taxonID	type	description	language	source
03D987AA3E7E200E9F37FB56670BFCE4.taxon	diagnosis	Diagnoses Imagoes of Pogonini can be differentiated from the rest of Trechinae by the following characters: umbilical series complete, formed by approximately 12 setae disposed from the humeri to the apical region. Basal elytra border complete. Last labial and maxillary palpomeres fusiform. Tarsi glabrous and with a longitudinal dorsal sulcus.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7E200E9F37FB56670BFCE4.taxon	description	Description Length between 4.0 and 8.0 mm from the anterior margin of the labrum to the apex of the elytra. Tegument glabrous; metallic in most species, although in some it is totally or partially depigmented. Microsculpture polygonal. Head with eyes protruding, medium- or large-sized and in general convex. Neck absent or poorly marked. Front wide, with two supraocular sulci more or less long and deep depending on the genus (very poorly marked in Sirdenus grayii). Two supraocular setae. Antennae long and filiform, pilose from the anterior half of the second antennomere; last antennomeres slightly wider than the first ones. A pair of clypeal setae; 6 dorsal setae in the anterior margin of the labrum (the external ones are longer than the inner ones). Mandibles with a small and thin seta inserted in the mandibular sulcus. Last labial and maxillary palpomeres fusiform, with small setae (except in the apical ones). Ligula with a pair of setae. Labium not fused with the prebasilar, with a central and conspicuous bifid-tooth provided with a pair of fine setae in the base. Pronotum clearly wider than the head (except in S. grayii), convex and of variable morphology: cordiform or sub-cordiform and more or less transverse; the lateral sinuosity and the posterior narrowing are more marked in some species than in others. Hind and posterior angles of the pronotum in general protruding; the lateral sulcus is narrow, slightly more explanate in some species. The pronotum has two pairs of setae, one inserted at its widest region and another inserted near the hind angles. Basal foveae in general large and limited by a carinula which presents different grades of development between the species (absent in S. grayii). Base of pronotum depressed, punctured (punctation is more marked in some species than in others) and almost rectilinear, always coincident to the basal elytra border. Elytra, in general, convex and parallel or sub-parallel. Shoulders not or hardly protruding; functional wings. Basal and lateral elytra border complete. Scutellum present. 8 th elytral striae punctured and well-marked until the apical region, where they are frequently very tenuous or obliterated; punctuation more or less deep depending on the species. Basal striole punctured, present between the elytral suture and the first stria. Umbilical series complete from the humeral to the apical region, formed by approximately 12 setae; the first 3 – 4 setae are inserted in the lateral sulcus while the rest are inserted along the 8 th stria. The 1 st stria presents a setigerous pore in its base. Up to 5 setigerous pores disposed along the 3 rd stria in all species (commonly 3); setigerous pores present also in the 5 th and / or 7 th striae in Pogonus (Pogonoidius) meridionalis. Fine preapical setae present at the end of the 2 nd and 5 th striae. Legs, in general, slender; of a different colour from the body in some cases. Tarsus with a longitudinal dorsal sulcus; protarsus of the males with the first and second tarsomeres dilated, pilose and provided with adhesive phanerae. Prosternum glabrous in Pogonus and Pogonoidius, pilose in Sirdenus. Metepimerum visible. Last abdominal sternite with 2 setigerous pores in the males and 4 in the females. Aedeagus with median lobe asymmetrical, and the basal orifice located on its right side. Form and length of the median lobe variable between species, with the apex, in general, rounded and directed down. The internal sac of the median lobe is covered by small squamae, visible under the microscope; in the resting state it is possible to observe folds and sclerotised pieces which are variable on its form and extension between the species and groups. Left and right parameres unequal, both of them with at least one seta inserted in the distal region (the number could vary between individuals and species). Left paramere well-developed, with a triangular or sub-triangular form. Right paramere reduced, narrow and more or less elongated; in some cases, it is elbowed or curved.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7E200E9F37FB56670BFCE4.taxon	distribution	Distribution The tribe Pogonini has a worldwide distribution; it is present in all zoogeographical regions, with the main diversity of species in the Palaearctic (Grebennikov & Bousquet 1999; Choi et al. 2015; Bousquet 2017).	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7E200E9F37FB56670BFCE4.taxon	discussion	Remarks The tribe Pogonini is a very well supported monophyletic group within the subfamily Trechinae, closely related to Tachyini Motschulsky, 1862, Bembidiini, Anillini and other tribes than with Trechini (Maddison & Ober 2011; Maddison 2012; Maddison et al. 2019). The species of Pogonini show, in general, predaceous, thermophilous and diurnal behaviours (Rueda & Montes 1987), although it is also possible to find them developing a sublapidicolous life (Mateu 1947). Detailed phenology and larval instars are unknown for most Pogonini (Matalin & Makarov 2008), but some data are available for a small number of species (Jeannel 1941; see Grebennikov & Bousquet 1999). Maybe the best-documented feature of the species of Pogonini is their strict relationship with saline environments. This trait, combined with the extended hygrophilous character of all Trechinae (Ortuño & Toribio 2005: 17 – 18), implies that the most common habitats of Pogonini are the coastal regions, salt marshes and saline continental waters. For this reason, pogonines have been widely considered as halobiont elements (Bousquet & Laplante 1997). It was suggested that this specialised lifestyle derived from the basal hygrophilous condition of the subfamily, and it was originated as a response to intense interspecific competition with other riparian relatives (Rueda & Montes 1987). The results of breeding Pogonini species in laboratory obtained by Grebennikov & Bousquet (1999) suggested that the halobiont condition of these ground beetles is shared both for the pre-imaginal and the imaginal stages. With respect to the study area, although the term “ halobiont ” has been applied with different criteria by Iberian entomologists (Vives & Vives 1978 a; Ortiz et al. 1989), a high grade of specialization and adaptation to the life in saline habitats (sometimes extreme saline) have been recognised for the local species (Vives & Vives 1981, 1986; Rueda & Montes 1987; Serrano et al. 1990; Novoa et al. 1998; Andújar et al. 2001, 2002; Pardo et al. 2008; among others).	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B200D9F66FB636043FA74.taxon	type_taxon	Type species Pogonus grayii Wollaston, 1862.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B200D9F37FC7C603CFB7E.taxon	type_taxon	Type species Pogonus filiformis Dejean, 1828.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B20039CA4FA6A670BFECF.taxon	description	Figs 1 A, 3 A, 4 A	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B20039CA4FA6A670BFECF.taxon	description	Tegument partially depigmented, testaceous. Pronotum longitudinal, as wide as or barely wider than the head. Elytra parallel and body almost cylindrical. Up to 4 setigerous pores with short setae along the 3 rd interstria. Legs shorter, in relation to the body, than in the rest of species.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B20039CA4FA6A670BFECF.taxon	materials_examined	Material examined PORTUGAL • 5 specs; Ribatejo, Herdade do Pinheiro; C. de Barros leg.; MNCN. SPAIN • 22 specs; Alicante, Albufera de Elche; Apr. 1934; C. Bolívar leg.; MNCN • 138 specs; same collection data as for preceding; no date; MNCN • 1 spec.; Alicante, Calpe; 7 Apr. 1918; Moroder leg.; MNCN • 3 ♂♂, 2 ♀♀; Alicante, Urbanova; 17 Apr. 1988; V. M. Ortuño leg.; VMO • 2 ♂♂, 1 ♀; same collection data as for preceding; 21 Apr. 1988; VMO • 2 ♂♂, 3 ♀♀; same collection data as for preceding; 28 Apr. 1988; VMO • 2 ♂♂, 1 ♀; same collection data as for preceding; 25 May 2001; VMO • 3 specs; Almería, Salinas de Cabo de Gata; 2 May 1842; E. Zarco leg.; MNCN • 1 spec.; Cádiz, San Fernando; 2 Jul. 1976; J. Ramírez leg.; MNCN • 2 specs; same collection data as for preceding; 13 Jul. 1976; J. Ramírez leg.; MNCN • 4 ♂♂, 8 ♀♀; Guadalajara, Santamera; 10 Aug. 2021; J. Muñoz-Santiago, V. M. Ortuño and E. Andrés Gómez leg.; JMS • 1 ♂, 2 ♀♀; same collection data as for preceding; 19 Aug. 2021; J. Muñoz-Santiago leg.; JMS.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B20039CA4FA6A670BFECF.taxon	description	Description Habitus (Fig. 1 A). Length: 4.0 – 5.0 mm. Head as wide as, or barely wider than the pronotum; supraocular sulcus short. Pronotum longitudinal, scarcely narrowed in its posterior region and with a long and smooth lateral sinuosity; hind angles obtuse, little or nothing protruding. Base of the pronotum without punctation; foveae little deep, no limited by a carinula but a longitudinal cleft. Elytra slightly wider than the pronotum. Striae with superficial punctation. The 9 th interstria is wider than the rest. Legs with a similar colour to the tegument. Prosternum covered by fine setae. Median lobe of the aedeagus rather short, with a straight base and the apex scarcely directed down (Fig. 3 A). Chorology The genus, subgenus and species S. grayii are distributed principally in the Palaearctic Region (Europe, Asia and North Africa) but also in the Afrotropical Region (Bousquet 2017). Sirdenus grayii also occurs in the Canary Islands and Cyprus (Putzeys 1874; Jeanne 1986). Ibero-Balearic distribution (Fig. 4 A): in the Iberian Peninsula, the species is principally represented along the Mediterranean coast, but it is also present at some meridional Atlantic coastal localities. The interior records correspond to saline lagoons and saltmarshes from the Ebro valley (NE Spain) and the Southern Sub-plateau (SE Spain). It is known, with no precise records, from Los Monegros Desert (NE Spain) and other saline environments of Aragon (NE Spain; Vives & Vives 1978 a, 1981, 1999; Vives 1998), Toledo (SE Spain; Vives & Vives 1981) and Alicante (E Spain; Vives & Vives 1978 a). We also examined collection material labelled as coming from “ Herdade do Pinheiro // Ribatejo ”, an inland locality from Portugal crossed by the Tagus River. In the Balearic Islands, Sirdenus grayii is present in Mallorca.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7B20039CA4FA6A670BFECF.taxon	biology_ecology	Biology Sirdenus grayii is present both in interior and coastal regions (Bolívar 1919; see Vives & Vives 1978 a, 1981). This species is frequently associated with plants of the Salicornioideae and Arthrocnemetea groups (Sauleda 1985). The maximum activity period principally occurs in spring, but it has been found from March to July (Sauleda 1985; Andújar et al. 2002). Sirdenus grayii develops a sub-lapidicolous to endogean life (Machado 1992): it occupies abandoned galleries excavated by some species of the genus Bledius Leach, 1819 (Coleoptera, Staphylinidae). Its morphological appearance – tegument partially depigmented, cylindrical and stylised body, relatively short legs – could be interpreted as adaptations of a gallery-dweller (Sauleda 1985). Bibliographic Ibero-Balearic records Aistleitner & Lencina 2018; Andújar et al. 2002; Baehr 1988; Bolívar 1919; Español 1964; Jeanne 1968; Putzeys 1874; Sauleda 1985; Serrano et al. 2015, 2021; Vives & Vives 1978 a; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520039F0FFE0B636AFAE3.taxon	type_taxon	type species Pogonus gracilis Dejean, 1828.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520039F0FFE0B636AFAE3.taxon	type_taxon	Type species Pogonus testaceus Dejean, 1828. Diagnoses Supraocular sulcus short and shallow, ending at the level of the anterior supraocular seta or barely surpassing it. Pronotum cordiform or sub-cordiform; elytra parallel or sub-parallel. Up to 3 setigerous pores along the 3 rd interstria; 9 th interstria wider than the 8 th.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520039F0FFE0B636AFAE3.taxon	description	Description Length: 4.0 – 6.0 mm. Tegument testaceous or brown (in the last case, with bronze sheen). Palps yellow, lighter than the testaceous legs and antennae. Pronotum little or nothing transverse; the anterior angles may be protruding or not, and occasionally have with tiny setae. Lateral sulcus of the pronotum narrow; hind angles hardly conspicuous. Base of the pronotum as wide as or less wide than the base of the elytra. Elytra generally convex. Legs testaceous, never darker than the body tegument. Prosternum covered by fine setae. Median lobe of the aedeagus slightly arcuate and clearly directed down. Chorology The genus is represented over the whole Palaearctic Region (Bousquet 2017). The Ibero-Balearic representatives of Pogonistes are distributed along the Mediterranean coast of the peninsula, in the saline environments of the Ebro valley and the Southern Sub-plateau and in the Balearic Islands.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520009F12FA176372FC3D.taxon	description	Figs 1 B, 3 B, 4 B	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520009F12FA176372FC3D.taxon	description	Tegument brown with bronze sheen; legs and antennae testaceous, contrasting with the rest of the body. Pronotum cordiform, with a marked sinuosity; anterior angles with 1 – 3 short and fine setae. Elytra subparallel slightly convex.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520009F12FA176372FC3D.taxon	materials_examined	Material examined SPAIN • 2 ♂♂, 2 ♀♀; Álava, Laguna de Carravalseca; 23 Mar. 1995; J. M. Marcos leg.; VMO • 1 ♂; Alicante, Calpe; 10 Apr. 1989; J. M. Beltrán leg.; VMO • 1 ♂; Alicante, Elche, El Hondo; 6 Mar. 1988; J. M. Beltrán leg.; VMO • 1 ♂; Alicante, Salinas de Calpe; 19 Apr. 1934; C. Bolívar leg.; MNCN • 9 ♂♂, 2 ♀♀; Alicante, Urbanova; 21 Apr. 1988; VMO • 1 ♀; same collection data as for preceding; 28 Apr. 1988; VMO • 2 ♂♂, 1 ♀; same collection data as for preceding; 2 Jun. 1995; V. M. Ortuño leg.; VMO • 1 ♀; same collection data as for preceding; 16 May 1996; VMO • 1 ♂; same collection data as for preceding; 25 May 2001; VMO • 1 ♂; Alicante, Santa Pola; 2 Mar. 1983; T. G. Sempere leg.; VMO • 1 ♂; Cádiz, Conil de la Frontera, El Palmar; 29 Aug. 2001; J. Navarro leg.; VMO • 1 ♂, 1 ♀; Huelva; Cardiel leg.; MNCN • 2 ♂♂, 1 ♀; Málaga, Fuente de Piedra; 3 Jun. 1987; J. Barandica leg.; VMO • 25 ♂♂, 34 ♀♀; Toledo, Villacañas; 22 May 1934; C. Bolívar leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520009F12FA176372FC3D.taxon	description	Description Habitus (Fig. 1 B). Length: 4.0 – 5.0 mm. Head with a similar colour to the body tegument. Anterior angles of the pronotum obtuse and barely protruding. Base of the pronotum clearly narrower than the anterior margin of the elytra; the punctuation of the base is hidden by longitudinal clefts, so the surface of this region presents a striated appearance. Elytra punctuation deep. Median lobe of the aedeagus (Fig. 3 B) narrower and with the apex more developed than in P. testaceus. Chorology Mediterranean coast of South Europe, from Spain to Greece (including Malta) and North Africa (Compte Sart 1968; Magrini & Schembri 1997; Bousquet 2017). Ibero-Balearic distribution (Fig. 4 B): in the Iberian Peninsula, the species is present discontinuously along the Mediterranean coast, from Cádiz to Tarragona (Spain), and in saline localities of the Ebro valley (NE Spain) and the Southern Sub-plateau (SE Spain). The species has been reported, with no precision, from Algarve (SW Portugal; Zaballos & Jeanne 1994), Los Monegros Desert (NE Spain; Vives & Vives 1978 a, 1981, 1999), saltmarshes of Alicante (E Spain; Vives & Vives 1978 a) and a saltmarsh in Toledo (SE Spain) (Vives & Vives 1981). We examined collection material from Huelva (SW Spain, with no precise locality), a territory from which P. gracilis has never previously been reported. In the Balearic Islands, Pogonistes gracilis is present in Mallorca, Menorca and Ibiza.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7520009F12FA176372FC3D.taxon	biology_ecology	Biology The species inhabits coastal and interior regions equally (Bolívar 1919; see Vives & Vives 1978 a, 1978 b, 1981). Imagoes are active during spring, principally from March to July, but some records are dated from other months such as November (Andújar et al. 2002). Pogonistes gracilis can be found in direct contact with water when it takes refuge under the rocks on the shores, in damp and saline environments where it occurs (Ortuño & Marcos 2003). Bibliographic Ibero-Balearic records Aistleitner & Lencina 2018; Andújar et al. 2002; Baehr 1988; Compte Sart 1968; Jeanne 1968, 1978; Ortiz et al. 1989; Ortuño & Marcos 2003; Serrano et al. 1990, 2015, 2021; Vives & Vives 1986, 1978 a, 1978 b.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7620079F56FC3A6165FD6D.taxon	description	Figs 1 C, 3 C, 4 C	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7620079F56FC3A6165FD6D.taxon	description	Tegument testaceous. Pronotum sub-cordiform, scarcely narrowed in its posterior region (smooth sinuosity); anterior angles protruding. Elytra parallel and convex, sub-cylindrical, with the posterior half slightly depigmented.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7620079F56FC3A6165FD6D.taxon	materials_examined	Material examined SPAIN • 1 ♂; Vuillefroy leg.; Pérez Arcas coll., MNCN • 1 ♂, 2 ♀♀; Alicante, Albufera de Elche; Apr. 1934; C. Bolívar leg.; MNCN • 3 ♂♂, 1 ♀; Alicante, Santa Pola; 29 Apr. 1985; T. G. Sempere leg.; VMO • 2 ♀♀; 17 Jan. 1983; VMO • 3 ♂♂; Valencia, Los Valles; Morote leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7620079F56FC3A6165FD6D.taxon	description	Description Habitus (Fig. 1 C). Length: 5.5 – 6.0 mm. Tegument with some regions more pigmented than others. Head brown, slightly darker than the rest of the body. Pronotum with the base as, or almost as, wide as the anterior margin of the elytra and with well-visible punctuation. Elytral striae with almost superficial punctutation, considerably more tenuous than in other species of the tribe. Legs slightly lighter than the rest of the body. Median lobe of the aedeagus (Fig. 3 C) wider than in P. gracilis and with the apex less developed. Chorology Species distributed along the West Mediterranean: the subspecies P. testaceus testaceus is present in North Africa (records from Argelia), the Iberian Peninsula and along the French coast (Español 1964; Bousquet 2017); the subspecies P. testaceus graecus (Apfelbeck, 1904) is known from Bulgaria and Greece (Bousquet 2017). Ibero-Balearic distribution (Fig. 4 C): in the Iberian Peninsula, the species is present along the Mediterranean coast ranging from Tarragona to Alicante (Spain). On the Balearic Islands, it is present in Menorca and Ibiza.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7620079F56FC3A6165FD6D.taxon	biology_ecology	Biology It was suggested that this species prefers the most high-saline environments (Sauleda 1985), which could explain why its distribution in the Iberian Peninsula is restricted to the Mediterranean coast, where it can be a common element (Español 1964). Imagoes present all year, with a peak of activity in spring (Sauleda 1985). Bibliographic Ibero-Balearic records Aistleitner & Lencina 2018; Compte Sart 1968; Español 1964; Jeanne 1968, 1978; Sauleda 1985; Serrano et al. 2015, 2021; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7120079F34FD69663EF836.taxon	type_taxon	Type species Carabus littoralis Duftschmid, 1812. Diagnoses Tegument with metallic sheen, at least in the head and pronotum. Supraocular sulcus deep and long, surpassing the level of the anterior supraocular seta and ending close to the posterior one. Pronotum always wider than the head, with different shape among species. Up to 5 setigerous pores along the 3 rd interstria (commonly 3). 8 th and 9 th interstriae of the same width.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7120079F34FD69663EF836.taxon	description	Description Length: 5.0 – 8.0 mm. Tegument with variable coloration from green to bronze (not always uniform between the parts of the body). Antennae and palps yellow, testaceous or brown. Pronotum cordiform or sub-cordiform, in general convex and more transverse in some species than in others. Anterior angles of the pronotum glabrous or with fine setae. Base as, or almost as, wide as the anterior margin of the elytra, with a marked and deep punctuation that is limited by a well-developed carinula. Except in P. riparius and P. meridionalis, the base is oblique near the hind angles, so they are obtuse and more or less protruding. Elytra parallel, sub-parallel or sub-oval, convex and wider than the pronotum. In all the species, the punctuation of the elytral striae is deep, except in the apical region. Interstriae other than the 3 rd glabrous, except in P. meridionalis, which also has setigerous pores in the 5 th and 7 th interstriae. Legs yellow, testaceous or brown. Median lobe of the aedeagus curved or arcuate, with the apex clearly directed down, except in P. littoralis. Sclerotised pieces of the internal sac rather massive in some species. Chorology Genus widely distributed in all geographic regions except the Neotropical; the Palaearctic Region is where it is best-represented (Bousquet & Laplante 1997; Bousquet 2017). Species of Pogonus are present both along the Atlantic and Mediterranean coasts of the Iberian Peninsula, the saline environments of its continental interior and in the Balearic Islands.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020069F1AFE9E6017FE43.taxon	type_taxon	type species Pogonus punctatulus Dejean, 1828.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020069F1AFE9E6017FE43.taxon	type_taxon	Type species Pogonus meridionalis Dejean, 1828.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020049CB4FD8860A1FC9A.taxon	description	Figs 1 D – E, 3 D, 4 D	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020049CB4FD8860A1FC9A.taxon	description	Tegument bronze in most Ibero-Balearic individuals, but blue with metallic sheen in the southern populations of the peninsula. Pronotum markedly convex, with the base clearly wider than the anterior margin and small setae inserted in the anterior angles. Elytra sub-oval and slightly convex. Setigerous pores present in the 5 th, 7 th or both interstriae, in addition to those present in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020049CB4FD8860A1FC9A.taxon	materials_examined	Material examined SPAIN • 1 ♂, 1 ♀, 2 specs; Pozuelo; MNCN • 2 ♂♂, 1 ♀; Alicante, Elche, El Hondo; 6 Mar. 1988; J. M. Beltrán leg.; VMO • 1 ♂, 1 ♀; same collection data as for preceding; 9 Mar. 1991; VMO • 3 ♂♂, 2 ♀♀; Alicante, ctra. [road] La Marina-Elche (km 8); 23 Mar. 1996; J. M. Beltrán leg.; VMO • 3 ♂♂, 5 ♀♀; Alicante, Salinas, Laguna de Salinas; 20 Feb. 1993; J. M. Beltrán leg.; VMO • 5 ♂♂, 5 ♀♀; same collection data as for preceding; 14 May 1993; VMO • 1 ♀; Alicante, Santa Pola; 2 Mar. 1983; T. G. Sempere leg.; VMO • 2 ♂♂; Cádiz, El Palmar, Conil de la Frontera; 29 Aug. 2001; J. Navarro leg.; VMO • 2 ♂♂, 2 ♀♀; Cádiz, Jerez; Lauffer leg.; MNCN • 1 ♂; same collection data as for preceding; L. Cepero leg.; MNCN • 1 ♂; Cádiz, Puerto de Santa María; Dieck leg.; MNCN • 1 ♂, 1 ♀; Lauffer leg.; MNCN • 3 ♂♂, 1 ♀; Ciudad Real, Pozuelo de Calatrava; La Fuente leg.; MNCN • 1 ♂, 1 ♀; same collection data as for preceding; no leg.; MNCN • 1 ♂; Cuenca, Uclés; Pantiel leg.; MNCN • 1 ♀; Huelva; May 1911; Cendrero leg.; MNCN • 2 ♂♂; Madrid, Colmenar de Oreja, Laguna de las Esteras; 8 Jul 2020; J. Muñoz-Santiago leg.; JMS • 1 ♀; Murcia, Cartagena; 5 Jan. 1924; Escalera leg.; MNCN • 2 ♂♂, 4 ♀♀; 7 Jan. 1925; MNCN • 2 ♂♂, 1 ♀; same collection data as for preceding; 22 Jan. 1905; G. Schramm leg.; MNCN • 2 ♂♂; same collection data as for preceding; Dec., no year; MNCN • 1 ♂, 2 ♀♀; same collection data as for preceding; Lauffer leg.; MNCN • 1 ♀; same collection data as for preceding; Ehlers leg.; MNCN • 1 ♀; same collection data as for preceding; Bolívar leg.; MNCN • 2 ♀♀; Toledo, Quero; 22 May 1929; Escalera leg.; MNCN • 1 ♂; 15 May 1999, Lauffer leg.; MNCN • 1 ♀; same collection data as for preceding; no date; MNCN • 3 ♂♂, 2 ♀♀; same collection data as for preceding; 22 May 1934; C. Bolívar leg.; MNCN • 1 ♂, 1 ♀; same collection data as for preceding; no date; no leg.; MNCN • 6 ♂♂, 5 ♀♀; Toledo, Quero, Laguna Grande; 9 Apr. 1993; V. M. Ortuño leg.; VMO • 1 ♂; Toledo, Quero, Laguna de Peña Hueca; 10 Jun. 1986; J. P. Zaballos leg.; VMO • 1 ♂, 2 ♀♀; Toledo, Villacañas, Salinas de Peña Hueca; 25 Apr. 1997; J. M. Beltrán leg.; VMO • 1 ♂; Toledo, Villacañas; 20 May 1934; C. Bolívar leg.; MNCN • 1 ♀; same collection data as for preceding; no date; C. Bolívar leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020049CB4FD8860A1FC9A.taxon	description	Description Habitus (Fig. 1 D – E). Length: 5.5 – 7.0 mm. Antennae and palps dark brown, lighter at the base of antennomeres (from the second to the last) and at the apex of the last palpomeres. Pronotum with short lateral sinuosity and the base sub-rectilinear; lateral sulcus narrow; hind angles well marked and straight or slightly sharp. Legs brown, with the femora darkened (in southern populations, almost the entire legs darkened). Median lobe of the aedeagus arcuate, with the apex rounded and not narrowed (Fig. 3 D). Chorology The subgenus Pogonoidius is well represented in the Palaearctic Region (Bousquet 2017). The species P. meridionalis is a Palaearctic element with a wide distribution from the Iberian Peninsula to Siberia (Bousquet 2017). Ibero-Balearic distribution (Fig. 4 D): in the Iberian Peninsula, the species is present in the meridional extreme, in the Segura basin (SE Spain), some localities of the Catalonian coast and in lagoons and saltmarshes of the Southern Sub-plateau (SE Spain) and Central Spain. It has been reported from Portugal, with no precise locality, by Serrano (2002).	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7020049CB4FD8860A1FC9A.taxon	biology_ecology	Biology The species can be found in saline areas colonised by plants of the Arthrocnemetea group and the genus Salicornia L. (Sauleda 1985; Vigna Taglianti & Bonavita 1994; J. Muñoz-Santiago pers. obs.) but also in non-saline environments, as it is less halobiont than other species of the genus (Sauleda 1985). This suggestion is supported by the presence of the species in the Embalse de Valmayor (Madrid, Spain; Ortuño & Toribio 1996), a freshwater body. The phenology of a population of P. meridionalis in Elton lake (Volgograd, Russia) was studied by Matalin & Makarov (2008): it had a peak of imaginal activity on spring (April – May), and the imagoes which emerged on one year coexisted with imagoes from the previous year, so it is not an annual species. The large number of imagoes in spring was also observed by Sauleda (1985). Bibliographic Ibero-Balearic records Aistleitner & Lencina 2018; Andújar et al. 2002; Baehr 1986; Cuní 1885; Jeanne 1968; López-Pérez et al. 2014; Mateu 1947; Ortuño & Toribio 1996; Prieto 2018; Sauleda 1985; Serrano et al. 2021; Vives & Vives 1976; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220049F08FCDF6199F983.taxon	type_taxon	type species Pogonus smaragdinus Waltl, 1835.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220049F08FCDF6199F983.taxon	type_taxon	type species Pogonus reticulatus Schaum, 1857.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220049F08FCDF6199F983.taxon	type_taxon	type species Pogonus riparius Dejean, 1828.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220049F08FCDF6199F983.taxon	type_taxon	Type species Carabus littoralis Duftschmid, 1812. Diagnoses Tegument with metallic sheen (at least in the head and pronotum). Up to 5 setigerous pores in the 3 rd interstria (commonly 3); the rest of interstriae are glabrous.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220049F08FCDF6199F983.taxon	description	Description Length: 5.0 – 8.0 mm. Tegument generally green or bronze, uniform in most of the species; in P. luridipennis and P. pallidipennis the elytra are a testaceous. Anterior angles of the pronotum glabrous. Base of the pronotum slightly curvilinear.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220049F08FCDF6199F983.taxon	distribution	Distribution The distribution range of the subgenus is similar to that of the genus.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220199C85F9C860B0FE37.taxon	description	Figs 1 F, 3 E, 4 E	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220199C85F9C860B0FE37.taxon	description	Tegument bronze or blue with metallic sheen. Pronotum cordiform; anterior region clearly punctured. Elytra generally wide and sub-oval. Up to 3 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220199C85F9C860B0FE37.taxon	materials_examined	Material examined PORTUGAL • 1 ♂; P. d’Oliveira leg.; Pérez Arcas coll.; MNCN • 4 ♀♀; Algarve, Faro; Paulino leg.; MNCN. SPAIN • 1 ♂; [A] Coruña; Bolívar leg.; MNCN • 1 ♂, 1 ♀; A Coruña, Boiro, Playa Mañons; 25 Jul. 1992; V. M. Ortuño leg.; VMO • 2 ♂♂; Albacete, Laguna de Pétrola; 18 Jul. 1989; J. M. Beltrán leg.; VMO • 2 ♂♂, 2 ♀♀; Albacete, Laguna del Salobralejo; 15 May 1999; J. M. Beltrán leg.; VMO • 1 ♂; Alicante, Calpe; 30 Dec. 1984; J. M. Beltrán leg.; VMO • 1 ♀; same collection data as for preceding; 21 Dec. 1985; VMO • 1 ♂; Alicante, Elche, El Hondo; 3 May 1986; J. M. Beltrán leg.; VMO • 1 ♂; same collection data as for preceding; 9 Mar. 1991; VMO • 2 ♂♂, 1 ♀; Alicante, Salinas, Laguna de Salinas; 14 May 1993; J. M. Beltrán leg.; VMO • 3 ♀♀; same collection data as for preceding; 26 Mar. 1994; VMO • 3 ♂♂, 2 ♀♀; Alicante, Salinas de Calpe; 22 Apr. 1934; C. Bolívar leg.; MNCN • 2 ♂♂; Alicante, Urbanova; 21 Apr. 1988; V. M. Ortuño leg.; VMO • 1 ♀; same collection data as for preceding; 23 Apr. 1988; VMO • 1 ♀; same collection data as for preceding; 28 Apr. 2002; VMO • 1 ♀; Alicante, Santa Pola; 17 Jan. 1983; T. G. Sempere leg.; VMO • 2 ♂♂; Alicante, Salinas de Santa Pola; 11 Feb. 1980; T. G. Sempere leg.; VMO • 1 ♀; Alicante, Santa Pola, srra. [mountain range]; 6 Mar. 1994; J. M. Beltrán leg.; VMO • 1 ♀; same collection data as for preceding; 24 Dec. 1994; VMO • 1 ♀; Alicante, Guardamar del Segura, Segura River; 30 Mar. 1996; J. M. Beltrán leg.; VMO • 1 ♂, 1 ♀; Barcelona, Faro de Llobregat; Oct. 1987; Bolívar leg.; MNCN • 1 ♂, 1 ♀; Cádiz; 1918; Smith leg.; MNCN • 1 ♂, 1 ♀; Cádiz, Conil de la Frontera, El Palmar; 29 Aug. 2001; J. Navarro leg.; VMO • 72 specs; Cádiz, Jerez, Laguna de Medina; 8 May 1982; Ramírez leg.; Ferrer coll.; MNCN • 2 specs; Cádiz, Palmones; 29 Apr. 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 6 Jul. 1976; J. Ramírez leg.; MNCN • 4 specs; Cádiz, Tarifa, Punta Paloma; 6 Jun. 1998; Ferrer coll; MNCN • 2 specs; Cádiz, San Fernando; 13 Jun. 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 15 Apr. 1976; J. Ramírez leg.; MNCN • 1 spec.; Cádiz, San Roque; 20 Jul 1974; J. Ramírez leg.; MNCN • 1 spec.; Cádiz, Zahara de los Atunes; 24 Apr. 1976; J. Ramírez leg.; MNCN • 4 ♂♂; Ciudad Real; Pozuelo de Calatrava; MNCN • 5 ♂♂, 1 ♀; Cuenca, Uclés; Pantel leg.; MNCN • 2 ♂♂; same collection data as for preceding; Lauffer leg.; MNCN • 1 ♀; same collection data as for preceding; no leg.; MNCN • 1 ♂; same collection data as for preceding; Var. leg.; MNCN • 1 ♂; Girona, Aiguamolls del Emporda; 25 Apr. 1990; V. M. Ortuño leg.; VMO • 1 ♀; Guadalajara, Imón, Salinas de Imón; 10 Aug. 2021; J. MuñozSantiago, V. M. Ortuño and E. Andrés Gómez leg.; JMS • 1 ♀; Guadalajara, Paredes de Sigüenza; 8 Mar. 1985; V. M. Ortuño leg.; VMO • 1 ♂, 1 ♀; Guadalajara, Santamera; 10 Aug. 2021; J. Muñoz-Santiago, V. M. Ortuño & E. Andrés Gómez leg.; JMS • 1 ♂, 1 ♀; same collection data as for preceding; 19 Aug. 2021; J. Muñoz-Santiago leg.; JMS • 1 spec.; Málaga, Sierra de las Nieves; 22 Jun. 1966; Ferrer coll.; MNCN • 1 ♀; Mallorca; Mas de Xaxars leg.; MNCN • 1 ♂, 1 ♀; Mallorca, Palma; Lauffer leg.; MNCN • 2 ♂♂; same collection data as for preceding; Moraguer leg.; MNCN • 2 ♂♂; Mallorca, Pollensa; H. J. Jorda leg.; MNCN • 1 ♂; Sevilla; Marvier leg.; MNCN • 1 ♀; Soria; Pérez Arcas coll.; MNCN • 1 spec.; Toledo, Quero; 12 May 1899; Lauffer leg.; MNCN • 2 ♂♂, 1 ♀; same collection data as for preceding; Jun. 1909; MNCN • 2 ♂♂, Toledo, Quero, Laguna Grande; 9 Apr. 1993; V. M. Oruño leg.; VMO • 2 ♂♂; Toledo, Villacañas, Laguna de Peña Hueca; 9 Apr. 1993; V. M. Ortuño leg.; VMO • 6 ♂♂, 5 ♀♀; Toledo, Villacañas; 24 May 1999; Lauffer leg.; MNCN • 1 ♂, 3 ♀♀; same collection data as for preceding; 20 May 1934; C. Bolívar leg.; MNCN • 9 ♂♂, 7 ♀♀; same collection data as for preceding; no date; MNCN • 1 spec.; same collection data as for preceding; MNCN • 1 ♂, 2 ♀♀; Madrid, Aranjuez; F. Moróder leg.; MNCN • 1 ♀; same collection data as for preceding; Sanz leg.; MNCN • 1 ♂, 1 ♀; same collection data as for preceding; Bolívar leg.; MNCN • 1 ♀; same collection data as for preceding; Lauffer leg.; MNCN • 1 ♂; same collection data as for preceding; no leg.; MNCN • 1 ♂; Madrid, Aranjuez, Laguna de Ontígola; 1 Jan. 1973; J. Serrano leg.; MNCN • 2 ♂♂; same collection data as for preceding; 2 Apr. 1975; J. Serrano leg.; MNCN • 1 ♂; Murcia, Cartagena; Dec., no year; G. Schramm leg.; MNCN • 1 ♂, 1 ♀; same collection data as for preceding; no date; no leg.; MNCN • 1 ♀; Valencia, El Puig; 11 Apr. 1987; J. M. Beltrán leg.; VMO • 1 ♂, 1 ♀; Valladolid; 29 Sep. 1921; Lezcano leg.; MNCN • 1 ♂; same collection data as for preceding; no date; Chevrolat leg.; Pérez Arcas coll.; MNCN • 2 ♂♂; Bolívar leg.; Pérez Arcas coll.; MNCN • 2 ♂♂, 3 ♀♀; Zamora, Lagunas de Villafáfila; 6 Feb. 1987; V. M. Ortuño leg.; VMO • 3 ♂♂; Zaragoza, Laguna de Gallocanta; 17 Jul 1981; J. Serrano leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220199C85F9C860B0FE37.taxon	description	Description Habitus (Fig. 1 F). Length: 5.0 – 7.0 mm. Antennae testaceous, with the distal half of the antennomere 2 and the base of the rest of antennomeres lighter (sometimes the last antennomeres are entirely lightened). Palps yellow, with the base of the last palpomere darkened. Anterior angles of the pronotum more marked than in other species. Lateral sulcus of the pronotum slightly widened. Legs testaceous, sometimes slightly yellow. Median lobe of the aedeagus arcuate; the internal sac presents a characteristic thornshaped sclerotised piece whose apex is directed to the dorsum, among others (Fig. 3 E). Chorology The species is distributed along the Mediterranean and Atlantic coasts from the Iberian Peninsula and North Africa to the North Sea and the Black Sea (Magrini & Schembri 1997; Tamadouni & Arahou 2016; Bousquet 2017). The subspecies P. c. viridanus occurs in the Iberian Peninsula and, with P. c. alticola Antoine, 1955, in North Africa (Bousquet 2017). The populations of the rest of Europe correspond to the subspecies P. c. chalceus (Marsham, 1802). Ibero-Balearic distribution (Fig. 4 E): in the Iberian Peninsula, P. c. viridanus is present at several localities of the Atlantic and Mediterranean coasts and in many interior saline lagoons and saltmarshes. The species is known from Los Monegros Desert (NE Spain), with no precise records (Vives & Vives 1978 a). We could examine collection material from unpublished but imprecise localities: Sierra de las Nieves Natural Park (Málaga, SE Spain), Coruña (NW Spain), Soria (NE Spain) and Valladolid (NW Spain). In the Balearic Islands, it is present in Mallorca, Menorca and Ibiza.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E7220199C85F9C860B0FE37.taxon	biology_ecology	Biology Species that inhabits both coastal and interior regions, as long as it finds the conditions that it requires (see Vives & Vives 1978 a), where it is very common (Andújar et al. 2002). It has been typically considered as a spring / summer-reproductive species, both in the Iberian Peninsula (Cárdenas et al. 1999) and other Palaearctic regions (Matalin & Makarov 2008). Following the information provided by some authors, the imagoes are present during all or most of the year, with a peak of activity between March and August (Sauleda 1985; Andújar et al. 2002). In addition, Ortuño & Marcos (2003) remark the scarcity – not absence – of individuals during the driest months. The phenotypical variability of this species had resulted in the descriptions of several subspecies and ‘ forms’ along the geographic regions where it is present (Carret 1903; Bousquet 2017; see Machado 1992). The dispersal ability of P. chalceus depends on its wing-polymorphism status (Desender 1989; Desender et al. 1998), with flightless individuals more frequent in Atlantic than in Mediterranean populations (Desender et al. 2000). This genetic difference could be related, according to Desender et al. (2000), with the stability of the environment: the Atlantic habitats, in general, are subject to fewer alterations than the Mediterranean ones, resulting in a smaller need for colonisation of new territories and, therefore, the existence of a higher proportion of flightless individuals. The genetic features of P. chalceus and their implications in the phenotype were widely studied and discussed by Van Belleghem (2014). Bibliographic Ibero-Balearic records Aistleitner & Lencina 2018; Andújar et al. 2001, 2002; Baehr 1986; Campos Gómez & Novoa 2006; Champion 1898; Chapman & Champion 1907; Compte Sart 1968; Desender et al. 2000; Eiroa et al. 1988; Herrera & Arricibita 1990; Jeanne 1968, 1978; López-Pérez et al. 2014; Mateu 1947; Novoa et al. 1998; Ortiz et al. 1989; Ortuño et al. 1997; Ortuño & Marcos 2003; Ortuño & Toribio 1996; Prieto 2018; Prieto Piloña & Valcárcel 1996; Putzeys 1874; Ramos-Abuín 1990; Ruiz-Tapiador & Zaballos 2001; Sauleda 1985; Santos et al. 1985; Serrano 1983; Serrano et al. 1990, 2015, 2021; Viñolas et al. 2019; Vives & Vives 1978 a, 1986; Zaballos 1986 b.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6F20189F41FE2366C9FD08.taxon	description	Figs 2 A, 3 F, 4 F	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6F20189F41FE2366C9FD08.taxon	description	Tegument dark brown, with a bronze sheen. Pronotum cordiform, transverse or sub-transverse, with the disc smooth. Elytra parallel or sub-parallel; up to 5 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6F20189F41FE2366C9FD08.taxon	materials_examined	Material examined SPAIN • 3 ♂♂, 1 ♀; Alicante, Albufera de Elche; C. Bolívar leg., MNCN • 2 ♂♂, 2 ♀♀; Alicante, Elche; MNCN • 1 ♂; Alicante, Salinas de Bonmatí; 16 Aug. 1985; V. M. Ortuño leg.; VMO • 1 ♀; same collection data as for preceding; 21 Aug. 1985; V. M. Ortuño leg.; VMO • 3 ♂♂, 1 ♀; same collection data as for preceding; 21 Jul. 1986; V. M. Ortuño leg.; VMO • 6 ♂♂, 2 ♀♀; Alicante, Salinas de Calpe; 22 Apr. 1934; C. Bolívar leg.; MNCN • 3 ♂♂, Alicante, Urbanova; 21 Apr. 1988; V. M. Ortuño leg.; VMO • 1 ♂; 23 Apr. 2002; V. M. Ortuño leg.; VMO • 1 ♂; Cádiz; F. Fernando leg.; Pérez Arcas coll.; MNCN • 2 specs; Cádiz, Jerez; Laguna de Medina, 8 May 1982; Ramírez leg.; MNCN • 1 spec.; Cádiz, Palmones; 29 Apr. 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 3 May 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 5 May 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 6 Jul. 1976; MNCN • 1 spec., Cádiz, Tarifa, Río del Valle; 18 Jul. 1977; J. Ramírez leg.; MNCN • 3 spec.; Cádiz, San Fernando; 13 Jul. 1976; J. Ramírez leg.; MNCN • 6 spec.; same collection data as for preceding; 15 Apr. 1976; MNCN • 2 spec.; Cádiz, Zahara de los Atunes; 24 Apr. 1976; J. Ramírez leg.; MNCN • 1 ♂, 1 ♀; Huelva; Cardiel leg.; MNCN • 2 ♂♂; same collection data as for preceding; Lauffer leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6F20189F41FE2366C9FD08.taxon	description	Description Habitus (Fig. 2 A). Length: 5.5 – 6.0 mm. Antennae and palps almost entirely testaceous or yellow (base of the first antennomere and insertion points of the palpomeres slightly darkened). Anterior angles of the pronotum not or hardly protruding; lateral sulcus widened. Legs entirely testaceous. Median lobe of the aedeagus arcuate and narrowed progressively until the apex, which is well developed and spatulashaped (Fig. 3 F). Among the sclerotised pieces that are found in the internal sac, the presence of a thick thorn directed to the dorsum should be noted, on which a helical scaly structure is superimposed. Chorology Species with a wide Palaearctic distribution, also present in the Afrotropical Region (Ratti 1983; Sauleda 1985; Bousquet 2017) and in the Canary Islands (Machado 1992). Ibero-Balearic distribution (Fig. 4 F): in the Iberian Peninsula, the species is widely present along the Atlantic coast of the meridional extreme, but also at some localities of the Mediterranean coast. Jeanne (1968) published a record from Las Hurdes (Cáceres, W Spain) that probably is incorrect as, according to the current chorological knowledge, it would represent a very isolated population. In the Balearic Islands P. gilvipes is present in Mallorca, Menorca and Ibiza.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6F20189F41FE2366C9FD08.taxon	biology_ecology	Biology Sauleda (1985) indicates that this is an uncommon species which can be found in areas colonised by plants of Arthrocnemetea all year except winter. Some comments on the larvae of this species have been published: Machado (1992) suggested that, at coastal localities, the preimaginal stages of P. gilvipes could develop under the remains of marine algae deposited by the sea on rocky shorelines. Bibliographic Ibero-Balearic records Baehr 1986; Jeanne 1968, 1978; López-Pérez et al. 2014; Palmer & Vives 1993; Putzeys 1874; Sauleda 1985; Serrano et al. 1986, 2015, 2021; Vives & Vives 1978 a; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6E201F9CABFD4C611FFD2B.taxon	description	Figs 2 B, 3 G, 5 A	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6E201F9CABFD4C611FFD2B.taxon	description	Body tegument, and commonly first antennomere, bronze. Pronotum cordiform and sub-transverse; anterior region with tenuous punctuation. Elytra parallel or sub-parallel; up to 3 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6E201F9CABFD4C611FFD2B.taxon	materials_examined	Material examined SPAIN • 1 ♀; Alicante, Elche, El Hondo; 21 Nov. 1988; J. M. Beltrán leg.; VMO • 2 ♂♂; Alicante, Salinas de Bonmatí; 21 Jul. 1986; V. M. Ortuño leg.; VMO • 1 spec.; Cádiz, Palmones; 29 Apr. 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 6 Jul. 1976; J. Ramírez leg.; MNCN • 1 ♀; Cádiz, Conil de la Frontera; 29 Aug. 1998; J. Navarro leg.; VMO • 1 ♂, 1 ♀; Cádiz, Playa de Sancti Petri; 1 Apr. 1987; V. M. Ortuño leg.; VMO • 2 specs; Cádiz, San Roque • 20 Jul. 1974; J. Ramírez leg.; MNCN • 4 specs; Cádiz, Zahara de los Atunes; 24 Apr. 1976; J. Ramírez leg.; MNCN • 1 ♀; Huelva; Cardiel leg.; MNCN • 1 spec.; Málaga, Sierra de las Nieves; 22 Jun. 1966; Ferrer leg.; MNCN • 2 ♂♂; Mallorca; Mas de Xaxars leg.; MNCN • 1 ♂, 1 ♀; Mallorca, Pollensa; H. J. Jorda leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6E201F9CABFD4C611FFD2B.taxon	description	Description Habitus (Fig. 2 B). Length: 5.5 – 6.5 mm. Antennae brown except the first antennomere. Palps testaceous, with the basal half of the last palpomere darkened. Lateral sulcus of the pronotum slightly widened. Legs testaceous. Median lobe of the aedeagus scarcely curved, with the apex little developed (Fig. 3 G). Chorology Species widely distributed along the Palaearctic Region (Compte Sart 1968, Ratti 1983; Tamadouni & Arahou 2016; Bousquet 2017). Ibero-Balearic distribution (Fig. 5 A): in the Iberian Peninsula, it is present along the southern Atlantic coast, in the meridional extreme, at some localities of the Mediterranean coast and in saline interior environments of Albacete (SE Spain). We examined collection material from the Sierra de las Nieves National Park (Málaga, SE Spain), a new record that could not been represented in the distribution map. In the Balearic Islands, it is known from Mallorca and Menorca.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6E201F9CABFD4C611FFD2B.taxon	biology_ecology	Biology There is few published information about the biology of this species. As in the case of P. riparius, Ratti (1994) suggested that, in some populations along the Adriatic coast, the different life phases of P. littoralis can overlap in the same year, so it cannot be exclusively considered an annual species. Bibliographic Ibero-Balearic records Andújar et al. 2002; Baehr 1986; Champion 1898; Jeanne 1968, 1978; López-Pérez et al. 2014; Putzeys 1874; Serrano et al. 2015, 2021; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E69201E9CA9FD206708FDB1.taxon	description	Figs 2 C, 3 H, 5 B	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E69201E9CA9FD206708FDB1.taxon	description	Tegument bicolored: head and pronotum with metallic sheen, bluish green or bronze, while the elytra and appendices have a testaceous colour. Pronotum markedly cordiform and transverse. Elytra wide, sub-parallel and less convex than in other species. Up to 3 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E69201E9CA9FD206708FDB1.taxon	materials_examined	Material examined GERMANY • 1 ♂, 1 ♀; Hamburg-Altona, Mittelbe Sülldorf (near Magdeburg); 4 Jun. 1993; Wrase leg.; VMO. SPAIN • 1 ♂; Huelva; Cardiel leg.; MNCN • 1 ♂; Toledo, Quero; 22 May 1929; Escalera leg.; MNCN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E69201E9CA9FD206708FDB1.taxon	description	Description Habitus (Fig. 2 C). Length: 6.0 – 7.0 mm. First antennomere slightly darker than the rest, that are uniformly testaceous. Palps yellow. Lateral sinuosity of the pronotum well-marked; anterior and hind angles in general protruding; lateral sulcus mildly widened. Surface of the anterior region of the pronotum slightly striated. Legs entirely testaceous. Median lobe of the aedeagus narrow and slightly arcuate, with the apex scarcely developed but clearly directed down (Fig. 3 H). Chorology Wide Palaearctic distribution from the Iberian Peninsula to Siberia, in Mediterranean, Atlantic coastal and interior localities; coast of the North Sea, Black Sea and Caspian Sea (Ratti 1983; Bousquet 2017). Ibero-Balearic distribution (Fig. 5 B): in the Iberian Peninsula, the species is known with precision from coastal localities of the meridional extreme and from saline environments of the Southern Subplateau (SE Spain) and the Ebro valley (NE Spain). Imprecise records of P. luridipennis exist from Los Monegros Desert (NE Spain; Vives & Vives 1981, 1999), a saltmarsh of Toledo (SE Spain; Vives & Vives 1981), Alicante and Barcelona (E Spain; Zaballos & Jeanne 1994). The species is not known from the Balearic Islands.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E69201E9CA9FD206708FDB1.taxon	biology_ecology	Biology Species associated with saline environments where plants of the genus Salicornia grow (Vigna Taglianti & Bonavita 1994). The preimaginal stages of P. luridipennis are known, unusually for Pogonini (see Jeannel 1941). Bibliographic Ibero-Balearic records Andújar et al. 2002; Jeanne 1978; Serrano 1983; Vives & Vives 1978 a, 1986; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E68201E9CA1FDA561E5F832.taxon	description	Figs 2 D, 3 I, 5 C	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E68201E9CA1FDA561E5F832.taxon	description	Tegument similar than in P. luridipennis. Pronotum weakly or not transverse, sub-cordiform. Elytra narrow, parallel and convex. Up to 3 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E68201E9CA1FDA561E5F832.taxon	materials_examined	Material examined No Ibero-Balearic material was available for study. FRANCE • 4 ♂♂, 7 ♀♀; Provence-Alpes-Côte d’Azur, Saintes-Maries-de-la-Mer; 28 Mar. 1974; G. Dubault leg.; VMO.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E68201E9CA1FDA561E5F832.taxon	description	Description Habitus (Fig. 2 D). Length: 7.0 – 8.0 mm. Antennomeres testaceous, but the 2 nd and 3 rd are slightly lighter than the rest. Palps yellow. Lateral sinuosity of the pronotum relatively long and smooth; lateral sulcus slightly widened. Anterior region of the pronotum slightly punctured; punctuation of the base deep, combined with marked longitudinal clefts that give the surface a striated appearance. Legs entirely testaceous. Median lobe of the aedeagus wide in its median region, clearly arcuate and with the apex narrow and relatively sharp (Fig. 3 I). Chorology Species present along the Mediterranean coast of the Iberian Peninsula, France and Italy (Bousquet 2017). Ibero-Balearic distribution (Fig. 5 C): in the Iberian Peninsula, the only precise records of this species come from some localities of the Catalan coast (NE Spain), but it has also been reported, with no precision, from the coast of Valencia (E Spain; Zaballos & Jeanne 1994).	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E68201E9CA1FDA561E5F832.taxon	biology_ecology	Biology The few published comments about its biology remark that, as in other Pogonini, P. pallidipennis has a sub-lapidicolous life close to wet and saline zones (it can also be collected under other solid objects that act as refuges) (Español 1964). Bibliographic Ibero-Balearic records Español 1964; Vives & Vives 1978 a; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6B201C9F43FE9E670BFE0E.taxon	description	Figs 2 E, 3 J, 5 D	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6B201C9F43FE9E670BFE0E.taxon	description	Body tegument and first antennomere with metallic sheen, bronze or blue. Pronotum sub-cordiform, convex and almost quadrangular, with the posterior region poorly narrowed and the base sub-rectilinear and markedly wider than the anterior margin. Elytra sub-parallel and convex. Up to 3 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6B201C9F43FE9E670BFE0E.taxon	materials_examined	Material examined No Ibero-Balearic material was available for study. GREECE • 2 ♂♂; Corfu, Korission Lagoon; 15 May 1992; Gudenzi leg.; PN • 3 ♀♀; Lefkada; 4 Jun. 2004; P. Neri leg.; PN • 1 ♂; Peloponnesus; Akra Arakos, Limno, Kalogria / Limanaki; 28 Apr. 1996; W. A. Marggi leg.; WAM • 1 ♂; Phthiotis, Lamia; 30 May 1999; W. A. Marggi leg.; WAM • 1 ♂, 1 ♀; Thessaly, Fanari, Limni Ptelia; 28 May 1999; W. A. Marggi leg.; WAM • 2 ♀♀; Thrace, Kavalla, Agiasma; 28 May 1999; W. A. Marggi leg.; WAM. ITALY • 3 ♂♂, 2 ♀♀; Basilicata, mouth of the River Sinni; 25 Apr. 1988; Gudenzi leg.; PN • 1 ♂, 1 ♀; Emilia-Romagna, Ravenna, Cervia; 7 Aug. 1983; Pavanello leg.; PN • 2 ♀♀; same collection data as for preceding; Jul. 1982; Pavanello leg.; PN • 1 ♂; Emilia-Romagna, Ravenna; 28 Apr. 1979; Gudenzi leg.; PN • 1 ♀; Emilia-Romagna, Ravenna, mouth of the River Savio; 11 May 2008; Gudenzi leg.; PN • 2 ♀♀; Emilia-Romagna, Ravenna, Pineta di San Vitale; 5 Jan. 1975; Gudenzi leg.; J. Salvigni coll.; PN • 1 ♂, 2 ♀♀; Puglia, Foggia, Lake Lesina; 11 Jul. 1986; Gudenzi leg.; PN • 2 ♀♀; Toscana, Grosseto, Marina di Alberesse; 28 Apr. 2017; P. Neri leg., PN.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6B201C9F43FE9E670BFE0E.taxon	description	Description Habitus (Fig. 2 E). Length: 6.0 – 8.0 mm. Antennae dark, with the base of the antennomeres (especially the 2 – 5) slightly lighter. Palps dark, but lighter in the base and / or apex. Anterior angles of the pronotum protruding; hind angles straight. Lateral sulcus of the pronotum marked and narrow. Surface of the pronotal disc slightly rugose. Legs brown, but femora the base and apex of the tibiae or the dorsal surface of the tarsi can be darkened (in this last case, sometimes with a slight metallic sheen). Median lobe of the aedeagus narrow, regularly arcuate from the base to the apex (Fig. 3 J). Chorology Species distributed along the Mediterranean coast of South Europe from Spain to Albania (Bousquet 2017), also present in Cyprus (Jeanne 1986). In Bulgaria, it is associated with the coast of the Black Sea (Gueorguiev & Gueorguiev 1995). Ibero-Balearic distribution (Fig. 5 D): in the Iberian Peninsula, the species is known from some coastal and saline environments of Alicante (E Spain) and Catalonia (NE Spain). Eiroa et al. (1988) reported one record from the Playa de Barra (Pontevedra, Galicia, NW Spain), but it was based on incorrect identification of P. chalceus (Campos Gómez & Novoa 2006). The species has been reported, with no precise locality, from the Balearic Islands (Menorca; Compte Sart 1968; Zaballos & Jeanne 1994).	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6B201C9F43FE9E670BFE0E.taxon	biology_ecology	Biology The species occupies environments where plants of Arthrocnemetea grow (Sauleda 1985). The only comments about its phenology come from Ratti (1994), who suggested that, in some populations along the Adriatic coast, the different life phases of this species can overlap in the same year, so it cannot be exclusively considered an annual species. Bibliographic Ibero-Balearic records Baehr 1986; Cuní 1885; Prieto Piloña & Valcárcel 1996; Sauleda 1985; Serrano et al. 2021; Toribio 2022.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6A201C9F57FE4B610CF897.taxon	description	Figs 2 F, 3 K, 5 E	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6A201C9F57FE4B610CF897.taxon	description	Tegument shiny, bluish or emerald green. Pronotum sub-cordiform and markedly convex; surface of the central disc smooth. Elytra sub-parallel and convex. Up to 3 setigerous pores in the 3 rd interstria.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6A201C9F57FE4B610CF897.taxon	materials_examined	Material examined SPAIN • 2 ♂♂, 2 ♀♀; Cádiz, San Fernando, Campo Soto; 11 Nov. 1993; P. Coello leg.; VMO • 2 ♂♂; Cádiz, Chiclana; 29 Mar. 1991; J. M. Beltrán leg.; VMO • 1 ♂; Cádiz, La Janda; 18 Oct. 1980; J. de Ferrer leg.; VMO • 1 spec.; Cádiz, Laguna de La Janda; 15 Jun. 1976; J. Ramírez leg.; MNCN • 1 ♀; Cádiz, Los Barrios; 27 Apr. 1976; J. de Ferrer leg.; VMO • 2 specs; Cádiz, Palmones; 27 Apr. 1976; J. Ramírez leg.; MNCN • 2 specs; same collection data as for preceding; 3 May 1976; J. Ramírez leg.; MNCN • 3 specs; same collection data as for preceding; 29 Apr. 1976; MNCN • 3 specs; Cádiz, San Fernando; 15 Apr. 1976; J. Ramírez leg.; MNCN • 1 spec.; Cádiz, Zahara de los Atunes; 24 Apr. 1976; J. Ramírez leg.; MNCN • 1 spec.; same collection data as for preceding; 22 May 1976; MNCN • 1 ♂, 1 ♀; Sevilla, Los Palacios y Fillafranca, Caño de la Vera; 30 Nov. 2002; J. Navarro leg.; VMO.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6A201C9F57FE4B610CF897.taxon	description	Description Habitus (Fig. 2 F). Length: 6.5 – 8.0 mm. Palps yellow; antennae entirely testaceous. Anterior angles of the pronotum rounded and not protruding; lateral sulcus narrow. Base of the pronotum punctured. Legs pallid testaceous or yellow. Median lobe of the aedeagus narrow, arcuate near the distal extreme and with claw-appearance since the apex is abruptly directed down (Fig. 3 K). Chorology Species with a narrow distribution: it is present only at coastal and saline interior localities of the meridional extreme of the Iberian Peninsula (Fig. 5 E) and in North Africa (Vives 1965; Bousquet 2017).	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
03D987AA3E6A201C9F57FE4B610CF897.taxon	biology_ecology	Biology There is no published information about the biology of this Pogonini. The first record of P. smaragdinus in an interior locality of the Iberian Peninsula corresponds to the Laguna de Herrera, a saline lagoon in Antequera (Málaga, Spain) (Zaballos, 1986 a). Bibliographic Ibero-Balearic records Champion 1898; Zaballos 1986 a; Zaballos & Jeanne 1994.	en	Muñoz-Santiago, José, Ortuño, Vicente M. (2022): Revision of the tribe Pogonini (Coleoptera, Carabidae, Trechinae) from the Ibero-Balearic region. European Journal of Taxonomy 846 (1): 75-109, DOI: https://doi.org/10.5852/ejt.2022.846.1967, URL: http://zoobank.org/d187046c-f9c9-4fb9-8192-a80d06cd0e47
