identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0E4D87D0FFEDFFBA2FEC0923FC888D9B.text	0E4D87D0FFEDFFBA2FEC0923FC888D9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hilliardiella elaeagnoides (DC.) N. Swelankomo & J. C. Manning	<div><p>Hilliardiella elaeagnoides (DC.) N.Swelankomo &amp; J.C.Manning in Swelankomo et al. (2016: 50).</p> <p>= Webbia elaeagnoides Candolle (1836: 73). = Cacalia elaeagnoides (DC.) Kuntze (1891: 968). = Vernonia elaeagnoides (DC.) Schultz Bipontinus (1844: 668), non Kunth (1818).</p> <p>Type: — SOUTH AFRICA. KwaZulu-Natal: Zwischen Omsamwubo und Omsamcaba, auf Grasfeldern, 1000–1500 fuss [between Umzimvubu/Mzimvubu and Umsikaba/Msikaba, on grassland, 305–457 m], (QDS: 3129BC/BD/DA Port St Johns), 18 February 1832, Drege 5077 (lectotype: G-DC, designated here; isolectotype: P, barcode P022661, all seen as images). (Lectotypes chosen as recommended by McNeill (2014). (Information on Drège localities from Drège (1843), Glen &amp; Germishuizen (2010) and labels on Paris specimens).</p> <p>= Vernonia kraussii Schultz Bipontinus (1844: 668); Sch.Bip. in Walpers (1843: 947), nom. nud. in syn.</p> <p>Type: — SOUTH AFRICA. KwaZulu-Natal: In graminosis ad radicem (Octobri) et in summitate mont. Tafelbergen [in grassy places at the foot and the top of Table Mountain], (QDS: 2930DA], August 1839, Krauss 462 (lectotype: MO, barcode MO 391699, designated here; isolectotypes: P, barcode P 022670, TUB, barcode 004988, all seen as images).</p> <p>Note:— Schultz Bipontinus (1844) cited two Krauss specimens: 48 and 462. Specimens of Krauss 462 are housed in MO, P, TUB (JSTOR 2022, P online). The TUB specimen with barcode 004988 has two labels attached to it, one label with the number 48 and the date October 1839, and the second label with the number 462 and the date August 1839. The MO specimen was selected as the lectotype to eliminate any confusion. Only one specimen of Krauss 48 was seen on the P online website.</p> <p>Rigid, robust plant up to 1 m high, with many rigid, erect stems from underground rootstock, with ovate to subrotund, pinnately veined leaves up to 20(–25) mm broad, with many capitula in large, open, corymbose synflorescences. For full description see Swelankomo et al. (2016). (Figs 1A, 2).</p> <p>Distribution: —Widespread in southern and tropical Africa.</p></div> 	https://treatment.plazi.org/id/0E4D87D0FFEDFFBA2FEC0923FC888D9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Herman, Paul P. J.	Herman, Paul P. J. (2022): Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety. Phytotaxa 570 (1): 29-42, DOI: 10.11646/phytotaxa.570.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.570.1.3
0E4D87D0FFE8FFBF2FEC0DEAFA7C8F07.text	0E4D87D0FFE8FFBF2FEC0DEAFA7C8F07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hilliardiella oligocephala (DC.) H. Robinson 1999	<div><p>Hilliardiella oligocephala (DC.) H. Robinson (1999: 230).</p> <p>= Webbia oligocephala Candolle (1836: 73). = Vernonia oligocephala (DC.) Sch.Bip. in Walpers (1843: 947). = Vernonia kraussii var. oligocephala (DC.) Harvey (1865: 51).</p> <p>Type: — SOUTH AFRICA. KwaZulu-Natal: ‘ Zwischen Omsamculo und Omcomas, in den Dünen und auf Grashöhen am Gestade, 200 ft, 19 März 1832’ [between Umzimkulu/Mzimkulu and Umkomaas, on grassy heights and the beach, 61 m], (QDS: 3030BB/BC/CB/ DA Port Shepstone), 19 March 1832, Drege 5076 (lectotype: G-DC, barcode G00464306, designated here; isolectotypes: P, barcodes P022663, P022664, all seen as images). (Lectotypes chosen as recommended by McNeill (2014).</p> <p>Slender perennial herbs with few soft, annual, herbaceous stems from small underground rootstock. Rootstock also with narrow fusiform tubers. Annual stems unbranched or rarely with one or two short side branches, faintly to distinctly ribbed, sparsely hairy to glabrescent basally, upwards densely silvery/whitish hairy, hairs short-stalked, medifixed T-shaped with arms parallel to surface or very rarely sometimes with long-stalked, asymmetrically, Y-shaped hairs with arms shorter than stalk and stalk with reddish transverse cell walls. Leaves alternate, sessile or very shortly petiolate, densely set to more scattered, discolorous, margin entire, often undulate and distantly, sparsely serrulate, adaxial surface sparsely hairy or glabrescent, hairs scattered over surface or confined to veins, often glandular punctate towards the apex, abaxial surface densely hairy, hairs silvery/whitish, hairs short-stalked, medifixed, T-shaped with arms parallel to surface, similar to those on stems. Capitula homogamous, discoid, very rarely heterogamous, discoid, few, on very short peduncles, arranged in dense, terminal, umbellate synflorescences or sometimes densely umbellateracemosely arranged, rarely synflorescence more open and lax. Peduncle densely hairy, often hidden by uppermost leaves. Involucre broadly obconical to campanulate. Involucral bracts arranged in (4)5 rows, imbricate, outer shorter than inner, densely white or silvery hairy and glandular-punctate. Receptacle epaleate. Disc florets bisexual, fertile, sometimes few outer florets female, actinomorphic; corolla pinkish or purple, infundibuliform. Anthers with (narrowly) ovate apical appendages, bases caudate. Style exserted, bifurcate; style branches pinkish purple, filiform, recurved, with acute hairs on outside of branches and onto the shaft below furcation, total inner surface stigmatic. Ovary/cypsela brown, ± obconical, ribbed, densely twin hairy and glandular punctate, with distinct carpopodium. Pappus dimorphic, an outer row of laciniate scales and inner row of barbellate bristles.</p> <p>Key to the varieties</p> <p>Leaves ovate, broadly ovate, cordate or narrowly ovate, elliptic or broadly elliptic, lowest pair often orbicular, up to 28 × 20(–25) mm (including short petiole), prominently pinnately veined........................................................................................ var. oligocephala</p> <p>Leaves narrowly elliptic, linear-elliptic, sometimes elliptic, rarely narrowly obovate or obovate, up to 45(–50) × 10(–12) mm (including short petiole), prominently 3(rarely 5)-veined from base, these raised on abaxial surface.............................................. var. besteri</p></div> 	https://treatment.plazi.org/id/0E4D87D0FFE8FFBF2FEC0DEAFA7C8F07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Herman, Paul P. J.	Herman, Paul P. J. (2022): Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety. Phytotaxa 570 (1): 29-42, DOI: 10.11646/phytotaxa.570.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.570.1.3
0E4D87D0FFE8FFB32FEC083FFE4D8215.text	0E4D87D0FFE8FFB32FEC083FFE4D8215.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hilliardiella oligocephala var. oligocephala var. oligocephala	<div><p>Hilliardiella oligocephala var. oligocephala</p> <p>Slender perennial herb with few soft, annual, herbaceous stems from small underground rootstock. Rootstock also with narrow fusiform tubers. Annual stems 80–330, very rarely to 380, mm high, unbranched or rarely with one or two short side branches, faintly to distinctly ribbed, sparsely hairy to glabrescent basally, upwards densely silvery/ whitish hairy, hairs short-stalked, medifixed T-shaped with arms parallel to surface or very rarely sometimes with long-stalked, asymmetrically, Y-shaped hairs with arms shorter than stalk and stalk with reddish transverse cell walls. Leaves alternate, sessile or very shortly petiolate, densely set to more scattered, discolorous, ovate, broadly ovate, cordate or narrowly ovate, elliptic or broadly elliptic, lowest pair often orbicular, 10–28 × 3.5–20.0(–25.0) mm (including petiole), apex acute, attenuate, mucronate apiculate, acuminate, obtuse or rounded, often shortly cuspidate, base cuneate, attenuate, obtuse, cordate or subcordate, margin entire, often undulate and distantly, sparsely serrulate, prominently pinnately veined, these raised on abaxial surface, on adaxial surface faint or sunken, adaxial surface sparsely hairy or glabrescent, hairs scattered over surface or confined to veins, often glandular punctate towards the apex, very rarely sometimes on adaxial side near apex and along margins of apex with long-stalked, asymmetrically, Y-shaped hairs with arms shorter than stalk and stalk with reddish transverse cell walls, abaxial surface densely hairy, hairs silvery/whitish, hairs short-stalked, medifixed, T-shaped with arms parallel to surface, similar to those on stems. Capitula homogamous, discoid, very rarely heterogamous, discoid, few, on very short peduncles, arranged in dense, terminal, umbellate synflorescences or sometimes densely umbellate-racemosely arranged, rarely synflorescence more open and lax. Peduncle (2–)3–12(–15), very rarely to 28, mm long, densely hairy, often hidden by uppermost leaves. Involucre broadly obconical to campanulate, 5–9(–10) mm wide across upper part. Involucral bracts arranged in (4)5 rows, imbricate, outer shorter than inner, densely white or silvery hairy and glandular-punctate; outer row lineartriangular or very narrowly ovate, (2.2–)2.5–4.0 × 0.2–0.5 mm, apex attenuate, usually cuspidate, usually reddish, hairy, glabrous or glabrescent, upper part of margin sometimes reddish, surface densely silvery hairy and glandular punctate, main vein sometimes obvious; second row linear-triangular, very narrowly ovate or rarely very narrowly obovate, 3.0–4.5(–4.8) × 0.4–0.6 mm, apex acuminate or attenuate, cuspidate, usually reddish, hairy, glabrous or glabrescent, upper part of margin sometimes reddish, surface densely silvery hairy and glandular punctate, main vein sometimes obvious and sometimes keeled in lower part; third row narrowly triangular, very narrowly ovate, narrowly elliptic or rarely narrowly obovate, (3.5–)4.0–5.4 × 0.5–1.4(–1.5) mm, apex attenuate or acuminate, cuspidate, usually reddish, sparsely hairy, glabrous or glabrescent, upper part of margin usually reddish or pink-purple; surface whitish hairy and glandular punctate, main vein usually visible, often keeled in lower half, inside surface with prominent main vein or sometimes 3-veined; fourth row narrowly ovate, narrowly elliptic or rarely narrowly obovate, 4.2–5.5(–6.0) × 0.5–1.4(–1.5) mm, apex acuminate or attenuate, cuspidate, usually reddish, glabrous or glabrescent, upper part of margin reddish or pink-purple, surface whitish hairy and glandular punctate, main vein usually visible, often keeled in lower half, inside surface with prominent main vein or sometimes 3-veined; fifth row narrowly ovate, narrowly elliptic or rarely narrowly obovate or narrowly oblong, (4.0–)4.5–5.5(–6.0) × 0.6–1.2 mm, apex attenuate, cuspidate, usually reddish, glabrous or glabrescent, upper part of margin reddish or pink-purple, surface sometimes reddish in upper part, whitish hairy and glandular punctate, main vein ± prominent and often keeled in lower half, inside surface with prominent main vein or sometimes 3-veined. Receptacle epaleate. Disc florets bisexual, fertile, sometimes few outer florets female, actinomorphic; corolla pinkish or purple, funnelform; tube 3.5–5.0(–5.5) mm long, upper part of tube with few sessile glands and T-shaped hairs, 5-lobed; lobes (1.5–)2.0–3.5(–4.0) mm long, apex acute, with sessile glands and T-shaped hairs but around apex with unicellular hairs. Anthers (1.5–)1.8–2.2(–2.5) mm long, with (narrowly) ovate apical appendages, bases caudate. Style (4.0–)4.5–6.5(–8.5) mm long, exserted, bifurcate; style branches pinkish purple, filiform, 1.0– 1.8 mm long, recurved, with acute hairs on outside of branches and onto the shaft below furcation, total inner surface stigmatic. Ovary/cypsela brown, ± obconical, 1.0– 1.5 mm long, ribbed, densely twin hairy and glandular punctate, with distinct carpopodium. Pappus dimorphic, an outer row of laciniate scales, 0.5–1.0 mm long and inner row of barbellate bristles, 3.5–5.0(–5.5) mm long. Flowering time: July to December with one record each for January, February and April. (Figs 3, 4A, B, E).</p> <p>Diagnostic characters:— Hilliardiella oligocephala var. oligocephala is a slender plant up to about 350 mm high, with only a few soft, herbaceous, slender stems from the underground rootstock, with narrowly or broadly ovate, cordate, elliptic or broadly elliptic, prominent pinnately veined, leaves and a few capitula arranged in dense terminal umbellate synflorescences. H. elaeagnoides is a rigid, robust plant up to 1 m high, with clumps of many rigid, erect stems from the underground rootstock, with many capitula in large, open corymbose synflorescences. Some specimens were seen with capitula more densely arranged in almost umbellate synflorescences, but their stems are still quite hard and rigid.</p> <p>Distribution:— Hilliardiella oligocephala var. oligocephala is endemic to South Africa and occurs in a small area around Barberton (Mpumalanga), through KwaZulu-Natal and into the Eastern Cape (Fig. 5). Not as common as H. elaeagnoides and usually overlooked because of its smaller stature.</p> <p>Additional specimens examined</p> <p>SOUTH AFRICA. Mpumalanga: Barberton District, ca. 9.8 km directly SW of Barberton, Farm Schultzenhorst 718JT, N facing mountain slope, grassland, 1618 m, (QDS: 2531CC Komatipoort), 9 December 2008, Bester 8832 (PRE). Mukwana Heights north of house, 2.4 km S of Barberton, burnt N facing mountain slope, 3700 ft [1128 m], (QDS: 2531CC Komatipoort), 18 August 1974, De Souza 434b (PRE). Barberton, Saddleback Mountain, 4000 ft [1219 m], (QDS: 2531CC Komatipoort), September 1889, Galpin 425 (PRE). Locality not found: “Brommers” – Mountains SE of Barberton, steep, rocky grassland, recently burnt, 10 July 1985, Onderstall 1247 (PRE).</p> <p>KwaZulu-Natal: Louwsburg Distr., Itala Nature Reserve, Potwe, mountain, 3800 ft [1158 m], (QDS: 2731AC Louwsburg), 8 December 1975, Brown &amp; Shapiro 34 (PRE). Ntonjaneni Distr., 3 ½ m [5.6 km] S of Ntonjaneni on road to Melmoth, dense grassveld, recently burnt, 3400 ft [1036 m], (QDS: 2831CB Nkandla), 8 October 1946, Codd 1796 (PRE). Ngoye Forest [Umgoye/Ongoye Forest], grassland between exposed rock faces, 1600 ft [488 m], (QDS: 2831DC Nkandla), 11 December 1973, Stirton 446 (PRE). Zululand, Hlabisa district, west of Charter’s Creek, amongst grasses in sandy soil near vlei, 100 ft [30 m], (QDS: 2832AB Mtubatuba), 5 December 1955, Ward 2847 (PRE). St Lucia, S of Lake, Dukuduku State forest, moist riverine forest/thicket, 20 m, (QDS: 2832AD Mtubatuba), 9 December 1989, Hobson 370 (PRE). Witbank, Balgowan, [Lion’s River Distr.], dolorite outcrop, grassland, 4800 ft [1463 m], (QDS: 2930AC Pietermaritzburg), 12 October 1964, Moll 1219 (PRE). Greytown, 3420 ft [1042 m], (QDS: 2930BA Pietermaritzburg), Oct-Nov 1931, Wylie s.n. TvlMus 33875 (PRE). Pietermaritzburg, Silverglen Nature Reserve, NE of Umlazi Nursery, recently burnt grassland, (QDS: 2930DD Pietermaritzburg), 21 October 1985, Ngwenya 181 (PRE). [Pinetown], Kloof, 1500 ft [457 m], (QDS: 2930DD Pietermaritzburg), 14 September 1893, Schlechter 3203 (PRE). Hill near Pinetown, 1500 ft [457 m], (QDS: 2930DD Pietermaritzburg), 29 August 1893, Wylie s.n. Nat. Herb. 495 (PRE, 2 specimens). Weza State Forest, foothills of Ngeli Mountain near Loveday [Lovedale?] lookout, grassland, (QDS: 3029DA Kokstad), 20 November 1986, Van Wyk 7561 (PRE). District Alexandra, Station Dumisa, Farm Friedenau, 600 m, (QDS: 3030AD Port Shepstone), 31 January 1908, Rudatis 235 (PRE). District Alexandra, Station Dumisa, Campbellton, 800 m, (QDS: 3030AD Port Shepstone), 8 November 1912, Rudatis 1745 (PRE). Dududu, 2 km W of the Cedars, gentle slope, recently burnt grassland, loamy soil, (QDS: 3030BA Port Shepstone), 5 November 1992, Sikhakhane 268 (PRE). Port Edward, Clearwater, (QDS: 3130AA Port Edward), 22 August 1982, Abbott 241 (PRE). Just outside Port Edward on road to Bizana, hillside, grassland, recently burned, (QDS: 3130AA Port Edward), 20 September 1996, Germishuizen 8659 (PRE).</p> <p>Vague locality: Zululand, 1–2000 ft [305–610 m], April 1895, Wylie / Medley Wood 5755 (PRE).</p> <p>Eastern Cape: Griqualand East, near Clydesdale, Mount Malowe, grassy slopes, 3500 ft [1069 m], (QDS: 3029BD Kokstad), November 1885, Tyson 2148 (PRE, 2 specimens). Bizana Commonage, grassveld, 2500 ft [762 m], (QDS: 3029DD Kokstad), 22 November 1945, Acocks 12257 (PRE). Bizana District, flats 0.5 miles [0.8 km] W of town, sour grassveld, 762 m, (QDS: 3029DD Kokstad), 22 November 1945, Story 587 (PRE). Ntsubane Forest Station near Lusikisiki, grassy plains, (QDS: 3129BC Port St Johns), 2 February 1929, Galpin 10993 (PRE). Transkei, Mkambati Nature Reserve, burnt grassland, 230 m, (QDS: 3129BD Port St Johns), 19 February 1985, Shackleton 228 (PRE). Transkei, 1.3 myl [2.1 km] vanaf die kantore by Mkambati op pad na die hospitaal, in onlangs gebrande grasveld [2.1 km from the offices at Mkambati on the way to the hospital, in recently burnt grassveld], (QDS: 3129BD Port St Johns), 9 July 1976, Van Wyk 1555 (PRE). Flora of Coffee Bay, Tembuland, on grassy slopes, (QDS: 3129CC Port St Johns), November 1919, Tyson s.n. PRE44465 (PRE). Bizana District, Umtentu mouth, open grassveld, 100 ft [30 m], (QDS: 3130AA Port Edward), 17 August 1953, Story 4171 (PRE). District Kentani, margin of woods, 1500 ft [457 m], (QDS: 3228CB Butterworth), 1901, Pegler 64 (PRE). Kentani, (QDS: 3228CB Butterworth), November 1906, Pegler s.n. TvlMus4126 (PRE).</p></div> 	https://treatment.plazi.org/id/0E4D87D0FFE8FFB32FEC083FFE4D8215	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Herman, Paul P. J.	Herman, Paul P. J. (2022): Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety. Phytotaxa 570 (1): 29-42, DOI: 10.11646/phytotaxa.570.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.570.1.3
0E4D87D0FFE5FFB72FEC0DD7FE778AC1.text	0E4D87D0FFE5FFB72FEC0DD7FE778AC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hilliardiella oligocephala var. besteri P. P. J. Herman 2022	<div><p>Hilliardiella oligocephala (DC.) H.Rob. var. besteri P.P.J.Herman, var. nov.</p> <p>Similar to H. oligocephala var. oligocephala but with narrowly elliptic to linear-elliptic leaves up to 10(–12) mm broad, 3(rarely 5)-veined, 3(5) veins prominently raised abaxially.</p> <p>Type: — SOUTH AFRICA. Mpumalanga: Barberton District, ± 8.5 km directly SSW of Barberton, Farm Ameide 717JT, NW of Shokhohlwa Peak, S hill slope, grassland, 1591 m, (QDS: 2531 CC Komatipoort), 9 December 2008, Bester 8817 (holotype PRE).</p> <p>Etymology:—Named for S.P. (Pieter) Bester, colleague and diligent plant collector.</p> <p>Slender perennial herb with few, soft, annual, herbaceous stems from small underground rootstock. Rootstock also with fusiform tubers. Annual stems (100–)120–250(–270) mm high, unbranched or rarely with one or two short side branches, faintly to distinctly ribbed, densely brown or silvery/whitish hairy, hairs short-stalked, medifixed Tshaped with arms parallel to surface. Leaves alternate, sessile or very shortly petiolate, discolorous, narrowly elliptic, linear-elliptic, sometimes elliptic, rarely narrowly obovate or obovate, (10–)12–45(–50) × (2.0–)2.5–10.0(–12.0) mm (including petiole), apex obtuse to acute, sometimes rounded or rarely attenuate, base cuneate or shortly attenuate, margin entire, prominently 3(rarely 5)-veined from base, these raised on abaxial surface, on adaxial surface faint or sunken, rarely some reticulate veins visible between main veins, adaxial surface sparsely to more densely hairy, abaxial surface densely hairy, hairs brownish or whitish, also short-stalked, medifixed T-shaped with arms parallel to surface, similar to those on stems. Capitula homogamous, discoid, few, on very short peduncles, arranged in dense, terminal, umbellate synflorescences. Peduncle (2.0–)2.5–14.0 mm long, densely hairy. Involucre cupuliform to campanulate, (5–) 6–10 mm wide across upper part. Involucral bracts arranged in 4 or 5 rows, imbricate, outer shorter than inner, densely white or silvery hairy and glandular-punctate; outer row narrowly triangular, linear-triangular, linear or narrowly ovate, (2.5–)3.0–4.0(–5.8) × 0.5–1.0 mm, apex usually cuspidate, reddish; second row linear-triangular, narrowly triangular, narrowly ovate or narrowly obovate, (3.0–)4.0–5.0(–6.5) × (0.8–)1.0–1.5(–2.0) mm, apex attenuate or cuspidate, upper part of margin and apex usually reddish, sometimes 3-veined, visible on inside; third row narrowly ovate or narrowly obovate, 4.0–5.0(–5.5) × 1.0–1.6(–2.0) mm, apex acute or attenuate, cuspidate, upper part of margin and apex usually reddish, usually 3-veined, visible on inside; fourth row narrowly ovate, narrowly elliptic or (very) narrowly obovate, (4–)5–6 × 1.0–1.5(–2.2) mm, apex acute or attenuate, cuspidate, apical part of margin and apex reddish, usually 3-veined, visible on inside; fifth row narrowly elliptic or narrowly obovate, (4.2–)5.5(–6.0) × 1.0– 1.5 mm, apex attenuate, cuspidate, apical part of margin and apex reddish, usually 3-veined, visible on inside. Receptacle epaleate. Disc florets bisexual, fertile, actinomorphic; corolla pinkish or purple, tubular; tube 3.5–4.5(–5.0) mm long, upper part of tube with few sessile glands and T-shaped hairs, 5-lobed; lobes 2–3 mm long, apex acute, with sessile glands and T-shaped hairs but around apex with unicellular hairs. Anthers (1.5–)2.0– 2.5 mm long, with (narrowly) ovate apical appendages, bases caudate. Style 4.0–6.0(–7.5) mm long, exserted, bifurcate; style branches reddish or purple, filiform, 1.0–1.5(–1.8) mm long, recurved, with acute hairs on outside of branches and onto the shaft below furcation, total inner surface stigmatic. Ovary/cypsela brown, ± obconical, 1.0– 1.8 mm long, densely twin hairy and glandular punctate, with distinct carpopodium. Pappus dimorphic, an outer row of laciniate scales, 0.5–1.0 mm long and inner row of barbellate bristles 4–5 mm long. Flowering time: (August)September to December. (Fig. 1B, C, 4C, D, F, 6).</p> <p>Diagnostic characters:— Hilliardiella oligocephala var. besteri has few, soft, slender, annual, herbaceous stems only up to 250(–270) mm high from the underground rootstock, narrowly elliptic to linear-elliptic, abaxially prominently raised 3(rarely 5)-veined leaves up to 10(–12) mm broad. Hilliardiella oligocephala var. oligocephala has narrowly or broadly ovate, cordate, elliptic or broadly elliptic, prominently raised, pinnately veined leaves.</p> <p>Distribution:— Hilliardiella oligocephala var. besteri is near endemic to Mpumalanga and occurs in a small area in Mpumalanga and in eSwatini (Swaziland) (Fig. 7).</p> <p>Additional specimens examined</p> <p>SWAZILAND [eSWATINI]. Havelock Mine, grassland, 4300 ft [1311 m], (QDS: 2531CC Komatipoort), September 1955, Miller 3013 (PRE). Havelock Mine, grassland, 4000 ft [1219 m], (QDS: 2531CC Komatipoort), October 1955, Miller 3057 (PRE). Mbabane District, Waverley Mine, 5000 ft [1524 m], (QDS: 2630BD Carolina), 17 October 1956, Compton 26095 (PRE). Mbabane District, Ngwenya Mountain, 5500 ft [1676 m], (QDS: 2632AA Bela Vista), 18 November 1956, Compton 26332 (PRE).</p> <p>SOUTH AFRICA. Mpumalanga: Pilgrim’s Rest District, Sabie, Mauchsberg, 4800–5800 ft [1463–1768 m], (QDS: 2530BA Lydenburg), 22 December 1932, Smuts &amp; Gillett 2271 (PRE), Smuts &amp; Gillett 2329 (PRE). Pilgrim’s Rest District, Mount Anderson, 7000 ft [2134 m], (QDS: 2530BA Lydenburg), 24 December 1932, Smuts &amp; Gillett 2406 (PRE). Barberton District, ± 11 km E of Barberton, Farm Twello, Sappi Plantation, moderate NW mountain slope, grassland, 1449 m, (QDS: 2531CC Komatipoort), 13 November 2014, Bester 12085 (PRE). Barberton District, at turn-off from R40 between Barberton and Josefsdal towards Diepgezet mine, ± 18 km SSE of Barberton, gentle NW mountain slope, grassland, 1455 m, (QDS: 2531CC Komatipoort), 26 November 2017, Bester 13858 (PRE). Barberton District, Oosterbeek farm, 5 km S of Barberton, grassland, 5200 ft [1585 m], (QDS: 2531CC Komatipoort), 15 November 1977, De Sousa 631 (PRE). Barberton, Angle station, hill above station, (QDS: 2531CC Komatipoort), 2 August 1978, Venter 4491 (PRE).</p> <p>Locality not found: Mpumalanga, Barberton, mountain tops near Bosch’s, 4000–4500 ft [1219–1372 m], September 1889, Galpin 557 (PRE).</p></div> 	https://treatment.plazi.org/id/0E4D87D0FFE5FFB72FEC0DD7FE778AC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Herman, Paul P. J.	Herman, Paul P. J. (2022): Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety. Phytotaxa 570 (1): 29-42, DOI: 10.11646/phytotaxa.570.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.570.1.3
