identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C287E8FFB2FF83FF4BFAB8FB93E5C9.text	03C287E8FFB2FF83FF4BFAB8FB93E5C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mecistops leptorhynchus (Bennett 1835)	<div><p>Mecistops leptorhynchus (Bennett, 1835)</p> <p>Figure 6A</p> <p>Crocodilus leptorhynchus Bennett, 1835: 129. Type locality: “apud Fernando Po ” (near Bioko; no populations of this species are known in Bioko). Holotype likely originated from Central Africa and shipped to England (where the type is housed) via Bioko (Shirley et al. 2018). Type locality corrected to Central Africa (Shirley et al. 2018).</p> <p>Mecistops leptorhynchus (Bennett, 1835): Shirley et al. 2018: 151.</p> <p>Distribution. It is widespread throughout Central Africa, from Gabon to Cameroon and the Central African Republic, eastward to Tanzania and Zambia. In Equatorial Guinea it occurs in Río Muni at Monte Alén National Park (Lasso et al. 2002). Indirect signs of this species were also found in Monte Mitra at Monte Alén National Park (Gonwouo &amp; Nsang 2005); the only specimen examined by us is from coastal Río Muni (Map 1A).</p> <p>Comments. Previously known as Mecistops cataphractus Cuvier, 1825. Shirley et al. (2018) recognized two species within M. cataphractus. Consequently, they resurrected M. leptorhynchus as a valid species from Central Africa, whereas M. cataphractus remains restricted to western Africa. There are no known populations of this species in Bioko. Although reports of large-sized crocodilians on the beaches of Bioko have been mentioned (Bocage 1895a), they were probably the result of accidental dispersal from the Central African mainland (Mertens 1964a). Mecistops leptorhynchus is harvested as bush-meat for local consumption in Río Muni (Figure 6A from Gabon) (Fa et al. 2002). In 1964, during a visit to Río Muni, the zoologist J.A. Valverde noted its low abundance (Valverde 2004).</p> <p>Specimens examined. One specimen. Río Muni: Mbini, 1985 (EBD 25129).</p></div> 	https://treatment.plazi.org/id/03C287E8FFB2FF83FF4BFAB8FB93E5C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB0FF81FF4BFF42FCB7E1CE.text	03C287E8FFB0FF81FF4BFF42FCB7E1CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Osteolaemus tetraspis Cope 1861	<div><p>Osteolaemus tetraspis Cope, 1861</p> <p>Figure 6B</p> <p>Osteolaemus tetraspis Cope, 1861a: 550. Type locality: “Ogobai River, Western Africa”, Ogooué River, Gabon.</p> <p>Crocodilus frontatus Murray, 1862: 213. Type locality: “Old Calabar”, Duke Town, Nigeria.</p> <p>Halcrosia afzelii Lilljeborg, 1867: 715. Type locality: “ Sierra Leone, West Africa ”.</p> <p>Halcrosia nigra Gray, 1867: 200. Type locality: “ Senegal, Gabon; Old Calabar; Ogobai River ”.</p> <p>Distribution. This species occurs from the Ogooué River Basin in Gabon to Nigeria and southwest of the Republic of the Congo. In Equatorial Guinea it has been recorded from Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra Forest (Gonwouo &amp; Nsang 2005) (Map 1B).</p> <p>Comments. The taxonomy of the genus Osteolaemus was a complex matter before the molecular characterization of several populations across its distribution (Eaton et al. 2009). Molecular works support three distinct phylogenetic species within Osteolaemus, two of them being O. tetraspis and O. osborni Schmidt, 1919 from the Congo Basin, while the third remains as an undescribed taxon from West Africa (Eaton et al. 2009; Franke et al. 2013; Shirley et al. 2015). Osteolaemus tetraspis is a intensively harvested species in Río Muni, collected for the bush-meat market (a specimen from Monte Alén is shown in Figure 6B) (Fa &amp; García Yuste 2001; Fa et al. 2002).</p> <p>Specimens examined. Thirteen specimens. Río Muni: likely from Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 4171); without specific locality (MNCN 45315, EBD 21046); Ayamiken (EBD 34302); Asonga, 20 April 1986 (EBD 20912); Niefang, Cuenca del Uolo, 25 March 1987 (EBD 24934); Niefang Norte (EBD 20282); San Joaquín de Ndyiacom, 25 March 1987 (EBD 24935), September 1987 (EBD 27485); Bata, 31 August 1986 (EBD 24933); Bata, bought at the airport (EBD 20281); Punta Mbonda (EBD 334); Río Aye, camping of Alena, 13 June 1964 (EBD 2897).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB0FF81FF4BFF42FCB7E1CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB0FF81FF4BFAF0FB86E574.text	03C287E8FFB0FF81FF4BFAF0FB86E574.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pelusios gabonensis (Dumeril 1856)	<div><p>Pelusios gabonensis (Duméril, 1856)</p> <p>Figure 4F</p> <p>Pentonyx gabonensis Duméril, 1856: 373. Type locality: “ Gabon ”.</p> <p>Sternotherus gabonensis (Duméril, 1856): Bocage 1866a: 40.</p> <p>Sternothaerus steindachneri Siebenrock, 1901: 6. Type locality: Madagascar.</p> <p>Pelusios gabonensis (Duméril, 1856): Schmidt 1919: 413.</p> <p>Distribution. Widespread from West Africa to Central Africa, southward to Angola. In Equatorial Guinea it has been recorded from Río Muni, at Monte Alén National Park (Lasso et al. 2002). It has been found very close to the eastern border between Gabon and Río Muni (localities not specified) (Iverson 1992; Rhodin et al. 2017) (Map 1C).</p> <p>Comments. This species does not show a marked intraspecific genetic differentiation across its range in Central Africa (Kindler et al. 2016). The figured specimen (Figure 4F) is from Monte Alén National Park.</p> <p>Specimens examined. No specimens were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFB0FF81FF4BFAF0FB86E574	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB0FF80FF4BF805FB64E3F6.text	03C287E8FFB0FF80FF4BF805FB64E3F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pelusios niger (Dumeril & Bibron 1835)	<div><p>Pelusios niger (Duméril &amp; Bibron, 1835)</p> <p>Sternotherus niger Duméril &amp; Bibron, 1835: 397. Type locality: “probablement… Madagascar ” (in error).</p> <p>Sternothaerus oxyrhinus Boulenger, 1897a: 919. Type locality: unknown; “probably…Tropical Africa”.</p> <p>Pelusios niger (Duméril &amp; Bibron, 1835): Schmidt 1919: 402.</p> <p>Sternothaerus heinrothi Kanberg, 1924: 195. Type locality: Cameroon.</p> <p>Distribution. Gulf of Guinea, from Benin to Gabon. In Equatorial Guinea it has been recorded from Río Muni (close to Bata and Dijbloho) (Iverson 1992; Luiselli et al. 2018; Rhodin et al. 2017) (Map 2A).</p> <p>Comments. The phylogenetic relationships of the genus Pelusios are discussed by Stuckas et al. (2013). Pelusios niger is the sister lineage to P. cupulatta Bour &amp; Maran, 2003 (Stuckas et al. 2013).</p> <p>Specimens examined. One specimen. Río Muni: Mbini-Senye, 30 June 1986 (EBD 20918).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB0FF80FF4BF805FB64E3F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB1FF80FF4BFD73FB62E792.text	03C287E8FFB1FF80FF4BFD73FB62E792.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kinixys erosa (Schweigger 1812)	<div><p>Kinixys erosa (Schweigger, 1812)</p> <p>Figure 4G–H</p> <p>Testudo erosa Schweigger, 1812: 321. Type locality: “America septentrionali” (in error) (Bour 2006).</p> <p>Kinixys castanea Bell, 1827: 398. Type locality: “in Africâ”.</p> <p>Cinixyse erosa (Schweigger, 1812): Duméril &amp; Bibron 1835: 165.</p> <p>Kinixys erosa (Schweigger, 1812): Gray 1831.</p> <p>Distribution. Extends across the forests of West and Central Africa, from Guinea and Sierra Leone to Uganda and southwards to northern Angola. In Equatorial Guinea it has been recorded from Río Muni (Iverson 1992) at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 2B).</p> <p>Comments. This species seems to be relatively abundant in Monte Mitra (Gonwouo &amp; Nsang 2005). Reports of harvesting of this species in Monte Mitra forests are provided by Fa &amp; García Yuste (2001). It is the sister species of K. homeana Bell, 1827 (Kindler et al. 2012).</p> <p>Specimens examined. Sixteen specimens. Río Muni: Bata, 20 May 1966 (EBD 2775–2777, EBD 2895–2896); Mbini-Senye, 03 June1986 (EBD 20917); Nava-Ebeñ (Bata), 10 June 1984 (EBD 20273); Miboman, 08 June 1984 (EBD 20276), 10 June 1984 (EBD 20274), 20 June 1984 (EBD 20277); Ebian, 20 June 1984 (EBD 20275); San Joaquín de Ndyiacom, Ayamiken, 01 August 1986 (EBD 24983); Niefang, 25 March 1987 (EBD 25123); Etom, Evinayong-Aconibe, 25 May 1987 (EBD 24982); Mongomo, 02 July 1984 (EBD 18585 –18586).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB1FF80FF4BFD73FB62E792	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB1FF80FF4BFA6CFB86E582.text	03C287E8FFB1FF80FF4BFA6CFB86E582.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kinixys homeana Bell 1827	<div><p>Kinixys homeana Bell, 1827</p> <p>Kinixys homeana Bell, 1827: 400. Type locality: “Africâ occidentali”.</p> <p>Cinixys homeana Bell, 1827: Duméril &amp; Bibron 1835: 161.</p> <p>Distribution. West and Central Africa, from Liberia to Central African Republic, with some dubious records in the east of the Democratic Republic of the Congo (Rhodin et al. 2017). In Equatorial Guinea it has been found in Monte Mitra at Monte Alén National Park (Gonwouo &amp; Nsang 2005) and it is expected to be found to the North-west of Río Muni, close to the border with Cameroon, where several records exist from the latter country (Iverson 1992; Rhodin et al. 2017) (Map 2C).</p> <p>Comments. This species is only known in Equatorial Guinea from a single specimen found at Monte Mitra (Gonwouo &amp; Nsang 2005). It is the sister species of K. erosa (Kindler et al. 2012).</p> <p>Specimens examined. No specimens were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFB1FF80FF4BFA6CFB86E582	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB4FF85FF4BFEF2FAE1E1A2.text	03C287E8FFB4FF85FF4BFEF2FAE1E1A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclanorbis elegans (Gray 1869)	<div><p>Cyclanorbis elegans (Gray, 1869)</p> <p>Baikiea elegans Gray, 1869: 222. Type locality: “Africa”. Restricted by lectotype designation at “ Niger River drainage in west Africa” by Webb (1975).</p> <p>Cyclanorbis elegans (Gray, 1869): Boulenger 1889: 272.</p> <p>Cyclanorbis oligotylus Siebenrock, 1902: 465. Type locality: “ Ägypten ”.</p> <p>Distribution. It has a wide but patchy distribution and is currently known from Ghana, Togo, Nigeria, Cameroon, Chad, Central African Republic, Sudan, South Sudan and Ethiopia (Mazuch et al. 2016; Rhodin et al. 2017). The presence of this species in Equatorial Guinea has been overlooked (Map 3A).</p> <p>Comments. The examined specimen (EBD 31388) stands out by its large size, exhibiting a carapace length of 51 cm. There are no previous records of this genus in Equatorial Guinea (Mazuch et al. 2016; Rhodin et al. 2017). This newly recorded population of C. elegans in Río Campo (North of Río Muni) represents the southernmost record for the genus. The label of the examined specimen indicates that it contained in its stomach dates of nipa palm (Nypa fruticans) with unidentified crabs, snails, and mud.</p> <p>Specimens examined. One specimen. Río Muni: Yengüe, Ntem River, 21 February 1988 (EBD 31388).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB4FF85FF4BFEF2FAE1E1A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB4FF85FF4BFBB7FB6DE513.text	03C287E8FFB4FF85FF4BFBB7FB6DE513.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trionyx triunguis (Forskal 1775)	<div><p>Trionyx triunguis (Forskål, 1775)</p> <p>Testudo triunguis Forskål, 1775: 9. Type locality: “In Nilo rarior” (Nile River, Egypt).</p> <p>Testudo striata Suckow, 1798: 37 (partim).</p> <p>Trionyx aegytiacus Geoffroy, 1809: 116 (nomen substitutum pro Testudo triunguis).</p> <p>Trionyx niloticus Gray, 1831: 46 (nomen substitutum pro Testudo triunguis).</p> <p>Trionyx aegytiacus Cuvier, 1831: 18. Type locality: Egypt.</p> <p>Trionyx labiatus Bell, 1837: unnumbered. Type locality: Sierra Leone.</p> <p>Trionyx mortoni Hallowell, 1844: 120. Type locality: Africa.</p> <p>Aspidonectes aspilus Cope, 1860a: 295. Type locality: “Rembo and Ovenga rivers, …Fernando Vas, Equatorial West Africa”. Fernan Vaz, Gabon.</p> <p>Fordia africana Gray, 1869: 219. Type locality: Nile River at Khartoum, Sudan.</p> <p>Trionyx triunguis rudolfianus Deraniyagala, 1948: 30. Type locality: Ferguson’s Gulf, Lake Rudolf, Kenya.</p> <p>Distribution. It occurs both in the eastern Mediterranean and in African regions south of the Sahara Desert. In the Mediterranean it ranges along the eastern coastal areas, from Turkey to Israel, whereas in Africa it extends from West Africa to East Africa, southwards to the north of Namibia and northwards to Egypt (Nile River). In Equatorial Guinea it has been recorded from Río Muni at Monte Mitra in Monte Alén National Park (Gonwouo &amp; Nsang 2005) and also it is expected to be found close to the Cameroonian border (Campo River) (Rhodin et al. 2017) (Map 3B).</p> <p>Comments. The current known record of this species is based on a single specimen from Monte Mitra (Gonwouo &amp; Nsang 2005).</p> <p>Specimens examined. No specimens of this species were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFB4FF85FF4BFBB7FB6DE513	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB5FF84FF4BFEBAFE0CE714.text	03C287E8FFB5FF84FF4BFEBAFE0CE714.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agama lebretoni Wagner, Barej & Schmitz 2009	<div><p>Agama lebretoni Wagner, Barej &amp; Schmitz, 2009</p> <p>Figure 4E, 7A–B</p> <p>Agama lebretoni Wagner, Barej &amp; Schmitz, 2009: 291. Type locality: Cameroon, north-east of Mamfe, Mukwecha, Amebisu, N 05°53.866′ E 009°33.495′.</p> <p>Distribution. It extends from the southeast border of Nigeria to Gabon. In Equatorial Guinea it has been recorded in Bioko (Drewes et al. 1999) at Basilé (Bocage 1895 a, 1903), Natividad (Hacienda Natividad) (Bocage 1895 a, 1903), Punta Frailes (Boulenger 1906), San Carlos (Mertens 1941) and Riaba (=Concepción) (Mertens 1941, 1964), and in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra Forest, where it specifically has been found in Bisun (Gonwouo &amp; Nsang 2005) (Map 3C).</p> <p>Comments. Populations from Equatorial Guinea were formerly considered part of Agama agama (Linnaeus, 1758) until the publication of the comprehensive taxonomic revisions carried out by Wagner et al. (2009a, b) for the genus. Agama lebretoni is recovered as the sister lineage to the clade formed by A. agama, A. paragama Grandison, 1968, A. picticauda (Peters, 1877), A. finchi Böhme, Wagner, Malonza, Lötters &amp; Köhler, 2005, A. parafricana Trape, Mediannikov &amp; Trape, 2012, and A. africana (Hallowell, 1844) (Leaché et al. 2014). Usually found in disturbed environments, it is uncommon in pristine forests (Lasso et al. 2002).</p> <p>Specimens examined. Thirty-one specimens. Bioko: Finca Carboneras, Malabo: 26 October 1983 (EBD 19683), 29 October 1983 (EBD 19684 –19685), July 1983 (EBD 19686), 14 November 1983 (EBD 19687), 2 November 1983 (EBD 19688), 28 October 1983 (EBD 19689, EBD 19692), 9 September 1983 (EBD 19690), 22 October 1983 (EBD 19691, EBD 19700), 11 November 1983 (EBD 19693), 9 December 1983 (EBD 19694), 7 December 1983 (EBD 19695), 23 November 1988 (EBD 19696), November 1988 (EBD 19697), 7 December 1983 (EBD 19698), 18 November 1983 (EBD 19699), 31 November 1983 (EBD 19701 –19702, EBD 19705), 14 November 1983 (EBD 19703), 5 November 1983 (EBD 19704), 31 October 1983 (EBD 19706); without specific locality (MNCN 4725– 4726). Río Muni: Bata, January 1933 (MNCN 19779–19780); Bome, Bata, 6 July 1987 (EBD 27483), 10 October 1987 (EBD 27486); without specific locality (MNCN 19717), 1948.</p> <p>CHAMAELEONIDAE</p></div> 	https://treatment.plazi.org/id/03C287E8FFB5FF84FF4BFEBAFE0CE714	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB5FF8BFF4BFA01FC83E54B.text	03C287E8FFB5FF8BFF4BFA01FC83E54B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaeleo dilepis Leach 1871	<div><p>Chamaeleo dilepis Leach, 1871</p> <p>Figure 4B</p> <p>Chamaeleo dilepis Leach, 1819: 493. Type locality: not directly stated in the original description, but according to the content of the article should be within the region formerly known as French Congo [currently could be located at Republic of the Congo fide Tilbury (2010) or Gabon fide Uetz et al. (2022) and Main et al. (2022)].</p> <p>Chamaeleo bilobus Kuhl, 1820: 104. Type locality: “Congo”, French Congo (= Republic of the Congo).</p> <p>Chamaeleo planiceps Merrem, 1820: 162. Type locality: Africa.</p> <p>Chamaeleon petersii Gray, 1865a: 470. Type locality: Mozambique.</p> <p>Chamaeleo petersii var. kirkii Gray, 1865b. Type locality: East Africa.</p> <p>Chamaeleo dilepis kirkii (Gray, 1865): Loveridge 1954.</p> <p>Chamaeleo capellii Bocage, 1866a: 42. Type locality: Benguela, Angola.</p> <p>Chamaeleo angusticoronatus Barbour, 1903: 61. Type locality: Zanzibar, Tanzania.</p> <p>Chamaeleon ruspolii Boettger, 1893: 116. Type locality: Ogaden Plateau, Somaliland.</p> <p>Chamaeleon isabellinus Günther, 1893: 556. Type locality: Shire highlands, Malawi.</p> <p>Chamaeleo dilepis ruspolii Boettger, 1893: Parker 1942: 79.</p> <p>Chamaeleo dilepis idjwiensis Loveridge, 1942: 363. Type locality: Upper Mulinga River, Idjwi Island, lake Kivu, Belgian Congo (Democratic Republic of the Congo).</p> <p>Chamaeleo dilepis isabellinus Günther, 1893: Lönnberg 1911: 19.</p> <p>Chamaeleo dilepis martensi Mertens, 1964b: 113. Type locality: Pemba Island, Tanzania.</p> <p>Chamaeleo dilepis petersii Gray, 1865: Loveridge 1953: 186.</p> <p>Chamaeleo dilepis var. quilensis Bocage, 1866a: 42. Type locality: Rio Quilo (north of Cabinda).</p> <p>Chamaeleon parvilobus Boulenger, 1887: 449. Type locality: Natal, Gaboon … Cameroon.</p> <p>Chamaeleon roperi Boulenger, 1890: 85. Type locality: Kilifi, north of Mombasa, Kenya.</p> <p>Chamaeleon dilepis var. roperi Boulenger, 1890: Werner 1902: 343.</p> <p>Distribution. Widely distributed across Africa. It ranges through Central, Eastern and Southern Africa, from Cameroon to Somalia and southward to South Africa. In Equatorial Guinea it has been recorded in Río Muni (Tilbury 2010) at Cabo San Juan (Boulenger 1905), Monte Alén National Park (Lasso et al. 2002), Bata, Cogo and from Río Campo to Río Benito (Pérez del Val 2001). The records of C. gracilis from Río Muni (Monte Alén National Park and Cabo San Juan) (Boulenger 1905; Lasso et al. 2002) are herein regarded as C. dilepis (see comments) (Map 4A).</p> <p>Comments. The identity of some published records of the genus Chamaeleo for Equatorial Guinea requires confirmation due to possible misidentifications between C. dilepis and C. gracilis.This latter species usually occurs in savanna habitats, except for some forest populations in the southern part of its range (which belong to the subspecies C. gracilis etiennei Schmidt, 1919). The range of C. gracilis overlaps with that of C. dilepis at the South and East of its distribution, but they can be morphologically distinguished by the presence and extension of the occipital lobes, and by the development of the parietal crest. Chamaeleo gracilis lacks occipital lobes or if present, they are very small and immobile, and shows the parietal crest weakly marked. In contrast, C. dilepis shows the occipital lobes well developed and the parietal crest is more distinct than in C. gracilis (Tilbury 2010). All the specimens examined by us in this study (N=70) present the occipital lobes well developed, surpassing the posterior border of the helmet, with a distinctive parietal crest. These examined traits are congruent with those of C. dilepis. Boulenger (1905) recorded C. gracilis based on the MNCN specimens collected by M. Martínez de la Escalera in Cabo San Juan, Río Muni. These series of specimens were examined by us and all of them show the mentioned diagnostic traits for C. dilepis. This is noteworthy, as on the other hand, Boulenger (1887) described and figured Chamaeleon parvilobus (currently a synonym of C. dilepis) (plate XXXIX, fig. 5) based on specimens from Natal, Gabon and Cameroon with identical head traits to those of the specimens examined by us from Río Muni. Additionally, the published records of C. gracilis for Monte Alén in Río Muni (Lasso et al. 2002) (some of them housed in the EBD) are herein re-identified and regarded as C. dilepis (Figure 4B). In this sense, we remove C. gracilis from the species list of Río Muni. Chamaeleo dilepis is a widespread species complex pending of a deep revisionary work (Klaver &amp; Böhme 1986; Tilbury 2010; Main et al. 2018, 2022). Specimen MNCN 5121 was misidentified as C. senegalensis Daudin, 1802 (Pérez del Val 2001). This specimen was examined and re-identified as C. dilepis.</p> <p>Specimens examined. Seventy specimens. Río Muni: San Joaquín de Ndyiacom, 1990 (EBD 31509 for eight specimens), 1 August 1986 (EBD 24987 –24993); Asonga, 23 October 1988 (EBD 27676); Asonga, Bata, 06 November 1984 (EBD 18373); Río Muni, without specific locality (EBD, no numbered, collector number for the three specimens: B3338, B3230); Dogo, 15 km East of Bata, 14 June 1964 (EBD 2722); 3 km North of Bata, 24 June 1964 (EBD 2726); Airport of Bata, 16 October 1965 (EBD 2728); Yengue, Río Muni, 08 July 1964 (EBD 2723); Macoriza, Ayamiken, Río Muni, 20 July 1964 (EBD 2724); Bata, 17 October 1965 (EBD 2729), 29 November 1985 (EBD 20815 –20816), 07 June 1986 (EBD 20815), 1965 (EBD 2727), 06 November 1984 (EBD 18375 –18377), 05 October 1984 (EBD 18382), no date (MNCN 5103); Surroundings of Bata, 23 April 1985 (EBD 20813 –20814); Mbini, 02 October 1984 (EBD 18374, EBD 18379); Ncoaben (=Ncoeakieñ), from Bata to Movo (EBD 18378); 16 km from Niefang, 26 October 1984 (EBD 18380); from Mbini to Bata, Km 3, 26 August 1984 (EBD 18391); From Djovo to Bata, Km 19, 10 June 1984 (EBD 18392); Alosa to Niefang, 11 July 1985 (EBD 18587 –18588); Bata, 14 July 1984 (EBD 18589); Alene Samtua, Niefang, 29 June 1984 (EBD 18590 –18592); Miboman, Km 27, 03 July 1984 (EBD 18593); Alosa-Niefang, 09 July 1984 (EBD 18594); Nam-elon, Km 13 from Bata to Miboban, 17 July 1984 (EBD 18595 –18596); Miboman, Km 27 from Bata to Movo, 21 June 1984 (EBD 18622), 01 September 1984 (EBD 18675), no date (EBD 18389); Colegio de la Salle, Bata, January 1965 (EBD 2725); San Joaquín de Ndyiacom, 1990 (EBD 31501 _B-8389); Río Muni, without specific locality, 1963 (EBD 0273), July 1891 (MNCN 5094), no date (MNCN 5098, MNCN 5102, MNCN 5107–5111); from Río Campo to Río Benito, 1885 (MNCN 5095–5096); Kogo, Río Muni, August 1940 (MNCN 5121).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB5FF8BFF4BFA01FC83E54B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB8FF89FF4BFF42FD22E1E1.text	03C287E8FFB8FF89FF4BFF42FD22E1E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhampholeon spectrum (Buchholz 1874)	<div><p>Rhampholeon spectrum (Buchholz, 1874)</p> <p>Chamaeleo spectrum Buchholz, 1874: 298. Type locality: “Victoria und Bonjongo, Kamerun ”, restricted to Bonjongo, Cameroon, by Mertens (1938: 43).</p> <p>Rhampholeon spectrum (Buchholz, 1874): Günther 1874: 443</p> <p>Rhampholeon affinis Steindachner, 1911: 178. Type locality: “Beni”, Democratic Republic of the Congo.</p> <p>Brookesia spectrum (Buchholz, 1874): Mertens 1938.</p> <p>Distribution. Central Africa. From eastern Nigeria to Gabon and south-western Central African Republic. In Equatorial Guinea it has been recorded in Bioko at Musola (Boulenger 1906) and Ureca (Mertens 1964a), and in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra Forest (Gonwouo &amp; Nsang 2005). Observations from iNaturalist are as follows: Caldera de Luba (03°21′58.05″N, 08°30′26.07″E and 03°20′06.06″N, 08°28′42.09″E) and Loreto Lake (03°24′21.08″N, 08°40′06.07″E) (Map 4B).</p> <p>Comments. Commonly found in Monte Alén National Park (Lasso et al. 2002), where it has been recorded from lowlands (350 m altitude) to higher elevations at 1,100 m (Gonwouo &amp; Nsang 2005).</p> <p>Specimens examined. Twenty-three specimens. Río Muni: imprecise locality (different specimens collected among Noayong, Etom and Engong), 1987 (EBD 24974 –24977); Etom, Aconibe (from Evinayong to Mongomo), 07 July 1987 (EBD 27487); Equatorial Guinea, without specific locality (EBD 20950 –20954); Mofup, Acurenam, 24 August 1984 (EBD 18700), 28 August 1984 (EBD 18702), 29 August 1984 (EBD 18676); Oveng, Acurenam, 27 August 1984 (EBD 18701); Evinayong (Aconibe), May 1987 (EBD 25066); Miboman, Km 27 from Bata to Movo, 08 July 1984 (EBD 18599, 19600–19603), 14 June 1984 (EBD 18621), 16 June 1984 (EBD 18607); Ncoaebeñ, Bata to Movo, Km 23, 17 July 1984 (EBD 18597).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB8FF89FF4BFF42FD22E1E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB8FF89FF4BFBFFFAA6E42E.text	03C287E8FFB8FF89FF4BFBFFFAA6E42E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioceros chapini (Witte 1964)	<div><p>Trioceros chapini (Witte, 1964)</p> <p>Chamaeleo chapini Witte, 1964: 1. Type locality: “near Lukolela, in the Territory of Bikoro, Equator District of the Province of Equator, Congo (former Belgian Congo)”, Democratic Republic of the Congo.</p> <p>Trioceros chapini (Witte, 1964): Tilbury &amp; Tolley 2009: 66.</p> <p>Distribution. This species occurs in the Democratic Republic of the Congo, Gabon and Equatorial Guinea. In Equatorial Guinea it has only been recorded in Río Muni at Monte Alén National Park (Watkins-Colwell &amp; De Leon 2003) (Map 4C).</p> <p>Comments. The only known specimen from Equatorial Guinea is an adult male found in the stomach of a Telothornis kirtlandi on 5 June 2002. The specimen is housed in the Yale Peabody Museum of Natural History with the catalogue number YPM 12638 (Watkins-Colwell &amp; De Leon 2003).</p> <p>Specimens examined. No specimens of this species have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFB8FF89FF4BFBFFFAA6E42E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFB8FF8EFF4BF9F3FD67E02B.text	03C287E8FFB8FF8EFF4BF9F3FD67E02B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioceros cristatus (Stutchbury 1837)	<div><p>Trioceros cristatus (Stutchbury, 1837)</p> <p>Figure 4A</p> <p>Chamaeleon cristatus Stutchbury, 1837: 361. Type locality: “banks of the River Gaboon ”. Gabon River, Gabon.</p> <p>Pterosaurus cristata (Stutchbury, 1837): Gray 1865: 349.</p> <p>Trioceros cristatus (Stutchbury, 1837): Tilbury &amp; Tolley 2009: 66.</p> <p>Distribution. This species ranges from Nigeria to the Republic of the Congo and Central African Republic. In Equatorial Guinea it has been recorded both in Bioko (Boulenger 1887; Bocage 1903) at Musola (Boulenger 1906; Mertens 1941) and San Carlos (Mertens 1964a), and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra Forest (Fa &amp; García Yuste 2001; Gonwouo &amp; Nsang 2005) (Map 5A).</p> <p>Comments. Reports of harvesting over this species in Monte Mitra forests are provided by Fa &amp; García Yuste (2001). Commonly found in Monte Alén National Park (Lasso et al. 2002).</p> <p>Specimens examined. Eighteen specimens. Equatorial Guinea, without specific locality, but likely part of the material examined by Boulenger (1905) from Cabo San Juan (MNCN 5105–5106). Río Muni: Ebebiyin, Río Muni, 1 July 1964 (EBD 2721); Mongomo, Río Muni, 16 April 1986 (EBD 20808 –20809); Surroundings of Bata, 29 November 1985 (EBD 20810); Miboman, 30 km from Bata, 14 October 1986 (EBD 20807); Río Lobo, Moca, Evinayong, 24 May 1986 (EBD 20811); Besabeba, 01 October 1985 (EBD 20812); Asonga, Bata, 27 September 1974 (EBD 18387); Colegio de la Salle, Bata, January 1965 (EBD 2720); Movo, 28 August 1984 (EBD 18394); Acurenam “Guemg”, 24 August 1984 (EBD 18677); Acoaseng (not found locality), Bata, 09 July 1987 (EBD 27489); from Bata to Niefang, 12 February 1988 (EBD 27675); San Joaquín de Ndyiacom, 1990 (EBD 31501 _ B-9070); without specific locality, 1963 (EBD 2719).</p> </div>	https://treatment.plazi.org/id/03C287E8FFB8FF8EFF4BF9F3FD67E02B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFBFFF8EFF4BFDF6FB7BE6A3.text	03C287E8FFBFFF8EFF4BFDF6FB7BE6A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioceros feae (Boulenger 1906)	<div><p>Trioceros feae (Boulenger, 1906)</p> <p>Chamaeleon feae Boulenger, 1906: 207. Type locality: “ Moka, Fernando Po, 1300–1500 m. altitude” (=Bioko).</p> <p>Chamaeleo montium feae Boulenger, 1906: Mertens 1964a: 212.</p> <p>Trioceros feae (Boulenger, 1906): Tilbury &amp; Tolley 2009: 66.</p> <p>Distribution. Endemic to Bioko. In Bioko it has been recorded at Moka (Boulenger 1906; Mertens 1964a) (Map 5B).</p> <p>Comments. It is usually found at high elevations (1200–1500 m) (Mertens 1964a). It is closely allied to T. camerunensis (Pook &amp; Wild 1997). Five specimens conserved at the EBD collection are included in the same jar with two specimens of two different species, a single one of T. cristatus and another one of C. dilepis, all of them labelled as coming from San Joaquín de Ndyiacom, Río Muni, and sharing the number EBD 31501. However, T. feae is endemic to Bioko and this case likely represents a labelling mistake.</p> <p>Specimens examined. Six specimens. Bioko: Moka, Bioko Sur, 23 July 1984 (EBD 18598). Labelling mistake, “San Joaquín de Ndyiacom” for the specimens B-9271, B-9595, B-9203, B-9363, B-9266 (all of them share the label EBD 31501 with a specimen of Trioceros cristatus and another one of Chamaeleo dilepis).</p> </div>	https://treatment.plazi.org/id/03C287E8FFBFFF8EFF4BFDF6FB7BE6A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFBFFF8DFF4BFB7EFAE2E2CB.text	03C287E8FFBFFF8DFF4BFB7EFAE2E2CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioceros oweni (Gray 1831)	<div><p>Trioceros oweni (Gray, 1831)</p> <p>Chamaeleo oweni Gray, 1831: 7. Type locality: Fernando Po (=Bioko).</p> <p>Chamaeleo tricornis Duméril &amp; Bibron, 1836: 227. Type locality: Fernando Po (=Bioko).</p> <p>Chamaeleo bibroni Martin, 1838: 65. Type locality: Fernando Po (=Bioko).</p> <p>Chamaeleo grayii Swainsson, 1839: 369 (nomen substitutum pro Chamaeleo oweni).</p> <p>Chamaeleo unicornis Mocquard, 1906: 464. Type locality: Ngomo, dans le bas Ogoue, Gabon.</p> <p>Chamaeleo oweni cristata Sternfeld, 1912: 259 (non Chamaeleo cristatus Stutchbury, 1837).</p> <p>Type locality: “Landana in Loango, nördlich der Kongomündung”, Democratic Republic of the Congo.</p> <p>Chamaeleo michelli Müller, 1913: 230. Type locality: “ Station Go am Rulei, unterer Uelle District ”, Democratic Republic of the Congo.</p> <p>Trioceros oweni (Gray, 1831): Tilbury &amp; Tolley 2009: 66.</p> <p>Distribution. It ranges across central Africa, from Cameroon to the East of the Democratic Republic of the Congo, southward to Cabinda (Angola). In Equatorial Guinea it has been recorded in Bioko (Gray 1831; Martínez y Sáez 1886; Boulenger 1887) at Bissé (Bocage 1895 a, 1903), Natividad (Hacienda Natividad) (Bocage 1895 a, 1903), San Carlos (Mertens 1941, 1964), Riaba (=Concepción) (Mertens 1964a; Pérez del Val 2001) and Sampaka, Malabo (Pérez del Val 2001), and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 5C).</p> <p>Comments. Rarely found in Monte Alén National Park (Lasso et al. 2002).</p> <p>Specimens examined. Nineteen specimens. Equatorial Guinea, without specific locality (MNCN 5099–5100, 5866; EBD, labels with the numbers 3313 and 3325). Río Muni: Bome, Bata, 25 May 1987 (EBD 24994); Bata, 19 October 1984 (EBD 18372); Aconibe, 7 July 1987 (EBD 27488); Río Muni, 1963 (EBD 2730); Etom, a 13 km de Aconibe, 22 April 1987 (EBD 24972 –24973); Noayong, Km 23, road from Evinayong to Aconibe, 13 September 1987 (EBD 24995). Bioko: Malabo, 13 October 1984 (EBD 18393); seminario de Banapá, Malabo, Bioko (EBD 22093 –22094); Finca Sampaka, Malabo, 1941 (MNCN 5101); Bioko, without specific locality (MNCN 5113); Bioko, without specific locality, 24 April 1886 (MNCN 5115); Riaba (= Concepción), August 1959 (MNCN 18458).</p> </div>	https://treatment.plazi.org/id/03C287E8FFBFFF8DFF4BFB7EFAE2E2CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFBCFF8DFF4BFDBFFABCE687.text	03C287E8FFBCFF8DFF4BFDBFFABCE687.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ancylodactylus koehleri (Mertens 1937)	<div><p>Ancylodactylus koehleri (Mertens, 1937)</p> <p>Figure 7C</p> <p>Cnemaspis köhleri Mertens, 1937. Type locality: Buéa, Cameroun.</p> <p>Cnemaspis africanus köhleri Mertens, 1937: Loveridge 1947: 90.</p> <p>Ancylodactylus koehleri (Mertens, 1937): Malonza &amp; Bauer 2022.</p> <p>Distribution. This species occurs in Cameroon and Equatorial Guinea. In Equatorial Guinea it has been recorded from Río Muni at Benito River based on a single female specimen held at the British Museum (Thys van den Audenaerde (1967) [cited as C. africana (Werner, 1896)]; Perret 1986) (Map 6A).</p> <p>Comments. The genus Ancylodactylus has been recently proposed to include all the African species formerly treated within Cnemaspis (Malonda &amp; Bauer, 2022). Thys van den Audenaerde (1967) recorded the specimen housed in the British Museum as C. africana, noticing that it was distinct from A. koehleri. However, after the revision of the genus Cnemaspis of Africa by Perret (1986), this specimen was treated as C. koehleri. A single specimen of the genus Ancylodactylus was collected on Bioko at Caldera de Luba, Hormigas Camp, by Rayna Bell in 2013 and held at the Herpetological Collection of the Cornell University Museum of Vertebrates (catalog number: R-0014927). Based on the morphological traits and the phenotype exhibited (Figure 7C), we identify it as A. koehleri, extending its distribution to Bioko.</p> <p>Specimens examined. No specimens of this taxon were found by us in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFBCFF8DFF4BFDBFFABCE687	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFBCFF8CFF4BFAD3FB4AE127.text	03C287E8FFBCFF8CFF4BFAD3FB4AE127.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus angulatus Hallowell 1852	<div><p>Hemidactylus angulatus Hallowell, 1852</p> <p>Figure 8A</p> <p>Hemidactylus angulatus Hallowell, 1854: 63. Type locality: “west coast of Africa”, corrected to Gabon fide Loveridge (1947).</p> <p>Hemidactylus cyanodactylus Girard, 1858: 284 (non Rafinesque). Type locality: “Port Praya, San Jago, Cape de Verde Islands ”.</p> <p>Hemidactylus stellatus Boulenger, 1885: 130. Type locality: “ West Africa ”.</p> <p>Hemidactylus brookii haitianus Meerwarth, 1901: 17. Type locality: “ Haiti, … Port au Prince ”.</p> <p>Hemidactylus brookii angulatus (Hallowell, 1852): Mertens 1955.</p> <p>Hemidactylus cyanogaster Schleich, 1987: 51 (nomen substitutum pro Gecus cyanodactylus Girard, 1858).</p> <p>Hemidactylus haitianus (Meerwarth, 1901): Powell et al. 1996.</p> <p>Hemidactylus leightoni Boulenger, 1911a: 19. Type locality: “ Honda, Magdalena R., Colombia, 300-400 feet ”.</p> <p>Hemidactylus neotropicalis Shreve, 1936: 270. Type locality: “ Puerto Witches, Santander, Colombia ”.</p> <p>Distribution. Widely distributed in Africa, from Mauritania and Cape Verde at the west, to Kenya and Tanzania at the east. Introduced in several Caribbean islands and coastal localities. In Equatorial Guinea it has been recorded from Bioko (Boulenger 1885; Bocage 1903) at Malabo (Mertens 1941; Pérez del Val 2001; Carranza &amp; Arnold 2006), and from Río Muni at Bata (Pérez del Val 2001) (Map 6B).</p> <p>Comments. Hemidactylus angulatus is probably a species complex endemic to Africa (Carranza &amp; Arnold 2006). The taxonomic history of this taxon has been confusing. Hemidactylus angulatus was long considered part of the polytypic taxon H. brookii Gray, 1845 [although Schmidt (1919) recognized it as a full species]. Thus, old records of this species from Equatorial Guinea were recorded under the names H. brookii (Boulenger 1885; Bocage 1903; Mertens 1941) and H. brookii angulatus (Mertens 1964a). With the revision by Kluge (1969), the populations from Africa of this species complex were treated as H. brooki angulatus. Carranza &amp; Arnold (2006) considered H. angulatus with specific status, separated from H. brookii, as they do not form a monophyletic group. In the same work, the single specimen sequenced from Bioko was genetically more similar to the Cuban specimens (considered as H. haitianus Meerwarth, 1901) than to the rest of African populations (Carranza &amp; Arnold 2006). Currently, the H. brookii species complex is limited to Asia [type locality restricted to Borneo by Mahony (2011)] (Carranza &amp; Arnold 2006; Bauer et al. 2010; Mahony 2011), whereas the H. angulatus species complex occurs in Africa, but also in the New World, where it has been introduced (Carranza &amp; Arnold 2006; Weiss &amp; Hedges 2007). Pérez del Val (2001) listed the specimens MNCN 15664 and MNCN 15666–15668 as H. mabouia, which are herein re-identified as H. angulatus. The specimens MNCN 15664 and MNCN 15666–15668, were supposedly collected in Annobon, together with a single specimen morphologically allied to H. longicephalus (MNCN 15665). Both species have never been recorded in Annobon despite the sampling efforts carried out on this small island (Jesus et al. 2003). Due to these doubts regarding the specimens MNCN 15664, MNCN 15666–15668, we do not consider H. angulatus as part of the Annobon fauna until further investigations confirm its presence in the island.</p> <p>Specimens examined. Eight specimens. Río Muni: Bata, 14 March 1933 (MNCN 4475–4476). Bioko: Malabo 16–24 January 1933 (MNCN 4473–4474). Annobon Island? (MNCN 15664, MNCN 15666–15668).</p> </div>	https://treatment.plazi.org/id/03C287E8FFBCFF8CFF4BFAD3FB4AE127	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFBDFF8CFF4BFCFBFB73E789.text	03C287E8FFBDFF8CFF4BFCFBFB73E789.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus aporus Boulenger 1906	<div><p>Hemidactylus aporus Boulenger, 1906</p> <p>Hemidactylus aporus Boulenger, 1906: 199. Type locality: “Anno Bon”, Annobon.</p> <p>Distribution. Endemic to Annobon (Map 6C).</p> <p>Comments. Only known from the type specimens, as no more individuals have been recorded since 1906 (Meiri et al. 2017). The eight syntypes are housed in the Museo civico di storia naturale Giacomo Doria (Capocaccia 1961b). Loveridge (1947) redescribed H. aporus confirming the absence of precloacal and femoral pores. It is noteworthy that the sympatric H. newtoni, another species endemic to Annobon, also lacks preanal and femoral pores. Loveridge (1947) provided some differences between these taxa based on the number of subdigital lamellae (seven vs. 11–12 lamellae in the fourth toe of H. aporus and H. newtoni respectively) and on the arrangement of the dorsal tubercles (regular vs. irregular arranged in H. aporus and H. newtoni, respectively). However, the taxonomic status of H. aporus is in need of revision (Ceríaco et al., 2022), as some authors suggest that it could be a synonym of H. newtoni or H. longicephalus (L.M.P. Ceríaco, pers. comm.). Boulenger (1906) mentioned that Hemidactylus aporus and H. longicephalus (as H. bocagii) are closely allied, with morphological differences mainly restricted to the presence/absence of preanal and femoral pores (Loveridge, 1947) (see H. longicephalus account).</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFBDFF8CFF4BFCFBFB73E789	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFBDFF8CFF4BF990FB73E570.text	03C287E8FFBDFF8CFF4BF990FB73E570.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus biokoensis Wagner, Leache & Fujita 2014	<div><p>Hemidactylus biokoensis Wagner, Leaché &amp; Fujita 2014</p> <p>Hemidactylus biokoensis Wagner, Leaché &amp; Fujita 2014: 9. Type locality: “ Ureca on Bioko Island, Equatorial Guinea ”.</p> <p>Distribution. Endemic to Bioko. In Bioko, it has been found at Malabo (Mertens 1941), San Carlos (= Luba) (Mertens 1964a; Carranza &amp; Arnold 2006) and Ureca (Mertens 1964a) (Map 7A).</p> <p>Comments. Since Boulenger (1885), it was formerly treated as H. fasciatus Gray, 1842, until the revision of Wagner et al. (2014). There is a record from iNaturalist in Playa Aleñá (03°31′21.07″N, 08°34′43.03″E).</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFBDFF8CFF4BF990FB73E570	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF80FFB1FF4BFF0AFCC6E049.text	03C287E8FF80FFB1FF4BFF0AFCC6E049.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus coalescens Wagner, Leache & Fujita 2014	<div><p>Hemidactylus coalescens Wagner, Leaché &amp; Fujita 2014</p> <p>Figure 8B</p> <p>Hemidactylus coalescens Wagner, Leaché &amp; Fujita 2014: 7. Type locality: “ Cameroon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.04515&amp;materialsCitation.latitude=2.3972" title="Search Plazi for locations around (long 10.04515/lat 2.3972)">Campo Region</a>, Nkoelon, 2.3972º N, 10.04515º E ″.</p> <p>Distribution. This species occurs from southern Cameroon to Gabon and Congo. In Equatorial Guinea it has been recorded from Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 7B).</p> <p>Comments. Populations from Equatorial Guinea were formerly treated as H. fasciatus Gray, 1842 until the revision of Wagner et al. (2014).</p> <p>Specimens examined. Eight specimens. Río Muni: Cabo San Juan, 17 August 1901, likely part of the material examined by Boulenger (1905) (MNCN 4504); boca del Río Aye, Río Muni, 1964 (EBD 2744); Yengüe, Río Muni, 9 July 1964 (EBD 2733–2734); Bata, 1987 (EBD 27506 –27509).</p> </div>	https://treatment.plazi.org/id/03C287E8FF80FFB1FF4BFF0AFCC6E049	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF80FFB1FF4BFD51FB73E6BE.text	03C287E8FF80FFB1FF4BFD51FB73E6BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus kamdemtohami Bauer & Pauwels 2002	<div><p>Hemidactylus kamdemtohami Bauer &amp; Pauwels, 2002</p> <p>Hemidactylus kamdemtohami Bauer &amp; Pauwels, 2002: 2. Type locality: “ Gabon, eastern slope of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.821111&amp;materialsCitation.latitude=-1.1755555" title="Search Plazi for locations around (long 11.821111/lat -1.1755555)">Mont Iboundji</a> [near 01°10′32″S, 11°49′16″E (coordinates of Boussimbi, closest village to the type locality)], alt. ca. 520 m asl ”.</p> <p>Distribution. This species has been recorded from Gabon, Cameroon and Equatorial Guinea. In Equatorial Guinea the only record is based on a single photographed specimen from Río Muni at Monte Alén National Park (01°39′25″N, 10°16′10″E) (Bauer &amp; Pauwels 2002) (Map 7C); but see comments.</p> <p>Comments. This species was reported in Equatorial Guinea based on a single specimen sighted and photographed in Monte Alén National Park under the bark of a tree, Dacryodes buettneri (Engl.) H.J. Lam (Burseraceae), at 1.5 m above the ground (Bauer &amp; Pauwels 2002). However, confirmation throughout a detailed morphological revision would be advisable, since at first sight it could have been misidentified for Hemidactylus muriceus.</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF80FFB1FF4BFD51FB73E6BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF80FFB6FF4BFB40FB09E373.text	03C287E8FF80FFB6FF4BFB40FB09E373.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus longicephalus Bocage 1873	<div><p>Hemidactylus longicephalus Bocage, 1873</p> <p>Figure 8C</p> <p>Hemidactylus longicephalus Bocage, 1873: 210. Type locality: Holotype lost from “Mossamedes (Capangombe) et de Catumbella, près de Benguella”. Neotype from “Kawa Camp, Kissama National Park (-9.18303°, 13.37063°, 136 m), Luanda Province, Republic of Angola ”.</p> <p>Hemidactylus bocagii Boulenger, 1885: 125 (nomen novum pro H. longiceps).</p> <p>Hemidactylus mabouia var. molleri Bedriaga, 1892: 739. Type locality: São Thomé.</p> <p>Distribution. Widespread from Angola to Cameroon and West of the Democratic Republic of the Congo. In Equatorial Guinea it had never been recorded (Map 8A).</p> <p>Comments. The presence of this species in Equatorial Guinea is currently based on a single specimen held at the Yale University Peabody Museum and catalogued as “YPM HERR 014399” from Los Altos de Nsork National Park, in southeastern Río Muni. This is the first record of this taxon from Equatorial Guinea. The specimen MNCN 15665, which is morphologically similar to H. longicephalus, was supposedly collected in Annobon together with some specimens of H. angulatus (MNCN 15664, MNCN 15666–15668). Altought H. longicephalus was introduced in São Tome (type locality of Hemidactylus mabouia var. molleri), this seems not to be the case in Annobon, since several field expeditions carried out to the island failed to record it. Remarkably, H. aporus, a closely allied species (based on morphology) to H. longicephalus fide Boulenger (1906) has been never recorded since the original description (see H. aporus account). Hemidactylus longicephalus presents 8–11 subdigital lamellae in the fourth toe, 14–18 regular rows of dorsal tubercles, and 4–8 precloacal pores in males, whereas H. aporus presents 7 lamellae in the fourth toe, 16–20 regular rows of dorsal tubercles, and absence of precloacal pores in males (Loveridge, 1947). The examined specimen MNCN 15665, a female (precloacal pores absent), has five subdigital lamellae in the first toe and nine in the fourth, and around 16 dorsal tubercles regularly arranged. The combination of these traits is mostly within the variability of H. longicephalus. Because of (1) the possibility of a labelling mistake, or (2) the taxonomic problem regarding the Annobon endemic H. aporus, we remain cautious and do not consider H. longicephalus as part of the Annobon fauna until more evidence is obtained.</p> <p>Specimens examined. One specimen. Annobon? (missing original label) (MNCN 15665).</p></div> 	https://treatment.plazi.org/id/03C287E8FF80FFB6FF4BFB40FB09E373	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF87FFB6FF4BFE1AFAC9E5C6.text	03C287E8FF87FFB6FF4BFE1AFAC9E5C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus mabouia (Moreau de Jonnes 1818)	<div><p>Hemidactylus mabouia (Moreau de Jonnès, 1818)</p> <p>Figure 9A</p> <p>Gekko mabouia Moreau de Jonnès, 1818: 138. Type locality: “en Amérique, dans les contrées continentales qui avoisinent au midi l’archipel des Antilles, et qu’il est également répandu dans les îles même de l’archipel, depuis la Trinité jusqu’à la Jamaïque ”, restricted to St. Vincent in the Lesser Antilles by Stejneger (1904).</p> <p>Gekko incanescens Wied-Neuwied, 1824: 101. Type locality: Brazil.</p> <p>Gecko tuberculosus Raddi, 1823: 63. Type locality: warmer climates throughout the world, even on remote oceanic islands.</p> <p>Gekko armatus Wied-Neuwied, 1824: 104. Type locality: Brazil.</p> <p>Gecko aculeatus Spix, 1825: 16. Type locality: Rio de Janeiro, Brazil.</p> <p>Gecko cruciger Spix, 1825: 16. Type locality: Bahia, Brazil.</p> <p>Hemidactylus mabouia (Moreau de Jonnès, 1818): Duméril &amp; Bibron 1836: 362.</p> <p>Hemidactylus frenatus var. calabaricus Boettger, 1878: 1. Type locality: Old Calabar, Nigeria.</p> <p>Distribution. Widely distributed in tropical areas both in the Old and New Worlds, including some Atlantic islands. In Equatorial Guinea it has been recorded in the islands of Bioko (at Malabo) (Carranza &amp; Arnold 2006) and Annobon (Jesus et al. 2003; Rocha et al. 2005), and in Río Muni at the village of Sendje in Monte Mitra, within the Monte Alén National Park (Gonwouo &amp; Nsang 2005) (Map 8B).</p> <p>Comments. One of the most widespread species of the genus, as it has been introduced in several tropical regions. Likely originated in East Africa. Based on mtDNA (12S and CytB gene fragments) this species is very uniform across most of its distribution, which encompass tropical Africa, including Bioko (Malabo), South America, North America (Florida) and some Caribbean areas (Carranza &amp; Arnold 2006; Rato et al. 2021). However, despite the widespread genetic uniformity of H. mabouia in most of its range, deep lineages and cryptic diversity across the eastern coast of Africa have been documented (Agarwal et al., 2021). The taxonomic status and relationships between some populations formerly recorded as H. mabouia and the Malagasy H. mercatorius Gray, 1842 are in need of revision (Rocha et al. 2005; Rocha et al. 2010a; Agarwal et al. 2021). Within this context, the results obtained by Rocha et al. (2005) and Rocha et al. (2010a) based on mtDNA (16S gene fragment) and nuDNA (cmos gene fragment), showed that populations from Annobon and some other Gulf of Guinea islands, such as São Tomé and Príncipe, are nested within a clade formed by populations of H. mercatorius from Madagascar, Comoros, Uganda, and the Islands of Mahé (Seychelles), Zanzibar and Pemba (Tanzania). As Hemidactylus mercatorius has been described from northern Madagascar [restricted to Seychelles by Loveridge (1947)], Rocha et al. (2010a) suggested the use of H. mercatorius for the populations of the islands of the Gulf of Guinea (Annobon, São Tomé and Príncipe), leaving the status of continental populations pending further studies. By contrast, the specimens from Bioko (Malabo) and São Tomé and Príncipe sequenced by Carranza &amp; Arnold (2006) did not cluster with two Malagasy samples of H. mercatorius, thus considering the Atlantic populations as H. mabouia (Rato et al. 2021), which is congruent with the results obtained by Agarwal et al. (2021). Until the taxonomic identities of these populations alongside with those from mainland Africa are properly investigated, we tentatively follow the application of the most conservative designation, H. mabouia, for the Equatorial Guinea populations (Agarwal et al. 2021). Ceríaco et al. (2018, 2022) used the name H. mabouia for the populations from the islands of the Gulf of Guinea. In Río Muni, the species frequents human settlements (Gonwouo &amp; Nsang 2005). Pérez del Val (2001) listed the preserved specimens MNCN 15664 and MNCN 15666–15668 as H. mabouia, which are herein re-identified as H. angulatus.</p> <p>Specimens examined. Thirty-five specimens. Río Muni: Bata, 19 September 1984 (EBD 18384); Alosa, Niefang, 8 July 1984 (EBD 18608 –18609); from Ebian to Bata, Km 19, 13 June 1984 (EBD 18664); Asonga, Bata (EBD 18669); Hotel Panáfrica, Bata, 9 June 1984 (EBD 18696); Finca de los Cocos, Bata, 15 April 1963 (EBD 2735–2739, EBD 2742–2743), 26 May 1964 (EBD 2747); Boca del Río Etembue, 9 June 1964 (EBD 2740–2741); Boca del Río Aye, Río Muni (EBD 2745–2746); San Joaquín de Ndyacom, 1990 (EBD 31500). Bioko: Malabo, 20 July 1986 (EBD 21395 –21396); Las Caracolas, Finca Carboneras, Malabo, 2 June 1984 (EBD 18683 –18695, EBD 22562).</p> </div>	https://treatment.plazi.org/id/03C287E8FF87FFB6FF4BFE1AFAC9E5C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF8AFFBBFF4BFF0AFB73E0AC.text	03C287E8FF8AFFBBFF4BFF0AFB73E0AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus muriceus Peters 1870	<div><p>Hemidactylus muriceus Peters, 1870</p> <p>Hemidactylus muriceus Peters, 1870: 641. Type locality: “Quitah, Gold Coast ”, Keta, Guinea (= Ghana).</p> <p>Hemidactylus intestinalis Werner, 1897: 263. Type locality: Togo.</p> <p>Distribution. Widespread from West Africa (Guinea Conakry) to Central Africa (Gabon, Democratic Republic of the Congo). In Equatorial Guinea it has been recorded from Río Muni, in Monte Alén National Park at the Laña River (Lasso et al. 2002) and at Monte Mitra Forest (Gonwouo &amp; Nsang 2005) (Map 8C).</p> <p>Comments. Lasso et al. (2002) recorded H. muriceus in the forest of the Laña River. Gonwouo &amp; Nsang (2005) reported the presence of H. muriceus (as H. intestinalis Werner, 1897, current junior synonym of H. muriceus) at 2.5 m above the ground in primary forest at Monte Mitra. Both records fall within Monte Alén National Park, where Hemidactylus kamdemtohami has been recorded occupying the same microhabitat (Bauer &amp; Pauwels 2002). However, as the record of H. kamdemtohami is based on a single photographed specimen morphologically close to the specimens of H. muriceus from Río Muni, a detailed study of Hemidactylus from Río Muni is needed.</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF8AFFBBFF4BFF0AFB73E0AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF8AFFBBFF4BFD6DFB73E65F.text	03C287E8FF8AFFBBFF4BFD6DFB73E65F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus newtoni Ferreira 1897	<div><p>Hemidactylus newtoni Ferreira, 1897</p> <p>Hemidactylus newtoni Ferreira, 1897: 251. Type locality: “ Ilha de Anno Bom”, Annobon.</p> <p>Distribution. Endemic to Annobon (Ferreira 1897; Ceríaco et al. 2018) (Map 9A).</p> <p>Comments. Specimens originally recorded in Annobon as H. mabouia by Bocage (1893:45) became the types of H. newtoni (Ferreira 1897). Hemidactylus newtoni is the sister taxon of a putative undescribed species from São Tomé (Jesus et al. 2005a), which actually seem to be H. longicephalus (Ceríaco et al. 2022). Based on direct observations, Jesus et al. (2005a) indicated that the introduced H. mabouia is much more commonly found than H. newtoni, which could be ecologically displaced.</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF8AFFBBFF4BFD6DFB73E65F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF8AFFBBFF4BFB22FB73E7DC.text	03C287E8FF8AFFBBFF4BFB22FB73E7DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus richardsonii (Gray 1845)	<div><p>Hemidactylus richardsonii (Gray, 1845)</p> <p>Velernesia richardsonii Gray, 1845: 156. Type locality: not stated.</p> <p>Hemidactylus richardsonii (Gray, 1845): Boulenger 1885: 143.</p> <p>Distribution. Central Africa, from Cameroon to Central African Republic, southward to the Republic of the Congo. In Equatorial Guinea (Río Muni) it has been recorded by Boulenger (1919) without specific locality (Map 9B).</p> <p>Comments. No exact locality is known for this species in Equatorial Guinea.</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF8AFFBBFF4BFB22FB73E7DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF8AFFB9FF4BF915FB73E373.text	03C287E8FF8AFFB9FF4BF915FB73E373.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus conraui Tornier 1902	<div><p>Lygodactylus conraui Tornier, 1902</p> <p>Figure 9B</p> <p>Lygodactylus conraui Tornier, 1902: 670. Type locality: “Bipindi…Fernando Po” (Bipindi, Cameroon, and Bioko, Equatorial Guinea).</p> <p>Lygodacutylus (sic) strongi Barbour &amp; Loveridge, 1927: 18. Type locality: Firestone Plantation, Du River, Liberia.</p> <p>Distribution. It extends from Liberia to Gabon. In Equatorial Guinea it has been recorded from Bioko (Tornier 1902) at Bahía de San Carlos (Boulenger 1906) and Casa Moritz, Malabo (Mertens 1964a), and from Río Muni at Monte Mitra in Monte Alén National Park (Gonwouo &amp; Nsang 2005) (Map 9C).</p> <p>Comments. Part of the L. fischeri group, which also contains L. fischeri and L. thomensis (Röll et al. 2010). The presence of its close relative species L. fischeri in Río Muni (Loveridge 1947), suggests that both taxa could be sympatric. They can be readily differentiated based on the number of rows of enlarged subcaudal scales, with two enlarged rows in L. fischeri and a single one in L. conraui (Boulenger 1890; Loveridge 1947; Trape et al. 2012).</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF8AFFB9FF4BF915FB73E373	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF88FFB9FF4BFE0FFB73E0F5.text	03C287E8FF88FFB9FF4BFE0FFB73E0F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus fischeri Boulenger 1890	<div><p>Lygodactylus fischeri Boulenger, 1890</p> <p>Lygodactylus fischeri Boulenger, 1890: 80. Type locality: “ Sierra Leone ”.</p> <p>Distribution. It extends from Sierra Leone to Gabon. In Equatorial Guinea it has been recorded from Río Muni at Río Benito (Boulenger 1900 [as L. capensis]; Loveridge 1947) (Map 10A).</p> <p>Comments. Part of the L. fischeri group, which also contains L. thomensis and L. conraui (Röll et al. 2010; Gippner et al. 2021). See comments on L. conraui.</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF88FFB9FF4BFE0FFB73E0F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF88FFB9FF4BFC84FD74E467.text	03C287E8FF88FFB9FF4BFC84FD74E467.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus thomensis (Peters 1881)	<div><p>Lygodactylus thomensis (Peters, 1881)</p> <p>Scalabotes thomensis Peters, 1881: 795. Type locality: Rolas Islet, St. Thomé.</p> <p>Lygodactylus thomensis (Peters, 1881): Boulenger, 1885: 161.</p> <p>Lygodactylus rolasi Greef, 1885: 48 (nomen substitutum pro Lygodactylus thomensis).</p> <p>Lygodactylus thomensis delicatus Pasteur, 1962: 608. Type locality: “Ile du Prince”.</p> <p>Lygodactylus thomensis wermuthi Pasteur, 1962: 610. Type locality: “ Annobon ”.</p> <p>Distribution. This species is distributed in the oceanic islands of the Gulf of Guinea: São Tomé, Príncipe, and Annobon. In Equatorial Guinea it occurs only in Annobon (Boulenger 1906; Jesus et al. 2003; Ceríaco et al. 2018), where it has been particularly recorded from the north-east shore of the Apot Lake (Rodríguez-Prieto et al. 2010), the northern part of the island and “Pueblo” (Boulenger 1906), and the west coast (Rodríguez-Prieto et al. 2010) (Map 10B).</p> <p>Comments. The populations from Annobon belong to the endemic subspecies L. thomensis wermuthi Pasteur, 1962, which is treated with full species status by some authors (Ceríaco et al. 2022). Phylogenetic relationships between the three different subspecies of L. thomensis are provided by Jesus et al. (2006). The population from Annobon is closely related to the populations from Príncipe (L. thomensis delicatus Pasteur, 1962), and both taxonomic units are sister to the nominal subspecies from São Tomé. In Annobon, Jesus et al. (2003) found this species exclusively in forests, where it seems to have a patchy distribution. Lygodactylus thomensis wermuthi has been found foraging on trunks, branches and on the floor, both in dry and wet forests (Rodríguez-Prieto et al. 2010). Egg-laying in Annobon has been reported in November within nests of solitary wasps, and the eggs of this species have a diameter of 6 mm (Rodríguez-Prieto et al. 2010).</p> <p>Specimens examined. Thirteen specimens. Annobon Island (=Pagalu), (1959?) (MNCN 23714–23724). Without specific locality (MNCN 46132–46133).</p></div> 	https://treatment.plazi.org/id/03C287E8FF88FFB9FF4BFC84FD74E467	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF89FFB8FF4BFF0AFC03E1B9.text	03C287E8FF89FFB8FF4BFF0AFC03E1B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lacertaspis gemmiventris (Sjostedt 1897)	<div><p>Lacertaspis gemmiventris (Sjöstedt, 1897)</p> <p>Figure 9C, 10A</p> <p>Lygosoma gemmiventris Sjöstedt, 1897: 56. Type locality: “Kamerungebirge, Mapanja, etwa 3000 Fuss”. Mount Cameroun, Mapanja, 1000 m a.s.l.</p> <p>Riopa gemmiventris (Sjöstedt, 1897): Mertens 1965: 221.</p> <p>Panaspis gemmiventris (Sjöstedt, 1897): Fuhn 1972: 72.</p> <p>Lacertaspis gemmiventris (Sjöstedt, 1897): Schmitz et al. 2005: 16.</p> <p>Distribution. Cameroon and Equatorial Guinea. In Equatorial Guinea it has been recorded in Bioko at Moka (Mertens 1964a) and between Moka and Malabo (Schmitz et al. 2005; Jesus et al. 2007; Medina et al. 2016) (Map 10C).</p> <p>Comments. Molecular sequences of some specimens from Bioko and housed in the CAS were provided by Schmitz et al. (2005) and Medina et al. (2016). Schmitz et al. (2005) suggested that the populations from Bioko could represent an undescribed species. Mertens (1964) figured the notable variability in the colour pattern within this species, which is based on three phenotypes: (1) patternless, uniformly coloured; (2) striped, with several longitudinal black lines on the dorsal surface; and (3) marbled. These three distinct phenotypes can be found together in a single locality, such as in Moka, Bioko (Mertens 1964a). The two specimens from Moka (Bioko) examined by us are either uniform (EBD 18605) or striped (EBD 18604), and the specimens from San Joaquín de Ndyacom, in northern Río Muni, exhibit the marbled (EBD 31523) and uniform (EBD 31514) phenotypes.</p> <p>Specimens examined. Five specimens. Bioko: Moka, 23 July 1984 (EBD 18604–18605); Banapá (EBD 23566). Río Muni: San Joaquín de Ndyiacom, 1990 (EBD 31514, EBD 31523).</p> </div>	https://treatment.plazi.org/id/03C287E8FF89FFB8FF4BFF0AFC03E1B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF89FFB8FF4BFC41FDA1E420.text	03C287E8FF89FFB8FF4BFC41FDA1E420.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lacertaspis reichenowii (Peters 1874)	<div><p>Lacertaspis reichenowii (Peters, 1874)</p> <p>Figure 10B</p> <p>Lygosoma (Mocoa) reichenowii Peters, 1874: 160. Type locality: “Camaroongebirge”. Mountains from Cameroon.</p> <p>Euprepes reichenowi (Peters, 1874): Reichenow 1874: 294.</p> <p>Riopa (Panaspis) reichenovei (Peters, 1874): Smith 1937: 229.</p> <p>Leiolopisma reichenovei (Peters, 1874) (sic): Mittleman 1952: 29.</p> <p>Panaspis reichenowi (Peters, 1874): Greer 1974: 29.</p> <p>Leptosiaphos (Lacertaspis) reichenowi (Peters, 1874): Broadley 1989: 441.</p> <p>Lacertaspis reichenowi (Peters, 1874): Perret 1975.</p> <p>Distribution. It extends across Central Africa, from Cameroon to Gabon and Central African Republic. In Equatorial Guinea it has been recorded both in Bioko at Musola (Boulenger 1906) and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 11A).</p> <p>Comments. Type species of the genus Lacertaspis.</p> <p>Specimens examined. One specimen. Río Muni: imprecise locality (from Nsork, Nvom or Campo River), 1986–1987 (EBD 25054 _B5284).</p> </div>	https://treatment.plazi.org/id/03C287E8FF89FFB8FF4BFC41FDA1E420	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF89FFBDFF4BF9FEFD05E357.text	03C287E8FF89FFBDFF4BF9FEFD05E357.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lacertaspis rohdei (Muller 1910)	<div><p>Lacertaspis rohdei (Müller, 1910)</p> <p>Figure 10C, 11A</p> <p>Lygosoma rohdei Müller, 1910: 585. Type locality: “Dibongo bei Edea”, Edéa, Dibongo, Cameroon.</p> <p>Lygosoma (Leiolopisma) digitatum Chabanaud, 1917a: 97. Type locality: Lambaréné, Gabon.</p> <p>Riopa rohdei (Müller, 1910): Mertens 1968: 77.</p> <p>Panaspis rohdei (Müller, 1910): Fuhn 1972: 263.</p> <p>Leptosiaphos (Lacertaspis) rohdei (Müller, 1910): Broadley 1989: 441.</p> <p>Lacertaspis rohdei (Müller, 1910): Perret 1975.</p> <p>Distribution. It extends across Central Africa, from Cameroon to Gabon and Republic of the Congo. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 11B).</p> <p>Comments. There are no records of Lacertaspis rohdei in Bioko. Mertens (1968) found colour differences between the populations of L. rohdei from Mt. Cameroon and Mt. Kupe. As Mertens (1968) did not compare nor mention specimens of L. rohdei from Bioko in his work, it seems that the reference to Mertens (1968) made by Schmitz et al. (2005) is a mistake.</p> <p>Specimens examined. Two specimens. Río Muni: imprecise locality (from Nsork, Nvom or Campo River), 1986–1987 (EBD 25052 _B-4925, EBD 25055).</p> </div>	https://treatment.plazi.org/id/03C287E8FF89FFBDFF4BF9FEFD05E357	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF8CFFBDFF4BFE63FAA6E698.text	03C287E8FF8CFFBDFF4BFE63FAA6E698.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptosiaphos vigintiserierum (Sjostedt 1897)	<div><p>Leptosiaphos cf. vigintiserierum (Sjöstedt, 1897)</p> <p>Lygosoma vigintiserierum Sjöstedt, 1897: 56. Type locality: “Kamerungebirge, Mapanja, etwa 3000 Fuss”. Mount Cameroun, Mapanja, 1000 m a.s.l.</p> <p>Riopa vigintiserierum (Sjöstedt, 1897): Mertens 1964: 224.</p> <p>Panaspis vingintiserierum (Sjöstedt, 1897): Fuhn 1972: 265.</p> <p>Leptosiaphos vigintiserierum (Sjöstedt, 1897): Schmitz et al. 2005: 18.</p> <p>Distribution. It occurs in Cameroon and Equatorial Guinea. In Equatorial Guinea it has been only recorded in Bioko (Smith 1937) at Moka (Mertens 1964a) and between Moka and Malabo (Schmitz et al. 2005) (Map 11C).</p> <p>Comments. This taxon represents a species complex (Schmitz et al. 2005). The identities of the populations of Leptosiaphos from Bioko are currently unknown. Smith (1937) and Mertens (1964) identified the Biokoan specimens as L. vigintiserierum. However, in the molecular analysis by Schmitz et al. (2005), the populations included of this taxon showed a marked genetic structure. The Biokoan population and a specimen from Cameroon (Mt. Nlonako) form a clade sister to the rest of the populations of L. vigintiserierum. As a result, the population from Bioko remained as Leptosiaphos sp. (Schmitz et al. 2005; Jesus et al. 2007) and is herein tentatively treated as Leptosiaphos cf. vigintiserierum until a detailed taxonomic study is published. Specimens of L. cf. vigintiserierum from Río Muni are housed in the Yale Peabody Museum (Connecticut) and in the Natural History Museum (London) (not examined by us). These specimens represent the first records of this taxon from Río Muni.</p> <p>Specimens examined. No specimens of this species have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF8CFFBDFF4BFE63FAA6E698	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF8CFFBDFF4BFAA4FB89E5CB.text	03C287E8FF8CFFBDFF4BFAA4FB89E5CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panaspis breviceps (Peters 1873)	<div><p>Panaspis breviceps (Peters, 1873)</p> <p>Figure 11B</p> <p>Euprepes (Mabuia) breviceps Peters, 1873: 604. Type locality: “ Gabun ”. Gabon.</p> <p>Lygosoma breviceps (Peters, 1873): Boulenger 1887: 300.</p> <p>Mabuia batesi Boulenger, 1900: 449. Type locality: “Benito River” Río Benito, Río Muni, Equatorial Guinea.</p> <p>Lygosoma (Leiolopisma) buchneri Werner, 1909: 62. Type locality: “ Kamerun ”.</p> <p>Lygosoma (Leiolopisma) auricillatum Chabanaud, 1917a: 98. Type locality: “ Congo: Lambaréné ”.</p> <p>Emoia breviceps (Peters, 1873): Loveridge 1936b: 71.</p> <p>Riopa (Panaspis) breviceps (Peters, 1873): Smith 1937: 229</p> <p>Panaspis breviceps (Peters, 1873): Mittleman 1952: 17–23.</p> <p>Distribution. Central Africa, from Cameroon to Central African Republic and southward to northern Angola. In Equatorial Guinea it has been recorded in Bioko at Basilé (Boulenger 1906), Ureca (Mertens 1964a), and at a beach (likely in Aleñá) across from the Island of Loros (Mertens 1964a), and in Río Muni at Benito River (Boulenger 1900; Perret 1973), and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra forest (Gonwouo &amp; Nsang 2005) (Map 12A).</p> <p>Comments. A hypothesis on the phylogenetic relationships of this species was provided by Schmitz et al. (2005). It is phylogenetically allied to Panaspis cf. togoensis from Cameroon (Schmitz et al. 2005).</p> <p>Specimens examined. Two specimens. Río Muni: San Joaquín de Ndyiacom, 1990 (EBD 31521 _B-9523); imprecise locality (from Nsork, Nvom or Campo River), 1986–1987 (EBD 25051).</p> </div>	https://treatment.plazi.org/id/03C287E8FF8CFFBDFF4BFAA4FB89E5CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF93FFA2FF4BFCCCFE9DE49C.text	03C287E8FF93FFA2FF4BFCCCFE9DE49C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mochlus fernandi (Burton 1836)	<div><p>Mochlus fernandi (Burton, 1836)</p> <p>Figure 12A</p> <p>Tiliqua fernandi Burton, 1836: 62. Type locality: “Fernando Po” (Bioko, Equatorial Guinea).</p> <p>Tiliqua nigriceps Kramer, 1979: 164. Type locality: “Goldküste”, Gold Coast = Ghana.</p> <p>Lygosoma fernandi (Burton, 1836): Boulenger 1887: 304.</p> <p>Lepidothyris fernandi (Burton, 1836): Cope 1892a: 233.</p> <p>Riopa fernandi (Burton, 1836): Loveridge 1936: 71.</p> <p>Mochlus fernandi (Burton, 1836): Mittleman 1952: 9.</p> <p>Plestiodon harlani Hallowell, 1845: 170. Type locality: “ Liberia, West coast of Africa ”.</p> <p>Distribution. It extends from Sierra Leone in West Africa to Cameroon and Gabon in Central Africa. In Equatorial Guinea it has been recorded in Bioko (Burton 1836; Boulenger 1887) at Natividad (Hacienda Natividad) (Bocage 1895 a, 1903), Punta Frailes (Boulenger 1906), Musola (Mertens 1941) and San Carlos (Mertens 1964a) (Map 12C).</p> <p>Comments. Wagner et al. (2009c) revised this species and their relatives as part of the “ Lygosoma fernandi species group” and designated a neotype (of Tiliqua fernandi Burton, 1836) from San Carlos, Bioko. This specimen, the current neotype, was previously recorded by Mertens (1964). It was included in the genus Mochlus by Freitas et al. (2019) while they synonymized the genus Lepidothyris Cope, 1892 with Mochlus. There are currently two recognized subspecies, M. fernandi harlani (Hallowell, 1844) from West Africa (from Sierra Leone to Ghana) and the nominal one from West-Central Africa (from Nigeria to Gabon) (Wagner et al. 2009c). We provide the only known record of M. fernandi for Río Muni, specifically at the airport of Bata. The other records of the genus Mochlus in Río Muni correspond to M. striatus.</p> <p>Specimens examined. Two specimens. Bioko: Moka (EBD 22170). Río Muni: Bata Airport, 9 November 1966 (EBD 2841).</p> </div>	https://treatment.plazi.org/id/03C287E8FF93FFA2FF4BFCCCFE9DE49C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF93FFA2FF4BFF0AFA3AE043.text	03C287E8FF93FFA2FF4BFF0AFA3AE043.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panaspis annobonensis (Fuhn 1972)	<div><p>Panaspis annobonensis (Fuhn, 1972)</p> <p>Figure 11C</p> <p>Panaspis africana annobonensis Fuhn, 1972. Type locality: “ Ile Annobon, Golfe de Guinée”.</p> <p>Panaspis annobonensis (Fuhn, 1972): Perret 1973: 605.</p> <p>Leptosiaphos annobonensis (Fuhn, 1972): Haft 1993: 59.</p> <p>Afroablepharus annobonensis (Fuhn, 1972): Jesus et al. 2007.</p> <p>Distribution. Endemic to Annobon (Fuhn 1972; Jesus et al. 2007; Soares et al. 2018; Ceríaco et al. 2018) (Map 12B).</p> <p>Comments. The species level status was supported by Jesus et al. (2007). It is more closely related to P. cabindae than to the other congeneric species from the Gulf of Guinea islands (Soares et al. 2018).</p> <p>Specimens examined. Eighteen specimens. Annobon: 1987 (EBD 27702 –27703); 10 November 1987 (EBD 27701, EBD 27699); Pico de Fuego, July 1978 (EBD 23580 –23583); “lowland”, July 1978 (EBD 23584 –23593).</p> </div>	https://treatment.plazi.org/id/03C287E8FF93FFA2FF4BFF0AFA3AE043	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF93FFA1FF4BF89DFEF1E0BA.text	03C287E8FF93FFA1FF4BF89DFEF1E0BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mochlus striatus (Hallowell 1854)	<div><p>Mochlus striatus (Hallowell, 1854)</p> <p>Figure 12B</p> <p>Euprepis striata Hallowell, 1854b: 98. Type locality: “ Liberia, wetern coast of Africa” in error. Corrected to Gabon by Hallowell (1860).</p> <p>Euprepes (Tiliqua) elegans Fischer, 1883. Type locality: “ Sierra Leone ” (preoccupied by Euprepis elegans Peters, 1854).</p> <p>Euprepes leoninus Fischer, 1884a: 7 (nomen novum pro E. elegans Fischer, 1883).</p> <p>Lepidothyris striatus (Hallowell, 1854): Wagner et al. 2009: 16.</p> <p>Mochlus striatus (Hallowell, 1854): Freitas et al. 2019: 1086.</p> <p>Distribution. Central Africa. From southern Cameroon to Gabon and the Republic of the Congo, and east to Central African Republic. In Equatorial Guinea it has been reported in Monte Alén National Park (Lasso et al. 2002) and Cabo San Juan (Boulenger 1905) as Lygosoma fernandi (Map 13A).</p> <p>Comments. We confirm the presence of this species for Equatorial Guinea. Two species of Mochlus occur in Río Muni, M. fernandi and M. striatus. Although syntopy between both species has been recorded in southern Cameroon (Wagner et al. 2009c), no syntopic populations have yet been found in Río Muni. Based on the examined specimens, Mochlus striatus seems to be more common in Río Muni than M. fernandi.</p> <p>Specimens examined. Five specimens. Río Muni: road from Aconibe to Asoc, 03 May 1987 (EBD 24996); without specific locality, 09 October 1966 (EBD 2840); Mongomo, Welé Nzas (EBD 20838); Mongomo, Welé Nzas, 16 April 1986 (EBD 20839); Cabo San Juan, likely part of the material examined by Boulenger (1905) (MNCN 5820).</p> </div>	https://treatment.plazi.org/id/03C287E8FF93FFA1FF4BF89DFEF1E0BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF90FFA0FF4BFCF7FCF7E2E7.text	03C287E8FF90FFA0FF4BFCF7FCF7E2E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis affinis (Gray 1838)	<div><p>Trachylepis affinis (Gray, 1838)</p> <p>Figure 12C</p> <p>Tiliqua affinis Gray, 1838: 289. Type locality: not stated.</p> <p>Euprepis blandingii Hallowell, 1844: 58. Type locality: “ Liberia, W. Africa ”.</p> <p>Euprepis raddoni Gray, 1845: 112. Type locality: “ West Africa ”.</p> <p>Euprepes (Euprepis) aeneofuscus Peters, 1864: 54. Type locality: “ Elmina ” (Ghana).</p> <p>Euprepes gracilis Bocage, 1872: 77. Type locality: “ Bissau ”.</p> <p>Euprepes stangeri Müller, 1882: 159. Type locality: “ Akropong ” (Ghana).</p> <p>Euprepes pantaenii Fischer, 1885: 90. Type locality: “a: Sierra Leone, b: Cameroon ”.</p> <p>Euprepes cupreus Fischer, 1886: 6. Type locality: “ Insel St. Thomé ”.</p> <p>Mabuia affinis (Gray, 1838): Boulenger 1887: 166.</p> <p>Euprepis affinis (Gray, 1838): Mausfeld-Lafdhiya et al. 2002 (by implication).</p> <p>Trachylepis affinis (Gray, 1838): Bauer 2003 (by implication).</p> <p>Distribution. Widely distributed from West Africa (Senegal) to Central Africa (South Sudan), southward to Angola, including small islands such as Príncipe. In Equatorial Guinea it has been recorded in Bioko at Pico Basilé (Bocage 1895 a, 1903), Punta Frailes (Boulenger 1906), Bococo (Mertens 1941), San Carlos de Luba (Mertens 1964a), and Malabo (Pérez del Val 2001), and in Río Muni at Río Benito (Boulenger 1901, Hoogmoed 1974), Cabo San Juan (Boulenger 1905), Evinayong (Hoogmoed 1974), and Monte Alén National Park (Lasso et al. 2002) including Monte Mitra forest (Gonwouo &amp; Nsang 2005) (Map 13B).</p> <p>Comments. This is the lizard most commonly found in Monte Mitra region (Gonwouo &amp; Nsang 2005). This taxon belongs to a species complex pending further systematic revisions. It contains two main lineages, one from West Africa (Ghana) and the other from Central-West Africa (Allen et al. 2019).</p> <p>Specimens examined. Eighty-three specimens. Bioko: without specific locality (MNCN 5786); Catedral Santa María, Malabo, 07 January 1973 (EBD 22177); Malabo 09-12 January 1933 (MNCN 5797–5798), January 1941 (MNCN 5806), 04–12 July 1957 (MNCN 9891); Ela Nguema, Malabo, 17 September 1984 (EBD 18678); Malabo, 1 November 1983 (EBD 22052), February, 1986 (EBD 20381), 20 June 1986 (EBD 21394, EBD 23541 –23543); Las Caracolas, Finca Carboneras, Malabo, 14 June 1984 (EBD 18703). Río Muni: Cabo San Juan (MNCN 5813), 17 August 1901 (MNCN 5811) [likely part of the material examined by Boulenger (1905)]; Asonga, Bata, 4 Agosto 1984 (EBD 18704 –18706), 23 April 1986 (EBD 20837); Mbini (a orillas del río Woro) (banks of the Uoro River), 17 June 1987 (EBD 27492 –27495); Covadonga, Corisco Island, 8 November 1985 (EBD 20955); Bome, Bata, 1 August 1987 (EBD 27490); San Joaquín de Ndyiacom, 1990 (EBD 31521 _B-8497); Equatorial Guinea (1989– 1990) (EBD 32032); Cogo, April-June 1964 (EBD 2814–2839); Finca de los Cocos, Cogo, Bata, from April to June 1964 (EBD 2803–2840); Mbini, 11 June 1987 (EBD 27691).</p> </div>	https://treatment.plazi.org/id/03C287E8FF90FFA0FF4BFCF7FCF7E2E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF91FFA0FF4BFEBAFD06E133.text	03C287E8FF91FFA0FF4BFEBAFD06E133.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis albilabris (Hallowell 1857)	<div><p>Trachylepis albilabris (Hallowell, 1857)</p> <p>Figure 13A</p> <p>Euprepes albilabris Hallowell, 1857: 51. Type locality: “Gaboon” Gabon.</p> <p>Mabuia albilabris (Hallowell, 1857): Boulenger 1905: 184.</p> <p>Euprepis albilabris Hallowell, 1857: Mausfeld-Lafdhiya et al. 2002 (by implication).</p> <p>Trachylepis albilabris (Hallowell, 1857): Bauer 2003 (by implication).</p> <p>Distribution. Widespread from West Africa (Guinea Conakry) to Central-East Africa (Uganda). In Equatorial Guinea it has been recorded in Río Muni at Cabo San Juan (Boulenger 1905; Hoogmoed 1974), Benito River (Hoogmoed 1974), Evinayong (Hoogmoed 1974), and Monte Alén National Park (Lasso et al. 2002) (Map 13C).</p> <p>Comments. This species is sister to T. aureogularis (Müller, 1885) from West Africa (Allen et al. 2019).</p> <p>Specimens examined. Two specimens. Río Muni: Cabo San Juan, likely part of the material examined by Boulenger (1905) (MNCN 5812, MNCN 5814).</p></div> 	https://treatment.plazi.org/id/03C287E8FF91FFA0FF4BFEBAFD06E133	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF91FFA0FF4BFCCFFE5CE577.text	03C287E8FF91FFA0FF4BFCCFFE5CE577.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis maculilabris (Gray 1845)	<div><p>Trachylepis maculilabris (Gray, 1845)</p> <p>Figure 13B</p> <p>Euprepis maculilabris Gray, 1845: 114. Type locality: “ West Africa ”.</p> <p>Euprepes notabilis Peters, 1879a: 36. Type locality: “ Pungo-Andongo ” (Malanje Province) and “ Chinchoxo ” (Cabinda Province).</p> <p>Mabuia (sic) maculilabris (Gray, 1845): Boulenger 1887: 164.</p> <p>Mabuia maculilabris major Sternfeld, 1912. Type locality: “Kamerun”, “ Sudan ”.</p> <p>Mabuia maculilabris var. kwidjwiensis Sternfeld, 1912. Type locality: “Kiwu-See”, “ Insel Wau im Kiwu-See” (Kivu lake).</p> <p>Mabuia maculilabris var. wauensis Sternfeld, 1912. Type locality: “Fort Beni” Fort Beni, Kivi, Democratic Republic of the Congo.</p> <p>Mabuia maculilabris var. schubotzi Sternfeld, 1912. Type locality: “Mawambi” and “Awakubi”, Democratic Republic of the Congo.</p> <p>Mabuia maculilabris var. graueri Sternfeld, 1912. Type locality: “Tanganyika” and “Lualaba” Democratic Republic of the Congo, “Kiwu-See” (Kivu lake).</p> <p>Mabuia maculilabris var. rohrbecki Sternfeld, 1912. Type locality: “ Nguru D. O. Africa ”, Tanzania.</p> <p>Mabuia maculilabris var. bergeri Sternfeld, 1912. Type locality: “ Mawambi ” (Zambia) and “Awakubi” (Democratic Republic of the Congo), “Fort Beni” (Fort Beni, Kivi, Democratic Republic of the Congo).</p> <p>Mabuya maculilabris (Gray, 1845): Schmidt 1919: 525.</p> <p>Mabuya maculilabris comorensis Loveridge, 1953: 200. Type locality: “ Chikwawa and Chiromo on the Shire River ”.</p> <p>Trachylepis maculilabris (Gray, 1845): Bauer 2003 (by implication).</p> <p>Distribution. Widespread from West Africa (Liberia) to East Africa (Ethiopia). In Equatorial Guinea it has been recorded from Monte Mitra forest at Monte Alén National Park (Gonwouo &amp; Nsang 2005) (Map 14A).</p> <p>Comments. This taxon belongs to a species complex pending further systematic revisions (Mausfeld-Lafdhiya et al. 2004; Jesus et al. 2005b; Rocha et al. 2010b; Ceríaco et al. 2016; Allen et al. 2019; Ceríaco et al. 2020b). It is phylogenetically related to T. polytropis (Allen et al. 2017, 2019). Gonwouo &amp; Nsang (2005) recorded T. maculilabris for the first time in Río Muni at Monte Mitra forest.</p> <p>Specimens examined. Nine specimens. Río Muni: San Joaquín de Ndyiacom, 1990 [EBD 31515, EBD 31522 (2 exx: B-8357, B-8358)]; Bome, Bata, 23 October 1987 (EBD 27692–27693, 27695–27697); Río Muni, without specific locality (EBD 25053).</p> </div>	https://treatment.plazi.org/id/03C287E8FF91FFA0FF4BFCCFFE5CE577	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9AFFABFF4BFA75FB2EE5C7.text	03C287E8FF9AFFABFF4BFA75FB2EE5C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Feylinia currori Gray 1845	<div><p>Feylinia currori Gray, 1845</p> <p>Figure 14A</p> <p>Feylinia currori Gray, 1845: 129. Type locality: “ Angola ”.</p> <p>Distribution. Widely distributed in Central Africa. It extends from Sierra Leone to western Kenya at Kakamega forest. In Equatorial Guinea it has been recorded in Río Muni (Brygoo &amp; Roux-Estève 1983) at Cabo San Juan (Boulenger 1905), Bata and Benito River (Wagner &amp; Schmitz 2006) and Monte Alén National Park (Lasso et al. 2002) (Map 15A).</p> <p>Comments. The examined specimen (MNCN 23911), a juvenile, presents 22 scale rows at mid-body and the third supralabial contacting the eye. These traits correspond to those of F. currori. Brygoo &amp; Roux-Estève (1983) reviewed this genus and Wagner &amp; Schmitz (2006) updated the geographic distribution of F. currori. Lasso et al. (2002) reported a single specimen in a cultivated field of Monte Alén National Park.</p> <p>Specimens examined. One specimen. Gulf of Guinea, Equatorial Guinea, without specific locality, but likely from Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 23911).</p></div> 	https://treatment.plazi.org/id/03C287E8FF9AFFABFF4BFA75FB2EE5C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9AFFABFF4BFF0AFE34E08F.text	03C287E8FF9AFFABFF4BFF0AFE34E08F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis ozorii (Bocage 1893)	<div><p>Trachylepis ozorii (Bocage, 1893)</p> <p>Figure 13C</p> <p>Mabuya ozorii Bocage, 1893a: 46. Type locality: “Anno-Bom”. Annobon.</p> <p>Mabuia ozorii Bocage, 1893: Sternfeld 1917.</p> <p>Trachylepis ozorii (Bocage, 1893): Bauer 2003 (by implication).</p> <p>Distribution. Endemic to Annobon (Bocage 1893a,b, 1897; Boulenger 1906; Jesus et al. 2003, 2005c) (Map 14B).</p> <p>Comments. The phylogenetic relationships of this species are not yet well studied. It is not closely related to any species from the Gulf of Guinea Islands (Jesus et al. 2005b). Based on some morphological traits, Ceríaco et al. (2016) tentatively suggested that this species could be related to T. polytropis and T. makolowodei Chirio, Ineich, Schmitz &amp; LeBreton, 2008.</p> <p>Specimens examined. Twenty specimens. Annobon: Aeropuerto de Anobon, 25 November 1987 (EBD 27822); Annobon, November 1987 (EBD 27808–27821, EBD 27823, EBD 27900), 10 November 1987 (EBD 27698, EBD 27700, EBD 27704).</p> </div>	https://treatment.plazi.org/id/03C287E8FF9AFFABFF4BFF0AFE34E08F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9AFFABFF4BFC92FB73E734.text	03C287E8FF9AFFABFF4BFC92FB73E734.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis polytropis (Boulenger 1903)	<div><p>Trachylepis polytropis (Boulenger, 1903)</p> <p>Mabuia polytropis Boulenger, 1903: 433. Type locality: “Benito River district, Gaboon”, Río Benito district, Río Muni, Equatorial Guinea.</p> <p>Mabuya polytropis (Boulenger, 1903): Schmidt 1919: 531.</p> <p>Euprepis polytropis (Boulenger, 1903): Mausfeld-Lafdhiya et al. 2002 (by implication).</p> <p>Trachylepis polytropis (Boulenger, 1903): Bauer 2003 (by implication).</p> <p>Distribution. This species ranges across West and Central Africa, from Liberia to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded from Bioko (Hoogmoed 1974) at Punta Frailes (Boulenger 1906), between San Carlos and Batete (Mertens 1964a), and at a beach (likely in Aleñá) across from the Island of Loros (Mertens 1964a), and from Río Muni at Benito River (Boulenger, 1903), and Monte Alén National Park (Lasso et al. 2002) including Monte Mitra forest (Gonwouo &amp; Nsang 2005) (Map 14C).</p> <p>Comments. This species is related to T. maculilabris (Allen et al. 2017, 2019).</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF9AFFABFF4BFC92FB73E734	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF98FFA9FF4BFF0AFAA6E058.text	03C287E8FF98FFA9FF4BFF0AFAA6E058.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Feylinia elegans (Hallowell 1892)	<div><p>Feylinia elegans (Hallowell, 1892)</p> <p>Acontias elegans Hallowell, 1854: 64. Type locality: “ Liberia ” or “ Gabon ” (see Schmidt [1919] and Brygoo &amp; Roux-Estève [1983]).</p> <p>Sphenorhina elegans (Hallowell, 1854): Hallowell 1857: 52.</p> <p>Anelytrops elegans (Hallowell, 1854): Bocage 1866a: 45.</p> <p>Feylinia elegans (Hallowell, 1854): Schmidt 1919: 605.</p> <p>Distribution. With the exception of the dubious type locality from Liberia in West Africa, this species is widespread across Central Africa, from Cameroon to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded in Río Muni at Bata (Brygoo &amp; Roux-Estève 1983) (Map 15B).</p> <p>Comments. A specimen from Bata is drawn in Brygoo &amp; Roux-Estève (1983), labelled as MNHN 90.359 and housed in the French National Museum of Natural History (MNHN).</p> <p>Specimens examined. No specimens of this species have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF98FFA9FF4BFF0AFAA6E058	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF98FFA9FF4BFC99FAA6E70C.text	03C287E8FF98FFA9FF4BFC99FAA6E70C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Melanoseps occidentalis (Peters 1877)	<div><p>Melanoseps occidentalis (Peters, 1877)</p> <p>Herpetosaura occidentalis Peters, 1877: 416. Type locality: “Cameruns”. Cameroon.</p> <p>Scelotes occidentalis (Peters, 1877): Werner 1898: 530.</p> <p>Melanoseps occidentalis (Peters, 1877): Tornier 1901: 62.</p> <p>Scelotes schebeni Sternfeld, 1910a. Type locality: “Windhuk”, Windhoek, Namibia.</p> <p>Melanoseps occidentalis zairensis Brygoo &amp; Roux-Esteve, 1982: 1187. Type locality: “Dekese”, Democratic Republic of the Congo.</p> <p>Distribution. It occurs in Central Africa, from Cameroon to Central African Republic and Angola. In Equatorial Guinea it has been recorded in Bioko at Musola (Boulenger 1906) (Map 15C).</p> <p>Comments. Morphologically similar and likely conspecific to Melanoseps poensis (Brygoo &amp; Roux-Esteve 1982). Further studies are needed to ascertain the identity and distribution of these taxa across Bioko, mainly through the revision of topotypic specimens of M. poensis, as the type series is lost (see comments in the account of M. poensis). The specimen recorded by Boulenger (1906) housed in the Museo Civico di Storia Naturale di Genova was examined by Brygoo &amp; Roux-Esteve (1982), supporting its identity as Melanoseps occidentalis.</p> <p>Specimens examined. No specimens of this species have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF98FFA9FF4BFC99FAA6E70C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF98FFA9FF4BFA0DFAA6E58E.text	03C287E8FF98FFA9FF4BFA0DFAA6E58E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Melanoseps poensis (Bocage 1895)	<div><p>Melanoseps poensis (Bocage, 1895)</p> <p>Scelotes poensis Bocage, 1895b: 16. Type locality: “Bissé, nas faldas do Pico de Santa Isabel ”. Basilé, foothill of Pico Basilé.</p> <p>Melanoseps poensis (Bocage, 1895): Brygoo &amp; Roux-Esteve 1982: 1171.</p> <p>Distribution. Endemic to Bioko. This taxon is only known from the type locality, close to Basilé, Bioko (Bocage 1895b, 1897, 1903; Boulenger 1906) (Map 16A).</p> <p>Comments. Treated as a synonym of Melanoseps occidentalis by Mertens (1964) without providing any justification. Morphologically allied and likely conspecific to Melanoseps occidentalis (Brygoo &amp; Roux-Esteve 1982), being the only difference between them is the number of supraciliary scales, five in M. poensis and four in M. occidentalis. Unfortunately, the types of Scelotes poensis were lost during the fire suffered in the Museu Bocage of Lisbon in 1977 (Brygoo &amp; Roux-Esteve 1982), hampering the solution of this taxonomic problem. The study of further collected topotypic specimens will be required in order to resolve its identity. Brygoo &amp; Roux-Esteve (1982) tentatively treated M. poensis as a valid species until more data are available but suggesting the likely conspecificity of both taxa.</p> <p>Specimens examined. No specimens of this species have been found in the collections examined.</p></div> 	https://treatment.plazi.org/id/03C287E8FF98FFA9FF4BFA0DFAA6E58E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9EFFAFFF4BFEF2FAA6E637.text	03C287E8FF9EFFAFFF4BFEF2FAA6E637.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monopeltis galeata (Hallowell 1852)	<div><p>Monopeltis galeata (Hallowell, 1852)</p> <p>Phractogonus galeatus Hallowell, 1852: 62. Type locality: “ Liberia, West Coast of Africa”; corrected to “Gaboon Country, West Africa” by Hallowell (1857).</p> <p>Monopeltis (Phractogonus) magnipartitus Peters, 1879b: 276. Type locality: “ Angeblich von dem Gabun ”.</p> <p>Lepidosternum galeatum (Hallowell, 1852): Strauch 1881: 121 (emendation).</p> <p>Lepidosternum dumerilii Strauch, 1881: 122. Type locality: “ West Afrika (Gabon)”.</p> <p>Lepidostenon magnipartitum Strauch, 1881: 124 (emendation).</p> <p>Monopeltis galeata (Hallowell, 1852): Boulenger, 1885: 384.</p> <p>Monopeltis unirostralis Mocquard, 1903: 210. Type locality: “ Gabon ”.</p> <p>Monopeltis boveei Mocquard, 1903: 211. Type locality: “ Fernand Vaz (Congo français)”.</p> <p>Distribution. Gabon and Equatorial Guinea. In Equatorial Guinea it has been recorded from Corisco (Gans &amp; Lehman 1973) (Map 16B).</p> <p>Comments. The genus Monopeltis is sister to the South African genus Dalophia (Gray, 1865) (Measey &amp; Tolley 2013). The phylogenetic relationships of M. galeata in relation to other congeneric species remain unknown. A comprehensive revision of this species was provided by Gans &amp; Lehman (1973). The single known specimen of M. galeata from Equatorial Guinea (Corisco Island), currently held at the ANSP and labelled as ANSP 9682, is a possible paralectotype (Gans &amp; Lehman 1973).</p> <p>Specimens examined. No specimens of this species have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FF9EFFAFFF4BFEF2FAA6E637	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9EFFAFFF4BFBD1FB1EE417.text	03C287E8FF9EFFAFFF4BFBD1FB1EE417.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monopeltis jugularis Peters 1880	<div><p>Monopeltis jugularis Peters, 1880</p> <p>Figure 14B</p> <p>Monopeltis (Phractogonus) jugularis Peters, 1880: 219. Type locality: “Westafrika”. West Africa.</p> <p>Lepidosternon jugulare (Peters, 1880): Strauch 1881: 124 (emendation).</p> <p>Lepidosternon koppenfelsii Strauch, 1881: 124. Type locality: “ Westafrika (Gabon)”.</p> <p>Monopeltis semipunctata Boettger, 1893: 89. Type locality: “ Kamerun ”.</p> <p>Distribution. Cameroon, Equatorial Guinea and likely Gabon. In Equatorial Guinea it has been recorded from Río Muni at Etembue (Gans &amp; Lehman 1973) (Map 16C).</p> <p>Comments. The phylogenetic relationships of this species in relation to other congeneric species remains unknown (Measey &amp; Tolley 2013). A comprehensive revision of this species was provided by Gans &amp; Lehman (1973).</p> <p>Specimens examined. One specimen. Río Muni: without specific locality (EBD 02842).</p> </div>	https://treatment.plazi.org/id/03C287E8FF9EFFAFFF4BFBD1FB1EE417	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9EFFACFF4BF961FA37E3C7.text	03C287E8FF9EFFACFF4BF961FA37E3C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gastropholis echinata (Cope 1862)	<div><p>Gastropholis echinata (Cope, 1862)</p> <p>Figure 14C</p> <p>Lacerta (Zootoca) echinata Cope, 1862: 189. Type locality: “West Africa”. Very likely Liberia according to Loveridge (1941).</p> <p>Lacerta hirticauda Vaillant, 1884: 344. Type locality: “Kinjabo” Kinjaho, Assini, Ghana.</p> <p>Lacerta langi Schmidt, 1919: 492. Type locality: “Medje”, Democratic Republic of the Congo.</p> <p>Centromastix echinatus (Cope, 1862): Laurent 1958: 118.</p> <p>Gastropholis echinata (Cope, 1862): Arnold 1989: 543.</p> <p>Distribution. West and Central Africa, from Liberia to the Northeast of the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded in Río Muni at Benito River (Boulenger 1921) and Bioko (Pérez del Val 2001) (but see comments) (Map 17A).</p> <p>Comments. Mertens (1964) did not include this species among the reptiles of Bioko. Later, Pérez del Val (2001) recorded a single specimen from the island (MNCN 6231). However, the collection locality of this specimen is tentative, and there are no recent confirmed records of this species in Bioko. In this sense, following Mertens (1964), we do not consider the presence of this species in Bioko. As this species inhabits the forest canopy, it is rarely observed and collected (Arnold 1989). Future prospections should be made to detect the presence (and confirm the Biokoan record) of this elusive species in Equatorial Guinea.</p> <p>Specimens examined. One specimen. Equatorial Guinea, without specific locality (Bioko?) (MNCN 6231).</p></div> 	https://treatment.plazi.org/id/03C287E8FF9EFFACFF4BF961FA37E3C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9DFFACFF4BFD13FDA7E61D.text	03C287E8FF9DFFACFF4BFD13FDA7E61D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holaspis guentheri Gray 1863	<div><p>Holaspis guentheri Gray, 1863</p> <p>Figure 15A</p> <p>Holaspis guentheri Gray, 1863: 153. Type locality: not stated.</p> <p>Distribution. It occurs in West and Central Africa, from Sierra Leone to Uganda. In Equatorial Guinea it has been recorded in Río Muni at Cabo San Juan (Boulenger 1905), Benito River (Boulenger 1921), and Monte Alén National Park (Pauwels &amp; Stévart 2007). Likely occurring in Bioko (Drewes et al. 1999) (Map 17B).</p> <p>Comments. Drewes et al. (1999) suggested the presence of this species on Bioko based on a possible sight (not collected) between Luba and Malabo.</p> <p>Specimens examined. One specimen. Río Muni: Cabo San Juan, likely part of the material examined by Boulenger (1905) (MNCN 7981).</p></div> 	https://treatment.plazi.org/id/03C287E8FF9DFFACFF4BFD13FDA7E61D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF9DFFACFF4BFB57FDA7E4E3.text	03C287E8FF9DFFACFF4BFB57FDA7E4E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Poromera fordii (Hallowell 1857)	<div><p>Poromera fordii (Hallowell, 1857)</p> <p>Figure 15B</p> <p>Tachydromus fordii Hallowell, 1857: 48. Type locality: “Gaboon”. Gabon.</p> <p>Poromera fordii (Hallowell, 1857): Boulenger 1887: 6.</p> <p>Distribution. It occurs in Central Africa, from Cameroon to Central African Republic, southward to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded in Bioko at Bonyoma (at 450 m a.s.l.) and Parador (North of Musola) (Mertens 1968), and in Río Muni at Cabo San Juan (Boulenger 1905), Benito River (Boulenger 1900, 1921), and Monte Alén National Park (Lasso et al. 2002) (Map 17C).</p> <p>Comments. Photographic records by Twan Leenders from Los Altos de Nsork National Park are available in “https://calphotos.berkeley.edu/”, likely belonging to one of the specimens catalogued as “YPM HERR 014402” and “YPM HERR 014402” held in the Yale University Peabody Museum.</p> <p>Specimens examined. One specimen. Río Muni: Cabo San Juan, likely part of the material examined by Boulenger (1905) (MNCN 7983).</p></div> 	https://treatment.plazi.org/id/03C287E8FF9DFFACFF4BFB57FDA7E4E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE2FFD3FF4BFF0AFD5EE78B.text	03C287E8FFE2FFD3FF4BFF0AFD5EE78B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Varanus niloticus (Linnaeus 1766)	<div><p>Varanus niloticus (Linnaeus, 1766)</p> <p>Lacerta monitor Linnaeus 1758: 201 (nomen rejectum).</p> <p>Lacerta nilotica Linnaeus, 1766: 369. Type locality: “in AEgypto” (Egypt). The holotype was a specimen mentioned by Hasselquist (1757), which is presumed lost (Marques et al. 2018).</p> <p>Stellio saurus Laurenti, 1768: 56. Type locality: “Zeylania ad littora maris”.</p> <p>Lacerta capensis Sparrman, 1783: 749. Type locality: “European settlement, called Agter Bruntjes-hoogte”, Hinterbruyntjes Cave, surroundings of Cape Town, South Africa.</p> <p>Lacertus tupinambis Lacépède, 1788: 251 (in part). Type locality: Cape of Good Hope and Senegal.</p> <p>Tupinambis elegans Daudin, 1802: 59 (in part). Type locality: “ Suriname ”.</p> <p>Tupinambis stellatus Daudin, 1802: 59 (in part). Type locality: from Senegal to Cape of Good Hope, restricted by Mertens (1942) to Senegal.</p> <p>Tupinambis ornatus Daudin, 1803: 353. Type locality: Malimbe, Cameroon.</p> <p>Monitor pulcher Leach in Bowdich, 1819: 493. Type locality: Fantee, Gold Coast, Ghana.</p> <p>Varanus niloticus (Linnaeus, 1766): Fitzinger, 1826: 50.</p> <p>Monitor elegans senegalensis Schlegel, 1844. Type locality: Senegal.</p> <p>Distribution. Widespread across sub-Saharan Africa, extending northward to Egypt. In Equatorial Guinea it has been recorded in Bioko (Boulenger 1885; Bocage 1895a) at San Carlos (Mertens 1941, 1942, 1964), Batete (Mertens 1941, 1942), Bococó (Mertens 1941), and at the road between San Carlos and Malabo (Mertens 1964a), and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 18A).</p> <p>Comments. Central African populations from the Gulf of Guinea were considered by some authors as a different species, namely Varanus ornatus (Daudin, 1803), which differs from the other African populations by some phenotypic traits (Böhme &amp; Ziegler 1997). However, based on molecular data, Dowel &amp; Hekkala (2016) found no genetic differences between V. niloticus and V. ornatus, recognizing the later as a synonym of V. niloticus. Thus, we follow Dowel &amp; Hekkala (2016) and consider the populations from the Gulf of Guinea a phenotypic morph of V. niloticus. This species is harvested as bush-meat for local consumption in Río Muni (Fa &amp; García Yuste 2001; Fa et al. 2002; Gonwouo &amp; Nsang 2005), where it is often found close to water bodies across a wide diversity of habitats; common around the Uoro and Laña rivers (Lasso et al. 2002).</p> <p>Specimens examined. Twenty-three specimens. Equatorial Guinea (MNCN 20834). Bioko: without precise locality (MNCN 8093, MNCN 8097, MNCN 23887–23888); without precise locality October 1897 (MNCN 45307); Finca Carboneras, 3 November 1983 (EBD, without number); Bioko Sur, 13 November 1983 (EBD 21390); Malabo, 24 January 1933 (MNCN 8098–8099). Río Muni: from Río Campo to Río Benito, Río Muni (MNCN 8095- 8096); San Joaquín de Ndyiacom, 1987 (EBD 27482); Miboman (30 km from Niefang), 1 February 1987 (EBD 25124), 01 August 1986 (EBD 25127); Besisebe, Río Ntem, 31 August 1986 (EBD 25126); Patio de Alosa, Niefang, 20 September 1986 (EBD 20913, EBD 20916), 20 August 1985 (EBD 21757); Miboman, Km 27, carretera Movo-Bata, 24 July 1985 (EBD 20914); Miboman, carretera Niefang, 08 July 1986 (EBD 20915); Miboman, December 1987 (EBD 27674); Bata, 17 May 1965 (EBD 2752).</p> </div>	https://treatment.plazi.org/id/03C287E8FFE2FFD3FF4BFF0AFD5EE78B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE2FFD2FF4BF8CAFB4EE1E9.text	03C287E8FFE2FFD2FF4BF8CAFB4EE1E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calabaria reinhardtii (Schlegel 1851)	<div><p>Calabaria reinhardtii (Schlegel, 1851)</p> <p>Figure 16A</p> <p>Eryx reinhardtii Schlegel, 1851: 1. Type locality: “Côte d’or” Ghana. Restricted to Aqua Pim (Akwapim), southern Ghana (Hughes &amp; Barry 1969).</p> <p>Calabaria fusca Gray, 1858: 155. Type locality: “Old Calabar, W. Africa ” Duke Town, Nigeria.</p> <p>Rhoptrura reinhardtii (Schlegel, 1848): Peters, 1858: 340.</p> <p>Rhoptrura petiti Sauvage, 1884: 202. Type locality: “l’ouest de l’Afrique”.</p> <p>Calabaria reinhardti (Schlegel, 1848): Boulenger 1893: 92.</p> <p>Distribution. It ranges from Liberia to eastern Democratic Republic of the Congo. In Equatorial Guinea it has been recorded from Bioko (Drewes et al. 1999) at Basilé, San Carlos (Bocage 1895 a, 1903; Boulenger 1906; Capocaccia 1961a), and Musola (Mertens 1941), and from Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002) (Map 18B).</p> <p>Comments. The genus Calabaria is currently monotypic and has a confusing taxonomic history. It was formerly placed in Pythonidae until molecular phylogenetic analyses clarified its evolutionary history as a member of Boidae (Pyron et al. 2014). Calabaria reinhardtii usually lives mainly inside superficial underground galleries and termite nests (Angelici et al. 2000). Despite its secretive behaviour, it is a well known species among the local people of Equatorial Guinea (Maximiliano Fero, pers. comm.).</p> <p>Specimens examined. Twenty-two specimens. Bioko: without precise locality (MNCN 22818–22819); San Carlos, Bioko, July 1909 (MNCN 22820). Río Muni: Mongomo, 16 April 1986 (EBD 20794, EBD 20797, EBD 20799); road from Mongomo to Aconibe, 29 November 1985 (EBD 20796); Mongomo (road to Añisok), 06 June 1986 (EBD 20798); road from Mfaman to Niefang, 10 March 1986 (EBD 20795); Miboman (Km 27, road to Bata), 15 February 1986; Miboman (30 km from Niefang), 01 February 1987 (EBD 25008); Sanduma, 27 February 1965 (EBD 2845); 10 km before the road of Alosa from Bata, 09 May 1966 (EBD 2846); San Joaquín de Ndyiacom, 01 August 1986 (EBD 25007); San Joaquín de Ndyiacon (road from Bata to Ayamiken), 01 February 1987 (EBD 25009), 03 February 1987 (EBD 25010), December 1987 (EBD 27753, EBD 27779), September 1987 (EBD 27484), 1990 (EBD 31492); Nvom (road from Aconibe to Asok), 08 May 1987 (EBD 25011); Equatorial Guinea, likely part of the material examined by Boulenger (1905) from Cabo San Juan (MNCN 22316, MNCN 22821).</p> </div>	https://treatment.plazi.org/id/03C287E8FFE2FFD2FF4BF8CAFB4EE1E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE3FFD2FF4BFAD1FF5BE541.text	03C287E8FFE3FFD2FF4BFAD1FF5BE541.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasypeltis fasciata Smith 1849	<div><p>Dasypeltis fasciata Smith, 1849</p> <p>Figure 16B</p> <p>Dasypeltis fasciata Smith, 1849: 73. Type locality: “ Sierra Leone ”.</p> <p>Dipsas carinatus Hallowell, 1844: 119 (nomen praeoccupatum).</p> <p>Dasypeltis macrops Boulenger, 1907: 324. Type locality: “ Efulen, Cameroon ”.</p> <p>Distribution. It is distributed from West Africa at Sierra Leone to East Africa at Democratic Republic of the Congo, and southward to Angola. There are no published records for Equatorial Guinea (Map 18C).</p> <p>Comments. We provide the first record of this species for Equatorial Guinea. The specimen housed in the MNCN was collected during the “Expedición Pittaluga”, which took place across Equatorial Guinea in 1909. During this expedition, Pedro Bengoa collected some specimens of Natriciteres fuliginoides and Python sebae at Río Utoche (southern Río Muni), close to the locality where the specimen MNCN 46190 of Dasypeltis fasciata was found (at Vermakoyo) three days earlier. Pittaluga (1910) provided a map with both locality names, currently out-of-date.</p> <p>Specimens examined. One specimen. Río Muni: “Wermakofo” (=Vermakoyo), 07 August 1909 (MNCN 46190).</p> </div>	https://treatment.plazi.org/id/03C287E8FFE3FFD2FF4BFAD1FF5BE541	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE4FFD5FF4BFF42FC5DE154.text	03C287E8FFE4FFD5FF4BFF42FC5DE154.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipsadoboa duchesnii (Boulenger 1901)	<div><p>Dipsadoboa duchesnii (Boulenger, 1901)</p> <p>Figure 16C</p> <p>Leptodira duchesnii Boulenger, 1901b: 10. Type locality: restricted by lectotype designation to “Mandungu, Itimbiri River, Congo-Kinshasa”. Mandongo, North-east of the Democratic Republic of the Congo (Hughes &amp; Barry 1969).</p> <p>Dipsodomorphus brevirostris Sternfeld, 1908a: 411. Type locality: “Yabassi”, Cameroon.</p> <p>Dipsadomorphus viridis Sternfeld, 1908a: 411. Type locality: “Bipindi”, Cameroon.</p> <p>Leptodira nigeriensis Werner, 1913: 27. Type locality: “ Südnigeria (Umgebung von Gana und Sagbama, Niger-Delta)”.</p> <p>Chamaetortus aulicus ellenbergeri Chabanaud, 1916: 375. Type locality: “Lambaréné”.</p> <p>Dipsadomorphus myops Chabanaud, 1917b: 453. Type locality: “ Gabon ”.</p> <p>Dipsoglyphophis duchesnii (Boulenger, 1901): Barbour &amp; Amaral 1927: 26.</p> <p>Dipsadoboa duchesnii (Boulenger, 1901): Bogert 1940: 64.</p> <p>Crotaphopeltis duchesnii duchesnii (Boulenger, 1901): Loveridge 1941: 122.</p> <p>Distribution. It ranges across West and Central Africa, from Nigeria to Uganda, south to Gabon and Republic of the Congo. In Equatorial Guinea it had never been recorded (Map 19A).</p> <p>Comments. We record this species for the first time in Equatorial Guinea based on two specimens from Río Muni. A detailed morphological revision was provided by Rasmussen (1989).</p> <p>Specimens examined. Two specimens. Río Muni: Miboman (road to Niefang), 06 September 1985 (EBD 20790); Miboman (road from Bata to Niefang), 28 July 1987 (EBD 27543).</p> </div>	https://treatment.plazi.org/id/03C287E8FFE4FFD5FF4BFF42FC5DE154	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE4FFD5FF4BFC31FB8FE771.text	03C287E8FFE4FFD5FF4BFC31FB8FE771.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipsadoboa underwoodi Rasmussen 1993	<div><p>Dipsadoboa underwoodi Rasmussen, 1993</p> <p>Figure 17A</p> <p>Dipsadoboa underwoodi Rasmussen, 1993: 169. Type locality: “Mukonjefarm, Mundame (04º35´N, 09º31´E), Cameroon ”.</p> <p>Distribution. It extends from Guinea to Republic of the Congo, south to Gabon. There are no previous published records for Equatorial Guinea (Map 19B).</p> <p>Comments. The specimen MNCN 22554 recorded as Dipsadoboa sp. by Pérez del Val (2001) is herein confidently identified as D. underwoodi. It represents the first record of this species for Equatorial Guinea (at Elobey Island). Rasmussen (1993) described and provided a detailed account of this species. The specimen MNCN 22554 lacks a great portion of the tail and has 193 ventrals and single subcaudals.</p> <p>Specimens examined. One specimen. Island of Elobey, 1885 (MNCN 22554).</p></div> 	https://treatment.plazi.org/id/03C287E8FFE4FFD5FF4BFC31FB8FE771	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE4FFD5FF4BFA14FB86E5C0.text	03C287E8FFE4FFD5FF4BFA14FB86E5C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipsadoboa unicolor Gunther 1858	<div><p>Dipsadoboa unicolor Günther, 1858</p> <p>Dipsadoboa unicolor Günther, 1858: 138. Type locality: “ West Africa ”.</p> <p>Dipsadoboa unicolor viridiventris Laurent, 1956a: 206. Type locality; “ Lubero, Kivu, Zaire ”.</p> <p>Distribution. It extends from Guinea-Bissau to Uganda. In Equatorial Guinea it has been recorded in Bioko (Günther 1896; Boulenger 1896; Rasmussen 1993) at Musola (Boulenger 1906) and in Río Muni at Benito River (Boulenger, 1900) and Monte Alén National Park (Lasso et al. 2002) (Map 19C).</p> <p>Comments. See account of Dipsadoboa viridis for the identity of Bocage’s (1895a) record from Bioko. Mertens (1964) highlighted some difficulties on the identification of D. viridis and D. unicolor, which were later examined by Rasmussen (1993). Branch et al. (2019) revealed that specimens morphologically identified as D. unicolor from Cameroon and Uganda do not form a monophyletic group, being a specimen of D. unicolor from Cameroon more closely related to D. underwoodi than to a specimen identified as D. unicolor from Uganda. Thus, it is likely that this taxon includes more species. Boulenger (1905) identified D. unicolor among the species collected in Cabo San Juan by M. Martínez de la Escalera. However, after the revision of these specimens of Dipsadoboa housed in the MNCN, we reidentified them as D. viridis based on several morphological traits (see the account of D. viridis for morphological details).</p> <p>Specimens examined. No specimens were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFE4FFD5FF4BFA14FB86E5C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFEAFFDBFF4BFF0AFE4DE7F2.text	03C287E8FFEAFFDBFF4BFF0AFE4DE7F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipsadoboa viridis (Peters 1869)	<div><p>Dipsadoboa viridis (Peters, 1869)</p> <p>Figure 17B</p> <p>Anoplodipsas viridis Peters, 1869: 442. Type locality: “Neu-Caledonien” (in error).</p> <p>Dipsadoboa assimilis Matschie, 1893: 173. Type locality: “ Togo, West-Afrika ”.</p> <p>Crotaphopeltis elongata Barbour, 1914: 95. Type locality: “southern Kamerun ”.</p> <p>Dipsadoboa elongata gracilis Laurent, 1956: 211. Type locality: “Teturi, 750 m, rive gauche de l’Ituri, Terr. de Mambasa, Ituri”.</p> <p>Dipsadoboa viridis (Peters, 1869): Rasmussen, 1993: 174.</p> <p>Distribution. This species extends from Guinea to the Democratic Republic of the Congo, Rwanda, and the Republic of the Congo. In Equatorial Guinea it has been recorded in Bioko (see Rasmussen 1993) at Basilé (Bocage 1895 a, 1903) and Musola (Boulenger 1906; Mertens 1941, 1964; Capocaccia 1961a), and in Río Muni at Cabo San Juan (Boulenger 1905; as D. unicolor), San Benito (=Río Benito) (Rasmussen 1993), and in Monte Mitra at Monte Alén National Park (Gonwouo &amp; Nsang 2005) (Map 20A).</p> <p>Comments. Due to its morphological similarity at first sight with D. unicolor, we provide some diagnostic traits of the examined specimens. In comparison to D. unicolor, in D. viridis the diameter of the eye is equal to or greater than the distance between eye and nostril, the frontal plate longer than broad, and there are 193–238 ventrals and 71–112 subcaudals. All the examined specimens are within the limits of variation of these diagnostic traits (Table 1). In contrast, D. unicolor has the diameter of the eye less than the distance between eye and nostril, 181–220 ventrals and 52–78 subcaudals. The specimens recorded by Bocage (1895a) from Bioko (ssp. D. v. assimilis Matschie, 1893, a current synonym of D. viridis) were re-examined and confirmed as D. viridis by Rasmussen (1993), who also verified the presence of D. unicolor in Bioko and provided a detailed account of D. viridis.</p> <p>Specimens examined. Six specimens. Río Muni: likely Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 23817, MNCN 50527); Miboman (road to Bata, Km 29), 15 November 1986 (EBD 20830); San Joaquín de Ndyiacom (Ayamiken), 01 August 1987 (EBD 25029, EBD 25059); San Joaquín de Ndyiacom, 1990 (EBD 31499).</p> </div>	https://treatment.plazi.org/id/03C287E8FFEAFFDBFF4BFF0AFE4DE7F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFEAFFD8FF4BF98CFEC0E32F.text	03C287E8FFEAFFD8FF4BF98CFEC0E32F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipsadoboa weileri (Lindholm 1905)	<div><p>Dipsadoboa weileri (Lindholm, 1905)</p> <p>Figure 17C</p> <p>Dipsadophidium weileri Lindholm, 1905: 186. Type locality: “Umgegend von Bibundi (Kamerun)”, vicinity of Bibundi, Cameroon.</p> <p>Dipsadoboa isolepis Boulenger, 1907: 325. Type locality: “Efulen”, Cameroon.</p> <p>Dipsadoboa weileri (Lindholm, 1905): Müller 1910: 607.</p> <p>Distribution. This species extends from Guinea to Sudan, Democratic Republic of the Congo, and Republic of Congo. In Equatorial Guinea it has been recorded from Río Muni at Benito River (Boulenger 1900; Rasmussen 1993) (Map 20B).</p> <p>Comments. Rasmussen (1993) provided a detailed account of this species. This species has been confused with D. unicolor; the record of D. unicolor from Benito River by Boulenger (1900) was reidentifyied as D. weileri (Rasmussen, 1993).</p> <p>Specimens examined. Six specimens. Río Muni: Miboman (road to Niefang), 29 March 1984 (EBD 20946); Miboman (30 km from Niefang), 01 February 1987 (EBD 25056); Miboman, April 1988 (EBD 27750), December 1987 (EBD 27751), without date (EBD 25065); San Joaquín de Ndyiacom (road from Bata to Ayamiken), 20 May 1987 (EBD 25057).</p> </div>	https://treatment.plazi.org/id/03C287E8FFEAFFD8FF4BF98CFEC0E32F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE9FFD8FF4BFE2BFCA3E6A3.text	03C287E8FFE9FFD8FF4BFE2BFCA3E6A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hapsidophrys lineatus Fischer 1856	<div><p>Hapsidophrys lineatus Fischer 1856</p> <p>Figure 18A</p> <p>Hapsidophrys lineatus Fischer, 1856. Type locality: “ Elmina, Ghana ”.</p> <p>Gastropyxis orientalis Werner, 1909: 55. Type locality: “ Deutsch-Ostafrika ”, German East Africa (= Burundi, Rwanda, and Tanzania).</p> <p>Distribution. This species extends from Sierra Leona to Uganda, southward to northern Angola. In Equatorial Guinea it has been recorded in Bioko (Bocage 1895 a, 1903; Drewes et al. 1999) at Ureca (Mertens 1964a), and in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005). There is a record from iNaturalist at Moka, Bioko (03°20′00.04″N, 08°40′20.08″E) (Map 20C).</p> <p>Comments. Martínez y Sáez (1886) and Pérez del Val (2001) recorded H. lineatus based on a misidentification of the specimens MNCN 20286–20289 and MNCN 20290–20293, which are herein reidentified as H. smaragdina. As a result, the first record of H. lineatus for Río Muni corresponds to Lasso et al. (2002). Mertens (1964) reported a case of predation over a juvenile of Amnirana albolabris (Hallowell, 1856) (Anura: Ranidae) from Ureca, Bioko.</p> <p>Specimens examined. Ten specimens. Equatorial Guinea (without specific locality) (EBD 25130). Río Muni: without specific locality (MNCN 23815); Miboman (from Bata to Niefang), March 1987 (EBD 25015), 28 July 1987 (EBD 27511); Miboman (road to Bata, Km 29), 15 November 1987 (EBD 20831, EBD 20833); Miboman (road from Movo to Niefang), August 1987 (EBD 27539); San Joaquín de Ndyiacom, 1990 (B9579; without EBD number); San Joaquín de Ndyiacom (road to Ayamiken), 25 June 1987 (EBD 27545); San Joaquín de Ndyiacom (road from Bata to Niefang), 27 July 1987 (EBD 27547).</p> </div>	https://treatment.plazi.org/id/03C287E8FFE9FFD8FF4BFE2BFCA3E6A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFE9FFDFFF4BFB7FFAD9E3C7.text	03C287E8FFE9FFDFFF4BFB7FFAD9E3C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hapsidophrys smaragdina (Schlegel 1837)	<div><p>Hapsidophrys smaragdina (Schlegel, 1837)</p> <p>Figure 18B</p> <p>Dendrophis smaragdina Schlegel, 1837: 237. Type locality: “Gold Coast”, Ghana.</p> <p>Leptophis gracilis Hallowell, 1844. Type locality: “ Liberia, W. Africa ”.</p> <p>Leptophis smaragdina (Schlegel, 1837): Hallowell 1854b: 100.</p> <p>Hapsidophrys coeruleus Fischer, 1856: 111. Type locality: none provided; but according to the general comments: “WestAfrika”.</p> <p>Gastropyxis smaragdina (Schlegel, 1837): Cope 1861b: 558.</p> <p>Ahaetulla smaragdina (Schlegel, 1837): Sclater 1891: 35.</p> <p>Hapsidophrys smaragdina (Schlegel, 1837): Boettger 1889: 279</p> <p>Tropidophidion steini Werner, 1902: 335. Type locality: “ Boké am Rio Nunez ”, Guinea.</p> <p>Distribution. This species extends from Gambia to Uganda, and southward to northern Angola. In Equatorial Guinea it has been recorded in Bioko (Boulenger 1894) at San Carlos and Natividad (Hacienda Natividad) (Bocage 1895 a, 1903; Mertens 1964a) and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 21A).</p> <p>Comments. One of the commonest snake species in Río Muni. The specimens erroneously recorded by Martínez y Sáez (1886) and Pérez del Val (2001) as H. lineatus (MNCN 20286–20289 and MNCN 20290–20293) are herein regarded as H. smaragdina.</p> <p>Specimens examined. Thirty-seven specimens. Equatorial Guinea (without specific locality), July 1891 (MNCN 23810). Island of Elobey (2 specimens) and Río Muni, from Río Campo to Río Benito (2 specimens) (MNCN 20286–20289). Río Muni: without specific locality, likely part of the material examined by Boulenger (1905) from Cabo San Juan (MNCN 22571, 22572, 22573–22575); San Joaquín de Ndyiacom, December 1987 (EBD 27733); San Joaquín de Ndyiacon (road from Bata to Ayamiken), August 1987 (EBD 27531), 20 May 1987 (EBD 24965), 03 February 1987 (EBD 25013 –25014); San Joaquín de Ndyiacom, 1990 (B9572, EBD 31502); Nvom (road from Aconibe to Asoc), 08 May 1987 (EBD 24964); Miboman (road from Bata to Niefang), March 1987 (EBD 24966); Asonga, Bata, 08 April 1986 (EBD 20785); Miboman (road to Bata, Km 29), 15 November 1986 (EBD 20832); Miboman (road from Bata to Niefang), 28 July 1987 (EBD 27535); Mongomo, 16 April 1986 (EBD 20786, EBD 20788); Suguisa, Niefang, 29 November 1985 (EBD 20787); Ayene to Niefang, 15 June 1986 (EBD 20941); Bata (EBD 23091); Colegio la Salle, Bata, 1967 (EBD 2859); Ncoaebeñ (=Ncoeakieñ), 16 June 1984 (EBD 18360); Niefang (road to Evinayong), 15 October 1984 (EBD 18366); Road from Movó to Bata, Km 14, 13 June 1984 (EBD 18371); Bata, Litoral (MNCN 22570). Bioko (MNCN 20290–20293; MNCN 22568–22569).</p> </div>	https://treatment.plazi.org/id/03C287E8FFE9FFDFFF4BFB7FFAD9E3C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFEEFFDFFF4BFD13FF5BE40E.text	03C287E8FFEEFFDFFF4BFD13FF5BE40E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothamnus carinatus (Anderson 1901)	<div><p>Philothamnus carinatus (Anderson, 1901)</p> <p>Figure 18C</p> <p>Chlorophis carinatus Andersson, 1901: 9. Type locality: “Mapanja, Cameroon ” and “ Cameroon ”. Restricted to Mapanja by Mertens (1964).</p> <p>Philothamnus nigrofasciatus Buchholz &amp; Peters, 1875: 199. Type locality: “Cameruns”.</p> <p>Philothamnus heterodermus carinatus Andersson, 1901: Loveridge 1951: 6.</p> <p>Distribution. This species is widespread from West Africa (Guinea Konakry) to East Africa (Kenya) and southward to southern Democratic Republic of the Congo. In Equatorial Guinea it has been recorded in Bioko at Bococo (Mertens 1941; Loveridge 1958), Moka (Mertens 1964a) and Bahía de San Carlos (Boulenger 1906; Capocaccia 1961a), and in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 21B).</p> <p>Comments. All the specimens examined by us have 13 scale rows at mid-body. This species was treated by some authors as a subspecies of P. heterodermus (Loveridge 1958). However, it has been suggested that P. carinatus could represent a species complex (Engelbrecht et al. 2019). It is formed by at least two main clades, one ranging across Gabon-western Republic of the Congo and the other occurring at the eastern of Democratic Republic of the Congo (Engelbrecht et al. 2019). The Philothamnus carinatus species complex is sister to P. heterodermus (Engelbrecht et al. 2019).</p> <p>The specimen recorded from Bahía de San Carlos, Bioko, by Boulenger (1906) was subsequently cited by Loveridge (1958) as P. heterodermus heterodermus and re-identified by Capocaccia (1961a) as P. heterodermus carinatus. Based on the number of scale rows at mid-body (13 scale rows), we follow Capocaccia (1961a) and Mertens (1964) in considering this specimen as P. carinatus.</p> <p>Specimens examined. Eleven specimens. Río Muni: Miboman, 1985–1986 (EBD 20927 –20930); Miboman (30 km from Niefang), 25 February 1987 (EBD 25018, EBD 25020 –25021); Miboman (road to Niefang), 28 July 1987 (EBD 27548); Miboman, 06 September 1986 (EBD 20792); Ayamiken, 01 August 1986 (EBD 25019, EBD 25022).</p> </div>	https://treatment.plazi.org/id/03C287E8FFEEFFDFFF4BFD13FF5BE40E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFEEFFDEFF4BF910FEF7E39F.text	03C287E8FFEEFFDEFF4BF910FEF7E39F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothamnus girardi Bocage 1893	<div><p>Philothamnus girardi Bocage, 1893</p> <p>Figure 19A</p> <p>Philothamnus girardi Bocage, 1893a: 46. Type locality: “Anno Bom Island, Guinea ”. Annobon, Equatorial Guinea.</p> <p>Distribution. Endemic to Annobon (Bocage, 1893 a,b; Loveridge 1958; Jesus et al. 2009) (Map 21C).</p> <p>Comments. Philothamnus girardi could probably be a junior synonym of P. dorsalis (Bocage, 1866a) (Engelbrecht et al. 2019). However, as previously reported (Loveridge 1958), all the specimens from Annobon examined by us have 13 scale rows at mid-body, instead of the 15 rows shown by P. dorsalis (Chippaux &amp; Jackson 2019). Philothamnus girardi has been treated by some authors as a subspecies of P. semivariegatus (Mertens 1934; Loveridge 1958), although this last taxon does not form a monophyletic unit and is not closely related to P. girardi (see Engelbrecht et al. 2019). Philothamnus thomensis from São Tomé Island is the sister species to the clade formed by P. girardi and P. dorsalis (Engelbrecht et al. 2019).</p> <p>Specimens examined. Fourteen specimens. Equatorial Guinea (without specific locality), November 1987 (EBD 27758). Annobon: Annobon, 1959 [MNCN 15834, MNCN 23167 (2 specimens)], 22 November 1987 (EBD 27762); San Antonio, Annobon, 19 November, 1987 (EBD 27766, EBD 27757); Annobon airport, 25 November 1987 (EBD 27769 –27772); San Pedro, Annobon, 11 November 1987 (EBD 27759); San Juan, Annobon (EBD 27760 –27761).</p> </div>	https://treatment.plazi.org/id/03C287E8FFEEFFDEFF4BF910FEF7E39F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFEFFFDEFF4BFE63FD3AE6BE.text	03C287E8FFEFFFDEFF4BFE63FD3AE6BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothamnus heterodermus (Hallowell 1857)	<div><p>Philothamnus heterodermus (Hallowell, 1857)</p> <p>Figure 19B</p> <p>Chlorophis heterodermus Hallowell, 1857: 54. Type locality: “Gaboon”, Gabon.</p> <p>Ahaetulla heteroderma (Hallowell, 1857): Günther 1863c: 285.</p> <p>Philothamnus heterodonta (Hallowell, 1857): Sauvage 1884: 201 (in error).</p> <p>Chlorophis cyaneus Hecht, 1929: 331. Type locality: “ Aioshöle am Nyong, Kamerun” Cameroon.</p> <p>Distribution. This species extends from West Africa (Guinea Bissau) to East Africa (Tanzania) and southward to northern Angola. In Equatorial Guinea it has been recorded in Río Muni at Bata (Pérez del Val 2001) and Monte Alén National Park (Lasso et at. 2002) (Map 22A).</p> <p>Comments. All the specimens examined by us present 15 scale rows at mid-body. Philothamnus heterodermus is sister to the P. carinatus species complex (Engelbrecht et al. 2019). An asymmetric pattern in the fragmentation of head temporal scales has been detected for some specimens, such as in MNCN 22864. See the account of P. carinatus for comments on the record of Boulenger (1906) from Bahía de San Carlos, Bioko.</p> <p>Specimens examined. Seven specimens. Río Muni: Bata, January 1933 (MNCN 22864); San Joaquín de Ndyiacom (road to Ayamiken), 01 August 1986 (EBD 25017), 25 June 1987 (EBD 27537); Miboman, 08 July 1986 (EBD 20793), 01 September 1984 (EBD 18358); Miboman (road from Movo to Niefang), 28 July 1987 (EBD 27544); Bata, December 1987 (EBD 27775).</p> </div>	https://treatment.plazi.org/id/03C287E8FFEFFFDEFF4BFE63FD3AE6BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFEFFFDEFF4BFB40FB48E5C7.text	03C287E8FFEFFFDEFF4BFB40FB48E5C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothamnus nitidus (Gunther 1863)	<div><p>Philothamnus nitidus (Günther, 1863)</p> <p>Figure 19C</p> <p>Ahaetulla nitida Günther, 1863c: 286. Type locality: “ Demerara ” Guyana (in error), corrected to “ Lagos ”, Nigeria by Boulenger (1894).</p> <p>Ahaetulla lagoensis Günther, 1872: 26. Type locality: “ Lagos ”, Nigeria.</p> <p>Philothamnus nitidus (Günther, 1863): Boulenger 1894: 100.</p> <p>Philothamnus nitidus loveridgei Laurent, 1960: 40. Type locality: Itula, Shabunda, Kivu, Democratic Republic of the Congo.</p> <p>Distribution. This species extends from Sierra Leone and Guinea to Congo, and southward to Democratic Republic of the Congo and Angola. In Equatorial Guinea it has been recorded in Bioko at San Carlos de Luba [as P. semivariegatus (Bocage 1895 a, 1903; Loveridge 1958)] and in Río Muni at Monte Alén National Park (Lasso et al. 2002). Loveridge (1958) suggested that Bocage’s (1895a) record of P. semivariegatus probably belongs to P. s. nitidus (currently P. nitidus) (Map 22B).</p> <p>Comments. Previous reports on P. semivariegatus in Bioko are herein treated as P. nitidus following Loveridge’s (1958) assumption on the identity of the Biokoan population [“ Fernando Po: San Carlos (Bocage, 1895a: more probably s. nitidus)” fide Loveridge (1958): 119]. Philothamnus nitidus is the sister taxon to P. angolensis, and both are nested within the paraphyletic assemblage of P. semivariegatus (Engelbrecht et al. 2019).</p> <p>Specimens examined. Ten specimens. Río Muni: Miboman, 1985–1986 (EBD 20931, EBD 20789), December 1987 (EBD 27734, EBD 27774); Miboman (road from Movo to Niefang), August 1987 (EBD 27536); Miboman (Km 27, road from Bata to Movo), 29 July 1985 (EBD 20791); Miboman (30 km from Niefang), 25 February 1987 (EBD 25021); Colegio La Salle, Bata, 1967 (B2859, without EBD number); San Joaquín de Ndyiacon (road to Bata), 28 July 1987 (EBD 27534); San Joaquín de Ndyiacom, 1990 (B 9580, without EBD number).</p> </div>	https://treatment.plazi.org/id/03C287E8FFEFFFDEFF4BFB40FB48E5C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFF0FFC1FF4BFF42FB23E64E.text	03C287E8FFF0FFC1FF4BFF42FB23E64E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thelotornis kirtlandii (Hallowell 1844)	<div><p>Thelotornis kirtlandii (Hallowell, 1844)</p> <p>Figure 20A</p> <p>Leptophis kirtlandii Hallowell, 1844: 62. Type locality: “ Liberia ”, Republic of Liberia.</p> <p>Oxybelis lecomtei Duméril &amp; Bibron, 1854: 821. Type locality: “ Gabon ”.</p> <p>Tragops rufulus Duméril &amp; Bibron, 1854: 827. Type locality: “ Sénégal ”.</p> <p>Dryophis kirtlandii (Hallowell, 1844): Hallowell 1854b: 100.</p> <p>Oxybelis violacea Fischer, 1856: 91. Type locality: “ Edina, Grand Bassa County, West-Afrika ”.</p> <p>Oxybelis kirtlandii (Hallowell, 1844): Hallowell 1857: 59.</p> <p>Thelotornis kirtlandii (Hallowell, 1844): Fischer 1884b: 11.</p> <p>Distribution. This species ranges across tropical forests from West Africa (Guinea Bissau) to East Africa at southern Sudan and Uganda, and south to Angola. In Equatorial Guinea it has been recorded in the islands of Bioko (Günther 1863a; Bocage 1895 a, 1903) at Bahía de San Carlos (Boulenger 1906; Loveridge 1944; Capocaccia 1961a) and Elobey (Pérez del Val 2001), and in Río Muni at Bata (Pérez del Val 2001), Cabo San Juan (Boulenger 1905; Pérez del Val 2001) and Monte Alén National Park (Lasso et al. 2002) (Map 22C).</p> <p>Comments. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.715016&amp;materialsCitation.latitude=3.3552778" title="Search Plazi for locations around (long 8.715016/lat 3.3552778)">We</a> provide the first record of this species for the Island of Corisco. The specimens MNCN 23257 and MNCN 23259 contained two and six eggs respectively (labelled as follows: MNCN 23260 for the two eggs of the female MNCN 23257, and MNCN 23258 for the six eggs of the female MNCN 23259). The eggs are elongated in shape; both MNCN 23260 are around 5,1 x 1,5 cm, whereas the six eggs (MNCN 23260) are around 4 x 1,8 cm. There is a record from iNaturalist at Riaba, Bioko (03°21′19.00″N, 08°42′54.06″E).</p> <p>Specimens examined. Twelve specimens. Island of Corisco (MNCN 23257, MNCN 23259). Island of Elobey (MNCN 23261). Bioko: 1876-1877 (MNCN 23264–23265). Río Muni: Bata, Litoral, July 1891 (MNCN 23262); Cabo San Juan, likely as part of the material examined by Boulenger (1905) (MNCN 23263); Miboman, December 1987 (EBD 27756); Miboman (road from Movo to Niefang), February 1987 (EBD 27540); San Joaquín de Ndyiacom (road to Ayamiken), 01 August 1986 (EBD 25012); San Joaquín de Ndyiacom, 1990 [2 specimens labelled as B9500 and B9632; both with a shared number (EBD 31495) with a specimen of Thrasops batesii].</p> </div>	https://treatment.plazi.org/id/03C287E8FFF0FFC1FF4BFF42FB23E64E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFF0FFC0FF4BFA88FECFE2CB.text	03C287E8FFF0FFC0FF4BFA88FECFE2CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thrasops aethiopissa Gunther 1862	<div><p>Thrasops aethiopissa Günther, 1862</p> <p>Figure 20B</p> <p>Rhamnophis aethiopissa Günther, 1862: 129. Type locality: “West Africa”.</p> <p>Crypsidomus aethiops (Günther, 1862): Boettger 1888: 64.</p> <p>Thrasops splendens Andersson, 1901: 11. Type locality: “Mapanja,… Bibundi”, Mapanja and Bibundi, Cameroon.</p> <p>Rhamnophis ituriensis Schmidt, 1923: 81. Type locality: Niapu, Democratic Republic of the Congo.</p> <p>Rhamnophis aethiopissa elgonensis Loveridge 1929: 24. Type locality: “ Yala (=Lukosa) River at the foot of Mount Elgon”.</p> <p>Thrasops aethiopissa aethiopissa (Günther, 1862): Trape &amp; Roux-Estève 1995: 40.</p> <p>Distribution. It is distributed from West Africa in Guinea to East Africa in Democratic Republic of the Congo and Uganda. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) and in Bioko (Boulenger 1896; Loveridge 1944) between San Carlos and Moka (Merterns 1964) (Map 23A).</p> <p>Comments. Treated by some authors within the genus Rhamnophis Günther, 1862 (see Chippaux &amp; Jackson 2019 for a discussion). The specimen EBD 27755 contained in its stomach a half digested frog of the species Chiromantis rufescens (Günther, 1869) (Anura: Rhacophoridae).</p> <p>Specimens examined. Sixteen specimens. Bioko: Malabo, July 1986 (EBD 20821). Río Muni: Nvom (road from Aconibe to Asoc), May 1987 (EBD 24953); Miboman (road from Bata to Niefang) (EBD 24954); Miboman (30 km from Bata), 14 October 1985 (EBD 20819); Miboman (road to Niefang, Km 30), 01 February 1987 (EBD 24958 –24959); Miboman (road to Movo), 1.753596º, 9.780538º, 01 September 1984 (EBD 18367); Miboman (road from Movo to Niefang), February 1987 (EBD 27541), 1987 (EBD 25058); Mongomo, 01 April 1985 (EBD 20822), 16 April 1986 (EBD 20921); San Joaquín de Ndyiacon (road from Bata to Ayamiken), 03 February 1987 (EBD 24960 –24961), February 1987 (EBD 27542); San Joaquín de Ndyiacom, April 1988 (EBD 27755), 1990 (B9544; without EBD label).</p> </div>	https://treatment.plazi.org/id/03C287E8FFF0FFC0FF4BFA88FECFE2CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFF1FFC0FF4BFED7FBDFE139.text	03C287E8FFF1FFC0FF4BFED7FBDFE139.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thrasops batesii Boulenger 1908	<div><p>Thrasops batesii Boulenger, 1908</p> <p>Figure 20C</p> <p>Thrasops batesii Boulenger, 1908: 93. Type locality: “ Efulen and Akok, Kribi River district, and Ja River district ”, Cameroon.</p> <p>Rhamnophis batesii (Boulenger, 1908): Schmidt 1923: 83.</p> <p>Distribution. It ranges from Cameroon to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 23B).</p> <p>Comments. Treated by some authors within the genus Rhamnophis Günther, 1862 (see Chippaux &amp; Jackson 2019 for a discussion).</p> <p>Specimens examined. Three specimens. Río Muni: Miboman (30 km from Niefang), 01 February 1987 (EBD 25035); Miboman (Km 27, road from Bata to Niefang) October 1987 (EBD 27732); San Joaquín de Ndyiacom, 1990 [B9574; shared label (EBD 31495) with two specimens of Thelotornis kirtlandii].</p> </div>	https://treatment.plazi.org/id/03C287E8FFF1FFC0FF4BFED7FBDFE139	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFF1FFC0FF4BFCC6FD54E419.text	03C287E8FFF1FFC0FF4BFCC6FD54E419.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thrasops flavigularis (Hallowell 1852)	<div><p>Thrasops flavigularis (Hallowell, 1852)</p> <p>Figure 21A–B</p> <p>Dendrophis flavigularis Hallowell, 1852: 205. Type locality: “ Gabon ” [Loveridge (1944) corrected the type locality to Gabon, originally given as “ Liberia, Western Africa”].</p> <p>Thrasops flavigularis (Hallowell, 1852): Hallowell 1857: 67.</p> <p>Hapsidophrys niger Günther, 1872: 25. Type locality: “Gaboon”.</p> <p>Thrasops pustulatus Buchholz &amp; Peters, 1875 in Peters (1875): 199. Type locality: “Cameruns; Mungo”.</p> <p>Distribution. It extends from Nigeria to the Democratic Republic of the Congo, south to northern Angola. In Equatorial Guinea it has been recorded both in Bioko at Moka (Boulenger 1906; Loveridge 1944; Capocaccia 1961a; Mertens 1964a) and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002) (Map 23C).</p> <p>Comments. The record of Dispholidus typus of Pérez del Val (2001) labelled as MNCN 19977, is a misidentification with a juvenile of T. flavigularis (Figure 21B). Because no confirmed records of D. typus exist in Equatorial Guinea, we remove it from the reptile species list of the country.</p> <p>Specimens examined. Eleven specimens. Bioko: Malabo, 3º45′N 08º47′E, 02 July 1987 (EBD 20829). Río Muni: Cabo San Juan [MNCN 9052 (recorded by Boulenger 1905), MNCN 19977]; Mongomo, Añisoc (EBD 25005 –25006); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.316667&amp;materialsCitation.latitude=1.6333333" title="Search Plazi for locations around (long 11.316667/lat 1.6333333)">Mongomo</a>, 1º38′N 11º19′E, 16 April 1986 (EBD 20826 –20827); Miboman (road from Bata to Niefang) (EBD 27552); Miboman, 30 km from Niefang (EBD 25036); Miboman (road to Niefang), 08 July 1986 (EBD 20828); Niefang, 01 June 1984 (EBD 18672).</p> </div>	https://treatment.plazi.org/id/03C287E8FFF1FFC0FF4BFCC6FD54E419	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFF1FFCBFF4BF958FE54E00F.text	03C287E8FFF1FFCBFF4BF958FE54E00F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxicodryas adamanteus Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Mwenebatu	<div><p>Toxicodryas adamanteus Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Mwenebatu, Mali, Badjedjea, Penner, Rödel, Rivera, Sterkhova, Johnson, Tapoandjou &amp; Brown, 2021</p> <p>Figure 21C</p> <p>Toxicodryas adamanteus Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Mwenebatu, Mali, Badjedjea, Penner, Rödel, Rivera, Sterkhova, Johnson, Tapoandjou &amp; Brown, 2021.</p> <p>Type locality: “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.2243&amp;materialsCitation.latitude=-0.7465" title="Search Plazi for locations around (long 18.2243/lat -0.7465)">Npenda village</a>, NE of Lake Tumba (00.7465°S, 18.2243°E, 311 m), Equateur Province, DRC ”.</p> <p>Distribution. It extends from the East of the Niger Delta to Kenya, southward to northern Angola (Greenbaum et al. 2021). In Equatorial Guinea it has been recorded both from Bioko (Boulenger 1896; Bocage 1903) and Río Muni, at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002) (Map 24A).</p> <p>Comments. A recently described species splitted from T. pulverulenta (Fisher, 1856) where it was formerly included (Greenbaum et al. 2021). There are no precise records of this species for Bioko (Mertens 1964a).</p> <p>Specimens examined. Fifteen specimens. Equatorial Guinea, without specific locality, but likely part of the material examined by Boulenger (1905) from Cabo San Juan (MNCN 23813). Río Muni: Miboman (30 km from Niefang), 01 February 1987 [EBD 25023 (includes 6 specimens)]; Miboman (road to Niefang), 7–8 July 1986 (EBD 20824 –20825), 6 September 1986 (EBD 20823); Miboman (road of Bata-Movo-Niefang), August 1987 (EBD 27530); Miboman, December 1987 (EBD 27738); Bata, December 1987 (EBD 27746); Bata (road of Movo-Miboman-Niefang), September 1987 (EBD 27518); San Joaquín de Ndyiacom (road from Bata to Ayamiken), 03 February 1987 (EBD 24978).</p> </div>	https://treatment.plazi.org/id/03C287E8FFF1FFCBFF4BF958FE54E00F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFAFFCBFF4BFD13FE05E4AE.text	03C287E8FFFAFFCBFF4BFD13FE05E4AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxicodryas blandingii (Hallowell 1844)	<div><p>Toxicodryas blandingii (Hallowell, 1844)</p> <p>Figure 22A</p> <p>Dipsas blandingii Hallowell, 1844: 170. Type locality: “ Liberia, West Africa ”.</p> <p>Triglyphodon fuscum Duméril, Bibron &amp; Duméril 1854: 1101. Type locality: “ Grand-Bassam, sur la Côte d’Ivoire (Guinée)”, Ivory Coast.</p> <p>Dipsas fasciata Fischer, 1856: 84. Type locality: “Peki (West-Afrika)”, Ghana.</p> <p>Dipsas valida Fischer, 1856: 87. Type locality: “ Edina (Grand Bassa County, West-Afrika)”, Liberia.</p> <p>Dipsas globiceps Fischer, 1856: 89. Type locality: “ Edina (Grand Bassa County, Liberia, West-Afrika)”.</p> <p>Toxicodryas blandingii (Hallowell, 1844): Hallowell 1857: 60.</p> <p>Dipsas fischeri Jan in Duméril, 1859: 212. Type locality: non provided.</p> <p>Triglyphodon fuscum var. obscurum Duméril, 1861: 211. Type locality: “Côte d’Or”, Ghana.</p> <p>Dipsas regalis Jan, 1871: 3 in Jan &amp; Sordelli (1870–1881). Type locality: “Côte d’Or”, Ghana.</p> <p>Dipsas globiceps var. tumboensis Müller, 1885: 688. Type locality: “Tumbo-Insel”, Guinea.</p> <p>Boiga blandingi occidentalis Stucki-Stirn, 1979: 377. Type locality: “Besongabang” Cameroon.</p> <p>Boiga blandingi subfulva Stucki-Stirn, 1979: 381. Type locality: Cameroon.</p> <p>Distribution. It ranges from Senegal to western Democratic Republic of the Congo (west of the confluence of the Congo and Ubangi Rivers) (Greenbaum et al. 2021). In Equatorial Guinea it has been recorded in Bioko at Bococo (Mertens 1941) and San Carlos (Mertens 1964a; Greenbaum et al. 2021), and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 24B).</p> <p>Comments. Populations located at the East of the confluence between Congo and Ubangi Rivers (not in Equatorial Guinea), formerly considered as Toxicodryas blandingii, have been recently separated from T. blandingii and described as T. vexator Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Mwenebatu, Mali, Badjedjea, Penner, Rödel, Rivera, Sterkhova, Johnson, Tapoandjou &amp; Brown, 2021 (Greenbaum et al. 2021). The examined specimen EBD 18673 contained a bird in its stomach.</p> <p>Specimens examined. Eleven specimens. Equatorial Guinea: without specific locality (MNCN 23819), October 1987 (EBD 27797). Bioko: without specific locality (MNCN 22315, MNCN 22555); Riaba (= Concepción) (EBD 31727). Río Muni: San Joaquín de Ndyiacom, 1990 (EBD 31493 –31494, EBD 31503); Nsung, 22 km from Bata, 05 June 1984 (EBD 18363); from Bata to Movo, Km 7, 06 June 1984 (EBD 18673); from Bata to Movo, Km 7, 16 June 1984 (EBD 18359).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFAFFCBFF4BFD13FE05E4AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFBFFCAFF4BFF0AFB2BE065.text	03C287E8FFFBFFCAFF4BFF0AFB2BE065.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grayia caesar (Gunther 1863)	<div><p>Grayia caesar (Günther, 1863)</p> <p>Figure 22B</p> <p>Xenurophis caesar Günther, 1863: 357. Type locality: “Fernando Po”, Bioko, Equatorial Guinea.</p> <p>Grayia caesar (Günther, 1863): Boulenger 1909: 944.</p> <p>Distribution. Central Africa, from Bioko and Cameroon to Democratic Republic of the Congo; it extends southward to Cabinda (northern Angola) (Chippaux &amp; Jackson 2019; Marques et al. 2018). In Equatorial Guinea it has been recorded from Bioko (Günther 1863a; Boulenger 1894; Bocage 1903) (Map 24C).</p> <p>Comments. There are no precise records of this species for Bioko. We provide the first records of this species for Río Muni.</p> <p>Specimens examined. Two specimens. Río Muni: San Joaquín de Ndyiacom (road from Bata to Ayamiken), February 1987 (EBD 25028); Miboman (30 km from Bata), 14 October 1985 (EBD 20945).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFBFFCAFF4BFF0AFB2BE065	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFBFFCAFF4BFD35FBB2E6CA.text	03C287E8FFFBFFCAFF4BFD35FBB2E6CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grayia ornata (Bocage 1866)	<div><p>Grayia ornata (Bocage, 1866)</p> <p>Figure 22C</p> <p>Macrophis ornatus Bocage, 1866b: 47. Type locality: “ Duque de Bragança dans l`interior? d`Angola”, Calandula, Angola.</p> <p>Grayia ornata (Bocage, 1866): Bocage 1895: 104.</p> <p>Distribution. This species ranges from Cameroon to the Democratic Republic of the Congo and Central African Republic, extending southward to Angola (Chippaux &amp; Jackson 2019; Marques et al. 2018). In Equatorial Guinea it has been recorded in Río Muni at Benito River (Boulenger, 1909) and Monte Alén National Park (Pauwels et al. 2000, Lasso et al. 2002) (Map 25A).</p> <p>Comments. A freshwater piscivorous species recorded in the streams of Monte Alén (Lasso et al. 2002). Pauwels et al. (2000) reported the fish Parauchenoglanis Boulenger, 1911 in the stomach of a young male specimen from Monte Alén, currently housed in the Muséum National d’Histoire Naturelle, and labelled as MNHN 1995.9386.</p> <p>Specimens examined. Seven specimens. Río Muni: San Joaquín de Ndyiacom (road from Bata to Ayamiken), 01 August 1987 (EBD 25060 –25061), 20 April 1987 (EBD 25062), 03 February 1987 (EBD 24950); road from San Joaquín de Ndyiacom to Ayamiken, 27 July 1987 (EBD 27510); Río Ñe, road from Evinayong to Aconibe, 08 May 1987 (EBD 24948); Miboman, road to Niefang, 29 March 1986 (EBD 20938).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFBFFCAFF4BFD35FBB2E6CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFBFFCAFF4BFAD7FCFDE5B8.text	03C287E8FFFBFFCAFF4BFAD7FCFDE5B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grayia smythii (Leach 1818)	<div><p>Grayia smythii (Leach, 1818)</p> <p>Figure 23A</p> <p>Coluber smythii Leach, 1818: 409. Type locality: “near Embomma”, Boma, Democratic Republic of the Congo.</p> <p>Coluber laevis Hallowell, 1844: 118. Type locality: Africa.</p> <p>Coronella triangularis Hallowell, 1854b. Type locality: Western Africa.</p> <p>Grayia silurophaga Günther, 1858: 51. Type locality: West Africa.</p> <p>Grayia smythii (Leach, 1818): Günther 1895: 525</p> <p>Distribution. Broadly distributed across sub-saharan Africa, from Senegal at the West to Kenya and Tanzania at the East, extending southward to Angola. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 25B).</p> <p>Comments. Notes regarding the type specimen/s and versions of the specific epithet are provided by Wallach et al. (2014) and Marques et al. (2018). A freshwater species recorded in the streams of Monte Alén National Park in Río Muni (Lasso et al. 2002).</p> <p>Specimens examined. Two specimens. Río Muni: Miboman, (Km 27, road from Movo to Bata) 15 November 1986 (EBD 20926); San Joaquín de Ndyiacom (EBD 31510).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFBFFCAFF4BFAD7FCFDE5B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFFFFCEFF4BFEBAFBD8E126.text	03C287E8FFFFFFCEFF4BFEBAFBD8E126.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afronatrix anoscopus (Cope 1861)	<div><p>Afronatrix anoscopus (Cope, 1861)</p> <p>Tropidonotus anoscopus Cope, 1861c: 299. Type locality: “ Cuba ” in error fide Loveridge (1941); corrected to Liberia (Loveridge 1941).</p> <p>Tropidonotus ferox Günther, 1863: 355. Type locality: “ Fernando Po ”, Bioko.</p> <p>Natrix anoscopus (Cope, 1861): Cope 1892b: 673.</p> <p>Helicops gendrii Boulenger, 1910: 512. Type locality: “ Labé, French Guinea ”.</p> <p>Natrix firestonei Taylor &amp; Weyer, 1958: 1210. Type locality: Liberia.</p> <p>Afronatrix anoscopus (Cope, 1861): Rossman &amp; Eberle 1977: 42.</p> <p>Distribution. This species ranges from Senegal to Cameroon. In Equatorial Guinea it has been recorded in Bioko (Günther 1863a; Boulenger 1893; Bocage 1903) (Map 25C).</p> <p>Comments. There are no precise localities for this species in Equatorial Guinea. The genus Afronatrix is sister to the clade comprising Hydraethiops Günther, 1872 and Helophis Witte &amp; Laurent, 1942 (Deepak et al. 2021).</p> <p>Specimens examined. No specimens have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFFFFFCEFF4BFEBAFBD8E126	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFFFFCEFF4BFC30FAB7E4B1.text	03C287E8FFFFFFCEFF4BFC30FAB7E4B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydraethiops melanogaster Gunther 1872	<div><p>Hydraethiops melanogaster Günther, 1872</p> <p>Figure 23B</p> <p>Hydraethiops melanogaster Günther, 1872: 28. Type locality: “ Gabon ”.</p> <p>Helicops marginatus Fischer, 1883: 11. Type locality: “Süd-Amerika”.</p> <p>Helicops lineofasciatus Sauvage, 1884: 203. Type locality: “ Majumba (Congo)”.</p> <p>Hydraethiops melanogaster Günther, 1872: Boulenger 1893: 281.</p> <p>Distribution. This species ranges from Cameroon to Republic Democratic of the Congo. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 26A).</p> <p>Comments. The genus Hydraethiops is sister to Heliophis (Deepak et al. 2021).</p> <p>Specimens examined. Forty specimens. Equatorial Guinea: without specific locality (EBD 25131). Río Muni: Miboman, December 1987 (EBD 27728 –27729); Miboman, 30 km from Bata, 14 October 1985 (EBD 20962), 14 October 1985 (EBD 20967); Miboman (Km 27, road from Movo to Bata), 15 November 1985 (EBD 20963 –20964), 15 February 1986 (EBD 20969), 29 July 1985 (EBD 22771); Miboman (road to Niefang), 06 October 1985 (EBD 20965 –20966), 29 March 1986 (EBD 20968, EBD 20970); Miboman (road from Bata to Niefang) (EBD 24937 – 24941), 28 July 1987 (EBD 27513, EBD 27554 –27555); Miboman (Km 30, road to Niefang), 01 February 1987 (EBD 24942 –24947); Miboman (road of Bata-Movo-Niefang), August 1987 (EBD 27522 –27529); Río Miong, Noayong, 23 km from Evinayong, 16 April 1987 (EBD 24936); Mongomo, 16 April 1986 (EBD 20961); Yengüe, 26 February 1988 (EBD 27763 –27764); province of Litoral (without specific locality) (EBD 27739).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFFFFCEFF4BFC30FAB7E4B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFFFFCDFF4BF881FB69E18A.text	03C287E8FFFFFFCDFF4BF881FB69E18A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Natriciteres fuliginoides (Gunther 1858)	<div><p>Natriciteres fuliginoides (Günther, 1858)</p> <p>Figure 23C</p> <p>Coronella fuliginoides Günther, 1858: 39. Type locality: “ West Africa ”.</p> <p>Mizodon longicauda Günther, 1863a: 352. Type locality: “ Fernando Po ”.</p> <p>Tropidonotus fuliginoides (Günther, 1858): Boulenger 1893: 217.</p> <p>Mizodon fuliginoides (Günther, 1858): Bocage 1895a: 75.</p> <p>Natrix fuliginoides (Günther, 1858): Loveridge 1936b: 21.</p> <p>Neusterophis fuliginoides (Günther, 1858): Bogert 1940: 33.</p> <p>Natriciteres fulinoides (Günther, 1858): (sic) Mattison 1995: 133</p> <p>Natriciteres fuliginoides (Günther, 1858): Broadley 1998a</p> <p>Distribution. This species is widespread across forests from Sierra Leone to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded from Bioko (Günther 1863a; Boulenger 1893) at Natividad (Hacienda Natividad), San Carlos (Bocage 1895 a, 1903; Loveridge 1958) and Musola (Boulenger 1906; Loveridge 1958; Capocaccia 1961a), and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002) (Map 26B).</p> <p>Comments. The finding of the exact locality of the specimen MNCN 23814, collected from “Río Utoche” was possible thanks to the old geographic map of Río Muni provided by Gustavo Pittaluga (Pittaluga 1910). All the examined specimens present 17 scale rows at mid-body.</p> <p>Specimens examined. Twenty-one specimens. Río Muni: without specific locality, but likely part of the material examined by Boulenger (1905) from Cabo San Juan (MNCN 23812, MNCN 23816); Río Utoche, 10 August 1909 (MNCN 23814); province of Litoral (without specific locality), 22 February 1988 (EBD 2774); Miboman, road to Niefang, 30 km from Bata, 14 October 1985 (EBD 20948); Miboman (road to Niefang), 29 March 1986 (EBD 20938), 14 October 1985 (EBD 20939), 08 July 1986 (EBD 20940), 08 July 1985 (EBD 20947); Miboman, October 1987 (EBD 27744); Miboman, 30 km from Niefang, 01 February 1987 (EBD 24979), without date (EBD 24980 – 24981); Miboman (from Movo to Niefang), 1987 (EBD 25063); Miboman, April 1988 (EBD 27727); Miboman (road from Bata to Niefang), 28 July 1987 (EBD 27551); San Joaquín de Ndyiacom, October 1987 (EBD 27743); Litoral province (without specific locality) (EBD27740 –27741); San Joaquín de Ndyiacom (road from Bata to Niefang), 25 June 1987 (EBD 27548); San Joaquín de Ndyiacom, September 1987 (EBD 27519).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFFFFCDFF4BF881FB69E18A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFFCFFF3FF4BFAB5FD33E3E3.text	03C287E8FFFCFFF3FF4BFAB5FD33E3E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aparallactus modestus (Gunther 1859)	<div><p>Aparallactus modestus (Günther, 1859)</p> <p>Figure 24A</p> <p>Elapops modestus Günther, 1859: 161. Type locality: “ West Africa ”.</p> <p>Pariaspis plumbeatra Cope, 1860b: 242. Type locality: “ Liberia ”.</p> <p>Elapops (Calamaria) petersii Jan, 1862: 32. Type locality: “ Costa d’ oro”.</p> <p>Aparallactus peraffinis Werner, 1897: 404. Type locality: “ Kamerun, Hinterland ”.</p> <p>Aparallactus modestus ubangensis Boulenger, 1897b: 279. Type locality: Zongo, Ubangi Rapids, Belgian Congo.</p> <p>Aparallactus flavitorques Boulenger, 1901b: 11. Type locality: “ Lubué, Kasai, Belgian Congo ”.</p> <p>Aparallactus dolloi Werner 1902: 346. Type locality: Banzyville, Ubangi River, Belgian Congo.</p> <p>Aparallactus congicus Werner 1902: 346. Type locality: Lingunda, Belgian Congo.</p> <p>Aparallactus batesii Boulenger, 1907: 325. Type locality: Kribi, French Cameroon.</p> <p>Aparallactus christyi Boulenger, 1910: 512. Type locality: Mabira Forest, Chagwe, Uganda.</p> <p>Aparallactus nigrocollaris Chabanaud, 1916: 377. Type locality: French Congo.</p> <p>Aparallactus nigrocollaris roucheti Chabanaud, 1916: 378. Type locality: French Congo.</p> <p>Guyomarchia unicolor Angel, 1923: 348. Type locality: French Congo (probably from near Sangha).</p> <p>Aparallactus graueri Werner 1924: 42. Type locality: “ Beni, Belgian Congo ”.</p> <p>Distribution. This species is widespread across the forests from Guinea to the Democratic Republic of the Congo and Uganda. In Equatorial Guinea it has been recorded from Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 26C).</p> <p>Comments. Two subspecies are currently recognized: A. modestus modestus and A. modestus ubangensis Boulenger, 1897. The latter only occurs from Uganda to the Democatric Republic of the Congo. The morphological traits of the Equatoguinean specimens examined by us are congruent with the nominal subspecies (i.e., only the sixth upper labial in contact with the parietal).</p> <p>Specimens examined. Thirteen specimens. Río Muni: Bata, 1987 (EBD 27517); Miboman (road from Bata to Niefang) (EBD 24968); Miboman (road to Niefang), 29 July 1984 (EBD 20944); Miboman (road Bata-MovoNiefang), August 1987 (EBD 27526, EBD 27538); San Joaquín de Ndyiacom, 01 August 1987 (EBD 25030 –25032); Miboman (Bata-Niefang) (EBD 25033 –25034); San Joaquín de Ndyiacom, 22 February 1988 (EBD 27737); San Joaquín de Ndyiacom (EBD 27532 –27533).</p> </div>	https://treatment.plazi.org/id/03C287E8FFFCFFF3FF4BFAB5FD33E3E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC2FFF3FF4BFDF7FE5AE708.text	03C287E8FFC2FFF3FF4BFDF7FE5AE708.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polemon collaris (Peters 1881)	<div><p>Polemon collaris (Peters, 1881)</p> <p>Figure 24B</p> <p>Microsoma collare Peters 1881b: 148. Type locality: “Macange, Cuango, West-Afrika”, Malanje, Angola.</p> <p>Miodon collaris collaris Witte &amp; Laurent, 1947: 70.</p> <p>Miodon collaris brevior Witte &amp; Laurent, 1947: 69. Type locality: “Vieux-Calabar”, Calabar, Nigeria.</p> <p>Miodon collaris longior Witte &amp; Laurent, 1947: 72. Type locality: “Medje (Uele)”.</p> <p>Polemon collaris (Peters 1881): Welch 1994: 99.</p> <p>Distribution. Widespread in Central Africa from Nigeria to Uganda, south to Angola. In Equatorial Guinea it has been recorded both in Bioko (Bocage 1895 a, 1903) and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 27A).</p> <p>Comments. Loveridge (1944) mentioned the presence of this species in Bioko (“Fernando Po: Spanish Guinea: Esong, near Bakossiberge ”); however, the locality provided corresponds to a region placed in Cameroon. Consequently, we exclude this record from this catalogue. At first sight, it can be confused with Polemon fulvicollis, but they are readily differentiated by the pholidosis pattern, among other traits. The specimens from the EBD B9272, B9273, EBD 18270 and EBD 20943, present the following numbers of subcaudals, respectively: 23, 17, 18 and 20. The number of ventral scales for the specimens EBD 18370 and EBD 20943 are 216 and 200, respectively.</p> <p>Specimens examined. Four specimens. Río Muni: San Joaquín de Ndyiacom, 1990 (EBD 31526 for two specimens with the field numbers B9272 and B9273); Moca, Evinayong, 15 June 1986 (EBD 20943); without specific locality (EBD 18370).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC2FFF3FF4BFDF7FE5AE708	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC2FFF1FF4BFA16FE05E283.text	03C287E8FFC2FFF1FF4BFA16FE05E283.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polemon notatus (Peters 1882)	<div><p>Polemon notatus (Peters, 1882)</p> <p>Figure 24C</p> <p>Microsoma notatum Peters 1882a: 127. Type locality: originally not stated. Designated as Cameroon by Chirio &amp; LeBreton (2007).</p> <p>Cynodontophis aemulans Werner 1902. Type locality: “ Congo ”.</p> <p>Polemon notatus aemulans Trape, J.F. &amp; R. Roux-Estève 1995: 43.</p> <p>Distribution. This species ranges from Cameroon to the western region of Central African Republic and Democratic Republic of the Congo. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 27B).</p> <p>Comments. Two subspecies are recognized on the basis of ventral scale number, P. notatus notatus and P. notatus aemulans (Werner, 1902). In Equatorial Guinea the nominal subspecies is present. The phylogenetic relationships of Aparallactinae are discussed by Portillo et al. (2018); among the included species of Polemon in this study, P. notatus represents the most phylogenetically differentiated lineage (Portillo et al. 2018).</p> <p>Specimens examined. One specimen. Río Muni: Monte Alén National Park (in the EBD collection, without EBD number).</p> <p>ATRACTASPIDINAE</p></div> 	https://treatment.plazi.org/id/03C287E8FFC2FFF1FF4BFA16FE05E283	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC0FFF1FF4BFEBAFAA6E126.text	03C287E8FFC0FFF1FF4BFEBAFAA6E126.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractaspis boulengeri Mocquard 1897	<div><p>Atractaspis boulengeri Mocquard, 1897</p> <p>Atractaspis boulengeri Mocquard, 1897: 16. Type locality: “les environs de Lambaréné, sur le bas Ogooué, Gabon ”.</p> <p>Atractaspis matschiensis Werner, 1897: 404. Type locality: “ Kamerun, Hinterland ”.</p> <p>Atractaspis boulengeri mixta Laurent, 1945. Type locality: “Mayombe”, Democratic Republic of the Congo.</p> <p>Atractaspis boulengeri schmidti Laurent, 1945: 333. Type locality: route Aketi-Buta (Uele), Mumbia (Ubangi).</p> <p>Atractaspis schultzei Sternfeld, 1917: 489. Type locality: “Mbio” Democratic Republic of the Congo.</p> <p>Atractaspis boulengeri vanderborghti Laurent, 1956. Type locality: Manguretshipa, Lubero (Kivu), Democratic Republic of the Congo.</p> <p>Distribution. It extends from Cameroon to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded from Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 27C).</p> <p>Comments. The subspecies present in Equatorial Guinea is A. boulengeri matschiensis Werner, 1897 (Laurent 1950; Lasso et al. 2002).</p> <p>Specimens examined. No specimens of this species have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFC0FFF1FF4BFEBAFAA6E126	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC0FFF1FF4BFCFBFBFAE724.text	03C287E8FFC0FFF1FF4BFCFBFBFAE724.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractaspis corpulenta (Hallowell 1854)	<div><p>Atractaspis corpulenta (Hallowell, 1854)</p> <p>Figure 25A</p> <p>Brachycranion corpulentum Hallowell, 1854b: 99. Type locality: “ Liberia, W. coast of Africa” in error fide Hughes &amp; Barry (1969). Corrected to Gabon (Hallowell 1857).</p> <p>Atractaspis corpulentus (Hallowell, 1854a): Hallowell 1857: 70</p> <p>Atractaspis leucura Mocquard, 1885: 15. Type locality: “Assinie” Ivory Coast.</p> <p>Atractaspis corpulenta kivuensis Laurent, 1958: 127. Type locality: “ Rivière Bibugwa, Kivu ”.</p> <p>Distribution. It extends from Guinea in West Africa to the eastern of the Democratic Republic of the Congo. In Equatorial Guinea it had never been recorded (Map 28A).</p> <p>Comments. We provide the first record of A. corpulenta for Equatorial Guinea.</p> <p>Specimens examined. One specimen. Río Muni: Miboman, 1988 (EBD 29751).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC0FFF1FF4BFCFBFBFAE724	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC0FFF1FF4BFAFAFE83E56D.text	03C287E8FFC0FFF1FF4BFAFAFE83E56D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractaspis reticulata Sjostedt 1896	<div><p>Atractaspis reticulata Sjöstedt, 1896</p> <p>Figure 25B</p> <p>Atractaspis reticulata Sjöstedt, 1896: 516. Type locality: “ Kamerun, Ekundu”, Cameroon (in error), corrected to Okundi, Nigeria (Chirio &amp; LeBreton 2007).</p> <p>Atractaspis heterochilus Boulenger, 1901: 13. Type locality: “environs d’Albertville, sur le Tanganika”, Democratic Republic of the Congo.</p> <p>Atractaspis reticulata brieni Laurent, 1956b: 256. Type locality: Ipamu, Democratic Rpeublic of the Congo.</p> <p>Distribution. This species ranges from Nigeria to Tanzania. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 28B).</p> <p>Comments. The subspecies currently recognized in Equatorial Guinea is A. reticulata heterochilus Boulenger, 1901 (Laurent 1950; Lasso et al. 2002).</p> <p>Specimens examined. One specimen. Río Muni: Miboman, Km 27, road from Bata to Movo, 15 February 1986 (EBD 20937).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC0FFF1FF4BFAFAFE83E56D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC6FFF7FF4BFD68FCF2E49E.text	03C287E8FFC6FFF7FF4BFD68FCF2E49E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dendroaspis jamesoni (Traill 1843)	<div><p>Dendroaspis jamesoni (Traill, 1843)</p> <p>Figure 25C</p> <p>Elaps jamesoni Traill, 1843: 53. Type locality: “ Demerara, Equinoxial America” in error, corrected to West Africa fide Mertens (1938).</p> <p>Dendraspis angusticeps Duméril, 1856: 558 (non Smith). Type locality: “Envoyé du Gabon ”.</p> <p>Dendraspis welwitschii Günther, 1865: 97. Type locality: “Golungo Alto, … Angola ”.</p> <p>Dinophis fasciolatus Fischer, 1885: 111. Type locality: “Westafrika”.</p> <p>Dendraspis jamesonii (Traill, 1843): Boettger 1888: 85.</p> <p>Dendraspis neglectus Bocage, 1903: 44. Type locality: “Fernão do Pó”, Bioko.</p> <p>Dendroaspis jamesoni kaimosae Loveridge, 1936a: 64. Type locality: “Kaimosi forest, …, Kakamega district, Nyanza Province, Kenya Colony”.</p> <p>Distribution. This species extends from Togo to Sudan. In Equatorial Guinea it has been recorded in Bioko (Boulenger 1896; Bocage 1903; Drewes et al. 1999) at Riaba (=Concepción) (Mertens 1941) and Ureca (Mertens 1964a), and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 29A).</p> <p>Comments. There are two currently recognized subspecies: Dendroaspis jamesoni jamesoni, widespread across most of the species range, including Equatorial Guinea, and D. j. kaimosae, restricted to eastern region of the species range, from the eastern of Democratic Republic of the Congo to Kenya and Uganda.</p> <p>Specimens examined. Fifteen specimens. Equatorial Guinea: without specific locality, but likely Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 22869). Río Muni: Río Utoche, 10 August 1909 (MNCN 22307); Cogo, 08 February 1984 (EBD 18674); Miboman (road to Niefang) (EBD 20920); Miboman, Km 27 road from Bata to Movo, 02 June 1985 (EBD 20778); Miboman (road to Niefang), 08 July 1986 (EBD 20776 –20777); Miboman (road from Bata to Niefang) (EBD 24969); Asonga, 17 August 1986 (EBD 24970); Miboman, December 1987 (EBD 27784 –27785); San Joaquín de Ndyiacom, April 1988 (EBD 27783); San Joaquín de Ndyiacom (road from Bata to Ayamiken), 03 February 1987 (EBD 24971); Colegio de la Salle, Bata, November 1960 (EBD 2871); Acurenam, 27 August 1984 (EBD 18369).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC6FFF7FF4BFD68FCF2E49E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC6FFF7FF4BFF42FBD8E026.text	03C287E8FFC6FFF7FF4BFF42FBD8E026.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Poecilopholis cameronensis Boulenger 1903	<div><p>Poecilopholis cameronensis Boulenger, 1903</p> <p>Poecilopholis cameronensis Boulenger, 1903: 352. Type locality: “Efulen, S. Cameroon ”, Efoulan, Cameroon.</p> <p>Distribution. This species has been found in Cameroon and in mainland Equatorial Guinea. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 28C).</p> <p>Comments. Currently, this species is only known from two localities, at the type locality in Cameroon (Boulenger 1903) and at Monte Alén National Park in Equatorial Guinea (Lasso et al. 2002).</p> <p>Specimens examined. No specimens have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFC6FFF7FF4BFF42FBD8E026	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC6FFF6FF4BF8DBFB73E3FB.text	03C287E8FFC6FFF6FF4BF8DBFB73E3FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Naja (Boulengerina) annulata Buchholz & Peters 1876	<div><p>Naja (Boulengerina) annulata Buchholz &amp; Peters, 1876</p> <p>Naja annulata Buchholz &amp; Peters, 1876: 119. Type locality: “Dorfe Mbusu (Eliva Sonange am Ogowe), Westküste Afrikas” Eliva Sonange, near Ogooué River, province of Ogooué-Maritime, Gabon.</p> <p>Aspidelaps bocagei Sauvage, 1884: 204. Type locality: Gabon and Majumba.</p> <p>Boulengerina annulata (Buchholz &amp; Peters, 1876): Schmidt 1923: 123.</p> <p>Boulengerina stormsi Dollo, 1886: 160. Type locality: “région du Tanganyka”.</p> <p>Distribution. This species ranges from Cameroon to the eastern of the Democratic Republic of the Congo, south to northern Angola. In Equatorial Guinea it has been recorded for Río Muni at Benito River (Boulenger 1900) and at Monte Alén National Park (Lasso et al. 2002) (Map 29B).</p> <p>Comments. Two subspecies are currently recognized: N. annulata annulata and N. annulata stormsi (Dollo, 1886). In Equatorial Guinea occurs the nominal taxon.</p> <p>Specimens examined. No specimens of this taxon were found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFC6FFF6FF4BF8DBFB73E3FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC7FFF6FF4BFD86FAA5E4D1.text	03C287E8FFC7FFF6FF4BFD86FAA5E4D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Naja (Boulengerina) melanoleuca Hallowell 1857	<div><p>Naja (Boulengerina) melanoleuca Hallowell, 1857</p> <p>Figure 26A</p> <p>Naja haje var. melanoleuca Hallowell, 1857: 61. Type locality: “Gaboon”, Gabon.</p> <p>Naja haje var. leucosticta Fischer, 1885: 115. Type locality: “von Cameroons, vom Gaboon und von Ogowé” Cameroon and Ogooué River, Gabon.</p> <p>Naia melanoleuca (Hallowell, 1857): Boulenger 1896: 376.</p> <p>Boulengerina melanoleuca (Hallowell, 1857): Wallach et al. 2014.</p> <p>Distribution. This species ranges from southwestern Nigeria to the east of the Albertine Rift Valley, south to northern Angola. In Equatorial Guinea it has been recorded in Bioko at Musola (Boulenger 1906; Capocaccia 1961a), Malabo (Mertens 1941), Moka and Ureca (Mertens 1964a), and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002) including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 29C).</p> <p>Comments. Recent taxonomic works (Ceríaco et al. 2017; Wüster et al. 2018) have unveiled an unexpected diversity within the former concept of N. melanoleuca, which currently includes N. subfulva Laurent, 1955, N. peroescobari Ceríaco, Marques, Schmitz &amp; Bauer, 2017, N. guineensis Broadley, Trape, Chirio, Ineich &amp; Wüster, 2018, N. savannula Broadley, Trape, Chirio &amp; Wüster, 2018 and N. melanoleuca. The specimens MNCN 22598, MNCN22693 and MNCN23187 were previously misidentified as Pseudohaje nigra Günther, 1858 (a species not present in Central Africa) (Pérez del Val 2001) and correspond to N. melanoleuca.</p> <p>Specimens examined. Thirty-nine specimens. Equatorial Guinea: without specific locality, but likely Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 22693); without specific locality, July 1891 (MNCN 23187). Island of Elobey (MNCN 22705). Río Muni: Bata (MNCN 22598); Bata 09 May 1966 (EBD 2867); Bata, road of Izagu (not found), 05 January 1967 (EBD 2868–2869); Miboman (road from Bata to Niefang), August 1987 (EBD 2752); Miboman, 17 June 1984 (EBD 20278), December 1987 [EBD 27730, EBD 27731 (for 2 specimens)], December 1987 (EBD 32045); Miboman (Km 30, road from Bata to Movo) (EBD 20783); Miboman (Km 27, road of Bata), 15 February 1986 (EBD 20781); Miboman, 30 Km from Bata, 14 October 1985 (EBD 20802), 14 October 1985 (EBD 20782); Miboman, 30 km from Niefang, 01 February 1987 (EBD 24998); Miboman (road to Niefang), 07 July 1986 (EBD 20779); San Joaquín de Ndyiacom, 01 August 1986 (EBD 24997); San Joaquín de Ndyiacom (road from Bata to Ayamiken), 20 May 1987 (EBD 25000); San Joaquín de Ndyiacom, December 1987 (EBD 27735); Bata, road to the airport, 20 November 1985 (EBD 20780); San Joaquín de Ndyiacom (EBD 32044), 16 February 1988 (EBD 32042), September 1987 (EBD 27798), April 1988 (EBD 27776, EBD 27778); 15 km from Bata, 20 July 1964 (EBD 2866); Bata, 01 May 1986 (EBD 20784); without specific locality (EBD 02864 –02865); Km 25, road from Bata to Ayamiken, 03 February 1987 (EBD 32037); Km 32, road from Bata to Niefang (EBD 27794); Asonga, 14 February 1988 (EBD 32043); Camp at río Ntem, 01 August 1986 (EBD 32038). Bioko: without specific locality, 1876–1877 (MNCN 22720–22723); November 1981 (without EBD number, R1796).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC7FFF6FF4BFD86FAA5E4D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC7FFF5FF4BF8A8FF5BE3D6.text	03C287E8FFC7FFF5FF4BF8A8FF5BE3D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Naja (Boulengerina) multifasciata (Werner 1902)	<div><p>Naja (Boulengerina) multifasciata (Werner, 1902)</p> <p>Figure 26B</p> <p>Naia multifasciata Werner, 1902: 347. Type locality: “der oberen Maringa.”; upper Moringa or Tshuapa River, Democratic Republic of the Congo.</p> <p>Naja anomala Sternfeld, 1917: 482. Type locality: “Assobam-Urwald” southern Cameroon.</p> <p>Elapechis duttoni Boulenger, 1904: 15. Type locality: “near Leopoldville”, Kinshasa, Democratic Republic of the Congo.</p> <p>Paranaja multifasciata (Werner, 1902): Harding &amp; Welch 1980.</p> <p>Naja multifasciata (Werner, 1902): Wüster et al. 2007: 445.</p> <p>Boulengerina multifasciata (Werner, 1902): Wallach et al. 2014: 121</p> <p>Distribution. This species ranges from Cameroon to western Democratic Republic of the Congo. In Equatorial Guinea it had never been recorded (Map 30A).</p> <p>Comments. We provide the first record of N. multifasciata for Equatorial Guinea.</p> <p>Specimens examined. One specimen. Río Muni: Nvom (road from Aconibe to Asoc), 08 May 1987 (EBD 25064).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC7FFF5FF4BF8A8FF5BE3D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC4FFF5FF4BFDE1FC93E6F3.text	03C287E8FFC4FFF5FF4BFDE1FC93E6F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohaje goldii (Boulenger 1895)	<div><p>Pseudohaje goldii (Boulenger, 1895)</p> <p>Figure 26C</p> <p>Naia goldii Boulenger, 1895: 34. Type locality: “near Asaba”, vicinity of Asaba, Nigeria.</p> <p>Naia yakomae Mocquard, 1895 (1896): 233. Type locality: “Yakoma-Abiras”.</p> <p>Naja goldii (Boulenger, 1895): Schmidt 1923: 130.</p> <p>Pseudohaje goldii (Boulenger, 1895): Bogert 1942: 4.</p> <p>Distribution. It extends from Ivory Coast to Kenya and southward to northern Angola.In Equatorial Guinea it has been recorded in Bioko at Moka Lake (Mertens 1941), Ureca, Moka, and between Moka and San Carlos (Mertens 1964a), and in Río Muni at Benito River (Boulenger 1900) and Monte Alén National Park (Lasso et al. 2002) (Map 30B).</p> <p>Comments. Pseudohaje goldii occurs across a wide elevational gradient, being one of the reptile species that inhabits from sea level to high altitude environments such as the surroundings of Moka Lake at 1800 m above sea level (Mertens 1964a).</p> <p>Specimens examined. Four specimens. Río Muni: San Joaquín de Ndyiacom (road from Bata to Ayamiken), 03 February 1987 (EBD 25004); Miboman (road of Bata) 04 March 1986 (EBD 20805); Miboman (road to Niefang), 29 March 1986 (EBD 20803), 08 July 1986 (EBD 20806).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC4FFF5FF4BFDE1FC93E6F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC4FFF9FF4BFA76FC7FE3C7.text	03C287E8FFC4FFF9FF4BFA76FC7FE3C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon olivaceus (Dumeril 1856)	<div><p>Boaedon olivaceus (Duméril, 1856)</p> <p>Figure 27A</p> <p>Holuropholis olivaceus Duméril, 1856: 466. Type locality: “Gaboon”, Gabon.</p> <p>Boodon poensis Günther, 1888: 330. Type locality: “ Fernando Po ” (= Bioko).</p> <p>Holuropholis olivaceus (Duméril, 1856): Boettger 1889: 279.</p> <p>Boodon olivaceus (Duméril, 1856): Boulenger 1893: 335.</p> <p>Boaedon olivaceus (Duméril, 1856): Loveridge 1957: 219.</p> <p>Boaedon olivaceus stirnensis Stucki-Stirn, 1979. Type locality: northern Cameroon.</p> <p>Lamprophis olivaceus (Duméril, 1856): Broadley 1983.</p> <p>Lamprophis olivacea (Duméril, 1856): Jackson 2008: 325.</p> <p>Distribution. It is widespread across the forests from Sierra Leona and Guinea in West Africa to Uganda in East Africa. In Equatorial Guinea it has been recorded in Bioko (Günther 1888; Boulenger 1893; Bocage 1903) at Malabo (Mertens 1941), San Carlos and Ureca (Mertens 1964a), and in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 30C).</p> <p>Comments. The specimen EBD 27754 contained a mouse in its stomach.</p> <p>Specimens examined. Nineteen specimens. Equatorial Guinea: estación Cori Laranetta (locality not found) (EBD 2849). Río Muni: Bata (EBD 2850); Miboman (road to Bata, Km 29), 15 November 1986 (EBD 20833); Miboman, 06 September 1985 (EBD 20836); Miboman (road to Movó), 01 September 1984 (EBD 18368); Mongomo, 06 April 1986 (EBD 20834), 17 April 1986 (EBD 20835); San Joaquín de Ndyiacom, 1999 (EBD 31498), 25 June 1987 (EBD 27806); San Joaquín de Ndyiacom (road from Bata to Niefang), 01 August 1986 (EBD 24967); San Joaquín de Ndyiacom, 1990 (EBD 31496, EBD 31498); three kilometres from Asonga, 20 June 1984 (EBD 18356). Bioko: without specific locality (MNCN 18373, MNCN 18374); road from Moka to Malabo, close to Patio Potao, 09 November 2003 (MNCN 50493); Bioko Sur, Luba, 08 June 1984 (EBD 18712); Banapá (two specimens housed in the EBD without calatogue number, P. Basilio leg.).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC4FFF9FF4BFA76FC7FE3C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC8FFF9FF4BFDDBFC58E733.text	03C287E8FFC8FFF9FF4BFDDBFC58E733.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon virgatus (Hallowell 1854)	<div><p>Boaedon virgatus (Hallowell, 1854)</p> <p>Figure 27B</p> <p>Coelopeltis virgata Hallowell, 1854b: 98. Type locality: “ Liberia, west coast of Africa”. Malnate (1971) mentioned “Gaboon” as type locality in his revision of the type specimens housed in the Academy of Natural Sciences, Philadelphia (ANSP).</p> <p>Boodon nigrum Fischer, 1856: 9. Type locality: “St. Thomé (West Afrika)”; likely in error fide Ceríaco et al. 2021: 24.</p> <p>Boodon ventralis Günther, 1888: 329. Type locality: “Old Calabar and…Ashantee” (= Calabar, Nigerica and Ghana, respectively).</p> <p>Boodon virgatus (Hallowell, 1854): Boulenger 1893: 331.</p> <p>Boaedon virgatum (Hallowell, 1854): Menzies 1966: 172.</p> <p>Lamprophis virgatus (Hallowell, 1854): Lawson 1993.</p> <p>Boaedon virgatus (Hallowell, 1854): Rödel &amp; Mahsberg 2000: 28.</p> <p>Distribution. This species extends from Guinea to Republic of the Congo. In Equatorial Guinea it has been recorded from Bioko (Wallach et al. 2014) and Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 31A).</p> <p>Comments. The specimens EBD 27754 and EBD 20935 contained each a mouse in their stomachs.</p> <p>Specimens examined. Ten specimens. Río Muni: Mbé, 15 km southwest of Pista de Alena, 22 May 1964 (EBD 2848); Colegio de la Salle, Bata, 30 June 1964 (EBD 2860); Miboman, October 1987 (EBD 27754); Miboman (road to Niefang), 08 July 1986 (EBD 20934); Miboman (road from Bata to Niefang), 07 July 1987 (EBD 27549 -27550); Bomudi, Bata, 23 April 1986 (EBD 20935); Bata-Niefang, Km 10, 03 February 1987 (EBD 25016); Bioko Sur, Luba, 08 June 1984 (EBD 18713); San Joaquín de Ndyiacom (EBD 31505).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC8FFF9FF4BFDDBFC58E733	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFC8FFF9FF4BFA07FAB2E567.text	03C287E8FFC8FFF9FF4BFA07FAB2E567.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bothrolycus ater Gunther 1874	<div><p>Bothrolycus ater Günther, 1874</p> <p>Figure 27C</p> <p>Bothrolycus ater Günther, 1874: 444. Type locality: “ Cameroon mountains”.</p> <p>Pseudoboodon albopunctatus Andersson, 1901: 6. Type locality: “ Mapanja, Cameroon ”.</p> <p>Pseudoboodon brevicaudatus Andersson, 1901: 8. Type locality: “ Mapanja, Cameroon ”.</p> <p>Distribution. It extends across Central Africa, from Cameroon to the East of Democratic Republic of the Congo. In Equatorial Guinea it has been recorded both in Bioko (Boulenger 1919) and Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 31B).</p> <p>Comments. No precise records are known from Bioko. Bothrolycus is a monotypic genus sister to Bothrophthalmus (Zaher et al. 2019).</p> <p>Specimens examined. One specimen. Río Muni: San Joaquín de Ndyiacom, 1990 (EBD 31496).</p> </div>	https://treatment.plazi.org/id/03C287E8FFC8FFF9FF4BFA07FAB2E567	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFCDFFFCFF4BFF42FD52E667.text	03C287E8FFCDFFFCFF4BFF42FD52E667.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bothrophthalmus brunneus Gunther 1863	<div><p>Bothrophthalmus brunneus Günther, 1863</p> <p>Figure 28A</p> <p>Bothrophthalmus lineatus brunneus Günther, 1863. Type locality: “Fernando Po”. Bioko, Equatorial Guinea.</p> <p>Distribution. Cameroon, Gabon, Republic of the Congo, and Equatorial Guinea. In Equatorial Guinea it has been recorded both in Bioko (Günther 1863a; Boulenger 1893; Bocage 1903; Mertens 1964a) and Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 31C).</p> <p>Comments. There are two species of Bothropthalmus currently recognized, namely B. brunneus and B. lineatus Peters, 1863. The taxonomic status of B. brunneus is controversial. The two species are distinguishable based on colouration only: B. brunneus presents a uniformly dark body pattern, whereas B. lineatus shows a red-black striped pattern (Schmidt 1923; Chippaux &amp; Jackson 2019). Boulenger (1893) included B. brunneus in the synonymy of B. lineatus, treating Günther’s taxon as a variety of the latter. Other authors have considered this taxon as a subspecies of B. lineatus (Schmidt 1923; Pauwels et al. 2002; Chippaux 2006) or treated it with full species status (Trape 1985; Meirte 1992; Pauwels &amp; David 2008a; Wallach et al. 2014). Recent molecular works included a single sample of B. lineatus from Uganda and another one of B. brunneus from Gabon, supporting them with full species status (Kelly et al. 2011; Zaher et al. 2019). However, in order to resolve this taxonomic problem, it will be required to improve the sampling for molecular analyses across most of the range of both species, including their type localities [the type locality of B. lineatus is the Coast of the Gulf of Guinea, but fide Hughes &amp; Barry (1969) the true type locality is likely in Ghana]. There are no precise records of this species in Bioko (Mertens 1964a). The examined specimen EBD 20925 contained an unidentified mouse in its stomach.</p> <p>Specimens examined. Eight specimens. Río Muni: Miboman, 16 April 1986 (EBD 20923), 08 July 1986 (EBD 20924 –20925); Miboman, 30 km from Bata, 14 October 1986 (EBD 20922); Miboman (road from Bata to Niefang), August 1987 (EBD 27514); Ncoeakieñ (road from Bata to Movo, Km 23), 19 June 1984 (EBD 18600); San Joaquín de Nydiacom, 01 August 1986 (EBD 25132 –25133).</p> </div>	https://treatment.plazi.org/id/03C287E8FFCDFFFCFF4BFF42FD52E667	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFCDFFFCFF4BFB70FAB5E53B.text	03C287E8FFCDFFFCFF4BFB70FAB5E53B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Buhoma depressiceps (Werner 1897)	<div><p>Buhoma depressiceps (Werner, 1897)</p> <p>Tropidonotus depressiceps Werner, 1897: 402. Type locality: “ Barombi, Cameroon ”.</p> <p>Geodipsas mapanjensis Anderson, 1901: 19. Type locality: “ Mapanja, Cameroon ”.</p> <p>Geodipsas depressiceps (Werner, 1897): Werner 1913: 27.</p> <p>Geodipsas depressiceps marlieri Laurent, 1956: 129. Type locality: “ Mwana, …, Terr. de Mwenga, Kivu ”.</p> <p>Buhoma depressiceps (Werner, 1897): Ziegler et al. 1997: 108.</p> <p>Distribution. This species ranges across central Africa, from Cameroon to Uganda. In Equatorial Guinea it has been recorded both in Bioko at Musola (Boulenger 1906; Capocaccia 1961a) and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 32A).</p> <p>Comments. Some authors recognize two subspecies (Spawls et al. 2018), the nominal B. depressiceps depressiceps, which is present in Equatorial Guinea, and B. depressiceps marlieri from Democratic Republic of the Congo, whereas other authors consider them with full species status (Chippaux &amp; Jackson 2019). The phylogenetic position of the genus Buhoma is a matter of discussion (Zaher et al. 2019).</p> <p>Specimens examined. No specimens of this taxon have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFCDFFFCFF4BFB70FAB5E53B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD2FFE3FF4BFF42FADDE043.text	03C287E8FFD2FFE3FF4BFF42FADDE043.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaelycus fasciatus (Gunther 1858)	<div><p>Chamaelycus fasciatus (Günther, 1858)</p> <p>Figure 28B</p> <p>Alopecion fasciatum Günther, 1858: 196. Type locality: “ West Africa ”.</p> <p>Lycophidium fasciatum (Günther, 1858): Boulenger 1893: 342.</p> <p>Oophilositum fasciatum (Günther, 1858): Parker 1933: 545.</p> <p>Chamaelycus fasciatus (Günther, 1858): Witte 1963: 635.</p> <p>Distribution. It extends from Senegal to the East of Democratic Republic of the Congo. In Equatorial Guinea it has been reported in Bioko at Bahía de San Carlos and Musola (Boulenger 1906; Capocaccia 1961a) (Map 32B).</p> <p>Comments. We provide the first record of this species for Río Muni.</p> <p>Specimens examined. One specimen. Río Muni: San Joaquín de Ndyiacom, October 1987 (EBD 27765).</p></div> 	https://treatment.plazi.org/id/03C287E8FFD2FFE3FF4BFF42FADDE043	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD2FFE3FF4BFC94FE88E68C.text	03C287E8FFD2FFE3FF4BFC94FE88E68C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonionotophis brussauxi (Mocquard 1889)	<div><p>Gonionotophis brussauxi (Mocquard, 1889)</p> <p>Figure 28C</p> <p>Gonionotus brussauxi Mocquard, 1889: 146. Type locality: Loudinia-Niari, Niari River, Republic of Congo.</p> <p>Gonionotus vossi Boettger, 1892: 418. Type locality: “ Kamerun ”.</p> <p>Simocephalus insignis Chabanaud, 1917b: 369. Type locality: “ Congo: Ogooùé ”.</p> <p>Gonionotophis brussauxi prigoginei Laurent, 1956a: 107. Type locality: Kamituga (Kivu).</p> <p>Distribution. This species ranges from Cameroon to the Democratic Republic of the Congo. In Equatorial Guinea it has been recorded from Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 32C).</p> <p>Comments. Type species of the genus Gonionotophis.</p> <p>Specimens examined. One specimen. Río Muni: Monte Alén National Park (in the EBD collection, without EBD number).</p></div> 	https://treatment.plazi.org/id/03C287E8FFD2FFE3FF4BFC94FE88E68C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD2FFE3FF4BFAC4FBAEE585.text	03C287E8FFD2FFE3FF4BFAC4FBAEE585.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hormonotus modestus (Dumeril, Bibron, and Dumeril 1854)	<div><p>Hormonotus modestus (Duméril, Bibron, and Duméril, 1854)</p> <p>Figure 29A</p> <p>Lamprophis modestus Duméril, Bibron &amp; Duméril, 1854: 429. Type locality: “côte de Guinée ”. Restricted to Dabocrom, Ghana fide Hughes &amp; Barry (1969).</p> <p>Hormonotus audax Hallowell, 1857: 56. Type locality: “Gaboon, west coast of Africa”.</p> <p>Heterolepis glaber Jan, 1863: 98. Type locality: “ Costa d’oro … Boutry, Africa occid.”.</p> <p>Boodon vossii Fischer, 1888: 3. Type locality: “ Kamerun ”.</p> <p>Hormonotus modestus (Duméril, Bibron, and Duméril, 1854): Boulenger 1893: 343.</p> <p>Distribution. This species extends from Guinea in West Africa to Uganda in East Africa. In Equatorial Guinea it has been recorded in Bioko (Boulenger 1893) at Natividad (Hacienda Natividad) (Bocage 1895 a, 1903; Mertens 1964a) and in Río Muni at Monte Mitra within Monte Alén National Park (Gonwouo &amp; Nsang 2005) (Map 33A).</p> <p>Comments. Hormonotus is a monotypic genus with not well-known phylogenetic affinities (Zaher et al. 2019).</p> <p>Specimens examined. Five specimens. Río Muni: Miboman, 30 km from Bata, 14 October 1985 (EBD 20932); San Joaquín de Ndyiacom (road from Bata to Ayamiken), February 1987 (EBD 24956 –24957); San Joaquín de Ndyiacom, 01 August 1986 (EBD 24955); Aconibe, 07 July 1987 (EBD 27512).</p> </div>	https://treatment.plazi.org/id/03C287E8FFD2FFE3FF4BFAC4FBAEE585	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD1FFE0FF4BFE9EFAACE0D9.text	03C287E8FFD1FFE0FF4BFE9EFAACE0D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limaformosa guirali (Mocquard 1887)	<div><p>Limaformosa guirali (Mocquard, 1887)</p> <p>Figure 29B</p> <p>Heterolepis guirali Mocquard, 1887: 23. Type locality: Niger and San-Benito, West Africa [may be Benito Point, Bendel State, South of Nigeria fide Wallach et al. (2014)].</p> <p>Simocephalus baumanni Sternfeld, 1908b: 93. Type locality: “ Togo ”.</p> <p>Mehelya guirali (Mocquard, 1887): Lönnberg 1910.</p> <p>Gonionotophis guirali (Mocquard, 1887): Kelly et al. 2011.</p> <p>Limaformosa guirali (Mocquard, 1887): Broadley et al. 2018.</p> <p>Distribution. This species ranges from Sierra Leona to the Democratic Republic of the Congo. This species had never been reported in Equatorial Guinea (Map 33B).</p> <p>Comments. We provided the first records of L. guirali for Equatorial Guinea.</p> <p>Specimens examined. Three specimens. Río Muni: San Joaquín de Ndyiacom, April 1988 (EBD 27782), 1990 (EBD 31513); San Joaquín de Ndyiacom (road from Bata to Ayamiken), 20 March 1987 (EBD 25003).</p> </div>	https://treatment.plazi.org/id/03C287E8FFD1FFE0FF4BFE9EFAACE0D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD1FFE5FF4BFCE9FD42E2E7.text	03C287E8FFD1FFE5FF4BFCE9FD42E2E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lycophidion laterale Hallowell 1857	<div><p>Lycophidion laterale Hallowell, 1857</p> <p>Figure 29C</p> <p>Lycophidion laterale Hallowell, 1857: 58. Type locality: “ Gabon ”.</p> <p>Lycophidium polylepis Boulenger, 1919: 21. Type locality: “Madié (Ituri)”.</p> <p>Distribution. It extends from Liberia to the East of the Democratic Republic of the Congo. It has been included in the fauna of Bioko (Wallach et al. 2014), although there are no precise confirmed records for the island. The occurrence of this species in Río Muni is herein confirmed (Map 33C).</p> <p>Comments. We herein provide the first confirmed record of this species for Río Muni. The body colour pattern of the Biokoan specimens housed at the MNCN and the one from Río Muni at the EBD, is congruent with the phenotype shown by some Cameroonian specimens of L. laterale (see a picture in Spawls et al. 2004), suggesting that this pattern is present, at least, in the coastal areas and surroundings from Cameroon to Río Muni. This pattern consists of several distinct and asymmetric paired large orange-yellowish spots, arranged along the entire dorsal surface, which in the examined specimens are usually fused, resulting in an irregular banded body pattern. Populations from Gabon, Republic of the Congo and Angola exhibit a different dorsal pattern, with a brown background, either patternless or with very small and isolated black dots arranged along the paravertebral axes (Guibé &amp; Roux-Estève 1972; Pauwels et al. 2017; Chippaux &amp; Jackson 2019). The specimens of Lycophidion collected by M. Iradier –currently housed in the MNCN– were referred in Iradier (1887: 464) but, unfortunately, with no data on their specific collection locality. Pérez del Val (2001) listed these specimens as L. irroratum (Leach, 1819) without any discussion, and tentatively suggested Río Muni as their collection locality (“ Rio Muni?”). However, the information reviewed at the documentation department of the MNCN regarding the housed specimens of Lycophidion, indicates that these specimens were bought by the MNCN to Manuel Iradier in 1878, and that they, among others, were collected in “ Fernando Poo ” (=Bioko). An old catalogue card, handwritten by Antonio de Zulueta, also indicates Bioko as the collection site. No more specimens of this species have been recorded nor found in other examined collections. Bocage (1895a) reported the presence of another congeneric species in Bioko, Lycophidion capense, based on two specimens, one of them from “Natividad” and the second one with unknown origin. Mertens (1964) argued that Bocage’s record could be an identification mistake for other species of Lycophidion or with Chamaelycus fasciatus, as the range of L. capense is characterized by open and dry savannas, away from the tropical forests of Central Africa. There is a chance that the specimens examined by Bocage (1895a) could belong to L. laterale but considering the striking pattern of the specimens from Equatorial Guinea (see below) it seems unlikely that Bocage (1895a) did not notice the differences between the patternless species L. capense and the banded L. laterale. Additional fieldwork is required to confirm the presence of L. laterale in Bioko.</p> <p>Specimens examined. Eight specimens. Río Muni: Miboman, December 1987 (EBD 27752). Bioko: without specific locality (1876–1877) (MNCN 23102–23108).</p> </div>	https://treatment.plazi.org/id/03C287E8FFD1FFE5FF4BFCE9FD42E2E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD4FFE5FF4BFEF3FA39E633.text	03C287E8FFD4FFE5FF4BFEF3FA39E633.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mehelya poensis (Smith 1847)	<div><p>Mehelya poensis (Smith, 1847)</p> <p>Figure 30A</p> <p>Heterolepis poensis Smith, 1847: 2. Type locality: Bioko.</p> <p>Heterolepis bicarinatus Duméril, Bibron &amp; Duméril, 1854: 422. Type locality: “cóte de Guinée ”.</p> <p>Mehelya poensis (Smith, 1847): Lönnberg 1910: 3.</p> <p>Distribution. This species extends from Sierra Leona to Uganda, southward to northern Angola. In Equatorial Guinea it has been recorded in Bioko (Smith 1847; Duméril et al. 1854; Bocage 1873; Martinez y Saez 1886; Boulenger 1893) at San Carlos (Bocage 1895 a, 1903; Loveridge 1939), Ureca, Musola, between Musola and Riaba (=Concepción) (Mertens 1964a), and in the island of Elobey (Pérez del Val 2001) (Map 34A).</p> <p>Comments. We provide the first records of this species for mainland Río Muni. Duméril et al. (1854) found a specimen of Trachylepis in the stomach of this species.</p> <p>Specimens examined. Fourteen specimens. Island of Elobey (MNCN 23115). Río Muni: San Joaquín de Ndyiacom, 01 August 1986 (EBD 25001 –25002), October 1987 (EBD 27748), 1990 (EBD 31524, EBD 31491); San Joaquín de Ndyiacom (road from Bata to Ayamiken), 03 February 1987 (EBD 24951); Bata, October 1987 (EBD 27749); Miboman (road from Bata to Niefang) (EBD 24952); Miboman (Km 27, road from Bata to Niefang) October 1987 (EBD 27747); Miboman (road to Niefang), 29 March 1986 (EBD 20841); Mongomo, 16 April 1986 (EBD 20840); Efulan (Acurenam), 24 September 1985 (EBD 20942); Asonga, 08 November 1984 (EBD 18383).</p> </div>	https://treatment.plazi.org/id/03C287E8FFD4FFE5FF4BFEF3FA39E633	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD4FFE5FF4BFBCFFBD8E40D.text	03C287E8FFD4FFE5FF4BFBCFFBD8E40D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mehelya stenophthalmus (Mocquard 1887)	<div><p>Mehelya stenophthalmus (Mocquard, 1887)</p> <p>Heterolepis stenophthalmus Mocquard, 1887: 16. Type locality: “Assinie” (=Assini, SE Ghana, 5º07′N, 3º17′W) and “cap Lopez” (=Cap Lopez, W Gabon, 0º38′N, 8º42′E) (see Wallach et al. 2014).</p> <p>Simocephalus stenophthalmus (Mocquard, 1887): Boulenger 1893: 347.</p> <p>Simocephalus rostralis Sternfeld, 1910b: 63 (fide Hallermann 2007: 162). Type locality: Cameroon.</p> <p>Gonionotophis microps Boulenger, 1911b: 370 (fide Loveridge 1957: 255). Type locality: “Bitye”, southern Cameroon.</p> <p>Mehelya stenophthalmus (Mocquard, 1887): Pitman 1974.</p> <p>Gonionotophis stenophthalmus (Mocquard, 1887): Kelly et al. 2011.</p> <p>Distribution. It ranges from Guinea Bissau to Uganda. In Equatorial Guinea it has been recorded in Río Muni at Monte Alén National Park in Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 34B).</p> <p>Comments. The only record of this species is based on a single specimen found under stacked planks of a research camp (Gonwouo &amp; Nsang 2005).</p> <p>Specimens examined. No specimens have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFD4FFE5FF4BFBCFFBD8E40D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD4FFE4FF4BF8F4FD43E156.text	03C287E8FFD4FFE4FF4BF8F4FD43E156.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Python sebae (Gmelin 1789)	<div><p>Python sebae (Gmelin, 1789)</p> <p>Figure 30B</p> <p>Coluber sebae Gmelin, 1789: 1118. Type locality: “Brasiliensis, è regione Guairâ”, in error, corrected to Africa fide FitzSimons (1962).</p> <p>Coluber speciosus Bonnaterre, 1790: 17. Type locality: “ Le Brésil ”, in error.</p> <p>Boa hieroglyphica Schneider, 1801: 266. Type locality: not directly stated.</p> <p>Python houttuyni Daudin, 1803b: 254. Type locality: not directly stated.</p> <p>Python sebae (Gmelin, 1789): Duméril &amp; Bibron 1844: 400.</p> <p>Python liberiensis Hallowell, 1845: 249. Type locality: “ Liberia, West Africa ”.</p> <p>Distribution. This species is widespread from Mauritania to Somalia and south to Zimbabwe. In Equatorial Guinea it has been recorded in Bioko at Basilé (Bocage 1895a) and Riaba (=Concepción) (Mertens 1941), in Elobey (Martínez y Sáez 1886; Pérez del Val 2001), and in Río Muni at Utoche River (Pérez del Val 2001) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 34C).</p> <p>Comments. Fa &amp; García Juste (2001) reported this species as part of the bush-meat market of Río Muni in Monte Mitra Forest. The finding of the exact locality of the specimen MNCN 23811 collected from “Río Utoche” was possible thanks to the old geographic map of Río Muni provided by Gustavo Pittaluga (Pittaluga 1910).</p> <p>Specimens examined. Fifteen specimens. Equatorial Guinea: without specific locality, 1986 (EBD 31634), without date (a single egg) (EBD 18715), without date (EBD 6167). Island of Elobey, before 1886 (MNCN 22790, MNCN 22792). Río Muni: San Joaquín de Ndyiacom, 25 June 1987 (EBD 27793); Utoche river, 1909 (MNCN 23811); Colegio la Salle, Bata, 1964 (EBD 2851–2852); “Encomoison” (locality not found, but likely corresponding to Micomeseng), 03 September 1984 (EBD 20279); Road from Bata to Movo, 03 September 1984 (EBD 29670); Miboman (road to Niefang), 08 July 1986 (EBD 20919); Bata, 05 June 1984 (EBD 18362). Bioko: without specific locality (MNCN 22794); Malabo, 1983 (EBD 29671).</p> </div>	https://treatment.plazi.org/id/03C287E8FFD4FFE4FF4BF8F4FD43E156	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD5FFE4FF4BFB13FE88E47F.text	03C287E8FFD5FFE4FF4BFB13FE88E47F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myriopholis perreti (Roux-Esteve 1979)	<div><p>Myriopholis perreti (Roux-Esteve, 1979)</p> <p>Figure 30C</p> <p>Leptotyphlops perreti Roux-Esteve, 1979: 464. Type locality: “ Ebolowa, Cameroun ”.</p> <p>Myriopholis perreti (Roux-Esteve, 1979): Adalsteinsson et al. 2009: 12.</p> <p>Distribution. This species occurs from southeastern Cameroon to the Republic of the Congo, including Gabon and Equatorial Guinea, where it has been recorded in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 35A).</p> <p>Comments. A poorly known species recorded only a handful of times (Roux-Esteve 1979; Trape &amp; RouxEstève 1995; Lasso et al. 2002; Pauwels &amp; David 2008b).</p> <p>Specimens examined. One specimen. Río Muni: Monte Alén National Park (in the EBD collection, without EBD number).</p></div> 	https://treatment.plazi.org/id/03C287E8FFD5FFE4FF4BFB13FE88E47F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD5FFE9FF4BF97AFBD8E357.text	03C287E8FFD5FFE9FF4BF97AFBD8E357.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricheilostoma sundewalli (Jan 1862)	<div><p>Tricheilostoma sundewalli (Jan, 1862)</p> <p>Stenostoma (Tricheilostoma) sundewalli Jan, 1862: 191. Type locality: unknown. Designated as West Africa (Jan &amp; Sordelli 1861).</p> <p>Glauconia gestri Boulenger, 1906: 15. Type locality: “Fernando Po (Basilé and Musola, 400-700m.)… Buea, Cameroon ”.</p> <p>Leptotyphlops sundevalli (Jan, 1862): Villiers 1950: 55.</p> <p>Guinea sundewalli (Jan, 1862): Adalsteinsson et al. 2009: 11.</p> <p>Distribution. This species extends from Ghana to southwestern Central African Republic. In Equatorial Guinea it has been recorded in Bioko at Basilé and Musola (Boulenger 1906) (Map 35B).</p> <p>Comments. The specimens from Bioko (Basilé and Musola) examined by Boulenger (1906), together with an additional specimen from Cameroun (Buea), represent the syntypes of Glauconia gestri Boulenger, 1906, currently a synonym of T. sundewalli.</p> <p>Specimens examined. No specimens have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFD5FFE9FF4BF97AFBD8E357	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD8FFE9FF4BFDDBFAB5E71B.text	03C287E8FFD8FFE9FF4BFDDBFAB5E71B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afrotyphlops angolensis (Bocage 1866)	<div><p>Afrotyphlops angolensis (Bocage, 1866)</p> <p>Onychocephalus angolensis Bocage, 1866b: 46, 65. Type locality: “le districto do Duque du Bragança, situado na latitude de Loanda, porém umas 75 leguas para o interior, Portuguesas Africa occidental”, Duque de Bragança (=Calandula), Malanje District, Angola.</p> <p>Typhlops congicus Boettger, 1887: 650. Type locality: “Povo Netonna prope urbem Banana, Congo” (=Povo Netonna, near Banana, Democratic Republic of the Congo).</p> <p>Typhlops adolfi Sternfeld, 1910c: 70. Type locality: “Fort Blus.” (=Fort Beni, Kivu, Democratic Republic of the Congo).</p> <p>Typhlops dubius Chabanaud, 1917b: 364. Type locality: “ Congo belge: volcans du Kivori (altitude 1500 mètres)” (=Kivu volcanoes, Democratic Republic of the Congo).</p> <p>Typhlops lestradei Witte, 1933: 206. Type locality: “Rubengera, Ruanda ”.</p> <p>Typhlops ochraceus Laurent, 1952: 269. Type locality: “Mulenge, Terr. D’Uvira, Kivu”.</p> <p>Typhlops angolensis irsaci Laurent, 1956: 57, 349. Type locality: “Lwiro, Terr. de Kabare, Kivu, Afrique centrale”.</p> <p>Typhlops angolensis polylepis Laurent, 1956: 56. Type locality: “Kiandjo, Terr. de Mwenga, Kivu, Afrique centrale”.</p> <p>Typhlops angolensis symoensi Laurent, 1960: 10. Type locality: “Rivière Ngovi, Territoire d’Uvira, Kivu, Congo oriental”.</p> <p>Rhinotyphlops angolensis (Bocage, 1866): Broadley &amp; Wallach 2000: 167</p> <p>Afrotyphlops angolensis (Bocage, 1866): Broadley &amp; Wallach 2009: 29.</p> <p>Distribution. It extends from Cameroon to western Kenya, and southward to Angola and Zambia. In Equatorial Guinea it has been recorded both in Bioko (Roux-Estève 1974) and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 35C).</p> <p>Comments. A comprehensive revision of this species was provided by Roux-Estève (1974).</p> <p>Specimens examined. No specimens of this taxon have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFD8FFE9FF4BFDDBFAB5E71B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD8FFE8FF4BFAE7FB84E623.text	03C287E8FFD8FFE8FF4BFAE7FB84E623.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afrotyphlops congestus (Dumeril & Bibron 1844)	<div><p>Afrotyphlops congestus (Duméril &amp; Bibron, 1844)</p> <p>Figure 31A–B</p> <p>Onychocephalus congestus Duméril &amp; Bibron, 1844: 334. Type locality: unknown.</p> <p>Onychophis barrowii Gray, 1845: 133. Type locality: “ India?” (in error fide Hahn 1980: 68).</p> <p>Typhlops crassatus Peters, 1881c: 50. Type locality: “Chinchoxo, durch die africanische Gesellschaft” (= Cabinda, Angola).</p> <p>Rhinotyphlops congestus (Duméril &amp; Bibron, 1844): Broadley &amp; Wallach 2000: 167.</p> <p>Afrotyphlops congestus (Duméril &amp; Bibron, 1844): Broadley &amp; Wallach 2009: 46.</p> <p>Distribution. It extends from southeastern Nigeria to the West of Uganda, southward to the Republic of the Congo. In Equatorial Guinea it has been recorded in Bioko at Basilé (Bocage 1895a; Bocage 1903; Roux-Estève 1974) and Moka (Mertens 1964a), and in Río Muni at Monte Alén National Park (Lasso et al. 2002) (Map 36A).</p> <p>Comments. Afrotyphlops congestus was formerly considered by some authors as a subspecies of A. punctatus (Leach, 1819) (Werner 1913; Roux-Estève 1974; see Broadley &amp; Wallach 2009). Some molecular works that included samples of both taxa, supported their independent species status (Vidal et al. 2010; Hedges et al. 2014; Pyron &amp; Wallach 2014; Nagy et al. 2015). In Wallach et al ’s. (2014) catalogue, A. punctatus is included in the fauna of Bioko, maybe as a result of a mistake coming from the previous status of A. congestus as a subspecies of A. punctatus. Taking into account (1) the high morphological similarity shown by A. punctatus and A. congestus, and (2) the ecology of these two species, in which A. punctatus usually occurs in savannah environments (Trape &amp; Baldé 2014) whereas A. congestus exclusively inhabits forests, we follow Roux-Estève (1974) in considering all the populations from Equatorial Guinea (both Bioko and Río Muni) as A. congestus, excluding A. punctatus from the reptile list of the country. Afrotyphlops congestus, like some other related species [e.g. A. punctatus, A. liberiensis (Hallowell, 1848)], presents two phenotypes, a “marbled” form and a “striped” form, the first one being more common in A. congestus (Roux-Estève 1974). Mertens (1964) reported the striped phenotype of A. congestus in Moka, Bioko. We found both phenotypes among the examined specimens either from Río Muni or Bioko: MNCN23354, MNCN23355, MNCN23357, EBD 20801, 27520, 24963 depict a marbled pattern, whereas MNCN23356 and EBD 2843 show a striped pattern. The pholidosis traits of the examined specimens fall within the variability of the species [midbody scale rows: 28 (MNCN23356), 31 (MNCN23357), and 30 (MNCN23354, MNCN23355); number of dorsal longitudinal scales: 332 (MNCN23356), 333 (MNCN23357), 383 (MNCN23355), 394 (MNCN23354)]. The examined specimens from Río Muni present a greater number of dorsal longitudinal scales (383–394) than the Biokoan examined specimens (332–333).</p> <p>There is a taxonomic question dealing with the striped phenotype, pending resolution until more data are available. Some putative undescribed species of Afrotyphlops are shown by Chirio &amp; LeBreton (2007). For example, the one named “ Rhinotyphlops sp.3 ”, is morphologically similar to some striped specimens of A. congestus. These authors noted the same pholidotic pattern in their undescribed form “ Rhinotyphlops sp.3 ” and in A. congestus. A molecular phylogeography including both the “marbled” and “striped” phenotypes of A. congestus, together with samples of specimens assignable to the putative new taxa depicted by Chirio &amp; LeBreton (2007) will be required in order to resolve this taxonomic conundrum. With the data at hand, we tentatively consider all the examined striped specimens as A. congestus.</p> <p>Specimens examined. Eight specimens. Río Muni: without specific locality, 1885 (MNCN 23354–23355). Nvom (road from Aconibe to Asoc), 08April 1987 (EBD 24963). Miboman (road from Movo to Niefang), September 1987 (EBD 27520). Miboman (Km 27, road from Bata to Movo), 02 June 1986 (EBD 20801). Enuc, 12 April 1965 (EBD 2843). Bioko: without specific locality, 1876–1877 (MNCN 23356–23357).</p> </div>	https://treatment.plazi.org/id/03C287E8FFD8FFE8FF4BFAE7FB84E623	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFD9FFEDFF4BFB37FAB5E373.text	03C287E8FFD9FFEDFF4BFB37FAB5E373.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Indotyphlops braminus (Daudin 1803)	<div><p>Indotyphlops braminus (Daudin, 1803)</p> <p>Eryx braminus Daudin, 1803c: 279. Type locality: Vizagapatam (Visakhapatnam), India.</p> <p>Tortrix russelii Merrem, 1820: 84. Type locality: “in India orientali”.</p> <p>Typhlops braminus (Daudin, 1803): Fitzinger 1826: 53.</p> <p>Tortrix braminus (Daudin, 1803): Schlegel 1839 in 1837–1844: 39.</p> <p>Argyrophis truncatus Gray, 1845: 138. Type locality: “ Philippines ”.</p> <p>Onychocephalus capensis Smith, 1846 in 1838–1849: 111 (unnumbered), pl. 51, fig. 3, pl. 54, figs 9–16. Type locality: “interior of Southern Africa”.</p> <p>Argyrophis braminus (Daudin, 1803): Blyth 1856: 712.</p> <p>Ophthalmidium tenue Hallowell, 1861: 497. Type locality: “ Hong-Kong, China ”.</p> <p>Typhlops (Typhlops) inconspicuus Jan, 1863: 11. Type locality: “ Madagascar ”.</p> <p>Typhlops (Typhlops) accedens Jan, 1863: 12. Type locality: unknown; East Indies fide Hahn 1980: 39.</p> <p>Typhlops (Typhlops) euproctus Boettger, 1882: 479. Type locality: “Lukubé, insula Nossi-Bé, Madagascariensis” (= Lokobe, Nosy Be Island, W Antsiranana Province, N Madagascar).</p> <p>Typhlops braminus arenicola Annandale, 1906: 192. Type locality: not stated; “Rámanád, Tamil Nadu State, southern India ” fide Das &amp; Gayen 2004: 97.</p> <p>Typhlops limbrickii Annandale, 1906: 193, pl. 9, figs. 3–3a. Type locality: “ Rámanád, (Tamil Nadu State), southern India.”</p> <p>Typhlops braminus pallidus Wall, 1909: 609. Type locality: “ Dibrugarh, south bank of Brahmaputra [River], Upper Assam, India ”.</p> <p>Glauconia braueri Sternfeld, 1910c: 69. Type locality: “ Bagamoyo, Deutsch-Ostafrika” (= Tanzania).</p> <p>Typhlops fletcheri Wall, 1919: 556, pl. 1 (upper right). Type locality: “ Nilgiri Hills, India ”.</p> <p>Typhlops khoratensis Taylor, 1962: 248, fig. 13A. Type locality: “ Muak Lek, Friendship Highway, Sara Buri Province, Thailand ”.</p> <p>Typhlops pseudosaurus Dryden &amp; Taylor, 1969: 270. Type locality: “ Harmon Village, Guam, Mariana Islands”.</p> <p>Indotyphlops braminus (Daudin, 1803): Hedges et al. 2014: 6.</p> <p>Virgotyphlops braminus (Daudin, 1803): Wallach 2020: 10.</p> <p>Distribution. Likely native of India or Sri Lanka; it is spread on all continents through human introductions. In Equatorial Guinea it has been recorded from Annobon at San Antonio outskirts (Map 36B).</p> <p>Comments. Recorded in Annobon by Jesus et al. (2003) at the surroundings of San Antonio, under rocks. It is a species complex (Sidharthan et al. 2022).</p> <p>Specimens examined. No specimens of this taxon have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFD9FFEDFF4BFB37FAB5E373	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFDCFFEDFF4BFAB6FCF6E4D7.text	03C287E8FFDCFFEDFF4BFAB6FCF6E4D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Causus lichtensteinii (Jan 1859)	<div><p>Causus lichtensteinii (Jan, 1859)</p> <p>Figure 32A</p> <p>Aspidelaps lichtensteinii Jan, 1859: 511. Type locality: “Côte-d’Or.” presumably Ghana (Wallach et al. 2014).</p> <p>Dinodipsas angulifera Peters, 1882b: 894. Type locality: “America australis”.</p> <p>Causus lichtensteini (Jan, 1859): Duméril 1859: 217.</p> <p>Distribution. This species extends from Guinea to Kenya and southward to Angola. In Equatorial Guinea it has been reported in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002) (Map 37A).</p> <p>Comments. Natural history aspects of this species were discussed by Ineich et al. (2006).</p> <p>Specimens examined. One specimen. Río Muni: without specific locality, but likely Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 46687).</p></div> 	https://treatment.plazi.org/id/03C287E8FFDCFFEDFF4BFAB6FCF6E4D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFDCFFEDFF4BFE47FE88E18B.text	03C287E8FFDCFFEDFF4BFE47FE88E18B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Letheobia caeca (Dumeril 1856)	<div><p>Letheobia caeca (Duméril, 1856)</p> <p>Figure 31C</p> <p>Onychocephalus caecus Duméril, 1856: 462. Type locality: “ Gabon ”.</p> <p>Letheobia caeca (Duméril, 1856): Cope 1868: 322.</p> <p>Rhinotyphlops caecus (Duméril, 1856): Roux-Estève 1974: 210.</p> <p>Distribution. It occurs from Nigeria and Cameroon to Gabon and the Republic of the Congo. In Equatorial Guinea it has been recorded in Bioko (Roux-Estève 1974) and in Río Muni (North-west), likely at Bata (Roux-Estève 1974), and at Monte Alén National Park as L. wittei (Lasso et al. 2002) (Map 36C).</p> <p>Comments. The specimen identified as L. wittei and recorded by Lasso et al. (2002), currently housed in the EBD collection, was re-identified as L. caeca.</p> <p>Specimens examined. One specimen. Río Muni: Monte Alén National Park (in the EBD collection, without EBD number).</p></div> 	https://treatment.plazi.org/id/03C287E8FFDCFFEDFF4BFE47FE88E18B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFDCFFECFF4BF8AAFBD8E373.text	03C287E8FFDCFFECFF4BF8AAFBD8E373.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Causus maculatus (Hallowell 1842)	<div><p>Causus maculatus (Hallowell, 1842)</p> <p>Distichurus maculatus Hallowell, 1842: 337, pl. 19. Type locality: “ Liberia, Western Africa ”.</p> <p>Causus maculatus (Hallowel, 1842): Bocage 1895c: 145.</p> <p>Distribution. Widespread from Mauritania to Angola, eastward to Ethiopia. In Equatorial Guinea it has been recorded from Bioko as C. rhombeatus (Lichtenstein, 1823) (Boulenger 1896; Bocage 1903) and from Río Muni at Monte Mitra in Monte Alén National Park (Gonwouo &amp; Nsang 2005) (Map 37B).</p> <p>Comments. There are no precise records of this species for Bioko (Mertens 1964a). Details of its natural history were provided by Ineich et al. (2006).</p> <p>Specimens examined. No specimens have been found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFDCFFECFF4BF8AAFBD8E373	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFDDFFECFF4BFDBFFB05E1A8.text	03C287E8FFDDFFECFF4BFDBFFB05E1A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheris hetfieldi Ceriaco, Marques, Bauer 2020	<div><p>Atheris hetfieldi Ceríaco, Marques, Bauer, 2020</p> <p>Atheris hetfieldi Ceríaco, Marques, Bauer, 2020. Type locality: “Moka road (3.38844º, 8.65878º, 1296 m), Bioko Island, Equatorial Guinea ”.</p> <p>Distribution. Endemic to Bioko.</p> <p>Comments. This species is currently known from two female specimens. The first one was reported from Moka (3.35081º, 8.66254º) by Capocaccia (1961a) as Atheris squamigera (Hallowell, 1854), but highlighting its singular morphological traits; this specimen represents the species’ paratype. The second one was recently reported and designated as the holotype of A. hetfieldi (Ceríaco et al. 2020a) (Map 37C).</p> <p>Specimens examined. No specimen of this taxon was found in the examined collections.</p></div> 	https://treatment.plazi.org/id/03C287E8FFDDFFECFF4BFDBFFB05E1A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFDDFFECFF4BFC76FB10E4AF.text	03C287E8FFDDFFECFF4BFC76FB10E4AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheris squamigera (Hallowell 1854)	<div><p>Atheris squamigera (Hallowell, 1854)</p> <p>Figure 32B</p> <p>Echis squamigera Hallowell, 1854a: 193. Type locality: “ Near the river Gaboon, Guinea”, Gabon River, Gabon.</p> <p>Poecilostolus burtonii Günther, 1863b: 20. Type locality: “ West Africa … Camaroon country”.</p> <p>Atheris lucani Rochebrune, 1885: 89. Type locality: “Locum Landana dictum” (untraceable locality).</p> <p>Atheris squamigera robusta Laurent, 1956. Type locality: “Nioka”, Congo Belge (Democratic Republic of the Congo).</p> <p>Distribution. This species extends from Nigeria to Kenya and southward to northern Angola. In Equatorial Guinea it has been reported from Bioko (Broadley 1998b; Drewes et al. 1999) at Moka (Boulenger 1906; Capocaccia 1961a; Mertens 1964a) and between Moka and Malabo (Mertens 1964a; Ceríaco et al. 2020a), and in Río Muni at Cabo San Juan (Boulenger 1905) and Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005). There is a record from iNaturalist close to Moka, Bioko (03°19′55.07″N, 08°38′39.04″E) (Map 38A).</p> <p>Comments. Based on cytochrome b gene sequences, Atheris squamigera is closely related to A. subocularis Fischer, 1888 (Lawson et al. 2001), and the two species form a clade sister to A. broadleyi Lawson, 1999 (Lawson et al. 2001). Phylogenetic relationships between these species and A. hetfieldi remain unknow.</p> <p>Specimens examined. Three specimens. Bioko: red swamp in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.514475&amp;materialsCitation.latitude=3.357775" title="Search Plazi for locations around (long 8.514475/lat 3.357775)">Caldera de Luba</a>, 03°21′27.99″N, 08°30′52.11″E, 27 November 2003 (MNCN 50275). Río Muni: without specific locality, but likely Cabo San Juan as part of the material examined by Boulenger (1905) (MNCN 22788); Ayamiken, July 1964 (EBD 2875).</p> </div>	https://treatment.plazi.org/id/03C287E8FFDDFFECFF4BFC76FB10E4AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FFDDFF17FF4BF8AAFAA9E053.text	03C287E8FFDDFF17FF4BF8AAFAA9E053.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bitis gabonica (Dumeril, Bibron & Dumeril 1854)	<div><p>Bitis gabonica (Duméril, Bibron &amp; Duméril, 1854)</p> <p>Figure 32C</p> <p>Echidna gabonica Duméril, Bibron &amp; Duméril, 1854: 1428. Type locality: “ Gabon ” by lectotype designation (Hughes &amp; Barry 1969: 1030).</p> <p>Bitis gabonica (Duméril, Bibron &amp; Duméril, 1854): Boulenger 1915: 222.</p> <p>Distribution. This species is distributed from southeastern Nigeria to western Kenya, southward to Angola and northern South Africa. In Equatorial Guinea it has been recorded from Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 38B).</p> <p>Comments. Fa &amp; García Juste (2001) reported this species in the bush-meat market of Río Muni at Monte Mitra Forest. There are some known sympatric populations of B. gabonica and B. nasicornis in Nigeria (Luiselli &amp; Akani 2003).</p> <p>Specimens examined. Eighteen specimens. Río Muni: San Joaquín de Ndyiacom, April 1988 (EBD 27745), December 1987 (EBD 27780); Monte Raíces, 10 October 1965 (EBD 2872); Miboman, 15 June 1984 (EBD 20269), 01 September 1984 (EBD 20260), 22 June 1984 (EBD 20270 –20271); Miboman, 30 km from Bata, 14 October 1985 (EBD 20772 –20774), 19 October 1985 (EBD 20767); Miboman, close to the town, 27 km from Movo, 06 September 1985 (EBD 20761); Mikan (road from Bata to Niefang), 21 September 1987 (EBD 27791); Road from Niefang to Miboman, 20 July 1986 (EBD 20771), 06 September 1985 (EBD 20769 –20770); Bata (Km 3, road to Mbimi), 15 October 1985 (EBD 20775); Monte de Bata (road to Niefang), 02 June 1985 (EBD 20768).</p> </div>	https://treatment.plazi.org/id/03C287E8FFDDFF17FF4BF8AAFAA9E053	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
03C287E8FF26FF16FF4BFD2EFE24E3BB.text	03C287E8FF26FF16FF4BFD2EFE24E3BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bitis nasicornis (Shaw & Nodder 1792)	<div><p>Bitis nasicornis (Shaw &amp; Nodder, 1792)</p> <p>Figure 33</p> <p>Coluber nasicornis Shaw &amp; Nodder, 1792: pl. 94, 3 pp. Type locality: “interior parts of Africa”. Restricted to coast of Guinea by Reinhardt (1843) and Ghana fide Hughes &amp; Barry (1969).</p> <p>Vipera hexacera Duméril, 1854: 1416. Type locality: Liberia dans l’Afrique occidentale (Guinée).</p> <p>Bitis nasicornis (Shaw &amp; Nodder, 1792): Boulenger 1915: 222.</p> <p>Distribution. This species ranges over the forests from Guinea in West Africa to Rwanda in East Africa and southward to the Republic of the Congo and Zambia. In Equatorial Guinea it has been recorded in Bioko (Boulenger 1896; Bocage 1903; Drewes et al. 1999) at Natividad (Hacienda Natividad) (Bocage 1895a), Moka (Boulenger 1906; Mertens 1941, 1964; Capocaccia 1961a), Bahía de San Carlos (Capocaccia 1961a), and Musola (Mertens 1941; Capocaccia 1961a), and in Río Muni at Monte Alén National Park (Lasso et al. 2002), including Monte Mitra (Gonwouo &amp; Nsang 2005) (Map 38C).</p> <p>Comments. As in the case of B. gabonica, in Equatorial Guinea this species is hunted and consumed by local people (Gonwouo &amp; Nsang 2005).</p> <p>Specimens examined. Twenty-seven specimens. Río Muni: Bata, 28 August 1983 (EBD 20763); “ Masasinga ”, Río Muni (locality not found) (MNCN 21684); Miboman, 03 September 1984 (EBD 20166), 18 June 1984 (EBD 20267), 01 September 1984 (EBD 20262, EBD 20264), 06 September 1985 (EBD 20761), December 1987 (EBD 27786), 01 September 1984 (EBD 20263 –20265), 29 July 1985 (EBD 20760), 09 April 1986 (EBD 20765); Miboman, 30 km from Bata, 19 October 1985 (EBD 20766); Mikan, road from Bata to Niefang, 21 November 1987 (EBD 2783); Moca, 24 July 1984 (EBD 20268); San Joaquín de Ndyiacom, 03 February 1987 (EBD 25122); San Joaquín de Ndyiacom (road from Bata to Ayamiken), September 1987 (EBD 27801 –27802); San Joaquín de Ndyiacom, 25 June 1987 (EBD 27804 –27805), September 1987 (EBD 27795); Playa Asonga, Bata, 25 August 1984 (EBD 18357); no specific locality (EBD 25177). Bioko: without specific locality (MNCN 22777–22778); Malabo, 1933 (MNCN 22779).</p> </div>	https://treatment.plazi.org/id/03C287E8FF26FF16FF4BFD2EFE24E3BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sánchez-Vialas, Alberto;Calvo-Revuelta, Marta;Riva, Ignacio De La	Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta, Riva, Ignacio De La (2022): Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197, DOI: https://doi.org/10.11646/zootaxa.5202.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5202.1.1
