taxonID	type	description	language	source
D875A26CFFD0BB182B96FC995DB00ED2.taxon	type_taxon	TYPE SPECIES. — Ptiloneura bidorsalis Enderlein, 1901, by original designation.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB182B96FC995DB00ED2.taxon	diagnosis	DIAGNOSIS. — See Enderlein (1901: 147) and García Aldrete et al. (2020: 293).	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB1E2BCDFB5F5C3A0F98.taxon	description	(Fig. 1) urn: lsid: zoobank. org: act: FA 5 B 33 B 0 - 07 C 9 - 4977 - A 47 D-A 7 D 40 F 9197 A 4	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB1E2BCDFB5F5C3A0F98.taxon	materials_examined	TYPE MATERIAL. — Holotype. Brazil • 1 ♂; Pará, Marabá, Serra Norte; 6 ° 05 ’ 03.1 ” S, 50 ° 10 ’ 35.5 ” W; XI. 1982; Michael Miles leg.; Light trap; INPA. HOLOTYPE CONDITION. — Mounted on microslide. The left fore- and hind- wings, legs, antennae, maxillary palps, left lateral sclerite of the hypandrium, side struts of the phallosome and remaining body parts missing.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB1E2BCDFB5F5C3A0F98.taxon	etymology	ETYMOLOGY. — The species honors Jacinto Campos Moreira de Castro, born in the state of Pará, father of Marcelo Cutrim, for his support and fellowship.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB1E2BCDFB5F5C3A0F98.taxon	distribution	GEOGRAPHICAL RECORDS. — Known only from the type locality, Brazil, Pará state, Marabá city, Serra Norte locality on Brazilian Amazon in northern Brazil.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB1E2BCDFB5F5C3A0F98.taxon	diagnosis	DIAGNOSIS. — Posterior process of the central sclerite of the hypandrium stout, mid-sized, slightly concave distally; posterior side side sclerites of the hypandrium long, curved, anteriorly rounded, posteriorly dilated, curved outward, acuminate (Fig. 1 E). Mesal endophallic sclerite of the phallosome transverse, with lateral posterior processes club-shaped, with inner borders denticulate (Fig. 1 F).	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD0BB1E2BCDFB5F5C3A0F98.taxon	description	DESCRIPTION Male (holotype) Color. Compound eyes black, ocelli hyaline; head pattern (Fig. 1 A). Forewings (Fig. 1 B) veins brown, pterostigma brown, slender brown marginal band from M 5 to A 2. Hindwings (Fig. 1 C) almost hyaline, with a pale brown spot at CuP. Morphology. Outer cusp of lacinial tips broad, with six short denticles (Fig. 1 D). Forewing pterostigma long (Fig. 1 B), narrow proximally, wider in the middle; areola postica wide, tall, apically rounded; Rs longer than its branches, almost straight, R 2 + 3 almost straight with a slight distal concave curve, R 4 + 5 with a median convex curve; M stem concave proximally, then almost straight, with five primary branches, M 5 distally forked, resulting in M 5 a and M 5 b. Hindwings (Fig. 1 C) Rs almost straight, R 2 + 3 straight, R 4 + 5 with a slight concavity distally; M stem concave proximally with four primary branches. Hypandrium (Fig. 1 E) of five sclerites, central sclerite broad, with anterior margin concave, with field of long macrosetae on each side of the longitudinal midline, that reach or surpass the apex of the median posterior process; posterior side processes long curved outward with a triangular posterior projection; lateral sclerite irregular. Phallosome (Fig. 1 F) relatively simple, external parameres proximally slender, curved inward, wider distally, with rounded apices, bearing a field of pores and a row of little teeth at the posterior margin. Three pairs of endophallic sclerites, an anterior pair long, curved inward, with rounded end; a mesal with anterior and posterior margin almost straight, laterally curved, and strong narrowing, from where it arises a posterior process, these almost straight, distally dilated; a postero-lateral pair, short, curved outward, proximally, and distally rounded, with a row of strong teeth along inner margin. Epiproct (Fig. 1 G) anteriorly convex, sides converging to posterior elongate process, setae absent. Paraprocts (Fig. 1 G) straight based, lateral margins converging to pointed apex, with sensory fields with 31 trichobothria on basal rosettes, setae as illustrated. Measurements (in microns). FW: 4551, HW: 3281, IO: 491, D: 536, d: 395, IO / d: 1.24, PO: 0.73.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD6BB1C2914F99E5C860BDB.taxon	description	(Fig. 2) urn: lsid: zoobank. org: act: 13 EFDBFB- 79 B 6 - 4 C 5 A-AF 4 A- 2 E 35 DD 9 B 941 F	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD6BB1C2914F99E5C860BDB.taxon	materials_examined	TYPE MATERIAL. — Holotype. Brazil • 1 ♂; Santa Catarina, Nova Teutônia; 27 ° 09 ’ 1 ” S, 52 ° 25 ’ 4 ” W; VI. 1972; Fritz Plaumann leg.; INPA. HOLOTYPE CONDITION. — Head, fore and hind wings, phallosome, hypandrium, epiproct, and paraproct mounted on microslide. The remaining body parts are preserved in 70 % ethanol.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD6BB1C2914F99E5C860BDB.taxon	etymology	ETYMOLOGY. — The species honors Leonardo de Assis Castro, son of Marcelo Cutrim.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD6BB1C2914F99E5C860BDB.taxon	distribution	GEOGRAPHICAL RECORDS. — Known only from the type locality, Brazil, Santa Catarina state, Nova Teutônia city, on Atlantic Forest in southeastern Brazil.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD6BB1C2914F99E5C860BDB.taxon	diagnosis	DIAGNOSIS. — Central sclerite of the hypandrium with two curved, postero-lateral processes, distally acuminate, curved outward resulting in a U-shaped posterior concavity; side sclerites short, almost rectangular (Fig. 2 G); forewings with a slender, marginal pigmented band from R 4 + 5 to areola postica (Fig. 2 B); mesal endophallic sclerite of phallosome transverse, with large, median, pointed process, and long, slender, postero-lateral processes (Fig. 2 F).	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD6BB1C2914F99E5C860BDB.taxon	description	DESCRIPTION Male (holotype) Color. Compound eyes black, ocelli hyaline; head pattern (Fig. 2 A). Scape and pedicel pale yellow, f 1 – f 3 pale brown. Mx 4 brown. Femora pale yellow; tibiae pale yellow with a brown band distally; tarsomeres 1 - 3 pale yellow. Forewing (Fig. 2 B) veins brown, pterostigma brown with a median hyaline area, a brown spot in R 2 + 3 and R 4 + 5 at wing margin. Hindwings (Fig. 2 C) hyaline. Morphology. Outer cusp of lacinial tips broad, with seven short denticles (Fig. 2 A). Right forewing (Fig. 2 B) pterostigma narrow proximally, wider in the middle; areola postica wide, tall, apically rounded; Rs shorter than its branches, slightly concave, R 2 + 3 almost straight, R 4 + 5 with a median concave curve; M stem concave proximally, then almost straight, with six primary branches. Right hindwing (Fig. 2 C) Rs almost straight, R 2 + 3 straight, R 4 + 5 with a median concave curve; M with four primary branches, M 2 forked. Left forewing (Fig. 2 D) differing from the right forewing in having M stem with five primary branches, M 5 forked, resulting in M 5 a and M 5 b. Left hindwing (Fig. 2 E) differing from the right hindwing in having the M stem with four primary branches. Hypandrium (Fig. 2 G) of three sclerites, central sclerite with with fields of long macrosetae on each side of the longitudinal midline, that do not reach the apices of the postero-lateral processes. Phallosome (Fig. 2 F) side struts slender, V-shaped, anteriorly independent, long, narrow, distal ends almost triangular as an outward directed hook, not fused posteriorly to external parameres, these slender, proximally dilated, elongated, narrowing distally, with a field of pores and an apical end inwardly directed. Four pairs of endophallic sclerites, an anterior small pair with two acuminate projections, the longest directed inward almost touching in the middle and the short, directed outward; a lateral bow-shaped pair curved inward, proximally wide, triangular, distally narrow, with a radular end area; a posterior pair, long, slightly wide proximally, almost straight, slightly curved outward distally and a mesal triangular sclerite, with two wide lateral process distally rounded. Epiproct (Fig. 2 H) rhomboid, with three short central setae, two long lateral and a set of posterior ones. Paraprocts straight based, broad, sensory fields with 33 trichobothria on basal rosettes, setae as illustrated (Fig. 2 H). Measurements (in microns). FW: 5503, HW: 3618, F: 1310, T: 2249, t 1: 879, t 2: 67, t 3: 165, f 1: 927, f 2: 904, f 3: 567, Mx 4: 270, IO: 545, D: 489, d: 317, IO / d: 1.71, PO: 0.58. VARIATION. — There are variations in the fore and hindwing venation in this specimen as seen in the right forewing (Fig. 2 B) with six primary branches differing from the left forewing (Fig. 2 D) with five primary branches with M 5 forked, resulting in M 5 a and M 5 b, and in the right hindwing (Fig. 2 C) with four primary branches with M 2 forked, resulting in M 2 a and M 2 b differing from the left hindwing (Fig. 2 E) with four simple primary branches.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD4BB1C2942FE5A5F630E8D.taxon	description	(Fig. 3)	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD4BB1C2942FE5A5F630E8D.taxon	materials_examined	MATERIAL EXAMINED. — Brazil • 1 ♂; Bahia, Camacan, Reserva Particular do Patrimônio Natural Serra Bonita; 15 ° 25 ’ 06 ” S, 39 ° 28 ’ 58 ” W; light trap; 15. II. 2013; Calor & Gomes leg.; MNRJ • 1 ♀; same data as for the male; MNRJ.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD4BB1C2942FE5A5F630E8D.taxon	distribution	GEOGRAPHICAL RECORDS. — Known only from the type locality, Brazil, Bahia state, Camacan city, Reserva Particular do Patrimônio Natural Serra Bonita, on Atlantic Forest in northeastern Brazil.	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
D875A26CFFD4BB1C2942FE5A5F630E8D.taxon	description	VARIATION. — It is the first record of this species with a variation in the forewing venation in which the stem of the vein M has six primary branches and a crossvein between vein A 2 and wing margin on the right and left forewings (Fig. 3 D, E). Also is the first record of a spur vein before the above mentioned crossvein, only in the right forewing (Fig. 3 D).	en	Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery, Aldrete (2022): The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species. Zoosystema 44 (20): 493-501, DOI: 10.5252/zoosystema2022v44a20
