taxonID	type	description	language	source
399B4B5BCC0356439F520B7C70952E91.taxon	materials_examined	Material examined. In addition to the holotype, we examined one female collected at the end of July 1900 during O. Neumann's and C. von Erlanger's expedition in " Hochebene Didda " (ZMB- 26913), which is one of the two syntypes of Leptopelis montanus Tiutenko & Zinenko, 2021 (= Pseudocassina ocellata), synonymised here with L. rugosus. As this specimen was mentioned as " the holotype " by Largen (2001) and has a collection date and a more precise locality than the other syntype (ZMB- 26914), which only states " Somaliand ", considered erroneous by (Largen 1977), we formally designate ZMB- 26913 as the lectotype of Pseudocassina ocellata here (see remarks below regarding the locality and taxonomic status). We also examined one female (16 - 130), collected on 12 July 2016 by J. Reyes-Velasco and S. Boissinot east of Mehal Meda (10.3316 ° N, 39.7812 ° E, 3265 m a. s. l.), one male (16 - 103), collected on 12 July 2016 by J. Reyes-Velasco and S. Boissinot north of Debre Sina (9.9894 ° N, 39.7452 ° E, 3017 m a. s. l.), three males (16 - 109, 16 - 116, 16 - 129), collected on 12 July 2016 by J. Reyes-Velasco and S. Boissinot east of Mehal Meda (10.16 - 10.33 ° N, 39.76 - 39.80 ° E, 3167 - 3265 m a. s. l.), five males (16 - 150, 16 - 151, 16 - 152, 16 - 153, 16 - 154), collected on 13 July 2016 by J. Reyes-Velasco and S. Boissinot east of Debre Birhan (9.6979 ° N, 39.5628 ° E, 2833 m a. s. l.), three males (16 - 164, 16 - 168, 16 - 172), collected on 14 July 2016 by J. Reyes-Velasco and S. Boissinot south of Fiche (9.73 - 9.75 ° N, 38.74 ° E, 2657 - 2726 m a. s. l.), five males (SB 530, SB 531, SB 532, SB 533, SB 540), collected on 1 and 2 July 2018 by S. Goutte and Y. Bourgeois east of Debre Birhan (9.6979 ° N, 39.5628 ° E, 2339 m a. s. l.), four males (SB 541, SB 542, SB 544, SB 545), collected on 2 July 2018 by S. Goutte and Y. Bourgeois between Debre Birhan and Ankober (9.6820 ° N, 39.7390 ° E, 3408 m a. s. l.), two females (SB 555, SB 558) and one male (SB 556), collected on 4 July 2018 by S. Goutte and Y. Bourgeois east of Mehal Meda (10.31 - 10.33 ° N, 39.78 - 39.80 ° E, 3337 - 3429 m a. s. l.) and three males (SB 608, SB 609, SB 610), collected on 11 July 2018 by S. Goutte and Y. Bourgeois south of Fiche (9.7307 ° N, 38.7439 ° E, 2365 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
399B4B5BCC0356439F520B7C70952E91.taxon	diagnosis	Diagnosis. A large (male (n = 22) SVL 38.7 + / - 2.7 mm, female (n = 2) SVL 52.4 + / - 0.2 mm) species of the Leptopelis gramineus species complex (Figs 5, 6). Robust, semi-fossorial form. It differs from other members of the Leptopelis gramineus species complex by the following combination of characters: (1) large tympanum (male TD / ED 0.57 + / - 0.11, female TD / ED 0.54), (2) long snout (male SL / HL 0.25 + / - 0.02, female SL / HL 0.27 + / - 0.01), (3) well-developed metatarsal tubercle (male MTL / FL 0.17 + / - 0.02, female MTL / FL 0.18 + / - 0.02), (4) ventrum lacking any brown spots, (5) yellow colouration on the side of the ventrum and the inner thighs almost always present.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
399B4B5BCC0356439F520B7C70952E91.taxon	description	Description of the holotype. Relatively large adult female (SVL 44.7 mm) in good condition of preservation (Fig. 5). Body robust and round. Head a third of body size in length, wider than long (HW / HL 1.21). Canthus rostralis obtuse and snout rounded and wide (IND / IOD 0.96). Nostril half-way between the tip of the snout and the eye (NS / SL 0.53). Tympanum partially hidden by flank skin rugosities and barely visible. Hind-limbs relatively long (TL / SVL 0.38 and THL / SVL 0.43). Finger and toe discs barely expanded, but distinct, ovoid. Finger formula: I <II <IV <III. Hand free of webbing. Foot longer than tibia (FL / TL 1.3). Inner metatarsal tubercle well-developed, oval in shape, 0.20 x foot length. Outer metatarsal tubercle absent. Toe formula: I <II <V <III <IV. Toe webbing formula (toe internal / external sides, number of phalanges webbed): Ie (1), IIi / e (1 - 1), IIIi / e (1 - 2), IVi / e (2 - 2), Vi (2). Skin of the dorsum, flanks and ventrum highly rugose.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
07AF3730BB605C0893F9A911DBAF0383.taxon	diagnosis	Diagnosis. Medium to large (male (n = 21) SVL 35.9 + / - 3.5 mm, female (n = 5) SVL 53.4 + / - 5.3 mm), robust semi-fossorial species of the Leptopelis gramineus species complex (Fig. 7). It differs from other members of the Leptopelis gramineus species complex by the following combination of characters: (1) short and robust hind-limbs (male TL / SVL 0.33 + / - 0.03, female TL / SVL 0.30 + / - 0.02), (2) well-developed metatarsal tubercle (male MTL / FL 0.17 + / - 0.03, female MTL / FL 0.16 + / - 0.02), (3) longer snout (male SL / HL 0.23 + / - 0.03, female SL / HL 0.24 + / - 0.02), (4) toe and fingertips not enlarged and (5) ventrum often with dark brown spots and / or yellowish sides.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
07AF3730BB605C0893F9A911DBAF0383.taxon	description	Description of the holotype. Medium size adult male (SVL 40.6 mm). Body robust (Fig. 7). Head a third of body size in length, wider than long (HW / HL 1.14). Snout wide (IND / IOD 0.87). Nostrils closer to the tip of the snout than the eyes (NS / SL 0.47). Canthus rostralis well-marked, but obtuse and loreal region slightly concave. Pupil vertical. Tympanum partially hidden on the posterior-dorsal edge, 0.40 x eye diameter. Fingers and toes robust with ovoid discs not expanded, but distinct. Finger formula: I <II <IV <III. Hand free of webbing. Hind-limbs short and robust (TL / SVL 0.31 and THL / SVL 0.38). Foot 1.32 x tibia length. Inner metatarsal tubercle present, oval in shape, 0.16 x foot length. Outer metatarsal tubercle absent. Toe formula: I <II <III <V <IV. Foot webbing minimal, except between toe III and toe IV where webbing extends to half-way between the first and the second phalanges. Skin of the dorsum, flanks and ventrum slightly rugose.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
07AF3730BB605C0893F9A911DBAF0383.taxon	etymology	Etymology. The specific name refers to the Shebelle River, as the distribution range of the species appears restricted to the Shebelle River Basin, with populations found both north and south of the source of the river (Fig. 1).	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
45DB2E04856C5F1B9153298EE20770DE.taxon	materials_examined	Other material examined. One female (SB 197) and 12 males (SB 186 - SB 196, SB 206), collected on 19 April 2018 by S. Goutte and J. Reyes-Velasco southeast of Kofele (7.0226 ° N, 38.8701 ° E, 2,561 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
45DB2E04856C5F1B9153298EE20770DE.taxon	diagnosis	Diagnosis. Small to medium-sized (male (n = 20) SVL 27.6 + / - 2.0 mm, female (n = 2) SVL 43.5 + / - 5.7 mm), robust arboreal species of the Leptopelis gramineus species complex (Fig. 8). It differs from other members of the Leptopelis gramineus species complex by the following combination of characters: (1) thin, elongated hind-limbs (male TL / SVL 0.36 + / - 0.02, female TL / SVL 0.37 + / - 0.03), (2) small metatarsal tubercle (male MTL / SVL 0.07 + / - 0.01, female MTL / SVL 0.06 + / - 0.00) (2) inter-orbital distance very short (male IOD / ED 0.88 + / - 0.16, female IOD / ED 0.69 + / - 0.12), (3) dorsal skin always smooth, except in females where it may be slightly rugose, (4) absence of yellow colouration on the ventrum or inner thighs.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
45DB2E04856C5F1B9153298EE20770DE.taxon	description	Description of the holotype. Small-sized male (SVL 28.9 mm) adult (Fig. 8). Body robust. Head a third of body size in length, wider than long (HW / HL 1.16). Snout angular and narrow (IND / IOD 0.75). Nostril beyond half-way between the tip of the snout and the eye (NS / SL 0.56). Canthus rostralis well-marked and loreal region slightly concave. Pupil vertical. Tympanum clearly visible, and round, 0.71 x eye diameter. Fingers and toes thin and elongated with discs barely expanded, but distinct, ovoid. Finger formula: I <II <IV <III. Hand free of webbing. Hind-limbs relatively long for the L. gramineus species complex (TL / SVL 0.38 and THL / SVL 0.42 vs. TL / SVL <0.37 and THL / SVL <0.40 in other species of the L. gramineus complex, except L. susanae). Foot 1.2 x tibia length. Inner metatarsal tubercle present, oval in shape, 0.21 x foot length. Outer metatarsal tubercle absent. Toe formula: I <II <V <III <IV. Foot webbing minimal. Skin of the dorsum and flanks smooth, ventrum rugose.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
45DB2E04856C5F1B9153298EE20770DE.taxon	etymology	Etymology. Leptopelis xeniae sp. nov. is named after Xenia Freilich, who conducted her doctoral research on Ethiopian anurans, including the Leptopelis gramineus complex.	en	Goutte, Sandra, Reyes-Velasco, Jacobo, Kassie, Abeje, Boissinot, Stephane (2022): Genetic and morphometric analyses of historical type specimens clarify the taxonomy of the Ethiopian Leptopelis gramineus species complex (Anura, Arthroleptidae). ZooKeys 1128: 63-97, DOI: http://dx.doi.org/10.3897/zookeys.1128.82176, URL: http://dx.doi.org/10.3897/zookeys.1128.82176
