taxonID	type	description	language	source
F501252A857F5FDE87B2D49B54B85614.taxon	description	Figs 1, 2	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
F501252A857F5FDE87B2D49B54B85614.taxon	description	Emended description. Gametophyte fragments 2.89 - 4.91 mm long, reddish to yellowish brown (Figs 1 A, B, 2 A), main shoots 0.46 - 1.00 mm wide, irregularly branched; branches 0.64 - 1.67 mm long and 0.28 - 0.92 mm wide. Stem reddish brown to dark brown, 37 - 65 μm in diameter. Lateral leaves incubous, (sub) horizontally spreading, alternate, imbricate to contiguous, complicate-trilobed. Dorsal lobe slightly concave to nearly flat (Figs 1 A-D, 2 A-C), ovate to elliptic, on main shoots 290 - 550 μm long x 200 - 380 μm wide, length: width ratio 1.2 - 1.5: 1; entire-margined, gradually narrowed towards apex or apex broadly rounded; dorsally extending 0.5 - 1.2 x the stem width beyond the farther edge of the stem. Lobe cells hexagonal, marginal cells (sub) isodiametric, 15 - 25 µm in diameter, medial cells slightly elongate, 16 - 36 μm long x 12 - 25 μm wide, up to 1.5 x as long as wide (Figs 1 C-E, 2 B, C); cell walls thin to moderately thickened, with small triangular trigones, no intermediate thickenings seen; one large mamilla per cell, ca. 5 µm high and 7.5 - 10 µm in diameter (Figs 1 E, 2 C), or leaf cells smooth. Ocelli not seen, but occasionally dorsal lobes with conspicuous large and elongated cells in the middle. Ventral lobule Frullania - type (Figs 1 A-D, 2), cylindrical, helmet-shaped to campanulate, inflated, broadest part near opening, 110 - 220 μm long x 80 - 210 μm wide, length-width ratio 0.8 - 2.2: 1, inserted in ca. 30 - 90 μm distance to stem, either nearly parallel to stem or obliquely positioned with upper part oriented towards stem and sometimes overlapping stem; opening not constricted, slightly emarginated, with short, acute tip on outer margin, dorsal and ventral margins angular, with one acute tooth in the middle (Figs 1 B-D, 2 C, D), tooth up to 10 µm (one cell) long, possibly bearing slime papillae at tip [seen on one tooth in GZG. BST. 22040]; lobule cells isodiametric, hexagonal, 11 - 24 μm in diameter, smooth to mamillose. Stylus narrow, lanceolate to linear, 30 - 40 µm long, base 2 - 3 cells wide with uniseriate apex of 1 - 3 cells. Underleaves distant to contiguous, 180 - 310 μm long x 50 - 150 μm wide, length: width ratio 2.1 - 3.6: 1, elliptic to elongate ovate, bilobed 0.3 - 0.8 x their length (Figs 1 F-H, 2 C, D), underleaf lobes lanceolate, (2) 4 - 5 (8) cells wide at base [not well visible on GZG. BST. 22040], gradually narrowing into a single cell wide filament, apically terminated by a slime papilla; lobes entire or weakly to strongly toothed or ciliate, with up to six 10 - 55 µm long teeth or cilia consisting of 1 - 3 uniseriate, quadrate to rectangular cells; teeth and cilia becoming successively smaller towards lobe tip; margins of lower lamina generally entire, occasionally crenulate due to protruding cells or with short teeth (Fig. 2 C, D); underleaf sinus more or less V-shaped with an acute to obtuse vertex. Rhizoids in bundles (Fig. 1 G, H), inserted just below sinus of underleaves, bundles up to 160 µm long and 20 - 30 µm wide at base; rhizoids ca. 10 µm in diameter. Sterile.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
63802BD00E76510C9DB68BD4753EA9FE.taxon	description	Figs 3, 4 A-E	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
63802BD00E76510C9DB68BD4753EA9FE.taxon	etymology	Etymology. The specific epithet honors the amber collector Patrick Mueller (Zweibruecken, Germany) who generously supports our research by providing numerous amber fossils for study.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
63802BD00E76510C9DB68BD4753EA9FE.taxon	diagnosis	Diagnosis. Gametophyte with zig-zagged stems and complicate bilobed lateral leaves; dorsal lobe oblong-elliptic with broadly rounded to obtuse apex; ventral lobule small, Lejeunea - type, longitudinally inserted, rounded rectangular to ovate with subacute to rounded apex, inner margin not ampliate, not exceeding stem; irregularly shaped gemmae produced on lobe margins.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
63802BD00E76510C9DB68BD4753EA9FE.taxon	description	Description. Unbranched gametophyte fragment ca. 4.84 mm long, yellowish to reddish brown (Fig. 3 A-D); main shoot 1.68 - 2.28 mm wide with leaves. Stem dark reddish brown, distinctly zig-zagged, ca. 60 µm wide [numbers of cell rows not clearly visible, possibly 4 - 5]; surface cells elongated, 2 - 3 x as long as wide; ventral and dorsal leaf-free strips ca. 2 cells wide. One rhizoid bundle visible on the lobule of a basal leaf (Fig. 3 E), bundle 7.5 - 10 µm wide, rhizoids up to 130 µm long. Foliation incubous, lateral leaves alternate, complicate bilobed, divided into large dorsal lobe and smaller ventral Lejeunea - type lobule enclosing the ventral leaf surface and forming a sharp postical keel (Fig. 3 D-F). Dorsal lobes imbricate, oblong-elliptic, longer than wide, nearly flat, spreading to more erect on upper part of shoot, but not obliquely patent (Fig. 3 A-D); margin entire to slightly crenulate due to gemmae production; postical margin slightly emarginated at end of keel, then regularly arched towards apex, apex broadly rounded to obtuse, antical margin regularly arched and nearly parallel to postical margin in the lobe middle, abruptly curved near stem and forming an angle of ca. 90 ° to the inner margin, inner margin not ampliate, extending onto the dorsal stem surface up to 0.5 - 1 x the stem width; lobe 550 - 600 µm long x 340 - 420 µm wide, length: width ratio 1.3 - 1.7: 1, length exterior to keel 410 - 510 µm, length of stem insertion ca. 170 - 190 µm [not clearly visible on most leaves]. Cells of lobe margin quadrate to rectangular, 10 - 20 µm long x 15 - 25 µm wide, long axis parallel with leaf margin; medial cells hexagonal, mostly isodiametric to occasionally weakly elongated, 20 - 35 µm long x 20 - 30 µm wide; basal cells of the same size or slightly larger than medial cells, 20 - 40 µm long x 20 - 35 µm wide (Fig. 4 A, B); cell walls thin, with small, triangular trigones possibly present at cell angles; free exterior wall of marginal cells unthickened. Ventral lobules small in relation to lobe (<0.2 x), rounded rectangular to ovate, insertion longitudinal, exterior margin curved, antical margin slightly curved or nearly straight, interior margin not ampliate, barely extending onto the stem surface (Fig. 3 D-F), apex subacute to broadly rounded; keel emerging at an angle of ca. 45 ° from the stem, slightly convex; lobule ca. 150 µm long x ca. 150 µm wide, length: width ratio 0.9 - 1.1: 1, keel length 140 - 160 µm, length of stem insertion 90 - 140 µm. Underleaves lacking. Asexual reproduction by gemmae produced from cells of leaf margin (Fig. 4 B-E, indicated by asterisks), gemmae possibly unistratose, subdiscoid to obcordate to irregularly thalloid as size increases, 20 - 110 µm long x 25 - 70 µm wide, first cell division seemingly periclinal (Fig. 4 C, E); stalk cell of gemmae rectangular. Sterile.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
9A7EE5A978C251DB8342F9B79B542784.taxon	description	Fig. 5 A-E	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
9A7EE5A978C251DB8342F9B79B542784.taxon	etymology	Etymology. The specific epithet refers to the village of Tanai, where Kachin amber is mined.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
9A7EE5A978C251DB8342F9B79B542784.taxon	diagnosis	Diagnosis. Gametophyte irregularly branched, branches similar to main shoot, Radula - type. Lateral leaves complicate bilobed; dorsal lobe oval to ovate with rounded apex, deeply emarginated at end of keel; free exterior wall of marginal cells distinctly thickened; ventral lobules Lejeunea - type, up to 0.4 x as big as dorsal lobes, ovate to rounded trapezoid, insertion longitudinal, antical free margin often reaching antical margin of lobe.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
9A7EE5A978C251DB8342F9B79B542784.taxon	description	Description. Branched gametophyte fragment ca. 4 mm long [tip of main shoot broken off], yellowish to reddish brown (Fig. 5 A-C); main shoot up to 2.12 mm wide with leaves. Branching irregular, Radula - type (Fig. 5 A, C); three intact branches with reduced leaves near their base, becoming main shoot-like, 2.6 - 2.72 mm long and 1.2 - 2.1 mm wide with leaves. Stem dark reddish brown, straight to slightly zig-zagged, on main shoot ca. 70 - 90 µm wide [strongly decomposed], on branches ca. 60 µm wide; surface cells not visible; ventral and dorsal leaf-free strips not visible. Rhizoids not seen. Foliation incubous, lateral leaves alternate, complicate bilobed, divided into large dorsal lobe and smaller ventral Lejeunea - type lobule enclosing the ventral leaf surface and forming a sharp postical keel (Fig. 5 B). Dorsal lobes imbricate, oval to ovate, insertion longitudinal, flat, spreading, not obliquely patent, riding onto dorsal stem surface; margin entire, postical margin deeply curved along inner half, first nearly straight then evenly curved towards apex along outer half, lobe apex broadly rounded, exterior antical margin evenly curved, interior margin curved and slightly ampliate, overlapping the stem up to 1 x the stem width beyond the father edge of the stem; lobes on main shoot 600 - 610 µm long x 440 - 510 µm wide, length: width ratio 1.2 - 1.4: 1, length exterior to keel 390 - 400 µm, length of stem insertion not clearly visible; lobes on branches 390 - 520 µm long x 300 - 360 µm wide, length exterior to keel 180 - 320 µm, length of stem insertion not visible. Marginal lobe cells quadrate to rectangular, 10 - 20 µm long x 15 - 25 µm wide, long axis either perpendicular or parallel with leaf margin (Fig. 5 E); medial cells (sub) isodiametric to slightly elongated, 15 - 30 µm long x 12.5 - 25 µm wide (Fig. 5 D); basal cells not visible; cell surfaces appearing smooth, but possible ornamentation visible in some parts (Fig. 5 D) [cells generally indistinct]; cell walls moderately thickened, with small triangular to subnodulose trigones, free exterior wall of marginal cells distinctly thickened (Fig. 5 E). Ventral lobules ovate to rounded trapezoid, up to 0.4 x as big as lobe, free antical margin often reaching antical margin of lobe (Fig. 5 B), insertion longitudinal, free exterior and antical margins nearly straight to curved, apex narrowly rounded to rounded, interior margin ampliate, ventrally extending up to 1 x the stem width beyond the farther edge of stem; keel emerging at an angle of ca. 45 ° from the stem, lobe deeply emarginated at end of keel; lobules on main stem 290 - 310 µm long x 340 - 370 µm wide, length: width ratio ca. 0.9: 1, keel length 210 - 230 µm, length of stem insertion not visible; on branches 270 - 330 µm long x 300 - 340 µm wide, length: width ratio 0.8 - 1: 1. Underleaves lacking. No asexual reproduction. Sterile.	en	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
