taxonID	type	description	language	source
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	description	(Figs 1 A, B, 2, 3, 6) www / zoobank. org / urn: lsid: zoobank. org: act: 6511 FDD 9 - 6 CC 6 - 4 B 05 - 92 C 4 - FEC 9 B 1731062	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	diagnosis	Diagnosis. Differs from all known species of the genus Sphrigodellus by uniformly distributed scales forming irregular pattern of grey and brown scales covering the body; prementum with four setae; constricted and truncate apex of median lobe, C-shaped lobes of endophallic sclerite; collum of spermatheca slender, 9 x longer than ramus; lamina of female tergite 8 with strongly sclerotized fields. From S. lineatus Yunakov, 2012 (yet another species from Udzungwa Scarp) S. griseus also differs by pterygia not projecting from outline of rostrum; anterior portion of epifrons with deep groove; 7 th funicular antennomere 2 x longer than wide; medial margin of metatibia serrate; parameres fused at base.	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	description	Description. Measurements: BL mf: 2.92 – 3.24 (3.05) mm; BW mf: 1.26 – 1.56 (1.37) mm; BH mf: 1.08 – 1.60 (1.30) mm; RL / RW mf: 1.20 – 1.33 (1.24); VW / ELD mf: 1 – 1.42 (1.19); PL / PW mf: 0.88 – 0.95 (0.91); EL / BH mf: 1.46 – 1.73 (1.59); EL / EW mf: 1.29 – 1.56 (1.43); EL / PL mf: 2.31 – 2.82 (2.58); BL / PL mf: 3.82 – 4.33 (4.11); BL / EL mf: 1.54 – 1.66 (1.59). Vestiture: Body densely covered with overlapping round striate scales, setose. Setae of head spatulate, forming rows along lateral margins of epifrons (recumbent or subrecumbent) and medial to eyes (erect). Subocular row consists of 3 – 4 setae. Anterior setal fringe on pronoum consists of 6 (3 + 3) erect setae; weakly differing from those on pronotal disc. Posterior setal fringe consists of spatulate setae, partly hidden by posterior margin of pronotum. Elytral intervals 1 – 6 each with distinct row of setae. Anterior half of elytra with rather short erect setae (as long as 0.5 of interval width), declivity with long, strongly erect, thick, truncated setae (as long as width of an interval). Ventral side rather sparsely squamose. Abdominal sternites with scattered scales; male anal ventrite with long hairs in apical portion. Metapleura densely squamose; basisternum and mesobasisternum bare. Antennal scape setose and squamose; setae very long, slender, acute, erect. External surface of femora and tibiae covered with overlapping scales and suberect setae, internal surface with sparse hairs; outer surface of distal portion of tibiae without scales; setae on legs slender, acute. Male metatibiae with sparse very small grooming brush; hairs short. Tarsi setose. Funicle setose, without scales; setae suberect as long as antennomere 8. Club densely tomentose. Coloration: Integument of body dark-brown to black; legs and antennae dark-brown. Background scaling consists of greyish-brown, and grey scales; spotty pattern obsolete, formed by grey and brown scales. Dark and light scales both with slightly pronounced pearly shine. Head brown, indistinctly greyish maculate. Pronotum with darkbrown background scaling and five longitudinal grey stripes: 1 discal (thin, obsolete) and 4 lateral (broad, rather distinct). Double lateral stripes occasionally merged in common broad stripe separated from ventral light scaling by very narrow brown striae. Green scales absent. Elytra with brown background scaling, alternate striped pattern indistinct. Interrupted grey stripes present at intervals 1, 3, 5, and 7. Some specimens with scattered indistinct grey marmorate pattern on elytral disc. Ventral side including meso- and metapleura with sparse grey scales. Sides of metasternum grey. Femora brown with grey bands in middle and proximal portion, and scattered grey scales occasionally merged in small spots. Tibiae with external surface brown with distal grey band. Head: Rostrum weakly elongate [RL / RWA: 1.2 – 1.32 (1.24)], parallel-sided. Pterygia not extended from outline of rostrum. Antennal sockets dorsal. Antennifer completely visible. Lateral carina convex. Epifrons parallel-sided, distinctly sinuate in middle, at level of antennal articulation 0.7 x as wide as width of vertex, distinctly sloping at sides, very steeply sloping anteriorly, significantly concave and deeply sulcate in anterior portion. Transverse sulcus deep, concealed by dense scaling on dorsal surface and partly open at sides. Anterior portion of epifrons with median sulcus, almost vertical. Frons vertical, glabrous, bare, with six frontal setae. Epistome vestigial, very narrow, transverse, surrounded by narrow carina, bearing two epistomal setae (Fig. 2 B). Prementum with four setae (Fig. 2 C). Eyes: orbicular, sublateral, strongly convex, highest posteriorly [VW / ELD: 1 – 1.42 (1,19)]. Vertex: broad, flat. Parietal socket deep, elongate. Antennae: Scape extending beyond anterior margin of pronotum, weakly, evenly curved, apically widened. Funicular antennomere 1 longer or as long as and wider than 2 nd, funicular antennomeres 3 – 7 oblong. Club spindle-shaped, 2.28 x longer than wide (Figs. 2 D, E). Thorax: Prothorax transverse [PL / PW: 0.87 – 0.95 (0.91)], evenly slightly convex at sides, not constricted, widest at middle. Disc strongly convex longitudinally and transversally. Posterior margin of pronotum slightly sinuate in middle part. Tergosternal and metepisternal sutures complete. Elytra: Elytra shape in male oblong-oval, in female oval [EL / EW: 1.29 – 1.56 (1.43)], anterior margin vertical, narrowly rounded apically; disc in both sexes strongly convex [EL / BH: 1.46 – 1.73 (1.59)]. Subscutellar callosity reaching 3 rd elytral interval, without tiny tubercles. Elytral intervals flat; interval 7 with distinct convex tubercle at base. Femora: edentate, swollen in middle part. Tibiae: Protibiae and mesotibiae almost straight, not widened at apex, medial margin C-shaped, serrate in distal portion (Fig. 2 G). Meso- and metatibiae with teeth on medial margin. Mucro well developed, thorn-shaped, acute (Fig. 2 H). Setal comb of protibia of sparse 10 – 14 brown setae weakly extending beyond outline of tibia. Bevel of metatibia narrowly enclosed, bare. Tarsi: Tarsomere 2 elongate, 1.5 x longer than wide; tarsomere 3 with two wide lobes; metatarsomere 5 extending beyond lobes of tarsomere 3 by length of last tarsomere. Claws connate in basal half. Abdomen: 1 st and 2 nd ventrites fused with distinct immovable suture. Posterior margin of 1 st ventrite straight, 2 nd ventrite 2 x longer than 3 rd, posterior margin of 2 nd ventrite straight. Male 5 th ventrite flat, without depression near apex, posterior margin broadly rounded. Female 5 th ventrite convex, posterior margin acute. Male genitalia: Aedeagus strongly sclerotized. Median lobe parallel-sided, as long as apodemes; apex constricted, truncate, dorso-ventrally swollen. Lateral margins of median lobe not fused dorsally. Internal sac without spiculate fields; endophallic sclerite large, consisting of two curved, rod-shaped lobes, connected by transverse bridge in proximal portion. Ostium triangular strongly sclerotized, protruded. Ventral membranes (ligulae) transparent (Fig. 3 A). Parameres fused in basal 2 / 3, basal piece of tegmen narrow, tegminal apodeme 0.5 x as long as apodeme of median lobe (Fig. 3 B). Sternite 8 small, rod-shaped. Apodeme of sternite 9 thick, 7.3 x longer than lamella (Fig. 3 C). Female genitalia: Coxites strongly sclerotized, with strongly sclerotized median baculi, evenly covered with pores, sensilla absent. Styli well developed, rod-shaped, bearing two setae (Fig. 3 E). Spermatheca moderately sclerotized, with short ramus; collum slender, 9 x longer than ramus. Corpus small, not swollen. Cornu slender, extended beyond corpus (Fig. 3 D). Tergite 8 subtrapezoid, fringe transformed to group of multiple, tenuous, short setae on posterior margin; lamina with numerous sclerotized fields (Fig. 3 G). Sternite 8 thick; lamella strongly sclerotized, knife-shaped, sharply narrowed and acute apically, posterior margin without setae; apodeme thick; caput strongly developed, club-shaped (Fig. 3 F).	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	distribution	Distribution. Tanzania: Udzungwa Mountains (Udzungwa Scarp) (Fig. 6).	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	biology_ecology	Ecology. The series of Sphrigodellus griseus was collected by Nikolai Scharf, Line Sørensen and colleagues during fieldwork by the ZMUC team in an undisturbed primary montane forest at 1800 – 1900 m southeast of Masisiwe village, above Kihanga stream, Udzungwa Scarp Forest Reserve, Iringa District, Tanzania (coordinates of the plots - 8.368250, 35.978222). The canopy fogging method using K- 10 Standard Thermal fogger was implemented (Sørensen 2004). The dominant trees of the canopy were: Parinari excelsa Sabine, Aphloia theiformis (Vahl) Bennett, Agauria salicifolia (Lam.) Hook., Allanblackia spp., and Albizia gummifera (Gmelin) Smith with a mixture of a few Tabernaemontana spp. (Sørensen 2004).	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	materials_examined	Material examined. Holotype, male (ZMUC) Tanzania, Iringa distr., Uzungwa Scarp Forest Reserve, 11 km SE Masisiwe village, 1800 m, S 8 ° 22 ′ E 35 ° 58 ′, 17 – 27. v. 1997 ZMUC / SI exp. / Zool. Mus. Copenhagen, Canopy Fogging, 27 May 1997, Fog. MA: 11 FB. Paratypes (13 specimens): 4 m, 8 f (ZMUC), same data as holotype; 1 f (ZMUC) Tanzania, Iringa distr., U [d] zungwa Scarp Forest Reserve, 11 km SE Masisiwe village, 1800 m, S 8 22 E 35 58, 17 – 27. v. 1997 ZMUC / SI exp. / Zool. Mus. Copenhagen, Canopy Fogging, 26 May 1997, Fog. MA: 10 DD.	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9AD455FF58FF08FAEFE49F.taxon	etymology	Etymology. The name is a Latin adjective describing the coloration of the body: “ griseus ” — grey.	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	description	(Figs 1 C – D, 4, 5, 6) www / zoobank. org / urn: lsid: zoobank. org: act: 748933 DF- 5 B 62 - 4 B 60 - B 7 ED- 9 E 731 EB 1670 F	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	diagnosis	Diagnosis. This species partly resembles minute species of the genus Dicasticus Pascoe, 1886 due to the well-developed granules on the disc of the pronotum, but significantly differs by gracile antennae; structure of abdominal sternite 8 of female; knife-shaped setae-less lamella, robust apodeme and well-developed caput; acute posterior margin of tergite 8 of female. It differs from all known species of the genus Sphrigodellus by black, glabrous, setiferous, flat granules on disc of the prothorax. It differs from S. lineatus Yunakov, 2012 by the presence of green scales covering intervals 1 – 2, posterior declivital portion of interval 3, and intervals 8 – 11. From S. nguruensis Yunakov, 2012, it also differs by the strongly developed transverse sulcus of rostrum, distinct median fovea, evenly convex eyes, and green 2 nd interval of elytra.	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	description	Description. Measurements. BL f: 4.65 mm, BW f: 2.3, BH f: 2, RL / RWA f: 1.17, VW / ELD f: 1.43, PL / PW f: 0.92, EL / BH f: 0.70, EL / EW f: 1.41. Vestiture: Body densely covered with overlapping rounded scales, setose. Setae of head spatulate, forming rows along lateral surfaces of epifrons (recumbent or subrecumbent) and medial to eyes (erect). Subocular row consisting of five setae. Pronotum uniformly covered with subrecumbent setae. Elytral intervals 1 – 7 each with distinct row of setae. Elytra in basal half with short subrecumbent setae (as long as 0.3 x width of interval), posterior declivity with long, erect weakly spatulate, truncate setae (as long as width of interval). Ventral side rather sparsely squamose. Abdominal sternites with dense rounded scales, metapleura densely squamose but scales not overlapping; basisternum and mesobasisternum bare. Antennal scape densely squamulate and setose, scales lanceolate, not overlapping; setae acute, subrecumbent. External surface of femora and tibiae covered with overlapping scales and subrecumbent setae; internal surface with sparse piliform scales and hairs; outer surface of distal portion of tibiae without scales; setae on femora slender, acute, setae of tibiae broad, truncate. Medial surface of male hind tibiae with vestigial grooming brush; hairs very short. Tarsi setose аnd scarcely squamose. Funicle setose; setae suberect 1.5 x longer than antennomere 8. Club densely tomentose. Coloration: Integument of body, legs and antennae dark-brown. Background scaling roseous-light-brown, with pearly shine; striped longitudinal pattern formed by light green scales. Head posteriad of eyes with green scales; subgenae partly covered with green scales. Pronotum with narrow greyish-roseous median and two sublateral longitudinal stripes; sides green. Elytra with alternate brown, roseous, and green longitudinal stripes. Intervals 1 – 2, posterior declivital portion of interval 3, and intervals 8 – 11 green; intervals 3, 5, and basal portion of 7, grey; interval 4 and basal portion of 6 roseous. Ventral side with sparse green scales. Main surface of femora with maculate pattern of brown and grey scales, proximal portion with green scales. External surface of tibiae with grey scales. Head: Rostrum parallel-sided. Pterygia not extended beyond rostrum. Antennal sockets dorsal. Antennifer partly-visible. Epifrons narrow, narrowed from base to posterior margin of antennifer, then abruptly widened apically; at the level of antennal articulation 0.62 x as wide as width of vertex, weakly convex longitudinally and transversally, with very narrow median carina almost completely concealed by scales. Transverse sulcus deep, partly concealed by dense scaling. Frons vertical, glabrous, bare, with six frontal setae. Epistome vestigial, surrounded by narrow carina, bearing two epistomal setae. Prementum with two setae. Eyes: sublateral, moderately convex, highest at mid-length [VW / ELD = 1.43]. Vertex: longitudinally and transversally convex. Median fovea strongly elongate. Antennae: Scape reaching middle of pronotum, weakly, evenly curved and widened, swollen in distal portion. Funicular antennomeres oblong; 1 st 0.85 x as long as 2 nd; 2 nd 3.50 x longer than wide; 3 rd 2.25 x longer than wide; 4 – 7 th 2 x longer than wide. Club spindle-shaped, 2.57 x longer than wide. Thorax: Prothorax transverse, evenly and slightly convex at sides, not constricted, widest at middle, bearing paired bilateral shallow depressions aside of disc. Disc with black, glabrous, scale-less, setiferous, flat granules. Size of granules decreases with distance from center of the disc. Posterior edge of pronotum truncate. Posterior setal fringe of spatulate setae, partly hidden by posterior edge of pronotum. Tergosternal and metepisternal sutures complete. Elytra: ovate, basal declivity vertical, slightly convex longitudinally and transversally. Femora: edentate, swollen in middle part. Tibiae: Protibiae weakly curved, not widened at the apex; medial margin weakly C-shaped, serrate in distal portion; denticles accompanied with narrow, dark-brown spine (Fig. 2 G). Size of the denticles decreases from mid-length of the tibia to apex. Meso- and metatibiae with medial margin dentate in distal portion. Mucro well developed, thorn-shaped, acute. Grooming brush moderately developed. Setal comb of protibia of 7 sparse black setae not extending beyond other margin of tibiae. Bevel of metatibia narrowly enclosed, setose, its surface covered with yellowish setae. Tarsi: Tarsomere 2 as long as width; tarsomere 3 with two wide lobes; tarsomere 5 of metatarsus extending beyond apical lobes of 3 rd by length of the lobes. Claws connate in basal half. Abdomen: 1 st and 2 nd ventrites fused, with distinct immovable suture. Posterior margin of 1 st ventrite straight, 5 th ventrite flat, posterior margin rounded. Female genitalia: Coxites weakly sclerotized, dorsal baculi strongly sclerotized, sensilla absent. Styli well developed, rod-shaped, each bearing 1 seta. Spermatheca moderately sclerotized, with swollen ramus, collum slender, 2 x as long as ramus; corpus slightly swollen; cornus slender, extended beyond corpus. Tergite 8 attenuate, posterior margin without setae. Sternite 8 thick; lamella strongly sclerotized, knife-shaped, sharply narrowed, acute, posterior margin without setae; apodeme thick; caput strongly developed, club-shaped (Fig. 5 C).	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	distribution	Distribution. Tanzania (Udzungwa Mountains: Ukami Forest) (Fig. 6).	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	biology_ecology	Ecology. The single specimen of Sphrigodellus marshalli was found in the early 1910 s in Ukami Forest (coordinates of the forest - 7.8834, 36.4001; now in Kilombero Nature Reserve, Udzungwa Mountains, Iringa District, Tanzania).	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	materials_examined	Material examined. Holotype, female, (NHMUK) ‘ Ukami / Germ. E. Afr. ’; ‘ Ellimenistes bellus Fst. ’; ‘ Cadoderus sp. n. ’; ‘ GAK Marshall / Coll. / B. M. 1950 - 255 ’.	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
03FDFF61FF9ED458FF58FD01FB86E116.taxon	etymology	Etymology. The name is dedicated to Sir Guy Anstruther Knox Marshall, a British entomologist who significantly contributed to the knowledge of weevils of Tropical Africa and Oriental Region.	en	Yunakov, Nikolai (2022): Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 2. Two new species of the genus Sphrigodellus Marshall, 1942 from Udzungwa Mountains, Tanzania. Zootaxa 5200 (2): 479-488, DOI: https://doi.org/10.11646/zootaxa.5200.5.5
