identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
11A6DFB8A6C45F52A2C8AD8CBF440CFF.text	11A6DFB8A6C45F52A2C8AD8CBF440CFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africophilus bartolozzii Rocchi 1991	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Africophilus bartolozzii Rocchi, 1991</p>
            <p>Type locality.</p>
            <p>Madagascar, Isalo National Park, Canyon des Singes.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-8; 8 ♂♂, 8 ♀♀: MAK-9; 1 ♀: MAK-36B; 1 ♀: MAK-39B.</p>
            <p>Distribution.</p>
            <p>Madagascar; previously known only from the sandstone massif of Isalo (Rocchi 1991).</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2H, I). This hygropetric species was collected only in inner Makay. Three of the sampling sites were situated in forested contexts and one site in a non-forested context. In the latter (MAK-9), the habitat was vertical rock walls with water film and crust of bryophytes and algae, at the bottom of a narrow and deep canyon. The other habitats were puddles and pools with sandy - stony bottom along small streams. Like other members of the genus, this species lives at the edge of small water bodies in the water film retained by capillarity on the substratum surface above water line, or on rock surface covered with a thin and slowly seeping water film, often close to cascades.</p>
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	https://treatment.plazi.org/id/11A6DFB8A6C45F52A2C8AD8CBF440CFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
5D89B900B7DA58F48AAA8EBA3B77BA90.text	5D89B900B7DA58F48AAA8EBA3B77BA90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Africophilus nesiotes Guignot 1951	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Africophilus nesiotes Guignot, 1951</p>
            <p>Type locality.</p>
            <p>Madagascar, Ambalavao region.</p>
            <p>Material examined.</p>
            <p>5 ♂♂, 8 ♀♀: MAK-5A; 1 ♀: MAK-5B; 1 ♀: MAK-8; 2 ♀♀: MAK-13; 3 ♂♂, 4 ♀♀: MAK-14A; 1 ♀: MAK-26; 1 ♀: MAK-39A; 2 ♂♂: MAK-40A</p>
            <p>Distribution.</p>
            <p>Sub-Saharan Africa, Madagascar (Guignot 1959-1961; Bertrand and Legros 1971; Franciscolo 1994; Bilardo et al. 2020). In Madagascar, widespread.</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2F, I, N). Similar to  A. bartolozzii (see above) but more common. </p>
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	https://treatment.plazi.org/id/5D89B900B7DA58F48AAA8EBA3B77BA90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
B0BE2A6E37BF5061A48F48FF7ACE6681.text	B0BE2A6E37BF5061A48F48FF7ACE6681.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bidessus longistriga Regimbart 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Bidessus longistriga 
Regimbart
, 1895
</p>
            <p>Type locality.</p>
            <p>Madagascar, Antsiranana,</p>
            <p>Material examined.</p>
            <p> 1 ♀: MAK-1A; 1 ♂: MAK-2; 2 ♀♀: MAK-19; 17  еxs .: MAK-42; 1  еx .: MAK-60; 1  еx .: MAK-61. </p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984;  Biström 1985; Rocchi 1991; Bergsten et al. 2020). </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C).  Bidessus longistriga was sampled at peripheral sites and at one inner massif site, in open areas, in permanent or temporary lentic environments, with water stagnant or slowly flowing, clear or turbid, with sand and/or clay bottom. It prefers habitats with at least some marginal vegetation, and is tolerant to anthropogenic disturbance. </p>
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	https://treatment.plazi.org/id/B0BE2A6E37BF5061A48F48FF7ACE6681	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
8BC18AA8D9845DC789C9CEA4DA688318.text	8BC18AA8D9845DC789C9CEA4DA688318.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bidessus perexiguus H. J. Kolbe 1883	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Bidessus perexiguus H. J. Kolbe, 1883</p>
            <p>Type locality.</p>
            <p>Madagascar, South inner part.</p>
            <p>Material examined.</p>
            <p>1 ex.: MAK-1A; 4 ♂♂, 1 ex.: MAK-2; 1 ♂: MAK-17; 82 exs.: MAK-18; 4 ♂♂, 25 exs.: MAK-19; 3 exs.: MAK-60; 1 ex.: MAK-61.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1959-1961;  Biström 1985; Bergsten et al. 2020). </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was collected mainly at peripheral sites. Its ecology is similar to that of  B. longistriga , and both species were several times sampled in the same habitats, but  B. perexiguus has a marked preference for small and very shallow water bodies without vegetation. Notably, this species was particularly abundant in a small muddy ditch only 5 cm deep, shaded under trees, without water flow and with turbid water, trampled by cattle and with no vegetation (MAK-18). </p>
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	https://treatment.plazi.org/id/8BC18AA8D9845DC789C9CEA4DA688318	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
F85261B6BA415FFB82DA26242DD476D8.text	F85261B6BA415FFB82DA26242DD476D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Canthydrus concolor Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Canthydrus concolor Sharp, 1882</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2.</p>
            <p>Distribution.</p>
            <p>Madagascar, widespread (Guignot 1959-1961; Bertrand and Legros 1971; Rocchi 1991; Nilsson 2011).</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species is widespread and generally common in Madagascar, in well-vegetated lentic or calm lotic environments. It is seemingly absent from inner Makay, and was sampled only once in a peripheral site. The habitat was a shallow insolated stream characterised by very weak water flow, sandy bottom, marked by anthropic disturbance (cattle trampling), sparse tufts of small  Cyperaceae and strong presence of filamentous green algae. </p>
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	https://treatment.plazi.org/id/F85261B6BA415FFB82DA26242DD476D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
9C9A5BB17ABA5957A42C8E8A74D696B5.text	9C9A5BB17ABA5957A42C8E8A74D696B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Canthydrus flavosignatus Régimbart 1903	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Canthydrus flavosignatus 
Regimbart
, 1903
</p>
            <p>Type locality.</p>
            <p>Madagascar, Fort-Dauphin, Ankara.</p>
            <p>Material examined.</p>
            <p>2 ♂♂: MAK-2; 1 ♀: MAK-19.</p>
            <p>Distribution.</p>
            <p>Zaire (Democratic Republic of the Congo), Madagascar (Guignot 1959-1961; Nilsson 2011).</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was harvested at two sites in the Makay periphery, both in deforested areas. One was the site described for  C. concolor and the other one was a large puddle, with water slowly flowing and with abundant rice straw debris, on a dirty road between two rice fields. </p>
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	https://treatment.plazi.org/id/9C9A5BB17ABA5957A42C8E8A74D696B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
4F3CE9CFAF9C5601851A980325CF150B.text	4F3CE9CFAF9C5601851A980325CF150B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Canthydrus guttula (Aube 1838)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Canthydrus guttula (
Aube
, 1838)
</p>
            <p>Type locality.</p>
            <p> La  Réunion ; Madagascar. </p>
            <p>Material examined.</p>
            <p>3 ♀: MAK-1A; 1 ♀: MAK-2; 1 ♀: MAK-11A; 52 ♂♂, 39 ♀♀: MAK-19; 1 ♂: MAK-20; 1 ♂, 2 ♀♀: MAK-38B; 1 ♂, 3 ♀♀: MAK-40A; 2 ♂♂, 4 ♀♀: MAK-41; 4♂♂, 8 ♀♀: MAK-42; 17 ♂♂, 11 ♀♀: MAK-60;; 1 ♂: MAK-61.</p>
            <p>Distribution.</p>
            <p>Madagascar and Mascarene Islands; widespread and common in Madagascar (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984; Rocchi 1991; Nilsson 2011).</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C, O). Collected mainly in peripheral but also in a few inner massif sites. This species is found in permanent or temporary lentic habitats as well as in the calm margins of slowly flowing water bodies, with at least some amount of clay or mud at the bottom. Although a few individuals were taken at some sites without any vegetation or significant accumulation of organic debris, the species is most abundant in well-vegetated habitats and/or with bottom heavily loaded with dead vegetal material. This species has a preference for open environments and is highly tolerant to anthropogenic perturbation (e.g., present in rice fields).</p>
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	https://treatment.plazi.org/id/4F3CE9CFAF9C5601851A980325CF150B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
4A890A281DD45A80A8DF5EB96F46B5B4.text	4A890A281DD45A80A8DF5EB96F46B5B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Canthydrus sp. Ma 5	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Canthydrus sp. Ma5</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-20; 1 ♂, 3 ♀♀: MAK-60; 3 ♀♀: MAK-61.</p>
            <p>Note.</p>
            <p> This species is very similar to  C. flavosignatus but smaller and with a slightly different shape of the apex of the median lobe of aedeagus in lateral view. The Malagasy species of  Noteridae are in great need of revision, and in the current state of knowledge we cannot assign a name to this species. </p>
            <p>Distribution.</p>
            <p>Madagascar. In addition to the specimens from the Makay, we also have specimens from Namoroka (north-eastern part of the island).</p>
            <p>Habitat in study area.</p>
            <p>Species collected in permanent lentic environments in open peripheral sites, with clay bottom, clear water and presence of vegetation. One of the collecting sites was a rice field.</p>
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	https://treatment.plazi.org/id/4A890A281DD45A80A8DF5EB96F46B5B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
2B4F30956035530D882599A222775A7E.text	2B4F30956035530D882599A222775A7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clypeodytes concivis Guignot 1955	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Clypeodytes concivis Guignot, 1955</p>
            <p>Type locality.</p>
            <p>Madagascar, Iharanandriana mountain.</p>
            <p>Material examined.</p>
            <p>3 ♂♂, 3 ♀♀: MAK-2; 1 ♂: MAK-61.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1955b;  Biström 1988a). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species was collected at two peripheral sites, a very slowly flowing and shallow stream, and a shallow stagnant pond, both in open environments. The mineral substratum was a mixture of sand and clay in the first case, and of clay and mud in the second case, with moderate amount of plant debris. The water was clear and there was a marginal vegetation of helophytes; at site MAK-2 with abundant filamentous green algae. At both sites, the biotope was trampled and enriched in nutrients by cattle.</p>
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	https://treatment.plazi.org/id/2B4F30956035530D882599A222775A7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
8439E0F8FDC4535B8F9E86129F95C634.text	8439E0F8FDC4535B8F9E86129F95C634.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clypeodytes insularis Guignot 1956	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Clypeodytes insularis Guignot, 1956</p>
            <p>Type locality.</p>
            <p>Madagascar, Bas Mangoky agricultural station.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-20.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread but not common (Guignot 1956;  Biström 1988a) </p>
            <p>Habitat in study area.</p>
            <p>A single specimen of this species was taken in a rice field on the banks of the Mangoky River (a peripheral site), at shallow depth. The bottom was composed of sand and clay with some plant debris; there was no visible inlet or outlet, the water was clear and there was a moderate presence of green algae. The environment was open with no trees in the surroundings.</p>
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	https://treatment.plazi.org/id/8439E0F8FDC4535B8F9E86129F95C634	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
3442DAA6A3BA514CA2D62B37E799A179.text	3442DAA6A3BA514CA2D62B37E799A179.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clypeodytes sp. Ma 3	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Clypeodytes sp. Ma3</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2.</p>
            <p>Distribution.</p>
            <p>Madagascar (known to us only from site MAK-2).</p>
            <p>Note.</p>
            <p> This is an undescribed species, close to  C. spangleri Biström , 1988 and  C. pseudolentus Biström , 1988 from continental Africa. </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was sampled only once in a peripheral site. The habitat was a shallow isolated stream characterised by very weak water flow, sandy bottom, marked anthropic disturbance (cattle trampling), sparse tufts of small  Cyperaceae and strong presence of filamentous green algae. </p>
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	https://treatment.plazi.org/id/3442DAA6A3BA514CA2D62B37E799A179	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
2676561C240C5DD3B82B666BA15DE068.text	2676561C240C5DD3B82B666BA15DE068.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus acamas Guignot 1955	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copelatus acamas Guignot, 1955</p>
            <p>Type locality.</p>
            <p>Madagascar, Isalo National Parc.</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-6; 3 ♂♂: MAK-7; 1 ♀: MAK-14A; 7 ♂♂, 13 ♀♀: MAK-30; 18 ♂♂, 17 ♀♀: MAK-32; 2 ♂♂, 3 ♀♀: MAK-34A; 2 ♂♂, 1 ♀: MAK-34B; 3 ♀♀: MAK-35A; 1 ♂: MAK-39A; 42 ♂♂, 25 ♀♀: MAK-45; 10 ♂♂, 7 ♀♀: MAK-46; 1 ♀: MAK-47; 26 ♂♂, 44 ♀♀: MAK-49; 10 ♂♂, 3 ♀♀: MAK-50; 1 ♀: MAK-52; 11 ♂♂, 5 ♀♀: MAK-53; 1 ♂, 2 ♀♀: MAK-54A; 4 ♂♂, 5 ♀♀: MAK-54B; 48 ♂♂, 57 ♀♀: MAK-59B; 5 ♂♂, 2 ♀♀: MAK-59C.</p>
            <p>Distribution.</p>
            <p>Madagascar; previously known only from the sandstone massif of Isalo (Guignot 1955a).</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2G, J, K, N). This species is very common in inner Makay and absent from peripheral sites. It was most often found in puddles and pools located in stream and river beds, as well as small water holes and springs; shaded or sun-exposed, with bottom of sand and/or sandstone, with or without vegetal debris. These water bodies were most often devoid of vegetation. The water was clear but often more or less heavily loaded with orange masses of iron bacteria. Almost all collection points where this species could be observed were in forested or semi-forested areas and all sites were relatively undisturbed.</p>
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	https://treatment.plazi.org/id/2676561C240C5DD3B82B666BA15DE068	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
4E7E203B937C530DB56C1C1E935CE1F6.text	4E7E203B937C530DB56C1C1E935CE1F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus andobonicus Guignot 1960	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copelatus andobonicus Guignot, 1960</p>
            <p>Type locality.</p>
            <p>Madagascar, Andobo, Antsingy forest.</p>
            <p>Material examined.</p>
            <p>4 ♀: MAK-12A; 2 ♀♀: MAK-33; 1 ♂: MAK-57.</p>
            <p>Distribution.</p>
            <p>Madagascar (Guignot 1960); species characteristic of dry deciduous forests in the western part of the island.</p>
            <p>Habitat in study area.</p>
            <p>This species was collected in lentic habitats (springs and small puddles), with clay or sandy-clay bottom and a lot of plant debris, located in forested or semi-forested areas and relatively unaffected by human disturbances.</p>
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	https://treatment.plazi.org/id/4E7E203B937C530DB56C1C1E935CE1F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
0232D3DD438F5881B147D45537BB1144.text	0232D3DD438F5881B147D45537BB1144.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus malavergnorum Manuel & Ramahandrison 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copelatus malavergnorum Manuel &amp; Ramahandrison sp. nov.</p>
            <p>Figs 3A, 4A-D</p>
            <p>Type locality.</p>
            <p>Madagascar, Toliara province, Malaimbandy municipality, Makay massif (northern part), ca. 20 km WSW Tsimazava, ca. 20°45'S, 45°31'E, altitude ca. 360 m a.s.l.</p>
            <p>Type material.</p>
            <p> Holotype ♂: "Madagascar. Ex-prov. Toliara / Makay massif, ca. 20 km / WSW Tsimazava / 20°44'42"S, 45°31'35"E / 11.IV.2018. Ramahandrison leg. [pr.] // Alt. 364 m. Blind channel / connected with the Sakapaly / River, with orange masses of / iron bacteria on the bottom. [pr.] // Holotype /  Copelatus malavergnorum sp. nov. / Manuel &amp; Ramahandrison, 2022 [red, pr.]" (MNHN). </p>
            <p>Diagnosis.</p>
            <p> This species belongs to the  Copelatus irinus -group and the  C. insuetus -complex (revised in Ranarilalatiana et al. 2019). It differs from  C. insuetus Guignot, 1941 by: smaller size; narrower and more parallel habitus, dorsally flatter; broader pronotum with lateral margins posteriorly more parallel; colour of dorsal and ventral surfaces paler; testaceous basal band of elytron broader; discal stria I on elytron more weakly impressed; strioles on postero-lateral region of pronotum and on metacoxal plate sparser; metacoxal lines shorter; medial lobe of aedeagus in lateral view with apical half of more even width and with apex less narrowly acute, in ventral view distinctly more evenly narrowed from base to apex. This species is externally similar to  C. vokoka Ranarilalatiana &amp; Bergsten, 2019, but differs by: habitus narrower and more parallel, dorsally flatter; pronotum broader and with lateral margins posteriorly more parallel; discal stria I on elytron more weakly impressed and anteriorly more strongly abbreviated; strioles on pronotum and metacoxal plates much sparser; metacoxal lines shorter; median lobe of aedeagus in lateral view with apical half broader and less strongly arcuate, in ventral view with apex twisted to the left (straight in  C. vokoka ). Finally, it differs from  C. kely Ranarilalatiana &amp; Bergsten, 2019 notably by the lateral margins of pronotum anteriorly more strongly convergent, the lateral margins of elytra posteriorly more strongly attenuated, the median lobe in lateral view with the distal half much thicker and the dorsal outline less strongly curved, and in ventro-apical view much more gradually narrowed (in  C. kely abruptly narrowed at ca. midlength from base to apex), subapically thicker and less strongly bent to the left. </p>
            <p>Description of holotype.</p>
            <p>Body elongated and parallel-sided (Fig. 3A), weakly convex dorsally. Pronotum broad (ratio between maximum width of pronotum and maximum body width ~ 0.97), with sides posteriorly subparallel. Head rufo-testaceous with only very faint infuscation between eyes. Ratio between interocular distance and maximum width of head ~ 0.66. Pronotum rufo-testaceous as head, with weak medial infuscation. Elytra brown with broad testaceous basal band; testaceous band very diffusely transitioning into darker colour posteriorly (Fig. 3A).</p>
            <p>Elytra with six discal and one submarginal striae. Stria I very weakly impressed. Striae I, V, and VI abbreviated anteriorly. Submarginal stria very weakly impressed, starting slightly before elytron midlength. Head, pronotum and elytra with fine reticulation and fine punctation. Posterolateral region of pronotum with few short and weakly impressed strioles.</p>
            <p>Ventral side rufo-testaceous, slightly darker laterally on metacoxal plate and on abdominal ventrites. Metacoxal plates with sparse and very fine short strioles; visible abdominal ventrites I-III with denser and longer very fine strioles. Prosternal process rather short and broad, with blunt apex. Metacoxal lines rather long, ending anteriorly at quite small distance from posterior margin of metaventrite, diverging anteriorly.</p>
            <p>Appendages: Antennae, palps and legs testaceous. Antennae particularly long (Fig. 3A). First three pro- and mesotarsomeres widened and ventrally equipped with suction cups; number of suction cups per articles (I-III) 7:4:4 on both pro- and mesotarsus. Protibia at base narrow, with bisinuate ventral margin, distally strongly broadened. Pro- and mesotarsal claws unmodified.</p>
            <p>Median lobe and parameres as in Fig. 4A-D.</p>
            <p>Female. Unknown.</p>
            <p>Measurements.</p>
            <p>TL 4.2 mm, TL without head 3.7 mm, MW 1.8 mm, ratio TL/MW 2.34.</p>
            <p>Etymology.</p>
            <p> This species is dedicated to the Malavergne family (Dominique, Catherine,  Clémence , Jacques, and Laurence,  Marie-José ) in recognition of their constant help and support to the first author during his PhD thesis. The species epithet is a name in the genitive plural. </p>
            <p>Distribution.</p>
            <p>So far known only from northern Makay in Madagascar.</p>
            <p>Habitat.</p>
            <p> The external morphology of this species (very narrow and parallel habitus, broad pronotum, pale colour, long antennae) suggests that it might be a semi-subterranean species. The habitat where the single specimen was found (MAK-44C) was a blind channel connected to River Sakapaly, in northern inner Makay. There was no apparent water flow but the bottom was covered with conspicuous orange masses of iron bacteria, which might be an indication of slow water seepage from underground. The substratum was sandy with moderate amount of decaying vegetal material and the water was red-brown coloured. This water body was fully shaded under trees in forest. There was no vegetation in the water but the surrounding forest floor displayed a typical hygrophilic vegetation of  Poaceae ,  Cyperus and  Pandanus . Other species of aquatic  Adephaga (all  Dytiscidae ) sampled at the same site:  Copelatus polystrigus ,  C. vigintistriatus ,  Hydrovatus acuminatus Motschulsky, 1860,  Laccophilus makay ,  Methles sp. Ma1,  M. sp. Ma5,  Neohydrocoptus sp. Ma3, and  Pachynectes sp. Ma1. </p>
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	https://treatment.plazi.org/id/0232D3DD438F5881B147D45537BB1144	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
F8647A4EB6A85115AE8E6557E98DF456.text	F8647A4EB6A85115AE8E6557E98DF456.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus polystrigus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copelatus polystrigus Sharp, 1882</p>
            <p> Copelatus marginalis =  C. marginalis Gschwendtner, 1932 </p>
            <p>Type locality.</p>
            <p>Madagascar, Senegal.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-11A; 1 ♂: MAK-11B; 30 ♂♂, 41 ♀♀: MAK-12A; 1 ♂: MAK-12B; 5 ♂♂, 1 ♀: MAK-12C; 2 ♂♂, 2 ♀♀: MAK-33; 2 ♂♂, 4 ♀♀: MAK-44A; 8 ♂♂, 22 ♀♀: MAK-44B; 23 ♂♂, 22 ♀♀: MAK-44C; 3 ♂♂, 8 ♀♀: MAK-56; 21 ♂♂, 16 ♀♀: MAK-57; 1 ♂: MAK-58.</p>
            <p>Distribution.</p>
            <p>Continental Africa south from Egypt and Sahara, Madagascar (Guignot 1959-1961; Bertrand and Legros 1971). In Madagascar, widespread and common.</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2O, P). This species was found in inner Makay, in pools and puddles, mainly in forest. These water bodies were shallow and did not exceed 1 m in depth. The mineral substratum was either clay or sand (or a mixture of both) and there was always a substantial quantity of plant debris.</p>
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	https://treatment.plazi.org/id/F8647A4EB6A85115AE8E6557E98DF456	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
57D17ECD30875951860B4ABBBE4CEAE7.text	57D17ECD30875951860B4ABBBE4CEAE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus ruficapillus Regimbart 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Copelatus ruficapillus 
Regimbart
, 1895
</p>
            <p>Type locality.</p>
            <p> Madagascar, Antsiranana, Montagne  d’Ambre , Ambohitra National Park. </p>
            <p>Material examined.</p>
            <p>6 ♂♂, 3 ♀♀: MAK-3; 4 ♂♂, 7 ♀♀: MAK-4; 1 ♂: MAK-7; 1 ♀: MAK-8; 6 ♂♂, 10 ♀♀: MAK-10; 1 ♀: MAK-11B; 2 ♂♂, 6 ♀♀: MAK-12A; 2 ♀♀: MAK-12B; 2 ♀♀: MAK-16; 1 ♀: MAK-25A; 1 ♂: MAK-25B; 3 ♂♂, 5 ♀♀: MAK-28; 1 ♂: MAK-29; 1 ♂, 1 ♀: MAK-33; 1 ♀: MAK-34B; 2 ♂♂, 2 ♀♀: MAK-35A; 4 ♂♂: MAK-38A; 1 ♂, 5 ♀♀: MAK-39A; 6 ♂♂, 5 ♀♀: MAK-39B; 1 ♀: MAK-45; 2 ♂♂: MAK-50.</p>
            <p>Distribution.</p>
            <p>Madagascar, widespread (Guignot 1959-1961; Bertrand and Legros 1971).</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2D, E, I-M, P). Similar to  C. acamas (see above). </p>
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	https://treatment.plazi.org/id/57D17ECD30875951860B4ABBBE4CEAE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
7250C5D625965E3198E4134C4285D58C.text	7250C5D625965E3198E4134C4285D58C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus vigintistriatus Fairmaire 1869	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copelatus vigintistriatus Fairmaire, 1869</p>
            <p>Type locality.</p>
            <p>Mayotte.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-44B; 2 ♂♂, 2 ♀♀: MAK-44C; 1 ♀: MAK-56; 2 ♀♀: MAK-60.</p>
            <p>Distribution.</p>
            <p>Madagascar (widespread), Mayotte (Guignot 1959-1961; Bertrand and Legros 1971).</p>
            <p>Habitat in study area.</p>
            <p>This species has been captured in a few inner massif sites all situated in northern Makay: a puddle, a blind channel, a small stream (these sites in forest) and an open marsh. These habitats had slightly turbid water and a mineral bottom ranging from clay to sand with moderate quantity of plant debris, and no vegetation.</p>
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	https://treatment.plazi.org/id/7250C5D625965E3198E4134C4285D58C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
DBD337F8D15C574DA2B62586C506AB51.text	DBD337F8D15C574DA2B62586C506AB51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus zanabato Manuel & Ramahandrison 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copelatus zanabato Manuel &amp; Ramahandrison sp. nov.</p>
            <p>Figs 3B, 4E-H</p>
            <p>Type locality.</p>
            <p>Madagascar, Toliara province, Malaimbandy municipality, Makay massif (northern part), ca. 21 km W of Tsimazava, ca. 20°42'S, 45°30'E, altitude ca. 430 m a.s.l.</p>
            <p>Type material.</p>
            <p> Holotype ♂: "Madagascar. Ex-prov. Toliara / Makay massif, ca. 21 km / W Tsimazava / 20°42'10"S, 45°30'18"E [pr.] // 12.IV.2018. Ramahandrison leg. / Alt. 433 m. Pool in a / very narrow and dark / canyon. [pr.] // Holotype /  Copelatus zanabato sp. nov. / Manuel &amp; Ramahandrison, 2022 [red, pr.]" (MNHN). Paratypes: 1 ♀: same as holotype. 1 ♂: "Madagascar. Ex-prov. Toliara / Makay massif, ca. 20 km / WSW Tsimazava / 20°43'54"S, 45°31'10"E / 14.IV.2018. Ramahandrison leg. [pr.] // Alt. 437 m. Small / pond in a canyon at / Ampasimaiky [pr.]" (CMM). Both paratypes with respective red label. </p>
            <p>Diagnosis.</p>
            <p> This species belongs to the  Copelatus erichsonii -group. It is externally rather similar to  C. acamas , from which it differs by distinctly smaller size; habitus narrower with sides more parallel, dorsally much flatter; discal stria IX on elytron more strongly abbreviated anteriorly; strioles on pronotum surface sparser and much more weakly impressed; shape of median lobe of aedeagus very different. Among  Copelatus species known from Madagascar, the aedeagus of  C. zanabato sp. nov. is most similar to that of  C. andobonicus . From the latter, the new species differs by: habitus narrower with sides more parallel, dorsally much flatter; pronotum paler and elytra with darker linear colouration following the striae much less contrasted with respect to paler background; strioles on pronotum surface denser, present on whole surface (in  C. andobonicus almost without striae in anterior disk region); median lobe of aedeagus in lateral view with broad flat protuberance on ventral side ca. halfway between base and apex (in  C. andobonicus with much smaller protuberance at ca. basal third) and apical third much broader, in ventral view with apical region much broader and evenly narrowed, twisted on the left farther from apex. </p>
            <p>Description of holotype.</p>
            <p>Body shape elongate oval, with sides subparallel (Fig. 3B), dorsally weakly convex. Pronotum sides evenly curved and converging from posterior angle. Ratio between maximum width of pronotum and maximum body width ~ 0.90. Head uniformly light rufo-testaceous. Ratio between interocular distance and maximum width of head ~ 0.68. Pronotum medially rufous, laterally colour becoming gradually rufo-testaceous. Elytra light chestnut brown, with darker linear colouration along striae (Fig. 3B).</p>
            <p>Elytra with ten well-impressed discal and one submarginal striae. Stria IX abbreviated anteriorly. Striae I and II diverging anteriorly. Submarginal stria starting slightly before elytron midlength, fragmented anteriorly. Head, pronotum and elytra with fine reticulation and fine punctation. Whole surface of pronotum with rather dense, short and fine strioles, in medial disk region strioles even finer.</p>
            <p>Ventral side uniformly rufo-testaceous. Metacoxal plates with moderately impressed short strioles; visible abdominal ventrites I-III with denser and longer very fine strioles. Prosternal process short and broad, with rounded apex. Metacoxal lines short, ending anteriorly at large distance from posterior margin of metaventrite, moderately diverging anteriorly.</p>
            <p>Appendages: Antennae, palps, forelegs and midlegs testaceous, hindlegs rufo-testaceous. First three pro- and mesotarsomeres widened and ventrally equipped with suction cups; number of suction cups per article (I-III) 7:4:4 on both pro- and mesotarsus. Protibia at base shortly narrow, with shallow protuberance along ventral margin, distally broadened. Pro- and mesotarsal claws unmodified.</p>
            <p>Median lobe and parameres as in Fig. 4E-H.</p>
            <p>Female. Strioles on pronotum surface denser. Pro- and mesotarsomeres and protibia unmodified.</p>
            <p>Measurements.</p>
            <p>Holotype: TL 6.35 mm, TL without head 5.7 mm, MW 2.9 mm, ratio TL/MW 2.20. Paratypes: TL 6.55-6.8 mm, TL without head 5.9-6.1 mm, MW 3.0-3.1 mm, ratio TL/MW 2.17.</p>
            <p>Variation.</p>
            <p>In the male paratype, strioles on pronotum are longer and more deeply impressed than in the holotype, and the metacoxal lines are slightly longer. In the female paratype, the elytral stria V is slightly abbreviated anteriorly.</p>
            <p>Etymology.</p>
            <p>The species name literally means "son of the rock" in Malagasy. It is an invariable name standing in apposition.</p>
            <p>Distribution.</p>
            <p>So far known only from northern Makay in Madagascar.</p>
            <p>Habitat.</p>
            <p> This species was collected at two sites located in two nearby canyons in northern inner Makay. Two specimens were sampled at site MAK-46, an isolated pool (~ 1 m  × 3 m) on the bottom of a narrow and dark canyon, and one specimen at site MAK-50, a stagnant temporary pond (~ 3 m  × 7 m) situated in a wider canyon and in a more open environment. Both habitats were characterised by sandy bottom with some plant debris, absence of visible inflow / outflow, somewhat turbid water and no vegetation. Other species of aquatic  Adephaga (all  Dytiscidae ) sampled at the same sites:  Copelatus acamas ,  C. ruficapillus ,  Cybister operosus Sharp, 1882,  Hydaticus sobrinus Aubé , 1838,  Hyphydrus separandus Régimbart , 1895,  Laccophilus makay ,  Pachynectes sp. Ma1, and  P. sp. Ma4. </p>
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	https://treatment.plazi.org/id/DBD337F8D15C574DA2B62586C506AB51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
970BF0489D895243BB0DD8AA02D45FC5.text	970BF0489D895243BB0DD8AA02D45FC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister cinctus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Cybister cinctus Sharp, 1882</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p> 1 ex.: MAK-1A; 1 ♂, 1 ♂: MAK-2; 1 ♂, 3 ♀♀: MAK-19; 4 ♂♂, 1  еx .: MAK-20; 1 ♀: MAK-23; 1 ♀: MAK-41. </p>
            <p>Distribution.</p>
            <p>Madagascar (widespread and common) (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984; Rocchi 1991; Bukontaite et al. 2015).</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species was collected only at peripheral sites, in various kinds of water bodies (lentic or slowly flowing) in open areas. It prefers habitats with at least some vegetation and is tolerant to anthropogenic pressure.</p>
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	https://treatment.plazi.org/id/970BF0489D895243BB0DD8AA02D45FC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
04747A9AEF8451F39CBE78770B59079E.text	04747A9AEF8451F39CBE78770B59079E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister operosus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Cybister operosus Sharp, 1882</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-50.</p>
            <p>Distribution.</p>
            <p>Madagascar (widespread but localised) (Guignot 1959-1961; Bukontaite et al. 2015).</p>
            <p>Habitat in study area.</p>
            <p>This species was captured only once, in northern inner Makay, in a large temporary pond located in a shallow open area on sand. This single capture does not reflect the usual habitat preferences of this species. According to our observations in Isalo and Ankarafantsika, this is a lotic species (an exceptional ecology for the genus) inhabiting the margins of streams and rivers with some vegetation and / or tree roots and / or plant debris, in well-preserved forested or semi-forested environments.</p>
            <p> Subfamily  Dytiscinae , tribe  Aciliini</p>
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	https://treatment.plazi.org/id/04747A9AEF8451F39CBE78770B59079E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
C694E79245DE5292855DFC6797AF2DDC.text	C694E79245DE5292855DFC6797AF2DDC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dineutus proximus Aube 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Dineutus proximus 
Aube
, 1838
</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂, 2 ♀♀: MAK-5B; 1 ♂, 1 ♀: MAK-5D; 2 ♂♂: MAK-13; 2 ♂♂, 3 ♀♀: MAK-14B; 2 ♂♂, 1 ♀: MAK _24; 3 ♂♂, 8 ♀ ♀: MAK-27; 1 ♀: MAK-30; 1 ♀: MAK-35C; 2 ♂♂, 2 ♀♀: MAK-37B; 2 ♀♀: MAK-40A; 2 ♂♂, 3 ♀♀: MAK-40B; 3 ♂♂, 3 ♀♀: MAK-48; 1 ♂, 2 ♀♀: MAK-55; 1 ♂, 1 ♀: MAK-59A;; 1 ♂, 3 ♀♀: MAK-59C.</p>
            <p>Distribution.</p>
            <p>Madagascar, widespread (Legros 1951; Brinck 1955; Bameul 1984).</p>
            <p>Habitat in study area.</p>
            <p>Collected only in inner Makay, in permanent lotic habitats (rivers and streams) with sandy bottom (in a few sites substrate was more rocky) and with clear water, in forested or semi-forested environmental context, with little or no anthropogenic disturbance.</p>
        </div>
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	https://treatment.plazi.org/id/C694E79245DE5292855DFC6797AF2DDC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
F678DC1E84F1534EA9E00CB26991E5BF.text	F678DC1E84F1534EA9E00CB26991E5BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dineutus sinuosipennis subsp. sinuosipennis Castelnau 1840	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Dineutus sinuosipennis sinuosipennis Castelnau, 1840</p>
            <p> Dineutus bidens =  D. bidens Vollenhoven, 1869;  D. denticulatus Régimbart , 1882. </p>
            <p>Type locality.</p>
            <p>Tibet (erroneous locality?).</p>
            <p>Material examined.</p>
            <p>1 ♂, 4 ♀♀: MAK-27; 3 ♂♂, 4 ♀♀: MAK-37B; 1 ♂, 1 ♀: MAK-40A; 1 ♂: MAK-40B; 1 ♀: MAK-48; 1 ♂, 5 ♀♀: MAK-52; 5 ♂♂, 9 ♀♀: MAK-55; 1 ♀: MAK-58.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Legros 1951; Brinck 1955; Bameul 1984). Another subspecies,  D. sinuosipennis comorensis Régimbart , 1892, is present in the Comoro archipelago (Brinck 1955; Wewalka 1980). </p>
            <p>Habitat in study area.</p>
            <p> Same as  D. proximus (both species often syntopic). This species is less abundant than  D. proximus in the Makay massif. </p>
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	https://treatment.plazi.org/id/F678DC1E84F1534EA9E00CB26991E5BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
4EF79FFA02295A3F8987AEBEF0A7CE01.text	4EF79FFA02295A3F8987AEBEF0A7CE01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eretes griseus (Fabricius 1781)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Eretes griseus (Fabricius, 1781)</p>
            <p> Eretes plicipennis =  E. plicipennis (Motschulsky, 1845);  E. succinctus (Klug, 1834). </p>
            <p>Type locality.</p>
            <p>India.</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-11B.</p>
            <p>Distribution.</p>
            <p>Southern half of the Palearctic region, Africa, Oriental region (Guignot 1959-1961; Miller 2002). In Madagascar, widespread.</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2P). This species was collected at a single site in central inner Makay, in an isolated shallow temporary puddle located in the middle of a large flat accumulation of sand in the outer part of river meander. The environment was totally open with no trees, the water was turbid and there was no vegetation. These habitat characteristics are very representative of the ecology of members of the genus  Eretes everywhere in the world (Miller 2002). </p>
            <p> Tribe  Hydaticini</p>
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	https://treatment.plazi.org/id/4EF79FFA02295A3F8987AEBEF0A7CE01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
9D6C4C3E08FE5BAC9EA6FAE4AED12CDF.text	9D6C4C3E08FE5BAC9EA6FAE4AED12CDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydaticus dorsiger Aube 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Hydaticus dorsiger 
Aube
, 1838
</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-11A; 1 ♀: MAK-11B; 1 ♂, 1 ♀: MAK-12A; 1 ♂: MAK-14A; 1 ♂: MAK-18; 8 ♂♂, 6 ♀♀: MAK-19; 1 ♂: MAK-23.</p>
            <p>Distribution.</p>
            <p> Whole tropical Africa to Arabia, Madagascar (where it is widespread and common) (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984; Rocchi 1991;  Hájek and Reiter 2014; Bukontaite et al. 2015). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2N-P). This species has been observed mainly at peripheral sites and in open areas, in lentic environments that were permanent or temporary, with or without slow water flow, with bottom often muddy, and with or without marginal vegetation. This species is tolerant to anthropogenic disturbance.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/9D6C4C3E08FE5BAC9EA6FAE4AED12CDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
D4413859D1245668B492926123E7AB90.text	D4413859D1245668B492926123E7AB90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydaticus exclamationis Aubé 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Hydaticus exclamationis 
Aube
, 1838
</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-11B.</p>
            <p>Distribution.</p>
            <p>Sub-Saharan Africa, Mauritius, Madagascar (Guignot 1959-1961; Bameul 1984; Bukontaite et al. 2015). In Madagascar, widespread and common, especially in lowlands.</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2P). This species was collected at a single site in south-central inner Makay (see description of the habitat above under  Eretes griseus ). It usually prefers lentic or slowly flowing habitats with at least some vegetation. </p>
        </div>
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</html>
	https://treatment.plazi.org/id/D4413859D1245668B492926123E7AB90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
FAD911BCBCBA561F9F1E800B170F82CE.text	FAD911BCBCBA561F9F1E800B170F82CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydaticus petitii Aube 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Hydaticus petitii 
Aube
, 1838
</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-52.</p>
            <p>Distribution.</p>
            <p>Madagascar, widespread (Guignot 1959-1961; Bukontaite et al. 2015).</p>
            <p>Habitat in study area.</p>
            <p> This species was collected once, in northern inner Makay, in a calm pool (~ 3 m  × 7 m) on the bottom of a canyon, with inflow and outflow from the nearby Ampasimaiky River. This pool was rather deep (&gt; 1 m), with bottom of sand covered with a thin layer of clay, almost without vegetal detritus, with moderately turbid water and no vegetation. The environment was semi-forested and the pond was surrounded by  Ravenea palm trees. As a rule this Malagasy endemic species is encountered in forest massifs in more or less undisturbed habitats. </p>
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	https://treatment.plazi.org/id/FAD911BCBCBA561F9F1E800B170F82CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
C64E289494575290A4271FBA2FDB2B2D.text	C64E289494575290A4271FBA2FDB2B2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydaticus servillianus Aube 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Hydaticus servillianus 
Aube
, 1838
</p>
            <p> Hydaticus discoidalis =  H. discoidalis Hope, 1843;  H. flavomarginatus Zimmermann, 1920). </p>
            <p>Type locality.</p>
            <p>South Africa, Western Cape, Cape of Good Hope.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-1A; 1 ♂: MAK-2; 13 ♂♂, 7 ♀♀: MAK-19; 1 ♂, 3 ♀♀: MAK-23; 2 ♂♂: MAK-40A; 1 ♂, 6 ♀♀: MAK-61; 1 ♂: MAK-62.</p>
            <p>Distribution.</p>
            <p> Sub-Saharan Africa, Madagascar (Guignot 1959-1961; Bameul 1984;  Hájek and Reiter 2014; Bukontaite et al. 2015). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species has been found in peripheral sites and at one inner massif site, in lentic and slowly flowing lotic habitats, mainly in open areas. The bottom comprised various amounts of sand and clay/mud and moderate to abundant plant debris. The water was clear to moderately turbid. The vegetation was variously developed. This eurytopic species is tolerant to anthropogenic perturbation.</p>
        </div>
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	https://treatment.plazi.org/id/C64E289494575290A4271FBA2FDB2B2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
10EFC99DF03056369E570F45361FB166.text	10EFC99DF03056369E570F45361FB166.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydaticus sobrinus Aube 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Hydaticus sobrinus 
Aube
, 1838
</p>
            <p> Hydaticus matruelis obliquevittatus =  Hydaticus matruelis var. obliquevittatus Régimbart , 1895). </p>
            <p>Type locality.</p>
            <p>Madagascar, Mascarene Islands (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984; Bukontaite et al. 2015).</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK- 8; 2 ♀: MAK-50; 2 ♀♀: MAK-53; 1 ♂: MAK-54A; 2 ♀♀: MAK-59C.</p>
            <p>Distribution.</p>
            <p> Mauritius, La  Réunion , Madagascar. In Madagascar, widespread. </p>
            <p>Habitat in study area.</p>
            <p>This species was collected only in inner Makay, in well-preserved forested or semi-forested areas. The habitats were isolated pools and very slowly flowing streams, with clear or slightly turbid water, sandy bottom (sometimes with stones), with moderate amount of plant debris and no vegetation.</p>
            <p> Subfamily  Hydroporinae , tribe  Bidessini</p>
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	https://treatment.plazi.org/id/10EFC99DF03056369E570F45361FB166	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
2978C8725947596A80975C7677064E6E.text	2978C8725947596A80975C7677064E6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydroglyphus capitatus (Regimbart 1895)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Hydroglyphus capitatus (
Regimbart
, 1895)
</p>
            <p> Hydroglyphus longivittis =  H. longivittis Régimbart , 1903. </p>
            <p>Type locality.</p>
            <p>Madagascar, Antsiranana.</p>
            <p>Material examined.</p>
            <p>13 ♂♂, 20 ♀: MAK-1A; 1 ♂, 1 ♀: MAK-4; 5 ♂♂: MAK-61.</p>
            <p>Distribution.</p>
            <p> Seychelles, Madagascar (Guignot 1959-1961;  Biström 1986; Rocchi 1991). In Madagascar, widespread and common particularly in lowlands. </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2E). This species was captured at two peripheral and at one inner massif sites, in highly contrasted habitats, including shallow puddles on the sandy banks of the Mangoky River, a shallow pond with clay-mud bottom and marginal helophytes, and (for the inner Makay site) a marginal spring on the bank of a river, full of orange masses of iron bacteria. The environment was open and strongly impacted by human activities in the two peripheral sites (where the species was more abundant); semi-forested and rather well preserved in the inner massif site.</p>
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	https://treatment.plazi.org/id/2978C8725947596A80975C7677064E6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
DB0D9FC7C9CF5BF1961404D6F65A8487.text	DB0D9FC7C9CF5BF1961404D6F65A8487.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydroglyphus geminodes (Regimbart 1895)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Hydroglyphus geminodes (
Regimbart
, 1895)
</p>
            <p> Hydroglyphus africanus =  H. africanus Régimbart , 1895. </p>
            <p>Type locality.</p>
            <p>Madagascar, Antsiranana.</p>
            <p>Material examined.</p>
            <p>6 ♂♂, 3 ♀: MAK-1A; 1 ♂, 2 ♀: MAK-2; 6 ♂♂, 3 ♀♀: MAK-4; 2 ♂♂, 10 ♀♀: MAK-17; 1 ♂: MAK-18; 4 ♂♂, 3 ♀♀: MAK-19; 4 ♂♂, 1 ♀: MAK-61.</p>
            <p>Distribution.</p>
            <p> Sub-Saharan Africa, Mauritius, La  Réunion , Madagascar (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984;  Biström 1986; Rocchi 1991). In Madagascar, widespread and common. </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C, E). This species has been found mainly at peripheral sites in open areas, at shallow depth in various kinds of water bodies (temporary or permanent lentic habitats with water stagnant or very slowly flowing). The water was generally clear and the bottom consisted of sand and/or clay, with some plant debris. There was either no vegetation or sparse helophytes and at some sites presence of filamentous green algae. This species is tolerant to anthropogenic disturbance.</p>
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	https://treatment.plazi.org/id/DB0D9FC7C9CF5BF1961404D6F65A8487	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
46B8072081F85E9C8EA81ED7D63CEC68.text	46B8072081F85E9C8EA81ED7D63CEC68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydroglyphus plagiatus (H. J. Kolbe 1883)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydroglyphus plagiatus (H. J. Kolbe, 1883)</p>
            <p>Type locality.</p>
            <p>Eastern part of Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-17.</p>
            <p>Distribution.</p>
            <p> Madagascar, common in the Central Highlands (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984,  Biström 1986; Rocchi 1991). </p>
            <p>Habitat in study area.</p>
            <p>This species seems very rare in the Makay since only one specimen was found, in an inner massif site located in a semi-forested environment. The habitat was a small, isolated, sun-exposed puddle on rock mass, on the bank of a river. The bottom consisted of sandstone, sand and clay without organic debris, the water was clear and there was no vegetation.</p>
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	https://treatment.plazi.org/id/46B8072081F85E9C8EA81ED7D63CEC68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
12E8DC35C5C4564599B80B9EA73E755B.text	12E8DC35C5C4564599B80B9EA73E755B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus acuminatus Motschulsky 1860	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus acuminatus Motschulsky, 1860</p>
            <p> Hydrovatus affinis =  H. affinis Régimbart , 1895;  H. badius (Clark, 1863);  H. consanguineus Régimbart , 1880;  H. ferrugineus Zimmermann, 1919;  H. humilis Sharp, 1882;  H. malaccae (Clark, 1863);  H. obscurus Motschulsky, 1860;  H. obscurus Régimbart , 1895;  H. sordidus Sharp, 1882. </p>
            <p>Type locality.</p>
            <p>South-East Asia (Indian continent).</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-1A; 6 ♂♂, 6 ♀♀: MAK-2; 1 ♀: MAK-3; 2 ♂♂, 1 ♀: MAK-19; 1 ♂, 2 ♀♀: MAK-21; 1 ♂: MAK-23; 1 ♀: MAK-44C; 1 ♂, 2 ♀♀: MAK-60; 1 ♂, 1 ♀: MAK-61; 1 ♂: MAK-62.</p>
            <p>Distribution.</p>
            <p> Sub-Saharan Africa, Madagascar, Seychelles, Turkey, Egypt, Arabian Peninsula, south-eastern Palearctic region from India to south Japan, Oriental region (  Biström 1996;  Hájek and Reiter 2014). In Madagascar, widespread and common in lowlands (absent from the Central Highlands). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B-D). This species is present in lentic and in slowly flowing lotic habitats. It was collected both at peripheral and inner massif sites. The bottom varied from clayey to sandy, with clear, red-brown or turbid water and with more or less abundant plant debris. This species has a clear preference for open environments and habitats with at least some vegetation and is highly tolerant to anthropogenic disturbance.</p>
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	https://treatment.plazi.org/id/12E8DC35C5C4564599B80B9EA73E755B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
F3DA6FBEC9A65B98AADEC3FF17226D44.text	F3DA6FBEC9A65B98AADEC3FF17226D44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus capnius Guignot 1950	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus capnius Guignot, 1950</p>
            <p>Type locality.</p>
            <p>Zambia (Congo Belge), Musosa.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2; 1 ♂, 1 ♀: MAK-42.</p>
            <p>Distribution.</p>
            <p> Sudan, Democratic Republic of the Congo, Madagascar (Guignot 1959-1961;  Biström 1996). Distribution within Madagascar poorly known, but probably restricted to lowlands. </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was found at only two peripheral sites, in open and non-forested environments, both markedly eutrophic. The first site was a shallow isolated stream characterised by very weak water flow, sandy bottom, marked anthropic disturbance (cattle trampling), sparse tufts of small  Cyperaceae and strong presence of filamentous green algae. The second site was an open marsh, relatively preserved from anthropogenic pressure, without water flow, with sandy bottom and moderate quantity of plant debris, moderately turbid water, sparse helophytes (  Cyperus ) and aquatic plants (  Nymphaea and  Polygonum ), floating ferns (  Azolla , an indicator of eutrophication) and moderate abundance of filamentous green algae. </p>
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	https://treatment.plazi.org/id/F3DA6FBEC9A65B98AADEC3FF17226D44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
215BFFDAF4E65F2BBDF71CF5EA762FCE.text	215BFFDAF4E65F2BBDF71CF5EA762FCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus crassicornis (H. J. Kolbe 1883)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus crassicornis (H. J. Kolbe, 1883)</p>
            <p>Type locality.</p>
            <p>Eastern part of Madagascar</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-42; 1 ♂: MAK-60; 1 ♂, 1 ♀: MAK-61.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1959-1961; Bertrand and Legros 1971;  Biström 1996). </p>
            <p>Habitat in study area.</p>
            <p>This species was collected at two peripheral and one inner massif sites. The habitats were located in open areas and were lentic water bodies (with or without slow water circulation) with sand-clay bottom and with plant debris, with water clear to moderately turbid, with discontinuous marginal belts of helophytes and at one site with filamentous green algae.</p>
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	https://treatment.plazi.org/id/215BFFDAF4E65F2BBDF71CF5EA762FCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
72C4C67510EF588CA29BC023ADBD671D.text	72C4C67510EF588CA29BC023ADBD671D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus cruentatus H. J. Kolbe 1883	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus cruentatus H. J. Kolbe, 1883</p>
            <p>Type locality.</p>
            <p>Eastern part of Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2; 1 ♀: MAK-42.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1959-1961;  Biström 1996). </p>
            <p>Habitat in study area.</p>
            <p> Same as  Hydrovatus capnius (see above). </p>
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	https://treatment.plazi.org/id/72C4C67510EF588CA29BC023ADBD671D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
B33BB39611B05624A29C7753FCC7D33C.text	B33BB39611B05624A29C7753FCC7D33C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus dentatus Bilardo & Rocchi 1990	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus dentatus Bilardo &amp; Rocchi, 1990</p>
            <p>Type locality.</p>
            <p>Zambia, Luangwa valley, Chibembe.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2.</p>
            <p>Distribution.</p>
            <p> Zambia, South Africa (  Biström 1996), Madagascar. For Madagascar, the first record is recent (Ranarilalatiana 2019) and was from Anjozorobe-Angavo in the Central Highlands. </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was sampled only once in a peripheral site. The habitat was a shallow insolated stream characterised by very weak water flow, sandy bottom, marked anthropic disturbance (cattle trampling), sparse tufts of small  Cyperaceae and strong presence of filamentous green algae. </p>
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	https://treatment.plazi.org/id/B33BB39611B05624A29C7753FCC7D33C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
407D478E96625322B13AFE3CCA3CCD16.text	407D478E96625322B13AFE3CCA3CCD16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus otiosus Guignot 1945	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus otiosus Guignot, 1945</p>
            <p>Type locality.</p>
            <p>Madagascar, Antananarivo, Ikopa River.</p>
            <p>Material examined.</p>
            <p>3 ♂♂, 2 ♀♀: MAK-2; 3 ♂♂, 1 ♀: MAK-62.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1959-1961; Bertrand and Legros 1971;  Biström 1996); common in the Central Highlands. </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was collected at only two peripheral sampling sites. The first site was the one described above for  H. dentatus . The second one was a canal at the edge of rice fields, with water slowly flowing, muddy bottom, water rather turbid, and without vegetation. Both sites were significantly impacted by anthropogenic disturbance. </p>
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	https://treatment.plazi.org/id/407D478E96625322B13AFE3CCA3CCD16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
6B4DE4274F2858D19DBC5790C5A7CC56.text	6B4DE4274F2858D19DBC5790C5A7CC56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus parvulus Régimbart 1900	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Hydrovatus parvulus 
Regimbart
, 1900
</p>
            <p> Hydrovatus noctivagus =  H. noctivagus Guignot, 1953;  H. ocnerus Guignot, 1958;  H. socors Guignot, 1954. </p>
            <p>Type locality.</p>
            <p>Madagascar, Antongil Bay.</p>
            <p>Material examined.</p>
            <p>2 ♂♂: MAK-2; 2 exs.: MAK-43; 3 exs.: MAK-61.</p>
            <p>Distribution.</p>
            <p> Sub-Saharan Africa, Madagascar (Guignot 1959-1961;  Biström 1996). In Madagascar, widespread and common. </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species was collected at three peripheral sampling sites, in open, non-forested environments. Two of the sites were lotic habitats (a shallow stream and the calm margin of a river) and one was lentic (a shallow pond ~ 25 m in diameter). In all cases, the bottom consisted of sand and clay, the water was clear, and the marginal zone where the beetles were collected was vegetated.</p>
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	https://treatment.plazi.org/id/6B4DE4274F2858D19DBC5790C5A7CC56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
517CA2364733571194697E15FD622F23.text	517CA2364733571194697E15FD622F23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus pictulus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus pictulus Sharp, 1882</p>
            <p> H. dilutus H. J. Kolbe, 1883;  H. scymnoides Régimbart , 1895. </p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>5 ♂♂, 1 ♀: MAK-2; 1 ♂: MAK-62.</p>
            <p>Distribution.</p>
            <p> Sub-Saharan Africa, Madagascar (Guignot 1959-1961;  Biström 1996). In Madagascar, widespread. </p>
            <p>Habitat in study area.</p>
            <p> Same as  Hydrovatus otiosus (see above). Throughout Madagascar, this species has an ecological optimum in the calm parts of rivers and streams or their satellite puddles and pools, with sandy or muddy bottom and few or no vegetation and is quite tolerant to anthropogenic disturbance. </p>
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	https://treatment.plazi.org/id/517CA2364733571194697E15FD622F23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
FD7EFCADC66E5335BE13072CFCD77A0E.text	FD7EFCADC66E5335BE13072CFCD77A0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus sp. Ma 7	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hydrovatus sp. Ma7</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-2.</p>
            <p>Note.</p>
            <p> This large species may be  H. confusus Régimbart , 1903, a species endemic to Madagascar, or  H. badeni Sharp, 1882, also present in continental Sub-Saharan Africa (  Biström 1996). </p>
            <p>Distribution.</p>
            <p>Unknown.</p>
            <p>Habitat in study area.</p>
            <p> Same as  Hydrovatus dentatus (see above) </p>
            <p> Tribe  Hyphydrini</p>
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	https://treatment.plazi.org/id/FD7EFCADC66E5335BE13072CFCD77A0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
EDDE447E959557E2AF3561562E242B6E.text	EDDE447E959557E2AF3561562E242B6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrovatus testudinarius Régimbart 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Hydrovatus testudinarius 
Regimbart
, 1895
</p>
            <p>Type locality.</p>
            <p>Madagascar, Antananarivo, Ambodinandohalo Lake.</p>
            <p>Material examined.</p>
            <p>7 ♂♂, 3 ♀♀: MAK-2.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread but not common (Guignot 1959-1961; Rocchi 1991;  Biström 1996). </p>
            <p>Habitat in study area.</p>
            <p> Same as  Hydrovatus dentatus (see above). </p>
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	https://treatment.plazi.org/id/EDDE447E959557E2AF3561562E242B6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
43165BFEDA9F5F5B8A68C0607CB0F45D.text	43165BFEDA9F5F5B8A68C0607CB0F45D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyphydrus separandus Regimbart 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Hyphydrus separandus 
Regimbart
, 1895
</p>
            <p> Hydrovatus oncodes =  H. oncodes Guignot, 1955. </p>
            <p>Type locality.</p>
            <p> Madagascar, Montagne  d’Ambre , Ambohitra National Park. </p>
            <p>Material examined.</p>
            <p>2 ♂♂, 8 ♀♀: MAK-3; 2 ♀♀: MAK-4; 30 exs.: MAK-5A; 8 exs.: MAK-5B; 15 exs.: MAK-5C; 85 exs.: MAK-5D; 5 ♂♂, 6 ♀♀: MAK-8; 1 ♀: MAK-10; 1 ♀: MAK-11B; 3 ♂♂, 5 ♀♀: MAK-14A; 1 ♀: MAK-16; 1 ♂, 1 ♀: MAK-25B; 4 exs.: MAK-28; 14 exs.: MAK-29; 1 ♂: MAK-35B; 1 ♂, 1 ♀: MAK-37A; 5 ♂♂: MAK-40A; 1 ex.: MAK-40B; 1 ♂: MAK-50; 2 ♂♂, 1 ♀: MAK-52; 1 ♂: MAK-59C.</p>
            <p>Distribution.</p>
            <p> Comoro Islands, Madagascar (Guignot 1959-1961; Wewalka 1980;  Biström 1982). In Madagascar, widespread. </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2D-F, I, L-N, P). This species was collected only in inner Makay where it is one of the most common species of aquatic  Adephaga . It was found in puddles, pools, ponds and slowly flowing streams, from a few centimetres to 150 cm in depth, with sandy or rocky bottom, clear water and more or less abundant plant debris, often without vegetation. Almost all collecting sites were located in forested or semi-forested environment. </p>
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	https://treatment.plazi.org/id/43165BFEDA9F5F5B8A68C0607CB0F45D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
BD3164EB861C58FCB91F40901497BCD4.text	BD3164EB861C58FCB91F40901497BCD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyphydrus stipes Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hyphydrus stipes Sharp, 1882</p>
            <p> Hydrovatus soarezicus =  H. soarezicus Alluaud, 1897. </p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-29.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (  Biström 1982; Rocchi 1991). </p>
            <p>Habitat in study area.</p>
            <p>This species was collected at a single site located in inner Makay, in a semi-forested small valley. The habitat was a sun-exposed isolated pool on sandstone rock, rather deep (70 cm), with bottom made up of sand and stones with a moderate quantity of plant debris, with clear water and no vegetation. This site was well preserved from anthropogenic disturbance.</p>
            <p> Tribe  Methlini</p>
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	https://treatment.plazi.org/id/BD3164EB861C58FCB91F40901497BCD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
BCD071B2A28E5785AC0B73522639A168.text	BCD071B2A28E5785AC0B73522639A168.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus addendus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laccophilus addendus Sharp, 1882</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-2; 1 ♀: MAK-5A; 28 ♂♂, 19 ♀♀: MAK-11A; 1 ♀: MAK-11B; 2 ♂♂, 1 ♀: MAK-19; 2 ♂♂, 1 ♀: MAK-25A; 1 ♂: MAK-26; 1 ♂: MAK-38A; 1 ♂, 1 ♀: MAK-62.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (Guignot 1959-1961; Bameul 1984; Rocchi 1991;  Biström et al. 2015). Previous records from outside Madagascar are considered uncertain by  Biström et al. (2015). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C, F, O, P). This species was collected at peripheral and inner massif sites, mainly in non-forested areas, in lentic or very slow flowing lotic habitats, most often sun-exposed but at a few sites partly shaded by trees. The bottom consisted of various amounts of sand, clay and mud, with moderate to abundant plant debris, and marginal vegetation was absent to well developed. The water was clear to rather turbid. This species occurs both in strongly anthropised contexts and in habitats preserved from anthropogenic pressure.</p>
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	https://treatment.plazi.org/id/BCD071B2A28E5785AC0B73522639A168	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
D6B6AE2BBC6C5DBFA2D2801DBFB1F78B.text	D6B6AE2BBC6C5DBFA2D2801DBFB1F78B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus flaveolus Regimbart 1906	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Laccophilus flaveolus 
Regimbart
, 1906
</p>
            <p> Laccophilus pampinatus =  L. pampinatus Guignot, 1941. </p>
            <p>Type locality.</p>
            <p>Kenya, Winam, Kavirondo Bay.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2.</p>
            <p>Distribution.</p>
            <p> Eastern Sub-Saharan Africa, Madagascar (Guignot 1959-1961; Bameul 1984;  Biström et al. 2015). </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was sampled only once at a peripheral site. The habitat was a shallow insolated stream characterised by very weak water flow, sandy bottom, marked anthropic disturbance (cattle trampling), sparse tufts of small  Cyperaceae and strong presence of filamentous green algae. </p>
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	https://treatment.plazi.org/id/D6B6AE2BBC6C5DBFA2D2801DBFB1F78B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
59F6BE32D147525C8F99CE88AFD08254.text	59F6BE32D147525C8F99CE88AFD08254.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus insularum Bistroem, Nilsson & Bergsten 2015	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Laccophilus insularum 
Bistroem
, Nilsson &amp; Bergsten, 2015
</p>
            <p>Type locality.</p>
            <p>Madagascar, Ankarafantsika National Park, Mahajanga, Boeny.</p>
            <p>Material examined.</p>
            <p>2 ♂♂, 2 ♀♀: MAK-5A.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread (  Biström et al. 2015). </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2F). This species was encountered only once, in south-central inner Makay. The habitat was a deep pond (dimensions about 30 m  × 15 m) at the bottom of a small canyon along the course of a stream. This water body was formed as a result of the collapse of downstream canyon walls (natural dam). It was shaded by the canyon walls and surrounded by trees (notably  Cyathea tree ferns) and hygrophilous vegetation. The habitat was furthermore characterised by clay bottom, with moderate amount of plant debris, clear water, and absence of marginal vegetation. This site was unaffected by anthropic disturbance. </p>
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	https://treatment.plazi.org/id/59F6BE32D147525C8F99CE88AFD08254	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
F7DAADE941DF5F1BB5C3F2EC89A8DC6A.text	F7DAADE941DF5F1BB5C3F2EC89A8DC6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus luctuosus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laccophilus luctuosus Sharp, 1882</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-21.</p>
            <p>Distribution.</p>
            <p> Madagascar (widespread in lowlands) (Guignot 1959-1961; Bertrand and Legros 1971; Rocchi 1991;  Biström et al. 2015). </p>
            <p>Habitat in study area.</p>
            <p> This species was encountered only once in a peripheral site located in an open area. The habitat was a small isolated temporary puddle with sandy bottom and clear water under  Phragmites , on the west bank of the Makaikely River. There was a moderate amount of plant debris, and no vegetation. </p>
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	https://treatment.plazi.org/id/F7DAADE941DF5F1BB5C3F2EC89A8DC6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
ACDD693E6E1B50E3BE82FCB09EECE56C.text	ACDD693E6E1B50E3BE82FCB09EECE56C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus makay Manuel & Ramahandrison 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laccophilus makay Manuel &amp; Ramahandrison, 2020</p>
            <p>Type locality.</p>
            <p>Madagascar, Toliara, Makay massif, 10.7 km NW of Tsivoky.</p>
            <p>Material examined.</p>
            <p>8 ♂♂, 7 ♀♀: MAK-3; 2 ♂♂, 1 ♀: MAK-4; 4 ♂♂, 1 ♀: MAK-5A; 3 ♂♂: MAK-5B; 2 ♂♂: MAK-5C; 7 ♂♂, 5 ♀♀: MAK-5D; 1 ♂, 2 ♀♀: MAK-6; 1 ♂, 1 ♀: MAK-7; 11 ♂♂, 9 ♀♀: MAK-8; 2 ♂♂, 4 ♀♀: MAK-10; 13 ♂♂, 6 ♀♀: MAK-14A; 3 ♂♂, 2 ♀♀: MAK-15; 1 ♂, 1 ♀: MAK-16; 1 ♂, 1 ♀: MAK-25A; 9 ♂♂, 22 ♀♀: MAK-25B; 1 ♂, 1 ♀: MAK-26; 4 ♂♂, 4 ♀♀: MAK-28; 1 ♀: MAK-29; 5 ♂♂, 2 ♀♀: MAK-30; 3 ♂♂, 5 ♀♀: MAK-31A; 5 ♂♂, 2 ♀♀: MAK-31B; 4 ♂♂, 1 ♀: MAK-31C; 12 ♂♂, 20 ♀♀: MAK-32; 1 ♂, 4 ♀♀: MAK-33; 1 ♂: MAK-34A; 2 ♂♂, 3 ♀♀: MAK-34B; 11 ♂♂, 3 ♀♀: MAK-35A; 4 ♂♂: MAK-35B; 5 ♂♂: MAK-35C; 6 ♂♂, 5 ♀♀: MAK-36B; 8 ♂♂, 11 ♀♀: MAK-38A; 1 ♂, 1 ♀: MAK-39A; 3 ♂♂, 1 ♀: MAK-39B; 1 ♂: MAK-44C; 68 exs.: MAK-45; 2 ♂♂, 5 ♀♀: MAK-46; 4 ♂♂, 6 ♀♀: MAK-47; 21 exs.: MAK-49; 130 exs.: MAK-50; 26 exs.: MAK-51; 111 exs.: MAK-52; 108 exs.: MAK-53; 16 ♂♂, 19 ♀♀: MAK-54A; 64 exs: MAK-54B; 8 ♂♂, 1 ♀: MAK-58; 37 exs.: MAK-59A; 5 ♂♂, 2 ♀♀: MAK-59B; 62 exs: MAK-59C.</p>
            <p>Distribution.</p>
            <p>So far endemic to the Makay massif, Madagascar (Manuel and Ramahandrison 2020).</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2D-G, I-N). This species was collected only at inner massif sites, in both south-central and northern Makay, where it is by far the most common and abundant species of aquatic  Adephaga . It was found in a wide diversity of lentic water bodies (puddles, pools, ponds, a blind river channel, etc.), isolated or with slow water renewal, as well as in very slowly flowing streams. The surrounding environment was forested or semi-forested and free from anthropisation. These habitats were further characterised by sandy bottom (sometimes with stones), various amounts of plant debris, clear water (but often with orange masses of iron bacteria), and marginal vegetation absent or poorly developed. </p>
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	https://treatment.plazi.org/id/ACDD693E6E1B50E3BE82FCB09EECE56C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
68D7D1208FFE542492C110A0E5A11522.text	68D7D1208FFE542492C110A0E5A11522.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus pallescens Regimbart 1903	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Laccophilus pallescens 
Regimbart
, 1903
</p>
            <p>Type locality.</p>
            <p>Southern Madagascar, Pays Androy.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-1A; 8 ♂♂, 5 ♀♀: MAK-2; 1 ♂: MAK-18; 1 ♂: MAK-19.</p>
            <p>Distribution.</p>
            <p> Sub Saharan-Africa, Arabian Peninsula, Madagascar (Guignot 1959-1961; Bertrand and Legros 1971;  Hájek and Reiter 2014;  Biström et al. 2015). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species was collected only at peripheral sites, in shallow lentic or slowly flowing lotic habitats, with or without marginal vegetation. The environment was open (non-forested), at some sites partly sheltered by sparse trees, and at most sites impacted by anthropogenic pressures (notably cattle trampling). The bottom varied from clay to sand and more or less abundant plant debris, and the water was clear to turbid.</p>
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	https://treatment.plazi.org/id/68D7D1208FFE542492C110A0E5A11522	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
63E8D9F3467B5892B70606E7B345E09E.text	63E8D9F3467B5892B70606E7B345E09E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus posticus Aube 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Laccophilus posticus 
Aube
, 1838
</p>
            <p>Type locality.</p>
            <p>Mascarene Islands, Mauritius.</p>
            <p>Material examined.</p>
            <p>5 ♂♂: MAK-1A; 1 ♂: MAK-2; 1 ♀: MAK-3; 3 ♀♀: MAK-4; 4 ♂♂, 3 ♀♀: MAK-11A; 1 ♂: MAK-11B; 1 ♂: MAK-17; 1 ♂: MAK-18; 243 exs.: MAK-19; 1 ♀: MAK-21; 3 ♂♂, 9 ♀♀: MAK-23; 1 ♀: MAK-28; 1 ♂, 2 ♀♀: MAK-40A; 11 ♂♂, 14 ♀♀: MAK-41; 37 exs.: MAK-42; 1 ♂: MAK-44A; 1 ♂: MAK-59C; 4 ♂♂, 6 ♀♀: MAK-60; 6 ♂♂, 1 ♀: MAK-62.</p>
            <p>Distribution.</p>
            <p> Mauritius, Aldabra, Madagascar (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984; Rocchi 1991;  Biström et al. 2015). In Madagascar, widespread and common in lowlands. </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B-E, O, P). This species was captured both at peripheral and inner massif sites, in various kinds of lentic habitats (puddles, pools, ponds, ditches) and in very slowly flowing streams. These habitats were most often located in open areas. The bottom variously consisted of sand, clay or stones, generally with plant debris. The water was clear to turbid and marginal vegetation was absent or variously developed. This species is highly tolerant to anthropogenic pressure.</p>
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	https://treatment.plazi.org/id/63E8D9F3467B5892B70606E7B345E09E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
AAA1740117905ED39A0AC36F4C128034.text	AAA1740117905ED39A0AC36F4C128034.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus rivulosus Klug 1833	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laccophilus rivulosus Klug, 1833</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-2; 2 ♂♂, 7 ♀♀: MAK-19.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread but not very common (Guignot 1959-1961; Bertrand and Legros 1971;  Biström et al. 2015). </p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was found at two sampling sites located in the periphery of the Makay massif: a shallow isolated stream characterised by very weak water flow, sandy bottom, sparse tufts of small  Cyperaceae and strong presence of filamentous green algae, and a large puddle partially sheltered by trees, with water slowly flowing and with abundant rice straw debris, on a dirty road between two rice fields. These habitats were situated in open environments and were impacted by anthropogenic disturbance (notably cattle trampling). </p>
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	https://treatment.plazi.org/id/AAA1740117905ED39A0AC36F4C128034	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
74F6D2CB69B65E919CC06727252E4ADD.text	74F6D2CB69B65E919CC06727252E4ADD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus seyrigi Guignot 1937	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Laccophilus seyrigi Guignot, 1937</p>
            <p>Type locality.</p>
            <p>Southern Madagascar, near Bekily.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-2.</p>
            <p>Distribution.</p>
            <p> South and south-western Madagascar (very rare); the present record is the first since the original description of the species (Guignot 1959-1961;  Biström et al. 2015). </p>
            <p>Habitat in study area.</p>
            <p> Same as  Laccophilus flaveolus (see above). </p>
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	https://treatment.plazi.org/id/74F6D2CB69B65E919CC06727252E4ADD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
B9211C721C09542EBEE1492B7B1A498D.text	B9211C721C09542EBEE1492B7B1A498D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus sp. Ma 19	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laccophilus sp. Ma19</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-21.</p>
            <p>Note.</p>
            <p> This is an undescribed species close to  L. lateralis Sharp, 1882. </p>
            <p>Distribution.</p>
            <p>Madagascar (widespread in lowlands).</p>
            <p>Habitat in study area.</p>
            <p> Same as  Laccophilus luctuosus (see above). </p>
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	https://treatment.plazi.org/id/B9211C721C09542EBEE1492B7B1A498D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
916F2599E9025AA5AD6FCBF320792047.text	916F2599E9025AA5AD6FCBF320792047.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccophilus transversovittatus Bistroem, Nilsson & Bergsten 2015	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Laccophilus transversovittatus 
Bistroem
, Nilsson &amp; Bergsten, 2015
</p>
            <p>Type locality.</p>
            <p>Madagascar, Isalo, Menamaty River.</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-12C; 1 ♀: MAK-26; 1 ♀: MAK-52.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread outside from the Central Highlands (  Biström et al. 2015). The record from Ankaratra in  Biström et al. (2015) is probably attributable to  L. rakouthae Manuel &amp; Ramahandrison, 2020. </p>
            <p>Habitat in study area.</p>
            <p> This species was found at three inner massif sites, two in south-central and one in northern Makay. The surrounding was forested or semi-forested and without visible anthropogenic disturbance. The habitats were: a small pond with water slowly renewed, just downstream from a spring, partly sheltered by trees, with clay bottom, with plant debris and clear water, with sparse helophytes (  Cyperus ); a slowly flowing stream, partly shaded, with sandy bottom, important accumulation of plant debris, clear water and sparse helophytes; and a small, isolated pool, partly shaded, with sandy bottom, no plant debris, tinted water, and no marginal vegetation. </p>
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	https://treatment.plazi.org/id/916F2599E9025AA5AD6FCBF320792047	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
370740EA691258498481CB483889DB19.text	370740EA691258498481CB483889DB19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liodessus luteopictus (Regimbart 1897)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Liodessus luteopictus (
Regimbart
, 1897)
</p>
            <p> Liodessus poecilopterus =  L. poecilopterus Régimbart , 1900. </p>
            <p>Type locality.</p>
            <p>Mascarene Islands, Mauritius, Curepipe.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-16.</p>
            <p>Distribution.</p>
            <p> Mauritius, La  Réunion , Comoros, Madagascar (Guignot 1959-1961; Bameul 1984;  Biström 1988b). In Madagascar, widespread. </p>
            <p>Habitat in study area.</p>
            <p> This species was sampled at a single inner massif site, a small pond, partly shaded, at the edge of a gallery forest close to River Andranomanintsy. The water was slowly renewed from the nearby river, the bottom was sandy with moderate plant debris and the water was clear. This small water body was filled in with subaquatic  Poaceae including a  Panicum species. </p>
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	https://treatment.plazi.org/id/370740EA691258498481CB483889DB19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
EA4B265D0BF952188497CF3CD498C2BF.text	EA4B265D0BF952188497CF3CD498C2BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Madaglymbus fairmairei (Zimmermann 1919)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Madaglymbus fairmairei (Zimmermann, 1919)</p>
            <p> Madaglymbus regimbartii =  Madaglymbus regimbartii Fairmaire, 1898. </p>
            <p>Type locality.</p>
            <p>Madagascar, Maevatanana.</p>
            <p>Material examined.</p>
            <p>1 ♂, 2 ♀: MAK-3; 2 ♂♂, 8 ♀♀: MAK-10; 1 ♂, 1 ♀: MAK-11B; 2 ♂♂, 2 ♀♀: MAK-12A; 1 ♂: MAK-15; 1 ♂, 1 ♀: MAK-16; 9 ♂♂, 1 ♀: MAK-25A; 3 ♂♂, 1 ♀: MAK-29; 1 ♂: MAK-35A; 1 ♀: MAK-37A; 4 ♂♂, 2 ♀♀: MAK-39A; 1 ♂: MAK-40B.</p>
            <p>Distribution.</p>
            <p>Madagascar (distribution within the island poorly known) (Guignot 1959-1961).</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2D, L, M, P). We met this species only in the central part of inner Makay, mainly in puddles and pools with or without water circulation and in small streams, with clear or turbid water, sandy bottom (at some sites with gravel and stones) and moderate to abundant plant debris or tree roots, in forested or semi-forested environments untouched by anthropogenic disturbance.</p>
            <p> Subfamily  Cybistrinae , tribe  Cybistrini</p>
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	https://treatment.plazi.org/id/EA4B265D0BF952188497CF3CD498C2BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
6AD22F636B675FD18A9934F1817CA2AB.text	6AD22F636B675FD18A9934F1817CA2AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Methles sp. Ma 1	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Methles sp. Ma1</p>
            <p>Material examined.</p>
            <p>4 ♂♂, 3 ♀♀: MAK-44C; 1 ♂, 2 ♀♀: MAK-60.</p>
            <p>Note.</p>
            <p> The Malagasy species of the genus  Methles are in great need of revision, and in the current state of knowledge we cannot assign a name to this species. </p>
            <p>Distribution.</p>
            <p>Unknown (but not confined to the Makay area).</p>
            <p>Habitat in study area.</p>
            <p> This species was collected at two inner massif sites in northern Makay, with very different habitat characteristics. One was a blind channel in forest connected to River Sakapaly; this site is the locus typicus of  Copelatus malavergnorum sp. nov. (see  “Habitat” under description of this species). The other site was an open marsh with vegetated margins (  Cyperaceae and  Polygonaceae ), with muddy bottom and water rather turbid, in semi-forested context near the Sakapaly River. Both sites were preserved from anthropogenic disturbance. </p>
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	https://treatment.plazi.org/id/6AD22F636B675FD18A9934F1817CA2AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
2F4450B01EC05F899F7FFFC21CD15C29.text	2F4450B01EC05F899F7FFFC21CD15C29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Methles sp. Ma 5	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Methles sp. Ma5</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-2; 3 ♂♂, 3 ♀♀: MAK-42; 1 ♀: MAK-43; 1 ♂: MAK-44C; 1 ♂, 3 ♀♀: MAK-60; 19 ♂♂, 24 ♀♀: MAK-61; 3 ♂♂, 1 ♀: MAK-62.</p>
            <p>Note.</p>
            <p> The Malagasy species of the genus  Methles are in great need of revision, and in the current state of knowledge we cannot assign a name to this species. </p>
            <p>Distribution.</p>
            <p>Unknown (but not confined to the Makay area).</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species is present in lentic and in slowly flowing lotic habitats. It was collected both at peripheral and inner massif sites. The bottom varied from clay to sandy, with more or less abundant plant debris. The water was either clear, red-brown or turbid. This species has a clear preference for open environments and habitats with at least some vegetation and is tolerant to anthropogenic disturbance.</p>
            <p> Subfamily  Laccophilinae , tribe  Laccophilini</p>
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	https://treatment.plazi.org/id/2F4450B01EC05F899F7FFFC21CD15C29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
85AC78A6944654C3B348E88464A8D466.text	85AC78A6944654C3B348E88464A8D466.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohydrocoptus seriatus (Sharp 1882)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Neohydrocoptus seriatus (Sharp, 1882)</p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p> 2 ♂♂: MAK-2; 1 ♀: MAK-11A; 8 ♂♂, 16 ♀♀: MAK-19; 1 ♂, 2 ♀♀: MAK-21; 20 exs.: MAK-23; 1 ♂: MAK-38B; 2 ♂♂, 1 ♀: MAK-42; 8 ♂♂, 10 ♀♀: MAK-43; 1 ♂, 4 ♀♀: MAK-44A; 1 ♀: MAK-44C; 9 ♂♂, 17 ♀♀: MAK-60; 2  еxs .: MAK-61. </p>
            <p>Distribution.</p>
            <p>Africa (Angola, Guinea, Guinea-Bissau, Mali), Madagascar, and Mascarene Islands (Guignot 1959-1961; Bertrand and Legros 1971; Rocchi 1991; Nilsson 2011). In Madagascar, widespread and common.</p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C, O). This species is present in lentic and in slowly flowing lotic habitats. It was collected both at peripheral and inner Makay sites. The bottom varied from clay to sandy, with clear, red-brown or turbid water and with more or less abundant plant debris. This species has a clear preference for open environments and habitats with at least some vegetation, and is highly tolerant to anthropogenic disturbance.</p>
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	https://treatment.plazi.org/id/85AC78A6944654C3B348E88464A8D466	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
72409EBF475351AAB71F9995E6006DB4.text	72409EBF475351AAB71F9995E6006DB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohydrocoptus sp. Ma 3	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Neohydrocoptus sp. Ma3</p>
            <p>Material examined.</p>
            <p>3 ♂♂, 2 ♀♀: MAK-43; 2 ♀♀: MAK-44C; 4 ♂♂, 1 ♀: MAK-56.</p>
            <p>Note.</p>
            <p> This species is smaller than  N. seriatus and the elytra do not bear additional serial groups of punctures beyond discal and lateral puncture rows. External features and the shape of the aedeagus evoke  N. aethiopicus (J. Balfour-Browne, 1961), a widespread sub-Saharan species, but examination of type material in the context of a revision will be necessary to confirm the identity of this species. </p>
            <p>Distribution.</p>
            <p>Madagascar, widespread but not very common.</p>
            <p>Habitat in study area.</p>
            <p>This species was sampled at three sites in inner Makay, in slowly flowing lotic habitats and a dead river arm, in forested or semi-forested contexts without anthropogenic disturbance. These biotopes had sandy bottoms with moderate abundance of plant debris. Two of the sites were surrounded by a well-developed hygrophilous vegetation and contained cyanobacteria.</p>
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	https://treatment.plazi.org/id/72409EBF475351AAB71F9995E6006DB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
C9D8363F971F52FFBF0E72324000CF28.text	C9D8363F971F52FFBF0E72324000CF28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neptosternus oblongus Regimbart 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Neptosternus oblongus 
Regimbart
, 1895
</p>
            <p>Type locality.</p>
            <p>Madagascar, Annanarivo (= Antananarivo?).</p>
            <p>Material examined.</p>
            <p>35 ♂♂, 50 ♀♀: MAK-5A; 3 ♂♂: MAK-5B.</p>
            <p>Distribution.</p>
            <p>Central and southern Madagascar (Guignot 1959-1961; Bilardo and Rocchi 2012). Distribution within the island poorly known.</p>
            <p>Habitat in study area.</p>
            <p> Same as  Laccophilus insularum (see above). </p>
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	https://treatment.plazi.org/id/C9D8363F971F52FFBF0E72324000CF28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
A3183F07BCE55CEB833F1F79B65C1A51.text	A3183F07BCE55CEB833F1F79B65C1A51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orectogyrus vicinus Regimbart 1892	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Orectogyrus vicinus 
Regimbart
, 1892
</p>
            <p>Type locality.</p>
            <p>Madagascar, Diego Suarez (Antsiranana), Isokitra.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-15; 2 ♂♂, 3 ♀♀: MAK-36A; 7 ♂♂, 6 ♀♀: MAK-37B; 1 ♀: MAK-40B; 1 ♂, 3 ♀♀: MAK-48.</p>
            <p>Distribution.</p>
            <p>Madagascar. Previously recorded only from the northern part of the island (Legros 1951; Brinck 1956; Gustafson et al. in press).</p>
            <p>Habitat in study area.</p>
            <p>Same as the two preceding species, with a stronger preference for forested and undisturbed habitats.</p>
            <p> Family  Haliplidae</p>
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	https://treatment.plazi.org/id/A3183F07BCE55CEB833F1F79B65C1A51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
154AA14CA1615C7290B9BB0864919B02.text	154AA14CA1615C7290B9BB0864919B02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pachynectes costulifer (Regimbart 1903)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Pachynectes costulifer (
Regimbart
, 1903)
</p>
            <p>Type locality.</p>
            <p>Madagascar, Imanombo.</p>
            <p>Material examined.</p>
            <p>4 ♂♂, 2 ♀♀: MAK-1A; 3 ♂♂, 11 ♀♀: MAK-1B; 12 ♂♂, 26 ♀♀: MAK-1C; 2 ♂♂: MAK-20; 2 ♂♂: MAK-22.</p>
            <p>Distribution.</p>
            <p> Madagascar, western and southern parts of the island (Guignot 1959-1961; Bertrand and Legros 1971;  Biström 1987; Rocchi 1991). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2A). This species was found only at peripheral sites located on the large sandy banks of the Mangoky River close to Beroroha, in small to large shallow isolated puddles, with sand or sand-clay bottom and clear water, without vegetation or with sparse vegetation, with or without presence of filamentous green algae.</p>
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	https://treatment.plazi.org/id/154AA14CA1615C7290B9BB0864919B02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
59D02A6C4DF957F398F2D27E3455827A.text	59D02A6C4DF957F398F2D27E3455827A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pachynectes sp. Ma 1	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Pachynectes sp. Ma1</p>
            <p>Material examined.</p>
            <p>5 ♂♂, 2 ♀♀: MAK-2; 25 ♂♂, 22 ♀♀: MAK-3; 8 ♂♂, 4 ♀♀: MAK-4; 3 ♂♂, 1 ♀: MAK-5A;; 8 exs.: MAK-5D; 2 ♀♀: MAK-8; 29 ♂♂, 14 ♀♀: MAK-14A; 5 ♂♂, 5 ♀♀: MAK-16; 24 exs.: MAK-28; 24 exs.: MAK-29; 3 exs.: MAK-36B; 91 exs.: MAK-37A; 3 exs: MAK-38B; 53 exs.: MAK-40A; 1 ex.: MAK-43; 1 ex.: MAK-44C; 2 exs.: MAK-45; 28 exs.: MAK-47; 5 exs.: MAK-49; 31 exs.: MAK-50; 15 exs.: MAK-51; 37 exs.: MAK-52; 10 exs.: MAK-53; 2 exs.: MAK-54B; 6 exs.: MAK-58; 39 exs.: MAK-59A; 25 exs.: MAK-59C.</p>
            <p>Note.</p>
            <p> This is a probably undescribed species close to  P. hygrotoides (  Régimbart , 1895). The Malagasy endemic genus  Pachynectes is currently being revised (J. Bergsten, pers. comm.) and many species are awaiting description. </p>
            <p>Distribution.</p>
            <p>Madagascar. The exact distribution of this species within the island remains to be established in the context of the upcoming revision, but it is not endemic to the Makay (sampled by us notably in the Isalo Massif).</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B-F, I, N). This is one of the most abundant species of aquatic  Adephaga in inner Makay, and it was also collected at one peripheral site (MAK-2). It was sampled in all kinds of lentic or very slowly flowing lotic habitats (puddles, pools, ponds, small streams, blind channels of rivers, etc.), with substrate sandy or stony, water generally clear, and in most cases without vegetation. This species has a preference for exposed or semi-shaded situations (majority of observations in semi-forested environments). </p>
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	https://treatment.plazi.org/id/59D02A6C4DF957F398F2D27E3455827A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
700E788A891A58FEA743455EE5FF89D3.text	700E788A891A58FEA743455EE5FF89D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pachynectes sp. Ma 4	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Pachynectes sp. Ma4</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-5A; 2 ♂♂: MAK-5D; 1 ♂, 1 ♂: MAK-14A; 1 ♂, 1 ♀: MAK-29; 1 ♀: MAK-45; 8 ♂♂, 4 ♀♀: MAK-50; 1 ♂, 4 ♀♀: MAK-51; 57 ♂♂, 37 ♀♀: MAK-52; 1 ♀: MAK-59C.</p>
            <p>Distribution.</p>
            <p>Madagascar. So far endemic to the Makay massif.</p>
            <p>Note.</p>
            <p>This is an undescribed species, rather large for the genus, and very close to another undescribed species which lives in the Isalo massif.</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2F, N). Similar to  Pachynectes sp. Ma1 (with which it was syntopic at all sites), but this species is rarer. </p>
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	https://treatment.plazi.org/id/700E788A891A58FEA743455EE5FF89D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
A056C291CD5A552A836320C951EE8811.text	A056C291CD5A552A836320C951EE8811.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Peltodytes quadratus Regimbart 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Peltodytes quadratus 
Regimbart
, 1895
</p>
            <p>Type locality.</p>
            <p>Madagascar, Antananarivo, Ambodinandohalo Lake.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-19; 1 ♂: MAK-41.</p>
            <p>Distribution.</p>
            <p>Madagascar, widespread (Guignot 1959-1961; Bertrand and Legros 1971; Bameul 1984; Rocchi 1991; van Vondel and Bergsten 2012).</p>
            <p>Habitat in study area.</p>
            <p> This species was only found at two sampling sites, both peripheral. One was a large puddle partially sheltered by trees, with water slowly flowing and with abundant rice straw debris, on a dirty road between two rice fields, and the other was a small puddle in open situation on the sandy bank of a river, with  Azolla aquatic ferns (eutrophication indicator). </p>
            <p> Family  Noteridae</p>
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	https://treatment.plazi.org/id/A056C291CD5A552A836320C951EE8811	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
2EA0903F3C6E595695CA7080D971E29B.text	2EA0903F3C6E595695CA7080D971E29B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philaccolus elongatus (Regimbart 1903)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Philaccolus elongatus (
Regimbart
, 1903)
</p>
            <p>Type locality.</p>
            <p>Madagascar, Sainte Marie Island.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2.</p>
            <p>Note.</p>
            <p> There is a complex of very similar species around  P. elongatus , and future studies may show that the specimen recorded here belong to a different species. </p>
            <p>Distribution.</p>
            <p>Madagascar (Guignot 1959-1961; Bertrand and Legros 1971). Distribution within the island poorly known.</p>
            <p>Habitat in study area.</p>
            <p> Same as  Laccophilus flaveolus (see above). </p>
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	https://treatment.plazi.org/id/2EA0903F3C6E595695CA7080D971E29B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
F1EFAE2510DA5DBE98052370F480F8E5.text	F1EFAE2510DA5DBE98052370F480F8E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseuduvarus sp. Ma 1	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Pseuduvarus sp. Ma1</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-2; 1 ♂: MAK-61.</p>
            <p>Distribution.</p>
            <p>Madagascar (widespread).</p>
            <p>Note.</p>
            <p> This species corresponds to  P. ornatipennis (  Régimbart , 1900), currently wrongly considered a junior synonym of  P. vitticollis (Boheman, 1848). A revision of the species of  Hydroglyphus /  Pseuduvarus is currently in preparation (J. Bergsten, pers. comm.). </p>
            <p>Habitat in study area.</p>
            <p> Same as  Clypeodytes concivis (see above). </p>
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	https://treatment.plazi.org/id/F1EFAE2510DA5DBE98052370F480F8E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
FFF287BE10D75137BFABE5FC147FA128.text	FFF287BE10D75137BFABE5FC147FA128.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhantaticus congestus (Klug 1833)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Rhantaticus congestus (Klug, 1833)</p>
            <p> Rhantaticus rochasi =  R. rochasi (Perroud &amp; Montrouzier, 1864);  R. signatipennis (Laporte, 1835). </p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♂: MAK-2; 2 ♀♀: MAK-19; 1 ♂: MAK-41.</p>
            <p>Distribution.</p>
            <p>From sub-Saharan Africa to Australia through tropical Asia and the Oriental region (Guignot 1959-1961; Bertrand and Legros 1971; Watts 1978; Bameul 1984; Hendrich et al. 2010). In Madagascar, widespread.</p>
            <p>Habitat in study area</p>
            <p> (Fig. 2B, C). This species was found at three sampling sites in the periphery of the Makay massif: a large puddle partially sheltered by trees, with water slowly flowing and with abundant rice straw debris, on a dirty road between two rice fields; a small puddle in open situation on the sandy bank of a river, with  Azolla aquatic ferns (eutrophication indicator); and a shallow isolated stream characterised by very weak water flow, sandy bottom, sparse tufts of small  Cyperaceae and strong presence of filamentous green algae. These habitats were all situated in open environments and more or less affected by anthropogenic disturbance. </p>
            <p> Tribe  Eretini</p>
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	https://treatment.plazi.org/id/FFF287BE10D75137BFABE5FC147FA128	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
85764FA04BC65B63A182738F86AEB2E7.text	85764FA04BC65B63A182738F86AEB2E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sternocanthus fabiennae (Bameul 1994)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Sternocanthus fabiennae (Bameul, 1994)</p>
            <p>Type locality.</p>
            <p>Madagascar, Mahajanga, Ambohimanatrika.</p>
            <p>Material examined.</p>
            <p>3 ♂♂, 2 ♀♀: MAK-19; 1 ♂, 1 ♀: MAK-41.</p>
            <p>Distribution.</p>
            <p>Madagascar (Bameul 1994; Nilsson 2011); distribution within the island poorly known.</p>
            <p>Habitat in study area.</p>
            <p>This species was collected at two peripheral sites in lentic habitats in areas with intense anthropogenic pressure: large puddle, with water slowly flowing and with abundant rice straw debris, on a dirty road between two rice fields; and puddle on the sandy banks of the Mangoky River.</p>
        </div>
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	https://treatment.plazi.org/id/85764FA04BC65B63A182738F86AEB2E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
234162579B8B5A2F88A2C15E174AF292.text	234162579B8B5A2F88A2C15E174AF292.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Synchortus asperatus (Fairmaire 1869)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Synchortus asperatus (Fairmaire, 1869)</p>
            <p> Synchortus duplicatus =  S. duplicatus Sharp, 1882. </p>
            <p>Type locality.</p>
            <p>Madagascar.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-21; 1 ♂: MAK-42.</p>
            <p>Distribution.</p>
            <p>Madagascar; widespread and common in lowlands (Guignot 1959-1961; Bertrand and Legros 1971; Nilsson 2011).</p>
            <p>Habitat in study area.</p>
            <p>This species was collected at two peripheral sites located in open areas, in temporary lentic habitats without water renewal, and with vegetation.</p>
            <p> Family  Dytiscidae</p>
            <p> Subfamily  Copelatinae , tribe  Copelatini</p>
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	https://treatment.plazi.org/id/234162579B8B5A2F88A2C15E174AF292	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
745802C4FD21537198D1E66838797117.text	745802C4FD21537198D1E66838797117.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Uvarus betsimisarakus (Guignot 1939)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Uvarus betsimisarakus (Guignot, 1939)</p>
            <p>Type locality.</p>
            <p>Madagascar, Maroantsetra.</p>
            <p>Material examined.</p>
            <p>1 ♂, 1 ♀: MAK-19.</p>
            <p>Distribution.</p>
            <p> Madagascar (Guignot 1959-1961;  Biström 1988c). Common in the Central Highlands. </p>
            <p>Habitat in study area.</p>
            <p>This species was collected only once, in a peripheral site located in a semi-open area impacted by human activities. The habitat was a large puddle partially sheltered by trees, with water slowly flowing and with abundant rice straw debris, on a dirty road between two rice fields.</p>
        </div>
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	https://treatment.plazi.org/id/745802C4FD21537198D1E66838797117	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
FC43306B09DC5932A4F86050F214CBD2.text	FC43306B09DC5932A4F86050F214CBD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Uvarus rivulorum (Regimbart 1895)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Uvarus rivulorum (
Regimbart
, 1895)
</p>
            <p> Uvarus cilunculus =  U. cilunculus Guignot, 1950. </p>
            <p>Type locality.</p>
            <p>Madagascar, Antsiranana.</p>
            <p>Material examined.</p>
            <p>1 ♀: MAK-1A; 2 ♀♀: MAK-2; 1 ♀: MAK-18; 4 ♂♂, 2 ♀♀: MAK-19; 4 exs.: MAK-60.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread in lowlands (Guignot 1959-1961; Rocchi 1991;  Biström 1988c). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2B, C). This species was collected mainly at peripheral sites, in shallow lentic or slowly flowing lotic habitats, with or without marginal vegetation. The environment was open (non-forested), at some sites partly sheltered by sparse trees, and at most sites impacted by anthropogenic pressures (notably cattle trampling). The bottom varied from clay to sand and more or less abundant plant debris; the water was clear to turbid.</p>
        </div>
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	https://treatment.plazi.org/id/FC43306B09DC5932A4F86050F214CBD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
B4643799AA5F5FC18A98F9952F0ABB25.text	B4643799AA5F5FC18A98F9952F0ABB25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yola costipennis (Fairmaire 1869)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Yola costipennis (Fairmaire, 1869)</p>
            <p>Type locality.</p>
            <p>Madagascar, Sainte Marie Island.</p>
            <p>Material examined.</p>
            <p>5 exs.: MAK-1A; 2 exs.: MAK-1C; 4 exs.: MAK-19; 6 exs.: MAK-20; 4 exs.: MAK-41; 1 ex.: MAK-42; 1 ex.: MAK-60; 3 exs.: MAK-62.</p>
            <p>Distribution.</p>
            <p> Madagascar, widespread and common (Guignot 1959-1961; Bertrand and Legros 1971;  Biström 1983; Bameul 1984; Rocchi 1991). </p>
            <p>Habitat in study area</p>
            <p>(Fig. 2A). This species was collected mainly at peripheral sites and always in open areas, often impacted by human activities. The habitats were various kinds of shallow lentic water bodies, most without but some with water circulation. They were sun-exposed (at one site partly sheltered by trees), with bottom of sand, clay, or a mix of sand and clay, with organic debris varying from absent to forming a thick layer above the mineral substratum, with water clear to moderately turbid; vegetation was absent or sparse, and filamentous green algae were either undetectable or variously developed.</p>
            <p> Tribe  Hydrovatini</p>
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	https://treatment.plazi.org/id/B4643799AA5F5FC18A98F9952F0ABB25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ramahandrison, Andriamirado Tahina;Rakouth, Bakolimalala;Manuel, Michael	Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala, Manuel, Michael (2022): The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae). ZooKeys 1127: 1-60, DOI: http://dx.doi.org/10.3897/zookeys.1127.85737, URL: http://dx.doi.org/10.3897/zookeys.1127.85737
