taxonID	type	description	language	source
03C087917C15FF98F8EEFDE0FC6FFE63.taxon	description	(Figs 1, 4 a – c)	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C15FF98F8EEFDE0FC6FFE63.taxon	materials_examined	Type material. Holotype: male (IEBR), VIETNAM: Son La Province, Thuan Chau District, Copia Nature Reserve, light trap, 28. iv. 2016, Hoang Vu Tru leg. Paratypes: 1 male (IEBR), the same data as holotype; 2 females (IEBR), the same locality as holotype, light trap, 01. v. 2016, Hoang Vu Tru leg; 1 female (IEBR), the same locality as holotype, UV light trap, 15. v. 2017, Pham Van Phu leg; 1 female (NMBS), LAOS NE: Houa Phan prov., 20 ° 13 ' 09 N – 103 ° 59 ' 53 - 140 ° 00 ' 03 E, 1480 - 1550 m, Phou Pane Mt, 09 - 16. VI. 2009, David Hauck leg, NHMB Basel NMPC Prague Laos 2009 expedition M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň; 3 males (NMPC), LAO-NE, Hua Phan prov., 20 ° 12 ' N 104 ° 01 ' E, Phu Phan Mt, 1500 - 1900 m, 17. V, Vit Kubáň, - 3. VI. 2017; 14 males, 9 females (NMPC), Laos NE, Houa Phan prov., 20 ° 13 ' 09 N – 103 ° 59 ' 54 '' - 140 ° 00 ' 03 '' E, 1480 - 1510 m, Phou Pane Mt, 22. IV- 14. V. 2008, Vit Kubáň leg.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C15FF98F8EEFDE0FC6FFE63.taxon	description	Description of the holotype (male). Body length: 16.5 mm (from anterior margin of clypeus to apex of elytra); body width: 8.6 mm (width across humeri). Color dark reddish brown on head, pronotum, scutellum; reddish brown to dark reddish brown on elytra, ventral surface, and legs; moderately shiny; body densely puncate, covered with hair-like setae (Fig. 1 a). Head. Surface densely punctate; punctures moderately large, each bearing a semierect seta; clypeus transverse, with apex moderately bilobed and strongly reflexed, slightly narrowing at base, length five times greater than width (Fig. 1 b). Clypeofrontal suture clearly defined, weakly sinuated. Frontal carina clearly defined, slightly irregular, reaching periocular margins. Basal area of head with small punctures, sparser punctate than forehead (Fig. 1 b). Antenna with 9 antennomeres, antennal club composed of 3 antennomeres, club slightly longer than antennomeres 2 – 6 combined (Fig. 1 e). Last segment of maxillary palp elongated, moderately flattened. Pronotum. Transverse; widest at basal third; pronotum with anterior margin moderately concave at lateral ends, basolateral angles obtuse and round; anterior half of lateral margin coarsely serrated, moderately reflexed, posterior half of lateral margin finely serrate, lateral margin with long sparse erect setae; anterior margin completely beaded, with long sparse erect setae (Fig. 1 b); basal margin smooth and flat; pubescence distinctly long; setiferous punctures as large as those on head, densely set, some confluent. Scutellum. Scutellum triangular, flat; punctures smaller than those on pronotum; lateral margins smooth, without punctures; 1.75 times wider than long; with some erect moderately long setae near apex (Fig. 1 d). Elytra. Discal surface densely rugose-punctate, moderately and long pubescent; widest at apical one third. Epipleuron with marginal pubescence. Legs. Protibiae tridentate, teeth equidistant, slightly sharp, basal tooth small, middle and apical teeth mostly equal; internal apical spur inserted in front of middle tooth, reaching middle of protarsomere 1; dorsal carina of protibia reaching the apex of apical tooth, and with a branch reaching the apex of second tooth (Fig. 1 g). Profemora, mesofemora and metafemora moderately covered with hair-like, long setae; metafemora slightly broadened anteriorly and posteriorly, slightly convex. Metatibiae with two apical spurs, nearly equal; upper spur reaching base of second tarsomere (Fig. 1 f). Ventral surface. Meso- and metasternum with dense, fine punctures and yellowish hair (Fig. 1 c). Abdomen: punctures at middle finer and sparser than laterally; covered with setiferous, short setae. Pygidium. Slightly convex, triangular, side margins bordered, apical margin thickened; dorsal surface covered with long and short setae; punctures not clear as those on elytra (Fig. 1 h). Aedeagus. Parameres symmetric (Figs 4 a – c), including four braches (two dorsal and two thin ventral branches), separated in dorsal, ventral, and lateral views; dorsal braches longer than ventral ones, their apex with one ventral spine for each; ventral branches apex sharp and curved outward. Variability. The male paratypes are almost identical to the holotype. Female. Darker and bigger than males (body length 18.4 – 18.6 mm; body width 9.0 – 9.2 mm). Antennomeres of club 0.65 x to 0.75 x shorter than in males. Upper spurs of metatibiae longer than basal second metatarsomere.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C15FF98F8EEFDE0FC6FFE63.taxon	discussion	Remarks. Miridiba apicespinosa, new species shares with M. taoi Li & Wang, 2015 pubescence on pronotum, elytra and scutellum distinctly long, and pronotum with anterior margin concave at lateral ends. However, it can be easily separated from M. taoi, endemic to the small island of Lanyu off Taiwan and known only up to now by the 3 typical males, by: pronotum widest at basal third (Fig. 1 b) (widest at middle in M. taoi); metatibial upper spur reaching base of second tarsomere in male (Fig. 1 f) (reaching basal one-third of second tarsomere in M. taoi). Parameres: apex of dorsal branches with ventral spines (without such spines in M. taoi), and ventral branches long and reaching dorsal ones (short and not reaching dorsal ones in M. taoi) (Figs 4 a – c). The morphological characters of M. apicespinosa, new species match the features of the morphotype “ Trichophora ” (Fig. 1), thus we classify the new species as a member of this morphotype in the nominal subgenus Miridiba.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C15FF98F8EEFDE0FC6FFE63.taxon	etymology	Etymology. This species is named after the ventral spines on apex of dorsal branches of parameres.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C15FF98F8EEFDE0FC6FFE63.taxon	distribution	Distribution. Laos (Houa Phan) and Vietnam (Son La Province).	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C17FF9EF8EEFD84FD76FEF3.taxon	description	(Figs 2, 4 d-f)	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C17FF9EF8EEFD84FD76FEF3.taxon	materials_examined	Type material. Holotype: male (IEBR), Kon Tum Province, Dak Glei District, Ngoc Linh Nature Reserve, light trap, 10. iii. 2019, Pham Van Phu leg.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C17FF9EF8EEFD84FD76FEF3.taxon	description	Description of the holotype (male): Body length: 18.0 mm (from anterior margin of clypeus to apex of elytra); body width: 8.4 mm (width across humeri). Color dark reddish brown on head, pronotum; yellowish brown to dark reddish brown on scutellum, elytra, ventral surface, and legs; moderately shiny; dorsal surface glabrous (Fig. 2 a). Head. Surface densely punctate; punctures large; clypeus transverse, with apex moderately bilobed and reflexed, widened at base, nearly five times as long as wide (Fig. 2 b). Clypeofrontal suture clearly defined, weakly sinuated (Fig. 2 b). Frontal carina distinctly raised, weakly curved backwards, reaching periocular margins. Basal area of head with small punctures, poorly impressed, and sparser than forehead (Fig. 2 b). Antenna with 10 antennomeres, antennal club composed of 3 antennomeres, club equal to length of antennomeres 2 – 6 combined (Fig. 2 e). Last segment of maxillary palp elongated, not flattened. Pronotum. Transversal, widest at basal third; pronotum with anterior margin without concavities at lateral ends, slightly protuberant; basolateral angles round; two fifth anterior of lateral margin crenate, narrow to broad, slightly reflexed, crenate areas with long sparse erect setae; anterior margin completely flanged (Fig. 2 b); basal margin smooth; setiferous punctures shallower and smaller than those on head, rarely confluent. Scutellum. Triangular, flat; punctures slightly smaller than those on pronotum; lateral margins smooth, without punctures; two times wider than long (Fig. 2 d). Elytra. Discal surface densely and finely punctate, sutural costa gradually broadening apically, widest on apical one third. Epipleuron with some short pubescence in basal part. Legs. Protibiae tridentate, middle tooth closer to basal than to apical one, tip of teeth rounded; basal tooth small, middle tooth two times smaller than apical one; internal apical spur inserted in front of middle tooth, reaching basal one-third of protarsomere 1; dorsal carina of protibia reaching the apex of apical tooth developed, and with a branch reaching the apex of second tooth weakly (Fig. 2 g). Profemora, mesofemora, and metafemora moderately covered with hair-like, long setae; metafemora slightly broadened anteriorly and posteriorly, slightly convex. Metatibiae with two apical spur, upper spur slightly longer than lower spur (Fig. 2 f). Ventral surface. Meso- and metasternum with dense fine punctures and yellowish hair (Fig. 2 c). Abdomen: first and second segments densenly covered with setiferous punctures, and moderately long setae. From third to anterior half of fifth abdomen with sparse punctures, meanwhile from posterior half of fifth to last abdomen with dense punctures, short and long setae mixed. Pygidium. Convex, triangular, margin roundly bordered, apical margin thick; punctures on dorsal surface identical to those on elytra; apical margin with moderately long yellowish erect setae (Fig. 2 h). Aedeagus. Parameres with dorsal and ventral branches completely fused in lateral view, and symmetric, apex unfortunately chipped (Figs 2 d – f). Aedeagus is similar to those of M. excavata Keith, 2021, M. quasisinensis Keith, 2020, M. sinensis (Hope, 1842), and M. pseudosinensis Keith, 2010 (see figs 2, 3 in Keith 2020 a; figs 7, 8 in Keith 2021; figs 2, 3 in Keith 2010; and figs 12 A – C in Gao & Coca-Abia 2021). Female. Unknown.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C17FF9EF8EEFD84FD76FEF3.taxon	discussion	Remarks. Miridiba ngoclinhensis, new species is closely similar to M. pseudosinensis but both species can be separated by the basal third of lateral margin of pronotum is smooth in M. ngoclinhensis (Fig. 1 b), but that is serrate in M. pseudosinensis (Fig. 5 a); and the shape of parameres (Figs 4 d-f, 5 b, c): the thin area (recessed area) at the anterior half of paramere is membrous and thiner, without inner expansion in dorsal view, slightly curved downward in M. ngoclinhensis. The morphological characters of M. ngoclinhensis, new species characterize the morphotype “ Sinensis ” (Figs 12 – 14, Gao & Coca-Abia 2021), thus we classify the new species as a member of this morphotype and belonging to the subgenus Pledina.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C17FF9EF8EEFD84FD76FEF3.taxon	etymology	Etymology. This species is named after the locality of the holotype (Ngoc Linh Nature Reserve).	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C17FF9EF8EEFD84FD76FEF3.taxon	distribution	Distribution. Vietnam (Kon Tum Province).	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C11FF9DF8EEFE74FD11FE1F.taxon	description	(Figs 3, 4 g-i)	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C11FF9DF8EEFE74FD11FE1F.taxon	materials_examined	Type material. Holotype: male (IEBR), Quang Nam Province, Nam Giang District, Cha Vai, Song Thanh, light trap, 28. iv. 2005, 520 m above sea level.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C11FF9DF8EEFE74FD11FE1F.taxon	description	Description of the holotype (male): Body length: 16.3 mm (from anterior margin of clypeus to apex of elytra); body width: 7.7 mm (width across humeri). Color dark reddish brown on head, pronotum, and scutellum; yellowish brown to dark reddish brown on elytra, ventral surface, and legs; moderately shiny; dorsal surface almost glabrous (Fig. 3 a). Head. Surface densely punctate; punctures large; clypeus transverse, with apex moderately bilobed and reflexed, widened at base, length five times greater wide (Fig. 3 b). Clypeofrontal suture visible, moderately sinuated. Frontal carina moderately raised, weakly curved backwards, reaching periocular margins (Fig. 3 b). Basal area of head with punctures smaller than those on forehead, punctures with short erect setae (Fig. 3 b). Antenna with 9 antennomeres, antennal club composed of 3 antennomeres, club longer than length of antennomeres 2 – 6 combined (2.5 / 1.7 mm). Last segment of maxillary palp elongated, dorsal surface with callus. Pronotum. Transversal, widest at middle; anterolateral angles moderately protuberant; basolateral angles round; lateral margin smooth; anterior margin completely bordered; basal margin finely flanged and smooth, without pubescence; setiferous punctures smaller than those on clypeus (Fig. 3 b). Scutellum. Triangular, flat; strongly and densely puncate except on side margins; nearly two times wider than long (Fig. 3 d). Elytra. Elytra nearly parallel, slightly widened posteriorly. Punctures dense near the base, less posteriad; base of elytra bearing several long setae. Epipleura end at one fifth posteriorly bearing a row of long setae. Legs. Protibiae tridentate, teeth equidistant, tip of teeth moderately round; basal tooth small, middle tooth two third as apical teeth; internal apical spur inserted in front of middle tooth, reaching basal one-third of protarsomere 1; dorsal carina of protibia reaching the apex of apical tooth, and with a branch reaching the apex of second tooth (Fig. 3 e). Profemora, mesofemora, and metafemora moderately covered with hair-like, long setae; metafemora strongly broadened anteriorly, slightly convex. Metatibia with two apical spurs, upper spur moderately longer than lower spur, and reaching one fourth of mesotarsomeres 2 (Fig. 3 f). Meso- and metatibiae with strong teeth on external margins and complete carina at apical third. Ventral surface. Meso- and metasternum with dense fine punctures and yellowish hair (Fig. 3 c). Abdomen: second and third visible abdomenal sternites densely covered with punctures, and long setae; those on fourth and fifth sternites sparserly punctate and setiferous, particularly on median parts; sixth sternite covered with longer setae. Pygidium. Slightly convex, triangular, margin bordered, apical margin moderately reflexed; dorsal surface with punctures identical to those on elytra with long and erect setae (Fig. 3 f). Aedeagus. Curved downwards, symmetrical in lateral view, bird's beak-like, without acicular expansion at base (Figs 4 g – i). Female. Unknown.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C11FF9DF8EEFE74FD11FE1F.taxon	discussion	Remarks. Miridiba quangnamensis, new species is closely similar to M. hanoiensis Keith, 2006 (Figs 5 d – f) but they can be separated by the following characters combined: posterior pronotal margin without pubesence in M. quangnamensis, new species (pubescent in M. hanoienis); clypeo-frontal suture visible in M. quangnamensis, new species (invisible in M. hanoiensis); antenna club longer than length of antennomeres 2 – 6 combined in M. quangnamensis, new species (Fig. 3 d) (almost equal in M. hanoiensis); anterolateral angles moderately protuberant in M. quangnamensis (Fig. 3 b) (nearly rectangular in M. hanoiensis – Fig. 5 d); median limits between sternites slightly opaque in in M. quangnamensis, new species (invisible in M. hanoiensis), and the shape of parameres (Figs 5 e, f; 4 g – i). The morphological characters of M. quangnamensis new species do not characterize any morphotype proposed by Gao & Coca-Abia (2021) and Gao et al. (2022), so the new species remains as incertae sedis in the genus Miridiba.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C11FF9DF8EEFE74FD11FE1F.taxon	etymology	Etymology. This species is named after the locality of the holotype (Quang Nam Province).	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C11FF9DF8EEFE74FD11FE1F.taxon	distribution	Distribution. Vietnam (Quang Nam Province).	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C13FF9CF8EEFA58FC40F9FE.taxon	materials_examined	Material examined. 16 males, 3 females (DKCC), Quang Nam, Tay Giang, Mt Axana, VII-VIII. 2021, local collector.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
03C087917C13FF9CF8EEFA58FC40F9FE.taxon	distribution	Distribution. Up to date, the species was known from the type series only collected in Yunnan (China). Specimens from Quang Nam represent the first record for Vietnam.	en	Pham, Phu Van, Keith, Denis, Pham, Nhi Thi, Cao, Nga Quynh Thi (2022): Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record. Zootaxa 5205 (4): 331-342, DOI: https://doi.org/10.11646/zootaxa.5205.4.2
