taxonID	type	description	language	source
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	description	urn: lsid: zoobank. org: act: F 159660 E- 5198 - 42 B 0 - 813 E- 70 AF 8770 C 1 D 0 Figs 1 – 12	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	diagnosis	Diagnosis Andrena deminuta sp. nov. can be placed in the subgenus Aciandrena Warncke, 1968 because of its small body size (Fig. 1), black integument, shagreened propodeal triangle (Fig. 4), simple male genitalia (Fig. 11), and yellow male clypeus (Fig. 8). It is closest to A. israelica Scheuchl & Pisanty, 2016 and A. judaea Scheuchl & Pisanty, 2016 because the facial foveae are relatively broad above and not substantially tapered below, in addition to their small body size of ca 5 mm, which makes them one of the smallest known members of the Aciandrena. Female material can be separated from A. judaea (alternative character state in parentheses) because in dorsal view the foveae are filled with brown hairs (vs whitish hairs) and the nervulus is strongly antefurcal (vs weakly antefurcal). It shares these characters with A. israelica and is therefore extremely similar. It can be separated by the stronger punctures on the clypeus (Fig. 2), punctures apically and laterally clearly present, separated by 0.5 – 1 puncture diameter (clypeal punctures fine, scattered, apically and laterally separated by 1 – 2 puncture diameters, not obviously contrasting underlying surface). Male material has a yellow clypeus (Fig. 8) and a strongly antefurcal nervulus, and therefore can be separated from both A. judaea (yellow clypeus, but weakly antefurcal nervulus) and A. israelica (black clypeus, strongly antefurcal nervulus). The genital capsule also differs from that of both taxa as it has pronounced and apically truncate gonocoxal teeth (Fig. 11), the apices therefore square-shaped (gonocoxal teeth weakly produced and apically rounded in A. judaea, gonocoxal teeth more produced but with rounded apices in A. israelica), the penis valves are basally broad and sharply narrow apically (penis valves comparatively narrower basally, narrowing less abruptly apically in A. israelica and A. judaea), and the outer margin of the gonostyli are more strongly emarginate (outer margin straight to very weakly emarginate, see illustrations in Pisanty et al. 2016).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	etymology	Etymology From the Latin adjective ‘ deminuta ’, meaning ‘ small, diminutive’, in reference to its small size.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	materials_examined	Material examined Holotype IRAN • ♂; Lorestan Province, Dorud Lanjaban env.; 33.419 ° N, 48.986 ° E; 960 m a. s. l.; [collected in May 2016 but no precise date on label]; M. Kafka leg.; OÖLM. Paratypes IRAN • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; M. Kafka leg.; OÖLM • 4 ♀♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM • 1 ♀; same collection data as for preceding; TJWC • 1 ♀; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM • 2 ♀♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 1 ♀; same collection data as for preceding; TJWC. Other material examined (aff. Andrena deminuta) IRAN • 1 ♂, 10 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 3 ♀♀; same collection data as for preceding; TJWC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	description	Description Female MEASUREMENTS. Body length 5 – 6 mm (Fig. 1). HEAD. 1.2 times as wide as long (Fig. 2). Clypeus dark, domed, with irregular large punctures, punctures separated by 0.5 – 3 puncture diameters, broad but unclear longitudinal impunctate midline present centrally; underlying surface uniformly shagreened, weakly shining. Process of labrum variable, generally narrowly trapezoidal, as broad as long or narrower than long, apical margin truncate. Face, gena, vertex, and scape with short whitish hairs. Gena equalling width of compound eye; ocelloccipital distance slightly less than diameter of lateral ocellus. Foveae dorsally broad, occupying half distance between compound eye and lateral ocellus, narrowed to half width below at level of antennal insertions, here separated from inner margin of compound eye by distance subequal to their own width; foveae filled with dark brown hairs (Fig. 3). Antennae basally dark, A (5) 6 – 12 strongly lightened orange below; A 3 subequal to A 4 + 5. MESOSOMA. Scutum and scutellum with very fine granular shagreen, weakly shining, sparsely punctate with fine punctures, punctures separated by 1 – 5 puncture diameters. Pronotum without humeral angle. Mesepisternum and lateral and dorsolateral faces of propodeum with fine granular shagreen, weakly shining. Propodeal triangle broad, internal surface with extremely fine raised network of circular carinae, thus resembling granular shagreenation, propodeal triangle therefore defined by change in surface sculpture (Fig. 4). Mesosoma with short sparse whitish hairs, propodeal corbiculae incomplete, dorsally composed of weakly plumose whitish hairs, internal surface with sparse whitish simple hairs. Legs dark, apical tarsal segments lightened orange, pubescence whitish. Tibial and femoral scopae and flocculus whitish. Hind tarsal claws unidentate. Wings hyaline, stigma orange with dark brown lateral margin, venation dark orange; nervulus strongly antefurcal, first recurrent vein enters second submarginal cell before its middle. METASOMA. Terga dark, marginal areas of T 2 – 4 slightly depressed, T 1 – 4 with apical margin lightened dark brown to yellow apically (Fig. 5). Terga finely microreticulate, strongest on T 1, becoming progressively weaker by T 4, weakly shining; tergal discs essentially impunctate (Fig. 6). Tergal discs with extremely scattered short whitish hairs, apical fringe of T 5 and hairs flanking pygidial plate golden. Pygidial plate narrowly rounded triangular, flat, internal surface weakly shagreened. Male MEASUREMENTS. Body length 5.5 mm (Fig. 7). HEAD. 1.3 times as wide as long. Clypeus yellow marked over majority of area, not extending fully to margins, laterally with two small black maculae (Fig. 8). Clypeus with irregular punctures, punctures separated by 0.5 – 3 puncture diameters, underlying surface uniformly shagreened, weakly shining. Process of labrum narrowly triangular, apical margin truncate. Face, vertex, and scape with whitish hairs, gena ventrally with long white hairs, equalling length of scape. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly less than diameter of lateral ocellus. Antennae dark basally, A 5 – 13 lightened orange below; A 3 exceeding A 4, this sub-square, shorter than A 4 + 5. MESOSOMA. Mesosoma structurally as in female (Fig. 9) with exception of hind tarsal claws with strong inner tooth, wing venation dark brown. METASOMA. Terga structurally as in female (Fig. 10). S 8 columnar, parallel-sided, apical margin truncate, ventral surface covered with short whitish hairs. Genital capsule compact, gonocoxae apically produced into strong square-ended teeth. Gonostyli broad, with strongly raised internal margin, apically flattened, slightly narrowed subapically. Penis valves broad basally, narrowing strongly at their midpoint (Fig. 11).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	discussion	Remarks Several specimens were examined from Fars Province (see below). Females were not separable from A. deminuta sp. nov., as the only apparent difference is that the tergal shagreenation is stronger, but the single male specimen had a divergent genital capsule. In the holotype male of A. deminuta, the gonocoxal teeth are elongate with clearly truncate and square-shaped apices (Fig. 11). However, in the specimen from Fars Province, the gonocoxal teeth are less strongly elongate, and apically more rounded (Fig. 12). Given the presence of cryptic Aciandrena species which cannot be morphologically separated in the female sex (Pisanty et al. 2022 a), it is unlikely that these belong to the same taxon. The material from Fars is not described until more material is available.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF97512ED9B0DD69FD5FFBF8.taxon	distribution	Distribution Southern Iran (provinces of Ilam and Lorestan).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	description	urn: lsid: zoobank. org: act: 33 AD 35 D 5 - FBE 2 - 4 BDC- 9 C 7 C- 0 B 17 C 62 EB 284 Figs 13 – 24	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	diagnosis	Diagnosis Andrena boustaniae sp. nov. can be placed in the subgenus Euandrena Hedicke, 1933 in the female sex because of the characteristically drop-shaped foveae which are narrowed below, and the simple scopal hairs. This subgenus is poorly understood in the Eastern Mediterranean to the Middle East (Praz et al. 2019; TJW, G. Pisanty & C. Praz unpublished data) and extreme care should be taken when identifying material in this region. Female material is superficially similar to taxa around A. allosa Warncke, 1975 (see Praz et al. 2019) due to the light scutal hairs (Fig. 13) and intermixed black and white hairs on the face (Fig. 15). However, females of A. boustaniae sp. nov. present several important characters, specifically the flattened clypeus with a slightly raised impunctate longitudinal ridge (Fig. 16), the broad, apically projecting, and medially emarginate process of the labrum (Fig. 18), the finely rugose propodeal triangle, the dorsally wide facial fovea (occupying over half the distance between the compound eye and the lateral ocellus), the dark hind tibiae with bicoloured tibial scopa (Fig. 13, white ventrally, black dorsally), the densely and clearly punctate scutum, the apically hyaline tergal margins (Fig. 20), and the clear white hair bands on T 2 – 4. In combination, these characters allow separation from all described Turkish, Caucasian, and Central Asian taxa of Euandrena. In the male sex, A. boustaniae sp. nov. has a unique genital capsule for Euandrena in this region, lacking gonocoxal teeth, the inner margins with an almost obtuse angle, thus superficially resembling the genitalia of certain species of A. (Graecandrena), gonocoxae slightly diverging medially, and the penis valves extremely broad at the base (Fig. 24). In the diverging gonocoxae, the genital capsule slightly resembles A. hermonella Scheuchl & Pisanty, 2016, but the lateral hyaline extensions of the penis valves are rounded, not forming angular projections.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	etymology	Etymology Named after Mira Boustani for her tireless work researching and cataloguing the understudied Lebanese bee fauna (Boustani et al. 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	materials_examined	Material examined Holotype IRAN • ♂; Yazd, Banadak-o sadat; 31.573 ° N, 54.204 ° E; 2103 m a. s. l.; 22 Mar. 2021; S. San leg.; OÖLM. Paratypes IRAN • 1 ♀; Yazd, Mehriz, Posht e Hosseinie Shohaday gomnam; 1500 m a. s. l.; 28 Feb. 2020; S. San leg.; OÖLM • 1 ♀; Yazd, Sakhvid, Mazraeh Khosraw; 2469 m a. s. l.; 27 Mar. 2019; S. San leg.; ICPI • 5 ♂♂; Yazd, Banadak-o sadat; 2103 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 1 ♂; same collection data as for preceding; TJWC • 1 ♂; same collection data as for preceding; OÖLM • 1 ♂; Yazd, Tezerjan, paen tar az Seyyed Mahmood; 1993 m a. s. l.; 6 Mar. 2020; S. San leg.; OÖLM • 1 ♂; Yazd, Jade konj-e kooh, dakal; 2019 m a. s. l.; 26 Mar. 2021; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	description	Description Female MEASUREMENTS. Body length 11 mm (Fig. 13). HEAD. 1.2 times as wide as long (Fig. 15). Clypeus dark, centrally flattened, irregularly punctate with unevenly sized punctures; punctures basally separated by <0.5 puncture diameter, becoming sparse apically, separated by 1 – 4 puncture diameters; apical rim of clypeus essentially impunctate; clypeus centrally with impunctate, longitudinal, and slightly raised ridge; underlying clypeal surface smooth and shining (Fig. 16). Process of labrum triangular-trapezoidal, strongly narrowing apically, with distinct, strongly bifurcate medioapical extension (Figs 17 – 18). Face centrally with white hairs on clypeus, supraclypeal area, scape, and frons, intermixing with black hairs laterally, dorsally, and ventrally. Gena ventrally with long white hairs, longest exceeding length of scape, intermixing with black hairs dorsally, hairs on vertex predominantly white. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly exceeding diameter of lateral ocellus. Foveae dorsally broad, occupying slightly over ½ distance between compound eye and lateral ocellus, narrowed to half width below at level of antennal insertions; foveae filled with black hairs. Antennae dark, A 4 – 12 slightly lightened below due to presence of grey cilia; A 3 exceeding A 4 + 5, A 4 sub-square, A 5 longer than broad, A 3 shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum with very fine and even shagreenation, shining; surface densely but slightly irregularly punctate with clear punctures, punctures separated by <0.5 – 1 puncture diameter (Fig. 14). Pronotum without humeral angle. Mesepisternum and lateral face of propodeum with fine shagreenation, weakly shining, surface covered with large flat punctures, becoming sparse centrally on face of propodeal corbicula. Propodeal triangle wide, well delineated laterally by fine raised carina, internal surface with raised longitudinal carina centrally, remaining surface with fine network of raised rugae, underlying surface more clearly shining than dorsolateral parts of propodeum (Fig. 19). Mesepisternum and majority of propodeum with long black hairs, propodeal corbicula complete, corbicular fringe dorsally composed of long black plumose hairs, becoming entirely white ventrally; internal surface of propodeal corbicula with simple white hairs. Scutum and scutellum dorsally with predominantly white hairs, some shorter black hairs intermixed laterally and centrally. Legs uniformly dark, pubescence white, fore femorae ventrally with long fringe of white plumose hairs, equalling length of scape. Tibial scopae bicoloured, white ventrally, blackish dorsally, composed of simple hairs; flocculus and femoral scopae unicolourous white. Hind tarsal claws with inner tooth. Wings hyaline, stigma orange centrally, dark brown laterally, venation dark brown; nervulus interstitial, first recurrent vein enters second submarginal cell beyond its middle. METASOMA. Terga dark, marginal areas of T 2 – 4 slightly depressed, T 1 – 4 with apical margin narrowly lightened yellow-hyaline (Fig. 20). Terga finely microreticulate, weakly shining, tergal discs deeply and densely punctate, punctures separated by 1 – 2 puncture diameters. T 1 – 2 on discs with loose long whitish hairs, laterally with additional scattered black hairs, discs of T 3 – 4 with short intermixed black and white hairs; T 2 – 4 apically with dense fringe of short white hairs, obscuring underlying surface. Apical fringe of T 5 black centrally, with long white hairs laterally, hairs flanking pygidial plate black; pygidial plate triangular, broadly rounded apically, with large and flat triangular raised area centrally. Male MEASUREMENTS. Body length 9 – 11 mm (Fig. 21). HEAD. 1.2 times as wide as long. Clypeus dark, very weakly domed, densely and evenly punctate, punctures separated by 0.5 – 1 puncture diameter; underlying surface smooth and shining (Fig. 22). Process of labrum as in female, though longitudinally compressed, with lateral wrinkles and ridges. Face more extensively dark haired than female, though still with white hairs on face centrally, on gena ventrally, and on vertex, longest hairs clearly exceeding length of scape. Gena broad, almost twice width of compound eye; ocelloccipital distance 1 ½ times diameter of lateral ocellus. Antennae dark, A 4 – 13 slightly lightened greyish-brown below due to presence of tiny cilia; A 3 slightly shorter than A 4. MESOSOMA. Scutum and scutellum more extensively shagreened than in female, dull except for small area on scutum centrally with reduced shagreenation, weakly shining. Mesosoma otherwise structurally as in female. Mesepisternum and propodeum with long, uniformly black hairs, without white hairs laterally, some scattered white hairs ventrally; scutum and scutellum with sparse long and intermixed black and white hairs, white hairs dominating. Legs dark, pubescence intermixed black and white, white hairs dominating on tibiae and tarsi. Hind tarsal claws with inner tooth. Wings hyaline, stigma orange centrally, dark brown laterally, venation dark brown; nervulus slightly antefurcal, first recurrent vein enters second submarginal cell beyond its middle. METASOMA. Terga dark, apical margins of T 2 – 5 slightly depressed, more extensively lightened hyalineyellow (Fig. 23). Tergal discs finely shagreened over majority of area, weakly shining, shagreen absent apically and on marginal areas, here smooth and shining; tergal discs evenly punctate, punctures separated by 2 puncture diameters. Discs of T 1 – 2 with sparse, long white hairs, becoming shorter and intermixed with black hairs on T 3 – 4, disc of T 5 with only short black hairs. S 8 columnar, apically truncate, laterally with short brown hairs. Genital capsule compact, gonocoxa with inner margin slightly diverging medially, inner angle rounded, slightly obtuse, without gonocoxal teeth. Gonostyli with apical blades spatulate, slightly flattened. Penis valves broad basally with large, rounded hyaline extensions laterally, apically narrowed (Fig. 24).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	discussion	Remarks Collected from flowers of Prunus (Amygdalus) spp. (Rosaceae) in February and March. It may therefore be associated with flowering trees in the early spring.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF935122D9BBDD9DFE17FAB1.taxon	distribution	Distribution Central Iran (Yazd).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	description	urn: lsid: zoobank. org: act: AC 3 B 3 A 80 - B 5 BC- 49 F 2 - 8256 - 37 BDEA 1 EF 690 Figs 25 – 30	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	diagnosis	Diagnosis Andrena oblata sp. nov. can be placed in the subgenus Euandrena in the female sex because of the characteristically drop-shaped foveae which are narrowed below, and the simple scopal hairs (Fig. 28). The centrally flattened clypeus places it close to two taxa, A. canuta Warncke, 1975 from Turkey, Armenia, and Azerbaijan and A. alijevi Osytshnjuk, 1986 from Azerbaijan. Andrena oblata can be separated from A. canuta because the clypeus is completely flat centrally (Fig. 26) (centrally with a longitudinal excavated groove), the clypeus itself is compact, not noticeably elongated (weakly elongated), and the tibial scopa is golden (tibial scopa white). Andrena oblata can be separated from A. alijevi by the facial fovea which dorsally occupy ⅓ of distance between lateral ocellus and compound eye (dorsally occupying ½ this distance), the uniformly golden hairs of the vertex, paraocular area, and scutum (hairs here greyish-brown with admixture of black hairs), and the golden tibial scopa (tibial scopa whitish with yellow-brown hairs at base). See also A. sani sp. nov. (below).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	etymology	Etymology From the Latin adjective ‘ oblatus ’, meaning ‘ flattened’, in reference to the flattened clypeus.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	materials_examined	Material examined Holotype IRAN • ♀; Yasouj, Darreh Grouh Firuz Abad, Faramarz Village; 30.945 ° N, 51.596 ° E; 6 May 2021; E. Rostami leg.; OÖLM. Paratypes IRAN • 2 ♀♀; Yasouj, Darreh Grouh Firuz Abad, Faramarz Village; 30.945 ° N, 51.596 ° E; 6 May 2021; E. Rostami leg.; OÖLM • 2 ♀♀; same collection data as for preceding; TJWC • 5 ♀♀; same collection data as for preceding; ICPI • 2 ♀♀; Yasouj, 2 km after Davood Abad Village; 1811 m a. s. l.; 6 May 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	description	Description Female MEASUREMENTS. Body length 9.5 – 11 mm (Fig. 25). HEAD. 1.2 times as wide as long. Clypeus dark, centrally flattened, with clear and variable punctures; punctures dense laterally, separated by 0.5 puncture diameter, becoming sparse and irregular centrally, separated by 1 – 4 puncture diameters (Fig. 26). Surface of clypeus laterally with weak shagreenation, smooth and shining over majority of area. Process of labrum trapezoidal with rounded corners, twice as broad as long. Face, scape, gena, and vertex with uniformly golden hairs, longest approximately half of length of scape. Gena slightly exceeding width of compound eye; ocelloccipital distance equalling diameter of lateral ocellus. Fovea narrow dorsally, occupying ⅓ of distance between lateral ocellus and compound eye, slightly narrowed below at level of antennal insertions; filled with dark brown hairs. Antennae dark, A 3 slightly exceeding A 4 + 5, A 4 sub-square, slightly wider than long, A 4 as long as broad. MESOSOMA. Scutum and scutellum with very fine and irregular shagreenation, surface shining to weakly shining; surface densely and clearly punctate, punctures separated by 0.5 – 1 puncture diameter (Fig. 27). Pronotum without humeral angle. Mesepisternum finely rugose, dull, lateral faces of propodeum with shagreened, weakly shining. Propodeal triangle poorly delineated with very fine and inconspicuous lateral carinae, internal surface with fine network of slightly raised rugae. Dorsolateral faces of propodeum strongly shagreened and densely punctate with large shallow punctures, punctures separated by 0.5 – 1 puncture diameter, propodeal triangle therefore distinguished by absence of punctures. Mesosoma with golden hairs, becoming lighter ventrally, longest on mesepisternum, not exceeding length of scape; propodeal corbicula with plumose hairs comprising dorsal fringe and weakly plumose hairs on internal surface. Legs uniformly dark, pubescence brownish basally to golden apically; tibial scopae brownishgolden, femoral scopae and flocculus lighter (Fig. 28). Hind tarsal claws with inner tooth. Wings hyaline, stigma orange, venation basally brown, becoming orange over majority of wing; nervulus weakly antefurcal to antefurcal, first recurrent vein enters second submarginal cell slightly beyond its middle. METASOMA. Terga dark, marginal areas of T 2 – 4 depressed, T 1 – 4 with apical margins narrowly lightened yellow-hyaline (Fig. 29). Terga faintly shagreened, weakly shining, clearly and deeply punctate, punctures separated by 1 – 2 puncture diameters, marginal areas weakly and obscurely punctate (Fig. 30). T 1 – 4 on discs with sparse golden hairs, on T 1 extending onto margin to form comparatively weak apical band, not obscuring underlying surface, T 2 – 4 with dense complete apical hairbands of long yellowish hairs, obscuring underlying surface. Apical fringe of T 5 and hairs flanking pygidial plate golden; pygidial plate narrowly triangular, with raised rounded longitudinal ridge centrally. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	discussion	Remarks All specimens were collected from Elaeagnus angustifolia L. (Elaeagnaceae) in May, with the majority of individuals having their scopae full of pollen. Further study is required to see if this is an important part of the pollen diet. Use of pollen of Elaeagnus has also been recorded in Andrena (Euandrena) nasica Lebedev, 1933, which is found in Kazakhstan, Turkmenistan, and Uzbekistan (Osytshnjuk et al. 2008). This species can easily be separated by its strongly elongated clypeus, fulvous tarsi and hind tibiae, its narrower foveae, and its much less densely punctate terga. However, the association with Elaeagnus and broadly similar morphology suggests that this plant and its associated fauna of Andrena should be more thoroughly studied in semidesert environments in Iran and Central Asia.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9F5127DA74DC57FE60FDFF.taxon	distribution	Distribution Southern Iran (Yasouj).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9A5138DA08DB9EFE17F85C.taxon	description	urn: lsid: zoobank. org: act: BD 70 A 72 C- 74 D 6 - 4 DE 7 - AF 60 - 57 D 0 A 3 E 9 BA 80 Figs 31 – 40	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9A5138DA08DB9EFE17F85C.taxon	diagnosis	Diagnosis Andrena sani sp. nov. can be placed in the subgenus Euandrena in the female sex because of the characteristically drop-shaped foveae which are narrowed below, and the simple scopal hairs. It is most similar to A. alijevi and to A. oblata sp. nov. because of the centrally flattened clypeus and thick yellow tergal hairbands (Fig. 35). Andrena sani can be separated from A. alijevi because of the uniformly light brown hairs on the vertex, paraocular area, and scutum (Figs 31 – 32) (hairs here greyish-brown with admixture of black hairs) and the slightly impressed longitudinal impunctate line on the clypeus surrounded by dense punctures (Fig. 33) (clypeus without impressed longitudinal line, centrally with scattered punctation). Andrena sani can be separated from A. oblata by the slightly impressed longitudinal impunctate line on the clypeus surrounded by dense punctures (clypeus without impressed longitudinal line, centrally with scattered punctation) and the weaker tergal puncturing, punctures here sparse and obscure (Fig. 36), separated by 2 – 4 puncture diameters (punctures dense, distinct, separated by 1 – 2 puncture diameters; compare Figs 30 and 36). Males are harder to diagnose (as for most Euandrena), but the combination of the characters A 3 = A 4, dark, non-metallic terga, predominantly black facial pubescence, non-elongated clypeus, scutum finely granulate and matt, terga finely shagreened with sparse indistinct punctures (Fig. 39), and simple genital capsule (Fig. 40) places it close to A. alijevi. It can be separated because of the slightly impressed impunctate longitudinal line on the clypeus, the clypeus otherwise entirely densely punctate (clypeus without longitudinal impunctate line, apical ⅓ smooth and shiny with scattered punctures) and the facial pubescence, which is extensively white centrally, black laterally (predominantly black, with occasional pale hairs).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9A5138DA08DB9EFE17F85C.taxon	etymology	Etymology Named after Sina San, who collected many of the new species described in this manuscript.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9A5138DA08DB9EFE17F85C.taxon	materials_examined	Material examined Holotype IRAN • ♀; Yazd, Sanij Baghe khaleghieh; 31.648 ° N, 54.015 ° E; 2060 m a. s. l.; 13 Mar. 2020; S. San leg.; OÖLM. Paratypes IRAN • 1 ♂; Yazd, Sakhvid, Mazraeh Khosraw; 2469 m a. s. l.; 27 Mar. 2019; S. San leg.; TJWC • 1 ♂; Yazd, Tezerjan, Hossein Abad, Seyyed mahmood; 1993 m a. s. l.; 5 Mar. 2020; S. San leg.; OÖLM • 1 ♀; Yazd, Yazd road to Sanij; 2621 m a. s. l.; 27 Mar. 2019; S. San leg.; TJWC • 2 ♀♀; Yazd, Taft, Tezerjan, Mazra ali, Agha seyyed; 2109 m a. s. l.; 24 Mar. 2019; S. San leg.; ICPI • 1 ♀; Yazd, Banadak-o sadat; 2103 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 1 ♂; Yazd, Mehriz, Shohaday gomnam; 1500 m a. s. l.; 8 May 2021; S. San leg.; ICPI • 1 ♂; Yazd, Tezerjan Bagh-e Agha seyyed; 2226 m a. s. l.; 5 May 2021; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9A5138DA08DB9EFE17F85C.taxon	description	Description Female MEASUREMENTS. Body length 9.5 – 10.5 mm (Fig. 31). HEAD. 1.2 times as wide as long (Fig. 32). Clypeus dark, flattened over majority of its area, densely punctate, punctures separated by 0.5 – 1 puncture diameter with exception of clear impunctate longitudinal line centrally with width of 2 puncture diameters; longitudinal line slightly impressed, giving impression of weak furrow (Fig. 33). Clypeus basally and laterally shagreened, weakly shining, becoming smooth and shiny apically. Process of labrum rounded-trapezoidal, twice as broad as long, apical margin very weakly emarginate. Face, gena, scape, and vertex with long pale brown hairs, longest equalling length of scape; face laterally and frons with some intermixed black hairs. Gena slightly exceeding width of compound eye; ocelloccipital distance equalling diameter of lateral ocellus. Fovea narrow dorsally, occupying just under ½ distance between lateral ocellus and compound eye, slightly narrowed below at level of antennal insertions; filled with dark brown hairs. Antennae dark, A 3 exceeds A 4 + 5, shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum shagreened, shagreenation slightly irregular, scutum duller laterally, weakly shining centrally; surface punctate, punctures separated by 1 – 2 puncture diameters (Fig. 34). Pronotum without humeral angle. Mesepisternum and lateral faces of propodeum with fine granular reticulation, weakly shining. Propodeal triangle narrow, weakly delineated laterally with fine carinae, internal surface with fine granular reticulation, dull, basally with slightly raised rugae. Dorsolateral faces of propodeum with weakly shining fine granular reticulation and fine raised rugosity, propodeal triangle therefore distinguished by absence of rugosity. Mesosoma with long light brownish hairs, becoming whitish ventrally, longest on mesepisternum, equalling length of scape; propodeal corbicula with plumose hairs comprising dorsal fringe, internal surface with sparse plumose hairs. Legs uniformly dark, pubescence dark to light brownish; tibial scopae light brownish, femoral scopae and flocculus whitish. Hind tarsal claws with inner tooth. Wings hyaline, stigma dark orange centrally with dark brown margin, venation dark brown to dark orange; nervulus interstitial to slightly antefurcal, first recurrent vein enters second submarginal cell at its midpoint. METASOMA. Terga dark, marginal areas of T 2 – 4 depressed, T 1 – 4 with apical margins widely lightened yellow-hyaline (Fig. 35). Terga strongly shagreened, weakly shining, with faint and obscure fine punctures, punctures separated by 2 – 4 puncture diameters, marginal areas basally punctate (Fig. 36). T 1 – 4 on discs with sparse long golden hairs, on T 1 extending onto margin to form comparatively weak apical band, not obscuring underlying surface, T 2 – 4 with dense complete hairband of long whitish hairs that obscures underlying surface. Apical fringe of T 5 whitish laterally, becoming dark brown centrally, hairs flanking pygidial plate dark brown; pygidial plate rounded triangular, with faint raised rounded longitudinal ridge centrally. Male MEASUREMENTS. Body length 8 – 8.5 mm (Fig. 37). HEAD. 1.2 times as wide as long. Clypeus dark, flattened over majority of its area, densely and uniformly punctate, punctures separated by 0.5 puncture diameter with exception of slightly impressed longitudinal impunctate line, width equal to 1 puncture diameter (Fig. 38). Clypeus surface weakly shining throughout. Process of labrum rounded-trapezoidal, apical margin broadly and weakly emarginate. Face centrally with extensive long white hairs from antennal insertions to apex of clypeus, replaced laterally by entirely black hairs that extend onto frons; longest hairs exceeding length of scape. Gena and vertex with predominantly whitish hairs, with occasional intermixed black hairs. Gena slightly elongate, exceeding width of compound eye; ocelloccipital distance equals diameter of lateral ocellus. Antennae dark, A 3 = A 4. MESOSOMA. Scutum and scutellum with fine granular shagreen, weakly shining, irregularly punctate with large shallow punctures, punctures separated by 1 – 3 puncture diameters. Structurally otherwise as in female. Mesosoma with long whitish-light brownish hairs, only propodeum with scattered individual black hairs intermixed. Legs dark, pubescence whitish with black hairs restricted to fore femorae apically. Hind tarsal claws with inner tooth. Wings as in female. METASOMA. Terga dark, T 1 – 4 with apical margins slightly depressed, apically lightened yellow-hyaline (Fig. 39). Tergal discs finely shagreened, weakly shining, shagreenation absent from marginal areas, these contrasting, more brightly shining. Tergal discs with obscure faint fine punctures, punctures separated by 2 – 4 puncture diameters. Disc of T 1 with long whitish hairs, T 2 – 5 with shorter whitish hairs, T 2 – 4 with thick white apical hairbands that obscure underlying surface laterally, becoming weaker medially. S 8 apically broadened, apex truncate, ventral surface covered with short brown hairs. Genital capsule simple, gonocoxa with very slightly produced rounded apical teeth, gonostyli apically spatulate, flattened, slightly broadened. Penis valves narrow, essentially parallel-sided (Fig. 40).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF9A5138DA08DB9EFE17F85C.taxon	distribution	Distribution Central Iran (Yazd).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF86513DDA42D81CFCB5FE03.taxon	description	urn: lsid: zoobank. org: act: 339546 A 8 - 6 F 7 C- 4250 - 8 E 6 B- 49017 F 4 EB 6 A 8 Figs 41 – 44	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF86513DDA42D81CFCB5FE03.taxon	diagnosis	Diagnosis Andrena elam sp. nov. can be placed in the subgenus Micrandrena Ashmead, 1899 because of its small body size (Fig. 41), black integument, and strongly rugose propodeal triangle. It has a domed clypeus with faint latitudinal striations (Fig. 42) (common in many Turkish Micrandrena and typically reminicient of A. simontornyella Noskiewicz, 1939), and shiny and densely punctate scutum (Fig. 43). This places it close to A. puffina Warncke, 1975 and A. stolida Warncke, 1975. Andrena elam can immediately be separated from both these taxa by the sculpturing of the terga which have extremely strong, raised microreticulation covering the entirety of T 1 and the discs of T 2 – 4 with large, dense punctures intermixed throughout, punctures particularly large and coarse laterally (Fig. 44). Andrena puffina has the terga finely and subtly punctate, and A. stolida has the terga with fine punctures that are not clearly visible against the underlying shagreen, not becoming noticeably coarser laterally. See also A. subviridula sp. nov. (see below).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF86513DDA42D81CFCB5FE03.taxon	etymology	Etymology Named after the ancient kingdom of Elam (ca 3000 – 500 BC, noun in apposition) that was largely found in the region that is now called Ilam Province.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF86513DDA42D81CFCB5FE03.taxon	materials_examined	Material examined Holotype IRAN • ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM. Paratypes IRAN • 9 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 16 ♀♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM • 5 ♀♀; same collection data as for preceding; TJWC • 2 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF86513DDA42D81CFCB5FE03.taxon	description	Description Female MEASUREMENTS. Body length 6.5 – 7 mm (Fig. 41). HEAD. 1.2 times as wide as long. Clypeus dark, domed, punctate, punctures separated by 0.5 – 3 puncture diameters. Clypeus surface microreticulate, centrally and basally with weakly raised microreticulation giving impression of weak latitudinal striations, becoming obscure apically; underlying surface weakly shining (Fig. 42). Process of labrum narrow, weakly tongue-shaped, slightly wider than long, apical margin weakly rounded. Face, gena, vertex, and scape with short whitish to brownish hairs. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly shorter than diameter of lateral ocellus. Foveae narrow, dorsally occupying ¼ distance between compound eye and lateral ocellus, subequal to width of flagellum, of uniform width throughout; foveae filled with light brownish hairs. Antennae dark, A 3 = A 4 + 5. MESOSOMA. Scutum and scutellum polished, smooth and shiny over majority of area, with shagreen in anterior 1 / 5 and laterally as scutum descends to pronotum and pronotal lobes, otherwise without shagreen (Fig. 43). Scutum and scutellum regularly and clearly punctate, punctures separated by 1 – 2 puncture diameters. Pronotum without humeral angle. Mesepisternum and lateral and dorsolateral faces of propodeum with fine granular shagreen, weakly shining; mesespisternum anteriorly and dorsolateral faces of propodeum with fine network of raised rugosity. Propodeal triangle delineated laterally with fine carinae, internal surface with fine network of rugae, thus weakly differentiated from sculpture of dorsolateral faces of propodeum. Mesosoma with moderately long whitish hairs; propodeal corbiculae thin, composed of sparse whitish plumose hairs, internal surface with numerous simple white hairs. Legs uniformly dark, pubescence whitish to light brownish; tibial scopae whitish ventrally, light brownish dorsally; femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Wings hyaline, stigma dark brown, venation dark orange to dark brown, nervulus interstitial to weakly antefurcal; first recurrent vein enters second submarginal cell at its middle. METASOMA. Terga dark, apical margins of T 1 – 4 slightly depressed, depression weakest on T 1, strongest on T 4, all marginal areas unicolourous black. Tergal discs with extremely dense raised microreticulation, tergal discs also densely punctate, punctures separated by 0.5 – 1 puncture diameter, punctures and microreticulation combining to form dense network of bead-like punctures separated by slightly raised margins; integument dull (Fig. 44). Entirety of T 1 covered with dense microreticulation, discs of T 2 – 4 with microreticulation, becoming progressively weaker, disc of T 4 without bead-like punctures, punctures distinct, without raised margins. Marginal areas of T 2 – 4 impunctate, without microreticulation, shagreened and weakly shining. T 2 – 4 medially almost hairless, laterally with weak whitish apical hair fringes, obscure. Tergal hairbands absent. Apical fringe of T 5 and hairs flanking pygidial plate dark brown; pygidial plate rounded triangular, flat. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF86513DDA42D81CFCB5FE03.taxon	distribution	Distribution Southern Iran (provinces of Ilam, Lorestan, Fars).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	description	urn: lsid: zoobank. org: act: 536 BF 1 F 3 - 74 B 7 - 49 E 7 - 9 CCF- 325 F 9359 B 54 E Figs 45 – 53	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	diagnosis	Diagnosis Andrena extenuata sp. nov. can be placed in the subgenus Micrandrena because of its small body size (Fig. 45), black integument, black male clypeus (Fig. 50), and strongly rugose propodeal triangle (Fig. 47). The propodeal triangle is long and finely rugose, and the facial foveae are very narrow throughout their entire length, which is consistent with the Andrena fumida species group (former subgenus Fumandrena Warncke, 1975 which now falls within a broad Micrandrena concept, Pisanty et al. 2022 b). Within the fumida - group and Micrandrena with consistently narrow foveae, A. extenuata is closest to A. protuber Pisanty, 2022 in the female sex because of the protruding clypeus which is centrally weakly punctate and shiny. However, the foveae of A. extenuata are noticeably narrower below, clearly narrower than the width of a flagellum (Fig. 46), linear and without a medial constriction (subequal to width of flagellum, with clear medial constriction), the scutellum is mirror-smooth (scutellum shagreened), and the terga are less densely punctate (Fig. 48) (with clear and moderately dense punctures). In the male sex, A. extenuata sp. nov. can be instantly recognised by its genital capsule, which is narrow with elongated penis valves and gonostyli (Fig. 53). Andrena protuber has a more compact and typical ‘ Micrandrena ’ genital capsule (see illustrations in Pisanty et al. 2022 a); that of A. extenuata is more similar to that of A. sandanskia Warncke, 1973 (elongate, but with gonostyli that broaden apically) or A. djelfensis Pérez, 1895 (gonostyli narrow and pointed, but with strongly developed gonocoxal teeth and gonostyli medially angled, Fig. 54).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	etymology	Etymology From the Latin adjective ‘ extenuatus ’, meaning ‘ thinned, reduced, diminished’ and referring to the narrow and pointed genital capsule.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	materials_examined	Material examined Holotype IRAN • ♂; Yasouj, Doposhteh, Dasht-e Rum; 30.589 ° N, 51.517 ° W; 2091 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; OÖLM. Paratypes IRAN • 3 ♀♀; Yasouj, Deli Bajak, Sepidar; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; ICPI • 3 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; ICPI • 2 ♀♀; same collection data as for preceding; OÖLM • 50 ♂♂, 5 ♀♀; Yazd, Banadak-o sadat; 2103 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 5 ♂♂, 2 ♀♀; same collection data as for preceding; OÖLM • 3 ♂♂, 2 ♀♀; same collection data as for preceding; TJWC • 3 ♂♂, 1 ♀; Yazd, Ghadam gah, Dehbala; 2175 m a. s. l.; 12 Mar. 2020; S. San leg.; ICPI • 1 ♀; Yazd, Hanza, Hosseinie; 1837 m a. s. l.; 6 Mar. 2020; S. San leg.; ICPI • 8 ♂♂; Yazd, Jade konj-e kooh, dakal; 2019 m a. s. l.; 26 Mar. 2021; S. San leg.; ICPI • 7 ♂♂, 1 ♀; Yazd, Khezr abad Ebteday Zorband; 1817 m a. s. l.; 9 Mar. 2020; S. San leg.; ICPI • 1 ♂, 2 ♀♀; Yazd, Khezr abad Zorband; 1844 m a. s. l.; 21 Mar. 2021; S. San leg.; ICPI • 3 ♂♂; Yazd, Khorashe, Mazre Tadayon; 1808 m a. s. l.; 13 Mar. 2020; S. San leg.; ICPI • 3 ♂♂, 1 ♀; Yazd, Mehriz, Hosseiniyeh, Shohaday gomnam; 1500 m a. s. l.; 28 Feb. 2020; S. San leg.; ICPI • 1 ♂; Yazd, Polis-e rah taft; 1399 m a. s. l.; 12 Mar. 2020; S. San leg.; ICPI • 1 ♂, 2 ♀♀; Yazd, Sakhvid, Mazraeh Khosraw; 2469 m a. s. l.; 27 Mar. 2019; S. San leg.; ICPI • 2 ♂♂; Yazd, Sanij khaleghieh; 2060 m a. s. l.; 13 Mar. 2020; S. San leg.; ICPI • 1 ♂; same collection data as for preceding; OÖLM • 1 ♂, 9 ♀♀; Yazd, Shahneh village, Zorband; 1890 m a. s. l.; 23 Mar. 2019; S. San leg.; ICPI • 4 ♀♀; Yazd, Taft, dare gahan; 1877 m a. s. l.; 21 Mar. 2021; S. San leg.; ICPI • 3 ♀♀; Yazd, Taft, Tezerjan, Hanza; 1949 m a. s. l.; 24 Mar. 2019; S. San leg.; ICPI • 3 ♂♂, 16 ♀♀; Yazd, Taft, Tezerjan, Hosseinabad; 2100 m a. s. l.; 22 Mar. 2019; S. San leg.; ICPI • 2 ♂♂; Yazd, Taft, Tezerjan, Mazraeh Ali, Agha Seyyed; 2109 m a. s. l.; 24 Mar. 2019; S. San leg.; ICPI • 1 ♂; same collection data as for preceding; OÖLM • 6 ♂♂; Yazd, Tezerjan Bagh-e Agha Seyyed; 2226 m a. s. l.; 12 Mar. 2020; S. San leg.; ICPI • 5 ♂♂, 16 ♀♀; Yazd, Tezerjan, Hossein Abad Seyyed Mahmood; 1993 m a. s. l.; 5 Mar. 2020; S. San leg.; ICPI • 5 ♂♂, 2 ♀♀; Yazd, Tezerjan, Seyyed Mahmood, 1993 m a. s. l.; 6 Mar. 2020; S. San leg.; ICPI. SYRIA • 14 ♂♂, 1 ♀; Bludan [Bloudan], 57 km NW of Damascus; 2000 m a. s. l.; 24 Apr. 1992; K. Warncke leg.; OÖLM • 9 ♀♀; Burg Baniyas / Mt. Hermon; 1500 m a. s. l.; 13 Apr. 1992; K. Warncke leg.; OÖLM • 1 ♀; Maalula [Maaloula], 60 km NE of Damascus; 1400 m a. s. l.; 14 Apr. 1992; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	description	Description Female MEASUREMENTS. Body length 6.5 – 7 mm (Fig. 45). HEAD. 1.1 times as wide as long. Clypeus dark, domed, irregularly punctate with large punctures, punctures laterally separated by 0.5 puncture diameter, becoming sparser centrally, here separated by up to 4 puncture diameters; clypeus centrally with broad longitudinal impunctate line (Fig. 46). Clypeus integument narrowly shagreened laterally, smooth and shiny over majority of area. Process of labrum quadrate, essentially parallel-sided, as long as broad, corners slightly rounded. Face, gena, vertex, and scape with long whitish hairs, longest not exceeding length of scape. Integument of paraocular area and frons with weak metallic green lustre, frons with weak longitudinal striations, with large, scattered punctures between individual rugae. Gena exceeding width of compound eye; ocelloccipital distance half diameter of lateral ocellus. Foveae extremely narrow, dorsally occupying 1 / 5 distance between compound eye and lateral ocellus, half width of flagellum, very slightly narrowed below at midpoint, otherwise essentially equally narrow throughout, extending slightly below level of antennal insertions. Antennae dark, A 7 – 12 slightly lightened brownish-red below; A 3 subequal to A 4 + 5. MESOSOMA. Scutum with fine and even microreticulation, weakly shining, surface irregularly punctate with moderately sized punctures, punctures separated by 1 – 4 puncture diameters. Scutellum strongly contrasting, narrowly shagreened laterally, remaining surface smooth and shining with small, isolated punctures. Pronotum without humeral angle. Mesepisternum and lateral face of propodeum with fine microreticulation, dull to weakly shining. Dorsal face of propodeum long, equalling length of scutellum plus metanotum; propodeal triangle occupying majority of surface, delineated laterally with fine carinae, internal surface covered by network of fine raised rugae (Fig. 47), remaining surface of dorsal face of propodeum microreticulate, dull. Mesepisternum, scutum, and scutellum with long whitish hairs, longest not exceeding length of scape; propodeal corbiculae thin, composed of sparse whitish plumose hairs, internal surface with numerous simple white hairs. Legs dark, apical tarsal segments slightly lightened dark brown, pubescence white, scopae and flocculus white. Hind tarsal claws with inner tooth. Wings hyaline, stigma orange, venation dark brown basally to orange apically, nervulus antefurcal, first recurrent vein enters second submarginal cell slightly before its middle. METASOMA. Terga dark, apical margins of T 2 – 4 slightly depressed, apically narrowly lightened yellow (Fig. 48). Terga finely microreticulate, weakly shining, microreticulation narrowly disappearing laterally on raised areas immediately above depressed margins, these areas shining. Terga essentially impunctate, with scattered and obscure hair-bearing points disappearing into microreticulation; T 2 – 4 with weak, medially interrupted whitish apical hair bands. Apical fringe of T 5 and hairs flanking pygidial plate whitish-golden; pygidial plate triangular, with truncate apex, with raised longitudinal ridge centrally. Male MEASUREMENTS. Body length 5.5 – 6 mm (Fig. 49). HEAD. 1.3 times as wide as long. Clypeus dark, weakly domed, with large shallow punctures, punctures separated by 0.5 – 1 puncture diameter except basally where a faint impunctate longitudinal line is visible (Fig. 50). Clypeus integument basally and centrally heavily microreticulate, becoming weaker apically, here slightly shining. Process of labrum tongue-shaped, slightly longer than broad, lateral margins weakly converging apically, apical margin straight. Face, gena, vertex, and scape with long black to brown hairs, longest subequal to length of scape. Gena slightly exceeding width of compound eye; ocelloccipital distance less than half diameter of lateral ocellus. Antennae dark, A 5 – 13 slightly lightened greyish-brown below due to presence of tiny cilia; A 3 exceeding A 4, shorter than A 4 + 5, A 4 sub-square. MESOSOMA. Scutum microreticulate, dull, surface with obscure shallow punctures; scutellum with weaker microreticulation, slightly greenish-metallic, punctures larger and more clearly visible. Structurally otherwise as in female. METASOMA. Terga dark, apical margins of T 2 – 4 slightly depressed, more extensively lightened hyalineyellow than in female (Fig. 51). Terga evenly microreticulate, weakly shining, presence of reduced shagreenation apicolaterally less pronounced than in female. Terga with scattered fine golden hairs, without hairbands or lateral fringes. S 8 columnar, slightly narrowed medially, with brown hairs laterally (Fig. 52). Genital capsule long, gonocoxa with very weakly pronounced gonocoxal teeth, gonostyli curved inwards apically, remaining essentially of same width throughout, apically slightly flattened, integument slightly brighter; penis valves long triangular, apically sharply pointed (Fig. 53).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	distribution	Distribution Syria (Anti-Lebanon mountain range) and southern and central Iran, from Yasouj to Yazd.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF805131DA55D8E0FDE2FD21.taxon	discussion	Remarks Collected from flowers of Acer monspessulanum L. (Sapindaceae), Pyrus glabra Boiss. (Rosaceae), and Prunus (Amygdalus) spp. (Rosaceae) from late February to April. It thus seems to be associated with flowering trees in the early spring.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8C5133D9BFDBA9FD19F9A5.taxon	description	urn: lsid: zoobank. org: act: D 3 E 4 C 96 A-F 24 F- 4 E 98 - 8 A 5 B-DD 2 E 17 D 9957 D Figs 55 – 60	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8C5133D9BFDBA9FD19F9A5.taxon	diagnosis	Diagnosis Andrena obsidiana sp. nov. can be place in the subgenus Micrandrena because of its small body size (Fig. 55), black integument, and strongly rugose propodeal triangle. Andrena obsidiana lacks a gradulus at the base of T 2 – 3 (Fig. 60), and has brilliant, completely unshagreened shiny terga. This unusual combination of characters separates it from all Middle Eastern species of Micrandrena. The only slightly similar species is A. omnilaevis Wood, 2020 from northern Spain and Portugal. Apart from the clear geographic separation, A. obsidiana can easily be separated by its narrower foveae which occupy ⅓ of the space between the compound eye and the lateral ocellus (occupying half of this space) and completely shiny scutum and terga (scutum and tergal discs with faint shagreenation).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8C5133D9BFDBA9FD19F9A5.taxon	etymology	Etymology From obsidian, a dark black igneous rock that can be polished to a brilliant shine, in reference to the shiny terga lacking in shagreenation or microreticulation.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8C5133D9BFDBA9FD19F9A5.taxon	materials_examined	Material examined Holotype IRAN • ♀; Ilam Province, Abda Man, Dinar Gaouh; 32.915 ° N, 47.301 ° E; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM. Paratypes IRAN • 44 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 8 ♀♀; same collection data as for preceding; TJWC • 3 ♀♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; M. Kafka leg.; OÖLM • 5 ♀♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; M. Kafka leg.; OÖLM • 2 ♀♀; Fars Province, Dast Arjan; 2040 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♀; Tehran, Ziyaran, Samgh Abad; 1900 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♀; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 7 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; M. Kafka leg.; OÖLM. TURKEY • 1 ♀; Hakkâri, Varegös / Mt. Sat; 1800 m a. s. l.; 21 May 1989; K. Warncke leg.; OÖLM • 1 ♀; Hakkâri, Varegös / Mt. Sat; 1700 m a. s. l.; 15 Jun. 1984; K. Warncke leg.; OÖLM • 1 ♀; Hakkâri, Tanin-Tanin Pass; 2300 m a. s. l.; 12 Jun. 1984; K. Warncke leg.; OÖLM • 1 ♀; Hakkâri, Tanin-Tanin Pass; 2300 m a. s. l.; 19 May 1989; K. Warncke leg.; OÖLM • 2 ♀♀; Hakkâri, Beytüşşebap; 25 May 1988; K. Warncke leg.; OÖLM • 13 ♀♀; Nemrut Dag [Dağı] / Adiyaman; 1500 m a. s. l.; 1 Jun. 1983; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8C5133D9BFDBA9FD19F9A5.taxon	description	Description Female MEASUREMENTS. Body length 6.5 – 7 mm (Fig. 55). HEAD. 1.3 times as wide as long. Clypeus dark, domed, irregularly punctate, punctures from touching to separated by 2 puncture diameters (Fig. 56). Clypeus integument basally with raised microreticulation, forming latitudinal wrinkes, becoming weaker centrally, absent in apical ⅓; clypeus weakly shining throughout. Process of labrum narrow, weakly tongue-shaped, slightly wider than long, apical margin weakly rounded. Face, gena, vertex, and scape with whitish to brownish hairs, longest on vertex, not equalling length of scape. Gena equalling width of compound eyes; ocelloccipital distance slightly less than diameter of lateral ocellus. Foveae narrow, dorsally occupying ¼ of distance between compound eye and lateral ocellus, equalling width of flagellum, slightly narrowed below at level of antennal insertions; foveae filled with light brownish hairs. A 3 = A 4 + 5. MESOSOMA. Scutum and scutellum brilliantly polished, dorsally without trace of shagreenation, smooth and shining (Fig. 57). Scutum with dense but irregularly-sized punctures, punctures separated by 1 – 3 puncture diameters (Fig. 58); scutellum with sparser punctation, punctures separated by 1 – 5 puncture diameters. Pronotum without humeral angle. Mesepisternum and lateral face of propodeum with fine granular shagreen, weakly shining. Propodeal triangle lateraly delineated with fine carinae, internal surface with network of raised rugosity, rugosity extending laterally over boundaries of propodeal triangle onto dorsolateral faces of propodeum, propodeal triangle therefore poorly defined. Mesosoma with short and scattered whitish hairs; propodeal corbiculae thin, dorsally composed of whitish-brownish plumose hairs, internal surface with numerous simple white hairs. Legs uniformly dark, pubescence whitish to light brownish, scopae and flocculus whitish. Hind tarsal claws with small inner tooth. Wings hyaline, stigma dark brown, venation light to dark brown; nervulus interstitial to weakly antefurcal, first recurrent vein enters second submarginal cell at its middle to slightly beyond its middle. METASOMA. Terga dark, apical margins of T 2 – 4 very slightly depressed, margins with extremely narrow rim slightly lightened dark red, essentially unicolourous with disc (Fig. 59). Gradulus absent at visible base of T 2 – 3. Terga without shagreen, uniformly polished, brilliant, smooth and shining throughout (Fig. 60). Discs of terga clearly punctate; T 1 with punctures separated by 2 – 3 puncture diameters centrally, laterally extremely sparse; T 2 – 4 punctures more regular, separated by 1 – 2 puncture diameters throughout; terga margins with scattered irregular weak and fine punctures. Tergal hair bands barely developed, T 2 – 4 laterally with weak and obscure hair fringes. Apical fringe of T 5 and hairs flanking pygidial plate dark golden to brownish; pygidial plate narrowly rounded triangular, flat, dull. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8C5133D9BFDBA9FD19F9A5.taxon	distribution	Distribution Southern (province of Adıyaman) and Eastern Turkey (province of Hakkâri) and Iran (provinces of West Azerbaijan, Tehran, Ilam, Lorestan, Fars).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8E5137D9A5DF5AFDF1FDF2.taxon	description	urn: lsid: zoobank. org: act: 97 FB 1684 - 4 A 57 - 4511 - ACE 3 - 67 A 05460053 A Figs 61 – 69	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8E5137D9A5DF5AFDF1FDF2.taxon	diagnosis	Diagnosis Andrena subviridula sp. nov. can be place in the subgenus Micrandrena because of its small body size (Fig. 61), black integument, black male clypeus (Fig. 66), and strongly rugose propodeal triangle. The propodeal triangle is long and finely rugose, and the facial foveae are very narrow throughout their entire length (Fig. 62), which is consistent with the Andrena fumida - group. In the female sex, it differs from eastern members of this group in the densely microreticulate terga with bead-like punctures (A. griseigena Warncke, 1975, with terga almost impunctate, without raised microreticulation) and the weakly shagreened and shining scutum (A. immaculata Warncke, 1975 and A. kopetica Osytshnjuk, 1993 with scutum uniformly dull). It is similar to A. elam sp. nov. because of the dense raised microreticulation of T 1 – 2 (Fig. 64), but the punctures of T 4 are faint and sparse, separated by 3 – 4 puncture diameters (punctures strong, separated by 1 – 2 puncture diameters; compare Figs 44 and 64) and the scutum has weak granular shagreenation (shagreenation absent, smooth and shining). In the male sex, A. subviridula has the long and dense ‘ beard’ of hairs on the clypeus that project ventrally that is typical of the fumida group species (Fig. 66), and closely resembles A. immaculata in general appearance. However, the genital capsule is different (lacking basally widened penis valves) and more closely resembles that of a A. tomora Warncke, 1975 and A. fabrella Pérez, 1903 (Fig. 70). However, the gonocoxae are not produced into apical points, the gonostyli are apically wider, not parallel-sided, and the inner margin of the gonostyli have a noticeable kink (Fig. 69, absent in A. fabrella, present in A. tomora but other previously mentioned characters apply).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8E5137D9A5DF5AFDF1FDF2.taxon	etymology	Etymology From the Latin prefix ‘ sub ’, meaning ‘ less’ and the Latin adjective ‘ viridulus ’, meaning ‘ green’, in reference to the subtle metallic green colouration of its scutal integument.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8E5137D9A5DF5AFDF1FDF2.taxon	materials_examined	Material examined Holotype IRAN • ♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 36.188 ° N, 51.307 ° E; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM. Paratypes IRAN • 8 ♂♂, 1 ♀; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM • 2 ♂♂, same collection data as for preceding; TJWC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8E5137D9A5DF5AFDF1FDF2.taxon	description	Description Female MEASUREMENTS. Body length 6 mm (Fig. 61). HEAD. 1.3 times as wide as long. Clypeus dark, domed, weakly punctate, punctures separated by 1 – 3 puncture diameters. Clypeus surface microreticulate, centrally and basally with weakly raised latitudinal microreticulation, giving impression of weak latitudinal striations, becoming obscure apically; underlying surface weakly shining (Fig. 62). Process of labrum narrow, slightly longer than wide, narrowly rounded-triangular. Face, gena, vertex, and scape with whitish to brownish hairs, frons laterally with intermixed dark brown hairs at level of antennal insertions. Gena exceeding width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Foveae extremely narrow, dorsally occupying 1 / 5 distance between compound eye and lateral ocellus, subequal to width of flagellum, uniformly wide, clearly extending below level of antennal insertions; foveae filled with dark brown hairs. Antennae dark, A 3 = A 4 + 5. MESOSOMA. Scutum and scutellum weakly shagreened over majority of area, weakly shining, shagreenation becoming stronger over anterior part of scutum, here dull. Scutum with weak and subtle metallic green reflections when viewed laterally. Scutum and scutellum irregularly but clearly punctate, punctures separated by 0.5 – 3 puncture diameters. Pronotum without humeral angle. Mesepisternum and lateral and dorsolateral faces of propodeum with fine granular shagreen, weakly shining; mesespisternum anteriorly and dorsolateral faces of propodeum with fine network of raised rugosity. Propodeal triangle laterally delineated with fine carinae, internal surface covered in fine network of raised rugosity, becoming sparser laterally; propodeal trinangle therefore defined by change in surface sculpture. Mesepisternum with moderately long whitish hairs, scutum and scutellum dorsally with short dark brownish hairs; propodeal corbiculae thin, composed of sparse whitish plumose hairs, internal surface with sparse simple white hairs. Legs dark, apical tarsal segments lightened dark orange, pubescence whitish to light brown; tibial scopae whitish, with scattered brown hairs dorsobasally, femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Wings hyaline, stigma dark orange to dark brown laterally, venation dark orange, nervulus interstitial; first recurrent vein enters second submarginal cell slightly before its middle. METASOMA. Terga dark, apical margins of T 2 – 4 slightly depressed, depression weakest on T 2, strongest on T 4, all marginal areas unicolourous black (Fig. 63). Tergal discs of T 1 – 3 with extremely dense raised microreticulation, tergal discs also densely punctate, punctures separated by 0.5 – 1 puncture diameter, punctures and microreticulation combining to form dense network of bead-like punctures separated by slightly raised margins; integument dull (Fig. 64). Entirety of T 1 covered with dense microreticulation, discs of T 2 – 3 with microreticulation, becoming progressively weaker. Disc of T 4 without bead-like punctures, punctures distinct, without raised margins, punctures separated by 3 – 4 puncture diameters. Marginal areas of T 2 – 4 impunctate, T 2 with fine microreticulation, T 3 – 4 without microreticulation, shagreened and weakly shining. T 2 – 3 laterally with weak whitish apical hair fringes, obscure, T 4 with hair fringe very sparse but complete. Apical fringe of T 5 and hairs flanking pygidial plate golden; pygidial plate rounded triangular, flat, with homogenous granular pattern, dull. Male MEASUREMENTS. Body length 5.5 – 6 mm (Fig. 65). HEAD. 1.3 times as wide as long. Clypeus structurally as in female, entirely covered in long, plumose, ventrally directed brownish hairs (Fig. 66). Process of labrum short, trapezoidal, smooth and shining, apical margin shallowly emarginate. Gena, vertex, and scape with long plumose brownish hairs, longest exceeding length of scape. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly less than half diameter of lateral ocellus. Antennae dark, A 3 sligtly exceeds A 4, shorter than A 4 + 5. MESOSOMA. Mesosoma structurally as in female (Fig. 67), though pubescence exceeding length of scape. METASOMA. Terga structurally as in female, though terga more densely and strongly punctate, punctures variable in size, separated by 0.5 – 1 puncture diameter (Fig. 68). S 8 columnar, apical margin truncate, ventrally covered in moderately long golden hairs. Genital capsule slightly elongate, gonocoxae medioapically truncate, right-angled, not produced into points. Gonostyli with rounded kink in inner margins basally, apically flattened, broadened, not parallel-sided. Penis valves relatively narrow basally, slightly narrowing apically (Fig. 69).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8E5137D9A5DF5AFDF1FDF2.taxon	distribution	Distribution Northern Iran (Tehran Province).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	description	urn: lsid: zoobank. org: act: AA 2 F 5 C 7 B-BAAC- 4 F 04 - 8 FA 9 - 4198 FA 4 E 564 E Figs 71 – 78	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	diagnosis	Diagnosis Andrena tabula sp. nov. can be placed in the subgenus Micrandrena because of its small body size (Fig. 71), black integument, black male clypeus (Fig. 76), and strongly rugose propodeal triangle. It has a strongly flattened clypeus (Fig. 72) and the male genitalia have penis valves that are strongly and conspicuously broadened at the base (Fig. 78), placing it close to A. saxonica Stoeckhert, 1935 and A. garzetta Warncke, 1975 (eastern limit Hatay Province, Turkey). Female A. tabula sp. nov. can be separated from A. saxonica by the comparatively more flattened clypeus (Fig. 72), flattened over the entirety of its surface (clypeus laterally weakly domed, flattened over ¾ of its surface) and the shinier scutum and scutellum, weakly shining with shagreenation that is comparatively weak (clearly and densely microreticulate, generally dull). In addition to the more flattened clypeus, they can be separated from A. garzetta by the much stronger sculpture and denser punctation of the scutum, punctures separated by 0.5 – 1 puncture diameter (finely and moderately punctate, punctures separated by 2 puncture diameters, underlying surface finely shagreened, weakly shining). In addition to the more clearly flattened clypeus, males can most obviously be separated by their genitalia (Fig. 78). Andrena tabula sp. nov. genitalia have the inner margin of the gonostyli raised but straight, more or less parallel with outer margin (A. saxonica with inner margin of gonostyli raised, curved towards penis valves, and with clear angle apically, not parallel with outer margin) and the gonocoxae are very weakly produced into apical points (A. garzetta with gonocoxae clearly produced into strong, apically extending points).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	etymology	Etymology From the Latin noun ‘ tabula ’, meaning ‘ tablet’ or ‘ board’, in reference to the extremely flat clypeus in both sexes.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	materials_examined	Material examined Holotype IRAN • ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 30.545 ° N, 51.610 ° E; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM. Paratypes IRAN • 4 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂, 1 ♀; same collection data as for preceding; TJWC • 1 ♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; M. Kafka leg.; OÖLM. TURKEY • 1 ♂, 1 ♀; Hakkâri, Suvari Halil-Pass; 2600 m a. s. l.; 15 Jun. 1981; K. Warncke leg.; OÖLM • 1 ♀; same collecting data as for preceeding; SMNHTAU • 1 ♀; Hakkâri, Suvari Halil-Pass; 2800 m a. s. l.; 14 Jun. 1984; K. Warncke leg.; OÖLM • 3 ♀♀; Hakkâri, Tanin-Tanin-Pass; 2500 m a. s. l.; 2 Jun. 1980; K. Warncke leg.; SMNHTAU • 1 ♀; Tanin-Tanin-Pass; 23 – 2600 m a. s. l.; 3 Jun. 1980; M. Schwarz leg.; OÖLM • 4 ♀♀; Tanin-Tanin-Pass; 1700 m a. s. l.; 12 Jun. 1984; K. Warncke leg.; OÖLM • 1 ♀; Tanin-Tanin-Pass; 2300 m a. s. l.; 19 May 1989; K. Warncke leg.; OÖLM • 1 ♀; Hakkâri, Mt. Sat; 2050 – 2450 m a. s. l.; 10 Jun. 1981; K. Warncke leg.; SMNHTAU • 1 ♀; Nemrut Dağı, 50 km NE of Kanta [Kahta]; 2 – 14 Jun. 1996; P. Jelinek leg.; OÖLM. Other material examined (Andrena saxonica) TURKEY • 1 ♀; Hakkâri, Beytüşşebap; 25 May 1988; K. Warncke leg.; OÖLM • 20 ♀♀; Hakkâri, Tanin-Tanin-Pass; 1700 m a. s. l.; 12 Jun. 1984; K. Warncke leg.; OÖLM • 1 ♂, 1 ♀; Kars, 10 km E of Karakurt; 1460 m a. s. l.; 31 May – 1 Jun 1988; K. Warncke leg.; OÖLM • 1 ♂, Kars, 20 km W of Sarikamis [Sarıkamış]; 2100 m a. s. l.; 29 May 1983; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	description	Description Female MEASUREMENTS. Body length 7 – 8 mm (Fig. 71). HEAD. 1.2 times as wide as long. Clypeus dark, strongly and conspicuously flattened over entire surface, with irregular and variably sized punctures, punctures separated by 1 – 4 puncture diameters, becoming gradually sparser centrally, central ⅓ of clypeus impunctate (Fig. 72). Clypeus integument with fine granular microreticulation, dull. Process of labrum trapezoidal, slightly wider than long. Face, gena, vertex, and scape with short whitish to brownish hairs. Gena equalling width of compound eyes; ocelloccipital distance slightly less than diameter of lateral ocellus. Foveae extremely narrow, dorsally occupying 1 / 5 distance between compound eye and lateral ocellus, slightly narrower than width of flagellum, uniform width throughout; foveae filled with whitish hairs. Antennae dark, A (4) 5 – 12 slightly lightened dark reddish-brown below; A 3 slightly exceeding A 4 + 5, clearly shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum shagreenation, weakly shining, regularly and densely punctate with clear punctures, punctures separated by 0.5 – 1 puncture diameter (Fig. 73). Pronotum without humeral angle. Mesepisternum, lateral and dorsolateral faces of propodeum with sparse network of raised reticulation, underlying surface dull to weakly shining in places. Propodeal triangle weakly delineated laterally by raised carinae, internal surface strongly and regularly rugose, clearly differing from irregular pattern of reticulation on dorsolateral faces of propodeum. Mesepisternum with moderately long whitish hairs, scutum and scutellum with sparse extremely short brownish hairs, scutellum with occasional scattered long dark brown-black hairs. Propodeal corbiculae thin, dorsally composed of whitish plumose hairs, internal surface with numerous simple white hairs. Legs uniformly dark, pubescence brownish, scopae and flocculus whitish to light brownish. Hind tarsal claws with small inner tooth. Wings hyaline, stigma dark orange to dark brown laterally, venation dark brown; nervulus interstitial, first recurrent vein enters second submarginal cell at its middle. METASOMA. Terga dark, apical margins of T 1 weakly, T 2 – 4 more strongly depressed, with extremely narrow hyaline rim (Fig. 74). Terga with discs strongly microreticulate, irregularly and very finely punctured, punctures fading into microreticulation, surface roughened, dull. Marginal areas of terga contrasting, lacking punctures, evenly and finely microreticulate, weakly shining. Terga laterally with sparse short hairs, T 2 – 4 with weak and obscure hair fringes. Apical fringe of T 5 and hairs flanking pygidial plate dark brown; pygidial plate rounded triangular, flattened with slightly raised apical rim, internal surface finely punctate, weakly shining. Male MEASUREMENTS. Body length 7 – 7.5 mm (Fig. 75). HEAD. 1.3 times as wide as long. Clypeus dark, strongly and conspicuously flattened over entire surface, with irregular punctures, punctures separated by 0.5 – 3 puncture diameters (Fig. 76). Clypeus integument with fine granular microreticulation, dull. Process of labrum trapezoidal, wider than long, apical margin weakly emarginate. Face, gena, vertex, and scape with short whitish to brownish hairs. Gena slightly exceeds width of compound eyes; ocelloccipital distance equals diameter of lateral ocellus. Antennae dark, A 3 slightly shorter than A 4 + 5. MESOSOMA. Scutum and scutellum shagreened, weakly shining (Fig. 77), structurally otherwise as in female. METASOMA. Terga dark, apical margins of T 1 weakly, T 2 – 4 strongly depressed, with extremely narrow hyaline rim. Tergal discs roughened, strongly microreticulate, densely punctate, punctures separated by 1 puncture diameter. Punctures absent from marginal areas, with microreticulation becoming weaker apically, weakly shining; marginal areas strongly contrasting tergal discs. Tergal discs with scattered irregular short white hairs. S 8 columnar, apically rounded, ventral surface covered with short brown hairs. Genital capsule large, gonocoxae very weakly produced into apical points. Gonostyli broad, slightly broadening apically with strongly truncate and square-shaped apices, apically flattened and broadly spatulate. Penis valves strongly broadened basally, strongly narrowing apically (Fig. 78).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	discussion	Remarks Some of the specimens from Turkey were separated by Warncke with type labels and were called ‘ Andrena galacta Warncke’, but he never published this name. Other Turkish specimens were discovered in undetermined material. For A. saxonica, though the maps of Gusenleitner & Schwarz (2002) show an eastern range limit in the Bosphorus, Turkey, the two taxa can be found in sympatry in eastern Turkey, based on the examination of previously undetermined material.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FF8A5108DA4BDB90FE6AF90D.taxon	distribution	Distribution Southern (province of Adiyaman) and Eastern Turkey (province of Hakkâri) and southern Iran (provinces of Ilam, Lorestan, Fars).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB5510DDA43DFE2FD1BFD8B.taxon	description	urn: lsid: zoobank. org: act: E 4762290 - 100 B- 4 C 49 - 94 F 9 - 5 DB 4273 D 990 B Figs 79 – 84	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB5510DDA43DFE2FD1BFD8B.taxon	diagnosis	Diagnosis Placement is tricky because of taxonomic confusion surrounding taxa previously placed in the subgenus Carandrena Warncke, 1968, which is now a strict synonym of Notandrena Pérez, 1890, but Carandrena sensu Warncke is polyphyletic (Pisanty et al. 2022 b). Andrena idigna sp. nov. has the general appearance of a Carandrena because of its broad head (Fig. 81), tergal hairbands (Fig. 84), and lack of any obvious derived features, one of the reasons why this former subgenus has been used as a ‘ waste basket’ for species that cannot be easily allocated elsewhere. Its integument lacks metallic reflections or red colouration on the terga, immediately separating it from almost all species in this group. It is therefore superficially similar to A. euzona Pérez, 1895, which also has dark terga. However, A. euzona has an extremely shiny scutum and is found outside the Carandrena + Notandrena group, whereas A. idigna has the scutum entirely and uniformly shagreened and dull. Though A. idigna lacks a clear humeral angle on the pronotum (faint hint of this trait present but obscure), a character that is usually present in the Notandrena, provisional placement in this subgenus is the most appropriate because of the broad head, shagreened scutum, shagreened and essentially impunctate terga, and tergal hairbands. Its dark terga, shagreened scutum, and unusual propodeal triangle, which is delineated laterally by fine carinae but with a narrow, longitudinal, and slightly impressed channel surrounded by irregular latitudinal rugae along its length, allow separation from all related taxa. This subgeneric placement should be reassessed following discovery of male specimens and DNA sampling.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB5510DDA43DFE2FD1BFD8B.taxon	etymology	Etymology From the Sumerian ‘ idigna ’ or ‘ idigina ’, meaning ‘ running water’, in reference to its locus typicus on the Karun River, the largest river in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB5510DDA43DFE2FD1BFD8B.taxon	materials_examined	Material examined Holotype IRAN • ♀; Mollasani, Ahwaz [Ahvaz]; 17 Feb. 1965; S. Tirgari leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB5510DDA43DFE2FD1BFD8B.taxon	description	Description Female MEASUREMENTS. Body length 9 mm (Fig. 79). HEAD. 1.4 times as wide as long (Fig. 80). Clypeus dark, weakly domed, densely punctate, punctures separated by 0.5 – 2 puncture diameters (Fig. 81). Clypeus with fine and dense granular shagreen, dull to weakly shining centrally. Process of labrum trapezoidal, two times as wide as long, corners rounded. Face, gena, vertex, and scape with moderately long white hairs, not exceeding length of scape. Gena slightly exceeding width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Foveae dorsally broad, occupying ⅔ rd of space between compound eye and lateral ocellus, very slightly narrowed below at level of antennal insertions; filled with whitish hairs. Antennae dark basally, A 5 – 12 lightened dark orange-red below; A 3 exceeding A 4 + 5, shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum densely and uniformly granularly shagreened, dull; surface with irregular and obscure punctures, punctures separated by 2 – 3 puncture diameters, not well-differentiated from underlying surface (Fig. 82). Pronotum without clear humeral angle, slightly angulate above, obscure. Mesepisternum and lateral and dorsolateral faces of propodeum with fine granular shagreen, dull; dorsolateral faces of propodeum with sparse network of fine, slightly raised rugae. Propodeal triangle declineated laterally with fine carinae, internal surface with narrow, longitudinal, and slightly impressed channel surrounded by irregular latitudinal rugae along its length. Remaining surface of propodeal triangle with fine granular shagreen, finer than on dorsolateral faces of propodeum; propodeal triangle thus defined by change in surface sculpture. Mesosoma with moderately long whitish to greyish hairs, not exceeding length of scape; propodeal corbiculae with weakly plumose hairs dorsally, inner surface with weakly plumose hairs. Legs dark, apical tarsal segments lightened dark orange; pubescence whitish. Scopae and flocculus white. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation orange, nervulus weakly antefurcal; first recurrent vein enters second submarginal cell at its middle. METASOMA. Terga dark, apical margins of T 2 – 4 slightly depressed, marginal areas of T 1 – 4 lightened dark red to yellow-hyaline apically (Fig. 83). Terga with uniform granular shagreen, weakly shining, essentially impunctate with only scattered and obscure hair-bearing punctures (Fig. 84). Tergal discs with sparse whitish hairs, longest on T 1, becoming progressively shorter apically. T 1 – 4 with white apical hair fringes, weak and interrupted on T 1, complete and dense on T 2 – 4, obscuring underlying surface. Fringe of T 5 and hairs flanking pygidial plate golden; pygidial plate broadly rounded, centrally densely punctate, with narrow impunctate rim. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB5510DDA43DFE2FD1BFD8B.taxon	distribution	Distribution South-western Iran (Khuzestan Province).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB0510FD9B5DB69FE60FB52.taxon	description	urn: lsid: zoobank. org: act: C 5 B 11605 - 577 C- 4490 - B 0 FE-B 29 E 187 A 8795 Figs 85 – 90	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB0510FD9B5DB69FE60FB52.taxon	diagnosis	Diagnosis Andrena flagrans sp. nov. can be placed in the subgenus Planiandrena Osytshnjuk, 1983 because of its very broad process of the labrum (Fig. 88), narrow foveae that tightly follow the inner margin of the compound eye (Fig. 87), broad and finely shagreened propodeal triangle with weakly indicated lateral margins, shiny terga (Fig. 90), hind tibial spur that is slightly broadened submedially, and pygidial plate with slightly raised central area. The subgenus was described from four Central Asian species (Osytshnjuk 1983 a), but it reaches to Israel (Pisanty et al. 2022 a). Andrena flagrans differs from Central Asian taxa because the clypeus is convex (not flattened or depressed) and is most similar to A. veterana Pisanty, 2022 and A. huma sp. nov. (see below for separation from this species). Andrena flagrans can easily be separated from A. veterana because the terga are entirely shiny (Fig. 90), without shagreen, discs impunctate (tergal discs of T 2 – 4 basally shagreened, sparsely and very finely punctured) and the propodeal triangle lacks short raised longitudinal rugae along its basal margin (propodeal triangle with short raised longitudinal rugae).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB0510FD9B5DB69FE60FB52.taxon	etymology	Etymology From the Latin present participle ‘ flagrans ’, meaning ‘ burning, blazing’, or ‘ shining’, in reference to the strongly shining terga.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB0510FD9B5DB69FE60FB52.taxon	materials_examined	Material examined Holotype IRAN • ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 30.545 ° N, 51.610 ° E; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM Paratypes IRAN • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; TJWC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB0510FD9B5DB69FE60FB52.taxon	description	Description Female MEASUREMENTS. Body length 10 mm (Fig. 85). HEAD. 1.2 times as wide as long (Fig. 86). Clypeus dark, weakly domed, laterally with large shallow punctures, punctures separated by 1 – 2 puncture diameters, centrally becoming sparser, separated by 3 – 4 puncture diameters; clypeus centrally with wide longitudinal impunctate line, becoming wider apically, apical third of clypeus without punctures (Fig. 87). Clypeus surface laterally and basally shagreened, becoming weaker apically, apical half of clypeus smooth and shining. Process of labrum short, broadly trapezoidal, four times as wide as long, covered in fine latitudinal wrinkles (Fig. 88). Face, gena, vertex, and scape with long whitish hairs, longest equalling length of scape. Gena equalling width of compound eye; ocelloccipital distance small, ½ diameter of lateral ocellus. Foveae narrow dorsally, occupying ⅓ of space between compound eye and lateral ocellus, narrowed slightly below at level of antennal insertions; filled with light brown hairs. Antennae dark, A 3 exceeding A 4 + 5, slightly shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum laterally shagreened, weakly shining, shagreen weaker centrally, here shining more strongly; surface irregularly punctate with shallow punctures, punctures separated by 1 – 3 puncture diameters, sparser centrally (Fig. 89). Pronotum without humeral angle. Mesepisternum with fine granular reticulation, weakly shining, covered in faint extremely shallow punctures with poorly defined margins. Internal part of propodeal corbiculae with fine granular reticulation, weakly shining. Propodeal triangle broad, not structurally delineated laterally but very slightly depressed, basally with very small and faint raised rugae, elsewhere with granular shagreen, weakly shining. Dorsolateral faces of propodeum with stronger shagreened, duller, propodeal triangle therefore defined by change in surface sculpture. Mesosoma with long whitish hairs, sparser dorsally, longest equalling length of scape; propodeal corbicula with weakly plumose hairs dorsally, inner surface with long simple whitish hairs. Legs dark, basitarsi and apical tarsal segments progressively lightened orange, pubescence whitish. Tibial scopae white with few brownish hairs dorsally at base, femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Hind tibial spur slightly broadened submedially. Wings hyaline, stigma and venation orange, nervulus interstitial; first recurrent vein enters second submarginal cell beyond its middle. METASOMA. Terga dark, marginal areas of T 2 – 4 very slightly depressed, T 1 – 4 apically narrowly lightened dark brown (Fig. 90). Terga extremely smooth and shiny, almost without shagreen; tergal discs essentially impunctate, tergal margins with small dense punctures, punctures separated by 1 – 2 puncture diameters. T 1 with scattered long whitish hairs, T 2 – 4 with sparse short hairs laterally, apical hairbands absent. Fringe of T 5 and hairs flanking pygidial plate golden; pygidial plate rounded triangular with slightly raised area centrally. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB0510FD9B5DB69FE60FB52.taxon	distribution	Distribution Southern Iran (Yasouj).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	description	urn: lsid: zoobank. org: act: F 848 A 003 - A 55 A- 448 F- 8 DF 0 - 22 E 51 F 524941 Figs 91 – 102	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	diagnosis	Diagnosis Like A. flagrans sp. nov., A. huma sp. nov. can be placed within the subgenus Planiandrena because of its very broad process of the labrum (Fig. 93), narrow foveae that tightly follow the inner margin of the compound eye, broad and finely shagreened propodeal triangle with weakly indicated lateral margins (Fig. 95), hind tibial spur that is slightly broadened submedially, and pygidial plate with slightly raised central area. Females are most similar to those of A. veterana and A. flagrans because of the slightly convex clypeus (Fig. 92). Andrena huma can easily be separated from both A. veterana and A. flagrans as T 1 is entirely shagreened (vs T 1 entirely shiny, without shagreen), the apical ⅓ of the clypeus is smooth and shining (vs entirely shagreened in A. veterana), the terga have strong apical hairbands (vs terga without apical hairbands in A. flagrans), the weak structure of the basal part of the propodeal triangle (vs propodeal triangle basally with strong longitudinal rugae in A. veterana), and the dark hairs on T 5 and flanking the pygidial plate (Fig. 96) (vs hairs here golden in A. flagrans). Males have a yellow-marked clypeus (Figs 98 – 99), putting them close to A. veterana and A. planirostris Morawitz, 1876, the only two known Planiandrena with a yellow clypeus in the male sex. Andrena huma sp. nov. can easily be separated from A. veterana because the genital capsule is elongate (Fig. 102), simple, the inner margin of the gonostyli lacks an angular extension, and the gonocoxal teeth are produced into acute points (genital capsule comparatively compact, inner margin of gonostyli with angular extension, gonocoxal teeth truncate, not forming clear apically projecting teeth). The genital capsule of A. huma is similar to that of A. planirostris, but the gonocoxal teeth are much more strongly pronounced and the penis valves are strongly broadened at the base before narrowing apically, therefore triangular (gonocoxal teeth only slightly pronounced, penis valves very weakly broadened basally, essentially parallel-sided, see illustrations in Osytshnjuk 1983 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	etymology	Etymology Named after the mythical Huma bird from Iranian legends and fables that was said to never alight upon the ground.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	materials_examined	Material examined Holotype IRAN • ♂; Yazd, Tezerjan, Hossein Abad Seyyed Mahmood; 31.624 ° N, 54.214 ° E; 1993 m a. s. l.; 5 – 6 Mar. 2020; S. San leg.; OÖLM. Paratypes GOLAN HEIGHTS • 2 ♂♂; Har Avital; 15 Mar. 1995; R. Kasher leg.; SMHNTAU. IRAN • 5 ♂♂; Yazd, Tezerjan, Hossein Abad Seyyed Mahmood; 31.624 ° N, 54.214 ° E; 1993 m a. s. l.; 5 – 6 Mar. 2020; S. San leg.; OÖLM • 3 ♂♂; same collection data as for preceding; TJWC • 19 ♂♂, 2 ♀♀; same collection data as for preceding; ICPI • 1 ♂, 3 ♀♀; Yazd, Banadak-o sadat; 2103 m a. s. l.; 22 Mar. 2021; S. San leg.; OÖLM • 1 ♂, 3 ♀♀; same collection data as for preceding; TJWC • 3 ♂♂, 14 ♀♀; same collection data as for preceding; ICPI • 2 ♂♂, 4 ♀♀; Yazd, Jade konj-e kooh, dakal; 2019 m a. s. l.; 26 Mar. 2021; S. San leg.; ICPI • 2 ♂♂; Yazd, Khezr abad Ebteday Zorband; 1817 m a. s. l.; 9 Mar. 2020; S. San leg.; ICPI • 1 ♀; Yazd, Khezr abad Zorband; 1844 m a. s. l.; 21 Mar. 2021; S. San leg.; ICPI • 6 ♂♂; Yazd, Sakhvid, Mazraeh Khosraw; 2469 m a. s. l.; 27 Mar. 2019; S. San leg.; ICPI • 1 ♀; Yazd, Sanij khaleghieh; 2060 m a. s. l.; 13 Mar. 2020; S. San leg.; ICPI • 1 ♂, 1 ♀; Yazd, Sanij; 2337 m a. s. l.; 22 Mar. 2019; S. San leg.; ICPI • 1 ♀; Yazd, Taft, Tezerjan, Hanza; 1949 m a. s. l.; 24 Mar. 2019; S. San leg.; ICPI • 6 ♀♀; Yazd, Taft, Tezerjan, Hosseinabad; 2100 m a. s. l.; 22 Mar. 2019; S. San leg.; ICPI • 1 ♀; Yazd, Taft, Tezerjan, Mazra ali, Agha Seyyed; 2109 m a. s. l.; 24 Mar. 2019; S. San leg.; ICPI • 1 ♀; Yazd, Tang e chenar, Az konj-e kooh; 2237 m a. s. l.; 23 Mar. 2021; S. San leg.; ICPI. SYRIA • 1 ♀; Burg Baniyas / Mt. Hermon; 1500 m a. s. l.; 14 Apr. 1992; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	description	Description Female MEASUREMENTS. Body length 10 – 11 mm (Fig. 91). HEAD. 1.2 times as wide as long. Clypeus dark, weakly domed, with large shallow punctures separated by 2 – 3 puncture diameters, becoming sparser apically, separated by 3 – 4 puncture diameters. Clypeus surface laterally and in basal half shagreened with network of fine slightly raised reticulation, weakly shining; in apical ⅓ smooth and shining, strongly contrasting with basal half (Fig. 92). Process of labrum short, three times as wide as long, surface with fine longitudinal wrinkles (Fig. 93). Face, gena, vertex, and scape with long white hairs, longest not exceeding length of scape; vertex and frons with some intermixed black hairs. Gena exceeding width of compound eye; ocelloccipital distance small, slightly less than diameter of lateral ocellus. Foveae narrow dorsally, occupying at most ⅓ of space between compound eye and lateral ocellus, narrowed below at level of antennal insertions; filled with dark brown hairs. Antennae dark, A 3 slightly exceeding A 4 + 5 + 6. MESOSOMA. Scutum and scutellum uniformly strongly shagreened to microreticulate, dull, surface with obscure shallow punctures, punctures separated by 0.5 – 2 puncture diameters (Fig. 94). Pronotum with hint of humeral angle, inconspicuous. Mesepisternum with fine network of raised reticulation, punctures with slightly raised ‘ crater-like’ rims, underlying surface weakly shining. Lateral faces of propodeum shagreened, weakly shining. Propodeal triangle broad, very subtly defined laterally with fine carinae, basally with short raised longitudinal rugae, extending slightly to central part of propodeal triangle; internal surface with granular shagreen, weakly shining (Fig. 95). Dorsolateral faces of propodeum with sparse raised reticulation, propodeal triangle therefore defined by change in surface sculpture. Mesosoma with long whitish hairs, sparser dorsally, longest not exceeding length of scape; propodeal corbicula with weakly plumose hairs dorsally, inner surface with long simple whitish hairs. Legs dark, pubescence light brownish to whitish. Tibial scopae white with numerous brownish hairs dorsally at base, femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Hind tibial spur slightly broadened submedially. Wings hyaline, stigma dark orange with dark brown margins, venation dark brown, nervulus interstitial; first recurrent vein enters second submarginal cell beyond its middle. METASOMA. Terga dark, marginal areas of T 1 – 4 slightly depressed, T 1 – 4 apically very narrowly lightened light brown-hyaline (Fig. 96). Terga shagreened, most strongly on T 1, becoming progressively weaker apically, weakly shining. Tergal discs and margins with scattered punctures, punctures separated by 1 – 5 puncture diameters. Tergal discs with scattered white hairs, longest on T 1, T 1 – 4 with dense apical fringes of white hairs that form hairbands, broadly interrupted on T 1 – 2, complete on T 3 – 4. Fringe of T 5 and hairs flanking pygidial plate dark brown; pygidial plate rounded triangular with slightly raised area centrally. Male MEASUREMENTS. Body length 9 – 10.5 mm (Fig. 97). HEAD. 1.2 times as wide as long (Fig. 98). Clypeus centrally with large triangular yellow mark, occupying majority of surface area, not extending completely to margins, centro-laterally with two black marks (Fig. 99). Clypeus weakly flattened, with weak and even punctures, punctures separated by 1 puncture diameter, slightly sparser centrally; underlying surface shagreened, weakly shining. Process of labrum broadly trapezoidal, three times as wide as long, fore margin widely and shallowly emarginate. Face centrally, clypeus, and scape with white hairs, forming strong and long fringe on apical margin of clypeus; face laterally with strongly contrasting black hairs along inner margin of compound eye. Gena with long white hairs ventrally, extending to vertex, black hairs intermixed dorsolaterally. Gena equalling width of compound eye; ocelloccipital distance slightly less than diameter of lateral ocellus. Antennae dark, A 3 exceeding A 4, slightly shorter than A 4 + 5. MESOSOMA. Mesosoma structurally as in female, though scutum duller, punctures more obscure (Fig. 100). Pubescence as in female, though lateral faces of propodeum with many intermixed black hairs. METASOMA. Terga dark, similar to female but with stronger shagreenation, additionally with slightly raised microreticulation on basal parts of terga, forming weak latitudinal wrinkles; terga therefore duller. Terga discs with stronger, more consistent, and more pronounced punctation, separated by 2 puncture diameters. Terga discs more extensively haired, with moderately long whitish hair, not obscuring underlying surface (Fig. 101). T 1 – 4 with apical white hairbands, slightly interrupted medially. S 8 weakly triangular, slightly narrowed medially, apical half parallel-sided, apically truncate. Genital capsule long, gonocoxa with strongly pronounced elongate teeth, gonostyli apically broadened. Penis valves broadened basally, laterally exceeding apices of gonocoxal teeth, narrowed apically (Fig. 102).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	discussion	Remarks In Iran, collected from flowers of Prunus (Amygdalus) spp. in March. It may also be associated with flowering trees.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB25103DA60DC30FB5AF93B.taxon	distribution	Distribution Central Iran (Yazd) and the Anti-Lebanon mountain chain (southern Syria, Golan Heights).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFBE5104DA43DFD9FE60FE0D.taxon	description	urn: lsid: zoobank. org: act: EDCCBC 04 - DFE 5 - 46 BD- 92 C 6 - 2 EA 819162 D 4 C Figs 103 – 108	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFBE5104DA43DFD9FE60FE0D.taxon	diagnosis	Diagnosis Andrena sella sp. nov. differs from the two previously described species of Planiandrena. It has the same very broad process of the labrum (Fig. 105) and narrow foveae that tightly follow the inner margin of the compound eye, but the propodeal triangle is noticeably narrower (Fig. 107), slightly depressed, and there is no obviously raised area centrally. The propodeal triangle does, however, still have basal rugae, a feature present in A. huma sp. nov. and absent in A. flagrans sp. nov. Because the shape of the head is so extremely similar, A. sella is confidently placed in the Planiandrena. It can be instantly separated from all described species of Planiandrena because the apical margin of T 1 and the entirety of T 2 are red marked (Fig. 108). Structurally, it can be separated by the combination of the shape of the propodeal triangle, which is slightly depressed and medially narrowed (broad and unnarrowed in A. huma and A. flagrans), by the clypeus, which is domed and highly polished over the majority of its area (Fig. 105) (shiny only in the apical ⅓ in A. huma and A. flagrans), and by the terga, which are shiny but clearly punctate throughout, both on the discs and the marginal areas (at least basally shagreened in A. huma, without shagreen but also with only occasional punctures in A. flagrans).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFBE5104DA43DFD9FE60FE0D.taxon	etymology	Etymology From the Latin noun ‘ sella ’, meaning ‘ saddle’, in reference to the red-banded abdomen.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFBE5104DA43DFD9FE60FE0D.taxon	materials_examined	Material examined Holotype IRAN • ♀; Yasouj, Deli Bajak, Sepidar; [30.518 ° N, 51.495 ° E]; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFBE5104DA43DFD9FE60FE0D.taxon	description	Description Female MEASUREMENTS. Body length 8.5 mm (Fig. 103). HEAD. 1.3 times as wide as long (Fig. 104). Clypeus dark, domed, with clear rounded punctures, punctures laterally separated by 0.5 – 1 puncture diameter; punctures becoming sparse and irregular centrally and apically, punctures here separated by 1 – 6 puncture diameters (Fig. 105). Clypeus surface laterally and basally with narrow area of slightly raised microreticulation, weakly shining; clypeus strongly polished and shining over majority of remaining area. Process of labrum short, rounded trapezoidal, four times as wide as long. Face, gena, vertex, and scape with long whitish hairs, frons and gena dorsolaterally with some intermixed black hairs, no hairs exceeding length of scape. Gena exceeding width of compound eye; ocelloccipital distance small, ½ diameter of lateral ocellus. Foveae narrow dorsally, occupying ⅓ of space between compound eye and lateral ocellus, narrowed below at level of antennal insertions; filled with dark brown hairs. Antennae dark, A 3 exceeding A 4 + 5, shorter than A 4 + 5 + 6. MESOSOMA. Scutum with dense granular microreticulation, weakly shining, with large shallow punctures, punctures separated by 2 – 3 puncture diameters (Fig. 106). Scutellum with shagreen restricted to lateral areas, smooth and shining over majority of area, with scattered punctures. Pronotum with hint of humeral angle, inconspicuous. Mesepisternum and lateral and dorsolateral faces of propodeum with granular shagreenation and fine network of raised reticulation, very weakly shining. Propodeal triangle slightly depressed, internal surface with weakly shining granular microreticulation; basally with network of short raised longitudinal rugae, centrally longer and extending into centre of propodeal triangle; propodeal triangle therefore defined by depressed surface area and change in surface sculpture (Fig. 107). Mesosoma with long whitish hairs, equalling length of scape, propodeal corbiculae with long white plumose hairs, internal surface with long white simple hairs. Legs uniformly dark, with whitish to light brownish pubescence; tibial scopae white with intermixed brown hairs dorsobasally. Femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Hind tibial spur parallel-sided. Wings hyaline, stigma dark orange, venation orange to dark orange, nervulus weakly antefurcal; first recurrent vein enters second submarginal cell beyond its middle. METASOMA. Apical margin of T 1 and disc of T 2 red marked, with exception of dark mark centrally on T 2 (Fig. 108). Remaining terga dark, apical margins of T 3 – 4 lightly lightened yellow-hyaline. Terga with very weak shagreenation, smooth and shining throughout. Terga clearly punctate, T 1 with punctures sparse, separated by 3 – 4 puncture diameters, T 2 – 4 more densely punctate, punctures separated by 1 – 2 puncture diameters. Tergal discs with extremely scattered short white hairs, T 1 – 4 with dense unbroken apical fringes of white hairs that obscure underlying integument. Fringe of T 5 and hairs flanking pygidial plate brown; pygidial plate rounded triangular, broadly flat without obvious structure. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFBE5104DA43DFD9FE60FE0D.taxon	distribution	Distribution Southern Iran (Yasouj).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB95119DA41D8E3FD92F85C.taxon	description	urn: lsid: zoobank. org: act: 86 FAABC 5 - 56 AB- 4758 - AC 10 - 8 E 045 BD 9 AC 28 Figs 109 – 120	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB95119DA41D8E3FD92F85C.taxon	diagnosis	Diagnosis Andrena bulbosa sp. nov. can swiftly be placed in the subgenus Ulandrena Warncke, 1968 by the combination of the inner hind tibial spur that is broadened slightly before its base, the yellow-marked male clypeus (Fig. 116), and generally enlarged genital capsule (Fig. 118). It belongs to the Andrena polita species group and is closest to the larger species with red-marked terga, specifically A. caspica Morawitz, 1886, A. unicincta Friese, 1899, and A. unifasciata Friese, 1899. Males can instantly be separated by the unique genitalia, which lack gonocoxal teeth (Fig. 118) (vs strongly produced and pointed in A. caspica and A. unifasciata, weakly produced and laterally diverging in A. unicincta). Females can be separated from A. unicincta by the shiny scutum (Fig. 112) (vs dull) and interstitial nervulus (vs postfurcal). Separation from A. caspica can be made by the smaller body size of 10 mm (12 – 13 mm) as well as the comparatively shinier scutum (vs only slightly shiny in A. caspica, comparative material required). It is therefore very similar to A. unifasciata, and based on the limited female material, separation from this taxon in the female sex is not currently possible. Similar red-marked female material of Ulandrena collected from Uludere in Şırnak Province (Turkey) by Klaus Warncke (16 Jun. 1981) is morphologically inseparable from Levantine specimens of A. unifasciata and the Iranian specimen of A. bulbosa sp. nov. Males must be found that can resolve the identity of specimens from south-eastern Turkey, which has biogeographical affinities with both the Levant and Iran. Molecular data (G. Pisanty pers. comm.) suggests that there may be cryptic species within this group that require clarification and delineation. Based on morphological knowledge, both A. unicincta and A. caspica range from the Levant to Iran, but male A. unifasciata are currently known only from the Levant. The nomenclatural and distributional situation may therefore change pending revision.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB95119DA41D8E3FD92F85C.taxon	etymology	Etymology From the Latin meaning ‘ bulbous’ in reference to the swollen genital capsule that separates this taxon from similar red-marked species of Ulandrena.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB95119DA41D8E3FD92F85C.taxon	materials_examined	Material examined Holotype IRAN • ♂; Fars, 15 km SE of Sarvestan; 1800 m a. s. l.; K. Warncke leg.; OÖLM. Paratype IRAN • 1 ♀; Fars, same collection data as for holotype; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB95119DA41D8E3FD92F85C.taxon	description	Description Female MEASUREMENTS. Body length 10 mm (Fig. 109). HEAD. 1.3 times as wide as long. Clypeus dark, weakly domed, regularly punctate with exception of narrow impunctate longitudinal midline, punctures otherwise separated by 0.5 – 1 puncture diameter; underlying surface faintly shagreened, shining (Fig. 110). Process of labrum broadly trapezoidal, apical margin with hint of emargination medially, essentially straight (Fig. 111). Face, gena, vertex, and scape with whitish hairs. Gena slightly exceeding width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Foveae dorsally occupying ½ distance between compound eye and lateral ocellus, slightly narrowed below at level of antennal insertions; foveae filled with whitish hairs. Antennae dark, A (4) 5 – 12 slightly lightened dark orange below; A 3 slightly exceeding A 4 + 5, shorter than A 4 + 5 + 6. MESOSOMA. Scutum and scutellum with dense and even punctation, punctures almost contiguous to separated by 0.5 puncture diameter; underlying surface generally smooth and shining, scutum shagreened anteriorly (Fig. 112). Pronotum without humeral angle. Mesepisternum and lateral face of propodeum with fine shagreen, weakly shining, mesepisternum with fine raised pattern of reticulation. Propodeal triangle weakly delineated laterally by fine carinae, internal surface with weak pattern of raised rugae basally, disappearing centrally, here with reticulation, dull. Propodeal triangle therefore weakly contrasting with dorsolateral faces of propodeum, these covered in fine but dense raised reticulation, dull. Mesosoma with whitish hairs, longest on mesepisternum; propodeal corbicula dorsally composed of white plumose hairs. Legs dark, apical tarsal segments lightened orange; pubescence white, tibial and femoral scopae and flocculus unicolourous white. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation dark orange, nervulus interstitial, first recurrent vein enters second submarginal cell slightly beyond its middle. METASOMA. Terga dark with extensive red maculations; red maculations form patches on T 1 basally and apically, T 2 – 3 almost entirely red-marked, T 4 basally and apically; tergal margins lightened yellowhyaline (Fig. 113). Terga T 1 – 4 weakly shagreened, shining; T 5 dull; all terga densely and uniformly punctate, punctures separated by 1 puncture diameter (Fig. 114). Tergal discs with very fine, short, and sparse whitish hairs, T 2 – 4 with weak white apical hair fringes that do not obscure underlying surface, widely interrupted. Apical fringe of T 5 and hairs flanking pygidial plate white to golden; pygidial plate rounded triangular, without surface sculpture. Male MEASUREMENTS. Body length 12 mm (Fig. 115). HEAD. 1.2 times as wide as long. Clypeus domed, entirely yellow-marked, yellow colouration extending onto lower paraocular areas; clypeus evenly punctate, punctures separated by 1 puncture diameter; underlying surface weakly shining (Fig. 116). Process of labrum rounded rectangular, broadly emarginate medially. Lower face and gena with white hairs, becoming golden-brown on vertex, scape, and frons. Gena exceeding width of compound eye; ocelloccipital distance 1 ½ times diameter of lateral ocellus. Antennae dark, A 4 – 13 slightly lightened dark brown below; A 3 slightly exceeding A 4 + 5, shorter than A 4 + 5 + 6. MESOSOMA. Mesosoma structurally as in female. Mesepisternum and propodeum with moderately thick long whitish hairs, becoming golden-brown dorsally on scutum and scutellum. Legs dark, apical tarsal segments lightened orange; pubescence whitish to brownish. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange; nervulus very slightly antefurcal, first recurrent vein enters second submarginal cell clearly beyond its middle. METASOMA. Terga dark, red maculations cover entirety of T 2 – 3, apical margins of T 1 and T 4; tergal margins otherwise lightened yellow-hyaline (Fig. 117). Tergal discs weakly shagreened, strongly shining; terga densely and evenly punctate, punctures separated by 1 – 1.5 puncture diameters. Tergal discs with fine golden pubescence, hairs longest on T 1, T 2 – 4 with complete apical hairbands composed of golden hairs, weakest on T 2, becoming progressively thicker to T 4, here obscuring underlying surface. S 8 short, triangular, ventral surface covered by short golden hairs that obscure underlying shape. Genital capsule large, gonocoxae with inner margin evenly rounded, lacking apical teeth (Fig. 118); in lateral view gonocoxae forming rounded 90 ° angle (Fig. 119). Gonostyli with apical blades flattened, sharply right-angled basally, and produced into ventrally projecting points. Penis valves mediolaterally with short hyaline extensions, apically produced into long narrow projection, very subtly bifurcate apically (Fig. 120).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFB95119DA41D8E3FD92F85C.taxon	distribution	Distribution Southern Iran (Fars Province).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	description	urn: lsid: zoobank. org: act: 334 B 5201 - 2518 - 44 BE- 9 F 05 - 18 D 14 CB 63906 Figs 121 – 126	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	diagnosis	Diagnosis Andrena hosseiniiae sp. nov. is similar to the previously described species of Planiandrena, but it cannot currently be placed there with confidence. It appears similar because the clypeus is domed and shining (Fig. 122), the foveae are dorsally narrow (though slightly constricted and separated from the inner margin of the compound eye medially), and there is a faintly raised longitudinal ridge centrally on the pygidial plate. However, it is smaller, the process of the labrum is not as wide and is generally more rounded (Fig. 123), the integument of the frons and paraocular areas have hints of metallic green reflections, the propodeal triangle is constricted medially, extremely narrow, and slightly depressed, therefore slightly ‘ T’ shaped (Fig. 125), and the hind tibial spur is straight, not broadened submedially. In the absence of male material, no subgenus is currently assigned. Andrena hosseiniiae can be recognised and separated from any Middle Eastern Andrena by this unusual combination of characters, specifically the domed and shining clypeus along with the unusual, medially constricted ‘ T’ shaped propodeal triangle.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	etymology	Etymology Named after Tahereh Hosseinii, who led much of the recent work in Iranian bee collection that is presented here.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	materials_examined	Material examined Holotype IRAN • ♀; Yasouj, Doposhteh, Dasht-e Rum; 30.589 ° N, 51.517 ° E; 2091 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; OÖLM. Paratypes IRAN • 1 ♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; OÖLM • 1 ♀; same collection data as for preceding; TJWC • 1 ♀; same collection data as for preceding; ICPI • 1 ♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 31 Mar. 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	description	Description Female MEASUREMENTS. Body length 7 – 7.5 mm (Fig. 121). HEAD. 1.1 times as wide as long. Clypeus dark, domed, with few irregular punctures, punctures laterally separated by 1 – 2 puncture diameters, very sparse centrally and apically, almost impunctate (Fig. 122). Clypeus surface basally and laterally with fine, slightly raised microreticulation, forming latitudinal wrinkles; remainder of surface with very weak shagreen, strongly shining. Process of labrum trapezoidal, twice as broad as long, corners rounded, apical margin very faintly emarginate (Fig. 123). Face, gena, vertex, and scape with sparse whitish hairs, not exceeding length of scape. Integument of frons and paraocular area with slight metallic green reflections. Gena equalling width of compound eye; ocelloccipital distance small, just over ½ diameter of lateral ocellus. Foveae dorsally narrow, occupying ⅓ of space between compound eye and lateral ocellus, medially narrowed, here separated from inner margin of compound eye by a distance subequal to its medial diameter; fovea returning to original width and inner margin of compound eye at level of antennal insertions; fovea filled with dark brown hairs. Antennae dark, A 3 = A 4 + 5 + 6, A 4 sub-square, slightly wider than long. MESOSOMA. Scutum with fine granular microreticulation, weakly shining, surface with obscure shallow punctures, punctures separated by 1 – 4 puncture diameters (Fig. 124). Scutellum strongly contrasting, shagreen limited to lateral areas, smooth and shining over majority of area, with scattered obscure punctures. Pronotum with hint of humeral angle, inconspicuous. Mesepisternum and lateral faces of propodeum with fine granular shagreenation, weakly shining. Propodeal triangle slightly depressed, internal area lower than dorsolateral faces of propodeum; basally with subtle longitudinal rugae extending onto internal surface, apically shagreened; triangle apically constricted, thus broadly ‘ T’ shaped when viewed dorsally (Fig. 125). Dorsolateral faces of propodeum with coarser granular shagreen, propodeal triangle therefore defined by change in surface sculpture. Mesosoma with long whitish hairs, longest not exceeding length of scape; propodeum dorsally and laterally with some intermixed black hairs, propodeal corbiculae dorsally with weakly plumose white hairs, internal surface with long white simple hairs. Legs uniformly dark, pubescence whitish; tibial and femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Hind tibial spur parallel-sided. Wings hyaline, stigma orange with dark brown margin, venation dark brown, nervulus antefurcal; first recurrent vein enters second submarginal cell at its middle to slightly beyond its middle. METASOMA. Terga dark, apical margin of T 1 very narrowly, of T 2 – 4 slightly more extensively lightened yellow-hyaline (Fig. 126). Terga very weakly shagreened, generally shining; terga essentially impunctate with extremely obscure and scattered fine punctures. Terga discs with scattered short white hairs, T 2 – 4 laterally with weak apical fringes of whitish hairs. Fringe of T 5 and hairs flanking pygidial plate dark brown; pygidial plate triangular with faintly raised longitudinal ridge centrally. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	discussion	Remarks All specimens were collected from Acer monspessulanum in April.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA5511ADA46D81CFE60FBA5.taxon	distribution	Distribution Southern Iran (Yasouj).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	description	urn: lsid: zoobank. org: act: 64 B 3 F 387 - 3 CCE- 4830 - 8473 - AD 5 C 20 F 83 A 21 Figs 127 – 132	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	diagnosis	Diagnosis Andrena rostamiae sp. nov. is challenging to place. It is partly similar to previously described species of Planiandrena because of the short and broad process of the labrum (Fig. 129) and the similarly sculptured though significantly wider clypeus (Fig. 128) that is wider than long (vs as wide as long or longer than wide in A. flagrans sp. nov., A. huma sp. nov., and A. sella sp. nov.). However, the foveae are substantially wider dorsally, occupying nearly half of the distance between the compound eye and the lateral ocellus (Fig. 130) (vs not occupying more than ¼ of this distance in the three comparison species). The pygidial plate is also flat and the hind tibial spur is unbroadened. It is therefore similar to A. hulae Pisanty, 2022 from northern Israel, which is also not possible to place in a subgenus at this time. Andrena rostamiae can be separated by the dark antennae (vs antennal segments 5 – 12 lightened orange below), the flattened clypeus (vs clypeus domed), the shorter ocelloccipital distance that is subequal to the diameter of a lateral ocellus (vs slightly greater than the diameter of a lateral ocellus), the smoother propodeal triangle that lacks basal rugae (vs propodeal triangle with short longitudinal basal rugae), and the impunctate scutum and scutellum (vs shallowly but distinctly punctate). Until male material is available, no firm decision on subgeneric placement is taken.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	etymology	Etymology Named after Ehlam Rostami, who collected many of the new species described in this manuscript.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	materials_examined	Material examined Holotype IRAN • ♀; Yasouj, Doposhteh, Dasht-e Rum; 30.589 ° N, 51.517 ° E; 2091 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; OÖLM. Paratypes IRAN • 2 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; OÖLM • 1 ♀; same collection data as for preceding; TJWC • 4 ♀♀; same collection data as for preceding; ICPI • 5 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; same collection data as for preceding; TJWC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	description	Description Female MEASUREMENTS. Body length 8.5 – 9.5 mm (Fig. 127). HEAD. 1.3 times as wide as long. Clypeus dark, broad, 1.5 times as broad as long, weakly flattened over majority of its area, covered with weak and irregular punctures, punctures separated by 1 – 4 puncture diameters; individual punctures slightly elongated latitudinally; poorly defined longitudinal impunctate line formed centrally (Fig. 128). Clypeus surface basally and laterally with fine, slightly raised microreticulation, forming latitudinal wrinkles; becoming weaker centrally and apically, here weakly shining. Process of labrum rectangular, short and broad, 4 times as wide as long; apical margin weakly and shallowly emarginate (Fig. 129). Face, gena, vertex, and scape with long whitish to light brownish hairs, not exceeding length of scape. Gena equalling width of compound eye; ocelloccipital distance small, just over ½ diameter of lateral ocellus. Fovea dorsally broad, occupying ½ of space between compound eye and lateral ocellus, medially slightly constricted, separated slightly from inner margin of compound eye; foveae filled with light brown hairs (Fig. 130). Antennae dark, A 3 slightly exceeding A 4 + 5, distinctly shorter than A 4 + 5 + 6. MESOSOMA. Scutum with dense granular microreticulation, dull, essentially impunctate; scutellum with similar but slightly weaker sculpturing, very weakly shining (Fig. 131). Pronotum with weak humeral angle, dorsolateral angle pronounced dorsally but becoming rounded ventrally, obscure. Mesepisternum, lateral and dorsolateral faces of propodeum with granular microreticulation, weakly shining. Propodeal triangle broad, laterally defined by narrow dark line lacking granular microreticulation, internal surface with same granular microreticulation as dorsolateral faces of propodeum, becoming coarser basally. Mesosoma covered in whitish hairs laterally and ventrally, becoming light brownish dorsally; propodeal corbiculae with light brownish weakly plumose hairs, internal surface with simple light brownish hairs. Legs dark, apical tarsal segments lightened dark brown, pubescence brownish-white to dark brown; tibial scopae light brownish, femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Hind tibial spur parallel-sided. Wings hyaline, stigma orange, venation dark orange, nervulus antefurcal; first recurrent vein enters second submarginal cell very strongly beyond its middle. METASOMA. Terga dark, marginal areas of T 2 – 4 very weakly depressed, marginal areas of T 1 – 4 lightened dark brown to narrowly yellow-hyaline apically (Fig. 132). Terga densely and finely microreticulate; microreticulation becoming weaker from mid-point of T 3 onwards; overall, terga dull to weakly shining apically. T 1 – 3 essentially impunctate, disc of T 4 with obscure scattered punctures. T 2 – 4 with weak apical hair fringes, broadly interrupted on T 2 – 3, complete on T 4. Fringe of T 5 and hairs flanking pygidial plate golden orange; pygidial plate broadly rounded apically, flattened, internal surface with dense scalelike microreticulation, strongly delineated by impunctate rim. Male Unknown.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	discussion	Remarks All specimens were collected from Acer monspessulanum in April.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA7511CDA5EDD59FE60F85C.taxon	distribution	Distribution Southern Iran (Yasouj).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9A8DCB3FED8F85C.taxon	materials_examined	Material examined IRAN • 1 ♀; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♂; Gilan Province, 20 km W of Astara; 640 m a. s. l.; 17 Jun. 2010; Mi. Halada leg.; OÖLM • 6 ♂♂, 3 ♀♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 3 ♂♂, 1 ♀; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM • 1 ♂; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9A8DCB3FED8F85C.taxon	distribution	Distribution Central and Eastern Europe through Turkey to northern Iran (Gusenleitner & Schwarz 2002; Scheuchl & Willner 2016).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9B5D82BFBA3FBA1.taxon	materials_examined	Material examined IRAN • 1 ♂; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Kurdistan Province, Paniran; 1450 m a. s. l.; 14 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9B5D82BFBA3FBA1.taxon	discussion	Remarks The taxonomic status of A. abbreviata is not completely clear, as many similar taxa have been described (Gusenleitner & Schwarz 2002). Though female specimens are quite variable, the male genital capsule is extremely similar if not identical across its distributional range. In the absence of genetic revisionary work, the broad species concept as shown in the distributional map presented by Gusenleitner & Schwarz (2002) is adopted here.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9B5D82BFBA3FBA1.taxon	distribution	Distribution South-eastern Europe, Ukraine, Russia (European part), Turkey, Cyprus, Levant, Caucasus, Iran, Kazakhstan (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9B9DD28FB01FACB.taxon	materials_examined	Material examined IRAN • 1 ♀; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA2511FD9B9DD28FB01FACB.taxon	distribution	Distribution Greece, Turkey, Armenia, Iran * (Gusenleitner & Schwarz 2002; Rasmont et al. 2017).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA3511ED982D81DFB2FFCB8.taxon	materials_examined	Material examined IRAN • 5 ♀♀; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 3 ♂♂, 8 ♀♀; Golestan, 40 km E of Minudasht NP; 750 m a. s. l.; 1 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Golestan, 70 km E of Minudasht; 1050 m a. s. l.; 12 Jun. 2010; Mi. Halada leg.; OÖLM • 2 ♂♂; Kerman Prov., Jupar [Joupar]; 1900 m a. s. l.; 1 Jun. 2010; Mi. Halada leg.; OÖLM • 2 ♂♂; Pasagrad env. [Pasargad]; 8 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA3511ED982D81DFB2FFCB8.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA3511ED9E9DA20FB23FA2A.taxon	materials_examined	Material examined JORDAN • 1 ♀; 20 km N of Karak; 1000 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 2 ♀♀; 30 km N of Tafila [At-Tafilah]; 2 May 1996; Ma. Halada leg.; OÖLM • 1 ♂; Al Karak env. [Kerak]; 6 Apr. 2013; M. Snižek leg.; OÖLM • 2 ♀♀; Kerak, 20 km N of Tafila; 1000 m a. s. l.; 19 Mar. 1988; L. Blank leg.; MSC • 2 ♂♂; Wadi-el-Mawjib; 20 Mar. 2009; V. Barták leg.; OÖLM. SYRIA • 1 ♂, 4 ♀♀; Damascus Airport; 29 Mar. 1988; L. Blank leg.; MSC • 1 ♀; Homs; 4 Jun. 1962; Kargi leg.; OÖLM • 1 ♀; 20 km E of Homs; 400 m a. s. l.; 1 Apr. 1988; S. M. Blank leg.; MSC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA3511ED9E9DA20FB23FA2A.taxon	distribution	Distribution Morocco, Portugal, Spain, Algeria, Tunisia, Italy (Sicily), Libya, Egypt, Israel and the West Bank, Jordan *, Syria *, Iran. Subspecies levantina is found in the eastern part of the range.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA35111D9AADCD2FE20FDC7.taxon	materials_examined	Material examined IRAN • 1 ♀; Kohgiluyeh and Boyer-Ahmad, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM • 17 ♂♂; Gilan Province, Tutkabon near Roodbar; 16 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Golnar, Ravar, Kerman; 23 May 2016; Z. Khajouee leg.; ICPI • 1 ♂; Kohgiluyeh and Boyer-Ahmad, Chenar; 2092 m a. s. l.; 15 Jun. 2018; F. Arannezhad leg.; ICPI • 1 ♂, 2 ♀♀; Yazd, Bafgh, Moellem square; 994 m a. s. l.; 31 May 2019; S. San leg.; ICPI • 12 ♀♀; Yazd, Dehno pajofar; 19 May 2021; S. San leg.; ICPI • 1 ♀; Yazd, Mehriz Posht e masjed, Abitaleb; 12 May 2020; S. San leg.; ICPI • 4 ♀♀; Kohgiluyeh and Boyer-Ahmad, Yasoj, Park Velayat; 1829 m a. s. l.; 20 Jun. 2018; N. Dehghan leg.; ICPI • 2 ♀♀; Yasouj, Zirtol; 20 Aug. 2016; A. Mirzapour leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA35111D9AADCD2FE20FDC7.taxon	discussion	Remarks The subgenus Taeniandrena is taxonomically complex and contains many cryptic species. It has recently undergone a substantial revision (Praz et al. 2022), with many changes for the West Palearctic fauna, including the recognition of A. afzeliella as a distinct taxon and the senior name of A. albofasciata Thomson, 1870.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFA35111D9AADCD2FE20FDC7.taxon	distribution	Distribution Confused due to historical synonymy with A. ovatula. Across Europe to Turkey, the Levant, and Iran (Praz et al. 2022).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAC5111D9BCDBB4FCC3FAD2.taxon	materials_examined	Material examined JORDAN • 1 ♀; Petra, Wadi Musa; 14 May 1995; K. Deneš leg.; OÖLM • 3 ♀♀; Wadi Rum; 4 May 1996; Mi. Halada leg.; OÖLM. SYRIA • 7 ♀♀; Ar-Raqqa, Mishirfeh; 4 Jun. 2000; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAC5111D9BCDBB4FCC3FAD2.taxon	discussion	Remarks The first currently valid taxon described from Iran, this taxon is found in arid, semi-desert habitats. Reported as new for Jordan and Syria.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAC5111D9BCDBB4FCC3FAD2.taxon	distribution	Distribution Morocco, Algeria, Tunisia, Libya, Egypt, Israel and the West Bank, Jordan *, Syria *, Iraq, Iran (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAC5111D9F5DC86FAEEF85C.taxon	materials_examined	Material examined TURKEY • 4 ♀♀; Hakkâri, Beytüşşebap; 25 May 1988; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAC5111D9F5DC86FAEEF85C.taxon	discussion	Remarks The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate the presence of A. albopicta in northern Iran. No specimens of A. albopicta could be found in the Warncke collection, but A. albopicta is present in eastern Turkey and so it is considered to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAC5111D9F5DC86FAEEF85C.taxon	distribution	Distribution Turkey, Russia (European part), Armenia, Iran (Gusenleitner & Schwarz 2002; Rasmont et al. 2017).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5110D945DA5CFD3BFAE1.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province, 10 km E of Kazerum; 1990 m a. s. l.; 23 May 2014; J. Halada leg.; OÖLM. TURKEY • 2 ♀♀; Mardin; 1000 m a. s. l.; 2 Jul. 1987; R. Hensen leg.; RMHN • 2 ♀♀; Batman, 15 km N of Midyat; 22 Jun. 1997; Ma. Halada leg.; OÖLM • 1 ♀; 40 km E of Midyat / Mardin; 900 m a. s. l.; 25 May 1983; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5110D945DA5CFD3BFAE1.taxon	distribution	Distribution Turkey * and Iran (Ariana et al. 2009 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5110D994D81DFB2FFCAA.taxon	materials_examined	Material examined IRAN • 3 ♂♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; Gilan Province, Tutkabon near Roodbar; 16 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Golestan Province, 45 km E of Minudasht; 960 m a. s. l.; 11 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Golestan, 70 km E of Minudasht; 1050 m a. s. l.; 12 Jun. 2010; OÖLM • 1 ♀; Mashhad; 1 Apr. 2010; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5110D994D81DFB2FFCAA.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5113D999DC6BFCBCFCC1.taxon	materials_examined	Material examined CRIMEA • 1 ♂, 3 ♀♀; Krim [Crimea], Karadagh [Kara Dag], Vodianja balka, Wald; 17 – 31 May 2002; Y. Budaschkin leg.; OÖLM. GEORGIA • 1 ♀; NC, N of Tbilisi, Magaroskarj env.; 19 Jun. 2014; M. Snižek leg.; OÖLM • 2 ♀♀; Tbilisi env.; 21 May 1973; Dr. Svozil leg.; OÖLM. IRAN • 1 ♀; Kohgiluyeh and Boyer-Ahmad Province, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun 2010; Mi. Halada leg.; OÖLM • 1 ♀; Esfahan Province, 40 km SE of Aligudars (Nowghan); 2250 m a. s. l.; 31 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Kerman Province, Sirač [? Sirch]; 1640 m a. s. l.; 2 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Lorestan Province, Tootmashour [Tut, Masur-e Abi]; 2000 m a. s. l.; 31 May 2010; Mi. Halada leg.; OÖLM. MONTENEGRO • 1 ♀; Durmitor Žabljak env.; 25 Jun. – 7 Jul. 1958; Bouček leg.; OÖLM. RUSSIA • 2 ♂♂, 2 ♀♀; N. Kaukasus, Umg. Pjatigorsk; 800 m a. s. l.; 25 – 30 May 2002; V. Pack leg.; OÖLM. UKRAINE • 1 ♀; Ivanofrankivskij rajon, 30 km E of Ivano-Frakivsk, Nizvev vill [Nyzhniv] Dnestr River; 7 Jun. 2016; M. Snižek leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5113D999DC6BFCBCFCC1.taxon	discussion	Remarks The presence of A. alutacea in Georgia is confirmed from female material (see doubts raised by SchmidEgger 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAD5113D999DC6BFCBCFCC1.taxon	distribution	Distribution Switzerland, Germany, Italy, Austria, Croatia, Poland, Latvia, Romania, Bulgaria, Montenegro *, North Macedonia, Greece, Turkey, Ukraine *, Crimea *, Russia, Georgia, Azerbaijan, Iran, and Turkmenistan (Schmid-Egger 2005; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5113D993DA8AFEDEFB53.taxon	materials_examined	Material examined IRAN • 1 ♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5113D993DA8AFEDEFB53.taxon	distribution	Distribution North Macedonia, Greece, Romania, Bulgaria, Turkey, Ukraine, Russia (European part), Lebanon, Iran * (Wood 2021 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5113D98DDC1BFD99F97F.taxon	materials_examined	Material examined IRAN • 2 ♂♂; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5113D98DDC1BFD99F97F.taxon	discussion	Remarks A taxon showing a distribution in both the Anti-Lebanon mountain range and the Zagros Mountains of western Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5113D98DDC1BFD99F97F.taxon	distribution	Distribution Syria and Iran * (Wood 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5112D9A9DFE0FCE6FDB9.taxon	materials_examined	Material examined IRAN • 1 ♂; Elburz, Pulour [Polour], 22 km N of di Ab Ali; 11 Jul. 1965; G. Soika and G. A. Mavromoustakis leg.; OÖLM. TURKEY • 1 ♀; Van, town walls; 28 Jun. 1993; Jiroušek leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5112D9A9DFE0FCE6FDB9.taxon	discussion	Remarks Described from a single female collected in Djulfa (now Julfa, Nakhchivan Autonomous Republic, Azerbaijan; Popov 1940). This town is directly on the border with Iran, and so the presence of this species in Iran is not unexpected.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAE5112D9A9DFE0FCE6FDB9.taxon	distribution	Distribution Turkey, Azerbaijan, Iran * (Popov 1940; Scheuchl 2010).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAF5169D9FBDB21FAF4FC71.taxon	materials_examined	Material examined Lectotypes CAUCASUS • ♀ (lectotype of Andrena assimilis); Caucasus; ZMHB (illustrated Figs 133 – 138). FRANCE • ♀ (lectotype of Andrena gallica); Mont-de-Marsan; MNHN (illustrated Le Divelec 2021). Syntypes ITALY • ♀ (syntype of Andrena gallica alpicola); Como-S. Cadenabbia; 1 – 31 Mar. 1903; ZMHB (illustrated Figs 139 – 140). IBERIA • ♀ (syntype of Andrena gallica iberica); Hispania; 189 [no further information]; ZHMB (illustrated Figs 141 – 142). CRIMEA • ♀ (syntype of Andrena gallica taurica); Krim [Crimea]; Nordmann leg.; Coll. Gerst. [Gerstaecker]; ZMHB (illustrated Figs 143 – 144). GERMANY • ♀ (syntype of Andrena gallica fulvitegularis); Mittenwalde, Machn. Weinbg.; 20 Jul. 1921; Hedicke leg.; ZMHB • ♀ [syntype of A. gallica fulvitegularis]; Gross Machnow; 21 Jul 1918; ZMHB. Other material ALBANIA • 1 ♀; Bizë, b. Shengjergji [Shëngjergj]; 1400 – 1500 m a. s. l.; 15 Jul. 1961; K. Warncke det. [as assimilis gallica]; OÖLM. AUSTRIA • 1 ♀; Oberweiden; 15 Apr. 1936; K. K. Warncke det. [as assimilis gallica]; R. Schmidt leg.; OÖLM. AZERBAIJAN • 1 ♀; Helendorff [renamed Khanlar, now Göygöl]; 1886; K. Warncke det. [as assimilis s. str.]; OÖLM. CZECH REPUBLIC • 1 ♀; Hovorany; 8 Aug. 1941; K. Warncke det. [as assimilis gallica]; V. Zavdil leg.; OÖLM. FRANCE • 3 ♀♀; Aude, Servies-en-Val; 5 Aug. 1970; R. T. Simon Thomas leg.; RMNH • 2 ♀♀; Montpellier; ZMHB • 1 ♀; Pyr. Or., Mont Louis; 1600 m a. s. l.; 20 Aug. 1957; M. C. and G. Kreuseman; RMNH. GERMANY • 6 ♂♂, 6 ♀♀; Niemegk, Fläming; 26 Jul. 1953; E. Heidenreich leg.; ZMHB. GREECE • 1 ♂, 1 ♀; Saloniki; 1 – 30 Apr. 1932; K. Warncke det.; Paduschin-Kattinger leg.; ZMHB. IRAN • 1 ♂; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Elburs [Alborz], Pelur [Polour]; ca 2000 m a. s. l.; Wagner leg.; ZMHB • 1 ♀; Elburs [Alborz], Rehne, Demavend [Damavand]; 2700 – 2600 m a. s. l.; Wagner leg.; ZMHB • 1 ♀; Elburz, 30 km di. of Ab Ali; 9 Jul. 1965; K. Warncke det. [as assimilis s. str.]; G. Soika and G. A. Mavromoustakis leg.; OÖLM • 1 ♀; Gilan Province, 15 km SE of Tutkabon; 1100 m a. s. l.; 9 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Golestan, 70 km E of Minudasht; 1050 m a. s. l.; 12 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Kerman Province, 20 km E of Ghobira; 1780 m a. s. l.; 5 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Seman Province, 10 km W of Damghan; 1200 m a. s. l.; 10 Jun. 2010; Mi. Halada leg.; OÖLM. KAZAKHSTAN • 1 ♀; Alma-Ata (Medeo); 12 Jul. 1977; K. Warncke det. [as assimilis s. str.]; K. Bleyl leg.; OÖLM. KYRGYZSTAN • 1 ♀; Alarca [Ala Archa National Park]; 2500 m a. s. l.; 27 May 1974; K. Warncke det. [as assimilis s. str.]; Rataj leg.; OÖLM • 1 ♀; Przewalsk [Karakol], Tian-Schan; K. Warncke det. [as assimilis gallica]; OÖLM. MOROCCO • 1 ♀; Dait Achlaf [Lac Hachlaf]; 27 Apr. 1987; K. Warncke det. [as assimilis barnei Cockerell, 1931]; M. Kraus leg.; OÖLM • 1 ♂; Ifrane; 22 Jul. 1932; K. Warncke det. [as assimilis barnei]; Ad. Nadig leg.; OÖLM. PORTUGAL • 1 ♂, 1 ♀; Estremadura, Apostiça; 28 Feb 1990; B. Souça leg.; MNHNC • 1 ♀; Baixo Alentejo, Vila Nova de Milfontes; 14 Jul. 1971; E. Scheuchl det.; MNHNC • 2 ♀♀; Carrapateira; 3 Apr. 2015; T. J. Wood leg.; TJWC. SPAIN • 1 ♀; Aragon, Albarracín; 21 Jul. 1924; K. Warncke det. [as assimilis gallica]; Zerny leg.; OÖLM • 1 ♀; Setcases (Girona); 1500 m a. s. l.; F. J. Ortiz-Sánchez leg.; OSCA (illustrated Fig. 145) • 3 ♀♀; Burgos, Hornilloyuso; 24 Apr. 1984; R. Leys leg.; RMNH • 2 ♀♀; Caceres, Villasbuenas de Gata; 1 Jul. 1988; M. Schwarz leg.; MSC • 2 ♀♀; Puerto de Somosierra (Madrid, Sierra de Guadarama); 1355 m a. s. l.; F. J. Ortiz-Sánchez leg.; OSCA (illustrated Fig. 146) • 4 ♀♀; Granada, Sierra Nevada, Trevélez to Refugio La Campiñuela; 1700 – 2400 m a. s. l.; T. J. Wood leg.; TJWC (illustrated Fig. 147) • 1 ♀; Madrid, Cercedilla; 21 Apr. 1973; J. A. W. Lucas leg.; RMNH • 1 ♂, 2 ♀♀; Ávila, 1 km S of Navacepeda de Tormes; 15 Jul. 2021; T. J. Wood leg.; TJWC (illustrated Fig. 148) • 1 ♀; Sierra de Guadarrama; 18 Aug. 1912; K. Warncke det. [as assimilis s. str.]; J. M. Dusmet y Alonso leg.; OÖLM • 1 ♀; Soria; 26 May 1965; K. Warncke det. [as assimilis gallica]; W. Linsenmaier leg.; OÖLM. RUSSIA • 1 ♀; Bijsk [Biysk]; 27 May 1921; ZMHB • 1 ♂ [syntype of A. gallica fulvitegularis]; Irkutsk; 28 Apr. – 11 May 1913; O. Hesse leg.; ZMHB • 1 ♀ [syntype of A. gallica fulvitegularis]; Irkutsk; 14 – 27 May 1913; O. Hesse leg.; ZMHB. SERBIA • 1 ♀; Fejértelep [Šušara]; K. Warncke det. [as assimilis s. str.]; OÖLM. SWITZERLAND • 2 ♀♀ [paratypes of A. gallica alpicola]; Sierre; 15 Jul. 1884; H. Friese leg.; ZMHB. TURKEY • 1 ♀; Çıldır, Kars; 2000 m a. s. l.; 19 Aug. 1991; K. Warncke det. [as assimilis s. str.]; OÖLM. UKRAINE • 1 ♂; Desna Nadinovka [Nadynivka]; 29 – 30 Apr. 2003; Y. Budaschkin leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAF5169D9FBDB21FAF4FC71.taxon	discussion	Remarks The taxonomic history of the two names A. assimilis and A. gallica is long and confused (SchmidEgger & Scheuchl 1997; Gusenleitner & Schwarz 2002), in part due to the unclear location of the type material of A. gallica, which was recently rediscovered (Le Divelec 2021). The putative criteria used to separate material was in the colour of the wing venation and the degree of infuscation of the wing itself (Osytshnjuk et al. 2008; Motyka et al. 2016). However, this character is highly variable. Examination of the type of A. assimilis in Berlin shows that, though the wings are somewhat infuscate, the veins are not entirely dark, with intermixed dark and light veins, particularly the basal vein (Fig. 137). In contrast, the type of A. gallica (and more broadly all examined specimens from southern France) has hyaline wings with almost uniformly golden veins (see illustrations in Le Divelec 2021). Problems with the proposed two-species classification can most easily be seen in Iberia. In the Pyrenees, the light form is present (e. g., A. gallica iberica, Fig. 140), extending across the northern parts of Spain (Fig. 145) and down the Atlantic coast as far as the Algarve in southern Portugal. However, in central and southern Spain forms with dark infuscate wings can be found that show both light and dark veins in the same wing (Madrid, Fig. 146). In the Sierra Nevada (Granada, at 2100 m a. s. l., Fig. 147) and the Sistema Central (Ávila, 1300 m a. s. l., Fig. 148) forms with extremely dark wings and wing venation can also be found. This increased melanism is even more apparent in Morocco, where populations in the Atlas Mountains (A. assimilis barnei) are entirely dark, including all of their body hair. Warncke treated A. gallica as a subspecies of A. assimilis, and in his distribution map (Gusenleitner & Schwarz 2002) he presented a clean picture of three separated populations: A. assimilis s. str. in the southern Balkans, Turkey, and the Caucasus; A. assimilis gallica in continental Europe and Russia; and A. assimilis barnei in Morocco. However, Warncke was inconsistent with his determination of material from this group, as specimens listed here from Azerbaijan, Iran, Kazakhstan, Kyrgyzstan, and Serbia with golden wing venation were determined as A. assimilis s. str. He also determined specimens from Spain as both assimilis s. str. and assimilis gallica, contradicting the clean picture presented in his distribution map. Likewise, Osytshnjuk et al. (2008) argue that both A. assimilis and A. gallica are present from Eastern Europe eastwards, Western and Central Europe containing only A. gallica. The situation is reminiscent of the taxonomic confusion present in Bombus (Thoracobombus) concerning B. laesus Morawitz, 1875 and B. mocsaryi Kriechbaumer, 1877, which vary in hair colour and are present in sympatry across much of Eastern Europe. Though varying in their hair colour, these taxa were recently found to belong to the same broad taxon (Brasero et al. 2021). Due to this extensive variation within A. assimilis, including in the type specimen itself, which does not unambiguously show uniformly dark wing venation, A. gallica is synonymised under A. assimilis. This includes the subspecies described by Bischoff (1922, Figs 139 – 144), which were previously synonymised under A. assimilis gallica by Warncke (1967) but were listed under A. gallica by Gusenleitner & Schwarz (2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFAF5169D9FBDB21FAF4FC71.taxon	distribution	Distribution Andrena assimilis is distributed across the West and Central Palearctic, from Morocco (absent from the rest of North Africa and the Levant) across continental Europe to Turkey, northern Iran *, Russia (European part to Eastern Siberia), and Central Asia (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a). It is unclear why Warncke did not include the Soika and Mavromoustakis record from northern Iran in his distribution maps for this species, since he clearly examined and determined it.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD45169D993DAFBFABDF9FE.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD45169D993DAFBFABDF9FE.taxon	distribution	Distribution Greece, Turkey, Cyprus, Israel and the West Bank, Georgia, Iran * (Schwenninger et al. 2015).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD45169D98FDF67FA83F85C.taxon	materials_examined	Material examined IRAN • 1 ♀; West Azerbaijan Province, Serou, NW Orumiye; 1800 m a. s. l.; S. Kadlec leg.; OÖLM • 1 ♀; Kohgiluyeh and Boyer-Ahmad, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Gilan, 5 km E of Rudbar; 400 m a. s. l.; 8 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD45169D98FDF67FA83F85C.taxon	distribution	Distribution Hungary, Romania, North Macedonia, Greece, Turkey, the Caucasus, Syria, Iran * (Wood 2021 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD55168D9BDDD24FAA0F939.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province, 10 km E of Kazerum; 1990 m a. s. l.; 23 May 2014; J. Halada leg.; OÖLM • 1 ♂, 1 ♀; Fars Province, Dasht Arjan; 2040 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD55168D9BDDD24FAA0F939.taxon	discussion	Remarks The subgenus Euandrena is highly taxonomically challenging, and requires much work. The presence of A. bicolor in Iran should be confirmed by barcoding, as A. bicolor is both a variable species, and also a species complex that contains cryptic taxa. A deep revision is required.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD55168D9BDDD24FAA0F939.taxon	distribution	Distribution Probably West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD55168D99CDB32FC8BFBBE.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province, Dasht Arjan; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ilam Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM • 3 ♀♀; Yasouj, Deli Bajak, Sepidar; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; ICPI. SYRIA • 1 ♂; Bloudan; 16 May 1995; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD55168D99CDB32FC8BFBBE.taxon	distribution	Distribution Turkey, Syria *, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD55168D9F4D81CFB8AFE4B.taxon	distribution	Distribution Azerbaijan and Iran (Gusenleitner & Schwarz 2002; Allahverdi et al. 2016).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD5516BD9D2DFA0FAA6FCC5.taxon	materials_examined	Material examined IRAN • 7 ♂♂, 21 ♀♀; Golestan, 70 km E of Minudasht; 1050 m a. s. l.; 12 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Kerman Province, 8 km N of Bardsir; 2050 m a. s. l.; 6 Jun. 2010; Mi. Halada leg.; OÖLM • 9 ♀♀; Kerman Province, Deh Bakri, Gebal Barez Mts; 1640 m a. s. l.; 3 Jun. 2010; Mi. Halada leg.; OÖLM • 2 ♂♂, 1 ♀; Khorasan Province, 10 km W of Raz, Koppe Dag [Kopet Dag]; 1200 m a. s. l.; 27 May 2014; J. Halada leg.; OÖLM • 2 ♂♂; Yazd, Abarkooh, Faraghe; 1698 m a. s. l.; 6 Mar. 2021; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD5516BD9D2DFA0FAA6FCC5.taxon	discussion	Remarks The taxonomic status of A. bimaculata and the many taxa that have been described and synonymised with it is highly complicated. Gusenleitner & Schwarz (2002) give an overview of this subject, but what is clear is that although a number of coloured female forms exist (e. g., morawitzi Thomson, 1872, atrorubicata Dours, 1872, aulica, bluethgeni Stoeckhert, 1930), the male genitalia associated with these forms are extremely similar, leading many authors to group taxa together into a broad A. bimaculata. This problem of morphological variability exists in many taxa found within the subgenus Plastandrena, and detailed molecular revision encompassing a wide geographic range is necessary to fully resolve their statuses.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD5516BD9D2DFA0FAA6FCC5.taxon	distribution	Distribution Unclear due to ongoing taxonomic confusion. Andrena bimaculata s. lat. is found throughout the West and Central Palearctic to Mongolia (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD6516BD99DDA8CFD0DFA01.taxon	materials_examined	Material examined IRAN • 1 ♂, 1 ♀; Elburz Mountains, 60 km E of Minudasht; 26 May 2007; O. Sauša leg.; OÖLM • 1 ♂; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Kirmanschahad, 70 km SE of Shahabad; 1200 m a. s. l.; 11 May 1976; Holzschuh leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD6516BD99DDA8CFD0DFA01.taxon	distribution	Distribution Italy and Austria eastwards to Turkey, the Caucasus, the Levant, Iran * (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD6516BD990DCC8FB6AF85C.taxon	materials_examined	Material examined IRAN • 2 ♂♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; Kohgiluyeh and Boyer-Ahmad Province, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM. JORDAN • 1 ♂, 1 ♀; Ajlun, S of Anjara; 27 Apr. 2002; M. Snižek leg.; TJWC • 1 ♂, 2 ♀♀; North Shuna; 29 Apr. 1996; Ma. Halada leg.; OÖLM. SYRIA • 1 ♀; 60 km S of Damascus, Khahab; 14 May 1996; Ma. Halada leg.; OÖLM • 1 ♀; Anata, 50 km SE of Suwayda; 20 – 21 May 1996; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD6516BD990DCC8FB6AF85C.taxon	distribution	Distribution Turkey, Israel and the West Bank, Jordan *, Syria *, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD98DD8D4FB14FD53.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD98DD8D4FB14FD53.taxon	distribution	Distribution Southern Europe to Turkey and the Near East, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD9ADD81DFF44FE23.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD993DADCFAFDF9BE.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province, Komehr; 2380 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM. JORDAN • 1 ♀; 20 km N of Karak; 1000 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 1 ♂, 3 ♀♀; 20 km NW of Amman; 420 m a. s. l.; 5 May 2006; K. Deneš leg.; OÖLM • 1 ♀; 20 km SW of Madaba; 1 – 31 May 2006; F. Kantner leg.; OÖLM • 1 ♂; 20 km SW of Madaba; 26 May 2006; Z. Kejval leg.; OÖLM • 1 ♀; 30 km NW of Aljun; 600 m a. s. l.; 29 Apr. 2006; K. Deneš leg.; OÖLM • 62 ♂♂, 6 ♀♀; North Shuna; 29 – 30 Apr. 1996; Ma. Halada leg.; OÖLM. SYRIA • 4 ♀♀; 30 km N of Dara, Nawa; 18 May 1996; Ma. Halada leg.; OÖLM • 1 ♀; Hasake [Al Hasakah]; 8 May 1996; Ma. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD993DADCFAFDF9BE.taxon	distribution	Distribution Turkey, Israel and the West Bank, Jordan *, Syria *, Armenia, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD9BBDA04FF44FC2B.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD9AFDF27FBA0F85C.taxon	materials_examined	Material examined ARMENIA • 1 ♂; Achtarak, near Mt. Arailer; 1 – 30 Jun. 2003; M. Můčka leg.; OÖLM. IRAN • 1 ♀; Kohgiluyeh and Boyer-Ahmad Province, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun 2010; Mi. Halada leg.; OÖLM • 1 ♂, 1 ♀; Fars Province, Dasht Arjan; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 4 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD7516AD9AFDF27FBA0F85C.taxon	distribution	Distribution Greece, Turkey, Syria, Armenia *, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD0516DD99FDC49FA98F9ED.taxon	distribution	Distribution Golan Heights, Lebanon, Turkey, Iran (Dubitzky 2006; Wood et al. 2020 a; Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD0516DD9AED81DFC20FA82.taxon	materials_examined	Material examined IRAN • 1 ♂, 3 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Kohgiluyeh and Boyer, Ahmad Prov., Likak, Gach Bolland; 1640 m a. s. l.; 7 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Kurdistan: Sonnate [Sonnat-e Olya]; 11 May 1976; Holzschuh leg.; OÖLM • 1 ♂, 7 ♀♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM. JORDAN • 2 ♀♀; 20 km S of North Shuna, Tall Al Arbatin; 19 Apr. 1996; Ma. Halada leg.; OÖLM • 1 ♂; Aljoun; 28 Apr. 2012; M. Kafka leg.; OÖLM. SYRIA • 1 ♂; Latakia, Saladinburg [Citadel of Saladin]; 900 m a. s. l.; 4 Apr. 1988; L. Blank leg.; OÖLM • 1 ♂; Tartus, 10 km E of Safita; 300 m a. s. l.; 3 Apr. 1988; L. Blank leg; TJWC. TURKEY • 1 ♂; Hakkâri, 10 km W of Uludere; 1000 m a. s. l.; 4 Jun. 1980; M. Schwarz leg.; OÖLM • 6 ♀♀; Uludere, Hakkâri; 1000 m a. s. l.; 16 Jun 1981; K. Warncke and M. Kraus leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD0516DD9AED81DFC20FA82.taxon	discussion	Remarks Molecular revision of the polita group of Ulandrena is required, particularly the red-marked species that includes A. caspica, as additional cryptic taxa may be present.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD0516DD9AED81DFC20FA82.taxon	distribution	Distribution Unclear due to taxonomic confusion, but probably Greece, Turkey, Israel and the West Bank, Jordan *, Syria *, the Caucasus, Russia (European part, Derbent), Iran *.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD0516CD9FCDF95FB2BFDB0.taxon	discussion	Remarks The exact identity of this species in Iran is not clear, as previous authors have not distinguished between A. barbareae Panzer, 1805 and A. danuvia Stoeckhert, 1950. This was the case for Warncke, and his distribution maps include dots for A. cineraria in northern Iran (Gusenleitner & Schwarz 2002). In Turkey, the only taxon is A. danuvia (Hazir et al. 2014; Scheuchl & Willner 2016), and this therefore may be the taxon present in Iran, but this cannot be verified without the inspection of specimens. Of the specimens collected by Baker, some were on the wing in July and are therefore clearly bivoltine, further suggesting that they are not A. cineraria in a narrow sense.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD0516CD9FCDF95FB2BFDB0.taxon	distribution	Distribution In a broad sense, Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516CD9F8DB39FE68FBC6.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province, Dasht Arjan; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM • 3 ♀♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM. JORDAN • 1 ♀; 15 km W of Jerash, Dibbin; 2 May 2006; K. Deneš leg.; OÖLM • 1 ♀; Aljoun; 6 May 2012; M. Kafka leg.; OÖLM • 1 ♀; Jerash, Dibbeen; 4 May 2012; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516CD9F8DB39FE68FBC6.taxon	distribution	Distribution Greece, North Macedonia, Romania, Bulgaria, Turkey, Cyprus, Russia (European part, Derbent), Israel and the West Bank, Jordan *, Iran *, Afghanistan (Warncke 1974 c; Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516CD9A0DD8EFD3FF95D.taxon	materials_examined	Material examined IRAN • 2 ♀♀; Elburz, Ov Zanak-Ski, 1 km N of Ab Ali; 2300 m a. s. l.; 11 Jul. 1965; K. Warncke det.; G. Soika and G. A. Mavromoustakis leg.; OÖLM • 1 ♀; Elburz, Pulour [Polour], 22 km N of di Ab Ali; 13 Jul. 1965; K. Warncke det.; G. Soika and G. A. Mavromoustakis leg.; OÖLM • 1 ♀; Elburz, Stausee [Resevoir] W Karaj, 1700 m a. s. l.; K. Warncke det.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516CD9A0DD8EFD3FF95D.taxon	discussion	Remarks Label data of these specimens were never published, but dots are present in the distribution map for A. clusia drawn by Warncke (Gusenleitner & Schwarz 2002). This species is therefore not reported as new for Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516CD9A0DD8EFD3FF95D.taxon	distribution	Distribution North Macedonia, Turkey, Armenia, Azerbaijan, Iran, Turkmenistan, Uzbekistan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516FD9FFDE06FAB3FDF1.taxon	materials_examined	Material examined IRAN • 1 ♂; Bakhtiyari Province, 20 km SE of Lordegan; 31 May 2014; J. Halada leg.; OÖLM • 1 ♂; Boyer-A. o. Kohg Prov., Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Lorestan Province, Tootmashour [Tut, Masur-e Abi]; 2000 m a. s. l.; 31 May 2010; Mi. Halada leg.; OÖLM • 1 ♂; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD1516FD9FFDE06FAB3FDF1.taxon	distribution	Distribution Mediterranean basin to Central Asia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD2516FD92DDFC2FD4EF85C.taxon	distribution	Distribution Turkey and Iran (Gusenleitner & Schwarz 2000).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD2516FD993DB66FC5EFB9D.taxon	materials_examined	Material examined IRAN • 1 ♀; Elburz Mountains, 60 km E of Minudasht; 26 May 2007; O. Sauša leg.; OÖLM • 1 ♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD2516FD993DB66FC5EFB9D.taxon	distribution	Distribution Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD2516FD986DD43FAA5F91D.taxon	materials_examined	Other material SYRIA • 1 ♂, 1 ♀; Aleppo, 500 m, Simeons-Kloster; 19 Apr. 1988; L. Blank leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD2516FD986DD43FAA5F91D.taxon	discussion	Remarks See Pisanty et al. (2022 a) for discussion of similar species.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD2516FD986DD43FAA5F91D.taxon	distribution	Distribution Syria *, Turkey, Azerbaijan, Iran, Afghanistan, Tajikistan (Schuberth et al. 2001; Wood 2021 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD3516ED99AD81DFE68FD05.taxon	materials_examined	Material examined IRAN • 1 ♂; West Azerbaijan, Bazargan; 1380 m a. s. l.; 11 May 1978; M. Kraus leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD3516ED99AD81DFE68FD05.taxon	distribution	Distribution Eastern Europe through Turkey and the Caucasus to Central Asia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD3516ED984DD04FC94F9D2.taxon	materials_examined	Material examined CYPRUS • 1 ♀; Kykkos; 800 m a. s. l.; 11 May 2014; M. Kafka leg.; OÖLM. IRAN • 7 ♀♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM. JORDAN • 1 ♀; 10 km W of Jarash; 1 May 1996; Ma. Halada leg.; OÖLM • 2 ♀♀; 30 km NW of Aljun; 600 m a. s. l.; 29 Apr. 2006; K. Deneš leg.; OÖLM • 2 ♀♀; Jarash env.; 1 May 1996; Ma. Halada leg.; OÖLM • 1 ♀; N. Shuna env.; 30 Apr. 1996; Mi. Halada leg.; OÖLM • 1 ♀; North Shuna; 20 – 22 Apr. 1996; Ma. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD3516ED984DD04FC94F9D2.taxon	distribution	Distribution North Macedonia, Greece, Turkey, Cyprus *, Israel and the West Bank, Jordan *, Syria, Iran * (Gusenleitner & Schwarz 2002; Pisanty et al. 2018).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD3516ED9F3DBCFFCBDFC51.taxon	materials_examined	Material examined IRAN • 1 ♂; Yasouj, Abshar; 2 Mar. 2011; H. Elahi and M. Mesavian leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD3516ED9F3DBCFFCBDFC51.taxon	distribution	Distribution Turkey and Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD35161D98BDF85FEC5FEC1.taxon	materials_examined	Literature records IRAN • 1 ♂; Mazanderan Province, Kujur, 2 km SE of Hassanabad; 500 m a. s. l.; 29 Apr. 1967; D. Baker leg.; SEMC; via GBIF occurrence 784990468. Material examined IRAN • 16 ♂♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD35161D98BDF85FEC5FEC1.taxon	distribution	Distribution Turkey, Israel and the West Bank, Syria, Russia (European part), Armenia, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDC5161D9B9DA6CFA9EF91C.taxon	materials_examined	Material examined IRAN • 1 ♂; Ali Sadr; 14 May 1999; K. Deneš leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM • 1 ♂; Luristan [Lorestan], 30 km SE of Khoramabad; 2000 m a. s. l.; 17 May 1976; C. Holzschuh leg.; OÖLM • 1 ♂; Luristan [Lorestan], 50 km SE of Khoramabad; 1700 m a. s. l.; 13 – 15 May 1976; C. Holzschuh leg.; OÖLM • 1 ♀; Tehran, NW Qazvin; 1500 m a. s. l.; 19 May 1976; C. Holzschuh leg.; OÖLM. JORDAN • 8 ♀♀; 20 km N of Karak; 1000 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 1 ♀; North Shuna env.; 29 – 30 Apr. 1996; Mi. Halada leg.; OÖLM • 1 ♀; S of At Tafila; 27 Mar. 2013; M. Snižek leg.; OÖLM. SYRIA • 1 ♀; 10 km SE of Suwayda, Kafr; 19 May 1996; Mi. Halada leg.; OÖLM • 3 ♂♂, 10 ♀♀; Hasake [Al Hasakah]; 8 May 1996; Ma. Halada leg.; OÖLM • 18 ♀♀; Salkhad env.; 6 May 1996; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDC5161D9B9DA6CFA9EF91C.taxon	distribution	Distribution Turkey, Cyprus, Israel and the West Bank, Jordan *, Syria *, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDC5161D984D889FD23FCE4.taxon	materials_examined	Material examined IRAN • 1 ♀; Kameran val., Kameran vil.; 7 Jun. 2013; V. Major leg.; OÖLM • 1 ♀; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDC5161D984D889FD23FCE4.taxon	distribution	Distribution Crimea, Russia (European part, Urals), Turkey, the Caucasus, Iran, Central Asia, Mongolia (Osytshnjuk et al. 2008; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDC5160D9A1DFDBFEE8FE4F.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM • 1 ♂, 2 ♀♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Kurdistan, 20 km N of Samandaj; 1500 m a. s. l.; 11 May 1976; Holzschuh leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDC5160D9A1DFDBFEE8FE4F.taxon	distribution	Distribution Turkey, Israel and the West Bank, Lebanon, Jordan, Syria, Iran * (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5160D9AADDAFFF44FAF0.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5160D9AFDB36FE45FB20.taxon	materials_examined	Material examined IRAN • 1 ♀; Nafch, Share kord [Chaharmahal and Bakhtiari]; 2 Aug. 2016; S. Nazari leg.; ICPI. TAJIKISTAN • 1 ♀; Dusambe [Dushanbe] env.; 14 Jun. 1966; K. Deneš leg.; OÖLM. UZBEKISTAN • 2 ♀♀; 35 km S of Sammarkand, Aman Kutan; 39.312 ° N, 66.947 ° E; 30 Jun. 1981; M. Kocourek leg.; OÖLM • 11 ♀♀; same collection data as for preceding; MSC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5160D9AFDB36FE45FB20.taxon	distribution	Distribution Not completely clear because of the recent separation of A. flavilabris Schenck, 1874 (see Manderey et al. 2008). Probably broadly West Palearctic, excluding desert regions (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008), Iran, Tajikistan *, Uzbekistan *. Cited from “ Central Asia ” by Schönitzer et al. (1995) but without details.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5160D9F3DC7FFDB7F8A2.taxon	materials_examined	Material examined IRAN • 1 ♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; TJWC • 1 ♂; Yazd, Khezr abad Zorband; 1844 m a. s. l.; 21 Mar. 2021; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5160D9F3DC7FFDB7F8A2.taxon	distribution	Distribution Turkey, Russia (European part), the Caucasus, Iran, Kazakhstan (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5163D989DE29FB70FEC6.taxon	materials_examined	Literature records Allahverdi et al. (2016); Ascher & Pickering (2021): Mazandaran, Kujur, 2 km SE of Hassanabad; 500 m a. s. l.; 30 Mar. 1966; D. Baker leg.; SEMC; via GBIF occurrence 784990193.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDD5163D989DE29FB70FEC6.taxon	distribution	Distribution Unclear because of confusion with A. mizorhina Warncke, 1975 which is found in the Levant (Pisanty et al. 2022 a). Probably restricted to Turkey, the Caucasus, Russia (European part), Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9F3DF56FC0BF85C.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9F3DF56FC0BF85C.taxon	discussion	Remarks Confusion exists between this taxon and A. statusa Gusenleitner, 1998, which may be conspecific (see Pisanty et al. 2018). Further study is required.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9F3DF56FC0BF85C.taxon	distribution	Distribution Israel, Turkey and Iran * (Gusenleitner 1998; Pisanty et al. 2018).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9B2DA75FB28FBE1.taxon	distribution	Distribution Greece, Syria, Turkey, Iran (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9B7DC81FF44F9A9.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9ECDD6FFD0EFADB.taxon	materials_examined	Material examined IRAN • 1 ♀; Tehran Province, Samgh Abad; 1900 m a. s. l.; 16 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9ECDD6FFD0EFADB.taxon	distribution	Distribution Turkey and Iran * (Scheuchl & Hazir 2012).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9BED883FD63FC88.taxon	materials_examined	Material examined IRAN • 1 ♂; Mazandaran Province, Kojuk; 36 ° 23 N, 51 ° 40 E; 7 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂; Mazdaran Province, 10 km S of Chaloos [Chalus]; 380 m a. s. l.; 15 Jun. 2010; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDE5163D9BED883FD63FC88.taxon	distribution	Distribution Unclear because of historical confusion with A. propinqua Schenck, 1853, but probably from Europe to Central Asia (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDF5162D99FD81DFEE8FD92.taxon	distribution	Distribution Turkey, Cyprus, Israel and the West Bank, Lebanon, Syria, Iran (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDF5165D9E9DB5AFE0FFB87.taxon	materials_examined	Material examined Lectotype SYRIA • ♀ (of lectotype of Andrena emesiana); Homs; MNHN (illustrated Figs 149 – 152). Holotype TURKEY • ♀ (holotype of Andrena recurvirostra); Beyşehir; 4 Jun. 1964; J. Gusenleitner leg.; OÖLM. Other material IRAN • 1 ♀; Lorestan Province, Tootmashour [Tut, Masur-e Abi]; 2000 m a. s. l.; 31 May 2010; Mi. Halada leg.; OÖLM. SYRIA • 1 ♀; Salkhad env.; 6 May 1996; Mi. Halada leg.; OÖLM. TURKEY • 1 ♂; Mus Dağı; 1 Jun. 1932; M. D. leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDF5165D9E9DB5AFE0FFB87.taxon	discussion	Remarks Warncke (1967) synonymised A. emesiana with A. erythrocnemis Morawitz, 1871 auct. (= A. griseobalteata Dours, 1872, see Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 b; Le Divelec 2021). As is typical of Pérez’s writing, the description itself is unhelpful, and does not allow diagnosis. Inspection of the female lectotype in Paris (designated by Warncke in 1967, though automatically the holotype since the taxon was described from a single specimen) shows that it is not a synonym of A. griseobalteata, as the body size is too small, the scutum is less densely punctate, the tarsi are orange, and the first tergum is extremely finely and densely punctate (Figs 150 – 152). It is instead the senior synonym of A. recurvirostra, which was later described from Turkey (Warncke 1975). Unification under the name A. emesiana clarifies this situation, and means that A. griseobalteata has never been recorded from the Levant, as mentioned by Grace (2010, as A. erythrocnemis), this mention probably deriving from Warncke’s incorrect synonymisation. Note, the name is misspelled on one of the specimen labels as ‘ emesiensis ’ (Fig. 149); the correct spelling is emesiana (Pérez 1911).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDF5165D9E9DB5AFE0FFB87.taxon	distribution	Distribution Turkey *, Syria, Iran *.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD85165D993DF50FEC1F85C.taxon	materials_examined	Material examined IRAN • 1 ♂; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD85165D993DF50FEC1F85C.taxon	distribution	Distribution Greece, North Macedonia, Romania, Ukraine, Bulgaria, Egypt, Turkey, the Caucasus, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD85165D981DD4CFC33F9AA.taxon	materials_examined	Literature records Ascher & Pickering (2021): Centr. Alborz, Kandavan Pass, 8 km above Siahbishe; 2400 m a. s. l.; 7 Jun. 1966; D. Baker leg.; SEMC; via GBIF occurrence 686218963.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD85165D981DD4CFC33F9AA.taxon	discussion	Remarks The presence of A. enslinella in Iran was indicated by the distribution maps of Warncke (Gusenleitner & Schwarz 2002), so it is not treated as new for the country.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD85165D981DD4CFC33F9AA.taxon	distribution	Distribution Central Europe eastwards to Russia (European part, Urals, Western Siberia), Turkey, the Caucasus, Iran (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95164D990D81DFB01FD74.taxon	materials_examined	Material examined IRAN • 1 ♀; Khuzistan, Shadegan; 15 Feb. 1956; K. Warncke det.; R. Schäuffele leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95164D990D81DFB01FD74.taxon	discussion	Remarks The distribution maps of Warncke included a dot in northern Iran, but not a dot in southern Iran from this specimen, which is nonetheless part of the Warncke collection. No other specimens from Iran could be located in his collection.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95164D990D81DFB01FD74.taxon	distribution	Distribution Tunisia, Libya, Egypt, Israel and the West Bank, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95164D9B8DBFCFD8BFA00.taxon	materials_examined	Material examined IRAN • 1 ♀; Shah Abas Kabir; 8 May 1972; K. Warncke det.; Bytinski-Salz leg.; OÖLM. IRAQ • 1 ♂, 3 ♀♀; Baghdad; 15 Feb. 1963; Seipka leg.; OÖLM. SYRIA • 1 ♂, 3 ♀♀; Palmyra; 400 m a. s. l.; 10 Apr. 1986; K. M. Guichard leg.; NHMUK • 3 ♀♀; 80 km E of Palmyra; 22 Apr. 1992; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95164D9B8DBFCFD8BFA00.taxon	discussion	Remarks This taxon is possibly synonymous with A. splendidicollis Morawitz, 1894, which is found in Central Asia (Osytshnjuk et al. 2005). Type revision is needed. Molecular analysis shows that A. euzona belongs to a currently undescribed subgenus, and not to the former subgenus Carandrena (= Notandrena, Pisanty et al. 2022 b). The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate the presence of A. euzona in Iran, so this taxon is not considered newly recorded here.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95164D9B8DBFCFD8BFA00.taxon	distribution	Distribution Morocco, Algeria, Tunisia, Libya, Egypt, Israel and the West Bank, Jordan, Syria *, Iraq *, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95166D9E4DCC8FAEFFDF1.taxon	materials_examined	Material examined Syntype PAKISTAN • ♀ (syntype of Andrena peshinica); Quetta; 1 – 30 Jun. 1902; C. G. Nurse leg.; NHMUK (illustrated Figs 153 – 156). Other material IRAN • 3 ♂♂; Kerman Province, 20 km E of Ghobira; 1780 m a. s. l.; 5 Jun. 2010; Mi. Halada leg.; OÖLM • 2 ♂♂; Kerman Province, 8 km N of Bardsir; 2050 m a. s. l.; 6 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Kerman Province, Deh Bakri, Gebal Barez Mts; 1640 m a. s. l.; 3 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂, 1 ♀; Kerman Province, Serač [Sirch]; 2 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Kerman, Ravar, garden of Davarpanah; 1164 m a. s. l.; 9 May 2011; A. Polaszek leg.; NHMUK • 2 ♂♂, 1 ♀; Persepolis env.; 9 May 1999; K. Deneš leg.; OÖLM • 1 ♂, 3 ♀♀; Yazd, Hanza, Hosseinie; 1837 m a. s. l.; 3 Jul. 2020; S. San leg.; ICPI. IRAQ • 1 ♂, 1 ♀; Abu Ghuraib [Abu Ghraib]; 5 – 16 May 1945; H. A. Hamid leg.; OÖLM. KAZAKHSTAN • 1 ♂, 2 ♀♀; Baigakum [Baygekum] bei Djulek Turkest; MNHN.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95166D9E4DCC8FAEFFDF1.taxon	discussion	Remarks Though the specimen of A. peshinica in the NHMUK is marked as a type, it is strictly a syntype, as Nurse never designated a type, and no subsequent authors have redescribed this taxon or designated a lectotype. Gusenleitner & Schwarz (2001) noted that this taxon is extremely close to A. eversmanni based on personal correspondence from Warncke and, as suspected by Popov (1949), A. peshinica is indeed synonymous with A. eversmanni. Examination of the syntype shows the same small body size, squamous hairs on the scutum, thick tergal hairbands, and light terminal fringe (Figs 153 – 156). The justification given by Nurse (1904) that it differed from A. eversmanni because of the red-coloured tergal segments is insufficient, as A. eversmanni often has red coloured terga in other parts of its range, for example in Iran where all examined specimens were at least partially red-marked, and in Kazakhstan where one examined female had red-marked terga, the other with black-marked terga. Note, the name is misspelled on the syntype label as ‘ pishinica ’; the correct spelling is peshinica (Nurse 1904).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFD95166D9E4DCC8FAEFFDF1.taxon	distribution	Distribution Turkey, Armenia, Iran, Iraq *, Russia (European part), Turkmenistan, Uzbekistan, Kazakhstan, Tajikistan, Pakistan *, Mongolia, China (Popov 1949; Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5166D986DB79FCF5FB78.taxon	materials_examined	Material examined CYPRUS • 2 ♀♀; Kykkos; 800 m a. s. l.; 11 May 2014; M. Kafka leg.; OÖLM. IRAN • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ham [Ilam] Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5166D986DB79FCF5FB78.taxon	discussion	Remarks Also newly reported from Cyprus, the range of this taxon has expanded substantially eastwards following recent revisions.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5166D986DB79FCF5FB78.taxon	distribution	Distribution Bulgaria, Cyprus *, Turkey, Lebanon, Syria, Israel and the West Bank, Jordan, Iran * (Gusenleitner & Schwarz 2002; Pisanty et al. 2018; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5166DA4FDDE0FEDBFACB.taxon	distribution	Distribution Syria and Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5166D9ADDCB2FBC6F882.taxon	materials_examined	Material examined CYPRUS • 4 ♂♂, 2 ♀♀; 23 Feb. – 10 Mar. 2017; A. Varnava leg.; AVC • 22 ♂♂, 15 ♀♀; Limassol, Yermasoyia Dam; 7 Mar. 2017; AVC.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5166D9ADDCB2FBC6F882.taxon	distribution	Distribution Turkey, Cyprus *, Israel and the West Bank, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFDB5179D987DE49FCCCFEE4.taxon	distribution	Distribution Iran, Turkmenistan, Uzbekistan (Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45179DA4FD861FA9FFDDD.taxon	distribution	Distribution Europe to Turkey, Syria, Jordan, Iran (Gusenleitner & Schwarz 2002; Scheuchl & Willner 2016).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45179D9BBDC3CFAC7F91F.taxon	discussion	Remarks Andrena flavipes is the most common Iranian Andrena, with 169 female and 201 male specimens examined during this revision from across Iran. In the interests of brevity, these are not detailed here.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45179D9BBDC3CFAC7F91F.taxon	distribution	Distribution West Palearctic to Central Asia, China, India (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45179D998DB9BFDE1FBE2.taxon	discussion	Remarks The distribution maps of Warncke (Gusenleitner & Schwarz 2002) may indicate the presence of A. figurata in northern Iran. It is unclear whether the Ascher & Pickering (2021) listing is based on this source. As A. figurata is known from eastern Turkey and the Caucasus it is included in the Iranian fauna.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45179D998DB9BFDE1FBE2.taxon	distribution	Distribution Italy eastwards to Russia (European part, Urals), south to Turkey, Cyprus, Iran (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45179D9AFDD6FFF44FAB7.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45178D9AADFC5FD47FE6A.taxon	discussion	Remarks Aliyev et al. (2017) report A. florea from Iran with no precise details. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate that A. florea is present in Iran. However, no specimens demonstrating this could be found in the Warncke collection. Because of the presence of this species in eastern Turkey and its ease of identification, we consider A. florea to be present in Iran, though this should be positively confirmed with specimens.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC45178D9AADFC5FD47FE6A.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D994DFFEFC1BF85D.taxon	materials_examined	Material examined IRAN • 16 ♀♀; Mazandarn Province, 15 km S of Alamdeh; 7 Jun. 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D994DFFEFC1BF85D.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002), including Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D986DC42FC5CF976.taxon	materials_examined	Literature records Allahverdi et al. (2016); Ascher & Pickering (2021): 15 km N of Gazvin; 19 Jul. 1975; K. Warncke det.; P. F. Torchio leg.; SEMC; via GBIF occurrence 658366176.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D986DC42FC5CF976.taxon	distribution	Distribution Slightly unclear because of confusion between A. forsterella and A. wilhelmi Schuberth, 1995 (see Schönitzer et al. 1995), but probably found from Italy, through the Balkans, to Turkey, Lebanon, Israel, Iran (Schönitzer et al. 1995; Wood et al. 2020 a; Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D984DAF3FCE4FA9A.taxon	distribution	Distribution Germany eastwards to Russia (European part, Urals, Western Siberia), Turkey, the Caucasus, Kazakhstan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D99CDB12FD47FC0B.taxon	materials_examined	Material examined IRAN • 2 ♀♀; Kurdistan Province, Paniran; 1450 m a. s. l.; 14 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Pass SW Chalus-Karaj / Mazandaran; 2470 m a. s. l.; 28 May 1978; M. Kraus leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC55178D99CDB12FD47FC0B.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD9E3DCA4FB72F91E.taxon	materials_examined	Material examined IRAN • 1 ♀; Elburz, Chur [Khur]; 2400 m a. s. l.; 7 – 12 Jun. 2005; V. Major leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD9E3DCA4FB72F91E.taxon	distribution	Distribution Turkey, Israel, Armenia, Azerbaijan, Iran * (Osytshnjuk et al. 2008; Pisanty et al. 2018).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD997D81DFDF6FB94.taxon	materials_examined	Material examined IRAN • 1 ♀; Gilan, 5 km E of Rudbar; 400 m a. s. l.; 8 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂, 2 ♀♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 1 ♂; Mazandaran Province, 20 km S of Amol; 640 m a. s. l.; 6 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂; Mazandaran, 20 km S of Nowshahr, Weysser [Veysar]; 1400 m a. s. l.; 9 – 14 Jun. 1977; C. Holzschuh and F. Ressl leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD997D81DFDF6FB94.taxon	discussion	Remarks Though Morice reported specimens from Iran as A. lucens, a synonym of A. nitidiuscula, we consider them to be A. fulvicornis because they were collected in February. Andrena nitidiuscula is strictly univoltine, flying in the summer, whereas A. fulvicornis is bivoltine, flying in the spring and the summer (Schmid-Egger & Doczkal 1995).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD997D81DFDF6FB94.taxon	distribution	Distribution Unclear because of the historical synonymy by Warncke with A. nitidiuscula Schenck, 1853 (see SchmidEgger & Doczkal 1995; Gusenleitner & Schwarz 2002). Andrena fulvicornis has a more southerly distribution in hotter and more Mediterranean areas, and is found across the Mediterranean basin to the Middle East (Wood et al. 2020 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD9A5DD5CFB75FA3C.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province, Dasht Arjan; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Kurdistan Province, Paniran; 1450 m a. s. l.; 14 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517BD9A5DD5CFB75FA3C.taxon	distribution	Distribution Italy, Slovenia, Croatia, Albania, North Macedonia, Greece, Bulgaria, Ukraine, Turkey, Israel and the West Bank, Lebanon, Jordan, Iran * (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517AD9B9DFC5FAB1FD56.taxon	materials_examined	Material examined IRAN • 1 ♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♂; Fars Province, Dast Arjan; 2040 m a. s. l.; 6 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Gilan Province, 15 km SE of Tutkabon; 1100 m a. s. l.; 9 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂; Kerman Prov., 25 km E of Jiroft (Mijan); 1650 m a. s. l.; 27 May 2014; J. Halada leg.; OÖLM • 1 ♀; Kerman Prov., Jupar [Joupar]; 1900 m a. s. l.; 1 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Kerman Province, 20 km E of Ghobira; 1780 m a. s. l.; 5 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Kerman Province, Deh Bakri, Gebal Barez Mts; 1640 m a. s. l.; 3 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Kerman Province, Kūh-e Madvār; 3 Jun. 2008; V. Major leg.; OÖLM • 6 ♂♂, 1 ♀; Mazdaran Province, 10 km S of Chaloos [Chalus]; 380 m a. s. l.; 15 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Sahne env. [Sahneh]; 13 May 1999; K. Deneš leg.; OÖLM • 1 ♀; Shiraz-Yasouj road (15 km to Yasouj); 20 May 2011; A. Monfared leg.; ICPI • 1 ♀; Yasouj, Gachsaran Shadega; 26 May 2009; A. Monfared leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC6517AD9B9DFC5FAB1FD56.taxon	distribution	Distribution West and Central Palearctic to India (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC7517AD9B0DCD3FAA1F85C.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province, Dast Arjan; 2040 m a. s. l.; 6 May 2016; M. Kafka leg.; OÖLM • 1 ♀; 20 km W of Neyriz / Fars Steppe; 1550 m a. s. l.; 18 May 1978; K. Warncke leg.; K. Warncke det.; OÖLM. JORDAN • 1 ♀; 20 km N of Karak; 1000 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 1 ♀; Al Karak env. [Kerak]; 16 Apr. 2002; M. Snižek leg.; OÖLM • 1 ♀; Aljoun; 6 May 2012; M. Kafka leg.; OÖLM • 3 ♂♂; Petra [Wadi Musa]; 14 May 1995; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC7517AD9B0DCD3FAA1F85C.taxon	distribution	Distribution Egypt, Israel and the West Bank, Jordan *, Syria, Turkey, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC7517AD9A5DD8BFAA0FA2B.taxon	distribution	Distribution Turkey and Iran (Gusenleitner & Schwarz 2002; Khodarahmi Ghahnavieh & Monfared 2019).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC7517AD998DA1FFD23FBC3.taxon	materials_examined	Material examined IRAN • 1 ♀; Kurdistan Province, Paniran; 1450 m a. s. l.; 14 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC7517AD998DA1FFD23FBC3.taxon	distribution	Distribution Bulgaria, Turkey, Israel and the West Bank, Lebanon, Jordan, Syria, Iran * (Gusenleitner & Schwarz 2002; Wood et al. 2020 a; Wood 2021 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC0517DD998D81DFB76FBF2.taxon	materials_examined	Material examined Holotype TURKEY • ♀; Mardin; MNHN. Other material IRAN • 2 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM • 2 ♂♂; Yasouj, Abshar; 2 Mar. 2011; H. Elahi & M. Mesavian leg.; ICPI • 1 ♂; Yasouj, Khakalan; 6 Jun. 2011; A. Monfared leg.; ICPI • 1 ♂; Yasouj, Tange Tamoradi; 2247 m a. s. l.; 9 Mar. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Vezg. [Kūh-e Vezg] Khakalan; 12 Jun. 2011; S. Hossein Zade leg.; ICPI. TURKEY • 3 ♀♀; Tanin-Tanin-Pass [Şırnak Province]; 1900 m a. s. l.; 3 Jun. 1980; K. Warncke det.; K. Warncke leg.; OÖLM • 1 ♀; Uludere, Hakkâri; 5 Apr. 1977; K. Warncke det.; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC0517DD998D81DFB76FBF2.taxon	distribution	Distribution Unclear, due to confusion with A. elmaria (see Gusenleitner 1998). Probably only confirmed from eastern Turkey and Iran *. Records from Israel and the West Bank (Warncke 1969) are A. elmaria (Pisanty et al. 2022 a); records from Cyprus (Varnava et al. 2020) are probably all referable to A. elmaria. Inspection of material previously assigned to these two species is necessary to clarify exact ranges.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC0517DD980DD65FA83F98A.taxon	discussion	Remarks The taxonomic status of A. gussakovskii requires investigation; as noted by Gusenleitner & Schwarz (2002) it is very similar to A. marmora Nurse, 1904, which was described from Pakistan and is found also in Israel and the West Bank (Warncke 1969). The two taxa may be conspecific.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC0517DD980DD65FA83F98A.taxon	distribution	Distribution Iran, Central Asia, Russia (Western Siberia) (Osytshnjuk et al. 2008; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC0517DD9FEDF7DFC5EF85C.taxon	materials_examined	Material examined IRAN • 1 ♀; West Azerbaijan, Bazargan; 1380 m a. s. l.; 11 May 1978; M. Kraus leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC0517DD9FEDF7DFC5EF85C.taxon	distribution	Distribution Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517CD9BBD81CFD3EFD69.taxon	discussion	Remarks The original records supporting these listings are not clear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate the presence of this species in northern Iran, so it is included in the fauna.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517CD9BBD81CFD3EFD69.taxon	distribution	Distribution North Africa and Southern Europe to Russia (European part), Turkey, Azerbaijan, Iran (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517CD9A4DA17FB2FFB25.taxon	materials_examined	Material examined IRAN • 1 ♂, 1 ♀; Tehran Province, 6 km N of Gačsar [Gachsar]; 30 May 2002; J. Prochâzka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517CD9A4DA17FB2FFB25.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517ED9E7DDA3FBDCFE71.taxon	materials_examined	Material examined Syntypes PAKISTAN • ♀ (syntype of Andrena cara); Peshin; 1 – 31 Mar. 1902; C. G. Nurse leg.; NHMUK (illustrated Figs 157 – 160) • ♂ (syntype of Andrena halictoides); Peshin; 1 – 31 Mar. 1902; C. G. Nurse leg.; NHMUK (illustrated Figs 161 – 164). Other material IRAN • 1 ♂, 2 ♀♀; Yasouj, Isfahan Road, Tange Seriz; 1687 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; ICPI. TAJIKISTAN • 1 ♀; Fajsabad [Faizobod], 60 km E of Duschanbe; 5 – 7 May 1991; J. Halada leg; OÖLM. TURKMENISTAN • 8 ♀♀; Nebit-Dag [Balkanabat], 25 km NW of Jebel; 28 Apr. 1993; K. Deneš leg.; OÖLM • 1 ♀; Repetek; 8 Apr. 1992; M. Snižek leg.; OÖLM. UZBEKISTAN • 1 ♀; 80 km SE of Karschi [Qarshi]; 31 Apr. 1991; J. Halada leg; OÖLM • 2 ♀♀; Derbent, 13 km SE of Karschi [Qarshi]; 1450 m a. s. l.; 2 May 1991; J. Halada leg; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517ED9E7DDA3FBDCFE71.taxon	discussion	Remarks See comments on A. minor Warncke, 1975 stat. nov. at the end of the annotated list.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC1517ED9E7DDA3FBDCFE71.taxon	distribution	Distribution Iran *, Turkmenistan, Uzbekistan, Tajikistan, Pakistan * (Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517ED99ADDB7FA9DF85C.taxon	materials_examined	Material examined IRAN • 1 ♂; Pass SW Chalus-Karaj / Mazandaran; 2470 m a. s. l.; 28 May 1978; M. Kraus leg.; OÖLM. JORDAN • 1 ♀; 20 km N of Amman; 620 m a. s. l.; 23 Apr. 2006; K. Deneš leg.; OÖLM • 2 ♀♀; 30 km NW of Aljun; 600 m a. s. l.; 29 Apr. 2006; K. Deneš leg.; OÖLM • 3 ♀♀; Aljun env.; 840 m a. s. l.; 1 May 2006; K. Deneš leg.; OÖLM. SYRIA • 1 ♂; occ. An Nasrah env. [Al-Nasrah]; 8 – 13 Apr. 2005; J. Saki leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517ED99ADDB7FA9DF85C.taxon	discussion	Remarks The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate the presence of A. hyemala in Iran, and one specimen could be found in the Warncke collection. It is unclear whether the listing of Ascher & Pickering (2021) is supported by an additional source. The status of A. hyemala repressa Warncke, 1975 is unclear and requires revision, as it may consistitute a distinct species.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517ED99ADDB7FA9DF85C.taxon	distribution	Distribution South-eastern Europe, Ukraine, Russia (European part), Turkey, Israel and the West Bank, Jordan *, Syria *, Iran, Turkmenistan, Kazakhstan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517EDA5ADBB6FE49FC9B.taxon	distribution	Distribution Golan Heights, Syria, Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517ED99DDA43FAF1FBCF.taxon	materials_examined	Material examined IRAN • 1 ♂; Meymeh / Esfahan [Isfahan]; 2000 m a. s. l.; 14 May 1978; M. Kraus leg.; K. Warncke det.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517ED99DDA43FAF1FBCF.taxon	distribution	Distribution Ukraine, Russia (European part), Turkey, Iran * (Osytshnjuk 1978; Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC3517ED9B8D8F9FF44FDCE.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5171D983DDB1FCBDF9B3.taxon	materials_examined	Material examined IRAN • 2 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 5 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM • 1 ♀; Tehran Province, Samgh Abad; 1900 m a. s. l.; 16 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5171D983DDB1FCBDF9B3.taxon	distribution	Distribution Turkey and Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5171D9B5DAE1FF44FBCB.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5171D9E8D81DFBB0FC7A.taxon	discussion	Remarks Morice (1921) lists A. hypopolia Pérez, which is not the correct species authority. This may by a similar situation to that of the type of A. gallica, which was described by Schmiedeknecht but the type material was collected by Pérez in southern France, and hence the type series remained in the Pérez collection in Paris (Le Divelec 2021). Moreover, the taxonomic situation is highly confused. Warncke treated A. hypopolia as a subspecies of A. numida Lepeletier, 1841 (Warncke 1967), but also described taxa such as A. numida albiscopa Warncke, 1967 from Turkey, and used several other names in combination to describe ‘ regional’ variation (see map in Gusenleitner & Schwarz 2002). It is the opinion of the lead author that all taxa should probably be synonymised under A. numida, and that he observed variation is predominantly one of colouration, not structural variation. For now, A. hypopolia is included on the Iranian list, but this may be subsumed into A. numida following molecular revision.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5171D9E8D81DFBB0FC7A.taxon	distribution	Distribution Unclear. Taking a broad concept with all taxa synonymised under A. numida, this taxon has a panMediterranean distribution (see map in Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5173D9A6DF3AFCFAFD73.taxon	materials_examined	Material examined Syntype KASHMIR • ♀ (syntype of Andrena patella); 6000 – 7000 feet a. s. l.; 1 – 31 May 1901; NHMUK (illustrated Figs 165 – 168). Other material IRAN • 1 ♂; Elburz Mountains, 60 km E of Minudasht; 1280 m a. s. l.; 26 May 2007; O. Sauša leg.; OÖLM • 1 ♂, 2 ♀♀; Elburz, Ov Zanak-Ski, 1 km N of Ab Ali; 2300 m a. s. l.; 11 Jul. 1965; K. Warncke det.; G. Soika and G. A. Mavromoustakis leg.; OÖLM • 1 ♂; Fars Province [Kohgiluyeh and BoyerAhmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM. KASHMIR • 1 ♀; Gulmarg; summer 1913; NHMUK • 1 ♀; West Himalaya, Khalatse [Khalsi], Kashmir; 4000 – 5000 m a. s. l.; 6 May 1939; K. Warncke det.; OÖLM. TURKEY • 1 ♂; Başaklı, Erzurum; 20 May 1972; K. Warncke det.; H. Ozbek leg.; OÖLM • 3 ♀♀; Hakkâri Şivelan; 18 May 1975; K. Warncke det.; K. Warncke leg.; OÖLM • 1 ♂; Tanyeri, Erzincan; 22 Apr. 1976; K. Warncke det.; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5173D9A6DF3AFCFAFD73.taxon	discussion	Remarks Andrena induta is closely related to A. infirma Morawitz, 1876, which is found in Kazakhstan, Turkmenistan, Uzbekistan, Kyrgyzstan, and Tajikistan (Osytshnjuk et al. 2008). Structurally, there do not appear to be clear differences, but A. induta has much brighter pubescence, and females can be separated by the light (induta) or dark (infirma) scopal hairs. Gusenleitner & Schwarz (2002) argue that A. induta may therefore simply be a colour form of A. infirma. However, more study is required before a synonymy can be made with confidence. In contrast, the type specimen of A. patella is clearly identical to A. induta, as it has the same strong raised longitudinal impunctate line on the clypeus (Fig. 167), and the scopae are golden (Figs 166, 168). It also falls within the known distributional range of A. induta, this taxon being known from the western Himalayas in Kashmir. A prudent first step is to synonymise A. patella with A. induta, and for further synonymy between A. infirma and A. induta to wait until type inspection is possible and ideally molecular work is conducted. As Warncke’s distribution maps included points in northern Iran, A. induta is not reported here as new for the country.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCC5173D9A6DF3AFCFAFD73.taxon	distribution	Distribution Turkey, Armenia, Iran, Uzbekistan, Tajikistan, Afghanistan, Kashmir, India (Warncke 1974 c, 1975 a; Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCE5173D9AADD24FEFEFADA.taxon	distribution	Distribution Iran, Turkmenistan (Osytshnjuk et al. 2005). Material from the Arabian Peninsula (Ascher & Pickering 2021) must be investigated because of a lack of taxonomic clarity regarding A. skorikovi Popov, 1940 (see below).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCE5173D9BEDC81FCE4F8A1.taxon	materials_examined	Material examined IRAN • 2 ♂♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM. SYRIA • 3 ♂♂, 1 ♀; Bloudan; 16 May 1995; K. Deneš leg.; OÖLM • 1 ♀; Maalula [Maaloula]; 17 May 1995; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCE5173D9BEDC81FCE4F8A1.taxon	distribution	Distribution Hungary, Albania, Romania, Bulgaria, Greece, Turkey, Syria *, Georgia, Armenia, Azerbaijan, Iran * (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCE5173D9A6DBFAFD44FBBD.taxon	distribution	Distribution Morocco, Algeria, Tunisia, Libya, Egypt, Israel and the West Bank, Turkey, Iran (Gusenleitner & Schwarz 2002; Khodaparast & Monfared 2012).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCE5172D991DE28FCBDFEE4.taxon	materials_examined	Material examined IRAN • 2 ♀♀; Kohgiluyeh and Boyer-Ahmad Province, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCE5172D991DE28FCBDFEE4.taxon	distribution	Distribution Turkey and Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5172D9B8D862FB2FFC5C.taxon	materials_examined	Material examined IRAN • 1 ♂; Bakhtiyari Province, 20 km SE of Lordegan; 31 May 2014; J. Halada leg.; OÖLM • 1 ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 28 May 2014; J. Halada leg.; OÖLM • 1 ♀; Kerman Prov., Jupar [Joupar]; 1900 m a. s. l.; 1 Jun. 2010; P. Tymer leg.; OÖLM • 3 ♂♂; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg; OÖLM • 2 ♂♂; Pasagrad env. [Pasargad]; 8 May 1999; K. Deneš leg.; OÖLM • 1 ♂; Yasouj, Vezg; 2159 m a. s. l.; 23 Mar. 2019; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5172D9B8D862FB2FFC5C.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5172D99BDD1AFB0FFA00.taxon	materials_examined	Material examined IRAN • 1 ♂; Elburz Mountains, 60 km E of Minudasht; 26 May 2007; O. Sauša leg.; OÖLM • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5172D99BDD1AFB0FFA00.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5175D9AFDCCEFE31FECC.taxon	materials_examined	Material examined SYRIA • 1 ♂; Bosra; 3 May 1995; K. Deneš leg.; OÖLM • 1 ♂; Hasake; 8 May 1996; Ma. Halada leg.; OÖLM • 29 ♂♂, 5 ♀♀; Latakia, Salahidin env. [Castle of Saladin]; 10 Jun. 2000; K. Deneš leg.; OÖLM • 1 ♂; Maalula [Maaloula]; 17 May 1995; K. Deneš leg.; OÖLM • 1 ♂; Ras al Basit; 19 May 1995; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5175D9AFDCCEFE31FECC.taxon	discussion	Remarks No primary literature could be found to support the record of Ascher & Pickering, but based on the distribution maps of Warncke (Gusenleitner & Schwarz 2002), A. lamiana is present in northwestern Iran. It is therefore included in the faunal list.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCF5175D9AFDCCEFE31FECC.taxon	distribution	Distribution North Macedonia, Greece, Turkey, Cyprus, Syria *, Iran (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC85175D9D8DAE0FD0DFB42.taxon	distribution	Distribution Iran and Turkmenistan (Ariana et al. 2009).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC85175D982DC0BFB5BF85C.taxon	materials_examined	Material examined TURKEY • 1 ♀; Hakkâri, S. Varegös / Mt. Sat; 2000 m a. s. l.; 17 Jun. 1984; K. Warncke leg.; OÖLM • 1 ♂; Hakkâri, Beytüşşebap; 25 May 1988; K. Warncke leg.; OÖLM • 3 ♀♀; Van, Muradiye env., 120 km NE of Van; 2000 m a. s. l.; K. Deneš leg.; OÖLM • 1 ♂; Van, Güzeldere Gecidi, 70 km SE of Van; 2730 m a. s. l.; 7 Jun. 2001; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC85175D982DC0BFB5BF85C.taxon	discussion	Remarks No primary literature could be found to support the record of Ascher & Pickering, but based on the distribution maps of Warncke (Gusenleitner & Schwarz 2002) as well as examined specimens from Hakkâri and Van provinces, A. laticeps is present in eastern Turkey very close to the border of Iran, so the species is considered to be present in north-western Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC85175D982DC0BFB5BF85C.taxon	distribution	Distribution Turkey, Georgia, Armenia, Iran (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC85175D99DD8B5FCE4FC7F.taxon	materials_examined	Material examined AFGHANISTAN • 2 ♀♀; Saripul [Sar-e Pol], Goslandi distr., Malekan village; 21 May 1996; G. G. M. Schulten leg.; RMNH. IRAN • 1 ♀; S. W. Persia, Escalera; K. Sefiet leg.; NHMUK.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC85175D99DD8B5FCE4FC7F.taxon	distribution	Distribution Portugal, Spain, North Macedonia, Greece, Turkey, Israel and the West Bank, Georgia, Armenia, Azerbaijan, Iran, Turkmenistan, Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan, Afghanistan * (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95174D9A0D81DFD47FBB2.taxon	materials_examined	Material examined IRAN • 1 ♂; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 2 ♀♀; Fars Province, 10 km E of Kazerum; 1990 m a. s. l.; 23 May 2014; J. Halada leg.; OÖLM • 1 ♀; Fars Province, Dast Arjan; 2040 m a. s. l.; 6 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Kohgiluyeh and Boyer, Ahmad Prov., Likak, Gach Bolland; 1640 m a. s. l.; 7 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Yasouj, Darreh Grouh Firuz Abad, Faramarz Village; 1960 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; ICPI • 11 ♂♂, 2 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 31 Mar. 2021; E. Rostami leg.; ICPI • 2 ♂♂, 11 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; ICPI • 10 ♂♂, 1 ♀; Yasouj, Tange Tamoradi; 2247 m a. s. l.; 21 Mar. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Vezg; 2159 m a. s. l.; 23 Mar. 2019; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95174D9A0D81DFD47FBB2.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95174D989DD24FC2DF9DA.taxon	materials_examined	Material examined IRAN • 1 ♀; Azer. e Garbi. Prov. [West Azerbaijan Province], Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Kerman Prov., Deh Bakri Pass; 2300 m a. s. l.; 28 May 2014; J. Halada leg.; OÖLM • 1 ♀; 15 km SE of Sarvestan / Fars; 1800 m a. s. l.; 17 May 1978; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95174D989DD24FC2DF9DA.taxon	discussion	Remarks Barcoded Iranian specimens were used in the analysis of Praz et al. (2022).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95174D989DD24FC2DF9DA.taxon	distribution	Distribution Romania, Bulgaria, North Macedonia, Greece, Turkey, Cyprus, Lebanon, Iran * (Gusenleitner & Schwarz 2002; Varnava et al. 2020; Wood 2021 b; Boustani et al. 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95177D9B7DF8CFB2FFE51.taxon	materials_examined	Material examined IRAN • 1 ♀; Gilan Province, 20 km W of Astara; 640 m a. s. l.; 17 Jun. 2010; Mi. Halada leg.; OÖLM • 2 ♀♀; Kamyaran, Asr abad; 17 Jul. 2017; H. Fatehi leg.; ICPI • 1 ♀; Kerman Province, Kuhha Qohrud, Hane-ye Sorkh [Gardaneh-ye]; 2700 m a. s. l.; V. Major leg.; OÖLM • 1 ♀; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; J. Halada leg.; OÖLM • 1 ♀; NE Iran, Kuh-e Sorgh-Hesar [?], 10 km S of Ataiye; 1700 m a. s. l.; 28 May 2003; V. Major leg.; OÖLM • 1 ♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFC95177D9B7DF8CFB2FFE51.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCA5177D987DD3FFC41FA92.taxon	distribution	Distribution Iran, Turkmenistan, Uzbekistan, Kazakhstan (Radchenko et al. 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCA5177D987DB18FD8BFBB0.taxon	discussion	Remarks The primary source for these listings is unclear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) include a dot on the Iran / Turkmenistan border, which may be within Iran. Given that A. limbata is present in eastern Turkey and the Caucasus, it is considered highly likely to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCA5177D987DB18FD8BFBB0.taxon	distribution	Distribution Southern Europe to Russia (European part, Urals), Turkey, the Caucasus, Iran, and Turkmenistan (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCA5176D99DDC59FB04FE0E.taxon	materials_examined	Material examined IRAN • 5 ♂♂; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 3 ♀♀; Boyer-A. o. Kohg Prov., Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada; OÖLM • 1 ♂, 3 ♀♀; Fars Province, 10 km E of Kazerum; 1990 m a. s. l.; 23 May 2014; J. Halada leg.; OÖLM • 1 ♂, 6 ♀♀; Fars Province, Dasht Arian; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 3 ♂♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 57 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 3 ♀♀; Kohgiluyeh and Boyer, Ahmad Prov., Likak, Gach Bolland; 1640 m a. s. l.; 7 May 2016; M. Kafka leg.; OÖLM • 4 ♂♂; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM • 1 ♂, 6 ♀♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Lorestan Province, Tootmashour [Tut, Masur-e Abi]; 2000 m a. s. l.; 31 May 2010; Mi. Halada leg.; OÖLM • 4 ♂♂; Mazandaran Province, Kojuk [36 ’ 23, 51 ’ 40]; 7 Jun. 2014; J. Halada leg.; OÖLM • 2 ♀♀; Pasagrad [Pasargad]; 8 May 1999; K. Deneš leg.; OÖLM • 1 ♂; Persepolis env.; 9 May 1999; K. Deneš leg.; OÖLM • 3 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 31 Mar. 2021; E. Rostami leg.; ICPI • 9 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 10 Mar. 2021; E. Rostami leg.; ICPI • 1 ♂; Yasouj, Tang Konareh, Dasht-e Rum; 1858 m a. s. l.; 30 Apr. 2021; E. Rostami leg.; ICPI • 1 ♂, 5 ♀♀; Yasouj, Tange Tamoradi; 2247 m a. s. l.; 21 Mar. 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCA5176D99DDC59FB04FE0E.taxon	distribution	Distribution Golan Heights, Turkey, and Iran * (Gusenleitner & Schwarz 2002; Pisanty et al. 2018).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D9E0D8F6FC36FD56.taxon	distribution	Distribution Israel and the West Bank, Jordan, Iran (Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D982DF68FB3BF85C.taxon	materials_examined	Material examined IRAN • 1 ♂, 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D982DF68FB3BF85C.taxon	distribution	Distribution Bulgaria, Greece, Turkey, Cyprus, Israel and the West Bank, Lebanon, Jordan, Syria, Iran *, Afghanistan (Warncke 1974 c; Gusenleitner & Schwarz 2002; Wood et al. 2020 a; Wood 2021 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D9A1DD8DFBF0F9E7.taxon	materials_examined	Material examined IRAN • 1 ♂, 1 ♀; Golestan, 70 km E of Minudasht; 1050 m a. s. l.; 12 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Yazd, Mehriz, Shohaday gomnam, 1500 m a. s. l.; 23 Mar. 2021; S. San leg.; ICPI • 1 ♀; Yazd, Mehriz, Shohaday gomnam; 1500 m a. s. l.; 8 May 2021; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D9A1DD8DFBF0F9E7.taxon	distribution	Distribution Turkey, Georgia, Armenia, Azerbaijan, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D9CDDA1EFB49FBC4.taxon	materials_examined	Material examined IRAN • 2 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM. SYRIA • 5 ♂♂, 2 ♀♀; Bludan, 57 km NW of Damascus; 24 Apr. 1992; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFCB5176D9CDDA1EFB49FBC4.taxon	distribution	Distribution (subspecies usura) Turkey, Cyprus, Lebanon, Syria *, Iran * (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D99CDFD3FAC7F85C.taxon	materials_examined	Material examined IRAN • 1 ♀; Kohgiluyeh and Boyer-Ahmad Province, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D99CDFD3FAC7F85C.taxon	distribution	Distribution Central Europe to Russia (European part), Turkey, the Caucasus, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D99CD81DFC5EFDFB.taxon	materials_examined	Material examined IRAN • 4 ♂♂, 1 ♀; Elburz Mountains, 60 km E of Minudasht; 26 May 2007; O. Sauša leg.; OÖLM • 10 ♀♀; Mazandaran Province, 3.2 km S of Kandelous; 1877 m a. s. l.; 4 Jun. 2015; M. Obořil leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D99CD81DFC5EFDFB.taxon	distribution	Distribution Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D989DB63FB80FB9A.taxon	discussion	Remarks The primary source for these listings is unclear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate that A. minutuloides is present in northwestern Iran. The species is common in the Caucasus region, so is considered to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D989DB63FB80FB9A.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D9B9DD42FE23F92B.taxon	materials_examined	Material examined IRAN • 3 ♀♀; Khuzestan, Dezful, Rozgah; 15 Mar. 2017; A. Monfared leg.; ICPI • 1 ♂, 9 ♀♀; Yazd, Abarkooh, Faraghe; 1698 m a. s. l.; 17 Mar. 2021; S. San leg.; ICPI • 1 ♂; Yazd, Hanza to Mehriz; 1928 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 1 ♀; Yazd, Mehriz, Shohaday gomnam; 1500 m a. s. l.; 23 Mar. 2021; S. San leg.; ICPI • 5 ♀♀; Yazd, Polis e rah taft, entehay kooche; 1399 m a. s. l.; 12 Mar. 2020; S. San leg.; ICPI • 1 ♂; Yazd, Taft, Tezerjan, Mazra ali, Agha Seyyed; 2109 m a. s. l.; 24 Mar. 2019; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF45149D9B9DD42FE23F92B.taxon	distribution	Distribution Egypt, Israel and the West Bank, Lebanon, Jordan, Syria, Turkey, Iran (Wood et al. 2020 a; Wood 2021 a; Kratochwil 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF55148D9A4DCBAFE40F978.taxon	distribution	Distribution Spain, Morocco, Algeria, Tunisia, Israel and the West Bank, Jordan, Iran (Gusenleitner & Schwarz 2002; Wood et al. 2020 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF55148D988DB3FFC11FA32.taxon	materials_examined	Material examined IRAN • 1 ♂; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Elburz, Pulour [Polour], 22 km N of di Ab Ali; 13 Jul. 1965; K. Warncke det.; G. Soika and G. A. Mavromoustakis leg.; OÖLM • 1 ♀; Fars Province [Kohgiluyeh and BoyerAhmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Isfahan, Semirom, Yasouj-semirom road, LahJaru; 2263 m a. s. l.; 30 Mar. 2018; A. Monfared leg.; ICPI • 2 ♂♂; Yasouj, Abshar; 2 Mar. 2011; H. Elahi and M. Mesavian; ICPI • 1 ♀; Yasouj, Khakalan; 6 Jun. 2011; A. Monfared leg.; ICPI • 1 ♂; Yasouj, Vezg. [Kūh-e Vezg]; 12 Apr. 2011; A. Monfared leg.; ICPI • 6 ♂♂; Yazd, Banadak-o sadat; 2103 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 1 ♂; Yazd, Ghadam gah, Dehbala; 2175 m a. s. l.; 12 Mar. 2020; S. San leg.; ICPI • 4 ♂♂; Yazd, Jade konj-e kooh, dakal; 2019 m a. s. l.; 26 Mar. 2021; S. San leg.; ICPI • 8 ♂♂, 2 ♀♀; Yazd, Mehriz to Tang Chenar; 1965 m a. s. l.; 30 Mar. 2021; S. San leg.; ICPI • 6 ♂♂; Yazd, Mehriz, Posht e Hosseinie Shohaday gomnam; 1500 m a. s. l.; 28 Feb. 2020; S. San leg.; ICPI • 4 ♂♂; Yazd, Shahneh village, Zorband; 1890 m a. s. l.; 23 Mar. 2019; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF55148D988DB3FFC11FA32.taxon	distribution	Distribution Turkey, Armenia, Iran, Turkmenistan, Uzbekistan, Kazakhstan, Afghanistan, Kyrgyzstan, China, Mongolia (Popov 1967; Ascher & Pickering 2021). Records from within the current political boundaries of Mongolia require confirmation (Y. Astafurova pers. comm.).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF55148D989D81DFAFCFDB0.taxon	materials_examined	Material examined IRAN • 1 ♂; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF55148D989D81DFAFCFDB0.taxon	distribution	Distribution Greece, Turkey, Cyprus, Lebanon, Syria, Iran * (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF5514BD9B6DFE7FB2FFDDB.taxon	materials_examined	Material examined IRAN • 2 ♂♂; Azer. e Garbi. Prov. [West Azerbaijan Province], Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♂; Golestan, 40 km E of Minudasht NP; 750 m a. s. l.; 1 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂; Khorasan Province, Kuh-e Gamal, 20 km W of Deyhuk; 1 Jun. 2005; V. Major leg.; OÖLM • 3 ♀♀; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM • 1 ♀; W Elburz, Kalardasht pl. NE of Rudbarak; 1350 m a. s. l.; 12 Aug. 1970; OÖLM • 3 ♀♀; Zagros, 15 km Semirom; 2600 m a. s. l.; 24 May 2013; V. Major leg.; OÖLM • 1 ♀; Zangan [Qazvin] Province, Aveq; 2500 m a. s. l.; 9 Jun. 2013; V. Major leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF5514BD9B6DFE7FB2FFDDB.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF6514BD999DB82FB2FFC26.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF6514BD9BDDFB2FB2FF85C.taxon	materials_examined	Literature records Osytshnjuk et al. (2008); Allahverdi et al. (2016); Ascher & Pickering (2021): Mazandaran, Sardabrud, 3 – 5 km above Rudbarak; 1500 m a. s. l.; 17 Apr. 1966; D. Baker leg.; SEMC; via GBIF occurrence 658462936.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF6514BD9BDDFB2FB2FF85C.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF6514BD994DAADFE0DF9CA.taxon	materials_examined	Material examined IRAN • 1 ♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 2 ♂♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM. JORDAN • 1 ♀; 15 km W of Madaba; 760 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 4 ♀♀; 20 km N of Amman; 620 m a. s. l.; 23 Apr. 2006; K. Deneš leg.; OÖLM • 2 ♂♂, 13 ♀♀; 20 km N of Karak; 1000 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 1 ♀; North Shuna env.; 29 Apr. 1996; Mi. Halada leg.; OÖLM. SYRIA • 1 ♀; Bosra; 3 May 1995; K. Deneš leg.; OÖLM • 1 ♀; Hasake [Al Hasakah]; 8 May 1996; Ma. Halada leg.; OÖLM • 1 ♂, 1 ♀; 20 km E of Homs; 400 m a. s. l.; 1 Apr. 1988; L. Blank leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF6514BD994DAADFE0DF9CA.taxon	distribution	Distribution Turkey, Cyprus, Israel and the West Bank, Jordan *, Syria *, Iran (Gusenleitner & Schwarz 2002; Scheuchl et al. 2011).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF7514AD9C2D81DFAF8FCB1.taxon	materials_examined	Material examined ARMENIA • 1 ♀; Mount Aragats; 11 Jun. 2017; M. Kasparek leg.; MKC. GEORGIA • 1 ♀; Dedoplistskaro [Dedoplis Tskaro], Eagle Canyon; 13 May 2016; M. Kasparek leg.; MKC. IRAN • 1 ♀; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May 2002; J. Prochazka leg.; OÖLM. SYRIA • 1 ♀; Kassab env. [Kesab]; 28 Mar. 1994; S. Becvar leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF7514AD9C2D81DFAF8FCB1.taxon	distribution	Distribution Greece, Turkey, Syria *, Georgia *, Armenia *, Iran * (Scheuchl & Hazir 2012; Rasmont et al. 2017).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF7514DD9FBDA3BFE2AFE0C.taxon	description	Figs 169 – 174	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF7514DD9FBDA3BFE2AFE0C.taxon	materials_examined	Material examined IRAN • 1 ♀; 10 km SW of Rayen / Kerman [Kūh-e Hazār mountain, 29.518 ° N, 57.271 ° E]; 2350 m a. s. l.; 25 May 1978; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF7514DD9FBDA3BFE2AFE0C.taxon	discussion	Remarks Andrena nitidicollis is an obscure species that was previously known only from desert habitat in southern Kazakhstan. Its phylogenetic affinities are not clear, being placed in the Graecandrena Warncke, 1968 by Gusenleitner & Schwarz (2002), but not by Osytshnjuk et al. (2008). Due to the rarity of this taxon, it is illustrated here (Figs 169 – 174). Through its combination of small body size (Fig. 169), shiny scutum with short, semi-squamous hairs (Fig. 171), orange hind tibiae, thick white tergal hairbands (Fig. 173 – 174), and predominantly smooth propodeal triangle (Fig. 172), A. nitidicollis can be compared to two sets of species in two subgenera: 1) A. (Aciandrena) pavonia Warncke, 1974, A. (Aciandrena) palmyriae Wood, 2021, and A. (Aciandrena) xera Pisanty, 2022 and 2) A. (Graecandrena) schwarzi Warncke, 1975. In group 1, the propodeal triangle is entirely shagreened, leading to placement in the subgenus Aciandrena, whereas in group 2, the propodeal triangle has basal rugae, which along with the male genitalia leads to placement in subgenus Graecandrena. The propodeal triangle of the Iranian specimen has a narrow area with basal rugae, making it somewhat intermediate and resulting in its incertae sedis placement in the absence of genetic data. Comparison with photographs of the type of A. nitidicollis (Astafurova et al. 2022) shows that the Iranian specimen is conspecific due to the combination of its completely smooth scutum (completely shagreened in A. pavonia, partially shagreened in A. xera), its truncate labral process that lacks an apical notch (deeply notched in A. palmyriae), scutum with sparse punctures centrally, separated by> 2 puncture diameters (separated by 1 – 2 puncture diameters in A. schwarzi), broad facial foveae that occupy almost the entire space between the compound eye and the lateral ocellus (occupying ½ to ¾ of this space in all other species), and finally and most importantly, the relatively strongly and deeply punctate lateral parts of T 3 – 4, punctures separated by 1 – 2 puncture diameters (punctures weaker, sparser, or absent in all other species). Its presence in Iran is reminiscent of A. longiceps, which has a predominantly Central Asian distribution, but which was recently recorded in Iran for the first time (Radchenko et al. 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF7514DD9FBDA3BFE2AFE0C.taxon	distribution	Distribution Iran *, Kazakhstan.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CDA51DB44FF44FD68.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CD998DA17FE31FC3B.taxon	distribution	Distribution Turkey and Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CD9F3DAA2FCF5FAF5.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 4 ♂♂; Yasouj, Deli Bajak, Sepidar; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; ICPI • 1 ♂; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CD9F3DAA2FCF5FAF5.taxon	distribution	Distribution Greece, Turkey, Israel and the West Bank, Lebanon, Jordan, Syria, Turkey, Iran * (Gusenleitner & Schwarz 2002; Pisanty et al. 2018; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CD9ADD81DFAD5FD9D.taxon	distribution	Distribution Central and Eastern Europe eastwards to Russia (European part, Urals), Turkey, the Caucasus, Iran, and Turkmenistan (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CD99EDC7DFEE6F90B.taxon	materials_examined	Material examined IRAN • 1 ♂; West Azerbaijan Province, Tazenkendkaragolchange [? Tazeh Kand-e-Nosrat]; 15 May 2001; J. Prochazka leg.; OÖLM • 4 ♂♂, 1 ♀; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM. SYRIA • 1 ♀; Aleppo, Simeons-Kloster; 500 m a. s. l.; 19 Apr. 1988; L. Blank leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514CD99EDC7DFEE6F90B.taxon	distribution	Distribution	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514FD9F2DE02FBFDFD53.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province, Dasht Arjan; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 6 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; 11 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Yasouj, Kakan [Fars Province]; 14 Feb. 2011; A. Monfared leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF1514FD9F2DE02FBFDFD53.taxon	distribution	Distribution Hungary, North Macedonia, Romania, Bulgaria, Ukraine, Turkey, Cyprus, Israel and the West Bank, Lebanon, Jordan, Syria, Iran * (Schwenninger et al. 2015; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF2514FD9F5DA1BFDB6F85C.taxon	materials_examined	Material examined Holotypes AZERBAIJAN • ♀ (holotype of A. oulskii); Baku; ZMHB. IRAN • ♀ (holotype of A. oulskii roseotincta); Khuzistan, Haft Tepe [Hafttapeh], southeast Shush; 22 Mar. 1956; K. Warncke det.; R. Schäuffele leg.; OÖLM. Other material IRAN • 1 ♂, 1 ♀; Kašan [Kashan]; 3 May 1999; A. Ariana det.; K. Deneš leg.; OÖLM • 2 ♀♀; Kohgiluyeh and Boyer-Ahmad, Yasoj, Semeron [Semerun]; 1529 m a. s. l.; 2 May 2019; S. Dowlatkhan leg.; ICPI • 2 ♂♂, 2 ♀♀; Kohgiluyeh and Boyer-Ahmad, Yasoj, Semron [Semerun]; 1895 m a. s. l.; 25 Jan. 2019; Z. Mohammadi leg.; ICPI • 1 ♀; Kohgiluyeh and Boyer-Ahmad, Yasuj, Kakan; 2326 m a. s. l.; 14 Apr. 2018; S. Mahdi and R. Esfandiari leg.; ICPI • 1 ♀; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM • 1 ♂, 4 ♀♀; Persepolis env.; 9 May 1999; A. Ariana det.; K. Deneš leg.; OÖLM • 1 ♂; South, Bandar-e Bušehr; 11 May 1999; K. Deneš leg.; OÖLM • 1 ♂; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF2514FD9F5DA1BFDB6F85C.taxon	discussion	Remarks The use of this name has been confused, but was recently clarified by Wood (2021 b) after type inspection. Material described as A. oulskii roseotincta by Warncke is not sufficiently differentiated to merit subspecific status. See also Pisanty et al. (2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF2514FD9F5DA1BFDB6F85C.taxon	distribution	Distribution Syria, Turkey, Azerbaijan, Iran (Warncke 1975), possibly Afghanistan, but the species concept used by Warncke (1974 c) is unclear.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF3514ED93BD81DFC4BFBC0.taxon	materials_examined	Material examined ALBANIA • 1 ♀; N Arras; 3 Jun. 2013; K. Deneš leg.; OÖLM • 1 ♂; Tepelenë; 27 May 2013; K. Deneš leg.; OÖLM. IRAN • 2 ♂♂, 6 ♀♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Gilan Province, 15 km SE of Tutkabon; 1100 m a. s. l.; J. Halada leg.; OÖLM • 1 ♂, 1 ♀; Kurdistan Province, Paniran; 1450 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 670 m a. s. l.; M. Kafka leg.; OÖLM • 3 ♀♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; M. Kafka leg.; OÖLM • 1 ♂, 5 ♀♀; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF3514ED93BD81DFC4BFBC0.taxon	discussion	Remarks Unpublished barcode data suggest that A. panurgimorpha is a complex of at least two distinct species (G. Pisanty pers. comm.).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF3514ED93BD81DFC4BFBC0.taxon	distribution	Distribution Albania *, Bulgaria, Greece, Ukraine, Turkey, Cyprus, Israel and the West Bank, Lebanon, Jordan, Syria, Iran (Gusenleitner & Schwarz 2002; Wood et al. 2020 a; Wood 2021 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF3514ED9F4DD88FAB3F9E1.taxon	materials_examined	Material examined IRAN • 1 ♀; Tehran Province, Samgh Abad; 1900 m a. s. l.; 15 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF3514ED9F4DD88FAB3F9E1.taxon	discussion	Remarks Following the revised concept of Poecilandrena (Pisanty et al. 2022 b), A. paradisaea is hereby removed from Poecilandrena and considered incertae sedis. The curved tibial spurs and atypical genitalia clearly place this species outside the Andrena labiata and A. viridescens species group concepts.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF3514ED9F4DD88FAB3F9E1.taxon	distribution	Distribution Israel and West Bank, Turkey, and Iran * (Gusenleitner & Schwarz 2002; Pisanty et al. 2018).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF35141D9FCDF69FEFBFDBC.taxon	materials_examined	Literature records Ascher & Pickering (2021): Mazandaran; coastal plain between Chalus and Shahsavar; 20 m a. s. l.; 10 Apr. 1965; D. Baker leg.; SEMC; via GBIF occurrence 784988907. Material examined IRAN • 1 ♀; Yasouj, Isfahan Road, Tange Seriz; 1687 m a. s. l.; 8 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Mazandaran, 20 km S of Nowshahr, Weysser [Veysar]; 1400 m a. s. l.; 9 – 14 Jun. 1977; C. Holzschuh and F. Ressl leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF35141D9FCDF69FEFBFDBC.taxon	discussion	Remarks The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate the presence of A. parviceps in Iran, so this species is therefore not reported as new.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF35141D9FCDF69FEFBFDBC.taxon	distribution	Distribution Southern Europe, Turkey, the Caucasus, Iran, Turkmenistan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFC5141D9B8DB24FF44FC9F.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFC5141D9ADDA46FB14FAFF.taxon	discussion	Remarks As for A. bimaculata and A. tibialis, members of the subgenus Plastandrena are taxonomically highly challenging, with a great deal of variation and many described taxa. A deep molecular revision of the A. pilipes group is needed to establish whether multiple good taxa exist. Andrena pilipes s. lat. is commonly recorded in Iran, and therefore for brevity additional records are not detailed here.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFC5141D9ADDA46FB14FAFF.taxon	distribution	Distribution In a broad sense, Palearctic (Gusenleitner & Schwarz 2002; Ascher & Picking 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFC5140D8B7DC66FD7BFC3E.taxon	materials_examined	Material examined IRAN • 55 ♂♂; Azer. e Garbi. Prov. [West Azerbaijan Province], Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 2 ♂♂, 1 ♀; Kohgiluyeh and Boyer-Ahmad Province, Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun 2010; Mi. Halada leg.; OÖLM • 1 ♂; Fars Province, Komehr; 2380 m a. s. l.; 9 Jun 2010; Mi. Halada leg.; OÖLM. RUSSIA • 1 ♀; Сталингр. обл. [Stalingrad obl. = Volgograd Oblast], Тингута [Tinguta]; 5 Aug. 1952; Г. Викторов leg. [G. Viktorov]; OÖLM. TURKEY • 1 ♀; Mardin; 1000 m a. s. l.; 2 Jul. 1987; R. Hensen leg.; RMNH • 1 ♂, 25 ♀♀; Burdur, 20 km SW of Burdur; 940 m a. s. l.; 7 Jul. 2006; M. Halada and M. Kadlecová leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFC5140D8B7DC66FD7BFC3E.taxon	discussion	Remarks The presence of this species in southern Russia is surprising, as it was not reported from Russia in the recent revision of the subgenus (Ariana et al. 2009 a). What is more surprising is that the collecting locality of Tinguta is the locus typicus for Andrena (Brachyandrena) limonii Osytshnjuk, 1983, the holotype of which is a female from Volograd reg., Tinguta, 13 Aug. 1954, leg. Razumova (see Osytshnjuk 1983 b; Osytshnjuk et al. 2005). The specimen examined here was found in boxes of undetermined material at Linz that were part of the Warncke collection. It was part of a series of specimens of various species that were collected by the Russian entomologists G. Viktorov, D. Panfilov, and L. Zimina. These specimens were borrowed by Warncke from the Zoological Museum of Moscow University (Y. Astafurova pers. comm.) at some point and never identified. The specimen itself perfectly matches material of A. pinguis from Turkey and Iran, and the two taxa cannot be confused, as A. pinguis has the mesepisternum coarsely areolate, whereas it is smooth in A. limonii. Ariana et al. (2009 a) did not examine the female type (in the Moscow State University collection), but they did examine a male from the locus typicus collected 3 Aug. 1954, leg. Razumova, which would therefore not be a paratype, unless this is a typographical error and the specimen was collected on 13 Aug. 1954, in which case it would be part of the type series (see Osytshnjuk 1983 b for details of paratypes). The holotype should be examined to ensure that it corresponds to A. limonii sensu Ariana et al., though given the strong morphological differences it seems more likely that the two taxa simply (though unexpectedly) occur in sympatry.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFC5140D8B7DC66FD7BFC3E.taxon	distribution	Distribution Russia *, Turkey, Iran * (Ariana et al. 2009 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFD5140D985DAA4FBDCFB5E.taxon	materials_examined	Material examined IRAN • 2 ♀♀; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFD5140D985DAA4FBDCFB5E.taxon	distribution	Distribution Unclear because of synonymy with A. dorsata; probably distributed from north-western Africa throughout Europe to Turkey (Gusenleitner & Schwarz 2002; Hazir et al. 2014).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFD5140D9EADC1BFF44F85D.taxon	materials_examined	Material examined JORDAN • 1 ♂; 30 km N of Tafila [At-Tafilah]; 2 May 1996; Ma. Halada leg.; OÖLM. LIBYA • 1 ♀; Cyrenaika, Brega; 4 – 31 Mar. 1958; K. M. Guichard leg.; NHMUK. SAUDI ARABIA • 1 ♀; S of Riyadh, Al Kharj; 20 Mar. 1980; K. M. Guichard leg.; NHMUK. SYRIA • 1 ♀; NW of Aleppo, Afrin; 23 Jun. 2000; M. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFD5140D9EADC1BFF44F85D.taxon	discussion	Remarks The subspecies succinea will be elevated to species status in a subsequent publication dealing with the Andrena fauna of the Iberian Peninsula.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFD5140D9EADC1BFF44F85D.taxon	distribution	Distribution (subspecies succinea) Morocco, Algeria, Tunisia, Libya *, Egypt, Israel and the West Bank, Jordan *, Syria *, Saudi Arabia *, Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5143D99FD81DFD38FD87.taxon	materials_examined	Material examined IRAN • 4 ♀♀; Fars Province, 15 km S of Dasht Arjan; 2261 m a. s. l.; 2 – 6 May 2016; M. Obořil leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5143D99FD81DFD38FD87.taxon	distribution	Distribution Greece, Turkey, Iran * (Warncke 1975).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5143D9F8DB4FFC64FC30.taxon	distribution	Distribution Greece, Ukraine, Russia (European part), Turkey, Cyprus, Israel and the West Bank, Lebanon, Syria, the Caucasus, Iran (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5143D98DDABAFB8EF9C0.taxon	materials_examined	Material examined ARMENIA • 1 ♀; S of Vedi-Chozrov; 4 Jun. 2003; M. Múčka leg.; OÖLM. TURKEY • 1 ♀; Kars, 10 km E of Karakurt; 1500 m a. s. l.; 28 May 1983; K. Warncke leg.; OÖLM • 1 ♀; Hakkâri, S. Beytüşşebap; 1300 m a. s. l.; 13 Jun. 1984; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5143D98DDABAFB8EF9C0.taxon	discussion	Remarks The source of the Ascher & Pickering listing is unclear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate that A. quadrimaculata is present in northern Iran, though no specimens could be found in the Warncke collection. The species does occur in eastern Turkey and Armenia, so it is considered highly likely to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5143D98DDABAFB8EF9C0.taxon	distribution	Distribution Albania, Turkey, Armenia, Azerbaijan, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5142D9FEDF8AFB8DFE5C.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Yasouj, Deli Bajak, Sepidar; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; ICPI • 2 ♂♂, 21 ♀♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 31 Mar. – 10 Apr. 2021; E. Rostami leg.; ICPI • 1 ♂; Yasouj, Tang Konareh, Dasht-e Rum; 1858 m a. s. l.; 29 Mar. 2021; E. Rostami leg.; ICPI • 7 ♂♂, 25 ♀♀; Yasouj, Tange Tamoradi; 2247 m a. s. l.; 9 – 29 Mar. 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5142D9FEDF8AFB8DFE5C.taxon	discussion	Remarks Collected from Acer monspessulanum in the Yasouj area, supporting the morphological link with Andrena dividicincta Pisanty, 2022 (Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFE5142D9FEDF8AFB8DFE5C.taxon	distribution	Distribution Greece, Turkey, Iran * (Gusenleitner & Schwarz 2002; Rasmont et al. 2017).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFF5142D9B8DD94FC7FF880.taxon	materials_examined	Material examined IRAN • 2 ♂♂; Boyer-A. o. Kohg Prov., Kuh Gol, near Sisakht; 2500 m a. s. l.; 9 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Fars Province, 10 km E of Kazerum; 1990 m a. s. l.; 23 May 2014; J. Halada leg.; OÖLM • 1 ♂; Fars Province, Dast Arjan; 2260 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Gilan Province, Tutkabon near Roodbar; 16 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM • 8 ♀♀; Yasouj, 3 km after Davood Abad Village; 1898 m a. s. l.; 6 Apr. 2021; E. Rostami leg.; ICPI • 3 ♂♂; Yasouj, Deli Bajak, Sepidar; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 31 Mar. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Khahkaloon; 2037 m a. s. l.; 25 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Sisakht, Mishi Spring; 2341 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFF5142D9B8DD94FC7FF880.taxon	distribution	Distribution Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFF5142D9E5DB04FC3AFBEC.taxon	materials_examined	Material examined IRAN • 1 ♀; Elburz Mountains, Kandelouz; 2100 m a. s. l.; O. Sauša leg.; OÖLM • 1 ♀; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Pass SW of Chalus-Karaj / Mazandaran; 2100 m a. s. l.; 28 May 1978; M. Kraus leg.; OÖLM • 1 ♀; Khorasan, 15 km W of Bodjnurd; 1300 m a. s. l.; 30 May 1977; C. Holzschuh leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFF5142D9E5DB04FC3AFBEC.taxon	distribution	Distribution Spain, France, Switzerland, Italy, Ukraine, Turkey, Russia (European part, Urals, Western Siberia, Eastern Siberia), Iran, Turkmenistan, Uzbekistan, Kazakhstan, Tajikistan (Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFF5142D9BBDE48FF44F85D.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85145D987DB89FEB9FC75.taxon	distribution	Distribution Central Europe to Turkey, Lebanon, the Caucasus, Iran (Gusenleitner & Schwarz 2002; Boustani et al. 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85145D99FDAFDFC0EF98F.taxon	materials_examined	Material examined IRAN • 3 ♀♀; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 3 ♀♀; Lorestan Province, Tootmashour [Tut, Masur-e Abi]; 2000 m a. s. l.; 31 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; Shiraz-Yasouj road (15 km to Yasouj); 20 May 2011; A. Monfared leg.; ICPI • 1 ♂; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85145D99FDAFDFC0EF98F.taxon	discussion	Remarks Praz et al. (2022) synonymised A. similis Smith, 1849 with A. russula.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85145D99FDAFDFC0EF98F.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002; Praz et al. 2022).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85145D9F8D81DFB16FDC1.taxon	materials_examined	Material examined IRAN • 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Yasouj, Doposhteh, Dasht-e Rum; 2091 m a. s. l.; 31 Mar. 2021; E. Rostami leg.; ICPI • 1 ♂; Yasouj, Tang Konareh, Dasht-e Rum; 1858 m a. s. l.; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85145D9F8D81DFB16FDC1.taxon	distribution	Distribution Turkey, Israel and the West Bank, Lebanon, Jordan, Syria, Iran * (Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85144D998DF77FB4AFEE5.taxon	materials_examined	Material examined IRAN • 1 ♂; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Yasouj, Deli Awlad Alimomen; 2068 m a. s. l.; 30 Apr. 2021; E. Rostami leg.; ICPI • 2 ♀♀; Yasouj, Sisakht, Mishi Spring; 2341 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Yazd, Tezerjan Bagh-e Agha Seyyed; 2226 m a. s. l.; 5 May 2021; S. San leg.; ICPI • 2 ♀♀; Pass E Neyriz / Fars; 18 May 1978; K. Warncke leg.; OÖLM. JORDAN • 1 ♀; Petra; 14 May 1995; E. Scheuchl det.; K. Deneš leg.; SMNHTAU • 1 ♀; Petra [Wadi Musa]; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF85144D998DF77FB4AFEE5.taxon	distribution	Distribution Greece, Turkey, Cyprus, Golan Heights, Jordan *, Iran * (Warncke 1975; Pisanty et al. 2018).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95144DA65D863FF44FDB5.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95144D991DC09FB67F97D.taxon	materials_examined	Material examined IRAN • 9 ♂♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; Gilan Province, 15 km SE of Tutkabon; 1100 m a. s. l.; 9 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95144D991DC09FB67F97D.taxon	distribution	Distribution West Palearctic to Turkmenistan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95144D9AFDB33FAC7FB42.taxon	materials_examined	Material examined IRAN • 1 ♀; Lar, Fars; 18 Apr. 2012; S. Khazraie leg.; ICPI • 1 ♀; Yasouj; 1 – 30 Apr. 2015; M. Davoodi leg.; ICPI • 1 ♂; Yazd, Khezr abad Karkhane asphalt; 1317 m a. s. l.; 21 Mar. 2021; S. San leg.; ICPI • 1 ♂; Yazd, Kouch e talar e yazd; 1200 m a. s. l.; 1 Apr. 2020; S. San leg.; ICPI. JORDAN • 1 ♀; Wadi Rum; 4 – 5 May 1996; Mi. Halada leg.; OÖLM. OMAN • 1 ♂; Hajar Mountains, Ghubrah Canyon; 7 Feb. 2016; J. Monks leg.; NHMUK • 1 ♂, 1 ♀; NW of Bahla, Al Ayshi; 6 Mar. 2017; M. Snižek leg.; OÖLM • 1 ♂, 1 ♀; Samed Ashan, Asswareej; 1 Dec. 2019; A. Al Jahdhami leg.; NHMUK. YEMEN • 1 ♂; San’a [Sana’a]; 7,900 feet [ca 2400 m]; 2 – 9 Oct. 1937; Dr. C. Rathjens; NHMUK • 1 ♂; W. Aden Prot., Jebel Jihaf [Djabal Djihaf]; ca. 7100 feet [2100 m]; 1 – 31 Oct. 1937; H. Scott and E. B. Britton; NHMUK.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95144D9AFDB33FAC7FB42.taxon	distribution	Distribution Spain (Canary Islands), Morocco, Algeria, France (Corsica), Italy (Sardinia), Tunisia, Libya, Egypt, Israel and the West Bank, Jordan *, Saudi Arabia, Yemen *, Iraq, Iran *, United Arab Emirates, Oman *, Afghanistan, Pakistan, India (Warncke 1974 c; Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95147D9AADFFBFB0FFEE6.taxon	materials_examined	Material examined IRAN • 1 ♀; Gilan, 5 km E of Rudbar; 400 m a. s. l.; 8 Jun. 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFF95147D9AADFFBFB0FFEE6.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D985D86EFBC9FD2F.taxon	materials_examined	Material examined IRAN • 1 ♂, 1 ♀; Yasouj, Deli Bajak, Sepidar; 2181 m a. s. l.; 23 Apr. 2021; E. Rostami leg.; ICPI • 1 ♀; Yasouj, Kakan [Fars Province]; 30 Mar. 2010; M. Najafi leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D985D86EFBC9FD2F.taxon	distribution	Distribution Golan Heights, Lebanon, Turkey, Iran * (Wood et al. 2020 a; Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D98CDBD7FCBDFB51.taxon	materials_examined	Material examined IRAN • 1 ♀; Elburz Mountains, 60 km E of Minudasht; 26 May 2007; O. Sauša leg.; OÖLM • 1 ♂, 18 ♀♀; Golestan, 40 km E of Minudasht NP; 750 m a. s. l.; 1 Jun. 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D98CDBD7FCBDFB51.taxon	discussion	Remarks The relationship between Andrena sedentaria and A. (Micrandrena) dmitrii Osytshnjuk, 1993, which was described from the Turkmenistani side of the Kopet Dag mountain range, is unclear and should be investigated, as the latter may be a junior synonym. As noted by Gusenleitner & Schwarz (2002), female A. sedentaria are very similar to A. luscinia, but the tergal margins are shagreened and with scattered punctures, a trait mentioned in their comments on A. dmitrii. Type revision is required.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D98CDBD7FCBDFB51.taxon	distribution	Distribution Turkey and Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D9BDDC19FF44FA2D.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D996DCD4FADFF85C.taxon	materials_examined	Material examined TURKEY • 1 ♀; Hakkâri, S. Varegös / Mt. Sat; 1700 m a. s. l.; 15 Jun. 1984; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D996DCD4FADFF85C.taxon	discussion	Remarks The source of the listing of Ascher & Pickering is unclear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate that A. seminuda is present in northwestern Iran. Given that A. seminuda is found in eastern Turkey, it is considered highly likely to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFA5147D996DCD4FADFF85C.taxon	distribution	Distribution Central Europe to Russia (European part), Turkey, the Caucasus, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D9CCDC47FC52F85C.taxon	materials_examined	Material examined TURKEY • 1 ♂, 1 ♀; Hakkâri, Tal S Gevria-Pass; 3000 m a. s. l.; 4 Aug. 1986; S. Blank and K. Warncke leg.; OÖLM. Literature records Central Alborz, Kandavan Pass, nr Pol-e-Zanguleh; 2200 m a. s. l.; 17 Aug. 1967; D. Baker leg.; SEMC; via GBIF occurrence 784989096.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D9CCDC47FC52F85C.taxon	discussion	Remarks This material is almost certainly referable to the subspecies sischkai that is known from high mountains in the Balkan Peninsula, eastern Turkey, and the North Caucasus. It, along with the other ‘ subspecies’ of A. simillima, is likely to represent a good taxon (Wood et al. 2020 b; Wood 2021 b; Le Divelec 2021), but a revision is required.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D9CCDC47FC52F85C.taxon	distribution	Distribution (subspecies sischkai) Greece, Bulgaria, Russia (Teberda), Turkey *, Iran * (Warncke 1988).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D9DBDA89FCDDFA9F.taxon	materials_examined	Material examined IRAN • 17 ♀♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 2 ♀♀; Ilam Province, Sar Joob; 1930 m a. s. l.; 13 May 2016; M. Kafka leg.; OÖLM • 5 ♀♀; Kurdistan Province, Paniran; 1450 m a. s. l.; 14 May 2016; M. Kafka leg.; OÖLM • 7 ♀♀; Tehran Province, Samgh Abad; 1900 m a. s. l.; 16 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D9DBDA89FCDDFA9F.taxon	distribution	Distribution (sillata s. str.) Greece, Turkey, Israel and the West Bank, Lebanon, Jordan; subspecies histrionica in Turkey and Iran * only (Gusenleitner & Schwarz 2002; Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D99FD81DFF44FCC1.taxon	discussion	Remarks The taxonomic status and placement of A. shakuensis is completely unclear. Originally described in the subgenus Plastandrena (Popov 1949), Gusenleitner & Schwarz (2002) commented that the type lacks the rugose areolate propodeal triangle that helps characterise this subgenus, suggesting placement in the subgenus? Poliandrena instead. Osytshnjuk et al. (2005) placed A. shakuensis in the subgenus Campylogaster, noting the squamous hairs on the scutum in the female sex. As the former subgenus Poliandrena has been found to be strongly polyphyletic, and since most taxa placed in Campylogaster probably do not belong there (Pisanty et al. 2022 b), type revision is required to establish the identity of A. shakuensis and its phylogenetic placement.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFFB5146D99FD81DFF44FCC1.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE45159D995D81DFE2CFC1D.taxon	discussion	Remarks The taxonomic status of A. skorikovi is unclear and should be resolved. It is known from a single female specimen collected in southern Iran (Shellali near Dizful, Arabistan [now Dezful, Khuzestan Province]) on 4 March 1904. In his description, Popov compared A. skorikovi to A. iranella, noting a strong similarity, but giving characters related to the density of tergal puncturing. Gusenleitner & Schwarz (2002) remark on the strong structural similarity, noting that the dark coloured A. skorikovi may represent the first generation of the red-coloured A. iranella (citing a specimen in their possession from July). However, Osytshnjuk et al. 2005 write that A. iranella flies in April and May, and Popov listed specimens from March – May in the type series; further collection is required to establish whether A. iranella and A. skorikovi are distinct taxa.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE45159D995D81DFE2CFC1D.taxon	distribution	Distribution Iran (Popov 1940).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE45159D986DC37FA9FF85D.taxon	materials_examined	Material examined IRAN • 2 ♀♀; Yazd, Abarkooh, Faraghe; 1698 m a. s. l.; 17 Mar. 2021; S. San leg.; ICPI • 1 ♀; Yazd, Hossein Abad to Banadak Sadat; 2067 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 2 ♀♀; Yazd, Mehriz, Bagh e Safa Khanoom; 1500 m a. s. l.; 7 Mar. 2020; S. San leg.; ICPI • 9 ♀♀; 30 km S of Yazd, Mt Sir Kuh; 7 May 1999; K. Deneš leg.; OÖLM • 1 ♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 22 ♂♂, 59 ♀♀; Persepolis env.; 9 May 1999; K. Deneš leg.; OÖLM • 2 ♂♂, 7 ♀♀; Tehran env.; 2 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE45159D986DC37FA9FF85D.taxon	discussion	Remarks The A. spreta complex is highly taxonomically challenging, with currently no clear interpretation (Gusenleitner & Schwarz 2002). A broad interpretation is taken here; much more taxonomic work is needed, including a molecular revision. The taxon A. (Micrandrena) aiderensis Osytshnjuk, 1993 was described from Aýdere on the Turkmenistani side of the Kopet Dag. This taxon should be included in future revisions of the group as it may be A. spreta (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE45159D986DC37FA9FF85D.taxon	distribution	Distribution Unclear, but in the opinion of the lead author there is a broad pan-Mediterranean A. spreta taxon.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE45159D9BBDAC6FDECFB4E.taxon	distribution	Distribution Morocco, Algeria, Tunisia, Libya, Egypt, Israel and the West Bank, Jordan, Syria, Iran (Gusenleitner & Schwarz 2002; Wood et al. 2020 b).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D9FEDF03FB12F85C.taxon	materials_examined	Material examined IRAN • 1 ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Tehran Province, 6 km N of Gačsar [Gachsar]; 29 May – 2 Jun. 2002; J. Prochazka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D9FEDF03FB12F85C.taxon	distribution	Distribution For A. symphyti Schmiedeknecht, 1883 in a broad sense, Europe to Turkey, the Levant, the Caucasus, Iran (Gusenleitner & Schwarz 2002; Scheuchl & Willner 2016; Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D980D81DFC9DFD6D.taxon	discussion	Remarks The source of the Ascher & Pickering listing is unclear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate a single recorded locality of A. stigmatica in northern Iran. No specimens could be found in the Warncke collection. The species is included in this list, but further investigation is required.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D980D81DFC9DFD6D.taxon	distribution	Distribution Crimea, Iran, Turkmenistan (Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D9F2DA14FDF1FAF4.taxon	materials_examined	Material examined ARMENIA • 1 ♀; Bjurakan [Byurakan] env.; 1500 m a. s. l.; 5 Jun. 1989; J. Strejček leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D9F2DA14FDF1FAF4.taxon	discussion	Remarks The original records supporting these listings are not clear. The distribution maps of Warncke (Gusenleitner & Schwarz 2002) indicate that A. stoeckhertella is present in northern Iran. Given its presence in the Caucasus, the taxon is considered to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D9F2DA14FDF1FAF4.taxon	distribution	Distribution Ukraine, Russia (European part), Turkey, Georgia, Armenia *, Azerbaijan, Iran (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE55158D986DC73FF44FA45.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD9A4DCE1FEFBF8A0.taxon	distribution	Distribution Turkey, Georgia, Armenia, Azerbaijan, Iran, Turkmenistan (Gusenleiter & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD9E7D8AFFC54FD6D.taxon	materials_examined	Material examined IRAN • 1 ♂; Lorestan Province, Dorud Lanjaban env.; 960 m a. s. l.; 10 May 2016; M. Kafka leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD9E7D8AFFC54FD6D.taxon	distribution	Distribution Turkey, Israel and the West Bank, Syria, Iran * (Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD9A4D81DFE31FE20.taxon	distribution	Distribution Turkey and Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD996DA14FBB8FA7A.taxon	materials_examined	Material examined IRAN • 1 ♂; Azerbaidjan [Province], 30 km SE of Shahpur; 1500 m a. s. l.; 4 Jun. 1978; C. Holzschuh leg.; OÖLM. TURKEY • 1 ♂, 2 ♀♀; Hakkâri, 5 km N of Oramar [river]; 1450 m a. s. l.; 16 Jun. 1984; K. Warncke leg.; OÖLM • 1 ♂; Hakkâri, Suvari Halil-Pass; 2300 m a. s. l.; 14 Jun. 1984; K. Warncke leg.; OÖLM • 9 ♂♂, 1 ♀; Hakkâri, S. Varegös / Mt Sat; 15 Jun. 1984; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD996DA14FBB8FA7A.taxon	discussion	Remarks Full distribution unclear because of historical synonymy with A. concinna Smith, 1853 (see Gusenleiter & Schwarz 2002). Probably restricted to the Caucasus and immediately neighbouring areas. It is also the likely senior synonym of A. carinata Morawitz, 1877, which was also described from the Caucasus.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515BD996DA14FBB8FA7A.taxon	distribution	Distribution Turkey, Georgia, Armenia, Azerbaijan, Iran (Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515AD9ACDE2EFE06FE53.taxon	materials_examined	Material examined IRAN • 1 ♀; Yasouj, Sisakht, Mishi Spring; 2341 m a. s. l.; 10 Apr. 2021; E. Rostami leg.; ICPI • 2 ♀♀; Yasouj, Tange Tamoradi; 2247 m a. s. l.; 21 – 29 Mar. 2021; E. Rostami leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE6515AD9ACDE2EFE06FE53.taxon	distribution	Distribution Southern Europe to Central Asia, Turkey, Golan Heights, Lebanon, Iran (Gusenleiter & Schwarz 2002; Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE7515AD987DB1BFAB7FB85.taxon	materials_examined	Material examined IRAN • 1 ♀; Rahatabad, Tehran; 18 Apr. 2011; A. Monfared leg.; ICPI • 1 ♂, 2 ♀♀; Yasouj, Vezg [Kūh-e Vezg]; 19 Apr. 2011; H. Elahi and M. Mesavian leg.; ICPI • 1 ♂, 1 ♀; Yazd, Abarkooh, Faraghe; 1698 m a. s. l.; 6 Mar. 2021; S. San leg.; ICPI • 2 ♀♀; Yazd, Tezerjan Bagh-e Agha Seyyed; 2226 m a. s. l.; 12 Jun. 2020; S. San leg.; ICPI • 1 ♀; Yazd, Tezerjan Bagh-e Agha Seyyed; 2226 m a. s. l.; 13 May 2021; S. San leg.; ICPI • 1 ♂; Yazd, Tezerjan, paen tar az Seyyed Mahmood; 1993 m a. s. l.; 6 Mar. 2020; S. San leg.; ICPI • 1 ♀; Yazd, Yazd road to Sanij; 2621 m a. s. l.; 27 Mar. 2019; S. San leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE7515AD987DB1BFAB7FB85.taxon	distribution	Distribution Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE7515AD992DD4DFC89FA76.taxon	distribution	Distribution Spain, Morocco, Algeria, Libya, Egypt, Israel, Syria according to the broad interpretation of Warncke (Gusenleitner & Schwarz 2002). Kratochwil (2015) split A. tiaretta into three taxa, though not all individuals (including Iranian material) could be placed within this system. In the absence of genetic data, a broad interpretation of this taxon is taken here.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE7515DD99FDCFEFB0CFEE6.taxon	materials_examined	Material examined IRAN • 1 ♂; Yasouj, Abshar; 2 – 8 Mar. 2011; H. Elahi and M. Mesavian leg.; ICPI • 1 ♂; Yasouj, Vezg; 19 Apr. 2011; H. Elahi and M. Mesavian leg.; ICPI.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE7515DD99FDCFEFB0CFEE6.taxon	discussion	Remarks The two male specimens examined here have the genital capsules of A. tibialis, but S 8 is noticeably broadened apically, and should be recorded as ‘ aff. tibialis ’. As noted by Gusenleitner & Schwarz (2002), A. tibialis shows considerable variation across its range, particularly in Turkey and the Middle East. A revision of this taxon is needed, preferably with molecular techniques.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE7515DD99FDCFEFB0CFEE6.taxon	distribution	Distribution In a broad sense, Palearctic (Gusenleitner & Schwarz 2002; Scheuchl & Willner 2016).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515DD992DC07FE57F973.taxon	materials_examined	Material examined IRAN • 1 ♀; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂; Pass SW of Chalus-Karaj / Mazandaran; 2470 m a. s. l.; 28 May 1978; K. Warncke det.; M. Kraus leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515DD992DC07FE57F973.taxon	distribution	Distribution Greece, Turkey, Cyprus, Israel and the West Bank, Iran * (Gusenleitner & Schwarz, 2002). Records from Lebanon, Jordan, and Syria reported by Wood et al. (2020 a) were incorrectly identified female A. cypria. Andrena torda is likely still present in these countries, but must be confirmed with unambiguously distinctive male material.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515DD9FADA1AFB84FB5F.taxon	materials_examined	Material examined IRAN • 1 ♀; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM • 1 ♂; Pass SW Chalus-Karaj / Mazandaran; 2470 m a. s. l.; 28 May 1978; K. Warncke det.; M. Kraus leg.; OÖLM. Literature records Centr. Alborz, Kandavan Pass, 8 km above Siahbishe; 2400 m a. s. l.; 10 Jun. 1967; D. Baker leg.; SEMC; via GBIF occurrence 658462817.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515DD9FADA1AFB84FB5F.taxon	distribution	Distribution Turkey, Armenia, Iran * (Osytshnjuk et al. 2008; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515DD934D86EFB6AFD52.taxon	materials_examined	Material examined IRAN • 4 ♀♀; Bakhtiyari Province, 20 km SE of Lordegan; 31 May 2014; J. Halada leg.; OÖLM • 2 ♀♀; Gilan Province, 20 km W of Astara; 640 m a. s. l.; 17 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂, 5 ♀♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 1 ♀; Lorestan Province, 10 km SW of Dorud; 1520 m a. s. l.; 20 May 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515DD934D86EFB6AFD52.taxon	distribution	Distribution Turkey, Israel and the West Bank, Jordan *, Syria *, Iran * (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515CD986DFFBFE4BFE58.taxon	materials_examined	Material examined IRAN • 1 ♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♂; Golestan Province, Gorgan env. [36 ° 82 ′ N, 54 ° 28 ′ E]; 7 Jun. 2009; J. Voříšek leg.; OÖLM • 1 ♀; Golestan, 70 km E of Minudasht; 1050 m a. s. l.; 12 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♀; Khorasan Province, 10 km W of Raz; 1200 m a. s. l; 27 May 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE0515CD986DFFBFE4BFE58.taxon	distribution	Distribution Central Europe to Central Asia and the Middle East (Warncke 1974 c; Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515CD9E7DF24FCF1F885.taxon	distribution	Distribution Golan Heights, Turkey, and Iran (Pisanty et al. 2022 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515CD98DDB01FBBFFBAA.taxon	materials_examined	Material examined IRAN • 1 ♂; Yasouj, Abshar; 2 Mar. 2011; H. Elahi and M. Mesavian leg.; ICPI • 1 ♂; Yasouj, Tange Tamoradi; 2247 m a. s. l.; 21 Mar. 2021; E. Rostami leg.; ICPI. Literature records Mazandaran; coastal plain between Chalus and Shahsavar; 20 m a. s. l.; 6 Mar. 1968; D. Baker leg.; SEMC; via GBIF occurrence 784989671.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515CD98DDB01FBBFFBAA.taxon	distribution	Distribution Not completely clear because of problems separating females from those of A. scotica Perkins, 1916 (see below), but it appears to be distributed across Europe into north-western Africa, Turkey, Lebanon, Golan Heights, Iran (Gusenleitner & Schwarz 2002; Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515CD9E6DD53FB0FF9B3.taxon	materials_examined	Material examined IRAN • 1 ♂; Chalus, Marzan Abad; 28 Apr. 1995; M. Kafka leg.; OÖLM • 1 ♀; Mazandaran Province, Kojuk [36 ° 23 ′ N, 51 ° 40 ′ E?; 7 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515CD9E6DD53FB0FF9B3.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515FD9FDDE4EFEE8FECC.taxon	materials_examined	Material examined IRAN • 1 ♀; Persepolis env.; 9 May 1999; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE1515FD9FDDE4EFEE8FECC.taxon	distribution	Distribution South-eastern Europe, Turkey, Cyprus, Israel and the West Bank, Lebanon, Jordan, Syria, Iran * (Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE2515FD9C6DDD8FB3BF886.taxon	materials_examined	Material examined IRAN • 1 ♂; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM. JORDAN • 1 ♀; 15 km W of Jerash, Dibbin; 2 May 2006; K. Deneš leg.; OÖLM • 1 ♂, 3 ♀♀; 20 km N of Karak; 1000 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 2 ♀♀; 30 km N of Tafila [At-Tafilah]; 2 May 1996; Ma. Halada leg.; OÖLM • 2 ♀♀; Ajlun S of Anjara; 27 Apr. 2002; M. Snižek leg.; OÖLM • 1 ♀; Amman; 26 Mar. 1986; M. Kraus leg.; OÖLM • 6 ♀♀; Irbid, Saham vill.; 19 – 25 Apr. 2003; I. Pljushtch leg.; OÖLM • 2 ♀♀; North Shuna; 20 Apr. 1996; M. Halada leg.; OÖLM. SYRIA • 2 ♂♂; 8 km N of Shaykh Miskin [Al Sheikh Maskin]; 28 Mar. 1994; S. Becvar leg.; OÖLM • 1 ♀; SW of Aleppo; 700 m a. s. l.; 7 Apr. 1988; L. Blank leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE2515FD9C6DDD8FB3BF886.taxon	distribution	Distribution Turkey, Israel and the West Bank, Jordan *, Syria *, Iran * (Scheuchl & Hazir 2012).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE2515FD9BDD8B5FB16FB17.taxon	materials_examined	Material examined IRAN • 1 ♂, 1 ♀; 50 – 70 km E of Minudasht, Golestan Forest; 450 – 700 m a. s. l.; 23 Apr. 1978; Holzschuh leg.; OÖLM. SYRIA • 1 ♀; Apamea; 29 Apr. 1995; K. Deneš leg.; OÖLM • 1 ♀; Ganawat [Qanawat]; 16 May 1995; K. Deneš leg.; OÖLM • 4 ♀♀; Jisr al-Shughur; 26 May 1996; Ma. Halada leg.; OÖLM. TURKEY • 1 ♂; Hakkâri, 5 km N of Oramar [river]; 1450 m a. s. l.; 11 Jun. 1981; K. Warncke leg.; OÖLM • 1 ♀; Şemdinli, Hakkâri; 1700 m a. s. l.; 12 Jun. 1981; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE2515FD9BDD8B5FB16FB17.taxon	discussion	Remarks Molecular revision of the polita group of Ulandrena is required, particularly the red-marked species that includes A. unicincta, as additional cryptic taxa may be present.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE2515FD9BDD8B5FB16FB17.taxon	distribution	Distribution Turkey, Israel and the West Bank, Lebanon, Jordan, Syria *, Iran (Wood et al. 2020 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE2515EDA4DDE4FFB69FECD.taxon	distribution	Distribution Europe, Russia (European part, Urals, Western Siberia, Eastern Siberia), the Caucasus, Iran, Central Asia (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008; Proshchalykin et al. 2017 a).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE3515ED980DACAFD47F9E8.taxon	materials_examined	Material examined IRAN • 1 ♀; East Azerbaijan Province, Sis, 10 km E of Shabestar; 1540 m a. s. l.; 19 Jun. 2010; Mi. Halada leg.; OÖLM • 1 ♂; Esfahan [Isfahan], Ardestan, Rahmatabad; 23 Jun. 2018; O. Mazlum leg.; OÖLM • 2 ♀♀; Fars Province, Dasht Arjan; 2040 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 2030 m a. s. l.; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♂, 3 ♀♀; Gilan Province, 15 km SE of Tutkabon; 1100 m a. s. l.; 9 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Kohgiluyeh and Boyer-Ahmad, Yasoj, Naregah; 1790 m a. s. l.; 12 May 2019; T. M. Mohamadnejad leg.; ICPI • 1 ♂; Kohgiluyeh and Boyer-Ahmad, Yasoj, Park Velayat; 1829 m a. s. l.; 17 Jun. 2018; Z. Samadi leg.; ICPI • 3 ♂♂; Kurdistan Province, Paniran; 1450 m a. s. l.; 14 May 2016; M. Kafka leg.; OÖLM • 3 ♂♂, 1 ♀; Mazandaran Prov., 10 km N of Gashar [Gachsar]; 2300 – 2700 m a. s. l.; 7 Jun. 2014; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE3515ED980DACAFD47F9E8.taxon	distribution	Distribution West Palearctic (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE3515ED9A4D8B7FB2FFC00.taxon	materials_examined	Material examined IRAN • 3 ♂♂; West Azerbaijan Province, Serou; 1650 m a. s. l.; 28 May 2010; Mi. Halada leg.; OÖLM • 1 ♀; Gilan Province, 15 km SE of Tutkabon; 1100 m a. s. l.; 9 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Gilan Province, 5 km E of Rudbar; 400 m a. s. l.; 8 Jun. 2014; J. Halada leg.; OÖLM • 1 ♀; Yasouj; 7 May 2012; A. Arsalannia leg.; OÖLM • 1 ♀; Yasouj, Kakan [Fars Province]; 22 Apr. 2011; A. Monfared leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE3515ED9A4D8B7FB2FFC00.taxon	distribution	Distribution West and Central Palearctic (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE35151D9A9DF91FD1EFE54.taxon	materials_examined	Material examined IRAN • 1 ♂, 3 ♀♀; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 2 ♂♂, 74 ♀♀; Persepolis env.; 9 May 1999; K. Deneš leg.; OÖLM • 2 ♂♂, 2 ♀♀; Yasouj; 24 May 2017; F. Arajpour leg.; ICPI • 1 ♀; Yasouj, Choram; 6 May 2011; S. Zarifi leg.; ICPI. TAJIKISTAN • 2 ♀♀; Vose, 120 km SE of Dushanbe; 12 May 1991; J. Halada leg.; OÖLM. UZBEKISTAN • 3 ♂♂; 80 km SE of Karschi [Qarshi]; 31 Apr. 1991; J. Halada leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE35151D9A9DF91FD1EFE54.taxon	distribution	Distribution West Palearctic to Russia (European part), Turkmenistan, Uzbekistan *, Tajikistan * (Gusenleitner & Schwarz 2002; Ascher & Pickering 2021).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEC5151D985DD71FE68F978.taxon	materials_examined	Material examined IRAN • 1 ♀; West [West Azerbaijan Province], Češme Bigar env. [Çeşmə, or Cheshmeh, 38.594 ° N, 45.119 ° E]; 14 May 1999; K. Deneš leg.; OÖLM. JORDAN • 1 ♀; 15 km E of Petra; 26 Apr. 2008; K. Deneš leg.; OÖLM • 1 ♀; 15 km W of Madaba; 760 m a. s. l.; 27 Apr. 2006; K. Deneš leg.; OÖLM • 2 ♀♀; Al Karak env. [Kerak]; 6 Apr. 2013; M. Snižek leg.; OÖLM • 8 ♀♀; Irbid, Saham vill.; 19 Apr. 2003; I. Pljushtch leg.; OÖLM • 1 ♂; Ma’an, Wadi Rum; 900 m a. s. l.; 17 Mar. 1988; L. Blank leg.; TJWC. SYRIA • 1 ♀; Apamea; 29 Apr. 1995; K. Deneš leg.; OÖLM • 5 ♀♀; Bosra; 3 May 1995; K. Deneš leg.; OÖLM • 1 ♀; Euphrat, Dura Europos; 10 Apr. 2001; J. Plass leg.; OÖLM • 1 ♀; Maalula [Maaloula]; 17 May 1995; K. Deneš leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEC5151D985DD71FE68F978.taxon	distribution	Distribution Turkey, Israel and the West Bank, Jordan *, Syria *, Iran *, Turkmenistan (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEC5151D9BFDFE5FF44F85D.taxon	distribution	Distribution Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEC5151D98BDB12FCFBFB8B.taxon	materials_examined	Material examined IRAN • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM. Literature records Ascher & Pickering (2021): Mazandaran; coastal plain between Chalus and Shahsavar; 20 m a. s. l.; 2 May 1965; D. Baker leg.; SEMC; via GBIF occurrence 784990358.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEC5151D98BDB12FCFBFB8B.taxon	discussion	Remarks The presence of A. viridescens in Iran was indicated by the distribution maps of Warncke (Gusenleitner & Schwarz 2002), so it is not treated as new for the country.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEC5151D98BDB12FCFBFB8B.taxon	distribution	Distribution Europe to Turkey, Iran (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFED5153D9A5D81CFCC2FCF4.taxon	description	Figs 175 – 176	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFED5153D9A5D81CFCC2FCF4.taxon	materials_examined	Material examined Holotype TURKEY • ♂ (holotype of Andrena zostera); Diyarbakır; 15 – 16 Apr. 1972; K. Warncke leg.; OÖLM. Other material IRAN • 1 ♂, 1 ♀; Fars Province [Kohgiluyeh and Boyer-Ahmad Province], Yasuj [Yasouj], Sarb-e Taveh [Sarab-e Taveh]; 4 May 2016; M. Kafka leg.; OÖLM • 1 ♀; Ilam Province, Abda Man, Dinar Gaouh; 1830 m a. s. l.; 12 May 2016; M. Kafka leg.; OÖLM • 3 ♂♂; Kašan [Kashan]; 3 May 1999; K. Deneš leg.; OÖLM • 1 ♀; Khuzistan, 30 km SE of Shush Ufer am Diss; 23 Mar. 1956; K. Warncke det.; R. Schäuffele leg.; OÖLM • 1 ♀; Persepolis env.; 9 May 1999; K. Deneš leg.; OÖLM • 3 ♂♂; Yazd, Hanza to Mehriz; 1928 m a. s. l.; 22 Mar. 2021; S. San leg.; ICPI • 8 ♂♂, 16 ♀♀; Yazd, Mehriz, Shohaday gomnam; 1500 m a. s. l.; 23 Mar. 2021; S. San leg.; ICPI • 4 ♀♀; Yazd, Mehriz esmat abad; 1500 m a. s. l.; 7 Mar. 2021; S. San leg.; ICPI • 1 ♀; Yazd, Mehriz, Bagh e Safa Khanoom; 1500 m a. s. l.; S. San leg.; ICPI • 2 ♀♀; Yazd, Mehriz, Khiaban e Valieasr, Kouch e sheikh kahn; 1502 m a. s. l.; 4 May 2020; S. San leg.; ICPI. IRAQ • 1 ♂; Mosul, edges of a river; 7 Apr. 1988; Olejníček leg.; OÖLM. SYRIA • 6 ♂♂; Damascus airport; 29 Mar. 1988; L. Blank leg.; MSC (genital capsule illustrated Fig. 175). TURKMENISTAN • 5 ♀♀; Kara-Kala; 29 Apr. – 3 May 1989; S. Bečvář leg.; OÖLM • 2 ♂♂; Ashabad [Ashgabat]; 20 – 22 Apr. 1989; S. Bečvář leg.; OÖLM. UZBEKISTAN • 1 ♂; 80 km SE of Karschi [Qarshi]; 31 Apr. 1991; J. Halada leg.; OÖLM • 1 ♂; Derbent, 13 km SE of Karschi [Qarshi]; 1450 m a. s. l.; 2 May 1991; J. Halada leg.; OÖLM (genital capsule illustrated Fig. 176).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFED5153D9A5D81CFCC2FCF4.taxon	discussion	Remarks Andrena zostera is very close to A. falcinella but can be separated by its genital capsule and weaker scutal puncturing in the female sex. In Central Asia, A. subsmaragdina is easily separable from other members of the former subgenus Carandrena (= Notandrena, see Pisanty et al. 2022 b) because it is the only member in this region with a yellow clypeus in the male sex. Comparison of the genital capsule to that of material from the Levant to Uzbekistan (Figs 175 – 176, also illustrated in Osytshnjuk 1984; Osytshnjuk et al. 2005) shows that they are identical, and are therefore synonymised. This synonymy joins the West and Central Palearctic populations of this taxon.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFED5153D9A5D81CFCC2FCF4.taxon	distribution	Distribution Israel and the West Bank, Syria *, Turkey, Azerbaijan, Iraq *, Iran, Turkmenistan, Uzbekistan, Tajikistan (Warncke 1975; Osytshnjuk 1984; Osytshnjuk et al. 2005).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEE5153D986DD47FB71FA75.taxon	discussion	Reason for exclusion Andrena curvungula is present in eastern Turkey (Gusenleitner & Schwarz 2002), and hence is likely present in northwestern Iran, but based on the photographs provided by Allahverdi et al. (2015) their specimens are not A. curvungula and are probably A. transitoria. Specimens were also collected from Brassicaceae, whereas A. curvungula is a specialist of Campanula L. (Campanulaceae).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEE5153D9BCDCFEFDD3F887.taxon	discussion	Reason for exclusion Andrena falsifica is a species of cool, Potentilla - rich (Rosaceae) grassland environments. It has never been recorded from Turkey or the Caucasus (Gusenleitner & Schwarz 2002), and so its presence in Iran would be a notable range extension.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEE5152D9BADE4FFAC7FEE7.taxon	discussion	Reason for exclusion This taxon is restricted to the Balkan Peninsula (Schönitzer et al. 1995; Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEF5152D992DD7FFC9EF99E.taxon	discussion	Reason for exclusion Andrena fuscipes is a specialist of Ericaceae, and has a broad Atlantic distribution across Europe and Russia (European part and Urals, Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2005; Proshchalykin et al. 2017 a), typically in areas with Calluna vulgaris (L.) Hull (Ericaceae). The specimens recorded by Allahverdi et al. (2015) were also collected in May, whereas A. fuscipes (along with other species of Cnemidandrena Hedicke, 1933) flies in the summer, predominantly in July and August. The specimen in the presented photographs appear to be A. flavipes.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEF5152D9A4D869FD37FD45.taxon	discussion	Reason for exclusion This taxon is known only from Greece, Crimea, and western Turkey (Gusenleitner & Schwarz 2002). It has never been recorded from eastern Turkey or the Caucasus. Its presence in Iran is considered unlikely, and the record presented by Allahverdi et al. (2016) may well be a different species of A. (Nobandrena), such as A. comptaeformis or A. nobilis.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEF5152D9F3DA0EFC09FBF6.taxon	discussion	Reason for exclusion This taxon is found in East Asia (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEF5155D9B1DF47FDCCFD5C.taxon	materials_examined	Material examined TURKEY • 1 ♂; Hakkâri Şivelan; 18 May 1975; K. Warncke leg.; OÖLM • 1 ♂; Hakkâri, 10 km SW of Yüksekova; 10 Jun. 1981; K. Warncke leg.; OÖLM • 1 ♀; Hakkâri, 25 km NE of Hakkâri; 2200 m a. s. l.; 30 May 1980; K. Warncke leg.; OÖLM • 2 ♀♀; Hakkâri, 25 km NW of Yüksekova; 2200 m a. s. l.; 30 May 1980; K. Warncke leg.; OÖLM • 1 ♂; Hakkâri, S. Varegös / Mt Sat; 1700 m a. s. l.; 15 Jun. 1984; K. Warncke leg.; OÖLM • 3 ♂♂; Hakkâri, S. Varegös / Mt Sat; 1800 m a. s. l.; 17 Jun. 1984; K. Warncke leg.; OÖLM • 2 ♂♂, 2 ♀♀; Karakurt / Arastal; 22 May 1975; K. Warncke leg.; OÖLM • 2 ♂♂; Kars, 10 km E of Karakurt; 1500 m a. s. l.; 28 May 1983; K. Warncke leg.; OÖLM • 53 ♂♂, 1 ♀; Kars, 20 km W of Sarikamis [Sarıkamış]; 2150 m a. s. l.; K. Warncke and M. Schwarz leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEF5155D9B1DF47FDCCFD5C.taxon	discussion	Reason for exclusion Andrena gravida is a species primarily of deciduous woodland in Europe to western Turkey (Gusenleitner & Schwarz 2002). Additionally, the taxon is present in eastern Turkey (Kars and Hakkâri provinces) based on previously unidentified specimens examined by the lead author that were collected by Klaus Warncke and Maximillian Schwarz (Linz). However, it is easy to confuse with other members of the former subgenus Zonandrena (= Melandrena Pérez, 1890, see Pisanty et al. 2022 b) such as A. gazella, which is present in Iran. The records presented by Allahverdi et al. (2016) also include specimens collected from late July, which is inconsistent with the ecology of A. gravida, which is univoltine and flies in the spring only, with the latest recorded date for examined Turkish specimens being June 17 th. Given the presence of this taxon in eastern Turkey, it may well be present in northwestern Iran, but this remains to be confirmed.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE85155D9ADDA19FD2EFBDC.taxon	materials_examined	Literature records Centr. Alborz, Kandavan Pass, 8 km above Siahbishe; 2400 m a. s. l.; 7 Jun. 1966; D. Baker leg.; SEMC; via GBIF occurrence 686218978.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE85155D9ADDA19FD2EFBDC.taxon	discussion	Reason for exclusion Andrena impunctata has an eastern range limit in central Turkey; it has never been recorded from eastern Turkey or the Caucasus (Gusenleitner & Schwarz 2002; Osytshnjuk et al. 2008). It is therefore considered unlikely to be present in Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE85155D991DF9AFC0FF85C.taxon	discussion	Reason for exclusion Andrena magunta is known from south-eastern Europe, western Turkey, Lebanon, and Israel (Gusenleitner & Schwarz 2002; Pisanty et al. 2018; Wood et al. 2020 a). It has never been recorded from eastern Turkey or the Caucasus. Given the very high difficulty of identifying Turkish and Middle Eastern Micrandrena, this record must be treated as unconfirmed.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE85155D99DDD99FEC6F9DD.taxon	materials_examined	Literature records Ascher & Pickering (2021): Mazandaran, Neka (25 km E of Sari); 7 Apr. 1968; D. Baker leg.; SEMC; via GBIF occurrence 684782674.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE85155D99DDD99FEC6F9DD.taxon	discussion	Reason for exclusion It is not clear whether the specimens collected from Iran by Baker were males or females. April is very early for this taxon, which in Europe typically flies in June and July. Given the extreme taxonomic challenge associated with the morphological identification of members of the subgenus Taeniandrena (Praz et al. 2022), and given that several taxa found in eastern Turkey were described after the capture of this specimen (e. g., A. producta Warncke, 1973, A. hova Warncke, 1975, A. solitaria Warncke, 1975), this taxon cannot be confirmed from Iran until male specimens with their distinctive genital capsule have been examined.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE95154D9BDDD73FAC1FA76.taxon	discussion	Reason for exclusion Based on the photographs provided by Allahverdi et al. (2015), their specimens are a species of A. (Brachyandrena). As was the case for the reported records of A. curvungula, their specimens were also collected from Brassicaceae, whereas A. pandellei is a specialist of Campanula (Campanulaceae).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE95154D9F9D81DFAC5FCB7.taxon	discussion	Reason for exclusion Andrena nanaeformis was described from Ukraine, and is sparingly distributed from east Germany and Austria eastwards to Russia (European part, Urals, Western Siberia, Eastern Siberia; Schmid-Egger & Scheuchl 1997; Gusenleitner & Schwarz 2002; Proshchalykin et al. 2017 a). It therefore has a distribution across the Eurasian cold steppe; Warncke did not record this taxon from the steppe of Turkey. Instead, he described two very similar species, A. luscinia and A. sedentaria (see above), that are found on the forest steppes of Turkey and Iran. Because of this different ecological distribution, we consider A. nanaeformis to be unconfirmed from Iran until this specimen can be carefully compared to Warncke’s type material.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE95154D99EDA3BFDEEFB88.taxon	discussion	Reason for exclusion See comments under A. fulvicornis.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFE95154DA42DCFAFDA0F85D.taxon	discussion	Reason for exclusion Warncke presented a distribution map (Gusenleitner & Schwarz 2002) that contained many taxa close to A. polita with a similar male genital capsule (e. g., including records from Israel and the West Bank and Libya from which true A. polita is absent). The dot present in Iran may be the basis for the listings of Aliyev et al. and Ascher & Pickering; either way, neither source presents precise details. This dot could be represented by A. caspica, A. unicincta, or true A. polita, which is present in eastern Turkey. What is clear is that specimens could not be located in the Warncke collection. True A. polita must be considered to be unconfirmed from Iran.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEA5157D9A8DF06FD7AF85C.taxon	discussion	Reason for exclusion The species concept of A. taraxaci has changed enormously, with many distinct taxa now being recognised after the broad concept adopted by Warncke (see Schwenninger 2015). As a result, it is unclear as to whether older publications apply a broad or narrow species concept, and so A. taraxaci s. str. is excluded from the Iranian fauna until specimens can conclusively demonstrate its presence (see comments under A. astica and A. orientana).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEA5157D9A2D81DFB11FD06.taxon	discussion	Reason for exclusion The taxa A. scotica and A. trimmerana have been endlessly confused (for use of the name scotica, this will be clarified in a manuscript, TJW et al. in prep.), as female material cannot be consistently separated morphologically. Confident determination can only be made from spring generation males. Andrena scotica must therefore be considered unconfirmed until such males can be examined.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEA5157D9A0DD85FC72FA55.taxon	discussion	Reason for exclusion Based on the photographs provided by Allahverdi et al. (2015), both female and male specimens identified as A. sericata are actually all males of A. truncatilabris.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEA5157D9AADBF0FB09FBD4.taxon	discussion	Reason for exclusion Andrena semilaevis has never been recorded from Turkey or the Caucasus, instead being found across temperate Europe to the European part of Russia, avoiding the Mediterranean basin (Gusenleitner & Schwarz 2002). The presence of A. semilaevis in Iran is considered extremely unlikely.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEB5156D98BD82BFB9DF85C.taxon	materials_examined	Material examined Holotype TURKEY • ♂ (holotype of Andrena cara minor); Diyarbakır; 15 – 16 Apr. 1972; K. Warncke leg.; OÖLM. Other material TURKEY • 2 ♂♂, 20 ♀♀; Halfeti env.; 3 – 5 May 1994; M. Halada leg.; OÖLM • 5 ♀♀; Birecik / Urfa; 19 Apr. 1984; K. Warncke leg.; OÖLM.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEB5156D98BD82BFB9DF85C.taxon	discussion	Remarks The status of species formerly placed in the subgenus Carandrena (= Notandrena) is problematic, both at a species level and also because the subgenus is polyphyletic (Pisanty et al. 2022 b). Osytshnjuk (1984, reproduced with additional data in Osytshnjuk et al. 2005) revised Central Asian Carandrena sensu Warncke. In her key, for males with black clypeal markings, she separated two taxa with the gena extended below into either a sharp projection (A. hieroglyphica) or a rounded projection (A. splendula), and described the latter as a new species. However, this conflicts with the work of Warncke. In his revision of Turkish Andrena, Warncke (1974 b, 1975) described a subspecies of A. cara with a rounded projection on the male gena. This character was illustrated by Warncke (1975), who treated the form with the sharp projection (Fig. 164) as A. cara cara, and the form with the rounded projection as A. cara minor. He must have treated A. halictoides as the male of A. cara, as no male for this taxon was described by Nurse (1904), though Warncke never published the synonym (e. g., Warncke 1967), which was later made by Gusenleitner & Schwarz (2002). The problem thus arises that A. cara and its male (described as A. halictoides) are synonymous with A. hieroglyphica because of this distinct genal spine. Andrena cara minor is therefore not a subspecies of this taxon, but a good species, and the senior synonym of A. splendula. The male genital capsules of A. minor and A. splendula are identical (see illustrations in Osytshnjuk 1984; Osytshnjuk et al. 2005). As Warncke’s collection and hence his types were not available to Osytshnjuk for study, or indeed the types of Nurse in London, this independent duplication of names by workers on the Turkish, Pakistani, and Central Asian faunas occurred, when the taxa themselves are found across similarly arid parts of these regions. For a wider classification, Andrena hieroglyphica is placed in incertae sedis; the extensive shiny areas that cover most of the scutum are strongly remenicient of A. euzona, which is found outside the Carandrena + Notandrena (Pisanty et al. 2022 b). Andrena minor is also placed as incertae sedis until genetic work can be conducted to fully define the limits of the new, more restricted subgenus Notandrena (Pisanty et al. 2022 b). More broadly, though A. cara was not the type species of the former subgenus Carandrena (type species: A. aerinifrons Dours, 1873; Warncke 1968), the likely reality that A. cara occurs outside of the expanded Notandrena sensu Pisanty et al. 2022 b further justifies the decision to synonymise these two subgenera in order to reduce nomenclatural confusion.	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
03AB87A2FFEB5156D98BD82BFB9DF85C.taxon	distribution	Distribution Turkey, Turkmenistan, Tajikistan (Osytshnjuk et al. 2005, as A. splendula).	en	Wood, Thomas J., Monfared, Alireza (2022): A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1-136, DOI: https://doi.org/10.5852/ejt.2022.843.1947, URL: http://dx.doi.org/10.5852/ejt.2022.843.1947
