs. 6 f – e). Few shells with soft parts remains were found in a sample from muddy bottom with gravel deeper than A. testae and A. cimicoides (at 1400 – 1500 m).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474862FFFDA0B1F99C43F600BA.taxon	description	(Fig. 7)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474862FFFDA0B1F99C43F600BA.taxon	materials_examined	Material examined. (9 empty shells and 10 with soft parts, in 11 samples): SPAIN • 1 sPc (juv) + 2 sh; 43 ° 48.421 ′ N, 008 ° 51.453 ′ W to 43 ° 49.160 ′ N, 008 ° 51.091 ′ W; 599 – 607 m; 10 – 20 SeP. 2003; DIVA-Artabria I DRN- 600 • 6 spc (4 juv) + 12 sh (4 juv); 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 1 sh; 43 ° 51.873 ′ N, 008 ° 53.683 ′ W to 43 ° 53.120 ′ N, 008 ° 53.301 ′ W; 802 – 788 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 800 • 1 sPc; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 1 sPc; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 2007) SARRI- 4 • 6 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 • 25 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 2 sh; 44 ° 09.896 ′ N, 008 ° 39.581 ′ W to 44 ° 10.129 ′ N, 008 ° 39.494 ′ W; 438 – 459 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 • 1 sPc; 44 ° 07.309 ′ N, 008 ° 48.289 ′ W to 44 ° 07.488 ′ N, 008 ° 48.047 ′ W; 445 – 438 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 - 5 • 1 sh; 44 ° 07.495 ′ N, 008 ° 46.999 ′ W to 44 ° 07.675 ′ N, 008 ° 45.225 ′ W; 416 – 407 m; 15 – 24 Jul. 2008; A SELVA AT- 11 - 2 • 1 sPc; 4211.73 ′ N, 009 ° 26.19 ′ W to 42 ° 11.73 ′ N, 009 ° 25.63 ′ W; 445 – 326 m; 14 – 25 Feb. 2009; FORSAGAL DRC- 6.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474862FFFDA0B1F99C43F600BA.taxon	discussion	Remarks. It is a common species of the continental slope in the north Atlantic, from Norway to Cape Verde Islands and in the Canary Islands (Bouchet & Warén 1993; Hoenselaar & Goud 1998; Hoffman et al. 2011), also present in the western Mediterranean (Gofas & Oliver 2011). Shells with soft parts were found in samples from 350 to 600 m in sandy bottoms, and some empty shells down to 1000 m depth. The strong sculpture of the shells forms knobs at the intersection of axial ribs and spiral cords (Fig. 7 a). Microsculpture of indistinct granulose spiral threads closely spaced (Figs. 7 e – g). Multispiral protoconch sculptured with irregular dashes forming spiral threads adapically (Figs. 7 b – c). Embryonic protoconch exhibiting a delightful reticular sculpture (Fig. 7 d).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474860FFFEA0B1F8FC466707DF.taxon	description	(Fig. 8)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474860FFFEA0B1F8FC466707DF.taxon	materials_examined	Material examined. (5 empty shells in the same sample): • 5 sh; 44 ° 09.896 ′ N, 008 ° 39.581 ′ W to 44 ° 10.129 ′ N, 008 ° 39.494 ′ W; 438 – 459 m; 15 – 24 Jul. 2008; A SELVA DRN- 11.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474860FFFEA0B1F8FC466707DF.taxon	discussion	Remarks. This species was only known previously from two specimens found off Portugal, one the holotype (from PORCUPINE Expedition, at 1809 m depth) and the other from THALASSA expedition at 810 m (Bouchet & Warén 1993). Therefore, the five shells found in one of the samples studied at about 450 m depth extend the knowledge on this rare species and expand its distribution area and bathymetric range. They are fusiform, up to 2.7 x 1.4 mm, with a high spire, whose general shape resembles some European species of Onoba (Figs. 8 a – b). Sculpture of weak spiral cords and axial ribs of similar width somewhat narrower than their respective interspaces. Microsculpture of fine striae (Fig. 8 c). Oval aperture with thin outer lip. Dome-shaped paucispiral protoconch (Figs. 8 d, g) with fine equidistant spiral threads starting from the nucleus (Fig. 8 h). Some details of its microsculpture are shown in Figure 8 e – i.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487EFFE1A0B1FDC546040356.taxon	description	(Fig. 9)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487EFFE1A0B1FDC546040356.taxon	materials_examined	Material examined. (4 empty shells and 6 with soft parts, in 6 samples): SPAIN • 1 sPc; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 2 sPc + 2 sh; 43 ° 36.544 ′ N, 009 ° 03.064 ′ W to 43 ° 36.816 ′ N, 009 ° 04.339 ′ W; 601 – 619 m; 17 – 28 SeP. 2004; VERTIDOS GA-EBS- 600 • 1 spc; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 2007) SARRI- 4 • 1 sh; 44 ° 08.516 ′ N, 008 ° 40.788 ′ W to 44 ° 09.688 ′ N, 008 ° 39.721 ′ W; 436 – 428 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 • 1 sPc + 1 sh; 44 ° 08.516 ′ N, 008 ° 40.788 ′ W to 44 ° 09.688 ′ N, 008 ° 39.721 ′ W; 436 – 428 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 - 2 • 1 spc; 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487EFFE1A0B1FDC546040356.taxon	discussion	Remarks. This species has been recorded from the southern part of the Bay of Biscay to off the Canaries and Mediterranean Sea (where the type locality is located) in a wide depth range, normally in 150 – 800 m and occasionally deeper down to 2770 m (Bouchet & Warén 1993). We have found six shells with soft parts and four empty shells in samples collected between 400 and 700 m in muddy sand bottoms. Alvania electa is a rather variable species, as was pointed out by Bouchet & Warén (1993), and a cryptic species-complex may be involved. Its complex synonymy was unraveled by Oliverio et al. (1992). This species is characterized by its deeply channeled suture, low number of sharp axial ribs, narrow umbilical chink, and large and blunt apex (Figs. 9 a, d – f). Protoconch sculpture with irregular microgranules separated by about six equidistant fine riblets (Figs. 9 b, c, g – i). The specimens here studied match with those figured by Gofas & Warén (1982, figs. 24 and 32 as Alvania deliciosa (Jeffreys, 1884), Oliverio et al. (1992, figs. 17 – 20) and Bouchet & Warén (1993, figs. 1412, 1415 and 1420). In Figure 10 a – d we show for comparison a shell (as Alvania sp.) from the Djibouti Bank (Alborán Sea) whose protoconch is like that of A. electa, but because its convex whorls and the higher number of axial ribs it resembles Alvania porcupinae Gofas & Warén, 1882. On the other hand, the Mediterranean Alvania elegantissima (Monterosato, 1875) has sometimes been confused with A. electa, but according to Oliverio et al. (1992) its convex whorls and protoconch without sculpture allow differentiate both species. Figure 10 e – g shows a shell from off the Italian coast of the Tyrrhenian Sea that may be Alvania elegantissima by having opisthocline axial ribs and a protoconch lacking sculpture.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487CFFE3A0B1FDC546940102.taxon	description	(Fig. 11)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487CFFE3A0B1FDC546940102.taxon	materials_examined	Material examined. (1 empty shell): SPAIN • 1 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 Sep. 2008; DIVA-Artabria II EBS- 04.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487CFFE3A0B1FDC546940102.taxon	discussion	Remarks. Alvania porcupinae is known from the Bay of Biscay to off Canary Islands (not found in the Mediterranean) in a wide depth range (800 – 4700 m) (Bouchet & Warén 1993; Hoenselaar & Goud 1998), and it is one of the few rissoids occurring at abyssal depths (Sysoev 2014). Within the material studied only one shell (Fig. 11 a) was found in a sample collected at 1050 m in depth. According to Bouchet & Warén (1993) A. porcupinae belongs to the same lineage as A. electa, A. adinogramma and A. cancapae, characterized by similar teleoconch features and a finely granulated protoconch. As in the case of A. electa, this species is rather variable (see Bouchet & Warén 1993 in figures 1424 – 1429) and needs a deep revision. The shell here studied is quite similar to the one of figure 1426 in Bouchet & Warén (1993) from off southern Morocco, bearing square reticulate sculpture (Figs. 11 a, g), but it is somewhat smaller (1.9 mm compared to 2.3 mm in the latter) and with a low number of spiral cords. Suture slightly channeled. About forty axial ribs in the body whorl, almost orthocline, somewhat narrower than the interspaces which adapically deviated by a slight subsutural depression. The ribs are interrupted at the level of the apertural insertion, disappearing at the base. Flat spiral cords, slightly wider than the ribs and somewhat narrower than their interspaces. Paucispiral dome protoconch (Figs. 11 b – d) with almost smooth surface and abrupt transition to the teleoconch (Fig. 11 f).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487DFFE4A0B1FCC0438F01B6.taxon	description	(Fig. 12)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487DFFE4A0B1FCC0438F01B6.taxon	materials_examined	Material examined. (2 empty shells in the same sample): SPAIN • 2 sh; 44 ° 14.929 ′ N, 008 ° 30.255 ′ W to 44 ° 15.367 ′ N, 008 ° 30.438 ′ W; 2121 – 2516 m; 15 – 24 Jul. 2008; A SELVA AT- 12.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487DFFE4A0B1FCC0438F01B6.taxon	discussion	Remarks. Alvania cancapae is a variable species known from off southern Morocco and off Madeira and Canary islands in the depth range 270 – 2000 m (Bouchet & Warén, 1993; Hoenselaar & Goud, 1998) with stronger sculpture in shallower specimens. The two shells (of about 2.0 x 1.3 mm) found in one of the samples (collected deeper than 2000 m) have a weak axial sculpture (Fig. 12 a), with very thin spiral threads narrower than interspaces overrunning the axial ribs (Figs. 12 e, f), and faint spiral cords on the periumbilical area of the body whorl, but hardly visible on the spire. Protoconch of the shells studied of 1.25 convex whorls (Figs. 12 b, c) whose surface is covered by minute granules (Fig. 12 d). This species resembles to Alvania adinogramma Bouchet & Warén, 1993, Alvania microstriata Hoenselaar & Goud, 1998 and Alavania seinensis Gofas, 2007, a group which seems restricted to the Lusitanian seamounts and Madeira (Gofas 2007). The present records represent a northward range extension of the species.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487AFFE4A0B1FF14437207D6.taxon	description	(Fig. 13)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487AFFE4A0B1FF14437207D6.taxon	materials_examined	Type material. Holotype: SPAIN • 1 emPty shell (Fig. 13); 42 ° 51.20 ′ N, 009 ° 43.85 ′ W to 42 ° 52.97 ′ N, 009 ° 44.25 ′ W; 1861 – 1177 m; 15 – 30 Sep. 2008; DIVA-Artabria II DRNp- 26; MHNS, deposit number: MHN-USC- 10118. Paratypes: SPAIN • 3 sh (ParatyPes 1 – 3); 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7; MHN-USC, deposit number: MHN-USC- 10118; • 5 sh (1 juv) (paratypes 4 – 8); 43 ° 19.92 ′ N, 009 ° 38.13 ′ W to 43 ° 21.25 ′ N, 009 ° 37.11 ′ W; 1267 – 1286 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 15; MNCN 15.05 / 200226. Type locality: Muxía Canyon off W Galicia (42 ° 51.20 ′ – 42 ° 52.97 ′ N; 009 ° 43.85 ′ – 009 ° 44.25 ′ W, 1177 – 1861 m).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487AFFE4A0B1FF14437207D6.taxon	etymology	Derivatio nominis: This species is dedicated to Dra. María Candás who has participated in the collection of samples in all oceanographic campaigns and has participated in the sorting of the samples and procedure of the species obtained.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487AFFE4A0B1FF14437207D6.taxon	description	Description. Small oval shell (1.8 x 1.1 mm in the holotype with somewhat more than 4 whorls) with moderately high conical sPire and blunt aPex (Fig. 13 a), aPical angle of about 42 °, colour white. APerture Pyriform, Pointed at union with penultimate whorl, height 0.7 mm (about 38 % of total height). Teleoconch with slightly convex whorls and channeled suture highlighted by a subsutural spiral depression (Fig. 13 o). Sculpture of weak, axial ribs almost orthocline, at about 10 ° with sPire axis; flat sPiral cords towards the base and minute sPiral striae (Figs. 13 j – k) also in the channeled suture (Fig. 13 e). On the body whorl there are about 30 ribs disappearing at the base. Rounded aperture with a thin edge, not thickened. Umbilicus open but partly covered by columellar lip; umbilical area with chink and coarse growth lines (Figs. 13 a, f, l). Paucispiral protoconch of just over a whorl (Figs. 13 b, c, g, m) with inconspicuous sculpture of very weak axial threads and scattered or slightly spirally aligned microtubercles (Figs. 13 d, i). At the base of the nucleus certain roughness can be observed under high magnification (Fig. 13 h). Soft parts and radulae unknown.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947487AFFE4A0B1FF14437207D6.taxon	discussion	Remarks. Alvania candasae sp. nov. seems to belong to the same group as A. electa, A. porcupinae, A. adinogramma Bouchet & Warén, 1993 and A. cancapae Bouchet & Warén, 1993. This last species, from off Morocco and Canary and Madeira Islands, is larger (more than 2 mm) with less numerous and more pronounced axial ribs, as well as A. electa, whose protoconch has a microsculpture of fine spiral threads and microgranules, different from that of the new species. Alvania porcupinae is also larger (more than 2 mm), its suture is not so channeled and its protoconch surface is smooth. A. adinogramma has more convex whorls, less ribbed suture, and a conspicuous spiral sculpture throughout the shell. Alvania adiaphoros Bouchet & Warén, 1993 is somewhat larger, with axial ribs more widely separated and not extending on the abapical part of the body whorl, and its channeled suture is less pronounced. Also, A. microstriata and A. seinensis, from Madeira and Seine Seamount, resemble this new species in their shape and size, but they have a shallow subsutural furrow instead of a channeled suture and their protoconch surface are densely covered by very fine microtubercles. Nine empty shells were found in samples collected between 900 and 1600 m.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474878FFE6A0B1FDC5464907F7.taxon	description	(Fig. 14)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474878FFE6A0B1FDC5464907F7.taxon	materials_examined	Type material. Holotype: SPAIN • 1 emPty shell (Figs. 14 d – h); 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000; MHNS, dePosit number: MHN-USC- 10117. Paratypes: SPAIN • 2 sh (ParatyPes 1 and 2); 43 ° 51.265 ′ N, 008 ° 54.480 ′ W to 43 ° 51.498 ′ N, 008 ° 53.123 ′ W; 827 – 819 m; 08 – 15 Sep. 2002; DIVA-Artabria I DRN- 800; MNCN 15.05 / 200227 • 1 sh; 43 ° 52.823 ′ N, 008 ° 56.151 ′ W to 43 ° 52.837 ′ N, 008 ° 55.597 ′ W; 988 – 920 m; 08 – 15 SeP. 2002; DIVA-Artabria I DRN- 1000 • 2 sh; 43 ° 47.188 ′ N, 008 ° 53.053 ′ W to 43 ° 55.312 ′ N, 008 ° 53.101 ′ W; 770 – 842 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 800 • 1 spc + 22 sh; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 1 sPc + 1 sh; 43 ° 53.575 ′ N, 008 ° 56.868 ′ W to 43 ° 54.015 ′ N, 008 ° 56.959 ′ W; 965 – 974 m; 10 – 20 SeP. 2003; DIVA-Artabria I DRN- 1000 • 12 sPc + 3 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 600 • 12 sh; 43 ° 51.873 ′ N, 008 ° 53.683 ′ W to 43 ° 53.120 ′ N, 008 ° 53.301 ′ W; 802 – 788 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 800 • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 720 m; SARRIDAL (2006 – 2007) SARRI- 1 • 14 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 • 2 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 1 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 04. Type locality: Ferrol Canyon off Galicia coast (43 ° 57.030 ′ - 43 ° 57.248 ′ N, 008 ° 54.795 ′ W- 00854.133 ′ W, 1132 – 1191 m).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474878FFE6A0B1FDC5464907F7.taxon	etymology	Derivatio nominis: This species is dedicated to Dra. Xela Cunha Veira who has participated in the collection of samples in all oceanographic campaigns and for the custody and handling of the samples studied.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474878FFE6A0B1FDC5464907F7.taxon	description	Description. Shell with a moderately high sPire with conical outline, aPical angle of about 45 ° and a blunt aPex, somewhat solid, with conspicuous reticulated sculpture (Fig. 14 a). Holotype (Fig. 14 d) of 1.6 x 1.0 mm with four convex whorls. Adult shells studied up to 1.7 x 1.2 mm. Well-marked, somewhat channeled suture. Colour white. Aperture pyriform, pointed at union with penultimate whorl, height 0.56 mm (about 46 % of total height). Umbilical area with chink and coarse growth lines. About 20 sharP, orthocline axial ribs, at about 10 ° with sPire axis, narrower than interspaces on the body whorl and disappearing toward the base. Thick spiral cords, also narrower than interspaces, more developed on the periumbilical area of the body whorl. Entire surface covered by regularly distributed spiral threads, much narrower than their interspaces, often interrupted where crossing growth lines (Fig. 14 e). Outer lip slightly opisthocline with a thin edge, thickened externally and not thickened inside. Inner lip thin, bordering a narrow umbilicus. Radulae, operculum and soft parts have not been described since they show great uniformity in species of this genus. Specimens are kept for possible future studies. Protoconch of 1.25 convex whorls (Figs. 14 g – h) whose surface is covered by minute granules irregularly arranged (Fig. 14 c) overlapping wide, diffuse spiral cords (Fig. 14 b).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474878FFE6A0B1FDC5464907F7.taxon	discussion	Remarks. About 75 shells and specimens were found in 10 samples collected between 400 and 1200 m. Most of the shells with soft parts were found in a sample at around 600 m depth in sandy bottom. No notable variability was observed. Alvania xelae sp. nov. has been believed to fall within the variability of A. electa, and corresponds to one of the shells illustrated under that name by Bouchet & Warén (1993, figs. 1416 – 1417 from THALASSA st X 362 at 385 – 600 m off north Spain coasts). Alvania xelae sp. nov. differs from A. electa by its more convex whorls, not so deeply channelled suture and especially by its protoconch with wide diffuse spiral cords (Fig. 14 b) instead of fine striae (Figs. 9 b – c). This sculpture of the protoconch also differentiates it from the other species of the group of A. electa (A. porcupinae, A. adinogramma, A. cancapae and A. candasae sp. nov.). On the other hand, A. seinensis from the Seine seamount has a shell protoconch similar to that of the new species but its teleoconch has much finer and more numerous ribs (more than 40 on the penultimate whorl) and a less marked spiral sculpture (Gofas 2007, fig. 7 E – F).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474879FFE7A0B1FB17448306E1.taxon	materials_examined	Type species: Gofasia galiciae Bouchet and Warén, 1993, by original designation.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474879FFE8A0B1FBA646190612.taxon	description	(Fig. 15)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474879FFE8A0B1FBA646190612.taxon	materials_examined	Material examined. (103 empty shells and 59 with soft parts, in 24 samples): SPAIN • 1 frg; 43 ° 47.188 ′ N, 008 ° 53.053 ′ W to 43 ° 55.312 ′ N, 008 ° 53.101 ′ W; 770 – 842 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 800 • 1 sh + 1 juv; 43 ° 57.030 ′ N, 00854.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 2 spc + 2 sh; 43 ° 48.421 ′ N, 008 ° 51.453 ′ W to 43 ° 49.160 ′ N, 008 ° 51.091 ′ W; 599 – 607 m; 10 – 20 SeP. 2003; DIVA-Artabria I DRN- 600 • 4 sh; 43 ° 53.575 ′ N, 008 ° 56.868 ′ W to 43 ° 54.015 ′ N, 008 ° 56.959 ′ W; 965 – 974 m; 10 – 20 Sep. 2003; DIVA-Artabria I DRN- 1000 • 1 spc + 2 sh; 43 ° 53.847 ′ N, 008 ° 57.324 ′ W to 43 ° 54.621 ′ N, 008 ° 57.261 ′ W; 993 – 1004 m; 10 – 20 SeP. 2003; DIVA-Artabria I AT- 1000 • 1 sPc; 43 ° 42.348 ′ N, 008 ° 45.889 ′ W to 43 ° 43 °. 269 ′ N, 008 ° 45.289 ′ W; 344 – 354 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 350 • 30 spc + 20 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 - 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 4 sPc + 5 sh; 43 ° 51.873 ′ N, 008 ° 53.683 ′ W to 43 ° 53.120 ′ N, 008 ° 53.301 ′ W; 802 – 788 m; 10 – 20 SeP. 2003; DIVAArtabria I EBS- 800 • 2 sh; 43 ° 38.812 ′ N, 009 ° 07.949 ′ W to 43 ° 39.841 ′ N, 009 ° 07.405 ′ W; 999 – 1001 m; 17 – 28 Sep. 2004; VERTIDOS GA-DRN- 1000 • 3 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 Sep. 2003; DIVA-Artabria I GA-EBS- 600 • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 07) SARRI- 3 • 20 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 3 sh; 44 ° 08.461 ′ N, 009 ° 04.634 ′ W to 44 ° 08.607 ′ N, 009 ° 04.413 ′ W; 917 – 896 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 - 1 • 15 spc + 25 sh; 44 ° 09.896 ′ N, 008 ° 39.581 ′ W to 44 ° 10.129 ′ N, 008 ° 39.494 ′ W; 438 – 459 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 • 2 sh; 44 ° 07.628 ′ N, 008 ° 45.871 ′ W to 44 ° 07.989 ′ N, 008 ° 45.542 ′ W; 412 – 411 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 - 4 • 2 sh; 44 ° 07.309 ′ N, 008 ° 48.289 ′ W to 44 ° 07.488 ′ N, 008 ° 48.047 ′ W; 445 – 438 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 - 5 • 1 sh; 43 ° 55.886 ′ N, 008 ° 54.846 ′ W to 43 ° 55.291 ′ N, 008 ° 55.65 ′ W; 933 – 930 m; 15 – 24 Jul. 2008; A SELVA DRN- 15 - 2 B • 1 spc + 2 sh; 44 ° 07.495 ′ N, 008 ° 46.999 ′ W to 44 ° 07.675 ′ N, 008 ° 45.225 ′ W; 416 – 407 m; 15 – 24 Jul. 2008; A SELVA AT- 11 - 2 • 2 spc + 1 sh; 44 ° 08.516 ′ N, 008 ° 40.788 ′ W to 44 ° 09.688 ′ N, 008 ° 39.721 ′ W; 436 – 428 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 • 1 sPc; 43 ° 58.84 ′ N, 008 ° 15.625 ′ W to 43 ° 59.621 ′ N, 008 ° 14.285 ′ W; 233 – 237 m; 15 – 24 Jul. 2008; A SELVA EBS- 20 • 2 sPc; 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1 • 1 sh; 43 ° 19.95 ′ N, 009 ° 34.77 ′ W to 43 ° 20.58 ′ N, 009 ° 33.91 ′ W; 972 – 839 m; 15 – 30 SeP. 2008; DIVA-Artabria II DRNr- 16 • 2 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 Sep. 2008; DIVA-Artabria II EBS- 04 • 1 sh; 43 ° 24.63 ′ N, 009 ° 31.55 ′ W to 43 ° 25.67 ′ N, 009 ° 30.73 ′ W; 880 – 827 m; 15 – 30 Sep. 2008; DIVA-Artabria II EBS- 08.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474879FFE8A0B1FBA646190612.taxon	discussion	Remarks. According to Bouchet & Warén (1993) this species is known from the continental slope of the Celtic Sea southwards to northern Spain. The depth range of the studied specimens (350 – 1000 m) coincides with that reported by these authors. Most shells with soft parts were found in a sample on sandy bottom at 600 m deep. Adult shells studied up to 2.8 mm with large aperture and rounded axial ribs (Figs. 15 a, d, g). Microsculpture of minute pores aligned in a spiral lines can be observed under high magnification (Figs. 15 b – c). Protoconch sculptured by spiral furrows (Figs. e – h). Bouchet & Warén (1993) included this species in the genus Frigidoalvania because of the similarity of its teleoconch and protoconch with other species of this genus. However, due its microsculpture of minute pores we propose here to include this species in the genus Gofasia, also because of its resemblance to Gofasia vanderlandi Bouchet & Warén, 1993, from which it differs by the sculpture of the protoconch. Microsculpture of minute pores is the most important character to separate the genus from allied genera. Therefore, the species of Frigidoalvania remain restricted to higher latitudes. Gofasia galiciae, known from the Galicia Bank and Ampère and Josephine seamount (Bouchet & Warén 1993; Gofas 2007), has not been found in our samples.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474874FFEAA0B1FC17426207F7.taxon	description	(Fig. 16)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474874FFEAA0B1FC17426207F7.taxon	materials_examined	Material examined. (61 empty shells and 51 with soft parts in 22 samples): Smooth form: SPAIN • 2 sh; 43 ° 47.188 ′ N, 008 ° 53.053 ′ W to 43 ° 55.312 ′ N, 008 ° 53.101 ′ W; 770 – 842 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 800 • 3 spc + 1 sh; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 1 spc; 43 ° 48.514 ′ N, 008 ° 51.439 ′ W to 43 ° 49.163 ′ N, 008 ° 51.157 ′ W; 616 – 616 m; 10 – 20 SeP. 2003; DIVA-Artabria I AT- 600 • 1 spc; 43 ° 41.590 ′ N, 008 ° 45.328 ′ W to 43 ° 42.396 ′ N, 008 ° 44.286 ′ W; 301 – 301 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 300 • 4 sPc + 2 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 600 • 1 sPc; 43 ° 38.812 ′ N, 009 ° 07.949 ′ W to 43 ° 39.841 ′ N, 009 ° 07.405 ′ W; 999 – 1001 m; 17 – 28 SeP. 2004; VERTIDOS GA-DRN- 1000 • 2 sh; polygon delimited by points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 07) SARRI- 3 • 16 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 • 9 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 2 sPc; 44 ° 09.896 ′ N, 008 ° 39.581 ′ W to 44 ° 10.129 ′ N, 008 ° 39.494 ′ W; 438 – 459 m; 15 – 24 Jul. 2008; A SELVA DRN- 11 • 1 sPc; 43 ° 56.478 ′ N, 008 ° 54.199 ′ W to 43 ° 55.934 ′ N, 008 ° 54.849 ′ W; 620 – 933 m; 15 – 24 Jul. 2008; A SELVA DRN- 15 - 2 • 2 sPc; 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1 • 1 sh; 43 ° 34.86 ′ N, 009 ° 21.95 ′ W to 43 ° 36.44 ′ N, 009 ° 20.71 ′ W; 1010 – 1112 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 04 • 1 sh; 43 ° 32.5 ′ N, 009 ° 25.37 ′ W to 43 ° 33.6 ′ N, 009 ° 24.5 ′ W; 848 – 1027 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 05 • 1 sh; 42 ° 55.76 ′ N, 009 ° 44.04 ′ W to 42 ° 55.79 ′ N, 009 ° 44.57 ′ W; 709 – 728 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 25. Sculptured form: SPAIN • 25 sPc + 10 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 600 • 2 sPc; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 3 sPc; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 2007) SARRI- 4 • 12 sh; 44 ° 08.65 ′ N, 008 ° 55.305 ′ W to 44 ° 08.771 ′ N, 008 ° 55.104 ′ W; 581 – 566 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 C • 2 sh; 44 ° 14.929 ′ N, 008 ° 30.255 ′ W to 44 ° 15.367 ′ N, 008 ° 30.438 ′ W; 2121 – 2516 m; 15 – 24 Jul. 2008; A SELVA AT- 12 • 6 sPc; 44 ° 08.516 ′ N, 008 ° 40.788 ′ W to 44 ° 09.688 ′ N, 008 ° 39.721 ′ W; 436 – 428 m; 15 – 24 Jul. 2008; A SELVA EBS- 11 • 1 sh; 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 – 24 Jul. 2008; A SELVA EBS- 30 - 1.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474874FFEAA0B1FC17426207F7.taxon	discussion	Remarks. Bouchet & Warén (1993) studied many shells of this species widespread from off northern Spain south to the Canary Islands in 155 – 1650 m depth range. Gofas (2007) pointed out that Pseudosetia amydralox is quite common on the uppermost part of the Lusitanian seamounts (the type locality is located at Gorringe Seamount, 305 – 320 m). We found several shells of this species with soft parts in samples from the deep range 300 – 1100 m mainly in sandy bottoms with phosphorites. The shells here studied show a remarkable variability, that ranges from practically smooth surface, with regularly convex whorls, lacking suture edges and a peripheral cord on the last whorl (Figs. 16 a, c, e), to shells with a conspicuous spiral sculpture, with suture edges and a peripheral cord, and plano-convex whorls (Figs. 16 k, o, n, r, s). The sculpture of the protoconchs also showed a certain degree of variability (Figs. 16 b, d – j, l, m, p, q). Not ruling out the possibility of cryptic species, we prefer for now to consider variability of a single species, that due to its geographic location it situated within the range of distribution of P. amydralox instead of that of the smooth Pseudosetia turgida (Jeffreys, 1870) or the more sculptured Pseudosetia azorica Bouchet & Warén, 1993.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474874FFEAA0B1FDC5443300B7.taxon	materials_examined	Type species: Rissoa turgida Jeffreys, 1870 by subsequent designation.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474872FFECA0B1FDC5441D00B7.taxon	materials_examined	Type species: Benthonella gaza Dall, 1889, by original designation.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474872FFEDA0B1FC17462F02EA.taxon	description	(Fig. 17)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474872FFEDA0B1FC17462F02EA.taxon	materials_examined	Material examined. (15 empty shells and 9 with soft parts in 7 samples): SPAIN • 3 sh + 2 spc; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 2 sPc; 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 150 • 1 sh + 3 sPc; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 4 sPc + 7 sh; 42 ° 45.9 ′ N, 009 ° 41.68 ′ W to 42 ° 47.00 ′ N, 009 ° 42 °. 12 ′ W; 1499 – 1373 m; 15 – 30 Sep. 2008; DIVA-Artabria II EBS- 27 • 3 sh; 42 ° 38.21 ′ N, 009 ° 51.46 ′ W to 42 ° 39.45 ′ N, 009 ° 50.73 ′ W; 1961 – 1971 m; 15 – 30 Sep. 2008; DIVA-Artabria II EBS- 28 • 1 sPc; 42 ° 27.66 ′ N, 009 ° 46.13 ′ W to 42 ° 29.08 ′ N, 009 ° 45.31 ′ W; 2033 – 2091 m; 15 – 30 SeP. 2008; DIVA-Artabria II EBS- 29 • 1 sh; 42 ° 29.55 ′ N, 01010.5 ′ W to 42 ° 30.08 ′ N, 01 ° 009.57 ′ W; 2798 – 2765 m; 14 – 25 Feb. 2009; FORSAGAL EBS- 4.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474872FFEDA0B1FC17462F02EA.taxon	discussion	Remarks. Benthonella tenella is the most common Atlantic and Mediterranean bathyal rissoid species, with the widest geographic and bathymetric range of all. It is distributed on both sides of the Atlantic, from Iceland to the Caribbean in the west and to the Macaronesian islands in the east, and in the whole Mediterranean basin, being also patchily present in the southern Atlantic (Bouchet & Warén 1993; Hoffman et al. 2011; Sysoev 2014; Negri & Corselli 2016). It is also common in the northeastern Atlantic seamounts (Gofas 2007). Its planktotrophic development with ontogenetic vertical migration allows the potential for large-scale dispersal (Rex & Etter 1990). Nevertheless, in the samples studied it has not been very abundant, having been found between 1400 and 2100 m. The few shells found in the samples belong to the smooth morph (Fig. 17 a). Multispiral protoconch with just over three whorls. The embryonic whorls show three fine spiral lines and microgranules in between (Figs. 17 d – e, g). Subsequent whorls (protoconch II) with two suprasutural spiral threads (Figs. 17 b, f), the upper one like a keel framing a flattened shoulder (Fig. 17 c).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474873FFEDA0B1FFA84462031B.taxon	materials_examined	Type species: Obtusella intersecta (S. Wood, 1857), by monotypy.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474873FFEDA0B1FEFB43E506D0.taxon	description	(Fig. 18)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474873FFEDA0B1FEFB43E506D0.taxon	materials_examined	Material examined. (46 empty shells and 29 with soft parts, in 9 samples): SPAIN • 1 sPc + 1 sh; 43 ° 40.192 ′ N, 008 ° 43.760 ′ W to 43 ° 40.943 ′ N, 008 ° 42 °. 366 ′ W; 207 – 212 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 200 • 1 sh; 43 ° 26.703 ′ N, 008 ° 30.669 ′ W to 43 ° 27.452 ′ N, 008 ° 29.612 ′ W; 103 – 102 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 100 • 5 sh; 43 ° 29.412 ′ N, 008 ° 28.362 ′ W to 43 ° 29.779 ′ N, 008 ° 28.240 ′ W; 110 – 111 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 110 • 1 sPc; 43 ° 32.026 ′ N, 008 ° 37.520 ′ W to 43 ° 32.806 ′ N, 008 ° 35.993 ′ W; 150 – 151 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 150 • 25 sPc + 15 sh; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 Sep. 2004; VERTIDOS AG-EBS- 150 • 1 sPc; 42 ° 29.455 ′ N, 009 ° 17.472 ′ W to 42 ° 31.292 ′ N, 009 ° 19.660 ′ W; 149 – 147 m; 17 – 28 SeP. 2004; VERTIDOS AG-AT- 150 • 2 sh; 42 ° 31.176 ′ N, 009 ° 23.380 ′ W to 42 ° 32.132 ′ N, 009 ° 24.151 ′ W; 242 – 248 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 250 • 1 sPc + 2 sh; 42 ° 15.780 ′ N, 009 ° 10.500 ′ W to 42 ° 17.100 ′ N, 009 ° 10.630 ′ W; 149 – 150 m; 17 – 28 SeP. 2004; VERTIDOS CH-EBS- 150 • 20 sh; 43 ° 58.84 ′ N, 008 ° 15.625 ′ W to 43 ° 59.621 ′ N, 008 ° 14.285 ′ W; 233 – 237 m; 15 – 24 Jul. 2008; A SELVA EBS- 20.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474873FFEDA0B1FEFB43E506D0.taxon	discussion	Remarks. This species is common in the outer part of the continental shelf off Europe from Iceland and Norway to Morocco (absent from the Baltic and eastern shores of North Sea) and the Mediterranean Sea (Bouchet & Warén 1993; Gofas & Oliver 2011). Shells with soft parts were found in samples from 147 to 212 m and were specially abundant in the sample AG-EBS- 150 collected at 150 m in a muddy sand bottom. These shells were very small, just over 1 mm, and globose (Figs. 18 a, d). Sculpture of closely spaced spiral striations crossed by growth lines (Fig. 18 c). Flattened protoconch (Figs. 18 b, e). High magnification reveals minute spirals in the embryonic whorl (Figs. 18 f – g).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474870FFEFA0B1F8664571033F.taxon	description	(Fig. 19)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474870FFEFA0B1F8664571033F.taxon	materials_examined	Material examined. (more than 2000 shells, most of them with soft parts, in 25 samples): SPAIN • 1 sPc; 43 ° 57.030 ′ N, 008 ° 54.795 ′ W to 43 ° 57.248 ′ N, 008 ° 54.133 ′ W; 1191 – 1132 m; 08 – 15 SeP. 2002; DIVA-Artabria I AT- 1000 • 250 spc + 100 sh; 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 150 • 10 sPc + 7 sh; 43 ° 40.192 ′ N, 008 ° 43.760 ′ W to 43 ° 40.943 ′ N, 008 ° 42.366 ′ W; 207 – 212 m; 08 – 15 Sep. 2002; DIVA-Artabria I EBS- 200 • 4 sPc + 2 sh; 43 ° 41.113 ′ N, 008 ° 44.297 ′ W to 43 ° 41.905 ′ N, 008 ° 43.078 ′ W; 256 – 258 m; 08 – 15 Sep. 2002; DIVA-Artabria I EBS- 250 • 5 sPc + 1 sh; 43 ° 41.689 ′ N, 008 ° 45.195 ′ W to 43 ° 42.556 ′ N, 008 ° 44.226 ′ W; 298 – 303 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 300 • 2 sPc; 43 ° 42.348 ′ N, 008 ° 45.889 ′ W to 43 ° 43.269 ′ N, 008 ° 45.289 ′ W; 344 – 354 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 350 • 1 sPc; 43 ° 45.892 ′ N, 008 ° 44.301 ′ W to 43 ° 46.966 ′ N, 008 ° 43.766 ′ W; 390 – 381 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 400 •> 300 spc + 100 sh; 43 ° 26.703 ′ N, 008 ° 30.669 ′ W to 43 ° 27.452 ′ N, 008 ° 29.612 ′ W; 103 – 102 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 100 •> 200 sPc + 100 sh; 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 Sep. 2002; DIVA-Artabria I EBS- 150 • 15 sPc + 2 sh; 43 ° 40.250 ′ N, 008 ° 43.755 ′ W to 43 ° 40.760 ′ N, 008 ° 42.120 ′ W; 207 – 197 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 200 • 9 sPc + 4 sh; 43 ° 41.590 ′ N, 008 ° 45.328 ′ W to 43 ° 42.396 ′ N, 008 ° 44.286 ′ W; 301 – 301 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 300 • 2 sPc; 43 ° 42.348 ′ N, 008 ° 45.889 ′ W to 43 ° 43.269 ′ N, 008 ° 45.289 ′ W; 344 – 354 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 350 • 3 sh; 43 ° 48.587 ′ N, 008 ° 51.402 ′ W to 43 ° 49.545 ′ N, 008 ° 51.197 ′ W; 610 – 598 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 600 • 15 sh; 43 ° 29.412 ′ N, 008 ° 28.362 ′ W to 43 ° 29.779 ′ N, 008 ° 28.240 ′ W; 110 – 111 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 110 • 4 sPc; 43 ° 32.026 ′ N, 008 ° 37.520 ′ W to 43 ° 32.806 ′ N, 008 ° 35.993 ′ W; 150 – 151 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 150 • 25 sPc + 10 sh; 43 ° 31.970 ′ N, 008 ° 37.670 ′ W to 43 ° 32.697 ′ N, 008 ° 3.171 ′ W; 151 – 151 m; 17 – 28 SeP. 2004; VERTIDOS GA-EBS- 150 • 5 sPc + 4 sh; 43 ° 36.222 ′ N, 008 ° 52.845 ′ W to 43 ° 36.797 ′ N, 008 ° 52.112 ′ W; 201 – 201 m; 17 – 28 Sep. 2004; VERTIDOS GA-EBS- 200 • 2 spc; 43 ° 35.611 ′ N, 008 ° 58.536 ′ W to 43 ° 36.104 ′ N, 008 ° 57.284 ′ W; 384 – 345 m; 17 – 28 Sep. 2004; VERTIDOS GA-EBS- 400 •> 100 sPc; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 150 • 80 sPc; 42 ° 31.176 ′ N, 009 ° 23.380 ′ W to 42 ° 32.132 ′ N, 009 ° 24.151 ′ W; 242 – 248 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 250 •> 600 sPc; 42 ° 50.507 ′ N, 009 ° 25.773 ′ W to 42 ° 48.810 ′ N, 009 ° 25.198 ′ W; 151 – 148 m; 17 – 28 SeP. 2004; VERTIDOS CA-EBS- 150 •> 100 spc; 42 ° 15.780 ′ N, 009 ° 10.500 ′ W to 42 ° 17.100 ′ N, 009 ° 10.630 ′ W; 149 – 150 m; 17 – 28 SeP. 2004; VERTIDOS CH-EBS- 150 • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 1 sh; 43 ° 31.727 ′ N, 008 ° 52.374 ′ W to 43 ° 32.678 ′ N, 008 ° 52.687 ′ W; 173 – 179 m; 15 – 24 Jul. 2008; A SELVA DRN- 23 • 53 spc; 43 ° 31.687 ′ N, 008 ° 52.188 ′ W to 43 ° 33.149 ′ N, 008 ° 53.142 ′ W; 171 – 182 m; 15 – 24 Jul. 2008; A SELVA EBS- 23 • 4 spc; 43 ° 48.252 ′ N, 008 ° 51.427 ′ W to 43 ° 49.707 ′ N, 008 ° 51.164 ′ W; 575 – 584 m; 15 - 24 Jul. 2008; A SELVA EBS- 30 - 1.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D379474870FFEFA0B1F8664571033F.taxon	discussion	Remarks. Obtusella macilenta is known from the continental shelf and slope from the southern part of the Biscay Bay to off Morocco and the Mediterranean (Gofas & Oliver 2011). It was the most abundant species in studied samples, along with A. testae, predominantly between 100 and 250 m on muddy sand bottoms, although one shell with soft parts was found in a sample collected at 1132 m in depth. It is even smaller than the previous species; the shells studied barely reach 1 mm and are less globose with weakest spiral sculpture (Figs. 19 a – b). Sinuous protoconch / teleoconch transition (Figs. 18 c – e) and embryonic shell with up to four delicate spires (Figs. 18 f – g).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484EFFD0A0B1FDC5456B0151.taxon	materials_examined	Type species: Rissoa philippi Aradas and Maggiore, 1844, by monotypy (renamed for Rissoa pusilla Philippi, 1836, preoccupied by Rissoa pusilla Grateloup, 1828).	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484EFFD1A0B1FC364550027E.taxon	description	(Fig. 20)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484EFFD1A0B1FC364550027E.taxon	materials_examined	Material examined. (12 empty shells and 23 with soft parts in 8 samples): SPAIN • 1 sPc; 43 ° 52.823 ′ N, 008 ° 56.151 ′ W to 43 ° 52.837 ′ N, 008 ° 55.597 ′ W; 988 – 920 m; 08 – 15 SeP. 2002; DIVA-Artabria I DRN- 1000 • 5 spc (1 juv); 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 150 • 1 sh; 43 ° 26.703 ′ N, 008 ° 30.669 ′ W to 43 ° 27.452 ′ N, 008 ° 29.612 ′ W; 103 – 102 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 100 • 1 sh; 43 ° 40.250 ′ N, 008 ° 43.755 ′ W to 43 ° 40.760 ′ N, 008 ° 42.120 ′ W; 207 – 197 m; 10 – 20 Sep. 2003; DIVA-Artabria I EBS- 200 • 5 sPc; 43 ° 29.412 ′ N, 008 ° 28.362 ′ W to 43 ° 29.779 ′ N, 008 ° 28.240 ′ W; 110 – 111 m; 17 – 28 Sep. 2004; VERTIDOS GA-AT- 110 • 10 sPc + 10 sh; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 150 • 1 sPc: 42 ° 31.176 ′ N, 009 ° 23.380 ′ W to 42 ° 32.132 ′ N, 009 ° 24.151 ′ W; 242 – 248 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 250 • 1 sPc; 42 ° 50.507 ′ N, 009 ° 25.773 ′ W to 42 ° 48.810 ′ N, 009 ° 25.198 ′ W; 151 – 148 m; 17 – 28 SeP. 2004; VERTIDOS CA-EBS- 150.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484EFFD1A0B1FC364550027E.taxon	discussion	Remarks. A species-complex can hide under the name Pusillina inconspicua (Gofas & Oliver 2011) due its variability, its discontinuous geographic distribution along the eastern Atlantic from Norway to Angola, including the Mediterranean, and due the different habitats where the species supposedly lives, from sublittoral to bathyal bottoms below 100 m depth, and even in brackish lagoon areas (Gofas & Oliver 2011). Cordeiro & Ávila (2015) also recorded P. inconspicua in the Azores. The shells found in the samples studied exhibited a remarkable variability ranging from weak axial ribs, thickened lip, curved columella and closed umbilicus (Fig. 20 a), to almost smooth surface with incised peripheral lines, thin lip, straightened columella and narrow umbilicus (Fig. 20 b). The protoconch bears a characteristic spiral microcord close to the suture (Figs. 20 c, g). Transition between protoconch I and II well delimited (Figs. 20 d – f). Shells with soft parts have been found in samples collected between 111 – 248 m depth on muddy sand bottoms, and one specimen at 920 m. It should be noted that some micromolluscs living in coastal algae could be swept by the currents away from the coast.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484FFFD1A0B1FB4045760733.taxon	description	(Fig. 21 d)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484FFFD1A0B1FB4045760733.taxon	materials_examined	Material examined. (3 juvenile shells in the same sample): SPAIN • 3 sh (juv); 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 150.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484FFFD1A0B1FB4045760733.taxon	discussion	Remarks. This species is known in northeastern Atlantic from the English Channel to Morocco, in sublittoral rocky bottoms (Gofas & Oliver 2011). It is common in Azores, Madeira and Canary Islands and it has been only recorded in the seamounts whose summit is shallower (Gorringe and Ampere) (Gofas 1990, 2007; Ávila & Malaquias 2003; Cordeiro & Ávila 2015). Only three juvenile shells have been found (from a single sample obtained at a depth of 150 m), one of them very eroded.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484FFFD1A0B1FF84459F0583.taxon	description	(Fig. 21 a – c)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484FFFD1A0B1FF84459F0583.taxon	materials_examined	Material examined. (23 empty shells and 2 with soft parts, in 8 samples): SPAIN • 1 sPc (juv); 43 ° 35.451 ′ N, 008 ° 34.432 ′ W to 43 ° 34.810 ′ N, 008 ° 35.407 ′ W; 153 – 151 m; 08 – 15 SeP. 2002; DIVA-Artabria I EBS- 150 • 2 sh; 43 ° 48.421 ′ N, 008 ° 51.453 ′ W to 43 ° 49.160 ′ N, 008 ° 51.091 ′ W; 599 – 607 m; 10 – 20 SeP. 2003; DIVA-Artabria I DRN- 600 • 1 sPc (juv); 43 ° 34.127 ′ N, 008 ° 36.562 ′ W to 43 ° 34.820 ′ N, 008 ° 35.585 ′ W; 152 – 149 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 150 • 6 sh; 43 ° 38.812 ′ N, 009 ° 07.949 ′ W to 43 ° 39.841 ′ N, 009 ° 07.405 ′ W; 999 – 1001 m; 17 – 28 Sep. 2004; VERTIDOS GA-DRN- 1000 • 6 frg; 43 ° 38.105 ′ N, 009 ° 02.006 ′ W to 43 ° 37.738 ′ N, 009 ° 00.905 ′ W; 836 – 826 m; 17 – 28 SeP. 2004; VERTIDOS GA-AT- 800 • 2 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 480 – 600 m; SARRIDAL (2006 – 2007) SARRI- 2 • 6 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 • 1 sh; 44 ° 08.461 ′ N, 009 ° 04.634 ′ W to 44 ° 08.607 ′ N, 009 ° 04.413 ′ W; 917 – 896 m; 15 – 24 Jul. 2008; A SELVA DRN- 7 - 1.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484FFFD1A0B1FF84459F0583.taxon	discussion	Remarks. It is a Mediterranean species with isolated records in the Atlantic coasts (Verduin 1976) usually in seagrass meadows (Gofas & Oliver 2011). Verduin (1976) pointed out the extreme variability of this species (with some local forms), whose precise delimitation is not clear. A thorough review of most species of genus Pusillina, including genetic analysis, is necessary to elucidate their taxonomic status. The presence of some juvenile shells in our samples, attributed with doubts to P. radiata (Figs. 21 a – b) is considered accidental; probably washed away by currents. Only two shells with soft parts were found in samples obtained at 150 m. The protoconch is shown in Figure 21 c.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484CFFD2A0B1FDC544110732.taxon	description	(Fig. 21 e)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484CFFD2A0B1FDC544110732.taxon	materials_examined	Material examined. (2 empty shells and 2 with soft parts, in 3 samples): SPAIN • 1 sPc (juv); 43 ° 40.250 ′ N, 008 ° 43.755 ′ W to 43 ° 40.760 ′ N, 008 ° 42.120 ′ W; 207 – 197 m; 10 – 20 SeP. 2003; DIVA-Artabria I EBS- 200 • 1 sPc (juv) + 1 sh; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 150 • 1 sh; 44 ° 00.806 ′ N, 008 ° 33.098 ′ W to 44 ° 01.967 ′ N, 008 ° 32.57 ′ W; 346 – 344 m; 15 – 24 Jul. 2008; A SELVA EBS- 14.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484CFFD2A0B1FDC544110732.taxon	discussion	Remarks. It is a very common sublittoral species, mainly in shallow waters, along all European and North African coasts from Norway to Morocco and throughout the Mediterranean, not present on the Macaronesian islands (Oliver et al. 2019). Very few eroded shells (Fig. 21 e) have been found in samples obtained at the depth range of 150 – 345 m, likely swept by the currents away from the coast.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484DFFD3A0B1FFDD42D70389.taxon	description	(Fig. 21 g)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484DFFD3A0B1FFDD42D70389.taxon	materials_examined	Material examined. (3 empty shells, in 2 samples): SPAIN • 2 sh; 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 150 • 1 sh; 44 ° 11.652 ′ N, 008 ° 58.152 ′ W to 44 ° 11.539 ′ N, 008 ° 57.574 ′ W; 908 – 1106 m; 15 – 24 Jul. 2008; A SELVA DRN- 7.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484DFFD3A0B1FFDD42D70389.taxon	discussion	Remarks. As in the previous species, Manzonia crassa is very common in sublittoral bottoms from the British Islands to Morocco (not present on the Macaronesian islands) and throughout the Mediterranean (Gofas & Oliver, 2011). Three eroded shells (Fig. 21 g) have been found in two of the samples likely swept by the currents away from the coast.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484DFFD3A0B1FDC543EA027E.taxon	description	(Fig. 21 f)	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484DFFD3A0B1FDC543EA027E.taxon	materials_examined	Material examined. (2 empty shells in 2 samples): SPAIN • 1 sh 42 ° 30.391 ′ N, 009 ° 19.517 ′ W to 42 ° 29.428 ′ N, 009 ° 17.924 ′ W; 147 – 148 m; 17 – 28 SeP. 2004; VERTIDOS AG-EBS- 150 • 1 sh; Polygon delimited by Points 44 ° 10.000 ′ N, 009 ° 00.000 ′ W / 44 ° 10.000 ′ N, 008 ° 35.000 ′ W / 44 ° 00.000 ′ N; 009 ° 00.000 ′ W / 44 ° 07.000 ′ N; 008 ° 35.000 ′ W; 417 – 668 m; SARRIDAL (2006 – 07) SARRI- 3.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
03D37947484DFFD3A0B1FDC543EA027E.taxon	discussion	Remarks. It is a common sublittoral NE Atlantic, usually from the intertidal zone to about 100 m depth, and rarely deeper (Fretter & Graham 1978; Nekhaev et al. 2014). Bouchet & Warén (1993) did not include it among the NE Atlantic bathyal species. Only two empty shells and one fragment were found from two samples. One of the shells from 417 – 668 m was well preserved (Fig. 21 f), which is surprising since O. semicostata is a shallow-water species.	en	Oliver, Joan Daniel, Gofas, Serge, Urgorri, Victoriano, Díaz-Agras, Guillermo, Templado, José (2022): Rissoidae gastropods of the outer continental shelf and slope off Galicia (NW Spain). Zootaxa 5196 (1): 1-45, DOI: https://doi.org/10.11646/zootaxa.5196.1.1
