taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
11A6DFB8A6C45F52A2C8AD8CBF440CFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
5D89B900B7DA58F48AAA8EBA3B77BA90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
B0BE2A6E37BF5061A48F48FF7ACE6681.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
8BC18AA8D9845DC789C9CEA4DA688318.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
F85261B6BA415FFB82DA26242DD476D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
9C9A5BB17ABA5957A42C8E8A74D696B5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
4F3CE9CFAF9C5601851A980325CF150B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
2B4F30956035530D882599A222775A7E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
3442DAA6A3BA514CA2D62B37E799A179.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
2676561C240C5DD3B82B666BA15DE068.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
0232D3DD438F5881B147D45537BB1144.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764729	https://doi.org/10.3897/zookeys.1127.85737.figure3	Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype).	Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype).	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
0232D3DD438F5881B147D45537BB1144.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764730	https://doi.org/10.3897/zookeys.1127.85737.figure4	Figure 4. Male genitalia. Scale bars: 200 µm A-D Copelatus malavergnorum sp. nov. (holotype) E-H Copelatus zanabato sp. nov. (holotype) A, E Median lobe of aedeagus in right lateral view B, F Median lobe of aedeagus in ventral view C, G Median lobe of aedeagus in left lateral view D, H Left paramere.	Figure 4. Male genitalia. Scale bars: 200 µm A-D Copelatus malavergnorum sp. nov. (holotype) E-H Copelatus zanabato sp. nov. (holotype) A, E Median lobe of aedeagus in right lateral view B, F Median lobe of aedeagus in ventral view C, G Median lobe of aedeagus in left lateral view D, H Left paramere.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
F8647A4EB6A85115AE8E6557E98DF456.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
57D17ECD30875951860B4ABBBE4CEAE7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
DBD337F8D15C574DA2B62586C506AB51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764729	https://doi.org/10.3897/zookeys.1127.85737.figure3	Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype).	Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype).	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
DBD337F8D15C574DA2B62586C506AB51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764730	https://doi.org/10.3897/zookeys.1127.85737.figure4	Figure 4. Male genitalia. Scale bars: 200 µm A-D Copelatus malavergnorum sp. nov. (holotype) E-H Copelatus zanabato sp. nov. (holotype) A, E Median lobe of aedeagus in right lateral view B, F Median lobe of aedeagus in ventral view C, G Median lobe of aedeagus in left lateral view D, H Left paramere.	Figure 4. Male genitalia. Scale bars: 200 µm A-D Copelatus malavergnorum sp. nov. (holotype) E-H Copelatus zanabato sp. nov. (holotype) A, E Median lobe of aedeagus in right lateral view B, F Median lobe of aedeagus in ventral view C, G Median lobe of aedeagus in left lateral view D, H Left paramere.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
970BF0489D895243BB0DD8AA02D45FC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
4EF79FFA02295A3F8987AEBEF0A7CE01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
9D6C4C3E08FE5BAC9EA6FAE4AED12CDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
D4413859D1245668B492926123E7AB90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
C64E289494575290A4271FBA2FDB2B2D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
2978C8725947596A80975C7677064E6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
DB0D9FC7C9CF5BF1961404D6F65A8487.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
12E8DC35C5C4564599B80B9EA73E755B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
F3DA6FBEC9A65B98AADEC3FF17226D44.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
B33BB39611B05624A29C7753FCC7D33C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
407D478E96625322B13AFE3CCA3CCD16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
6B4DE4274F2858D19DBC5790C5A7CC56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
43165BFEDA9F5F5B8A68C0607CB0F45D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
BCD071B2A28E5785AC0B73522639A168.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
D6B6AE2BBC6C5DBFA2D2801DBFB1F78B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
59F6BE32D147525C8F99CE88AFD08254.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
ACDD693E6E1B50E3BE82FCB09EECE56C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
68D7D1208FFE542492C110A0E5A11522.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
63E8D9F3467B5892B70606E7B345E09E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
AAA1740117905ED39A0AC36F4C128034.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
EA4B265D0BF952188497CF3CD498C2BF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
2F4450B01EC05F899F7FFFC21CD15C29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
85AC78A6944654C3B348E88464A8D466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
154AA14CA1615C7290B9BB0864919B02.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
59D02A6C4DF957F398F2D27E3455827A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
700E788A891A58FEA743455EE5FF89D3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
FFF287BE10D75137BFABE5FC147FA128.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
FC43306B09DC5932A4F86050F214CBD2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
B4643799AA5F5FC18A98F9952F0ABB25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/764728	https://doi.org/10.3897/zookeys.1127.85737.figure2	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	Figure 2. Representative habitats of aquatic Adephaga in Makay. Sites located in the peripheral area: A large shallow puddle on the sandy bank of the Mangoky River in Beroroa (MAK- 1 C), habitat of Pachynectes costulifer, Yola costipennis B, C shallow, slowly flowing stream in semi-open area, with sandy bottom and high density of green algae (MAK- 2), habitat of Bidessus longistriga, B. perexiguus, Canthydrus concolor, C. flavosignatus, C. guttula, Clypeodytes concivis, C. sp. Ma 3, Cybister cinctus, Hydaticus servillianus, Hydroglyphus geminodes, Hydrovatus acuminatus, H. capnius, H. cruentatus, H. dentatus, H. otiosus, H. parvulus, H. pictulus, H. testudinarius, H. sp. Ma 7, Laccophilus addendus, L. flaveolus, L. pallescens, L. posticus, L. rivulosus, L. seyrigi, Methles sp. Ma 5, Neohydrocoptus seriatus, Pachynectes sp. Ma 1, Philaccolus elongatus, Pseuduvarus sp. Ma 1, Rhantaticus congestus, Uvarus rivulorum. Sites located in inner Makay: D puddle (on the right) on the sandy bank of the Andranomanintsy River (left half of the picture) (MAK- 3), habitat of Copelatus ruficapillus, Hydrovatus acuminatus, Hyphydrus separandus, Laccophilus makay, L. posticus, Madaglymbus fairmairei, Pachynectes sp. Ma 1 E spring on the bank of the Andranomanintsy River (MAK- 4), habitat of Copelatus ruficapillus, Hydroglyphus capitatus, H. geminodes, Hyphydrus separandus, Laccophilus makay, L. posticus, Pachynectes sp. Ma 1 F deep pond above natural dam in the bottom of a canyon (MAK- 5 A), habitat of Africophilus nesiotes, Hyphydrus separandus, L. addendus, Laccophilus insularum, L. makay, Neptosternus oblongus, Pachynectes sp. Ma 1, P. sp. Ma 4 G edge of small shallow stream, in the bottom of a deep strongly embanked canyon, with orange deposit of iron bacteria (MAK- 6), habitat of Copelatus acamas and Laccophilus makay H vertical rock wall with water film and crust of bryophytes and algae, in the bottom of a deep canyon (MAK- 9), habitat of Africophilus bartolozzii I large quiet and shaded pool in forest, along streamlet, with masses of tree roots and bottom of sand, gravel and stones (MAK- 8), habitat of Africophilus bartolozzii, A. nesiotes, Copelatus ruficapillus, Hydaticus sobrinus, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1 J (context) K (close-up) Puddle with orange masses of iron bacteria, in stream bed, in the bottom of a deep strongly embanked canyon (MAK- 7), habitat of Copelatus acamas, C. ruficapillus, Laccophilus makay L (context) M (close-up) Small pool with clay-sandy bottom and plant debris, in a small canyon, in gallery forest (MAK- 10), habitat of Copelatus ruficapillus, Hyphydrus separandus, Laccophilus makay, Madaglymbus fairmairei N small shaded pool, with orange masses of iron bacteria, in gallery forest against the wall of a canyon (MAK- 14 A), habitat of Africophilus nesiotes, Copelatus acamas, Hydaticus dorsiger, Hyphydrus separandus, Laccophilus makay, Pachynectes sp. Ma 1, P. sp. Ma 4 O small muddy pond, with sparse vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 A), habitat of Canthydrus guttula, Copelatus polystrigus, Hydaticus dorsiger, Laccophilus addendus, L. posticus, Neohydrocoptus seriatus P puddle with turbid water and without vegetation, in open area on the sandy banks of the Menapanda River (MAK- 11 - B), habitat of Copelatus polystrigus, C. ruficapillus, Eretes griseus, Hydaticus dorsiger, H. exclamationis, Hyphydrus separandus, Laccophilus addendus, L. posticus, Madaglymbus fairmairei.	2022-11-02	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael		Zenodo	biologists	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael			
