identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5ACD1B7601DB53ECBD6EA8FCC61AC359.text	5ACD1B7601DB53ECBD6EA8FCC61AC359.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calymperites burmensis Heinrichs, Schäf. - Verw., Hedenäs, Ignatov & A. R. Schmidt	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Calymperites burmensis Heinrichs, 
Schaef
.-Verw., 
Hedenaes
, Ignatov &amp; A.R.Schmidt
</p>
            <p> Calymperites burmensis Heinrichs,  Schäf .-Verw.,  Hedenäs , Ignatov &amp; A.  R. Schmidt; in Cretaceous Research 51: 261, figs 1, 2. 2014. </p>
            <p>Holotype.</p>
            <p>AMNH Bu ASJH-2, Amber collection of the Division of Invertebrate Zoology of the American Museum of Natural History, New York, USA.</p>
            <p>Additional specimens investigated.</p>
            <p> BuB4339, BuB4398, Patrick  Müller Amber Collection. </p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Remarks.</p>
            <p>To date, this acrocarpous moss was only known from one specimen (Suppl. material 1: Fig. S1A; Heinrichs et al. 2014). As far as the disintegrated state of the new fossils allows to ascertain, they are in good accordance with the type. In both specimens the leaves are erect-spreading and slightly crisped, their bases are oblong-ovate and sheathing the stem before they taper into lanceolate upper regions, and the costa is single and excurrent on intact leaves (Suppl. material 1: Fig. S1B-D). The apices are acute to awned in BuB4398 (Suppl. material 1: Fig. S1B), while they are mostly broken in BuB4339 (Suppl. material 1: Fig. S1C, D). It is not visible if the leaves are serrulate, or if large hyaline cells are present near the apex. Furthermore, the leaf cells are not clearly visible for the largest part, but they appear to be small and quadrate on upper parts of the lamina (Suppl. material 1: Fig. S1D) and more elongated on lower parts (Suppl. material 1: Fig. S1B).</p>
            <p> Phylum  MARCHANTIOPHYTA</p>
            <p> Class  JUNGERMANNIOPSIDA</p>
            <p> Subclass  JUNGERMANNIIDAE</p>
            <p> Order  PORELLALES</p>
            <p> Family  FRULLANIACEAE</p>
            <p> Genus  Frullania</p>
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	https://treatment.plazi.org/id/5ACD1B7601DB53ECBD6EA8FCC61AC359	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
5997E6384C3A549493B66B779091AE26.text	5997E6384C3A549493B66B779091AE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Frullania baerlocheri Heinrichs, M. E. Reiner, K. Feldberg, von Konrat, Hentschel, Váňa & A. R. Schmidt	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Frullania baerlocheri Heinrichs, M.E.Reiner, K.Feldberg, von Konrat, Hentschel, 
Vana
&amp; A.R.Schmidt
</p>
            <p> Frullania baerlocheri Heinrichs, M.E.Reiner, K.Feldberg, von Konrat, Hentschel,  Váňa &amp; A.  R. Schmidt; in Review of Palaeobotany and Palynology 169: 26, plate IV, figs 2, 3. 2012. </p>
            <p> Frullania pinnata =  Frullania pinnata Heinrichs, K.Feldberg,  Schäf .-Verw. &amp; M.Krings; in Cretaceous Research 78: 57, figs 1-3. 2017.-Holotype: GZG.BST.21963, Geoscience Centre (GZG) at the University of  Göttingen , Germany; syn. fide Li et al., 2021a. </p>
            <p>Holotype.</p>
            <p>AMNH Bu-FB 1 g, Amber collection of the Division of Invertebrate Zoology of the American Museum of Natural History, New York, USA.</p>
            <p>Additional specimens investigated.</p>
            <p> SNSB-BSPG 2021 XII 1 (  Müller BuB1874; syninclusion  Protofrullania cornigera ), Bavarian State Collection for Palaeontology and Geology, Munich, Germany; GZG.BST.22015 (  Müller BuB3538), GZG.BST.22039 (  Müller BuB4220; with gynoecia), Geoscience Centre (GZG) at the University of  Göttingen , Germany; PB22712, PB23288 (with gynoecia), Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences; BuB3681, BuB3682, Patrick  Müller Amber Collection; ZMNH155044a (syninclusion  Radula heinrichsii ), ZMNH155047b, c (syninclusion  Frullania kachinensis ), Zhejiang Museum of Natural History, China. </p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Remarks.</p>
            <p>The new fossil specimen is in good accordance with the type as well as subsequently described material and represents the third fertile specimen (Suppl. material 1: Fig. S2A-E; compare descriptions in Heinrichs et al. 2012, 2017b; Feldberg et al. 2021b; Li et al. 2021a). Two unfertilized gynoecia are situated on short side branches similar to specimen PB23288 (Li et al. 2021a), but they are less far developed and most characters are obscured (Suppl. material 1: Fig. S2A, C). A subinvolucral lateral leaf with an explanate lobule is well visible (Suppl. material 1: Fig. S2D, indicated by black arrow), whereas the bracteoles and bracts are concealed for the largest part. The typical acute lobe tip can be seen on one bract (Suppl. material 1: Fig. S2E, indicated by white arrow).</p>
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	https://treatment.plazi.org/id/5997E6384C3A549493B66B779091AE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
D05E5B840E855D699A8F5B5BC0F9BE83.text	D05E5B840E855D699A8F5B5BC0F9BE83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Frullania cretacea Hentschel, A. R. Schmidt & Heinrichs	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Frullania cretacea Hentschel, A.R.Schmidt &amp; Heinrichs</p>
            <p> Frullania cretacea Hentschel, A.  R. Schmidt &amp; Heinrichs; in Cryptogamie, Bryologie 30: 326, figs 1-10. 2009. </p>
            <p>Holotype.</p>
            <p>AMNH B-011, Amber collection of the Division of Invertebrate Zoology of the American Museum of Natural History, New York, USA.</p>
            <p>Additional specimens investigated.</p>
            <p> AMNH Bu-FB 1 a-f (AMNH Bu-FB 1 b with gynoecium), AMNH Bu-FB 51, Amber collection of the Division of Invertebrate Zoology of the American Museum of Natural History, New York, USA; GZG.BST.22016, (  Müller BuB3533), GZG.BST.22017 (  Müller BuB1190), Geoscience Centre (GZG) at the University of  Göttingen , Germany; PB23687 Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences; BuB1772, BuB3530, BuB4411, Patrick  Müller Amber Collection; F3157/BU/CJW,  Jörg Wunderlich Amber Collection. </p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Remarks.</p>
            <p>This small specimen is in good accordance with the type and previously described material (Suppl. material 1: Fig. S2F, G; compare descriptions in Hentschel et al. 2009a; Heinrichs et al. 2012; Feldberg et al. 2021b). The typical rectangular to ovate underleaves with two long apical cilia (Suppl. material 1: Fig. S2G) and the lobules, which are inserted parallel to the stem to slightly oblique, are well visible.</p>
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	https://treatment.plazi.org/id/D05E5B840E855D699A8F5B5BC0F9BE83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
F501252A857F5FDE87B2D49B54B85614.text	F501252A857F5FDE87B2D49B54B85614.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Frullania kachinensis Y. Li, Y. - D. Wang & K. Feldberg	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Frullania kachinensis Y.Li, Y.-D.Wang &amp; K.Feldberg</p>
            <p>Figs 1, 2</p>
            <p> Frullania kachinensis Y.Li, Y.-D.Wang &amp; K.Feldberg; in Geological Journal 56: 5048, figs 1-3. 2021. </p>
            <p>Holotype.</p>
            <p>PB22711a, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences.</p>
            <p>Paratypes.</p>
            <p>PB22711c, d, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences; ZMNH155047a, d, e, ZMNH155048c, Zhejiang Museum of Natural History, China.</p>
            <p>Additional specimens investigated.</p>
            <p> GZG.BST.22040 (  Müller BuB4431), Geoscience Centre (GZG) at the University of  Göttingen , Germany. </p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Emended diagnosis.</p>
            <p> Sterile shoots irregularly branched. Foliation incubous; lateral leaves complicate-trilobed, dorsal lobe ovate to elliptic with rounded apex and mamillose cells, ventral lobule  Frullania -type, cylindrical, helmet-shaped to campanulate, with toothed margins, stylus narrow lanceolate to linear. Underleaves distant to contiguous, elliptic to elongate ovate, bilobed 0.3-0.8  × their length; lobes lanceolate, entire-margined to strongly toothed or ciliate, sinus more or less V-shaped. Rhizoids in bundles, inserted just below sinus. </p>
            <p>Emended description.</p>
            <p> Gametophyte fragments 2.89-4.91 mm long, reddish to yellowish brown (Figs 1A, B, 2A), main shoots 0.46-1.00 mm wide, irregularly branched; branches 0.64-1.67 mm long and 0.28-0.92 mm wide. Stem reddish brown to dark brown, 37-65  μm in diameter. Lateral leaves incubous, (sub)horizontally spreading, alternate, imbricate to contiguous, complicate-trilobed. Dorsal lobe slightly concave to nearly flat (Figs 1A-D, 2A-C), ovate to elliptic, on main shoots 290-550  μm long  × 200-380  μm wide, length:width ratio 1.2-1.5:1; entire-margined, gradually narrowed towards apex or apex broadly rounded; dorsally extending 0.5-1.2  × the stem width beyond the farther edge of the stem. Lobe cells hexagonal, marginal cells (sub)isodiametric, 15-25  µm in diameter, medial cells slightly elongate, 16-36  μm long  × 12-25  μm wide, up to 1.5  × as long as wide (Figs 1C-E, 2B, C); cell walls thin to moderately thickened, with small triangular trigones, no intermediate thickenings seen; one large mamilla per cell, ca. 5  µm high and 7.5-10  µm in diameter (Figs 1E, 2C), or leaf cells smooth. Ocelli not seen, but occasionally dorsal lobes with conspicuous large and elongated cells in the middle. Ventral lobule  Frullania -type (Figs 1A-D, 2), cylindrical, helmet-shaped to campanulate, inflated, broadest part near opening, 110-220  μm long  × 80-210  μm wide, length-width ratio 0.8-2.2:1, inserted in ca. 30-90  μm distance to stem, either nearly parallel to stem or obliquely positioned with upper part oriented towards stem and sometimes overlapping stem; opening not constricted, slightly emarginated, with short, acute tip on outer margin, dorsal and ventral margins angular, with one acute tooth in the middle (Figs 1B-D, 2C, D), tooth up to 10  µm (one cell) long, possibly bearing slime papillae at tip [seen on one tooth in GZG.BST.22040]; lobule cells isodiametric, hexagonal, 11-24  μm in diameter, smooth to mamillose. Stylus narrow, lanceolate to linear, 30-40  µm long, base 2-3 cells wide with uniseriate apex of 1-3 cells. Underleaves distant to contiguous, 180-310  μm long  × 50-150  μm wide, length:width ratio 2.1-3.6:1, elliptic to elongate ovate, bilobed 0.3-0.8  × their length (Figs 1F-H, 2C, D), underleaf lobes lanceolate, (2)4-5(8) cells wide at base [not well visible on GZG.BST.22040], gradually narrowing into a single cell wide filament, apically terminated by a slime papilla; lobes entire or weakly to strongly toothed or ciliate, with up to six 10-55  µm long teeth or cilia consisting of 1-3 uniseriate, quadrate to rectangular cells; teeth and cilia becoming successively smaller towards lobe tip; margins of lower lamina generally entire, occasionally crenulate due to protruding cells or with short teeth (Fig. 2C, D); underleaf sinus more or less V-shaped with an acute to obtuse vertex. Rhizoids in bundles (Fig. 1G, H), inserted just below sinus of underleaves, bundles up to 160  µm long and 20-30  µm wide at base; rhizoids ca. 10  µm in diameter. Sterile. </p>
            <p>Remarks.</p>
            <p> The new fossil of  F. kachinensis (Fig. 1) is in rather good accordance with the type (Fig. 2), but some characters are more conspicuous. Most notable are the rhizoid bundles inserted directly below the underleaf sinus as well as the strongly toothed to ciliate underleaf lobes (Fig. 1G, H) and the toothed lobule margins (Fig. 1B-D, indicated by white arrows). A reinvestigation of the holotype also revealed toothed lobule margins (Fig. 2C, D, indicated by white arrows) and the absence of acute lobe apices (Fig. 2B, indicated by black arrow and dashed line). All lobes are gradually narrowed towards the apex or have a broadly rounded apex. The presence of rhizoids in the type material could not be ascertained. </p>
            <p> Genus  Protofrullania</p>
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	https://treatment.plazi.org/id/F501252A857F5FDE87B2D49B54B85614	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
767F363D90DE530D919E7A164273D425.text	767F363D90DE530D919E7A164273D425.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protofrullania cornigera Heinrichs 2017	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Protofrullania cornigera Heinrichs</p>
            <p> Protofrullania cornigera Heinrichs; in Cretaceous Research 74: 225, figs 1, 2. 2017. </p>
            <p>Holotype.</p>
            <p> GZG.BST.21956, Geoscience Centre (GZG) at the University of  Göttingen , Germany. </p>
            <p>Additional specimens investigated.</p>
            <p> SNSB-BSPG 2021 XII 1 (  Müller BuB1874; syninclusion  Frullania baerlocheri ), SNSB-BSPG 2021 XII 2 (  Müller BuB1893; with juvenile gynoecium), SNSB-BSPG 2021 XII 3 (  Müller BuB1897), Bavarian State Collection for Palaeontology and Geology, Munich, Germany; GZG.BST.22018 (  Müller BuB1999; with perianth), GZG.BST.22019 (  Müller BuB3534; with oval underleaves), GZG.BST.22020 (  Müller BuB3535; with gynoecium), GZG.BST.22021 (  Müller BuB3537; with androecia and oval underleaves), GZG.BST.22022 (  Müller BuB3677; with oval underleaves), Geoscience Centre (GZG) at the University of  Göttingen , Germany; PB22707, PB22711b (syninclusion  Frullania kachinensis ), PB23289, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences; BuB3521, BuB3536, BuB3675, BuB3679 (with oval underleaves), BuB3811, BuB3812, BuB4333-4338, BuB4394 (with oval underleaves), BUB4408a, b, BUB4409, BUB4413 (with oval underleaves), Patrick  Müller Amber Collection; F3251/BU/CJW,  Jörg Wunderlich Amber Collection. </p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Remarks.</p>
            <p> All specimens are in good accordance with the type and previously described material (Suppl. material 1: Figs S3-S5; compare descriptions in Heinrichs et al. 2017a; Feldberg et al. 2021b) and show the characteristic helmet-shaped to campanulate  Frullania -type lobules which are often large in relation to the lobes (Suppl. material 1: Figs S3D, S4F, S5B, D, F). As in previously investigated material, the form of the underleaves is very heterogenous (Feldberg et al. 2021b). Most are elongate rectangular to strap-shaped and taper progressively towards the apex (Suppl. material 1: Figs S3D, S4B, S5B). Below the apex they are usually somewhat increasing with the uppermost sector being obtriangular or irregularly rounded and often carrying rhizoid bundles and marginal teeth or cilia. Some underleaves on main shoots and occasionally also on primary branches are ovate to obovate to rectangular with the widest part in the middle and often less prominent cilia (Suppl. material 1: Figs S4F, S5F). One specimen (BuB4334) has a very distinct flagelliform branch (Suppl. material 1: Fig. S3E). </p>
            <p> Family  RADULACEAE</p>
            <p> Genus  Radula</p>
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	https://treatment.plazi.org/id/767F363D90DE530D919E7A164273D425	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
63802BD00E76510C9DB68BD4753EA9FE.text	63802BD00E76510C9DB68BD4753EA9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Radula patrickmuelleri K. Feldberg, Schaef. - Verw. & M. A. M. Renner 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Radula patrickmuelleri K.Feldberg, 
Schaef
.-Verw. &amp; M.A.M.Renner
 sp. nov.</p>
            <p>Figs 3, 4A-E</p>
            <p>Holotype.</p>
            <p> GZG.BST.22041 (  Müller BuB4395), Geoscience Centre (GZG) at the University of  Göttingen , Germany. </p>
            <p>Etymology.</p>
            <p> The specific epithet honors the amber collector Patrick  Müller (  Zweibrücken , Germany) who generously supports our research by providing numerous amber fossils for study. </p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Diagnosis.</p>
            <p> Gametophyte with zig-zagged stems and complicate bilobed lateral leaves; dorsal lobe oblong-elliptic with broadly rounded to obtuse apex; ventral lobule small,  Lejeunea -type, longitudinally inserted, rounded rectangular to ovate with subacute to rounded apex, inner margin not ampliate, not exceeding stem; irregularly shaped gemmae produced on lobe margins. </p>
            <p>Description.</p>
            <p> Unbranched gametophyte fragment ca. 4.84 mm long, yellowish to reddish brown (Fig. 3A-D); main shoot 1.68-2.28 mm wide with leaves. Stem dark reddish brown, distinctly zig-zagged, ca. 60  µm wide [numbers of cell rows not clearly visible, possibly 4-5]; surface cells elongated, 2-3  × as long as wide; ventral and dorsal leaf-free strips ca. 2 cells wide. One rhizoid bundle visible on the lobule of a basal leaf (Fig. 3E), bundle 7.5-10  µm wide, rhizoids up to 130  µm long. Foliation incubous, lateral leaves alternate, complicate bilobed, divided into large dorsal lobe and smaller ventral  Lejeunea -type lobule enclosing the ventral leaf surface and forming a sharp postical keel (Fig. 3D-F). Dorsal lobes imbricate, oblong-elliptic, longer than wide, nearly flat, spreading to more erect on upper part of shoot, but not obliquely patent (Fig. 3A-D); margin entire to slightly crenulate due to gemmae production; postical margin slightly emarginated at end of keel, then regularly arched towards apex, apex broadly rounded to obtuse, antical margin regularly arched and nearly parallel to postical margin in the lobe middle, abruptly curved near stem and forming an angle of ca. 90° to the inner margin, inner margin not ampliate, extending onto the dorsal stem surface up to 0.5-1  × the stem width; lobe 550-600  µm long  × 340-420  µm wide, length:width ratio 1.3-1.7:1, length exterior to keel 410-510  µm , length of stem insertion ca. 170-190  µm [not clearly visible on most leaves]. Cells of lobe margin quadrate to rectangular, 10-20  µm long  × 15-25  µm wide, long axis parallel with leaf margin; medial cells hexagonal, mostly isodiametric to occasionally weakly elongated, 20-35  µm long  × 20-30  µm wide; basal cells of the same size or slightly larger than medial cells, 20-40  µm long  × 20-35  µm wide (Fig. 4A, B); cell walls thin, with small, triangular trigones possibly present at cell angles; free exterior wall of marginal cells unthickened. Ventral lobules small in relation to lobe (&lt;0.2  × ), rounded rectangular to ovate, insertion longitudinal, exterior margin curved, antical margin slightly curved or nearly straight, interior margin not ampliate, barely extending onto the stem surface (Fig. 3D-F), apex subacute to broadly rounded; keel emerging at an angle of ca. 45° from the stem, slightly convex; lobule ca. 150  µm long  × ca. 150  µm wide, length:width ratio 0.9-1.1:1, keel length 140-160  µm , length of stem insertion 90-140  µm . Underleaves lacking. Asexual reproduction by gemmae produced from cells of leaf margin (Fig. 4B-E, indicated by asterisks), gemmae possibly unistratose, subdiscoid to obcordate to irregularly thalloid as size increases, 20-110  µm long  × 25-70  µm wide, first cell division seemingly periclinal (Fig. 4C, E); stalk cell of gemmae rectangular. Sterile. </p>
            <p>Remarks.</p>
            <p> This fossil consists of a short, sterile gametophyte fragment but shows all relevant characters to delimitate it from other  Radula fossils from Kachin amber (Figs 3, 4A-E). The small, rounded rectangular to ovate lobules allow to differentiate it from  R. heinrichsii as well as  R. tanaiensis which have much larger lobules (Fig. 5A, B), and the apically rounded lobes with nearly parallel margins distinguish it from  R. cretacea (Fig. 4F). </p>
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	https://treatment.plazi.org/id/63802BD00E76510C9DB68BD4753EA9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
C8A27576D84D5626A8EB4BF207EC0AB7.text	C8A27576D84D5626A8EB4BF207EC0AB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Radula sp.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Radula sp.</p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Specimen investigated.</p>
            <p> GZG.BST.22043 (  Müller BuB4420), Geoscience Centre (GZG) at the University of  Göttingen , Germany. </p>
            <p>Remarks.</p>
            <p> The new fossil GZG.BST.22043 is not very well preserved, and the identification is difficult. It strongly resembles  R. cretacea (Fig. 4F-H) in having smooth lobe cells (Suppl. material 1: Fig. S6E, F), quadrate to trapeziform lobules (Suppl. material 1: Fig. S6C, D, indicated by arrows), which are very small in relation to the lobes, and globose gemmae on the lobe margins (Suppl. material 1: Fig. S6F). However, the new fossil has obliquely patent leaves on the upper part of the shoot, whereas the posture of the lower leaves is not as dorsoventrally erect and more similar to  R. cretacea (Figs 5F, Suppl. material 1: S6A-D). It also differs in having slightly narrower leaf lobes with a longer acuminate apex (Suppl. material 1: Fig. S6), though the obliqueness of the lobes makes this difficult to assess. </p>
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	https://treatment.plazi.org/id/C8A27576D84D5626A8EB4BF207EC0AB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
9A7EE5A978C251DB8342F9B79B542784.text	9A7EE5A978C251DB8342F9B79B542784.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Radula tanaiensis K. Feldberg, Schaef. - Verw. & M. A. M. Renner 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Radula tanaiensis K.Feldberg, 
Schaef
.-Verw. &amp; M.A.M.Renner
 sp. nov.</p>
            <p>Fig. 5A-E</p>
            <p>Holotype.</p>
            <p> GZG.BST.22042 (  Müller BuB4329), Geoscience Centre (GZG) at the University of  Göttingen , Germany. </p>
            <p>Etymology.</p>
            <p>The specific epithet refers to the village of Tanai, where Kachin amber is mined.</p>
            <p>Locality and horizon.</p>
            <p>Amber mines southwest of the village of Tanai ca. 105 km north of Myitkyina in Kachin State, northern Myanmar, fossil enclosed in upper Albian-lower Cenomanian Kachin amber.</p>
            <p>Diagnosis.</p>
            <p> Gametophyte irregularly branched, branches similar to main shoot,  Radula -type. Lateral leaves complicate bilobed; dorsal lobe oval to ovate with rounded apex, deeply emarginated at end of keel; free exterior wall of marginal cells distinctly thickened; ventral lobules  Lejeunea -type, up to 0.4  × as big as dorsal lobes, ovate to rounded trapezoid, insertion longitudinal, antical free margin often reaching antical margin of lobe. </p>
            <p>Description.</p>
            <p> Branched gametophyte fragment ca. 4 mm long [tip of main shoot broken off], yellowish to reddish brown (Fig. 5A-C); main shoot up to 2.12 mm wide with leaves. Branching irregular,  Radula -type (Fig. 5A, C); three intact branches with reduced leaves near their base, becoming main shoot-like, 2.6-2.72 mm long and 1.2-2.1 mm wide with leaves. Stem dark reddish brown, straight to slightly zig-zagged, on main shoot ca. 70-90  µm wide [strongly decomposed], on branches ca. 60  µm wide; surface cells not visible; ventral and dorsal leaf-free strips not visible. Rhizoids not seen. Foliation incubous, lateral leaves alternate, complicate bilobed, divided into large dorsal lobe and smaller ventral  Lejeunea -type lobule enclosing the ventral leaf surface and forming a sharp postical keel (Fig. 5B). Dorsal lobes imbricate, oval to ovate, insertion longitudinal, flat, spreading, not obliquely patent, riding onto dorsal stem surface; margin entire, postical margin deeply curved along inner half, first nearly straight then evenly curved towards apex along outer half, lobe apex broadly rounded, exterior antical margin evenly curved, interior margin curved and slightly ampliate, overlapping the stem up to 1  × the stem width beyond the father edge of the stem; lobes on main shoot 600-610  µm long  × 440-510  µm wide, length:width ratio 1.2-1.4:1, length exterior to keel 390-400  µm , length of stem insertion not clearly visible; lobes on branches 390-520  µm long  × 300-360  µm wide, length exterior to keel 180-320  µm , length of stem insertion not visible. Marginal lobe cells quadrate to rectangular, 10-20  µm long  × 15-25  µm wide, long axis either perpendicular or parallel with leaf margin (Fig. 5E); medial cells (sub)isodiametric to slightly elongated, 15-30  µm long  × 12.5-25  µm wide (Fig. 5D); basal cells not visible; cell surfaces appearing smooth, but possible ornamentation visible in some parts (Fig. 5D) [cells generally indistinct]; cell walls moderately thickened, with small triangular to subnodulose trigones, free exterior wall of marginal cells distinctly thickened (Fig. 5E). Ventral lobules ovate to rounded trapezoid, up to 0.4  × as big as lobe, free antical margin often reaching antical margin of lobe (Fig. 5B), insertion longitudinal, free exterior and antical margins nearly straight to curved, apex narrowly rounded to rounded, interior margin ampliate, ventrally extending up to 1  × the stem width beyond the farther edge of stem; keel emerging at an angle of ca. 45° from the stem, lobe deeply emarginated at end of keel; lobules on main stem 290-310  µm long  × 340-370  µm wide, length:width ratio ca. 0.9:1, keel length 210-230  µm , length of stem insertion not visible; on branches 270-330  µm long  × 300-340  µm wide, length:width ratio 0.8-1:1. Underleaves lacking. No asexual reproduction. Sterile. </p>
            <p>Remarks.</p>
            <p> The new fossil material consists of a short shoot fragment with the apex broken off and three main shoot-like  Radula -type branches. It is not very well preserved, and many branches and leaves are broken (Fig. 5A-E). The lobules are generally very large in relation to the lobes (Fig. 5A, B) and differentiate this fossil clearly from  R. patrickmuelleri as well as the much smaller  R. cretacea whose lobules are less than 0.2  × the lobe size (Figs 3, 4F). Superficially the new species looks similar to  R. heinrichsii (Fig. 5F, G; compare descriptions in Feldberg et al. 2021b), but the lobules of  R. tanaiensis are up to 0.4  × the lobe size and their antical margin often reaches the antical margin of the lobe (Fig. 5B), while the lobules of  R. heinrichsii remain smaller and are more rounded (Fig. 5F). The lobe cells are somewhat indistinct, but it is clearly visible that the outer walls of the marginal cells are distinctly thickened (Fig. 5E) whereas those of  R. heinrichsii are thinner (Fig. 5G). </p>
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	https://treatment.plazi.org/id/9A7EE5A978C251DB8342F9B79B542784	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Feldberg, Kathrin;Schaefer-Verwimp, Alfons;Li, Ya;Renner, Matt A. M.	Feldberg, Kathrin, Schaefer-Verwimp, Alfons, Li, Ya, Renner, Matt A. M. (2022): Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae). Fossil Record 25 (1): 213-230, DOI: http://dx.doi.org/10.3897/fr.25.82362, URL: http://dx.doi.org/10.3897/fr.25.82362
