taxonID	type	description	language	source
AE7687EAFFBBFFADFF4359EBFA08F87B.taxon	type_taxon	(type species by original designation: Trichoserica fulvopubens Reitter, 1896); synonymized by Nomura 1972: 110.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFBBFFADFF4359EBFA08F87B.taxon	type_taxon	Type species: Serica brunnea (Linnaeus, 1758) (by monotypy).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFBBFFADFF4359EBFA08F87B.taxon	description	Taxonomic revision and new species Key to subgenera and species groups of Serica in framework of the genera of Sericini with tri-lamellate antennal club occurring in continental South-East Asia (♂♂):	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFBBFFADFF4359EBFA08F87B.taxon	discussion	Key to the species of Serica (sensu lato) of continental South-East Asia (♂♂):	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB6FFA0FF4359EBFD1EFED6.taxon	description	Figures 1 A – D, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB6FFA0FF4359EBFD1EFED6.taxon	materials_examined	Type material examined. Holotype: ♀ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 11.4. 1938 (Fukien) / Type Serica longula G. Frey 1972 ” (ZMFK). Paratype: 1 ♀ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 12.5. 1938 (Fukien) / Type Serica longula G. Frey 1972 ” (ZMFK). Additional material examined. 1 ♂ “ Chine 28. VII. 46 Kuatun, Fukien leg. Tscheng-Sen ” (MHNG), 2 ♂♂ “ Kuatun, Fukien China, 15.6.46 leg. Tschung-Sen / ex. coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ♂, 1 ♀ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 11.5. 1938 (Fukien) ” (NMPC), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 11.4. 1938 (Fukien) ” (NMPC), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 30.4. 1938 (Fukien) ” (NMPC), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 8.5. 1938 (Fukien) ” (NMPC), 1 ♀ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 29.5. 1938 (Fukien) ” (NMPC), 1 ♀ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 26.5. 1938 (Fukien) ” (NMPC), 1 ♂ “ China: Fukien 06. V. 1938 leg. Klapperich ZFMK Bonn ” (ZFMK), 1 ♂ “ San’gang, Mt. Wuyishan, Fujian, 16 - 21. Ⅴ. 2004, leg. Yuan Caixia, Lijing ” (HBUM), 1 ♂ “ Huangxizhou, Mt. Wuyishan, Fujian, 27. Ⅴ. 2004, leg. Yuan Caixia, Lijing ” (HBUM).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB6FFA0FF4359EBFD1EFED6.taxon	description	Redescription (male). Length: 8.2 mm, length of elytra: 6.4 mm, width: 4.6 mm. Body elongate, dorsal face yellow and dull, glabrous, abdomen brown, antenna yellow. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface flat, finely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), very finely sparsely punctate, with a terminal seta. Frons with fine, dense punctures, dull toment anteriorly weaker, glabrous except for a few setae beside eyes. Eyes moderately large, ratio of diameter / interocular width: 0.62. Antenna composed of 9 antennomeres, club with three antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum transverse and subtrapezoidal, widest at base, lateral margins evenly and weakly convex, weakly convergent anteriorly; anterior angles moderately produced and rectangular; posterior angles blunt, moderately rounded at tip; anterior margin weakly convex, marginal line widely interrupted medially; basal margin without marginal line; surface finely and densely punctate, glabrous; anterior and lateral margins without setae. Scutellum wide, triangular, with coarse and moderately dense punctures, glabrous. Elytra oblong, widest at posterior quarter, striae distinctly impressed, finely and densely punctate; intervals weakly convex, with fine, moderately dense punctures concentrated along striae; odd intervals with a few fine, erect, short setae, otherwise glabrous; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, almost completely glabrous; metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise glabrous; penultimate sternite apically with a shiny smooth short chitinous border and a wide transversal shallow impression at middle. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.19. Pygidium convex, yellow but dull, with fine, moderately dense punctures and a few longer setae on apical half. Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, entirely serrate, dorsal margin also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width / length: 1 / 3.8, dorsally sharply carinate, with two groups of spines, basal one shortly before middle, apical one at four fifths of metatibial length; in basal half with a few punctures and short fine spines; lateral face longitudinally convex, with moderately dense and coarse punctures, glabrous; ventral margin serrate, with two widely distant spines; medial face finely sparsely punctate and with sparse short setae, apex interiorly near tarsal articulation shallowly concave. Tarsomeres irregularly partly coarsely punctate dorsally, ventrally with sparse, short setae; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere ventrally glabrous, little longer than following tarsomere and almost twice as long as dorsal tibial spur. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 1 A – C. Habitus: Fig. 1 D.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB4FFA1FF4358CBFF45FD1A.taxon	description	Figures 1 E – H, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB4FFA1FF4358CBFF45FD1A.taxon	materials_examined	Type material examined. Holotype: ♂ “ Mts. Tianmushan, Zhejiang, 8. VI. 1964, 1200 m, leg. Chen Tailu / LW- 153 ” (IZAS). Paratypes: 1 ♂, 3 ♀♀ “ Mts. Gutianshan, Zhejiang, 1 - 4. V. 2010, 446 - 715 m, leg. Liu Chongling, No. G 24 ha- 16 / LW- 915 ” (IZAS), 1 ♂ “ China Zhejiang Prov. Kaihua Co. Mt. Gutian alt. 400 - 500 m 29 ° 14 ’ N, 118 ° 08 ’ E 19 - VI- 2013 Yu Y, - M leg. ” (SNUC), 1 ♂ “ China: Zhejiang, Gutianshan NNR. 2010, CSP 14, 639 m 118.14 ° E, 29.25 ° N local collector ” (NME), 1 ♂ “ China: Zhejiang, Gutianshan NNR. 2010, CSP 10, 670 m 118.16 ° E, 29.25 ° N local collector ” (NME), 1 ♀ “ China: Zhejiang, Gutianshan NNR. 2010, CSP 24, 366 m 118.13 ° E, 29.21 ° N local collector ” (NME), 1 ♀ “ China: Zhejiang, Gutianshan NNR. 2010, local collector / CSP 15: 2010 618 m 118.13 ° E, 29.25 ° N ” (NME), 2 ♂♂, 1 ♀ “ China: Zhejiang, Gutianshan NNR 2010, CSP 11, 647 m 118.14 ° E, 29.27 ° N ” (NME, ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB4FFA1FF4358CBFF45FD1A.taxon	description	Description of the holotype. Length: 8.8 mm, length of elytra: 6.5 mm, width: 4.6 mm. Body elongate, dorsal face yellow and dull, glabrous, abdomen brown, antenna yellow. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface convex, finely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), very finely sparsely punctate, with a terminal seta. Frons with fine, dense punctures, dull toment anteriorly weaker, glabrous except for a few setae beside eyes. Eyes moderately large, ratio of diameter / interocular width: 0.6. Antenna composed of 9 antennomeres, club with three antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum transverse and subtrapezoidal, widest at base, lateral margins evenly and weakly convex, weakly convergent anteriorly; anterior angles moderately produced and rectangular; posterior angles blunt, moderately rounded at tip; anterior margin weakly convex, fine marginal line complete medially; basal margin without marginal line; surface finely and densely punctate, glabrous; anterior and lateral margins without setae. Scutellum wide, triangular, with fine and sparse punctures, glabrous. Elytra oblong, widest at posterior quarter, striae distinctly impressed, finely and densely punctate; intervals weakly convex, with fine, moderately dense punctures concentrated along striae; odd intervals with a few fine, erect, short setae, otherwise glabrous; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, almost completely glabrous; metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise glabrous; penultimate sternite apically with a shiny smooth short chitinous border and a wide transversal shallow impression at middle. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.37. Pygidium moderately convex, yellow but dull, with fine, moderately dense punctures and a few longer setae on apical half. Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, entirely serrate, dorsal margin also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width / length: 1 / 3.8, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length; in basal half with a few punctures and short fine spines; lateral face longitudinally convex, with moderately dense and coarse punctures, glabrous; ventral margin serrate, with two widely distant spines; medial face finely sparsely punctate and with sparse short setae, apex interiorly near tarsal articulation shallowly concave. Tarsomeres irregularly partly coarsely punctate dorsally, ventrally with sparse, short setae; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere ventrally glabrous, little shorter than following two tarsomeres combined and almost twice as long as dorsal tibial spur. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 1 E – G. Habitus: Fig. 1 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB4FFA1FF4358CBFF45FD1A.taxon	diagnosis	Diagnosis. Serica tianmushan Ahrens, Fabrizi & Liu, new species differs from the very similar Serica longula Frey, 1972 by the shape of parameres: the left paramere has a large arched basal lobe, the right paramere is compacter and shorter.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB4FFA1FF4358CBFF45FD1A.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from its type locality Tianmushan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB4FFA1FF4358CBFF45FD1A.taxon	description	Variation. Length: 8.6 – 8.9 mm, length of elytra: 6.0 – 6.6 mm, width: 4.2 – 4.9 mm. Head, pronotum and scutellum in some specimens greenish yellow. Female: antennal club short, little shorter than remaining antennomeres combined; pygidium strongly evenly convex; eyes little smaller than in male, ratio diameter / interocular width: 0.55.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB5FFA6FF435B87FBF1FCA6.taxon	description	Figures 1 I – M, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB5FFA6FF435B87FBF1FCA6.taxon	materials_examined	Type material examined. Holotype: ♂ “ China: Zhejiang, Anji County Longwang Shan, parking square to Dianmufeng, 30 ° 23 ’ 59 ’’ N, 119 ° 26 ’ 26 ’’ E, 1300 - 1450 m, 14. v. 2013, C. - C. Dai & X. - B. Song leg. / 1107 Asia Sericini spec. ” (SNUC). Paratype: 1 ♂ “ China: S. Zhejiang, Qingyuan Mt. nr. Liyang village, 27 ° 49 ’ 38 ’’ N, 119 ° 11 ’ 22 ’’ E, leaf litter, sifted, 29. iv. 2014 990 - 1160 m, Peng, Song, Yan & Yu leg. ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB5FFA6FF435B87FBF1FCA6.taxon	description	Description of the holotype. Length: 9.2 mm, length of elytra: 7.2 mm, width: 5.4 mm. Body elongate, dorsal and ventral face yellowish brown, dorsal face shiny and glabrous, antenna yellow. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins convex and convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface convex, coarsely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus moderately long and narrow (1 / 3 of ocular diameter), coarsely densely punctate, with a terminal seta. Frons with fine, dense punctures, completely shiny, glabrous except for a few setae beside eyes. Eyes moderately large, ratio of diameter / interocular width: 0.68. Antenna composed of 9 antennomeres, club with three antennomeres, strongly reflexed, three times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum transverse and subtrapezoidal, widest at base, lateral margins in basal two thirds straight and weakly convergent anteriorly, anteriorly more convex; anterior angles moderately produced and rectangular, strongly rounded in the tip; posterior angles blunt; anterior margin convex, robust marginal line complete medially; basal margin without marginal line; surface coarsely and densely punctate, glabrous; anterior and lateral margins with short sparse setae. Scutellum wide, triangular, with fine and dense punctures, glabrous, basal midline impunctate. Elytra oblong, widest at posterior quarter, striae distinctly impressed, finely and densely punctate; intervals weakly convex, with fine, moderately dense punctures concentrated along striae; intervals glabrous; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose; apical border chitinous, without a rim of microtrichomes. Ventral surface dull, metasternal disc and abdominal sternites except the last one shiny; metasternum and metacoxa with large and dense punctures, almost completely glabrous; metacoxa with several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise glabrous; ultimate sternite with a few long additional setae. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.12. Pygidium moderately convex, yellow shiny, with fine, moderately dense punctures, along the apical margin dull and with a few short setae. Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur shiny, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, entirely serrate, dorsal margin also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width / length: 1 / 3.8, dorsally sharply carinate, with only one distal group of spines, basal one reduced to a single robust seta at the middle of metatibial length, apical one at four fifths of metatibial length; in basal half with a short fine spines in row; lateral face longitudinally convex, with a few sparse and coarse punctures, glabrous; ventral margin serrate, with three long setae of which distal two are more distant; medial face coarsely densely punctate and slightly rugose, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres irregularly sparsely and finely coarsely punctate dorsally, ventrally with sparse, short setae; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; metatarsomeres ventrally glabrous, first metatarsomere little longer than following tarsomere and little longer then dorsal tibial spur. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with sharply pointed basal tooth. Aedeagus: Fig. 1 I – L. Habitus: Fig. 1 M. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB5FFA6FF435B87FBF1FCA6.taxon	diagnosis	Diagnosis. Serica longwang Ahrens, Fabrizi & Liu, new species differs from the very similar Serica longula Frey, 1972 and all other species of the Serica sensu lato assemblage by the lacking membraneous border of apical elytra, the competely shiny surface, shape of aedeagus: both parameres possess large basal lobes.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB5FFA6FF435B87FBF1FCA6.taxon	etymology	Etymology. The name of the new species, Longwang (noun in apposition), is derived from its type locality Longwang Shan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB5FFA6FF435B87FBF1FCA6.taxon	description	Variation. Length: 9.0 – 9.2 mm, length of elytra: 6.6 – 7.2 mm, width: 4.6 – 5.4 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB2FFA4FF435A20FBF1FED6.taxon	description	Figures 2 A – D, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB2FFA4FF435A20FBF1FED6.taxon	materials_examined	Type material examined. Holotype: ♂ “ China-Guangdong, Dagiao env., 1000 - 1200 m 24 ° 54 ’ N - 113 ° 01 ’ E, 1 - 3. v. 2002, Dr. R. Fencl lgt. / Coll. P. Pacholatko / 591 Sericini: Asia spec. ” (CP). Paratype: 1 ♂ “ Mts. Dadongshan, Lianxian, Guangdong, 25. Ⅴ. 1997, 1500 - 1700 m, leg. Chen Zhenguang ” (LSSYU).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB2FFA4FF435A20FBF1FED6.taxon	description	Description of the holotype. Length: 8.9 mm, length of elytra: 6.6 mm, width: 4.8 mm. Body elongate, dorsal face yellow and dull, glabrous, ventral face, frons and anterior portion of pronotum brown, antenna yellow. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface flat, finely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), very finely sparsely punctate, with a terminal seta. Frons with fine, dense punctures, dull toment anteriorly weaker, glabrous except for a few setae beside eyes. Eyes moderately large, ratio of diameter / interocular width: 0.68. Antenna composed of 9 antennomeres, club with three antennomeres, straight, 1.3 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum transverse and subtrapezoidal, widest at base, lateral margins in basal half almost straight, weakly convergent anteriorly, in anterior half convex and convergent; anterior angles moderately produced but sharp; posterior angles blunt, moderately rounded at tip; anterior margin weakly convex, marginal line complete medially; basal margin without marginal line; surface finely and densely punctate, with minute setae in punctures; anterior and lateral margins sparsely setose. Scutellum wide, triangular, with fine and dense punctures each bearing a minute seta, base at middle impunctate. Elytra oblong, widest at posterior quarter, striae distinctly impressed, finely and densely punctate; intervals weakly convex, with fine, moderately dense punctures concentrated along striae; lateral intervals with a few fine, erect, short setae, otherwise glabrous; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, almost completely glabrous; metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise glabrous; penultimate sternite apically with a shiny smooth short chitinous border. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.21. Pygidium convex, yellow but dull, with fine, moderately dense punctures and a few longer setae on apical half. Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, entirely serrate, dorsal margin also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width / length: 1 / 4.0, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one shortly behind three quarters of metatibial length; in basal half with a few punctures and short fine spines; lateral face longitudinally convex, with moderately dense and coarse punctures, glabrous; ventral margin serrate, with two widely distant spines; medial face finely sparsely punctate and with sparse short setae, apex interiorly near tarsal articulation shallowly concave. Tarsomeres irregularly partly coarsely punctate dorsally, ventrally with sparse, short setae; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere ventrally glabrous, little longer than following tarsomere and almost twice as long as dorsal tibial spur. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 2 A – C. Habitus: Fig. 2 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB2FFA4FF435A20FBF1FED6.taxon	diagnosis	Diagnosis. Serica daqiaoana Ahrens, Fabrizi & Liu, new species differs from the rather similar Serica longula Frey, 1972 in shape of aedeagus: the phallobase is lacking the distal lateral processes on each side, while the parameres differ significantly in length; the right paramere is almost twice as long as the left having a long basal lobe.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB2FFA4FF435A20FBF1FED6.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from its type locality Daqiao (Dagiao).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB2FFA4FF435A20FBF1FED6.taxon	description	Variation. Length: 8.9 – 9.1 mm, length of elytra: 6.6 – 7.4 mm, width: 4.8 – 5.3 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB0FFA5FF4358CAFE86FDAA.taxon	description	Figures 2 E – H, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB0FFA5FF4358CAFE86FDAA.taxon	materials_examined	Type material examined. Holotype: ♂ “ China: NW Guizhou prov. Fanjingshan, 35 km N of Jiankou, 3 - 4 Jun 1996 1200 m, Bolm lgt. / Coll. P. Pacholatko Invt. No. / 727 Sericini: Asia spec. ” (CP).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB0FFA5FF4358CAFE86FDAA.taxon	description	Description of the holotype. Length: 8.0 mm, length of elytra: 6.0 mm, width: 4.1 mm. Body elongate, dorsal face yellow and dull, glabrous, ventral face, frons and anterior portion of pronotum brown, antenna yellow. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface weakly convex, finely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), finely sparsely punctate, without a terminal seta. Frons with fine, dense punctures, dull toment anteriorly weaker, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter / interocular width: 0.58. Antenna composed of 9 antennomeres, club with three antennomeres, straight, as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum transverse and subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly; anterior angles moderately produced but sharp; posterior angles convex; anterior margin weakly convex, fine marginal line complete medially; basal margin without marginal line; surface finely and densely punctate, with minute setae in punctures; anterior and lateral margins sparsely setose. Scutellum wide, triangular, with fine and dense punctures each bearing a minute seta, base at middle impunctate. Elytra oblong, widest at posterior quarter, striae distinctly impressed, finely and densely punctate; intervals weakly convex, with fine, moderately dense punctures concentrated along striae; lateral intervals with a few fine, erect, short setae, otherwise glabrous; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of microtrichomes. Ventral surface dull, metasternum and metacoxa with large and dense punctures, almost completely glabrous; metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise glabrous; penultimate sternite apically with a shiny smooth short chitinous border. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.27. Pygidium strongly convex in apical half, yellow and dull, with fine, moderately dense punctures and a few longer setae on apical half. Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, entirely serrate, dorsal margin also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width / length: 1 / 4.0, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length; in basal half with a few punctures and short fine spines; lateral face longitudinally convex, with moderately dense and coarse punctures, glabrous; ventral margin serrate, with two widely distant spines; medial face finely sparsely punctate and with sparse short setae, apex interiorly near tarsal articulation shallowly concave. Tarsomeres irregularly partly coarsely punctate dorsally, ventrally with sparse, short setae; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere ventrally glabrous, little longer than following tarsomere and one third of its length longer than dorsal tibial spur. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 2 E – G. Habitus: Fig. 2 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB0FFA5FF4358CAFE86FDAA.taxon	diagnosis	Diagnosis. Serica jiankouensis Ahrens, Fabrizi & Liu, new species differs from the similar S. daqiaoana Ahrens, Fabrizi & Liu, new species in the shorter antennal club (male) and the shape of aedeagus: the phallobase is ventrally at apex more widened, the left paramere in S. jiankouensis Ahrens, Fabrizi & Liu, new species is slightly more compact, with the basal lobe distinctly shorter; the angle of the dorsal margin in the right paramere is more pronounced and the distal dorsal margin (lateral view) is concavely emarginated rather than straight as in S. daqiaoana Ahrens, Fabrizi & Liu, new species which has the angle of the dorsal margin more blunt; furthermore, the right paramere possesses in S. jiankouensis Ahrens, Fabrizi & Liu, new species a short and narrow basal lobe which is lacking in S. daqiaoana Ahrens, Fabrizi & Liu, new species.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB0FFA5FF4358CAFE86FDAA.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from its occurrence close to Jiankou.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB1FFBBFF435B17FD1EFBA2.taxon	description	Figures 3 A – D, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB1FFBBFF435B17FD1EFBA2.taxon	materials_examined	Type material examined. Holotype: ♂ “ Kuatun 2300 m 27,40 n. Br. 117,40 ö. L. J. Klapperich 4.6.1938 (Fukien) / Ophthalmoserica Type klapperichi G. Frey 1972 / Trichoserica klapperichi ” (ZFMK). Paratypes: 3 ♂♂, 1 ♀ “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 14.6.1938 (Fukien) / Paratype Ophthalmoserica klapperichi G. Frey 1972 ” (ZFMK, CF), 1 ♀ “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 22.6.1938 (Fukien) / Paratype Ophthalmoserica klapperichi G. Frey 1972 / ♀ ” (ZFMK), 1 ♀ “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 21.6.1938 (Fukien) / Paratype Ophthalmoserica klapperichi G. Frey 1972 / ♀ ” (ZFMK), 1 ♀ “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 15.6.1938 (Fukien) / Paratype Ophthalmoserica klapperichi G. Frey 1972 ” (CF). Additional material examined. 1 ♂ “ Chine 10. V. 46 Kuatun, Fukien leg. Tschung-Sen ” (MHNG), 2 ex. “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 14.6.1938 (Fukien) / ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun 2300 m 27, 40 n. Br. 117,40 ö. L. J. Klapperich 21.6.1938 (Fukien) / ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun, Fukien China 24.5.46 (Tschung Sen.) / ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun, Fukien China 15.7.46 Tschung-Sen / ex. Coll. V. Balthasar Natinal Museum Prague, Czech Republic ” (NMPC), 1 ♂ “ Neicujiang, Longsheng, Guangxi, 6. VI. 1963, 840 m, leg. Wang Shuyong ” (IZAS), 1 ♂ “ Mts. Tianpingshan, Huaping, Guangxi, 5. VI. 1963, leg. Yang Jikun ” (CAU), 1 ♂ “ Mt. Tianpingshan, Longsheng, Guangxi, 24. VI. 1982, leg. Wang Xinli ” (CAU), 1 ♂ “ Mt. Tianpingshan, Huaping, Guangxi, 5. VI. 1982, leg. Yang Jikun ” (CAU).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFB1FFBBFF435B17FD1EFBA2.taxon	description	Redescription (holotype). Length: 7.4 mm, length of elytra: 5.1 mm, width: 4.4 mm. Body oval, dorsal face light reddish-brown and dull, elytra partly with iridescent shine, head shiny, with dense, erect, moderately long setae, antenna yellow. Labroclypeus subrectangular, little wider than long, widest at middle, lateral margins convex and moderately convergent the base and to moderately rounded anterior angles; lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface flat, finely and densely punctate, with dense, moderately long, erect setae; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), finely densely punctate, with 2 – 3 terminal setae. Frons with fine and coarse, dense punctures, with dense, moderately long, erect setae. Eyes moderately large, ratio of diameter / interocular width: 0.67. Antenna composed of 9 antennomeres, club with three antennomeres, strongly reflexed, twice as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins in basal half straight and subparallel, in anterior half strongly convex and convergent anteriorly; anterior angles weakly produced and blunt; posterior angles blunt; anterior margin weakly convex, marginal line widely interrupted medially; basal margin without marginal line; surface coarsely and densely punctate, with moderately long, erect setae mixed with short adpressed ones; anterior and lateral margins with dense setae. Hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with coarse and moderately dense punctures, with short adpressed setae. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate; intervals convex at middle, with fine, dense punctures concentrated along striae, on middle of interval almost impunctate; intervals with dense, moderately long setae being directed posteriorly; epipleural margin robust, ending before rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a rim of microtrichomes. Ventral surface shiny along middle, dull at sides, metasternum and metacoxa with fine and moderately dense punctures, sparsely setose; metacoxa with fine adpressed setae, except for a several long setae laterally. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.63. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise with fine dense setae; penultimate sternite apically without distinct chitinous border. Pygidium strongly convex, dull, with fine, dense punctures and dense, long setae. Legs moderately wide and short; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur shiny, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, smooth, dorsal posterior margin also not serrate, finely setose. Metatibia moderately wide and short, widest at apex, ratio width / length: 1 / 2.8; dorsal margin sharply carinate, with two groups of spines, basal one at one third, apical one two thirds of metatibial length; basally with a few fine setae; lateral face longitudinally convex, with dense and coarse punctures, and moderately long setae in the punctures; ventral margin serrate, with three equidistant robust setae; medial face coarsely but sparsely punctate and with sparse minute setae in punctures, apex interiorly near tarsal articulation bluntly truncate and shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere dorsally and laterally also moderately carinate, distinctly longer than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin bluntly extended medially. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 3 A – C. Habitus: Fig. 3 D.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAFFFB8FF435D1FFBF1FABE.taxon	description	Figures 3 E – H, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAFFFB8FF435D1FFBF1FABE.taxon	materials_examined	Type material examined. Holotype: ♂ “ CH, Guizhou prov., ~ 650 m, Jiangkou (ca 50 km SW), 27 ° 32.83 ’ N 108 ° 36,49 ’ E, Shidu vill. Env., 29. vi. - 6. vii. 2001, C. Holzschuh leg. / 826 Sericini: Asia spec. ” (CP). Paratypes: 2 ♂♂ “ CH-Guizhou NE 27. V. - 3. VI. 20 km NW of Jiangkou, 1995 Fanjing Shan-Kuaichang E. Jendek & O. Sausa leg. / Coll. P. Pacholatko Invt. No. / CS 24 / 573 Sericini: Asia spec. ” (CP, CA), 1 ♂ “ Libo, Ⅴ. 1995 ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAFFFB8FF435D1FFBF1FABE.taxon	description	Description of the holotype. Length: 7.1 mm, length of elytra: 4.8 mm, width: 4.1 mm. Body oval, dorsal face light reddish-brown and dull, head shiny, with dense, erect, moderately long setae, antenna yellow. Labroclypeus subrectangular, little wider than long, widest at middle, lateral margins convex and moderately convergent the base and to moderately rounded anterior angles; lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin distinctly emarginate medially; surface flat, finely and densely punctate, with dense, moderately long, erect setae; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), finely densely punctate, with 2 – 3 terminal setae. Frons with fine and coarse, dense punctures, with dense, moderately long, erect setae. Eyes small, ratio of diameter / interocular width: 0.58. Antenna composed of 9 antennomeres, club with three antennomeres, strongly reflexed, twice as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins in basal half slightly concave and subparallel, in anterior half strongly convex and convergent anteriorly; anterior angles weakly produced and blunt; posterior angles blunt; anterior margin weakly convex, marginal line widely interrupted medially; basal margin without marginal line; surface coarsely and densely punctate, with moderately long, erect setae mixed with short adpressed ones; anterior and lateral margins with dense setae. Hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with coarse and moderately dense punctures, with short adpressed setae. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate; intervals convex at middle, with fine, dense punctures concentrated along striae, on middle of interval almost impunctate; intervals with dense, moderately long setae being directed posteriorly; epipleural margin robust, ending at rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a rim of microtrichomes. Ventral surface dull, metasternum and metacoxa with fine and dense punctures, densely setose; metacoxa with fine, dense, adpressed setae, with numerous long setae laterally. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.49. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise with fine dense setae; penultimate sternite apically without distinct chitinous border. Pygidium strongly convex, dull, with fine, dense punctures and dense, long setae. Legs moderately wide and short; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur shiny, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, smooth, dorsal posterior margin also not serrate, finely setose. Metatibia moderately wide and short, widest at apex, ratio width / length: 1 / 3.0; dorsal margin sharply carinate, with two groups of spines, basal one at first quarter, apical one two thirds of metatibial length; basally with a few fine setae; lateral face longitudinally convex, with dense and coarse punctures, and moderately long setae in the punctures; ventral margin serrate, with three almost equidistant robust setae; medial face coarsely but sparsely punctate and with sparse minute setae in punctures, apex interiorly near tarsal articulation bluntly truncate and shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere dorsally and laterally also moderately carinate, as long as following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin bluntly extended medially. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 3 E – G. Habitus: Fig. 3 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAFFFB8FF435D1FFBF1FABE.taxon	diagnosis	Diagnosis. Serica shiduensis Ahrens, Fabrizi & Liu, new species differs from the similar S. klapperichi Frey, 1972 by the more complete dull surface, the slightly concave lateral margin of pronotum, and the shape of aedeagus. The latter is characterized in Serica shiduensis Ahrens, Fabrizi & Liu, new species by the shorter and more ventrally positioned left lateral apical apophysis of phallobase which has a short median apophysis and dorsolateral right margin less extended, by the apically much more widened right paramere, and by the basal lobe of the right paramere being folded twice in the middle, while the one of S. klapperichi is folded only once.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAFFFB8FF435D1FFBF1FABE.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from its type locality, Shidu village.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAFFFB8FF435D1FFBF1FABE.taxon	description	Variation. Length: 7.0 – 7.1 mm, length of elytra: 4.8 – 5.1 mm, width: 4.1 – 5.3 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFACFFB9FF435C23FDEAF8C2.taxon	description	Figures 3 I – L, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFACFFB9FF435C23FDEAF8C2.taxon	materials_examined	Type material examined. Holotype: ♂ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 11.5.1938 (Fukien) / Type Ophthalmoserica excisa G. Frey 1972 ” (ZFMK) [aedeagus lost from attached cartoon]. Paratypes: 1 ♀ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 11.5.1938 (Fukien) / Paratype Ophthalmoserica excisa G. Frey 1972 ” (ZFMK), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 11.4.1938 (Fukien) / Paratype Ophthalmoserica excisa G. Frey 1972 ” (ZFMK [specimen photographed]), 1 ♂, 1 ♀ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 12.5.1938 (Fukien) / Paratype Ophthalmoserica excisa G. Frey 1972 / Trichoserica excisa ” (CF), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 5.5.1938 (Fukien) / Paratype Ophthalmoserica excisa G. Frey 1972 ” (ZFMK, CF), 5 ♂♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 11.5.1938 (Fukien) / Paratype Ophthalmoserica excisa G. Frey 1972 ” (ZFMK, CF). Additional material examined. 1 ex. “ Chine 26. V. 46 Kuatun Fukien leg. Tschung-Sen ” (MHNG), 1 ex. “ Chine 21. IV. 46 Kuatun Fukien leg. Tschung-Sen ” (MHNG), 1 ex. “ Chine 1. VI. 46 Kuatun Fukien leg. TschungSen ” (MHNG), 3 ex. “ Kuatun, Fukien China 14.5.46 leg. Tschung-Sen ” (NMPC), 1 ex. “ Kuatun, Fukien China 15.5.46 leg. Tschung-Sen ” (NMPC), 1 ex. “ Kuatun, Fukien China 25.5.46 leg. Tschung-Sen ” (NMPC), 1 ex. “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 11.4. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 8.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 6 ex. “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 11.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 12.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Ta-stien-Lou Chasseurs Thibetains 1896 / Museum Paris ex. Coll. R. Oberthur ” (MHNP), 1 ♂ “ Guanping, Tongmuguan, Chong’an, Fujian, 22. V. 1960, 800 - 900 m, leg. Jiang Shengqiao ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFACFFB9FF435C23FDEAF8C2.taxon	description	Redescription of the holotype. Length: 6.2 mm, length of elytra: 4.5 mm, width: 3.6 mm. Body oval, dorsal face yellow and shiny, head and ventral surface dark brown, shiny, with dense, backwards-bent and moderately long setae, antenna yellow. Labroclypeus subrectangular, little wider than long, widest at base, lateral margins subparallel in basal half, anteriorly strongly convex and moderately convergent to strongly rounded anterior angles; lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; anterior margin deeply but narrowly emarginate medially; surface flat, finely and densely punctate, with dense, long, erect setae; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and fine (1 / 4 of ocular diameter), finely densely punctate, with numerous setae. Frons with fine, dense punctures, with dense, long setae directed posteriorly. Eyes large, ratio of diameter / interocular width: 0.83. Antenna composed of 9 antennomeres, club with three antennomeres, strongly reflexed, 2.5 times as long as remaining antennomeres combined. Mentum anteriorly elevated and flattened. Pronotum subtrapezoidal, widest at base, lateral margins in basal half straight and subparallel, in anterior half strongly convex and convergent anteriorly; anterior angles weakly produced and blunt, well-rounded in the tip; posterior angles blunt; anterior margin weakly convex, marginal line fine but complete; basal margin without marginal line; surface finely and densely punctate, with long, backwards-directed setae mixed with short adpressed ones; anterior and lateral margins with dense setae. Hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with coarse and dense punctures, with short adpressed setae. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate; intervals flat, with fine, dense punctures; intervals with dense, long setae being directed posteriorly; epipleural margin robust, ending before rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a rim of microtrichomes. Ventral surface shiny, metasternum and metacoxa with fine and dense punctures, densely finely setose; metacoxa with fine adpressed setae, except for a several long setae laterally. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.38. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, otherwise with fine dense setae; penultimate sternite apically without distinct chitinous border. Pygidium moderately convex, shiny, with fine, dense punctures and dense, long setae. Legs moderately wide and long; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur shiny, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin straight, not widened in apical half, smooth, dorsal posterior margin also not serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width / length: 1 / 3.4; dorsal margin sharply carinate, with two groups of spines, basal one at one third, apical one two thirds of metatibial length; basally with a few fine setae; lateral face longitudinally convex, with dense and coarse punctures, and moderately long setae in the punctures; ventral margin serrate, with four equidistant robust setae; medial face coarsely but sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, laterally also moderately carinate; first metatarsomere distinctly shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin bluntly extended medially. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 3 I – K. Habitus: Fig. 3 L.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAAFFBCFF4359EBFCBCFDAA.taxon	description	Figures 4 A – D, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAAFFBCFF4359EBFCBCFDAA.taxon	materials_examined	Type material examined. Holotype: ♂ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 19.5. 1938 (Fukien) / Type Serica adspersa G. Frey 1972 ” (ZMFK). Paratypes: 1 ♀ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 19.5. 1938 (Fukien) / Paratype Serica adspersa G. Frey 1972 ” (ZMFK), 1 ♀ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 17.5. 1938 (Fukien) / Type Serica adspersa G. Frey 1972 ” (ZMFK), 1 ♀ “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 24.5. 1938 (Fukien) / Type Serica adspersa G. Frey 1972 ” (ZMFK). Additional material examined. 3 ex. “ Chine 21. IV. 46 Kuatun, Fukien leg. Tschung-Sen ” (MHNG), 2 ex. “ Chine 1. V. 46 Kuatun, Fukien leg. Tschung-Sen ” (MHNG), 1 ex. “ Chine 12. VI. 46 Kuatun, Fukien leg. Tschung-Sen ” (MHNG), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 19.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 15.6. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 3.6. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 2 ex. “ China, Fujian c., 21. - 22. VI. Ziyungdongshan, NW slopes 25 ° 46 ’ N 117 ° 20 ’ E, 900 - 1100 m Jaroslav Turna leg., 2007 ” (ZFMK), 1 ♂ “ China, W Jiangxi Jinggang Shan-Ciping 2 - 14. VI. 1994 E. Jendek & O. Sausa leg. / CS 78 ” (CP), 2 ♂♂, 1 ♀ “ Huangkeng Aotou, Jianyang, Fujian, 6. V. 1960, 750 - 950 m, leg. Zhang Yiran ” (IZAS), 1 ♀ “ Sangang, Chong’anxingcun, Fujian, 26. V. 1960, 750 m, leg. Jiang Shengqiao ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFAAFFBCFF4359EBFCBCFDAA.taxon	description	Description of the holotype. Length: 10.6 mm, length of elytra: 7.5 mm, width: 5.6 mm. Body oval, reddish brown, elytra and pygidium yellowish brown, elytra with dense small dark spots, dull, partly with greenish toment, antenna yellow; dorsal surface almost glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex and strongly convergent anteriorly; anterior angles moderately rounded; anterior margin distinctly emarginate medially; margins weakly reflexed; surface flat and moderately shiny, on base broadly dull, finely and densely, partly rugosely punctate, with a few sparse erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and flat, twice as wide as long; ocular canthus short and slender (1 / 3 of ocular diameter), finely densely punctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, glabrous, with a few fine setae beside eyes. Eyes small, ratio diameter / interocular width: 0.58. Antenna with 9 antennomeres; club with three antennomeres, 0.9 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum weakly produced medially, narrowly but distinctly emarginate medially. Pronotum narrow and subtrapezoidal, widest at base, lateral margins almost entirely straight, evenly and weakly convex and well convergent anteriorly; anterior angles well produced and sharp; posterior angles blunt, weakly rounded; anterior margin straight, marginal line widely lacking; surface densely finely punctate, with minute setae in punctures; anterior and lateral margins sparsely setose; hypomeron carinate at base, moderately produced ventrally. Scutellum moderately large, triangular, finely and densely punctate, with minute setae in punctures. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals weakly convex, with fine, sparse punctures concentrated along striae; with a few fine, short setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, glabrous (sparse setae in available specimens probably abraded); apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, glabrous; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.37. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being a quarter as long as sternite. Pygidium weakly convex and dull, finely and densely punctate, without smooth midline, with sparse, short setae on apical half. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Posterior legs in holotype lacking. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle. Aedeagus: Fig. 4 A – C. Habitus: Fig. 4 D. Variation. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.0, dorsally sharply carinate, with only one group of spines at three quarters of metatibial length, basally with a few single, robust setae in row beside the dorsal margin; external face densely and finely punctate, without longitudinal wrinkles, with short or minute setae in punctures; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally not carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protarsal claws asymmetrical, basal tooth of inner claw sharply pointed at apex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA8FFBDFF435B17FF42FC5E.taxon	description	Figures 4 E – H, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA8FFBDFF435B17FF42FC5E.taxon	materials_examined	Type material examined. Holotype: ♂ “ Hte Birmanie Mines des Rubies 1200 m 2300 m Doherty 1890 / Ophthalmoserica solivaga type Brsk. / Museum Paris ex. Coll. R. Oberthür / Type ” (MNHN). Additional material examined. Myanmar: 1 ♂ “ N. E. Burma Kambaiti, 7000 ft. 19 / 4. 1934 R. Malaise / 3296 E 91 ” (NHRS), 1 ♂ “ N. E. Burma Kambaiti, 2000 m 28 / 5. 1934 R. Malaise / 3299 E 91 ” (NHRS), 1 ♂ “ N. E. Burma Kambaiti, 7000 ft. 12 / 5. 1934 R. Malaise / Serica solivaga Determined from description G. J. A / 3169 E 91 ” (NHRS), 2 ex. “ N. E. Burma Kambaiti 7000 ft. 8 / 4.1934 / N. E. Burma R. Malaise B. M. 1945 - 71 ” (BMNH), 2 ex. “ N. E. Burma Kambaiti 7000 ft. 18 / 4.1934 / N. E. Burma R. Malaise B. M. 1945 - 71 ” (BMNH), 1 ex. “ N. E. Burma Kambaiti 7000 ft. 3 - 7 / 5.1934 / N. E. Burma R. Malaise B. M. 1945 - 71 ” (BMNH), 1 ex. “ N. E. Burma Kambaiti 7000 ft. 28 / 5.1934 / N. E. Burma R. Malaise B. M. 1945 - 71 ” (BMNH), 1 ex. “ N. E. Burma Kambaiti 7000 ft. 13 / 5.1934 / N. E. Burma R. Malaise B. M. 1945 - 71 ” (BMNH), 18 ex. “ Myanmar (Burma) Provinz Kanchin State, Ca. 20 km N von Panwar, 23.05.2006, 2180 m, N 25 ° 43 ’ 30.2 ’’ E 098 ° 23 ’ 35.3 ’’, Nachtfang, leg. M. Langer, S. Naumann & S. Löffler Coll. Michael Langer ” (ZFMK), 1 ♂ “ Myanmar: Danasho 2800 m, Minshia vi- 2004 leg. Yin et al. ” (ZFMK). China: 4 ♂♂ “ China Yunnan prov. Gaologongshan mts. 90 km W of Baoshan S. Becvar leg. 26 - 29. v. 1995 ” (CP), 3 ex. “ China: E-Yunnan; Damaidi 2500 m, Guangnan near Vietnam VII- 2003 leg. Li et al. ” (ZFMK), 2 ex. “ China, W Yunnan prov., mts. 60 Km E Tengchong, 2300 m, 14. - 19. v. 2006 S. Murzin & I. Shokin leg. ” (CP), 1 ♂ “ Yunnan 2200 - 2500 m 24.57 N 98.45 E 8. - 16 / 5 Gaoligong mts. Vit Kuban leg. 1995 ” (CP), 1 ♂ “ Hetouzhai, Jinping, Yunnan, 12. V. 1956, 1600 - 1700 m, leg. Huang Keren etc. ” (IZAS), 1 ♂ “ Hetouzhai, Jinping, Yunnan, 14. V. 1956, 1700 m, light trap, leg. Huang Keren ” (IZAS), 2 ♂♂ “ Jinping, Yunnan, 18. V. 2006, 2000 m, leg. Cui Jianxin ” (CAU), 1 ♂ “ Yingjiang, Yunnan, 13,14. IV. 1980, 1700 m, leg. Gao Ping ” (IZAS), 3 ♂♂ “ Yingjiang, Yunnan, 15. IV. 1980, 1700 m, leg. Gao Ping ” (IZAS), 3 ♂♂ “ Hetouzhai, Jinping, Yunnan, 9, 14. V. 1956, 1700 m, light trap, leg. Huang Keren etc. ” (IZAS), 1 ♂ “ Jinping, Yunnan, 18. V. 2006, 2000 m, leg. Cui Jianxin ” (CAU).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA8FFBDFF435B17FF42FC5E.taxon	description	Redescription of the holotype. Length: 8.5 mm, elytral length: 5.5 mm, width: 4.5 mm. Body oblong, dark brown, dull, legs brown, antenna yellow, dorsal surface almost glabrous. Labroclypeus narrow, subtrapezoidal, lateral margins straight and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin weakly sinuate medially, margins strongly reflexed; surface moderately shiny, flat, finely and densely punctate, with a few erect setae; frontoclypeal suture indistinct, curved and slightly elevated; ocular canthus moderately long and narrow (1 / 3 of ocular width), impunctate, with a short terminal seta. Frons with fine, moderately dense punctures, with a few long setae besides eyes. Antenna with 10 antennomeres, club with three antennomeres, strongly reflexed, 2.2 times as long as remaining antennomeres combined. Eyes large, ratio diameter / interocular width: 0.88. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins evenly moderately convex and convergent anteriorly; anterior angles blunt and moderately produced; posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface finely and moderately densely punctate, with minute seta in punctures. Hypomeron indistinctly carinate and not produced ventrally. Scutellum wide, triangular, sparsely punctate, with minute setae in punctures. Elytra narrow, widest in posterior third, striae distinctly impressed, finely punctate; intervals weakly convex, finely and densely punctate, punctures concentrated along striae; intervals almost glabrous apart from minute setae in punctures, odd intervals with a few single short setae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, densely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face dull, coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long or robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.4. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, coarsely and densely punctate, almost glabrous. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur iridescent, with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, ratio width / length: 1 / 4.2, sharply carinate dorsally, with two groups of spines, basal one shortly before middle, apical one at almost four fifths of metatibial length, basally with a few fine and short setae; lateral face longitudinally convex, finely and sparsely punctate, glabrous; apex interiorly sharply truncate (in an angle of ca. 45 ° to tibial axis) near tarsal articulation. Tarsomeres impunctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere distinctly shorter than following two tarsomeres combined and one third of its length longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetric, interior tarsal claws bluntly truncate. Aedeagus: Fig. 4 E – G. Habitus: 4 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA8FFBDFF435B17FF42FC5E.taxon	discussion	Remarks. The record of S. solivaga from “ northern Manchuria ” of Sawada (1939) refers to Serica rosinae Pic, 1904.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA9FFB2FF435D43FF41FC16.taxon	description	Figures 4 I – L, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA9FFB2FF435D43FF41FC16.taxon	materials_examined	Type material examined. Holotype: ♂ “ Mengsong, Menglongbanna, Yunnan, 27. IV. 1958, 1600 m, leg. Wang Shuyong / LW- 172 ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA9FFB2FF435D43FF41FC16.taxon	description	Description of the holotype. Length: 7.6 mm, elytral length: 5.5 mm, width: 4.2 mm. Body oblong, dark brown, dull, legs brown, antenna yellow, dorsal surface almost glabrous. Labroclypeus narrow, subtrapezoidal, lateral margins straight and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin weakly sinuate medially, margins strongly reflexed; surface moderately shiny, flat, finely and densely punctate, with a few erect setae; frontoclypeal suture indistinct, curved and slightly elevated; ocular canthus moderately long and narrow (1 / 3 of ocular width), impunctate, with a short terminal seta. Frons with fine, moderately dense punctures, with a few long setae besides eyes. Antenna with 10 antennomeres, club with three antennomeres, strongly reflexed, 2.5 times as long as remaining antennomeres combined. Eyes very large, ratio diameter / interocular width: 0.98. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins evenly moderately convex and convergent anteriorly; anterior angles blunt and moderately produced; posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface finely and moderately densely punctate, with minute seta in punctures. Hypomeron indistinctly carinate and not produced ventrally. Scutellum wide, triangular, sparsely punctate, with minute setae in punctures. Elytra narrow, widest at middle, striae distinctly impressed, finely punctate; intervals weakly convex, finely and densely punctate, punctures concentrated along striae; intervals almost glabrous apart from minute setae in punctures, odd intervals with a few single short setae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, densely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face dull, coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long or robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.79. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, coarsely and densely punctate, almost glabrous, with a few short setae along apical margin. Legs moderately long and wide, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur almost dull, with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia moderately long and wide, ratio width / length: 1 / 3.2, sharply carinate dorsally, with two groups of spines, basal one shortly before middle, apical one at almost four fifths of metatibial length, basally with a few fine and short setae; lateral face longitudinally convex, finely and sparsely punctate, glabrous; ventral margin with three robust spines of which the two distal ones are more distant to each other; mesal face impunctate and glabrous; apex interiorly sharply truncate (in an angle of ca. 45 ° to tibial axis) near tarsal articulation. Tarsomeres impunctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere as long as following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetric, interior tarsal claws bluntly truncate. Aedeagus: Fig. 4 I – K. Habitus: 4 L. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA9FFB2FF435D43FF41FC16.taxon	diagnosis	Diagnosis. Serica mengsongana Ahrens, Fabrizi & Liu, new species differs from the similar S. solivaga Brenske, 1898 by the wider and shorter hind legs, including metatibia, and the morphology of aedeagus: the apex of the phallobase is more straight distally and less bent as in S. solivaga; the internal lobe of the left paramere is in S. mengsongana Ahrens, Fabrizi & Liu, new species mesal rather than basal, as in S. solivaga; and the right paramere in the new species has interiorly a small sharp spine-like hook before apex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA9FFB2FF435D43FF41FC16.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from its type locality, Mengsong.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB2FF435A90FAADFAF6.taxon	description	Figures 5 A – D, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB2FF435A90FAADFAF6.taxon	materials_examined	Type material examined. Syntypes: 1 ♂ “ N. E. Burma Kambaiti 7000 ft. 25 - 27 / 4 / 1934 R. Malaise / ♂ / Cotype / N. E. Burma R. Malaise B. M. 1945 - 71 / Serica lupina co-type Arrow ” (BMNH), 1 ♂ “ N. E. Burma Kambaiti 7000 ft. 25 - 27 / 4 / 1934 R. Malaise / ♂ / Serica lupina co-type Arrow ” (NHRS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB2FF435A90FAADFAF6.taxon	discussion	Remarks. Unfortunately, we had no borrowed type specimens or any additional specimens for a redescription at hand when finalizing this revision. Images shown here date back to 2007 when D. A. visited the NHRS.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB3FF435CEBFF52FB12.taxon	description	Figures 5 E – H, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB3FF435CEBFF52FB12.taxon	materials_examined	Type material examined. Holotype: ♂ “ China: E-Yunnan; Damaidi 2500 m, Guangnan near Vietnam VII- 2003 leg. Li et al. / 717 Sericini Asia spec. ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB3FF435CEBFF52FB12.taxon	description	Description of the holotype. Length: 8.4 mm, elytral length: 5.9 mm, width: 4.6 mm. Body oblong, dark brown, dull, legs brown, antenna yellow, with numerous short and long setae on dorsal surface. Labroclypeus wide, trapezoidal, lateral margins convex and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin weakly sinuate medially, margins strongly reflexed; surface weakly convex, finely and densely punctate, with numerous erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 2 short terminal setae. Frons with coarse, dense punctures and numerous long setae. Antenna with nine antennomeres, club with three antennomeres, as long as remaining antennomeres combined. Eyes small, ratio diameter / interocular width: 0.53. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and weakly convergent, in anterior half evenly convex and convergent anteriorly; anterior angles sharp and distinctly produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface coarsely and densely punctate, with numerous short and longer adjacent or erect yellow setae, otherwise punctures with microscopic setae only. Hypomeron carinate but not produced ventrally. Scutellum wide, triangular, punctation as on pronotum, with numerous short adjacent setae. Elytra narrow, widest in posterior third, striae distinctly impressed, finely punctate, odd intervals weakly convex, even ones flat, finely and densely punctate, punctures concentrated along striae in odd intervals, elytra covered with numerous long adjacent setae apart from minute setae in punctures; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long or robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.7. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium moderately convex, coarsely and densely punctate, with dense, long setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia moderately long and wide, ratio width / length: 1 / 3.6, sharply carinate dorsally, with two groups of spines, basal one at three fifths, apical one at four fifths of metatibial length, basally with a few robust and short setae in line; lateral face longitudinally convex, finely and superficially sparsely punctate; apex concavely sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere as long as following two tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 5 E – G. Habitus: 5 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB3FF435CEBFF52FB12.taxon	diagnosis	Diagnosis. Serica guangnanensis Ahrens, Fabrizi & Liu, new species differs from Serica lupina Arrow, 1946 by the short antennal club in male, which is as long as remaining antennomeres combined, and of course by the shape of the aedeagus: phallobase apically deeply emarginated between the insertino of parameres and parameres subequal in length being moderatly long and curved dorsally.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA6FFB3FF435CEBFF52FB12.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from its type locality Guangnan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA7FFB1FF435D8FFD9AFA82.taxon	description	Figures 5 I – L, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA7FFB1FF435D8FFD9AFA82.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, Yunnan prov. 1. - 19.7.1992, Heishui 35 km N Lijiang 27.13 N; 100.19 E lgt. S. Becvar / 557 Sericini: Asia spec. ” (ZFMK). Paratypes: 2 ♂♂ “ China, Yunnan prov. Heishui 35 km N Lijiang 18.6. - 4.7.1992, 27.13 N; 100.19 E lgt. S. Becvar ” (CP), 1 ♂ “ Mts. Yulongshan, Lijiang, Yunnan, 27. VI. 1962, leg. Song Shimei ” (IZAS), 2 ♂♂ “ China, Yunnan prov.; 1993 Heishui 35 km N Lijiang; 27.13 N; 100.19 E; S. Becvar leg.; 18. vi. - 4. vii. / 859 Sericini: Asia spec. ” (CP), 1 ♂ “ Yunnan 1800 - 2500 m 25.10 N 100.21 E Weishan mt. 22. - 25.6.92 David Kral lgt. ” (NMPC), 2 ♂♂, 4 ♀♀ “ China: N-Yunnan, Baiyungshan (Rai Railing Mts.) 2400 m, Yong Ren, VII- 2003 leg. Ying et al. ” (ZFMK), 1 ♂ “ Nongkesuo, Bijie, Guizhou, 1970 s, No. 113, leg. Yang / LW- 191 ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA7FFB1FF435D8FFD9AFA82.taxon	description	Description of the holotype. Length: 9.1 mm, elytral length: 6.6 mm, width: 4.4 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with numerous short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins weakly convex and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin moderately sinuate medially, margins strongly reflexed; surface flat, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 3 short terminal setae. Frons with dull toment, with coarse, dense punctures and numerous long setae. Antenna with nine antennomeres, club with three antennomeres, twice as long as remaining antennomeres combined, moderately reflexed. Eyes large, ratio diameter / interocular width: 0.74. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and subparallel, in anterior half convex and convergent anteriorly; anterior angles almost blunt and weakly produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with mixed coarse and fine, dense punctures, with numerous short and longer erect or forward-directed yellow setae. Hypomeron very weakly carinate ventrally. Scutellum narrow and long, triangular, with fine, dense punctation, at base with a smooth impunctate triangular area, with numerous short adjacent setae. Elytra narrow, elongate, widest in posterior third, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, elytra with dense, long erect setae, minute setae absent; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.5. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, finely and densely punctate, with dense, long setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 4.76, sharply carinate dorsally, with four groups of spines, basal one at first quarter, second one at middle, third one at two thrids, apical one at four fifths of metatibial length, basally glabrous; lateral face longitudinally convex, finely and irregularly densely punctate, glabrous; apex concavely and narrowly sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 5 I – K. Habitus: 5 L.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA7FFB1FF435D8FFD9AFA82.taxon	diagnosis	Diagnosis. Serica longipes Ahrens, Fabrizi & Liu, new species differs from S. guangnanensis Ahrens, Fabrizi & Liu, new species by the densely and erectly setose dorsal body surface, the more slender metatibiae, and the shape of parameres, which are shorter and in lateral view almost straight.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA7FFB1FF435D8FFD9AFA82.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from the Latin combined words ‘ long ’ (i. e., long) and ‘ pes ’ (feed / leg), with reference on the long legs of the species.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA7FFB1FF435D8FFD9AFA82.taxon	description	Variation. Length: 9.0 – 9.8 mm, elytral length: 6.5 – 8.2 mm, width: 4.4 – 5.2 mm. Female: antennal club short, as long as remaining antennomeres combined; pygidium weakly convex; eyes much smaller than in male, ratio diameter / interocular width: 0.55.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA5FFB7FF435CFFFC2CFC16.taxon	description	Figures 6 A – D, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA5FFB7FF435CFFFC2CFC16.taxon	materials_examined	Type material examined. Holotype: ♂ “ Nanhua, Yunnan, 7. V. 1982, 2400 m, leg. Yu Peiyu ” (IZAS). Paratypes: 9 ♂♂, 1 ♀ “ China, W Yunnan prov., mts. 60 km E Tengchong, 2300 m, 14. - 19. v. 2006, S. Murzin & I. Shokhin ” (CP, ZFMK), 3 ♂♂, 5 ♀♀ “ China, W Yunnan prov., mts. 60 km E Tengchong, 2300 m, 19. - 22. v. 2006, S. Murzin & I. Shokhin ” (CP, ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA5FFB7FF435CFFFC2CFC16.taxon	description	Description of the holotype. Length: 8.4 mm, elytral length: 6.0 mm, width: 4.4 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with numerous short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins convex and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin moderately sinuate medially, margins strongly reflexed; surface flat, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 3 short terminal setae. Frons with dull toment, with coarse, dense punctures and numerous long setae. Antenna with nine antennomeres, club with three antennomeres, twice as long as remaining antennomeres combined, moderately reflexed. Eyes large, ratio diameter / interocular width: 0.83. Mentum convexly elevated and flattened anteriorly. Pronotum widest in posterior third, lateral margins evenly convex and convergent anteriorly and posteriorly; anterior angles almost blunt and weakly produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with mixed coarse and fine, dense punctures, with numerous short and longer erect or forward-directed yellow setae. Hypomeron very weakly carinate ventrally. Scutellum narrow and long, triangular, with fine, dense punctation, at base with a smooth impunctate triangular area, with numerous short adjacent setae. Elytra narrow, elongate, widest in posterior third, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, elytra with dense, long erect setae, minute setae absent; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.54. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, finely and densely punctate, with dense, long setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 3.9, sharply carinate dorsally, with four groups of spines, basal one at first quarter, second one at middle, third one at two thrids, apical one at four fifths of metatibial length, basally glabrous; lateral face longitudinally convex, finely and irregularly densely punctate, glabrous; apex concavely and narrowly sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 6 A – C. Habitus: 6 D.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA5FFB7FF435CFFFC2CFC16.taxon	diagnosis	Diagnosis. Serica nanhua Ahrens, Fabrizi & Liu, new species differs from Serica longipes Ahrens, Fabrizi & Liu, new species by the distinctly less curved left paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA5FFB7FF435CFFFC2CFC16.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from the type locality, Nanhua.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA5FFB7FF435CFFFC2CFC16.taxon	description	Variation. Length: 8.4 – 9.2 mm, elytral length: 6.0 – 6.9 mm, width: 4.4 – 4.9 mm. The shape of pronotum may considerably variate, in some specimens widest at base with margin in basal half straight and subparallel, as in S. longipes. Female: antennal club short, as long as remaining antennomeres combined; pygidium weakly convex; eyes much smaller than in male, ratio diameter / interocular width: 0.55.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA3FFB4FF435A90FBD8FC5E.taxon	description	Figures 6 E – H, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA3FFB4FF435A90FBD8FC5E.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, Yunnan prov. Heishui 35 km N Lijiang 18.6. - 4.7.1992, 27.13 N; 100.19 E lgt. S. Becvar / 1135 Sericini: Asia spec. ” (ZFMK). Paratypes: 2 ♂♂ “ China, Yunnan prov. Heishui 35 km N Lijiang 18.6. - 4.7.1992, 27.13 N; 100.19 E lgt. S. Becvar ” (CP), 2 ♂♂ “ China, Yunnan prov. Heishui 35 km N Lijiang 18.6. - 4.7.1993, 27.13 N; 100.19 E lgt. S. Becvar ” (ZFMK), 1 ♂ “ Mts. Yulongshan, Lijiang, Yunnan, 28. VI. 1962, leg. Song Shimei ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA3FFB4FF435A90FBD8FC5E.taxon	description	Description of the holotype. Length: 7.8 mm, elytral length: 6.1 mm, width: 4.1 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with sparse short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins straight and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin moderately sinuate medially, margins strongly reflexed; surface flat, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 3 short terminal setae. Frons with dull toment, with coarse, dense punctures and a few long setae. Antenna with nine antennomeres, club with three antennomeres, 2.5 times as long as remaining antennomeres combined, strongly reflexed. Eyes moderately large, ratio diameter / interocular width: 0.73. Mentum convexly elevated and flattened anteriorly. Pronotum widest at middle, lateral margins in basal half straight and subparallel, in anterior half convex and convergent anteriorly, only little narrowed towards base; anterior angles almost blunt and weakly produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with mixed coarse and fine, dense punctures, with a few short and longer erect or forward-directed yellow setae. Hypomeron very weakly carinate ventrally. Scutellum narrow and long, triangular, with fine, dense punctation, at base with a smooth impunctate triangular area, with a very few short setae. Elytra narrow, elongate, widest in posterior quarter, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, elytra with moderately dense, long and erect setae in row on each sides of odd intervals, minute setae absent; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long setae, otherwise almost glabrous. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.55. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium strongly convex, finely and densely punctate, with moderately dense, long setae on apical half. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 3.9, sharply carinate dorsally, with only one group of spines at four fifths of metatibial length, basally of it beside the dorsal margin with a 3 – 4 single robust setae over the entire metatibial length; lateral face longitudinally convex, finely and irregularly densely punctate, glabrous; apex concavely and narrowly sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 6 E – G. Habitus: Fig. 6 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA3FFB4FF435A90FBD8FC5E.taxon	diagnosis	Diagnosis. Serica allolongipes Ahrens, Fabrizi & Liu, new species differs from Serica longipes Ahrens, Fabrizi & Liu, new species by the much less dense pilosity on dorsal surface and the shape of parameres. The right paramere is bluntly widened before apex and almost pointed at apex, while in S. longipes the lateral blunt extension is absent and the apex is distinctly rounded. Furthermore, the left paramere is in basal half much more convexly widened.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA3FFB4FF435A90FBD8FC5E.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from the combined Greek prefix ‘ allo- ’ (different, other) and the species name ‘ longipes ’, with reference on the similarity to this synoccurring species.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA3FFB4FF435A90FBD8FC5E.taxon	description	Variation. Length: 7.8 – 9.4 mm, elytral length: 6.1 – 6.6 mm, width: 4.1 – 4.5 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA0FF8AFF435D48FBD8FC5E.taxon	description	Figures 7 A – E, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA0FF8AFF435D48FBD8FC5E.taxon	materials_examined	Type material examined. Holotype ♂ “ China: N-Yunnan, Baiyungshan (Rai Railing Mts.) 2400 m, Yong Ren, VII- 2003 leg. Ying et al. ” (ZFMK). Paratype: 1 ♂ “ China: N-Yunnan, Baiyungshan (Rai Railing Mts.) 2400 m, Yong Ren, VII- 2003 leg. Ying et al. ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA0FF8AFF435D48FBD8FC5E.taxon	description	Description of the holotype. Length: 7.8 mm, elytral length: 5.6 mm, width: 4.9 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with sparse short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins straight and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin moderately sinuate medially, margins strongly reflexed; surface weakly convex, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 3 short terminal setae. Frons with dull toment, with coarse, dense punctures and numerous long erect setae. Antenna with nine antennomeres, club with three antennomeres, 2.2 times as long as remaining antennomeres combined, strongly reflexed. Eyes small, ratio diameter / interocular width: 0.54. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and weakly convergent towards middle, in anterior half margins convex and convergent anteriorly; anterior angles almost blunt and moderately produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with mixed coarse and fine, dense punctures, with a few short and longer yellow setae. Hypomeron very weakly carinate ventrally. Scutellum narrow and long, triangular, with fine, dense punctation, at base with a smooth impunctate triangular area, with a very few short setae. Elytra narrow, elongate, widest in posterior quarter, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, elytra with moderately dense, long and erect setae in row on each sides of odd intervals, minute setae absent; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long setae, otherwise almost glabrous. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.62. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, finely and densely punctate, with a few sparse, long setae on apical half. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 4.1, sharply carinate dorsally, with two groups of robust spines, basal one shortly behind middle apical one at four fifths of metatibial length, basally with a few single robust setae; lateral face longitudinally convex, coarsely and densely punctate, glabrous; apex concavely and narrowly sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 7 A – D. Habitus: 7 E. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA0FF8AFF435D48FBD8FC5E.taxon	diagnosis	Diagnosis. Serica paralongipes Ahrens, Fabrizi & Liu, new species differs from sympatric Serica longipes Ahrens, Fabrizi & Liu, new species by the much less dense pilosity on dorsal surface and the left paramere which is narrower and sickle-shaped, almost circular in cross section over almost its entire length. From S. allolongipes Ahrens, Fabrizi & Liu, new species Serica paralongipes Ahrens, Fabrizi & Liu, new species differs by the simply rounded apex of right almost straight paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA0FF8AFF435D48FBD8FC5E.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from the combined Greek prefix ‘ para - ’ (false) and the species name ‘ longipes ’, with reference on the similarity to this synoccurring species.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFA0FF8AFF435D48FBD8FC5E.taxon	description	Variation. Length: 7.6 – 7.8 mm, elytral length: 5.5 – 5.6 mm, width: 4.8 – 4.9 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9EFF8BFF435D43FBD8FC16.taxon	description	Figures 7 F – J, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9EFF8BFF435D43FBD8FC16.taxon	materials_examined	Type material examined. Holotype: ♂ “ Yunnan 2200 - 2500 m 24.57 N 98.45 E 8 - 16 / 5 Gaoligong mts. Vit Kuban leg. 1995 / 1136 Sericini: Asia spec. ” (CP). Paratypes: 1 ♂ “ Yunnan 2200 - 2500 m 24.57 N 98.45 E 8 - 16 / 5 Gaoligong mts. Vit Kuban leg. 1995 ” (ZFMK), 2 ♂♂ “ N-Vietnam, Lao Cai Prov., Hoang Lien NP, Tram Ton, N 22 ° 21.197 ’ E 103 ° 46.513 ’, 1800 - 2050 m 13. - 16.5.2015, leg. A. Skale ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9EFF8BFF435D43FBD8FC16.taxon	description	Description of the holotype. Length: 9.6 mm, elytral length: 6.9 mm, width: 4.9 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with numerous short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins stright and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin moderately sinuate medially, margins strongly reflexed; surface flat, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 3 short terminal setae. Frons with dull toment, with coarse, dense punctures and numerous long setae. Antenna with nine antennomeres, club with three antennomeres, three times as long as remaining antennomeres combined, strongly reflexed. Eyes small, ratio diameter / interocular width: 0.58. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and weakly convergent, in anterior half convex and distinctly convergent anteriorly; anterior angles almost blunt, weakly rounded at tip, and weakly produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with mixed coarse and fine, dense punctures, with numerous short and longer erect or forward-directed yellow setae. Hypomeron very weakly carinate ventrally. Scutellum narrow and long, triangular, with fine, dense punctation, at base with a smooth impunctate triangular area, with numerous short adjacent setae. Elytra narrow, elongate, widest in posterior quarter, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, elytra with moderately dense, long and erect setae in row on each sides of odd intervals, minute setae absent; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.47. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, finely and densely punctate, with a few short setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 4.1, sharply carinate dorsally, with four groups of spines, basal one at first quarter, second one at middle, third one at two thrids, apical one at four fifths of metatibial length, basally glabrous; lateral face longitudinally convex, finely and irregularly densely punctate, glabrous; apex concavely and narrowly sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 7 F – I. Habitus: Fig. 7 J. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9EFF8BFF435D43FBD8FC16.taxon	diagnosis	Diagnosis. Serica gaoligong Ahrens, Fabrizi & Liu, new species differs from Serica longipes Ahrens, Fabrizi & Liu, new species by the shape of parameres: the right paramere is more strongly bent at middle and sarply pointed at apex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9EFF8BFF435D43FBD8FC16.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from its type locality (Gaoligong).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9EFF8BFF435D43FBD8FC16.taxon	description	Variation. Length: 9.1 – 9.6 mm, elytral length: 6.8 – 6.9 mm, width: 4.8 – 4.9 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9FFF88FF435A8BFACCFCA6.taxon	description	Figures 7 K – N, 22	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9FFF88FF435A8BFACCFCA6.taxon	materials_examined	Type material examined. Holotype: ♂ “ Myanmar: Danasho 2800 m, Minshia vi- 2004 leg. Yin et al. / Phil # 11 - 2004: 4. / 1134 Sericini: Asia spec. ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9FFF88FF435A8BFACCFCA6.taxon	description	Description of the holotype. Length: 9.8 mm, elytral length: 7.0 mm, width: 4.8 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with sparse short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins straight and moderately convergent, producing a distinct angle with ocular canthus; anterior angles strongly rounded; anterior margin moderately sinuate medially, margins strongly reflexed; surface weakly convex, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 3 short terminal setae. Frons with dull toment, with coarse, dense punctures and a few long setae. Antenna with nine antennomeres, club with three antennomeres, 2.2 times as long as remaining antennomeres combined, strongly reflexed. Eyes moderately large, ratio diameter / interocular width: 0.62. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and weakly convergent, in anterior half convex and convergent anteriorly; anterior angles rectangular and weakly produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with mixed coarse and fine, dense punctures, with a few short and longer erect or forward-directed yellow setae. Hypomeron very weakly carinate ventrally. Scutellum narrow and long, triangular, with fine, dense punctation, at base with a smooth impunctate triangular area, with a very few short setae. Elytra narrow, elongate, widest at middle, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae, elytra with moderately dense, long and erect setae in row on each sides of odd intervals, minute setae absent; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, only metasternal plate and lateral metacoxa with a few long setae, otherwise almost glabrous. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.57. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise glabrous. Pygidium moderately convex, finely and densely punctate, with moderately dense, long setae on apical half. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 4.6, sharply carinate dorsally, with two groups of robust spines, basal one shortly behind middle, apical one at four fifths of metatibial length, in basal half with a two further small robust setae beside the dorsal margin; lateral face longitudinally convex, finely and moderately densely punctate, glabrous; apex concavely and narrowly sinuate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 7 K – M. Habitus: Fig. 7 N. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9FFF88FF435A8BFACCFCA6.taxon	diagnosis	Diagnosis. Serica yini Ahrens, Fabrizi & Liu, new species differs from Serica allolongipes Ahrens, Fabrizi & Liu, new species by the shape of parameres. The apically bluntly widened right paramere has before apex an additional sharp lateral tooth and almost bluntly truncate at apex; the left paramere is robust over its entirly length and shorter, abruptly pointed at apex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9FFF88FF435A8BFACCFCA6.taxon	etymology	Etymology. The new species (adjective in nominative case) is named after one of its collectors, Mr. Yin.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9CFF8EFF435A1BFE2FFC32.taxon	description	Figures 8 A – D, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9CFF8EFF435A1BFE2FFC32.taxon	materials_examined	Type material examined. Holotype: ♂ “ China-Shaanxi, Taibashan Range, 1900 m, Houzhenzi vill. env., 1 - 12. VII. 1999, 33 ° 53 N, 107 ° 49 E, V. Siniaev & A. Plutenko lgt. / 833 Sericini: Asia spec. ” (CP). Paratypes: 4 ♂♂ “ China-Shaanxi, Taibashan Range, 1900 m, Houzhenzi vill. env., 1 - 12. VII. 1999, 33 ° 53 N, 107 ° 49 E, V. Siniaev & A. Plutenko lgt. ” (CP, ZFMK), 27 ♂♂, 2 ♀♀ “ China: Daxue Shan Mts., Sichuan, 40 km W Mianning, 7. - 8. vi. 1999, 2750 m, 28 ° 34 ’ N, 102 ° 00 ’ E, V. Siniaev & A. Plutenko lgt. ” (CP, ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9CFF8EFF435A1BFE2FFC32.taxon	description	Description of the holotype. Length: 9.2 mm, elytral length: 6.8 mm, width: 4.6 mm. Body oblong, dark yellowish brown, frons dark brown, dull, legs brown, antenna yellow, except a few short setae on head dorsal surface glabrous. Labroclypeus wide, trapezoidal, lateral margins weakly convex and convergent, producing a blunt angle with ocular canthus; anterior angles strongly convex; anterior margin distinctly emarginate medially; margins moderately reflexed; surface weakly convex, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture distinct, moderately curved; ocular canthus short and moderately narrow (1 / 4 of ocular width), smooth and glabrous, without terminal setae. Frons with dull toment, with fine, moderately dense punctures, anteriorly with a few long setae. Antenna with nine antennomeres, club with three antennomeres, 2.7 times as long as remaining antennomeres combined, distinctly reflexed. Eyes moderately large, ratio diameter / interocular width: 0.69. Mentum convexly elevated and flattened anteriorly. Pronotum moderately wide, widest at base, lateral margins evenly modertely convex and convergent anteriorly; anterior angles almost blunt, weakly produced, posterior angles blunt; anterior margin weakly convex, marginal line fine but complete; basal margin without marginal line; surface with coarse, dense punctures and glabrous, also without minute setae. Hypomeron not carinate ventrally. Scutellum narrow and long, triangular, with fine but very dense punctation, glabrous, basally with a triangular smooth area. Elytra elongate and narrow, widest in posterior third; striae distinctly impressed, finely punctate; intervals weakly convex, finely and densely punctate, punctures concentrated along striae; glabrous, lateral intervals with a few moderately long, adpressed or erect setae; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a very fine membraneous rim of short microtrichomes. Ventral face finely and densely punctate, glabrous. Metacoxae laterally with a few longer setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.43. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise also glabrous. Pygidium almost flat, finely and densely punctate, with moderately dense, long setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia long and narrow, widest at apex, ratio width / length: 1 / 4.4, sharply carinate dorsally, with two groups of spines, basal one at three fifths, apical one at four fifths of metatibial length, basally with a number of robust single setae; lateral face longitudinally convex, finely and moderately densely punctate, glabrous; apex shallowly concavely truncate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and little shorter than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 8 A – C. Habitus: Fig. 8 D.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9CFF8EFF435A1BFE2FFC32.taxon	diagnosis	Diagnosis. Serica paralupina Ahrens, Fabrizi & Liu, new species differs from Serica lupina Arrow, 1946 and all other species of this species group by the lack of dense dorsal pilosity, and, of course, by the shape of aedeagus: the apical phallobase is strongly asymmetric having the left paramere inserted more basally and displaced ventrally under the apical aedeagus.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9CFF8EFF435A1BFE2FFC32.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derrived from the combined Gree prefix ‘ pseudo - ’ (false) and the species name ‘ lupina ’, with reference to the similarity of the species to Serica lupina Arrow, 1946.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9CFF8EFF435A1BFE2FFC32.taxon	description	Variation. Length: 8.8 – 9.2 mm, elytral length: 6.4 – 6.8 mm, width: 4.4 – 4.6 mm. Female: Antennal club short, as long as reminaing antennomeres combined; eyes smaller than in male, ratio diameter / interocular width: 0.56; pygidium weakly convex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9AFF8FFF435AAFFE0AFC16.taxon	description	Figures 8 E – H, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9AFF8FFF435AAFFE0AFC16.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, NW Yunnan prov., Baishuitai, (San Ba terasses) 2600 m, 4. - 6. vi. 2006, S. Murzin & I. Shokhin / 851 Sericini: Asia spec. ” (CP). Paratypes: 13 ♂♂, 3 ♀♀ “ China, NW Yunnan prov., Baishuitai, (San Ba terasses) 2600 m, 4. - 6. vi. 2006, S. Murzin & I. Shokhin ” (CP, ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9AFF8FFF435AAFFE0AFC16.taxon	description	Description of the holotype. Length: 7.6 mm, elytral length: 5.5 mm, width: 4.4 mm. Body oblong, dark reddish brown, dull, legs brown, antenna yellow, with numerous short and long setae on dorsal surface. Labroclypeus moderately wide, trapezoidal, lateral margins straight and moderately convergent, producing a distinct angle with ocular canthus; anterior angles almost blunt, weakly rounded; anterior margin straight, only very weakly concave medially, margins strongly reflexed; surface weakly convex, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture indistinct, curved; ocular canthus moderately short and narrow (1 / 3 of ocular width), finely and densely punctate, with 1 – 2 short terminal setae. Frons with dull toment, with coarse, dense punctures and numerous short adpressed and a few long, setae bent posteriorly. Antenna with nine antennomeres, club with three antennomeres, 2.5 times as long as remaining antennomeres combined, distinctly reflexed. Eyes moderately large, ratio diameter / interocular width: 0.64. Mentum convexly elevated and flattened anteriorly. Pronotum widest at base, lateral margins in basal half straight and subparallel, in anterior half convex and convergent anteriorly; anterior angles rectangular and moderately produced, posterior angles blunt; anterior margin strongly convex, marginal line complete; basal margin without marginal line; surface with coarse, dense punctures and numerous short setae bent posteriorly and laterally dense longer yellow setae bent anteriorly. Hypomeron robustly carinate ventrally, carina produced ventrally. Scutellum narrow and long, triangular, with fine dense punctation, with numerous short adjacent setae. Elytra narrow, widest in posterior third, striae distinctly impressed, finely punctate, intervals weakly convex, finely and densely punctate, punctures concentrated along striae; with dense, short setae on disc and long erect setae bent partly anteriorly; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, with moderately dense short adpressed setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.46. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise short adpressed setae in the punctures. Pygidium moderately convex, finely and densely punctate, with dense, long setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia moderately long and narrow, widest at apex, ratio width / length: 1 / 3.8, sharply carinate dorsally, with two groups of spines, basal one at middle, apical one at five sixths of metatibial length, basally with a number of robust single setae; lateral face longitudinally convex, coarsely and irregularly punctate, with minute adpressed setae; apex shallowly emarginate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 8 E – G. Habitus: Fig. 8 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9AFF8FFF435AAFFE0AFC16.taxon	diagnosis	Diagnosis. Serica baishuitaiensis Ahrens, Fabrizi & Liu, new species differs from Serica guangnanensis Ahrens, Fabrizi & Liu, new species by the more densely setose dorsal body surface, the longer antennal club (male), and the shape of parameres, which are evenly curved ventrally (lateral view) rather than being curved dorsally as in S. guangnanensis Ahrens, Fabrizi & Liu, new species	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9AFF8FFF435AAFFE0AFC16.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from the type locality, Baishuitai.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9AFF8FFF435AAFFE0AFC16.taxon	description	Variation. Length: 7.6 – 9.1 mm, elytral length: 5.5 – 6.9 mm, width: 4.4 – 5.1 mm. Female: antennal club short, as long as remaining antennomeres combined; pygidium weakly convex; eyes smaller than in male, ratio diameter / interocular width: 0.54.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9BFF8CFF435A8BFED7FCA6.taxon	description	Figures 8 I – L, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9BFF8CFF435A8BFED7FCA6.taxon	materials_examined	Type material examined. Holotype: ♂ “ Shaanxi Huangbaiyuan 2012 - VI- 17 / LW- 1034 ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9BFF8CFF435A8BFED7FCA6.taxon	description	Description of the holotype. Length: 7.4 mm, elytral length: 5.7 mm, width: 4.5 mm. Body oblong, reddish brown, dull, legs brown, antenna yellow, dorsal surface covered with dense erect and moderately long setae. Labroclypeus wide, subrectangular, lateral margins weakly convex and weakly convergent, producing a blunt angle with ocular canthus; anterior angles strongly convex; anterior margin shallowly emarginate medially; margins moderately reflexed; surface weakly convex, finely and densely punctate, with numerous long and erect setae; frontoclypeal suture distinct, moderately curved; ocular canthus moderately long and narrow (1 / 3 of ocular width), finely punctate, without terminal seta. Frons with dull toment, with fine, moderately dense punctures, anteriorly and beside eyes with a few long setae. Antenna with nine antennomeres, club with three antennomeres, three times as long as remaining antennomeres combined, distinctly reflexed; antennomeres 4 to 6 short and transverse. Eyes moderately large, ratio diameter / interocular width: 0.7. Mentum convexly elevated and flattened anteriorly. Pronotum moderately wide, widest at base, lateral margins evenly modertely convex and convergent anteriorly; anterior angles almost blunt, weakly produced, posterior angles blunt; anterior margin weakly convex, marginal line fine but complete; basal margin without marginal line; surface with coarse, dense punctures and moderately dense erect setae, partly in punctures with minute adpressed, fine setae. Hypomeron carinate ventrally, not produced ventrally. Scutellum narrow and long, triangular, with coarse, very dense punctation and short adpressed setae, basally with a triangular smooth area. Elytra elongate and narrow, widest in posterior third; striae distinctly impressed, finely punctate; intervals weakly convex, coarsely and densely almost evenly punctate; with moderately dense, erect setae, which are dense on lateral intervals, partly with ninute adpressed setae in punctures; epipleural margin robust, ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a very fine membraneous rim of short microtrichomes. Ventral face finely and densely punctate, with short and dense setae. Metacoxae glabrous, only laterally with a few longer setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.44. Abdominal sternites finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta, otherwise with short fine setae in punctures. Pygidium almost flat, finely and densely punctate, with dense, long setae. Legs long and narrow, shiny; femora finely and sparsely punctate, with two longitudinal rows of setae. Metafemur with anterior margin acute, without adjacent serrated line; posterior ventral margin almost straight, weakly widened in apical half, ventrally and dorsally not serrated, sparsely setose. Metatibia moderately long and narrow, widest at apex, ratio width / length: 1 / 3.5, sharply carinate dorsally, with three groups of spines, basal one shortly before the middle, medium one at two thirds, apical one at four fifths of metatibial length, basally and along a longitudinal blunt carina beside the dorsal margin with a number of short, robust, single setae; lateral face almost flat, coarsely and densely punctate, with a few short setae; apex shallowly concavely truncate interiorly near tarsal articulation. Tarsomeres finely punctate dorsally, with moderately dense, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and little shorter than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetric, interior tarsal claws sharply pointed. Aedeagus: Fig. 8 I – K. Habitus: Fig. 8 L. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9BFF8CFF435A8BFED7FCA6.taxon	diagnosis	Diagnosis. Serica huangbaiyuanensis Ahrens, Fabrizi & Liu, new species differs from the other setose species of the Serica lupina group by the shorter legs, and by the basal lobe of right paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF9BFF8CFF435A8BFED7FCA6.taxon	etymology	Etymology. The name of the new species (adjective in nominative case) is derived from the type locality, Huangbaiyuan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF98FF82FF435A20FC2BFD8E.taxon	description	Figures 9 A – D, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF98FF82FF435A20FC2BFD8E.taxon	materials_examined	Type material examined. Holotype: ♂ “ China: Yunnan Prov., 8 km NW Gudong, 28. vi. 2016 Yunfeng Shan NP Lyzu temple, at light 25 ° 22.6 ’ N, 98 ° 24.4 ’ E, 2350 m J. Hájek & J. Růžička leg. / 1106 Asia Sericini spec. ” (NMPC). Paratypes: 2 ♂♂, 6 ♀♀ “ Myanmar (Burma) Provinz Kachin State Three River Junction (Thone chaung sone) N 26 ° 23 ’ 13.5 ’’ E 098 ° 39 ’ 36.3 ’’ 2. X. 2010 (H = 2044 m; LF) leg. Michael Langer, S. Naumann & S. Löffler ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF98FF82FF435A20FC2BFD8E.taxon	description	Description of the holotype. Length: 8.0 mm, length of elytra: 6.4 mm, width: 4.8 mm. Body elongate eggshaped, dark brown, partly with greenish toment or shine, elytra partly yellowish brown with dark brown spots, antenna and ventral surface including femora yellowish, tibiae brown; dorsal surface dull and almost glabrous except a few single short setae on pronotum and elytra. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins convex and convergent anteriorly, anterior angles acute; anterior margin deeply and widely emarginate medially; margins weakly reflexed; surface flat and moderately shiny, finely and moderately densely punctate, glabrous; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye small and flat, 1.3 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, without terminal seta. Frons completely dull and flat, with fine and dense punctures, almost completely glabrous, with a few short setae beside eyes. Eyes small, ratio diameter / interocular width: 0.48. Antenna with 10 antennomeres; club with three antennomeres, as long as remaining antennomeres combined and straight. Mentum strongly elevated, convex. Labrum strongly produced along the middle, deeply emarginate medially. Pronotum narrow and subtrapezoidal, widest at base, lateral margins evenly weakly convex and convergent anteriorly; anterior angles distinctly produced and acute; posterior angles blunt; anterior margin with a wide marginal line and weakly produced medially; surface densely and coarsely punctate, with sparse short yellow setae bent posteriorly and partly with minute setae in punctures; lateral and anterior margins sparsely setose; hypomeron carinate at base, not produced ventrally. Scutellum large, triangular, finely and densely punctate, on midline punctures less dense, glabrous. Elytra oval, widest in posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, moderately dense punctures concentrated along striae; dark spots impunctate; with a few single, fine, short, white, adpressed setae, without longer, erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin chitinous, without a rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, almost glabrous, with a few single setae on metasternal disc; metacoxa glabrous, except a few short lateral setae; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous and very superficially sparsely punctate, ultimate sternite densely shortly setose; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.44. Pygidium strongly convex and dull, yellow with small dark spots, finely and densely punctate, smooth midline narrow, with sparse short and longer setae on apical third. Legs slender; femora dull, with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin not serrated and only weakly widened at apex, dorsal posterior margin completely but finely serrated, in basal half with a few long setae which are one third as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.1, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, without longitudinal wrinkles; ventral margin serrated, with two extremely widely separated robust setae (abraded in holotype, but recognizable by their insertion point); medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally carinate, with a strongly serrated ridge ventrally; first metatarsomere little longer than following tarsomere and as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle (lateral protibial teeth in holotype strongly abraded); anterior claws symmetrical, basal tooth of inner claw bluntly truncate as the external one. Aedeagus: Fig. 9 A – C. Habitus: Fig. 9 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF98FF82FF435A20FC2BFD8E.taxon	diagnosis	Diagnosis. Serica jirii Ahrens, Fabrizi & Liu, new species strongly resembles the species of the Serica nigroguttata group. However, S. jirii differs from them by the more robust and more convex body, the deeply and widely emarginate labrum, the lacking membraneous rim of apical elytral margin, the dark preapical elytral spot, as well as by the short parameres lacking the framing of left paramere by the ventral membrane of the right one.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF98FF82FF435A20FC2BFD8E.taxon	etymology	Etymology. The species is named (noun in singular genitive case) after one of its collectors Jiří Hájek (NMPC).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF98FF82FF435A20FC2BFD8E.taxon	description	Variation. Length: 8.0 – 8.6 mm, length of elytra: 6.4 – 6.5 mm, width: 4.6 – 4.8 mm. Female: club only little shorter than in male; eyes as large as in male; pygidium weakly convex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF96FF83FF435BF3FBF1FC32.taxon	description	Figures 9 E – H, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF96FF83FF435BF3FBF1FC32.taxon	materials_examined	Type material examined. Holotype: ♂ “ Myanmar (Burma) Provinz Kachin State, Pangwa env. (LF) (H = 2450 m) 28. IX. 2010 N 25 ° 37 ’ 42.3 ’’ E 098 ° 23 ’ 22.5 ’’ leg. Michael Langer, S. Naumann & S. Löffler / 1108 Asia Sericini spec. ” (ZFMK). Paratype: 1 ♂ “ Myanmar (Burma) Provinz Kachin State, Pangwa env. (LF) (H = 2450 m) 28. IX. 2010 N 25 ° 37 ’ 42.3 ’’ E 098 ° 23 ’ 22.5 ’’ leg. Michael Langer, S. Naumann & S. Löffler ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF96FF83FF435BF3FBF1FC32.taxon	description	Description of the holotype. Length: 7.9 mm, length of elytra: 6.1 mm, width: 4.9 mm. Body elongate eggshaped, yellowish brown, head, pronotum, and scutellum dark brown, dorsal surface except dull frons shiny, partly with greenish shine, elytra with dark brown spots, antenna yellow; dorsal surface almost glabrous except a dense short erect setae on anterior pronotal disc and head. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and moderately convergent anteriorly, anterior angles moderately rounded; anterior margin distinctly but not deeply emarginate medially; margins weakly reflexed; surface flat and shiny, finely and moderately densely punctate, with a few short erect setae; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye small and flat, 1.3 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with sparse, short erect setae. Eyes small, ratio diameter / interocular width: 0.5. Antenna with 10 antennomeres; club with three antennomeres, 1.1 times as long as remaining antennomeres combined and straight. Mentum strongly elevated, convex. Labrum moderately produced along middle, moderately emarginate medially. Pronotum wide and subtrapezoidal, widest at base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles distinctly produced and acute; posterior angles strongly rounded; anterior margin with a wide marginal line and weakly produced medially; surface transversely impressed behind anterior margin and at middle resulting in two indistinct transversal sulci on each side of the pronotum, one anterior one posterior; surface irregularly densely and finely punctate, some areas on sulci lacking punctures, on anterior pronotal disc and beside anterior margin with dense short yellow setae partly erect partly bent posteriorly, minute setae in punctures absent; lateral and anterior margins sparsely setose; hypomeron indistinctly carinate at base. Scutellum large, triangular, finely and irregularly sparsely punctate, glabrous. Elytra oval, widest in posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures concentrated along striae fusing partly to transverse wrinkles; dark spots impunctate; intervals with a few single, fine, short setae, otherwise glabrous; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin very narrowly membraneous, with an extremely fine rime of minute microtrichomes (magnification 100 x). Ventral surface shiny, finely and densely punctate, almost glabrous, with a few single setae on metasternal disc; metacoxa glabrous, except a few short lateral setae; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous and very superficially sparsely punctate, ultimate sternite densely shortly setose; penultimate sternite at posterior margin with membraneous rim a quarter as long as sternite. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.28. Pygidium moderately convex and shiny, yellow with small dark spots, finely and moderately densely punctate, smooth midline narrow, with sparse long setae on apical third. Legs slender; femora dull, with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin apically finely serrated and only weakly widened, dorsal posterior margin completely but finely serrated, in basal half with a few long setae which are one third as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.2, dorsally sharply carinate, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, without longitudinal wrinkles; ventral margin serrated, with two extremely widely separated robust setae; medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally carinate, with a strongly serrated ridge ventrally; first metatarsomere little longer than following tarsomere and as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws symmetrical, basal tooth of inner claw bluntly truncate as the external one. Aedeagus: Fig. 9 E – G. Habitus: Fig. 9 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF96FF83FF435BF3FBF1FC32.taxon	diagnosis	Diagnosis. Serica pangwa Ahrens, Fabrizi & Liu, new species differs from S. jirii Ahrens, Fabrizi & Liu, new species by the densly setose pronotal disc, the more transverse pronotum, the more oblong and less convex body, and last not least by the shape of aedeagus. The parameres, particularly the left one, are significantly longer in S. pangwa than in S. jirii.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF96FF83FF435BF3FBF1FC32.taxon	etymology	Etymology. The species is named (noun in apposition) after its type locality Pangwa.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF96FF83FF435BF3FBF1FC32.taxon	description	Variation. Length: 7.9 – 8.0 mm, length of elytra: 6.0 – 6.1 mm, width: 4.5 – 4.9 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF97FF81FF435AB4FBE7FD8E.taxon	description	Figures 10 A – D, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF97FF81FF435AB4FBE7FD8E.taxon	materials_examined	Type material examined. Holotype: ♂ “ China: Zhejiang Prov., W Tiammu Shan (Mts.) reserve Immortal Peak; mountainous low forest, esp. On flowering Castanea sequini, 5. - 6. vii. 2017, 1500 m, 30 ° 20 ’ 58.5 ’’ N, 119 ° 25 ’ 26.5 ’’ E J. Hájek & J. Růžička leg. / 1059 Asia Sericini spec. ” (NMPC). Paratypes: 1 ♂ “ 23. III. 35 O. Piel / T’iemmu Shan Musée Heude / Serica / Pachyserica sp 22 ” (IZAS), 1 ♂ “ Mt. Sha hu shan, Hangzhou, Zhejiang Province 1957. VI. 30 / Pachyserica sp 20 ” (IZAS), 1 ♂ “ Mt. Bao shi, Hangzhou, Zhejiang 1957. VI. 4 ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF97FF81FF435AB4FBE7FD8E.taxon	description	Description of the holotype. Length: 8.8 mm, length of elytra: 6.5 mm, width: 4.9 mm. Body elongate, dark brown, dull, partly with greenish toment, elytra in punctures reddish brown, antenna and legs reddish brown; dorsal surface almost glabrous except a single and short, white adpressed setae on pronotum and elytra. Labroclypeus subrectangular, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles moderately convex; anterior margin weakly emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, partly punctures fusing to transverse wrinkles, with a few erect setae; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye small and flat, 1.5 times as wide as long; ocular canthus very short and slender (1 / 6 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes large, ratio diameter / interocular width: 0.79. Antenna with 9 antennomeres; club with three antennomeres, 1.3 times as long as remaining antennomeres combined and weakly reflexed. Mentum strongly elevated anteriorly, convex. Labrum weakly produced along middle, moderately emarginate medially. Pronotum transverse and subrectangular, widest in posterior third, lateral margins evenly weakly convex and convergent anteriorly as well posteriorly; anterior angles weakly produced and blunt, moderately rounded in the tip; posterior angles moderately rounded; anterior margin with robust marginal line and strongly produced medially; surface densely and finely punctate, along midline narrowly impunctate, with short or minute, pale, adpressed setae in punctures; lateral margins sparsely setose; hypomeron not carinate at base. Scutellum large, triangular, slightly impressed on each side, finely and densely punctate, on basal midline smooth, with minute setae in punctures. Elytra oval, widest in posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, moderately dense punctures concentrated along striae; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin chitinous, without a rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, metasternal disc densely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.25. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise finely and shortly setose and finely densely punctate; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Pygidium strongly convex and dull, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with sparse short, mostly adpressed setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 3.6, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with robust longitudinal wrinkles; ventral margin serrated, with three robust setae of which the two distal ones are widely separated; medial face glabrous and impunctate, but with robust longitudinal wrinkles; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and concavely truncate at apex. Aedeagus: Fig. 12 A – C. Habitus: Fig. 12 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF97FF81FF435AB4FBE7FD8E.taxon	diagnosis	Diagnosis. Serica jani Ahrens, Fabrizi & Liu, new species strongly resembles the species of the subgenus Serica sharing the lacking carina of basal hypomeron and the asymmetric protarsal claws. Both characters are known to have been developed convergently several times within some groups, so more evidence is needed for the proper assignment of this species. Serica jani differs from the species of the subgenus Serica by the short, robust, and strongly arched phallobase (lateral view) which in this form is unknown in the subgenus Serica. The species of the latter do have a dorsoventrally flattened phallobase.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF97FF81FF435AB4FBE7FD8E.taxon	etymology	Etymology. The species is named (noun in singular genitive case) after one of its collectors Jan Růžička (Prague).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF97FF81FF435AB4FBE7FD8E.taxon	description	Variation. Length: 8.8 – 10.0 mm, length of elytra: 6.5 – 7.6 mm, width: 4.9 – 5.9 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF95FF86FF435BF3FABDFE62.taxon	description	Figures 10 E – H, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF95FF86FF435BF3FABDFE62.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, SW Zhejiang, 12. V. - 5. VI. Fangyangshan, Huangmao Jian 27 ° 53 ’ N 119 ° 11 ’ E, 1500 - 1850 m Jaroslav Turna leg., 2008 ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF95FF86FF435BF3FABDFE62.taxon	description	Description of the holotype. Length: 7.5 mm, length of elytra: 6.0 mm, width: 4.1 mm. Body elongate, dark brown, dull, partly with greenish toment, legs reddish brown, elytra yellow brown with dark spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except a single and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles weakly convex; anterior margin widely shallowly emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, partly punctures fusing to transverse wrinkles, with numerous erect setae; frontoclypeal suture finely incised, weakly curved and slightly elevated; smooth area anterior to eye small and convex, as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes moderately large, ratio diameter / interocular width: 0.61. Antenna with 9 antennomeres; club with three antennomeres, 1.3 times as long as remaining antennomeres combined and weakly reflexed. Mentum strongly elevated anteriorly, convex. Labrum strongly produced along middle, narrowly but deeply emarginate medially. Pronotum narrow and subtrapezoidal, widest at base, lateral margins evenly and weakly convex and convergent anteriorly; anterior angles weakly produced and sharply pointed; posterior angles blunt; anterior margin straight, with robust marginal line; surface densely and finely punctate, along midline narrowly impunctate in basal half, with short or minute, pale, adpressed setae in punctures; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, slightly impressed on each side, finely and densely punctate, with minute setae in punctures. Elytra oval, widest at middle, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, densely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.38. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise finely densely punctate and finely shortly setose at middle of sternites, laterally setae less dense; penultimate sternite at posterior margin with a wide membraneous rim being one quarter as long as sternite. Pygidium strongly convex and dull, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with sparse, moderately long setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Tarsi dorsally impunctate, ventrally with sparse short setae. Hind legs missing in holotype. Protibia moderately long, bidentate, external margin only weakly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and concavely truncate at apex. Aedeagus: Fig. 10 E – G. Habitus: Fig. 10 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF95FF86FF435BF3FABDFE62.taxon	diagnosis	Diagnosis. Serica jaroslavi Ahrens, Fabrizi & Liu, new species resembles somewhat the species of the Serica nigroguttata group, however, the dark preapical elytral spot is missing in this new species and the parameres are strongly different in length as often is the case in the subgenus Serica. From Serica albisetis Ahrens, 2005, which also possesses a basal lobe at the right paramere, Serica jaroslavi Ahrens, Fabrizi & Liu, new species differs also by the extremely short left paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF95FF86FF435BF3FABDFE62.taxon	etymology	Etymology. The species is named (noun in singular genitive case) after its collector, Jaroslav Turna.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF92FF87FF435B5FFA87FDAA.taxon	description	Figures 11 A – D, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF92FF87FF435B5FFA87FDAA.taxon	materials_examined	Type material examined. Holotype: ♂ “ Mts. Shengtangshan, Jinxiu, Guangxi, 18. V. 1999, 1970 m, leg. Gao Mingyuan / LW- 157 / 1145 Asia Sericini spec. ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF92FF87FF435B5FFA87FDAA.taxon	description	Description of the holotype. Length: 8.4 mm, length of elytra: 6.1 mm, width: 4.2 mm. Body elongate, dark brown, dull, partly with greenish toment, legs reddish brown, elytra yellow brown with dark spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except a single and short, white adpressed setae on pronotum and elytra. Labroclypeus subrectangular, little wider than long, widest at base, lateral margins straight and subparallel, anterior angles moderately convex; anterior margin widely and distinctly emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture finely incised, weakly curved and slightly elevated; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with a few fine and long setae beside eyes and numerous short erect ones on disc. Eyes moderately large, ratio diameter / interocular width: 0.54. Antenna with 9 antennomeres; club with three antennomeres, 1.7 times as long as remaining antennomeres combined and moderately reflexed. Mentum strongly elevated anteriorly, convex. Labrum moderately produced along middle, distinctly emarginate medially. Pronotum wide but subtrapezoidal, widest at base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles moderately produced and sharply pointed; posterior angles strongly rounded; anterior margin straight, with robust but indistinct marginal line; surface densely and finely punctate, along midline narrowly impunctate in basal half, with minute, pale, adpressed setae in punctures, on sides with a few short setae; anterior and lateral margins sparsely setose; hypomeron carinate at base, not produced ventrally. Scutellum moderately large, triangular, finely and densely punctate, on midline impunctate, with minute setae in punctures. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, densely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.29. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous except on the ultimate sternite being finely shortly setose; penultimate sternite at posterior margin with a wide membraneous rim being one quarter as long as sternite. Pygidium moderately convex and dull, dark, finely and densely punctate, smooth midline narrow, with sparse, moderately long setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.7, dorsally sharply carinate, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with a few superficial longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face glabrous and impunctate; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and concavely truncate at apex. Aedeagus: Fig. 11 A – C. Habitus: Fig. 11 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF92FF87FF435B5FFA87FDAA.taxon	diagnosis	Diagnosis. Serica shengtangshan Ahrens, Fabrizi & Liu, new species differs from the similar S. albisetis Ahrens, 2005 by the left paramere having its dorsomedial tooth at basal third (not shortly before the apex as in S. albisetis).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF92FF87FF435B5FFA87FDAA.taxon	etymology	Etymology. The species is named (noun in apposition) after its type locality, Mts. Shengtangshan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF93FF84FF435B17FB7EFDAA.taxon	description	Figures 11 E – I, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF93FF84FF435B17FB7EFDAA.taxon	materials_examined	Type material examined. Holotype: ♂ “ Zhenba, Shaanxi, 19. VII. 1985, 1200 m, leg. Li Fasheng / 1144 Asia Sericini spec. ” (CAU).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF93FF84FF435B17FB7EFDAA.taxon	description	Description of the holotype. Length: 9.0 mm, length of elytra: 6.5 mm, width: 5.0 mm. Body oval, dark brown, dull, partly with greenish toment, legs reddish brown, elytra yellowish brown with dark spots, in punctures lighter, antenna yellow; dorsal surface with numerous and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles strongly convex; anterior margin distinctly emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, with numerous erect setae; frontoclypeal suture finely incised, weakly curved and slightly elevated; smooth area anterior to eye small and convex, as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes moderately large, ratio diameter / interocular width: 0.55. Antenna with 9 antennomeres; club with three antennomeres, 1.5 times as long as remaining antennomeres combined and weakly reflexed. Mentum strongly elevated anteriorly, convex. Labrum moderately produced medially, deeply emarginate medially. Pronotum wide and subtrapezoidal, widest at middle, lateral margins evenly and strongly convex and convergent anteriorly and posteriorly; anterior angles weakly produced and almost rectangular; posterior angles blunt; anterior margin straight, with robust marginal line; surface irregularly densely and finely punctate, along midline narrowly impunctate; with short, pale, adpressed setae, partly with minute setae in punctures; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, finely and densely punctate, with minute setae in punctures, impunctate on median base. Elytra oval, widest at middle, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, densely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.35. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise finely densely punctate and finely shortly setose at middle of sternites, laterally setae less dense; penultimate sternite at posterior margin with a membraneous rim being one quarter as long as sternite. Pygidium moderately convex and dull, dark, finely and densely punctate, smooth midline narrow, with sparse, moderately long setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur dull, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with robust longitudinal wrinkles; ventral margin serrated, with three robust setae of which the two distal ones are widely separated; medial face glabrous and impunctate, but with robust longitudinal wrinkles; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres lacking in holotype. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and concavely truncate at apex. Aedeagus: Fig. 11 E – H. Habitus: Fig. 11 I. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF93FF84FF435B17FB7EFDAA.taxon	diagnosis	Diagnosis. Serica zhenba Ahrens, Fabrizi & Liu, new species differs from the similar S. nanjiangana Ahrens, 2005 by the longer left paramere being more than half as long as the right paramere, and by the narrower basal lobe of the right paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF93FF84FF435B17FB7EFDAA.taxon	etymology	Etymology. The species is named (noun in apposition) after its type locality, Zhenba, Shaanxi.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF91FF9AFF4359EBFBE7FED6.taxon	description	Figures 12 A – D, 21	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF91FF9AFF4359EBFBE7FED6.taxon	materials_examined	Type material examined. Holotype: ♂ “ Jinzhong Road, Jinxiu, Guangxi, 10. V. 1999, 1100 m, leg. Yang Xingke / 1140 Asia Sericini spec. ” (IZAS). Paratype: 1 ♂ “ Datianding, Xinyi, Guangdong, 31. V. 1984 / 1704 m, light trap, leg. Liao Subai / LW- 100 ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF91FF9AFF4359EBFBE7FED6.taxon	description	Description of the holotype. Length: 9.5 mm, length of elytra: 7.4 mm, width: 5.1 mm. Body elongate, dark brown, dull, partly with greenish toment, legs reddish brown, elytra yellow brown with dark spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except a single and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins convex and weakly convergent, anterior angles moderately convex; anterior margin distinctly emarginate medially; margins moderately reflexed; surface flat and shiny, finely and densely punctate, with transverse wrinkles and a few fine erect setae anteriorly; frontoclypeal suture finely incised, almost straight and slightly elevated; smooth area anterior to eye small and convex, 1.2 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, without terminal seta. Frons completely dull and flat, with fine and dense punctures, with moderately dense, fine and short setae being bent posteriorly on disc and beside eyes beside eyes. Eyes small, ratio diameter / interocular width: 0.56. Antenna with 9 antennomeres; club with three antennomeres, 1.2 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, convex. Labrum moderately produced along middle, distinctly emarginate medially. Pronotum wide but subtrapezoidal, widest shortly before base, lateral margins in anterior half weakly convex, almost straight and strongly convergent anteriorly, in basal half strongly convex; anterior angles moderately produced and sharp; posterior angles strongly rounded; anterior margin convexly produced medially, with robust, distinct marginal line; surface densely and finely punctate, with numerous short, white, scale-like, adpressed setae, partly only with minute setae; anterior and lateral margins almost glabrous; hypomeron not carinate at base. Scutellum moderately large, triangular, finely and densely punctate, median base impunctate, pilosity as in pronotum. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, moderately dense punctures; with a few single, scale-like, short, white, adpressed setae on intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, densely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.29. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous except on the ultimate sternite being finely shortly setose; penultimate sternite at posterior margin with a wide membraneous rim being one quarter as long as sternite. Pygidium strongly convex but short, dull, dark, finely and densely punctate, smooth midline basally wide, with sparse, white, scale-like setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.2, dorsally sharply carinate, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with robust longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face glabrous, impunctate but wrinkled; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, with a few fine punctures dorsally. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and almost twice as long as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; protarsal claws lacking in holo- and paratype. Aedeagus: Fig. 12 A – C. Habitus: Fig. 12 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF91FF9AFF4359EBFBE7FED6.taxon	diagnosis	Diagnosis. Serica dissimillima Ahrens, Fabrizi & Liu, new species differs from all other Serica species by the particular shape of aedeagus, having the parameres strongly asymmetrically inserted and a dorsomedial apical apophysis in phallobase covering the base of the parameres (dorsal view). In general appearance it resemlbles strongly to the other Chinese Taiwanoserica species, however, the parameres are simple and not subdivided in dorsal and ventral lobe; also, the hypomeron of prothorax in not carinate, a character which is typical for Serica sensu stricto.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF91FF9AFF4359EBFBE7FED6.taxon	etymology	Etymology. The name of the new species is derrived from superlative of ‘ dissimile ’ (Latin), dissimilar, with reference to the dissimilar aedeagus morphology.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF91FF9AFF4359EBFBE7FED6.taxon	description	Variation. Length: 9.5 – 10.9 mm, length of elytra: 7.4 – 8.4 mm, width: 5.1 – 6.1 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8EFF98FF435FC6FA08FDD6.taxon	description	Figures 12 E – H, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8EFF98FF435FC6FA08FDD6.taxon	materials_examined	Type material examined. Holotype: ♂ “ Mts. Maoershan, Guangxi, 15. VII. 1985, 1100 m, leg. Wang Ziqing / LW- 140 / 1147 Asia Sericini spec. ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8EFF98FF435FC6FA08FDD6.taxon	description	Description of the holotype. Length: 8.1 mm, length of elytra: 6.2 mm, width: 4.8 mm. Body elongate, dark brown, dull, partly with greenish shine or toment, labroclypeus, anterior frons and anterior pronotum shiny; ventral surface and legs reddish brown; elytra with yellow spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except a single and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly convergent, anterior angles blunt; anterior margin widely and distinctly emarginate medially; margins moderately reflexed; surface flat, finely and densely punctate, with numerous erect setae; frontoclypeal suture finely incised, weakly curved and slightly elevated; smooth area anterior to eye small and convex, as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons dull only in posterior half, flat, with fine and dense punctures, with fine and long setae beside eyes and on disc. Eyes small, ratio diameter / interocular width: 0.55. Antenna with 10 antennomeres; club with three antennomeres, 0.8 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, convex. Labrum moderately produced along middle, distinctly emarginate medially. Pronotum wide but subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly, and slightly also posteriorly; anterior angles moderately produced and sharp; posterior angles blunt; anterior margin medially strongly convex, with indistinct marginal line; surface finely and densely but irregularly punctate, along midline narrowly impunctate in basal half, partly punctures fusing with each other rugosely, with a shallow impression on each side on disc; surface with minute or short, pale, adpressed setae in punctures; anterior and lateral margins sparsely setose; along base narrowly with dull toment; hypomeron carinate at base, weakly produced ventrally. Scutellum moderately large, triangular, finely and very densely punctate, on basal midline impunctate, with minute setae in punctures. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, erect setae on each interval; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, almost glabrous; metacoxa with a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.45. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being one quarter as long as sternite. Pygidium moderately convex and dull, dark with numerous yellow spots, finely and densely punctate, smooth midline narrow, with sparse, moderately long setae on apical half. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Posterior legs missing in holotype. Protibia moderately long, bidentate, external margin smooth, weakly extended at middle; anterior claws symmetrical, basal tooth of inner claw sharply pointed at apex. Aedeagus: Fig. 12 E – G. Habitus: Fig. 12 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8EFF98FF435FC6FA08FDD6.taxon	diagnosis	Diagnosis. Serica ziqingi Ahrens, Fabrizi & Liu, new species differs from the other Serica sensu lato species by the short antennal club in male, the particular mrohology of aedeagus, with the parameres bein almost as long as the phallobase and being both curved externally. It resembles somewhat the species of the Serica nigroguttata group or Taiwanoserica, but from both it differs by the shiny head and pronotum.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8EFF98FF435FC6FA08FDD6.taxon	etymology	Etymology. The species is named (noun in nominative genitive case) after its collector, Wang Ziqing.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8CFF9EFF435BC4FA05FE1F.taxon	description	Figures 13 A – D, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8CFF9EFF435BC4FA05FE1F.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, W Yunnan prov., mts. 60 km E Tengchong, 2200 m, 19. - 22. v. 2006, S. Murzin & I. Shokhin / 853 Sericini: Asia spec. ” (CP). Paratypes: 3 ♂♂, 1 ♀ “ China, W Yunnan prov., mts. 60 km E Tengchong, 2200 m, 19. - 22. v. 2006, S. Murzin & I. Shokhin ” (CP, ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8CFF9EFF435BC4FA05FE1F.taxon	description	Description of the holotype. Length: 9.4 mm, length of elytra: 7.2 mm, width: 5.4 mm. Body elongate eggshaped, dark brown, partly with greenish toment or shine, elytra with dark spots, antenna yellowish, legs brown; dorsal surface dull and densely covered with short yellow setae being bent posteriorly. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins almost straight and convergent anteriorly, anterior angles almost acute but rounded in tip, anteriorly deeply but not widely emarginate medially; margins moderately reflexed; surface flat and shiny, finely and densely punctate, finely densely setose, behind anterior margin glabrous; frontoclypeal suture finely incised, weakly convex; smooth area anterior to eye large and flat, approximately twice as wide as long; ocular canthus moderately long and slender (1 / 3 of ocular diameter), with a few fine punctures bearing each a short seta. Frons completely dull and flat, with fine and very dense punctures, evenly covered with short yellow setae being bent backwards and longer erect setae. Eyes small, ratio diameter / interocular width: 0.49. Antenna with ten antennomeres; antennomeres three to five slightly wider than long, antennomere six and seven transverse and short; club with three antennomeres, 1.2 times as long as remaining antennomeres combined and straight. Mentum weakly elevated, anteriorly flattened. Labrum transverse, short, strongly produced along the middle, moderately emarginate medially. Pronotum subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles distinctly produced and acute; posterior angles strongly rounded; anterior margin medially with a fine marginal line and very weakly produced medially; surface on disc densely and coarsely punctate, almost glabrous, punctures on sides less dense and with dense short yellow setae; lateral margins sparsely setose; hypomeron carinate at base, but not produced ventrally. Scutellum large, triangular, finely and very densely punctate, pilosity as in lateral pronotum. Elytra oval, widest at middle, striae indistinctly impressed, finely and densely punctate; intervals flat, with fine, very dense punctures; dark spots impunctate or with less dense punctures, with fine, short pilosity as in pronotum, without longer, erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical margin with a very fine membranous rim composed of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, including metacoxa and abdominal sternites densely shortly setose; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise shortly setose as rest of ventral surface; ultimate sternite at middle 1.5 times as long as penultimate one. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.44. Pygidium strongly convex and dull, finely and densely punctate, smooth midline narrow, with dense short setae, without long, erect setae. Legs slender; femora with two longitudinal rows of setae, coarsely and densely punctate between the rows, with robust setae on basal half; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge, posterior margin ventrally serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as width of metafemur. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.9, dorsally sharply carinate, with two groups of spines, basal group shortly before the middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face impunctate; ventral margin serrated, with two very widely separated robust setae; medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur; mesotarsomeres not carinate. Protibia moderately long, bidentate, external margin serrate, with a few small teeth at base, anterior claws symmetrical, basal tooth of inner claw bluntly truncate as the external one. Aedeagus: Fig. 13 A – C. Habitus: Fig. 7 D.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8CFF9EFF435BC4FA05FE1F.taxon	diagnosis	Diagnosis. Serica tengchongana Ahrens, Fabrizi & Liu, new species differs from S. deuvei Ahrens, 2005 and S. hirtella Ahrens, 2005 by the serrated external margin of protibia, as well as by the different shape of parameres: the right paramere is strongly bent at middle (lateral view) being subdivided into a dorsal lobe and a ventral distal one.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8CFF9EFF435BC4FA05FE1F.taxon	etymology	Etymology. The species is named (adjective in nominative singular case) according to its vicinity to Tengchong.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8CFF9EFF435BC4FA05FE1F.taxon	description	Variation. Length: 9.4 – 12.11 mm, length of elytra: 7.2 – 9.5 mm, width: 5.4 mm. Female: antennal club short, little shorter than remaining antennomeres combined; pygidium at apex strongly convex; eyes as large as in male.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8AFF9FFF435882FC39FDD3.taxon	description	Figures 13 E – H, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8AFF9FFF435882FC39FDD3.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, W Yunnan prov., mts. 60 km E Tengchong, 2200 m, 19. - 22. v. 2006, S. Murzin & I. Shokhin / 854 Sericini: Asia spec. ” (CP).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8AFF9FFF435882FC39FDD3.taxon	description	Description of the holotype. Length: 9.2 mm, length of elytra: 7.0 mm, width: 5.5 mm. Body elongate egg-shaped, dark brown, partly with greenish toment or shine, elytra with dark spots, antenna yellowish, legs brown; dorsal surface dull and densely covered with short yellow setae being bent posteriorly. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles acute; anterior margin deeply and widely emarginate medially; margins weakly reflexed; surface flat and shiny, finely and densely punctate, finely densely setose, behind anterior margin glabrous; frontoclypeal suture finely incised, weakly convex; smooth area anterior to eye large and flat, as wide as long; ocular canthus moderately long and slender (1 / 3 of ocular diameter), with a few fine punctures bearing each a short seta. Frons completely dull and flat, with fine and very dense punctures, evenly covered with short yellow setae being bent backwards and longer erect setae. Eyes small, ratio diameter / interocular width: 0.56. Antenna with ten antennomeres; antennomeres three to five slightly wider than long, antennomere six and seven transverse and short; club with three antennomeres, 1.2 times as long as remaining antennomeres combined and straight. Mentum weakly elevated, anteriorly flattened. Labrum transverse, short, strongly produced along the middle, moderately emarginate medially. Pronotum subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles distinctly produced and acute; posterior angles strongly rounded; anterior margin medially with a fine marginal line and very weakly produced medially; surface densely and coarsely punctate, with dense short yellow setae bent posteriorly; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, finely and very densely punctate, basal midline smooth, pilosity as in lateral pronotum. Elytra oval, widest at middle, striae indistinctly impressed, finely and densely punctate; intervals flat, with fine, very dense punctures; dark spots impunctate or with less dense punctures, with fine, short pilosity as in pronotum, without longer, erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical margin with a very fine membranous rim composed of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, including metacoxa and abdominal sternites densely shortly setose; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise shortly setose as rest of ventral surface; ultimate sternite at middle 1.5 times as long as penultimate one. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.49. Pygidium strongly convex and dull, finely and densely punctate, smooth midline narrow, with dense short setae, without long, erect setae. Legs slender; femora with two longitudinal rows of setae, coarsely and densely punctate between the rows, with robust setae on basal half; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge, posterior margin ventrally serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as width of metafemur. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.4, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face impunctate, on dorsal portion with a few superficial wrinkles; ventral margin serrated, with two very widely separated robust setae; medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur; mesotarsomeres not carinate. Protibia moderately long, bidentate, external margin serrate, with a few small teeth at base, anterior claws symmetrical, basal tooth of inner claw bluntly truncate as the external one. Aedeagus: Fig. 13 E – G. Habitus: Fig. 13 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8AFF9FFF435882FC39FDD3.taxon	diagnosis	Diagnosis. Serica alltengchongensis Ahrens, Fabrizi & Liu, new species differs from S. tengchongensis Ahrens, Fabrizi & Liu, new species by the shape of parameres: the right paramere has a separately developed, large basal lobe, the left paramere is slightly longer than that of S. tengchongensis and it is blade-like widened in apical half.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8AFF9FFF435882FC39FDD3.taxon	etymology	Etymology. The name of the species is derived from the combined Greek prefix ‘ allo - ’ (other) and the species name ‘ tengchongensis ’, with reference to the similarity to this species.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8BFF9DFF435BCFFF73FEAF.taxon	description	Figures 14 A – D, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8BFF9DFF435BCFFF73FEAF.taxon	materials_examined	Type material examined. Holotype: ♂ “ Myanmar (Burma) Provinz Kachin State; Mt. Emaw Bum nach Kamphant; Waldcamp-Holzmeiler 26 ° 09 ‘ 232 ‘‘ N, 98 ° 31 ‘ 164 ‘‘ E; 2358 m, 28.5.2006; leg. Michael Langer, Stefan Naumann, Swen Löffler; Coll. M. Langer / 1130 Asia Sericini spec. ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8BFF9DFF435BCFFF73FEAF.taxon	description	Description of the holotype. Length: 7.8 mm, length of elytra: 6.1 mm, width: 4.9 mm. Body elongate eggshaped, dark reddish brown, partly with greenish toment, elytra with dark spots, antenna yellowish, legs brown; dorsal surface dull and densely covered with short yellow setae being bent posteriorly. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles acute; anterior margin deeply and widely emarginate medially; margins weakly reflexed; surface flat and shiny, finely and densely punctate, finely densely setose, behind anterior margin glabrous; frontoclypeal suture finely incised, weakly convex; smooth area anterior to eye large and flat, as wide as long; ocular canthus moderately long and slender (1 / 3 of ocular diameter), with a few fine punctures bearing each a short seta. Frons completely dull and flat, with fine and very dense punctures, evenly covered with short yellow setae being bent backwards and longer erect setae. Eyes small, ratio diameter / interocular width: 0.47. Antenna with ten antennomeres; antennomeres three to five slightly wider than long, antennomere six and seven transverse and short; club with three antennomeres, 1.2 times as long as remaining antennomeres combined and straight. Mentum weakly elevated, anteriorly flattened. Labrum transverse, short, strongly produced along the middle, moderately emarginate medially. Pronotum subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles distinctly produced and acute; posterior angles strongly rounded; anterior margin medially with a fine marginal line and very weakly produced medially; surface densely and coarsely punctate, with dense short yellow setae bent posteriorly; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, finely and very densely punctate, basal midline smooth, pilosity as in lateral pronotum. Elytra oval, widest at middle, striae indistinctly impressed, finely and densely punctate; intervals flat, with fine, very dense punctures; dark spots impunctate or with less dense punctures, with fine, short pilosity as in pronotum, without longer, erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical margin with a very fine membranous rim composed of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, including metacoxa and abdominal sternites densely shortly setose; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise shortly setose as rest of ventral surface; ultimate sternite at middle 1.5 times as long as penultimate one. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.54. Pygidium strongly convex and dull, finely and densely punctate, smooth midline narrow, with dense short setae, without long, erect setae. Legs slender; femora with two longitudinal rows of setae, coarsely and densely punctate between the rows, with robust setae on basal half; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge, posterior margin ventrally serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as width of metafemur. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.5, dorsally sharply carinate, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face impunctate, on dorsal portion with a few superficial wrinkles; ventral margin serrated, with two very widely separated robust setae; medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur; mesotarsomeres not carinate. Protibia moderately long, bidentate, external margin serrate, with a few small teeth at base, anterior claws symmetrical, basal tooth of inner claw bluntly truncate as the external one. Aedeagus: Fig. 14 A – C. Habitus: Fig. 14 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8BFF9DFF435BCFFF73FEAF.taxon	diagnosis	Diagnosis. Serica emaw Ahrens, Fabrizi & Liu, new species differs from S. langeri Ahrens & Fabrizi, 2011 by the shape of parameres: the right paramere is wider also in subapical portion and more concavely arched; the left paramere is at base much more narrow (lateral view).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF8BFF9DFF435BCFFF73FEAF.taxon	etymology	Etymology. The name of the species (noun in apposition) is derived from it type locality close to the Mt. Emaw.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF89FF93FF435812FA09FE8C.taxon	description	Figures 14 E – H, 23	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF89FF93FF435812FA09FE8C.taxon	materials_examined	Type material examined. Holotype: ♂ “ Guance Zhan, Wuyipeng, Wolongzhen, Sichuan, 24. VII. 2004, 2560 m, leg. Zhang Yong ” (IZAS). Paratypes: 1 ♂, 1 ♀ “ Ebian County, 25. VI. 1981, leg. Shan Shigui ” (IZAS), 1 ♂ “ Mts. Maoershan, Guangxi, 14. VII. 1985, 1900 m, leg. Liao Subai ” (IZAS), 1 ♂ “ Wolong Guangou, Wolong, Sichuan, 20. VII. 2004, 2150 m, light trap, leg. Zhang Yong ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF89FF93FF435812FA09FE8C.taxon	description	Description of the holotype. Length: 9.4 mm, length of elytra: 7.1 mm, width: 5.1 mm. Body elongate eggshaped, dark reddish brown, partly with greenish toment, elytra with dark spots, even intervals almost completely dark, antenna yellowish, legs brown; dorsal surface dull and densely covered with short yellow setae being bent posteriorly. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles acute; anterior margin deeply and widely emarginate medially; margins weakly reflexed; surface flat and shiny, finely and densely punctate, finely densely setose, behind anterior margin glabrous; frontoclypeal suture finely incised, weakly convex; smooth area anterior to eye large and flat, as wide as long; ocular canthus moderately long and slender (1 / 3 of ocular diameter), with a few fine punctures bearing each a short seta. Frons completely dull and flat, with fine and very dense punctures, evenly covered with short yellow setae being bent backwards and longer erect setae. Eyes small, ratio diameter / interocular width: 0.5. Antenna with ten antennomeres; antennomeres three to five slightly wider than long, antennomere six and seven transverse and short; club with three antennomeres, 1.1 times as long as remaining antennomeres combined and straight. Mentum weakly elevated, anteriorly flattened. Labrum transverse, short, strongly produced along the middle, moderately emarginate medially. Pronotum subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly; anterior angles distinctly produced and acute; posterior angles strongly rounded; anterior margin medially with a fine marginal line and very weakly produced medially; surface densely and coarsely punctate, with dense short yellow setae bent posteriorly; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, finely and very densely punctate, basal midline smooth, pilosity as in lateral pronotum. Elytra oval, widest at middle, striae indistinctly impressed, finely and densely punctate; intervals flat, with fine, very dense punctures; dark spots impunctate or with less dense punctures, with fine, short pilosity as in pronotum, without longer, erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical margin with a very fine membranous rim composed of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, including metacoxa and abdominal sternites densely shortly setose; abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise shortly setose as rest of ventral surface; ultimate sternite at middle 1.5 times as long as penultimate one. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.38. Pygidium moderately convex and dull, finely and densely punctate, smooth midline narrow, with dense short setae. Legs slender; femora with two longitudinal rows of setae, coarsely and densely punctate between the rows, with robust setae on basal half; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge, posterior margin ventrally serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as width of metafemur. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.6, dorsally sharply carinate, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face impunctate, with dense superficial longitudinal wrinkles; ventral margin serrated, with two very widely separated robust setae; medial face glabrous and smooth; apex interiorly near tarsal articulation concavely truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and almost twice as long as dorsal tibial spur; mesotarsomeres not carinate. Protibia moderately long, bidentate, external margin serrate, with a few small teeth at base, anterior claws symmetrical, basal tooth of inner claw bluntly truncate as the external one. Aedeagus: Fig. 14 E – G. Habitus: Fig. 14 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF89FF93FF435812FA09FE8C.taxon	diagnosis	Diagnosis. Serica ventrituberculata Ahrens, Fabrizi & Liu, new species differs from S. bicornis Ahrens, 2005 by the shape of parameres: the right paramere is almost straight (lateral view) and the left paramere has its lateral tooth almost at middle rather than before apex, the basal portion is almost twice as wide as the dorsal one (while it is almost subequal in width in S. bicornis); the phallobase is dorsally without tubercles, but instead it has ventrally before apex a pair of robust tubercles.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF89FF93FF435812FA09FE8C.taxon	etymology	Etymology. The name of the species (adjective in nominative case singular) is derived from the combined Latin words ‘ ventris ’ (venter) and ‘ tuberculatus ’ (i. e., having tubercles), with reference on the two ventral tubercles of phallobase.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF89FF93FF435812FA09FE8C.taxon	description	Variation. Length: 8.1 – 10.0 mm, length of elytra: 6.5 – 7.6 mm, width: 4.5 – 5.1 mm. Female: antennal club short, little shorter than remaining antennomeres combined; pygidium at apex strongly convex; eyes as large as in male.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF87FF90FF4358F0FBF1FE46.taxon	description	Figures 15 A – D, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF87FF90FF4358F0FBF1FE46.taxon	materials_examined	Type material examined. Holotype: ♂ “ Yuheba, Daozhen, Guizhou, 25. V. 2004, leg. Yu Yang / LW- 131 / 1143 Sericini Asia spec. ” (HBUM). Paratype: 1 ♂ “ Yuheba, Daozhen, Guizhou, 25. V. 2004, leg. Yu Yang ” (HBUM).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF87FF90FF4358F0FBF1FE46.taxon	description	Description of the holotype. Length: 8.2 mm, length of elytra: 6.2 mm, width: 5.2 mm. Body elongate, dark brown, dull, partly with greenish toment or shine, elytra with several dark impunctate spots and a large round dark preapical spot, antenna yellow; dorsal surface densely covered with short adpressed scale-like, white setae. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins in basal half straight, in apical half strongly convex and convergent towards strongly rounded anterior angles; anterior margin deeply emarginate medially; margins moderately reflexed; surface flat and shiny, finely and densely punctate, with numerous fine erect setae; frontoclypeal suture finely incised, weakly curved but indistinct under thick toment; smooth area anterior to eye small, as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, punctures with moderately dense, adpressed, white, scale-like setae and with long fine erect setae. Eyes small, ratio diameter / interocular width: 0.45. Antenna with 10 antennomeres; club with three antennomeres, 1.2 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, convex. Labrum moderately produced along middle, deeply emarginate medially. Pronotum wide but subtrapezoidal, widest at base, lateral margins straight and strongly convergent anteriorly; anterior angles moderately produced and strongly convex; posterior angles strongly rounded; anterior margin straight, without distinct marginal line; surface irregularly densely and finely punctate, partly punctures less dense, along basal midline broadly impunctate and glabrous, otherwise with dense, short, white, scale-like, adpressed setae, partly on disc only with minute setae; anterior and lateral margins sparsely setose; hypomeron carinate at base, distinctly produced ventrally. Scutellum large, triangular, finely and densely punctate, median base impunctate, pilosity as in pronotum. Elytra oval, widest at middle, striae finely impressed, finely and densely punctate; intervals flat, with fine, moderately dense punctures; pilosity as in pronotum, dark spots impunctate and glabrous; epipleural edge fine but at middle strongly widened and concavely reflexed, ending at moderately curved external apical angle of elytra, epipleura densely setose; with a blunt, transverse preapical carina before narrowly membraneous apical margin, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, including metacoxa densely setose, with long fine setae and dense white scale-like setae; metacoxa additionally with a few robust lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.3. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise pilosity as in the rest of the ventral surface; penultimate sternite at posterior margin with a wide membraneous rim being one quarter as long as sternite. Pygidium strongly convex, dull, dark, finely and densely punctate, with long setae along apical margin, otherwise pilosity as in abdominal sternites. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 3.67, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with a few short longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face glabrous, impunctate but wrinkled; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, with fine elongate punctures dorsally. Metatarsomeres ventrally glabrous, laterally carinate, with a strongly serrated ridge ventrally; first metatarsomere distinctly longer than following two tarsomeres combined and longer than twice as long as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth not extended laterally at middle; protarsal claws asymmetric, basal tooth of interior claw small lobiform, convex at apex. Aedeagus: Fig. 15 A – C. Habitus: Fig. 15 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF87FF90FF4358F0FBF1FE46.taxon	diagnosis	Diagnosis. Serica yuheba Ahrens, Fabrizi & Liu, new species differs from all other species of the Serica nigroguttata group by the strongly reflexed lateral margin of the elytra, as well as the strong mesal tooth at the middle of the left paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF87FF90FF4358F0FBF1FE46.taxon	etymology	Etymology. The new species is named (noun in apposition) after its type locality, Yuheba.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF87FF90FF4358F0FBF1FE46.taxon	description	Variation. Length: 8.2 – 8.3 mm, length of elytra: 6.2 – 6.3 mm, width: 5.2 – 5.3 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF85FF96FF4359EBFB67FED7.taxon	description	Figures 15 E – H, 24	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF85FF96FF4359EBFB67FED7.taxon	materials_examined	Type material examined. Holotype: ♂ “ China, W Yunnan prov. mts. 20 km SW Baoshan, 2400 m, 23. - 25. v. 2006, S. Murzin & Shokin leg. / 1150 Asia Sericini spec. ” (CP). Paratypes: 1 ♂, 1 ♀ “ China, W Yunnan prov. mts. 20 km SW Baoshan, 2400 m, 23. - 25. v. 2006, S. Murzin & Shokin leg. ” (CP, ZFMK), 1 ♂ “ S. China, Yunnan prov. 18 km SW Baoshan, 2100 m, 18. - 19. v. 2003 leg. S. Murzin ” (CP).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF85FF96FF4359EBFB67FED7.taxon	description	Description of the holotype. Length: 6.9 mm, length of elytra: 4.9 mm, width: 3.5 mm. Body elongate, brown, legs slightly lighter, surface dull, elytra with several slightly darker impunctate spots, and partly iridescent, antenna yellow; dorsal surface densely covered with short, pale setae. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins convex and convergent towards strongly rounded anterior angles; anterior margin distinctly emarginate medially; margins moderately reflexed; surface flat and shiny, finely and densely punctate, with numerous fine erect setae; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye small, twice as wide as long; ocular canthus short and slender (1 / 3 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, punctures with moderately dense, adpressed, white, setae and with a few pale long erect ones. Eyes small, ratio diameter / interocular width: 0.59. Antenna with 10 antennomeres; club with three antennomeres, 1.3 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, convex. Labrum moderately produced along middle, moderately emarginate medially. Pronotum wide but subtrapezoidal, widest at base, lateral margins in basal half straight and subparallel, in anterior half strongly convex and convergent anteriorly; anterior angles moderately produced and blunt; posterior angles blunt; anterior margin straight, with distinct marginal line; surface irregularly densely and finely punctate, along midline impunctate and glabrous, otherwise with dense, short, white, adpressed setae, on disc with numerous long, pale, erect setae; anterior and lateral margins sparsely setose; hypomeron carinate at base, not produced ventrally. Scutellum large, triangular, finely and very densely punctate, median base impunctate, with very dense, adpressed, short to minute setae in punctures. Elytra oval, widest at middle, striae finely impressed, finely and densely punctate; intervals flat, with fine, moderately dense punctures; punctures with pale, short adpressed setae, on odd intervals with a few singe robust white and erect setae; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, with fine, dense, adpressed, pale setae; metacoxa with minute setae in punctures, additionally with a few robust lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.5. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise pilosity as in the rest of the ventral surface; penultimate sternite at posterior margin with a wide membraneous rim being one quarter as long as sternite. Pygidium moderately convex, dull, dark, finely and densely punctate, with long pale setae margin, otherwise with fine white adpressed pilosity. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.3, dorsally sharply carinate, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with a few short longitudinal wrinkles on dorsal portion; ventral margin serrated, with two moderately separated robust setae; medial face glabrous, superficially finely punctate; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, with sparse fine punctures dorsally. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth not extended laterally at middle; protarsal claws asymmetric, basal tooth of interior claw widened and bluntly truncate at apex. Aedeagus: Fig. 15 E – G. Habitus: Fig. 15 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF85FF96FF4359EBFB67FED7.taxon	diagnosis	Diagnosis. Serica baoshan Ahrens, Fabrizi & Liu, new species differs from the similar Serica inaequalis Ahrens, 2007 by the dorsoventrally more flattened apical phallobase (lateral view), the more narrow left paramere, and the right paramere which is only before the apex bluntly extended to a lateral tooth (rather than at middle as in S. inaequalis).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF85FF96FF4359EBFB67FED7.taxon	etymology	Etymology. The new species is named (noun in apposition) after its type locality, Baoshan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF85FF96FF4359EBFB67FED7.taxon	description	Variation. Length: 6.1 – 6.9 mm, length of elytra: 4.2 – 4.9 mm, width: 3.4 – 3.5 mm. Female: Antennal club little shorter than remaining antennomeres combined, pygidium weakly convex, eyes as large as in male.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF96FF4358CAFA08F92B.taxon	discussion	Remarks. Taiwanoserica Nomura, 1974 was originally described as genus from Taiwan, from where a number of endemic species have so far been reported (Kobayashi 1983, 1988, 1993; Kobayashi & Yu 2000; Ahrens 2002, 2006, 2007 b; Ahrens & Bezděk 2016). Based on results of a morphology-based phylogeny (Ahrens 2007 c) it was ranked subsequently as a subgenus of Serica (Ahrens 2007 b). Taiwanoserica was later, based on a morphologybased phylogeny (Ahrens 2009), also reported from western China. In subsequent molecular phylogenies Taiwanoserica resulted to be sister group of the Serica (s. str.) (Liu et al. 2015 b; Eberle et al. 2017), thus its subgenus rank was confirmed. Herein, we describe six additional new species from western China, northern Vietnam and Taiwan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF94FF435FA3FBF1FA56.taxon	description	Figures 16 A – D, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF94FF435FA3FBF1FA56.taxon	materials_examined	Type material examined. Holotype: ♂ “ Guizhou Kuankuoshui Nature Reserve, Tea Farm, 2010 - VIII- 10, 1527 m, N: 28.22; E: 107.15 / LW- 1365 ” (ZFMK). Paratypes: 2 ♂♂ “ Pingbaying, Xianfeng, Hubei, 22. VII. 1999, leg. Zheng Leyi ” (NKU), 1 ♂ “ Mt. Fanjingshan, Guizhou, 27. VII. 2001, 1300 m, leg. Zhao Fang / LW- 956 ” (CAU), 1 ♂ “ Leigongshan Forestry Farm, Guizhou, 14. IX. 2005, leg. Wang Jiliang, Gaochao / LW- 087 ” (HBUM). # Species from Taiwan not included here.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF94FF435FA3FBF1FA56.taxon	description	Description of the holotype. Length: 8.6 mm, length of elytra: 6.5 mm, width: 5.0 mm. Body oval, dark brown, dull, partly with greenish toment, ventral surface and legs reddish brown, elytra yellow brown with numerous dark spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except some single and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, preapically abruptly angled; anterior angles blunt; anterior margin deeply emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, partly punctures fusing to transverse wrinkles, with numerous erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes small, ratio diameter / interocular width: 0.57. Antenna with 10 antennomeres; club with three antennomeres, as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum strongly produced along middle, narrowly but deeply emarginate medially. Pronotum narrow and subtrapezoidal, widest at base, lateral margins evenly and weakly convex and convergent anteriorly; anterior angles well produced and sharp, weakly rounded in the tip; posterior angles weakly rounded; anterior margin straight, with robust and complete marginal line; surface irregularly, partly densely and partly sparsely punctate, sparse punctures finer, along midline narrowly impunctate, with short or minute, white, adpressed setae; lateral margins sparsely setose; hypomeron carinate at base, moderately produced ventrally. Scutellum large, triangular, finely and densely punctate, with minute setae in punctures, on midline punctures less dense. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x) (left elytra of holotype broken at middle, apical portion missing). Ventral surface dull, finely and densely punctate, sparsely setose, widely glabrous; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.49. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Pygidium moderately convex and dull, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with sparse, moderately long setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 3.8, dorsally sharply carinate, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, with some superficial longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 16 A – C. Habitus: Fig. 16 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF94FF435FA3FBF1FA56.taxon	diagnosis	Diagnosis. Serica kuankuoshuiensis Ahrens, Fabrizi & Liu, new species resembles somewhat Serica silviae Ahrens, 2009, both species share the sclerotized and spinose endophallus which is visible between parameres. Both species differ significantly in the shape of parameres, in Serica kuankuoshuiensis the dorsal lobe of the left paramere is massive and the ventral lobe only filiform and small.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF94FF435FA3FBF1FA56.taxon	etymology	Etymology. The species is named (adjective in singular nominative case) after its type locality, Kuankuoshui Nature Reserve.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF82FF94FF435FA3FBF1FA56.taxon	description	Variation. Length: 8.4 – 8.6 mm, length of elytra: 6.5 – 6.6 mm, width: 5.0 – 5.8 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF80FF95FF435F4BFF40F8FE.taxon	description	Figure 16 E – H, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF80FF95FF435F4BFF40F8FE.taxon	materials_examined	Type material examined. Holotype: ♂ “ Coll. I. R. Sc. N. B. Vietnam: Pia Oac Mt. NR 22 ° 36 ’ N / 105 ° 53 ’ E 04 - 06 - VIII- 2010 / I. G. 31.668 Light Trap Leg. J. Coinstant & P. Limbourg / 1048 Sericini Asia spec. ” (ISNB). Paratypes: 7 ♂♂, 7 ♀♀ “ Coll. I. R. Sc. N. B. Vietnam: Pia Oac Mt. NR 22 ° 36 ’ N / 105 ° 53 ’ E 04 - 06 - VIII- 2010 / I. G. 31.668 Light Trap Leg. J. Coinstant & P. Limbourg ” (ISNB, ZFMK), 1 ♂ “ Coll. I. R. Sc. N. B. Vietnam: Cuc Phong 11 - 18 - VIII- 2010 I. G. 31.668 Leg. J. Coinstant & P. Limbourg ” (ISNB).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF80FF95FF435F4BFF40F8FE.taxon	description	Description of the holotype. Length: 8.9 mm, length of elytra: 6.5 mm, width: 5.2 mm. Body oval, dark brown, iridescent and moderately shiny, partly with greenish toment or shine, ventral surface and legs reddish brown, elytra yellow brown with numerous dark spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except some short, pale setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins strongly convex and convergent anteriorly; anterior angles moderately rounded; anterior margin deeply emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, partly punctures fusing to transverse wrinkles, with numerous erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes very small, ratio diameter / interocular width: 0.48. Antenna with 10 antennomeres; club with three antennomeres, 1.1 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum strongly produced along middle, narrowly but deeply emarginate medially. Pronotum narrow and subtrapezoidal, widest shortly before base, lateral margins evenly and weakly convex and convergent anteriorly; anterior angles distinctly produced and sharp; posterior angles strongly rounded; anterior margin straight, without marginal line; surface finely densely punctate, without smooth midline, with short or minute, fine, erect, pale setae; lateral margins sparsely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum large, triangular, finely and densely punctate, with minute setae in punctures, on midline narrowly impunctate. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, erect setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose (but setae in holotype abraded); apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, on metasternal disc densely setose, otherwise sparsely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.48. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being a quarter as long as sternite. Pygidium strongly convex and moderately shiny, dark, in punctures yellow, finely and densely punctate, smooth midline wide, with moderately dense, long and short setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.5, dorsally sharply carinate, with two groups of spines, basal group shortly behind the basal third, apical group at two thirds of metatibial length, basally with a few single, fine setae; external face sparsely and finely punctate, without wrinkles, glabrous; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally sparsely finely punctate. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 16 E – G. Habitus: Fig. 16 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF80FF95FF435F4BFF40F8FE.taxon	diagnosis	Diagnosis. Serica limbourgi Ahrens, Fabrizi & Liu, new species is in shape of aedeagus most similar to Serica sigipinensis Ahrens, 2009. Serica limbourgi differs by the bluntly widened ventral phallobase, and the strongly arched dorsal lobe of both parameres.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF80FF95FF435F4BFF40F8FE.taxon	etymology	Etymology. The new species is named (substantive in singular genitive case) after one of its collectors, Poul Limbourg.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFF80FF95FF435F4BFF40F8FE.taxon	description	Variation. Length: 8.9 – 10.4 mm, length of elytra: 6.5 – 8.1 mm, width: 5.2 – 5.8 mm. Female: antennal club short, little shorter than remaining antennomeres combined; pygidium at apex strongly convex; eyes as large as in male.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFEFFE8FF4359EBFD0DFF66.taxon	description	Figures 17 A – E, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFEFFE8FF4359EBFD0DFF66.taxon	materials_examined	Type material examined. Holotype: ♂ “ Mts. Tianpingshan, Sangzhi, Hunan, 14. VIII. 1988, 350 m, leg. Liu Hong / LW- 037 / 1142 Sericini Asia spec. ” (IZAS). Paratypes: 1 ♂ “ Mts. Tianpingshan, Sangzhi, Hunan, 14. VIII. 1988, 350 m, leg. Liu Hong ” (IZAS), 2 ♂♂ “ Mts. Tianpingshan, Sangzhi, Hunan, 13. VIII. 1988, 1640 m, leg. Yang Xingke ” (IZAS), 1 ♂ “ Mts. Tianpingshan, Sangzhi, Hunan, 12. VIII. 1988, 1650 m, leg. Yang Xingke ” (ZFMK), 1 ♂ “ Wang’erbao, Wanxian, Sichuan, 11. VIII. 1993, 1200 m, leg. Chen Xiaolin ” (IZAS), 1 ♂ “ Fenshuiling, Mts. Hefeng, Hubei, 30. VII. 1989, leg. Sun Baowen ” (IZAS), 1 ♂ “ Fenshuiling, Mts. Hefeng, Hubei, 30. VII. 1989, 1250 m, leg. Sun Baowen ” (IZAS), 1 ♀ “ Fenshuiling Forestry Farm, Mts. Hefeng, Hubei, 30. VII. 1989, 1200 m, leg. Sun Baowen ” (IZAS), 1 ♀ “ Fenshuiling Forestry Farm, Mts. Hefeng, Hubei, 30. VII. 1989, 1250 m, leg. Sun Baowen ” (IZAS), 1 ♂ “ Mts. Jinfoshan, Nanchuan, Chongqing, 23 - 24. VII. 2003, leg. Liu Yushuang, Yuan Caixia ” (HBUM), 1 ♂ “ Mts. Jinfoshan, Nanchuan, Chongqing, 24 - 28. VII. 2003, leg. Liu Yushuang, Yuan Caixia ” (HBUM), 1 ♂ “ Huguosi Temple, Mts. Fanjingshan, Guizhou, 14. VIII. 2001, leg. Pang Hong ” (LYYSU), 1 ♂ “ Tiechanghuang?, 11. VIII. 1989, 1450 m, leg. Yang Longlong ” (IZAS), 1 ♂ “ Wangerbao, Wanxian, Sichuan, 11. VIII. 1993, 1200 m, light trap, leg. Chen Xiaolin, Sun Baowen / LW- 049 ” (IZAS), 1 ♂ “ Wangerbao, Wanxian, Sichuan, 14. VIII. 1993, 1200 m, light trap, leg. Chen Xiaolin, Sun Baowen ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFEFFE8FF4359EBFD0DFF66.taxon	description	Description of the holotype. Length: 8.8 mm, length of elytra: 6.5 mm, width: 4.9 mm. Body oval, dark brown, dull, partly with greenish toment, ventral surface and legs reddish brown, elytra yellow brown with numerous dark spots, in punctures lighter, antenna yellow; dorsal surface almost glabrous except some single and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly, preapically abruptly angled; anterior angles blunt; anterior margin deeply emarginate medially; margins moderately reflexed; surface flat and moderately shiny, finely and densely punctate, partly punctures fusing to transverse wrinkles, with a few erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, without a short terminal seta. Frons completely dull and flat, with fine and dense punctures, posteriorly with numerous fine and short setae. Eyes small, ratio diameter / interocular width: 0.56. Antenna with 10 antennomeres; club with three antennomeres, little longer than remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum strongly produced along middle, narrowly but deeply emarginate medially. Pronotum moderately narrow and subtrapezoidal, widest at base, lateral margins evenly and weakly convex and convergent anteriorly; anterior angles well produced, weakly rounded in the tip; posterior angles blunt; anterior margin straight, robust marginal line widely interrupted medially; surface irregularly, partly densely and partly sparsely punctate, sparse punctures finer, along midline narrowly impunctate and medially on each side with a shallow impression, with sparse, short or minute, white, adpressed setae; lateral margins sparsely setose; hypomeron carinate at base, moderately produced ventrally. Scutellum large, triangular, finely and densely punctate, with minute setae in punctures, on basal midline impunctate. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, sparsely setose, widely glabrous; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.34. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Pygidium strongly convex and dull, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with sparse, moderately long setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 3.6; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face densely and coarsely but superficially punctate, without longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 17 A – D. Habitus: Fig. 17 E.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFEFFE8FF4359EBFD0DFF66.taxon	diagnosis	Diagnosis. Serica tianpingshanensis Ahrens, Fabrizi & Liu, new species differs clearly from the closely related and very similar Serica kuankuoshuiensis Ahrens, Fabrizi & Liu, new species in the shape of parameres: the dorsal lobe of the left paramere is much smaller and shorter, the basal lobe of the right paramere is much shorter and directed basally rather being bend distally.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFEFFE8FF4359EBFD0DFF66.taxon	etymology	Etymology. The species is named (adjective in singular nominative case) after its type locality, Mts. Tianpingshan.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFEFFE8FF4359EBFD0DFF66.taxon	description	Variation. Length: 6.9 – 8.8 mm, length of elytra: 5.2 – 6.5 mm, width: 4.2 – 4.9 mm. Female: Eyes and antennal club as in male; pygidium a little less convex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFCFFE9FF43585AFF45FEF3.taxon	description	Figures 17 F – I, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFCFFE9FF43585AFF45FEF3.taxon	materials_examined	Type material examined. Holotype: ♂ “ Mt. Wuyanling, Taishun, Zhejiang, 3. VIII. 2005, 650 m, leg. Wang Zhiliang / LW- 120 / 1149 Asia Sericini spec. ” (CAU).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFCFFE9FF43585AFF45FEF3.taxon	description	Description of the holotype. Length: 8.2 mm, length of elytra: 6.1 mm, width: 4.9 mm. Body oval, dark reddish brown, dull, elytra with yellow spots, antenna yellow; dorsal surface almost glabrous except some single and short, white setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins strongly convex and convergent anteriorly; anterior angles sharp; anterior margin narrowly and deeply emarginate medially; margins moderately reflexed; surface weakly convex and moderately shiny, finely and densely punctate, with transverse wrinkles and numerous fine and erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus moderately short and slender (1 / 3 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short as well as a few longer, posteriorly bent setae. Eyes small, ratio diameter / interocular width: 0.55. Antenna with 10 antennomeres; club with three antennomeres, little shorter than remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum strongly produced along middle, narrowly and deeply emarginate medially. Pronotum wide and subtrapezoidal, widest at base, lateral margins evenly convex and convergent anteriorly; anterior angles weakly produced, sharp; posterior angles moderately rounded; anterior margin straight, marginal broad but medially widely incomplete; surface irregularly, densely and finely punctate, with minute setae in punctures and a few pale adpressed setae on disc; anterior and lateral margins densely setose; hypomeron carinate at base, weakly produced ventrally. Scutellum moderately large, triangular, finely and densely punctate, with minute setae in punctures, at median base impunctate. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, short, white setae on all intervals, otherwise with minute white setae in punctures; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, sparsely setose, widely glabrous; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.3. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Pygidium strongly convex and dull, dark, midline impunctate, finely and densely punctate, with sparse, short setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.1; dorsal margin sharply carinate, with two groups of spines, basal group at basal third, apical group at two thirds of metatibial length, basally with a few single, robust setae; external face sparsely and finely punctate, without longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally finely and moderately densely punctate. Metatarsomeres ventrally glabrous, laterally and dorsally weakly carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 17 F – H. Habitus: Fig. 17 I. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFCFFE9FF43585AFF45FEF3.taxon	diagnosis	Diagnosis. Serica taishun Ahrens, Fabrizi & Liu, new species differs from Serica silviae Ahrens, 2009 by the absent longer, distally directed dorsal lobe of the left paramere, and in having the dorsal lobe of the right paramere more robust than the ventral one (it is the opposite in S. silviae).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFCFFE9FF43585AFF45FEF3.taxon	etymology	Etymology. The species is named (noun in apposition) after its occurrence close to the city Taishun (Zhejiang).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFDFFEEFF4358EEFBF1FD8E.taxon	description	Figures 18 A – D, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFDFFEEFF4358EEFBF1FD8E.taxon	materials_examined	Type material examined. Holotype: ♂ “ Huangjing, Luzhou, Sichuan, 16. VII. 2002, leg. Bai Ming, Wang Jianfeng / LW- 167 / 1146 Asia Sericini spec. ” (HBUM). Paratypes: 1 ♂ “ Mts. Mianxuling, Fanjingshan, Guizhou, 29. VII. 2001, 1900 m, leg. Dong Kangzhen ” (IZAS), 1 ♂ “ Sixiangping, Jiangkou, Guizhou, 1. VIII. 2001, leg. Ren Guodong / LW- 141 ” (HBUM).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFDFFEEFF4358EEFBF1FD8E.taxon	description	Description of the holotype. Length: 7.6 mm, length of elytra: 5.6 mm, width: 4.2 mm. Body oval, dark reddish brown, dull, elytra yellow brown with numerous dark spots, partly with greenish toment, ventral surface and legs reddish brown, antenna yellow; dorsal surface almost glabrous except some single and short, white adpressed setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent anteriorly; anterior angles blunt; anterior margin deeply emarginate medially; margins moderately reflexed; surface weakly convex and moderately shiny, finely and densely punctate, partly punctures fusing to transverse wrinkles, with numerous erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes small, ratio diameter / interocular width: 0.53. Antenna with 10 antennomeres; club with three antennomeres, 1.2 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum strongly produced along middle, narrowly but deeply emarginate medially. Pronotum moderately narrow and subtrapezoidal, widest shortly behind middle, lateral margins evenly convex and convergent anteriorly and posteriorly; anterior angles weakly produced, sharp; posterior angles blunt; anterior margin straight, robust marginal line complete; surface irregularly, densely and coarsely punctate, along basal midline narrowly impunctate and medially on each side with a shallow impression, with minute setae in punctures otherwise glabrous; lateral margins sparsely setose; hypomeron carinate at base, not produced ventrally. Scutellum large, triangular, finely and densely punctate, with minute setae in punctures, on basal midline impunctate. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, white, adpressed setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, sparsely setose, widely glabrous; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.4. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Pygidium strongly convex and dull, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with sparse, long and short setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.0; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face densely and coarsely but superficially punctate, without longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally smooth. Metatarsomeres ventrally glabrous, laterally weakly carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 18 A – C. Habitus: Fig. 18 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFDFFEEFF4358EEFBF1FD8E.taxon	diagnosis	Diagnosis. Serica huangjing Ahrens, Fabrizi & Liu, new species resembles in shape of aedeagus to Serica sigipinensis Ahrens, 2009. Serica huangjing differs by the strongly asymmetric parameres which strongly differ in length; their ventral lobes are finer while the dorsal obe of the left paramere is strongly bent laterally (dorsal view) before apex.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFDFFEEFF4358EEFBF1FD8E.taxon	etymology	Etymology. The species is named (noun in apposition) after its type locality, Huangjing.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFDFFEEFF4358EEFBF1FD8E.taxon	description	Variation. Length: 7.5 – 7.6 mm, length of elytra: 5.4 – 5.6 mm, width: 4.1 – 4.2 mm.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFBFFEFFF4359EBFCCFF86E.taxon	description	Figures 18 E – H, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFBFFEFFF4359EBFCCFF86E.taxon	materials_examined	Type material examined. Holotype: ♂ “ Hailuogou, Luding, Sichuan, 6. VIII. 2004, 1500 - 1600 m, leg. Zhang Yong / LW- 178 / 1148 Asia Sericini spec. ” (IZAS). Paratype: 1 ♂ “ Hailuogou, Luding, Sichuan, 9. VIII. 2004, leg. Wan Xia, Zhang Yong / LW- 190 ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFBFFEFFF4359EBFCCFF86E.taxon	description	Description of the holotype. Length: 8.0 mm, length of elytra: 6.0 mm, width: 4.4 mm. Body oval, dark reddish brown, dull, elytra with yellow spots, ventral surface and legs reddish brown, antenna yellow; dorsal surface almost glabrous except some single and short, white setae on pronotum and elytra. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins almost straight and weakly convergent anteriorly; anterior angles moderately rounded; anterior margin widely but moderately deeply emarginate medially; margins moderately reflexed; surface weakly convex and shiny, finely and densely punctate, with numerous fine and erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.5 times as wide as long; ocular canthus short and slender (1 / 5 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and dense punctures, with numerous fine and short, erect setae. Eyes small, ratio diameter / interocular width: 0.5. Antenna with 10 antennomeres; club with three antennomeres, as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum moderately produced along middle, narrowly and deeply emarginate medially. Pronotum wide and subtrapezoidal, widest shortly before base, lateral margins evenly convex and convergent anteriorly; anterior angles weakly produced, sharp; posterior angles strongly rounded; anterior margin straight, marginal line fine but complete; surface irregularly, densely and finely punctate, with minute setae in punctures, with a few pale adpressed setae on disc; lateral margins sparsely setose; hypomeron carinate at base, not produced ventrally. Scutellum large, triangular, finely and densely punctate, with minute setae in punctures, on basal midline almost punctate. Elytra oval, widest at posterior third, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, short, white setae on odd intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose; apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, sparsely setose, widely glabrous; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.34. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being one third as long as sternite. Pygidium weakly convex and dull, dark, finely and sparsely punctate, with sparse, short setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.3; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, basally with a few single, robust setae; external face densely and coarsely but superficially punctate, without longitudinal wrinkles; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally finely and moderately densely punctate. Metatarsomeres ventrally glabrous, laterally and dorsally weakly carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 18 E – G. Habitus: Fig. 18 H. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFBFFEFFF4359EBFCCFF86E.taxon	diagnosis	Diagnosis. Serica hailuogou Ahrens, Fabrizi & Liu, new species differs from Serica huangjing Ahrens, Fabrizi & Liu, new species by the strongly curved (dorsal view) dorsal lobe of the right paramere, and the preapically abrubtly bent dorsal lobe of the left paramere.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFBFFEFFF4359EBFCCFF86E.taxon	etymology	Etymology. The species is named (noun in apposition) after its type locality, Hailuogou.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFFBFFEFFF4359EBFCCFF86E.taxon	description	Variation. The paratype has the same size as the holotype.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF8FFECFF4359EBFA0CF86E.taxon	description	Figures 19 A – D, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF8FFECFF4359EBFA0CF86E.taxon	materials_examined	Type material examined. Holotype: ♂ “ Taiwan, Kaosuing Hsien, Kuanshan trail at Kaunshanchi Riv. 2400 m 20. VII. 93 A. Smetana (T 158) ” (CMNC).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF8FFECFF4359EBFA0CF86E.taxon	description	Description of the holotype. Length: 9.4 mm, length of elytra: 7.1 mm, width: 5.4 mm. Body elongate, dark brown, dull, antenna yellow, base and margins of pronotum lighter; dorsal surface almost glabrous except some short, white setae on elytra. Labroclypeus short, subtrapezoidal, much wider than long, widest at base, lateral margins strongly convex and convergent anteriorly; anterior angles moderately rounded; anterior margin deeply widely emarginate medially; margins moderately reflexed; surface weakly convex and shiny, finely and densely punctate, without wrinkles, with numerous erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.2 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and sparse punctures, with a few fine and short, erect setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter / interocular width: 0.5. Antenna with 10 antennomeres; club with three antennomeres, 1.1 times as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum moderately produced along middle, weakly emarginate medially. Pronotum wide and subtrapezoidal, widest at base, lateral margins evenly and strongly convex and strongly convergent anteriorly; anterior angles distinctly produced and sharp; posterior angles strongly rounded; anterior margin straight, with robust and complete marginal line; surface finely densely punctate, without smooth midline, with a few single fine pale setae on the sides, otherwise only with minute setae; lateral and anterior margins densely setose; hypomeron carinate at base, moderately produced ventrally. Scutellum small, triangular, finely and irregularly sparsely punctate, with minute setae in punctures. Elytra elongate, widest at posterior fifth, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, adpressed white setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose (but setae in holotype abraded); apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, sparsely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.31. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being a quarter as long as sternite. Pygidium strongly convex and moderately shiny, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with moderately dense, long and short setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.3, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at four fifths of metatibial length, basally with a few single, fine setae; external face sparsely and finely punctate, without wrinkles, glabrous; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally impunctate. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 19 A – C. Habitus: Fig. 19 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF8FFECFF4359EBFA0CF86E.taxon	diagnosis	Diagnosis. Serica smetanai Ahrens, Fabrizi & Liu, new species is in shape of aedeagus most similar to Serica avoata Nomura, 1974. Serica smetanai differs by the more strongly arched dorsal lobes of parameres which are of the same length.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF8FFECFF4359EBFA0CF86E.taxon	etymology	Etymology. The new species is named (substantive in singular genitive case) after its collector, Aleš Smetana.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF9FFE2FF435F4FFE0AF8C2.taxon	description	Figure 19 E – H, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF9FFE2FF435F4FFE0AF8C2.taxon	materials_examined	Type material examined. Holotype: ♂ “ Taiwan, Hualien Prov. Yuanfeng 2750 m, 18. VI. 1997 B. Herczig & L. Ronkay ” (HNHM). Paratype: 1 ♀ “ Taiwan, Hualien Prov. Yuanfeng 2750 m, 18. VI. 1997 B. Herczig & L. Ronkay ” (HNHM).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF9FFE2FF435F4FFE0AF8C2.taxon	description	Description of the holotype. Length: 9.0 mm, length of elytra: 6.4 mm, width: 4.5 mm. Body elongate, dark brown, dull, elytra with yellowish-brown with dark brown to reddish brown spots, antenna yellow, base and margins of pronotum lighter; dorsal surface almost glabrous except some short, white setae on elytra. Labroclypeus short, subtrapezoidal, little wider than long, widest at base, lateral margins straight and convergent anteriorly; anterior angles strongly rounded; anterior margin widely emarginate medially; margins moderately reflexed; surface weakly convex and shiny, finely and densely punctate, without wrinkles, with numerous erect setae; frontoclypeal suture finely incised, weakly bent medially; smooth area anterior to eye small and convex, 1.2 times as wide as long; ocular canthus short and slender (1 / 4 of ocular diameter), impunctate, with a short terminal seta. Frons completely dull and flat, with fine and sparse punctures, with a few fine and short, erect setae beside eyes, behind frontoclypeal suture, and on disc. Eyes very small, ratio diameter / interocular width: 0.45. Antenna with 10 antennomeres; club with three antennomeres, as long as remaining antennomeres combined and straight. Mentum strongly elevated anteriorly, flattened anteriorly. Labrum moderately produced along middle, deeply emarginate medially. Pronotum narrow and subtrapezoidal, widest at base, lateral margins in basal half subparallel and almost straight, at middle convexly bent, in anterior half again straight and convergent anteriorly; anterior angles distinctly produced and sharp; posterior angles strongly rounded; anterior margin straight, with robust and complete marginal line; surface finely densely punctate, without smooth midline, with a few single fine pale setae on sides, otherwise only with minute setae; lateral and anterior margins densely setose; hypomeron carinate at base, moderately produced ventrally. Scutellum small, triangular, finely and irregularly sparsely punctate, with minute setae in punctures, base impunctate. Elytra elongate, widest at posterior fifth, striae finely impressed, finely and densely punctate; intervals flat, with fine, sparse punctures; with a few single, fine, short, adpressed white setae on all intervals; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose (but setae in holotype abraded); apical margin narrowly membraneous, with a fine rime of minute microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, sparsely setose; metacoxa glabrous, except a few short lateral setae. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.42. Abdominal sternites with a transverse row of coarse punctures, each bearing a long seta, otherwise glabrous; penultimate sternite at posterior margin with a wide membraneous rim being a quarter as long as sternite. Pygidium strongly convex and moderately shiny, dark, in punctures yellow, finely and densely punctate, smooth midline narrow, with moderately dense, long and short setae. Legs slender, shiny, pro- and mesofemur dull; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows. Metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior edge; ventral posterior margin serrated and only weakly widened at apex, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as metafemur width. Metatibia slender and long, widest shortly before apex, ratio of width / length: 1 / 4.4, dorsally sharply carinate, with two groups of spines, basal group at middle, apical group at four fifths of metatibial length, basally with a few single, fine setae; external face sparsely and finely punctate, without wrinkles, glabrous; ventral margin serrated, with two widely separated robust setae; medial face finely sparsely punctate, glabrous; apex interiorly near tarsal articulation bluntly truncate. Tarsomeres ventrally with sparse, minute setae, dorsally impunctate. Metatarsomeres ventrally glabrous, laterally and dorsally carinate, with a few indistinct dorsal punctures, a strongly serrated ridge ventrally; first metatarsomere little shorter than following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, external margin smooth but bluntly extended at middle; anterior claws asymmetrical, basal tooth of inner claw widened and sharply pointed at apex. Aedeagus: Fig. 19 E – G. Habitus: Fig. 19 H.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF9FFE2FF435F4FFE0AF8C2.taxon	diagnosis	Diagnosis. Serica pseudogracilipes Ahrens, Fabrizi & Liu, new species is in shape of aedeagus very similar to Serica niitakana Sawada, 1939, S. yui Kobayashi, 1993 and S. gracilipes Nomura, 1974, both occurrring also in Taiwan. Serica pseudogracilipes differs from the latter two by the finer parameres including their dorsal lobes (dorsal view), which are at middle stronger bent than in Serica yui but not as strong as in S. gracilipes (lateral view). From Serica niitakana it differs by the wider aedeagus and the strongly curved parameres (both dosal view).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF9FFE2FF435F4FFE0AF8C2.taxon	etymology	Etymology. The name of new species is derived from the combined Greek prefix ‘ pseudo- ’ (substantive in singular genitive case) and the species name ‘ gracilipes ’, with references to the similarity of the species to S. gracilipes Nomura, 1974.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF9FFE2FF435F4FFE0AF8C2.taxon	description	Variation. Length: 9.0 – 9.5 mm, length of elytra: 6.4 – 6.9 mm, width: 4.5 – 5.0 mm. Female: antennal club short, as long as remaining antennomeres combined; pygidium weakly convex; eyes smaller than in male, ratio diameter / interocular width: 0.48.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF7FFE1FF4359EBFEA8FACE.taxon	description	Figures 20 A – D, 25	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF7FFE1FF4359EBFEA8FACE.taxon	materials_examined	Type material examined. Holotype: ♂ “ China-Guangdong, Daqiao env., 1000 - 1200 m 24 ° 54 ’ N - 113 ° 01 ’ E, 1 - 3. v. 2002, Dr. R. Fencl lgt. / Coll. P. Pacholatko / 1151 Sericini: Asia spec. ” (CP).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF7FFE1FF4359EBFEA8FACE.taxon	description	Description of the holotype. Length: 7.8 mm, length of elytra: 5.4 mm, width: 4.2 mm. Body oblong, dorsal surface yellow, frons and ventral surface darker brown, legs and antenna yellowish brown, dorsal surface entirely shiny and glabrous. Labroclypeus subtrapezoidal and moderately wide, widest at base; lateral margins convex and convergent to moderately rounded anterior angles; lateral margin and ocular canthus producing a distinct blunt angle; margins weakly reflexed; anterior margin distinctly emarginate medially; surface flat and shiny, finely and very densely punctate, distance between punctures less than their diameter, anteriorly with a few short, erect setae. Frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye short, approximately twice as wide as long; ocular canthus moderately long and slender, finely and densely punctate, with a short terminal seta. Frons with fine and dense punctures, entirely glabrous. Eyes very moderately large, ratio of diameter / interocular width: 0.63. Antenna with nine antennomeres; club with three antennomeres, almost 2.3 times as long as remaining antennomeres combined, strongly reflexed. Mentum elevated and flattened anteriorly. Labrum slightly produced and shallowly emarginate medially. Pronotum wide and transverse, widest at base, lateral margins moderately evenly convex and narrowed anteriorly; anterior angles moderately produced and sharp, posterior angles blunt but strongly rounded at tip; anterior margin strongly and convexly produced medially with a distinct and broad marginal line; basal margin without marginal line; surface with dense and fine punctures, partly punctures fusing to wrinkles, glabrous; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base, not produced ventrally. Scutellum narrow and long, with fine and moderately dense punctures, impunctate along the middle, glabrous. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate; intervals weakly convex, second interval flat, with fine and dense punctures concentrated along striae; surface glabrous except for a few fine setae on odd intervals; epipleural border robust, weakly reflexed, ending at strongly curved external apical angle; epipleura densely setaceous; apical border membraneous, with a very fine rim of short microtrichomes (visible at 100 x magnification). Ventral surface partly dull or shiny, with dense, large punctures, sparsely setose, only on metasternal disc with a few longer setae. Mesosternum between mesocoxae almost as wide as mesofemur, with irregularly scattered, strong setae. Ratio of length of metepisternum / metacoxa: 1 / 1.1. Metacoxa glabrous, laterally with a few fine setae. Abdominal sternites with fine, dense punctation, each with indistinct transversal row of coarse punctures bearing a short seta; penultimate sternite with a shallow and short median furrow and a pair of blunt protuberances beside it. Pygidium dull and strongly convex, finely and densely punctate, without smooth midline, with sparse short setae and a few longer setae adjacent to apical margin. Legs slender, shiny; femora with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur shiny, anterior margin acute, without a submarginal serrated line; posterior ventral margin straight, with a few strong setae medially, weakly widened in apical half and serrate; dorsal posterior margin completely serrated, with short setae. Metatibia slender and long, widest shortly before apex, ratio width / length: 1 / 4.2; dorsal margin sharply carinate, with one group of spines (basal group of spines reduced) at four-fifths of metatibial length, basally with a few single spines in punctures; external face beside dorsal margin longitudinally roof-like carinate, impunctate but with some longitudinal, very superficial wrinkles; ventral margin finely serrate, with two widely separated robust setae; medial face sparsely but coarsely punctate, without wrinkles; apex bluntly truncate interiorly near tarsal articulation. Tarsomeres glabrous and impunctate dorsally, with sparse, short setae ventrally; metatarsomeres dorsally with weak longitudinal impressions, ventrally with a strongly serrated ridge and a fine longitudinal carina laterally; first metatarsomere little longer than second, one third of its length longer than dorsal tibial spur. Protibia moderately long, bidentate; protarsal claws symmetrical, basal tooth of interior claw sharply pointed as the external one. Aedeagus: Parameres asymmetrical, without long basal lobes. Aedeagus: Fig. 20 A – C. Habitus: Fig. 20 D. Female unknown.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF7FFE1FF4359EBFEA8FACE.taxon	diagnosis	Diagnosis. Nipponoserica daqiao Ahrens, Fabrizi & Liu, new species differs from all shiny Nipponoserica species by its asymmetric parameres.	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF7FFE1FF4359EBFEA8FACE.taxon	etymology	Etymology. The new species is named (noun in apposition) after the type locality Daqiao. New records:	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF5FFE1FF435CD6FB62F9E6.taxon	materials_examined	Additional material material: 1 ♂ “ Yaojiaping, Lushui, Yunnan, 2. VI. 1981, 2500 m, leg. Zhang Xuezhong ” (IZAS), 1 ♂ “ Yunlong, Yunnan, 20. VI. 1981, 2450 m, leg. Liao Subai ” (IZAS), 1 ♂ “ China, C-Yunnan, SE of Jongdong, Ailao Shan, 24 ° 09 ’ N, 101 ° 22 ’ E 2600 - 2900 m, 14. - 19. VI. 2014, pitf. Tr., leg. Reuter ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF5FFE1FF435FDAFB33F82A.taxon	materials_examined	Additional material material: 4 ex. “ China, W Yunnan prov., mts. 60 Km E Tengchong, 2300 m, 14. - 19. v. 2006 S. Murzin & I. Shokin leg. ” (CP), 1 ex. “ China: Yunnan [CH 07 - 13 A] Baoshan Pref., Gaoligong Shan, E pass, 36 km SE Tengchong, 2200 m, 24 ° 49 ’ 32 ’’ N, 98 ° 46 ’ 06 ’’ E, decid. Forest, litter, wood, fungi sifted, 4. VI. 2007, leg. A. Pütz ” (CA).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF5FFE1FF435FDAFB33F82A.taxon	discussion	Remark. The species is for first time recorded from China (see Ahrens & Fabrizi 2009).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF2FFE6FF435C19FE50F9BC.taxon	materials_examined	Additional material examined. 4 ex. “ Myanmar (Burma) Provinz Kanchin State, Ca. 20 km N von Kanphant, 23.05.2006, 2180 m, N 25 ° 43 ’ 30.2 ’’ E 098 ° 23 ’ 35.3 ’’, Nachtfang, leg. M. Langer, S. Naumann & S. Löffler Coll. Michael Langer ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF2FFE6FF435F1DFC5CF8E0.taxon	materials_examined	Additional material examined. 1 ♂ “ China, Yunnan prov. Heishui 35 km N Lijiang 18.6 - 4.7.1993, 27.13 N; 100.19 E lgt. S. Becvar ” (ZFMK), 2 ♂♂, 1 ♀ “ Yaojiaping, Lushui, Yunnan, 2. VI. 1981, 2500 m, light trap, leg. Wang Shuyong ” (IZAS), 1 ♂ “ Mts. Yulongshan, Lijiang, Yunnan, 28. VI. 1962, leg. Song Shimei ” (IZAS), 1 ♂, 2 ♀♀ “ Yaojiaping, Lushui, Yunnan, 2. VI. 1981, 2500 m, light trap, leg. Zhang Xuezhong, Wang Shuyong ” (IZAS), 1 ♂ “ Qingtian Ge, Weixi, Yunnan, 23. VII. 1981, light trap, leg. Wang Shuyong ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF3FFE7FF435FFFFEEBF8B9.taxon	materials_examined	Additional material examined. 1 ♂ “ Mts. Ailaoshan, Jingdong, Yunnan, 7 - 9. VIII. 2009, leg. Xu Jishan, Zhang Liuxiang ” (HBUM).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF3FFE4FF435E27FC6DFED6.taxon	materials_examined	Additional material examined. 1 ex. “ Kuatun (2300 m) 27,40 n. Br. 117,40 ö. L. J. Klapperich 28.4. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 12.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 10.6. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun Fukien China, 16.5.46 (Tschung Sen.) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 2 ex. “ China Fukien 02. V. 1938 leg. Klapperich ZFMK Bonn ” (ZFMK), 4 ex. “ China Fukien 03. V. 1938 leg. Klapperich ZFMK Bonn ” (ZFMK), 1 ♂ “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 3.5.1938 (Fukien) ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF3FFE7FF435C1FFCD7F981.taxon	materials_examined	Additional material examined. 1 ♂ “ Mts. Zhibenshan, Yunlong, Yunnan, 20. VI. 1981, 2500 m, leg. Zhang Xuezhong ” (IZAS), 1 ♂ “ Pantiange, Weixi, Yunnan, 23. VII. 1981, 2500 m, light trap, leg. Wang Shuyong ” (IZAS), 1 ♂ “ Sanshenggou, Wolong, Sichuan, 9. VIII. 1983, 2500 m, leg. Niu Chenlai ” (IZAS), 1 ♂ “ Mts. Zhibenshan, Yunlong, Yunnan, 20. VI. 1981, 2500 m, leg. Zhang Xuezhong ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF0FFE4FF4358CBFDE1FD8E.taxon	materials_examined	Additional material examined. 432 ex. “ China: Sichuan; Wolong Reserve, Sigulian Shan, 31 º 09 ’ N 103 º 06 ’ E v. 2006, 1500 - 1800 m V. Siniaev ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF0FFE4FF435BF3FC86FC8A.taxon	materials_examined	Additional material examined. 4 ♂♂, 21 ♀♀ “ China: Sichuan; Wolong Reserve, Sigulian Shan, 31 º 09 ’ N 103 º 06 ’ E v. 2006, 1500 - 1800 m V. Siniaev ” (ZFMK), 1 ex. “ China-Shaanxi, Daba Shan, Shou Man vil., 32 ° 14 ’ N, 108 ° 34 ’ E, 25. v. - 14. vi. 2000, 1000 m, Siniaev & Plutenko leg. ” (CP).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF435FBBFD3CF8FE.taxon	materials_examined	Additional material examined. 1 ♂ “ CH-Guizhou NE 27. V. - 3. VI. 20 km NW of Jiankou 1995 Fanjing Shan-Kuaichang E. Jendek & O. Sausa leg. / CS 11 ” (CP).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF435DFBFAF9FB5A.taxon	materials_examined	Additional material examined. 2 ♂♂, 2 ♀♀ “ Miaotaizi, Shaanxi, 6. IX. 1991, leg. Tian Rungang ” (MNAFU).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF435F6FFA00F9C6.taxon	materials_examined	Additional material examined. 1 ♂ “ Jiulaoci, Mts. Emeishan, Sichuan, 28. VII. 1957, leg. Huang Keren ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF435C47FF65FA72.taxon	materials_examined	Additional material examined. 1 ♂ “ Wolong, Wenchuan, Sichuan, 29. VII. 1983, 1920 m, leg. Wang Shuyong ” (IZAS).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF4359EBFCA2FD8E.taxon	materials_examined	Additional material examined. China: 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 1.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 8.6. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 2.6. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun (2300 m) 27, 40 n. Br. 117,40 ö. L. J. Klapperich 26.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ Kuatun Fukien China 11.5.46 (Tschung-Sen.) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 2 ♂♂ “ China, Guangdong prov. Nanling National Nature Reserve Dadongshan, 18. - 21. iv. 2013, (border of mixed forest, at light) 24 ° 56.0 ’ N, 112 ° 42. # E, 690 m J. Hájek & J. Růžička leg. ” (NMPC). Vietnam: 1 ♂ “ Vietnam, Vinh Phu, Tam Dao; 02. - 04. V. 1993; leg. M. Hori ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF435AF7FE50FB86.taxon	materials_examined	Additional material examined. 6 ex. “ Myanmar (Burma) Provinz Kanchin State, Mt. Emaw Bum nach Kanphant, 28.05.2006, N 26 ° 09 ’ 23.2 ’’ E 098 ° 31 ’ 16.4 ’’ Waldcamp Holzmeiler, leg. M. Langer, S. Naumann & S. Loeffler Coll. Michael Langer ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
AE7687EAFFF1FFE5FF435BF3FBF6FC8A.taxon	materials_examined	Additional material examined. 2 ♂♂ “ Laos-NE Houa Phan prov., 20 ° 12.138 ’ N, 104 ° 00.621 ’ E, Phu Phan Mt., 1750 m, 17.5. - 3.6.2007, Vit Kubán leg. ” (ZFMK), 1 ♂ “ Laos-NE Houa Phan prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E, 1300 - 1900 m, leg. C. Holzschuh – ZFMK Ankauf 2012 ” (ZFMK).	en	Ahrens, Dirk, Fabrizi, Silvia, Bai, Ming, Liu, Wangang (2022): Taxonomic revision of Serica MacLeay, 1819 (sensu lato) from China and adjacent areas (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), with updates on Nipponoserica Nomura, 1972. Zootaxa 5186 (1): 1-83, DOI: https://doi.org/10.11646/zootaxa.5186.1.1
