taxonID	type	description	language	source
8D1487CFFFADF926FF665A58FA27FE8B.taxon	description	(Fig. 2: An 13, A 98, A 99, B 01; Fig. 3; Fig. 4: A 98)	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFADF926FF665A58FA27FE8B.taxon	description	Kerana sumata Fruhstorfer, 1911. Type locality: Nias. Kerana savara Fruhstorfer, 1911. Type locality: Borneo.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFADF926FF665A58FA27FE8B.taxon	description	Male genitalia (Fig. 3). In lateral view, uncus a bit shorter than tegumen, slightly convex, distal end pointed, lower branch triangular; gnathos arm-like, transverse part a little longer than longitudinal part; saccus about 1 / 3 the length of vinculum. In dorsal view, distal half of uncus bifid and U-shaped, tip of both branches with a notch. In lateral view, dorsal edge of valva concave, with a rounded distal shoulder; distal edge of valva rounded; harpe triangular, its outer side vertical. Distal branch of valva needle shaped in lateral view, while bifid and decorated with tiny sawteeth in dorsal view, its distal triangular lobe pointed. In lateral view of aedeagus, suprazonal sheath slightly curved downwards, about 2 / 5 the length of subzonal sheath. Juxta bifid and V-shaped, basal edge concave. Female genitalia (Fig. 4: A 98). In lateral view, lamella antevaginal almost bent to a right angle, its distal part triangular. In ventral view, lamella antevaginal nearly U-shaped, ostium bursae semi-sclerotized. In front view of lamella postvaginalis, its dorsal edge shallowly concave.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFADF926FF665A58FA27FE8B.taxon	discussion	Remarks. Evans (1949) stated that “ the male (of othonias) is unknown and it seems not unlikely that othonias may prove to be a dimorphic form of armatus. ” However, Maruyama (1985, 1991) treated othonias as a subspecies of nigrita based on the behavior and the color of the ventral side, and synonymized maura with othonias without providing any scientifically sound evidence. de Jong & Treadaway (2007) also pointed out that the female of nigrita in Borneo is dimorphic, and one form is othonias. In the present study, the NJ tree (Fig. 1) shows that othonias is a separate species rather than a form of armatus or a subspecies of nigrita, and maura is conspecific with othonias. The specimen of “ Ancistroides nigrita ” with GenBank accession number HM 862403 and coordinate 7.533 N, 99.783 E (pointing to a locality in Peninsular Thailand) is probably a representative of maura. The publication year of othonias was recorded as 1876 by Evans (1949) but 1878 by Maruyama (1991). Likewise, the publication year of maura was recorded as 1880 by Evans (1949) and de Jong & Treadaway (2007), but 1886 by Maruyama (1991). According to Bridges (1994), Maruyama (1991) ’ s record is correct in both cases.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFADF926FF665A58FA27FE8B.taxon	distribution	Distribution. S. Myanmar, Thailand, Laos, S. Vietnam, Malay Peninsula, Nias, Sipora, Sumatra, Bangka, Borneo.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFABF927FF665A58FDD0FEF1.taxon	description	(Fig. 2: A 97; Fig. 5: A 97) Hesperia nigrita Latreille, [1824]: 788. Type locality: Java. Kerana vasuba Fruhstorfer, 1911. Type locality: Java.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFABF927FF665A58FDD0FEF1.taxon	description	In the nominate subspecies, the forewing length is 21 – 23 mm (n = 1) in male (Evans 1949). The male genitalia (Fig. 5: A 97) are similar to those of Ancistroides othonias, except for: harpe elongated, reaches dorsal edge of valva; distal branch of valva thicker and finger shaped in lateral view, with a wider distal triangular lobe in dorsal view; subzonal sheath of aedeagus longer and more bent upwards in lateral view.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFABF927FF665A58FDD0FEF1.taxon	discussion	Remarks. Eliot (1992) stated that “ (in maura) the marginal areas of both wings are broadly paler on the underside ”. This description is contradictory with that by Evans (1949). Judging from the photos (Eliot 1992: plate 55, fig. 6, 7), the taxon examined by Eliot (1992) should belong to Ancistroides nigrita nigrita. Distribution. Malay Peninsula, Java.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFAAF927FF665D30FB4CFCB2.taxon	description	(Fig. 2: An 4, An 5; Fig. 4: An 5; Fig. 5: An 4) Nisoniades diocles Moore, [1866]: 787. Type locality: Bengal.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFAAF927FF665D30FB4CFCB2.taxon	distribution	Distribution. Sikkim, Bhutan, Assam, Bengal, N. Shan St., Karens, S. Guangxi, Hainan Island.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFAAF924FF665B4CFE57FD8D.taxon	description	(Fig. 2: An 10, An 11; Fig. 4: An 11; Fig. 6)	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFAAF924FF665B4CFE57FD8D.taxon	description	Male genitalia (Fig. 6) can be distinguished from those of Ancistroides othonias (Fig. 3) and A. nigrita (Fig. 5) by the following characters: in lateral view, uncus as long as tegumen, lower branch hooked and well developed; transverse part of gnathos obviously longer than longitudinal part; each side of uncus curved inwards in dorsal view, distal half V-shaped; in lateral view of valva, dorsal edge wavy, distal edge protrudent; harpe rubost and tapered; distal branch of valva finger-like and pointed in lateral view, while bifid and decorated with tiny spines in dorsal view, its distal triangular lobe much wider; suprazonal sheath of aedeagus bent downwards; juxta with deeper furcation, basal edge flat. Female genitalia (Fig. 4: An 11) are similar to those of Ancistroides othonias (Fig. 4: A 98) and A. nigrita (Fig. 4: An 5), but can be recognized by: distal part of lamella antevaginal elongated and tapered into a blunt point in lateral view; distal edge of lamella postvaginalis deeply concave.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFAAF924FF665B4CFE57FD8D.taxon	distribution	Distribution. Philippines.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFA9F925FF665958FCA5FEF1.taxon	description	(Fig. 2: A 95, An 8; Fig. 4: An 8; Fig. 7)	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFA9F925FF665958FCA5FEF1.taxon	description	Male genitalia (Fig. 7). In lateral view, uncus shorter than tegumen, basal part raised, distal end pointed, lower branch short and triangular; gnathos arm-like, transverse part a little longer than longitudinal part; saccus about 2 / 5 the length of vinculum. In dorsal view, uncus tapered, distal half bifid and narrowly U-shaped, tip of both branches rounded. In lateral view of valva, dorsal margin flat, distal part bifid into a protrudent shoulder and a sickle shaped branch which bears sawteech on inner edge; harpe triangular, vestigial. Aedeagus remarkably robust, left half of dorsal edge with a wide and deep gap; in lateral view, suprazonal sheath slightly curved downwards, about 3 / 5 the length of subzonal sheath. Juxta shield-shaped, with a pair of slender and pointed dorsal arms. Female genitalia (Fig. 4: An 8). In lateral view, lamella antevaginal nearly oval, strongly developed; antrum very robust. In ventral view, a wide gap between two strong arms of lamella antevaginal, ostium bursae well sclerotized. Lamella postvaginalis trapezoidal, middle of distal edge with a shallow V-shaped notch.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFA9F925FF665958FCA5FEF1.taxon	distribution	Distribution. S. Myanmar, Peninsular Thailand, W. Malaysia, Sumatra, Batoe, Sipora, Borneo. Remarks. According to Evans (1949), there is the other subspecies, viz. Ancistroides armatus niasana (Evans, 1926) which is endemic to Nias. The band on the dorsal side of the forewing is obsolete in male, and reduced to separated suffused red spots in female (Evans 1949: 281).	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFA8F92AFF66586CFD9CFE41.taxon	description	(Fig. 2: A 96; Fig. 8)	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFA8F92AFF66586CFD9CFE41.taxon	description	Male genitalia (Fig. 8). In lateral view, uncus as long as tegumen, basal part hump-shaped, distal end pointed, lower branch finger-like; gnathos curved upwards; saccus about 1 / 2 the length of vinculum. In dorsal view, middle of basal uncus concave, a pair of distal branches widely U-shaped, tip of both branches pointed. In lateral view of valva, dorsal edge flat, ventral edge evenly curved, tip of distal branch rounded; harpe slant, distal half short fingershaped, blunt. In dorsal view, distal branch of valva trapezoidal, with its proximal inner angle sharply pointed. Aedeagus elongated S-shaped in lateral view, suprazonal sheath a little shorter than 1 / 2 the length of subzonal sheath. Juxta deeply bifid and U-shaped, basal edge concave.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
8D1487CFFFA8F92AFF66586CFD9CFE41.taxon	distribution	Distribution. Timor, Sulawesi.	en	Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki, Li, Wenwei (2022): Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae). Zootaxa 5205 (5): 481-490, DOI: https://doi.org/10.11646/zootaxa.5205.5.5
