identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038387EAFFB9AC6312F11F8CFA4CF85E.text	038387EAFFB9AC6312F11F8CFA4CF85E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius addarcis Prokofiev 2022	<div><p>Maechidius addarcis Prokofiev, new species</p> <p>Fig. 1</p> <p>Type material. Holotype, male (Fig. 1A): “ New Guinea / Doron xi.1921 / R. Neill Rothschild / B.M. 1939-I ” (BMNH). Paratypes: 3 females, same data as for the holotype (BMNH).</p> <p>Description of the holotype. Total body length 8.5 mm, greatest width 4.2 mm. Dark reddish-brown with legs more reddish; antennae and palpi reddish; setation pale. Body (including dorsal surface of head but not labroclypeus) is almost completely covered with microscopical velvety pubescence. Antennae 9-segmented, club 3-lamellate. Labroclypeus broadly concave anteriorly, its lateral margins strongly bisinuate in both dorsal and lateral views; anterolateral angles small and pointed, triangular, weakly reflexed; canthus angulate. Dorsal surface of labroclypeus coarsely punctured, punctures deep, annular, with dense microscopical velvety pubescence; intervening spaces variable, smooth and glossy. Canthus is tightly covered with dense microscopical pubescence, with few moderately long erect setae. Underside of labroclypeus rather densely setose. Frons flat; punctures similar to those on labroclypeus, but much densely arranged; intervening spaces very narrow; setae erect to semi-erect, short, commensurable with diameter of punctures. Pronotum transverse with anterior margin concave and bisinuate; anterolateral angles acute; sides of pronotum broadly rounded in anterior two-thirds and deeply concave in posterior third; basal margin strongly convex; posterolateral angles almost straight. Lateral margins of pronotum crenulate, with about 20 poorly expressed crenulae, better developed along middle widest part of pronotum; a very short seta (often worn) present between every two crenulae. Pronotal punctures large, annular to hexagonal; intervening spaces very narrow; setation (largely worn) similar to that on head and elytra. Hypomeron separated from prosternum by moderately high emarginate carina bearing long setae; antennal pockets rather deep; punctures large, annular; intervening spaces generally smaller than punctures; setae moderately long in anterior half, becoming short, not surpassing diameter of puncture in posterior half. Scutellar shield apically rounded, with dense and rather coarse annular puncturation; setae probably worn. Elytra weakly widened posteriad, broadly rounded apically, with four weak longitudinal carinae splitting to a series of sparsely sitting granules (two rows between sutural margin and humeral umbo, third row running from humerus toward apical umbo, fourth one extending forward from apical umbo along lateral margin of elytron); sutural joint not elevated. Punctures of elytral disc narrow, incision-shaped; intervening spaces generally smaller than length of punctures; setae adpressed to semi-erect, not or hardly surpassing length of corresponding punctures. Pygidium flat, with shallow annular punctures; intervening spaces very narrow; setae semi-erect, similar to those on elytra. Thorax and abdominal ventrites with small and shallow annular punctures; intervening spaces smaller than diameter of punctures; setae adpressed, generally exceeding diameter of punctures in length on thorax and not surpassing it on abdominal ventrites. Ventral profile of abdomen slightly convex. Protibia (Fig. 1B) weakly widened distally, with two longitudinal carinae dorsally and two strong apical teeth directed outward; basal tooth weak. Protibial spur short, straight, somewhat inclined ventrally. Metatibial spurs short and straight, pointed; lower spur thicker and twice longer than upper one. Tarsomeres not elongated, setae not forming brush-like patches along their ventral margin. Tarsal claws with pulvilli. Aedeagus and spiculum gastrale, as on Figs. 1C–F; parameres symmetrical, with tips rectangular, crossing each other in default position. Spiculum gastrale flat along most of its length, only the base of its shaft strongly reflexed (Fig. 1F).</p> <p>Females. Total body length 8.0– 9.5 mm, greatest width 4.0– 4.5 mm. Secondary sexual differences not expressed except the comparatively shorter antennal lamellae, more pronounced basal tooth of protibia (Fig. 1G) and longer spurs on all tibiae. Metatibial spurs only slightly different in length, weakly clavate at tips. Sides of pronotum before posterolateral angles variably excavated. Longitudinal carinae on elytra variably developed, from continuous along almost whole length of elytra to expressed at their base and near umbones only.</p> <p>Differential diagnosis. This species is keyed to the couplet 15 in Telnov (2020), but can be easily distinguished from both M. subcostatus Heller, 1895 and M. similis Telnov, 2020 by dense coating of microscopical velvety pubescence and in the unique shape of parameres with tips crossing each other in default position. It is superficially somewhat similar to the members of the former genus Paramaechidius or M. pauxillus group as of Telnov (2020) but can be easily differentiated from them by absence of the clavate setae on the dorsum, presence of the tarsal pulvilli in both sexes, the dorsally flat frons and different aedeagus.</p> <p>Etymology. The species epithet is created from the Latin words ad (to) and arx (tip), due to the conjoining tips of the parameres.</p></div> 	https://treatment.plazi.org/id/038387EAFFB9AC6312F11F8CFA4CF85E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFBAAC6112F11A55F8FAFBF0.text	038387EAFFBAAC6112F11A55F8FAFBF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius astrolabius Prokofiev 2022	<div><p>Maechidius astrolabius Prokofiev, new species</p> <p>Fig. 2</p> <p>Type material. Holotype, male (Fig. 2A): “N. Guinea / Astrolabe-B. / Rhode ” (ZMHB).</p> <p>Description of the holotype. Total body length 7.5 mm, greatest width 4.2 mm. Dark reddish-brown, glossy from above; antennae and palpi rusty-brown, much lighter than body; setation pale. Antennae 9-segmented, club 3-lamellate. Labroclypeus broadly concave anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views; anterolateral angles distinctly protruding, reflexed; canthus angulate. Dorsal surface of labroclypeus densely punctured, punctures deep, annular, approximately 1.5 times less than those on frons; setation apparently worn. Underside of labroclypeus minutely punctured, punctures setigerous. Frons weakly convex. Punctures on frons very large and deep, ovoid, with dense microscopical velvety pubescence; intervening spaces very narrow, convex, smooth and glossy; setae (largely worn) moderate in length, ×0.75–1.5 length of puncture. Pronotum transverse, with anterior margin concave and anterolateral angles strongly protruding, acute. Sides of pronotum smoothly converging anteriad in anterior third, almost parallel-sided in posterior two-thirds; basal margin convex, more strongly before scutellar shield; posterolateral angles obtuse. Lateral margins of pronotum distinctly crenulate, with 16 crenulae becoming slightly larger in middle third of side-length; a short seta present between every two crenulae (often worn). Punctures and setation of pronotum similar to those on head, but punctures on disc slightly larger; intervening spaces very narrow, ridge-like, smooth and glossy, forming an allusion of longitudinal ridges in oblique light. Hypomeron separated from prosternum by moderately high emarginate carina bearing moderately long setae; antennal pockets conspicuous but not deep. Punctures of hypomeron large, annular, setigerous; setae as long as punctures; intervening spaces much smaller than diameter of puncture, shagreened, matt. Scutellar shield deeply and densely punctured, apically pointed; intervening spaces much smaller than diameter of punctures, shagreened, matt. Elytra moderately widened posteriad, broadly rounded apically, with vague traces of two longitudinal carinae between sutural carina and humeral umbones. Punctures of elytral disc long and narrow, incision-shaped, becoming smaller toward side-margins; intervening spaces wrinkled, finely reticulate, glossy, generally larger than length of punctures; setae adpressed, not surpassing length of corresponding punctures. Pygidium flat, with large and dense annular punctures; intervening spaces very narrow, shagreened, matt; setae semi-erect, ×1.0–1.5 diameter of puncture, becoming longer toward apical margin. Thorax and abdominal ventrites shagreened and matt except the smooth and glossy disc of metasternum; punctures annular, larger on disc of metasternum and in middle third of abdominal ventrites, becoming smaller laterally; intervening spaces generally smaller than diameter of punctures; setae not or slightly surpassing diameter of corresponding punctures. Ventral profile of abdomen slightly convex. Protibia (Fig. 2B) widened distally, with two longitudinal carinae dorsally and three outer teeth, basal tooth very weak; protibial spur long and pointed, straight along most of its length but slightly curved at base. Metatibial spurs with blunt tips, lower spur only slightly longer than upper one and weakly curved. All tarsomeres short, first two metatarsomeres of same length. Tarsal claws with pulvilli. Aedeagus, as on Figs. 2C–D; parameres somewhat asymmetrical, with overlapping tips. Spiculum gastrale was damaged (apparently thin and flat).</p> <p>Female unknown.</p> <p>Differential diagnosis. This species is keyed to the couplet 4 in Telnov (2020) and resembles M. acutus (Narakusumo &amp; Balke, 2019), M. humeralis Heller, 1914, and M. paralellicollis Moser, 1920 in many respects. However, M. astrolabius, new species can be easily separated from all aforementioned species by the larger-sized ovoid punctures on the head and pronotum covered with the fur-like micropubescence (vs. annular and lacking micropubescence) and by the bigger denticles along the lateral sides of the pronotum (16 vs. 19–25 in the compared species). It is more similar to M. acutus and M. humeralis by the presence of the vague longitudinal carinae on elytral disc and by the transversely wrinkled intervening spaces between the punctures on the elyta (both absent in M. paralellicollis) but differs from all these species in the shape of aedeagus. The aedeagus of the new species is most similar to that in M. acutus (Narakusumo &amp; Balke, 2019: figs. 1e–f), differing mainly by its asymmetrical parameres.</p> <p>Etymology. This species is named from its type locality, Astrolabe Bay.</p></div> 	https://treatment.plazi.org/id/038387EAFFBAAC6112F11A55F8FAFBF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFBCAC6412F11A6DFACEFF7C.text	038387EAFFBCAC6412F11A6DFACEFF7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius bombycinus Prokofiev 2022	<div><p>Maechidius bombycinus Prokofiev, new species</p> <p>Figs. 3–4</p> <p>Type material. Holotype, male (Fig. 3A): “ D. Neue-Guinea / Huon-Golf / Bukaua / Prof. Neuhauss S.” (ZMHB).</p> <p>Additional material examined. 1 female: “D. N. Guinea / Sattelberg” (ZMHB); 1 female: “D. N. Guinea / Sattelberg XII.08 / Prof. Neuhauss S.” (ZMHB); 1 female: “ Htld. V. Finschhafer I. 30 / L. Wagner S.” (ZMHB); 1 female: “D. N. Guinea / 2.VIII.12 Hauptlager / Kais. Augustafl. Exp. / Bürgers S.G.” (ZMHB); 1 female: “D. N. Guinea / Standlager a. Aprilfluss 19.-26.ix.12 / Kais. Augustafl. Exp. / Bürgers S.G.” (ZMHB); 1 female: “D. N. Guinea 152 / Standlager a. Aprilfluss / 15.-17.ix.12 Kais. Augustafl. Exp. / Bürgers S.G.” (ZMHB); 1 female: “D. N. Guinea 175 / Standlager a. Aprilfluss 8.-9.x.12 / Kais. Augustafl. Exp. / Bürgers S.G.” (ZMHB).</p> <p>Description of the holotype. Total body length 6.3 mm, greatest width 3.0 mm. Reddish-brown, silky; antennae yellowish-brown; setation pale. Antennae 9-segmented, club 3-lamellate. Labroclypeus deeply emarginate anteriorly, its lateral margins deeply and regularly concave between anterolateral angles and canthus in dorsal view, sinuous in lateral view; anterolateral angles distinctly protruding, lobate, reflexed; canthus broadly rounded in dorsal view. Dorsal surface of labroclypeus densely and rather finely punctured, punctures annular, setigerous, setae adpressed, somewhat shorter than on frons. Underside of labroclypeus sparsely setose. Frons flat. Punctures on frons moderately large, dense, annular; intervening spaces narrow and microreticulate; setae long, of inequal length, subequal to diameter of corresponding punctures or up to 2 its diameters in length. Pronotum transverse, with anterior margin concave and slightly bisinuate, anterolateral angles strongly protruding, acute. Pronotum widest across middle, its sides slightly more convergent anteriad than posteriad; basal margin bisinuate, convex before scutellar shield; posterolateral angles obtuse. Lateral margins of pronotum indistinctly crenulate, with crenulae almost confluent and uncountable in anterior half, becoming more obvious in posterior half of their length; a long seta (about half-length of compound eye) present between every two crenulae. Pronotal punctures moderately large, annular; intervening spaces smaller than punctures, with microreticulate sculpture giving a silky appearance; punctures setigerous, with setae not surpassing diameter of punctures. Hypomeron separated from prosternum by low continuous carina bearing moderately long setae; antennal pockets weakly expressed. Punctures of hypomeron moderately large and sparse, annular; their setae probably completely worn; intervening spaces very delicately microreticulate. Scutellar shield densely and somewhat irregularly punctured, apically pointed; punctures slightly smaller than on pronotum; intervening spaces generally smaller than diameter of punctures, microreticulate. Elytra moderately widened posteriad, subtruncate apically, without longitudinal carinae. Punctures of elytral disc long and narrow, incision-shaped; intervening spaces generally larger than length of punctures, with conspicuous microreticulate sculpture giving a silky appearance. Setae on elytral disc adpressed, not surpassing length of corresponding punctures, becoming longer (up to 1.5 length of punctures) and more erect toward the sides and apex. Pygidium weakly convex, with large annular punctures, more sparsely arranged along mid-line; intervening spaces generally smaller than diameter of punctures (except along mid-line), glossy, with microreticulation obvious laterobasally only; setae semi-erect, ×1.2–1.5 diameter of puncture, becoming longer toward apical margin. Meso- and metasternum bearing moderately small annular punctures of similar sizes; intervening spaces variable, distinctly microreticulate; setae expressed in punctures on metasternal disc only, adpressed, variable surpassing diameter of punctures. Abdominal ventrites distinctly microreticulate between moderately large annular punctures arranged in transverse rows. Ventral profile of abdomen weakly concave. Protibia widened distally, with two longitudinal carinae dorsally, two conspicuous apical teeth and very poorly expressed basal tooth; protibial spur absent, but apical inner side of protibia forming an angular extension at place of spur (Fig. 3B). Metatibial spurs slightly inequal in length, almost straight, pointed. All tarsomeres ordinary. Tarsal claws with large pulvilli. Aedeagus and spiculum gastrale, as on Figs. 3C–F.</p> <p>Females (tentatively referred). Total body length 6.0– 6.5 mm, greatest width 2.8–3.0 mm. Dark to pale reddishbrown, sometimes with greenish tint on dorsal head and pronotum in oblique light (Fig. 4A). Labroclypeus from weakly concave to almost straight anteriorly (Figs. 4B–C), its lateral margins weakly convex to weakly sinuous in dorsal and lateral view; anterolateral angles poorly expressed, obtuse, weakly reflexed; canthus broadly rounded in dorsal view. Microreticulate sculpture on pronotum and elytra less pronounced than in male holotype. Hypomeral carina less sharp than in male holotype. Elytral setae comparably longer and thicker than in male holotype, those near sutural joint grouping in double parallel rows in some specimens (maximal expression as shown on Fig. 4A). No conspicuous elytral carinae, but some intervening spaces variably thickened in some cases. Protibia with two strong apical teeth and variably expressed basal tooth (but usually strong); protibial spur absent, with no angular extension at apical inner side of protibia (Fig. 4D). Metatibial spurs inequal in length, lower one almost straight, 1.5 longer than upper one, distinctly clavate; upper spur slightly curved, blunt but not expanded at tip. Tarsal claws with large pulvilli. Anal valvi comma-shaped, with rather long setae at their distal tips.</p> <p>Remarks. Identification of the females is tentative and somewhat disputable.Although most of their differences from the male holotype are connected with the sexual dimorphism, the less pronounced hypomeral carina and less conspicuous microsculpture of the dorsal body may not be this case. Elytral setae trend to group in two to three double rows on each side along the suture in some females, reminiscent the pattern of M. nepenthephilus but more localized (cf. Telnov, 2020: fig. 65). Since no males associated with these females are known, they were not included in the type series and are provisionally referred to this species.</p> <p>Differential diagnosis. Although this species is keyed to the couplet 19 in Telnov’s (2020) key, it is most similar and apparently closely related to M. lobaticeps Frey, 1969 from the opposed couplet. In contrast to M. lobaticeps, the new species has distinctly microreticulate, opaque and silky dorsal surface (vs. smooth and shiny in M. lobaticeps). Nevertheless, the other characters in both species are very similar and the aedeagi show only slight differences: the parameres are more parallel-sided in dorsal view with more attenuated tips in the lateral view in the new species in comparison with M. lobaticeps (Telnov 2020: figs. 649–651). Within the species with microreticulate dorsal surface, the new species is most similar to M. nepenthephilus Telnov, 2020, but can be distinguished by more pronounced and lobate anterolateral angles of the male labroclypeus, elytral puncturation not forming the irregularly paired rows of punctures (at least in male), by less regular puncturation of the pygidium and shape of the parameres (Telnov 2020: figs. 679–680).</p> <p>Etymology. The species is named from the Latin adjective bombycinus, a, um (silky), in reference to the strongly opaque integument.</p></div> 	https://treatment.plazi.org/id/038387EAFFBCAC6412F11A6DFACEFF7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFBEAC6A12F11A59FA43FECC.text	038387EAFFBEAC6A12F11A59FA43FECC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius echinoides Prokofiev 2022	<div><p>Maechidius echinoides Prokofiev, new species</p> <p>Fig. 5</p> <p>Type material. Holotype, male (Fig. 5A): “Mäanderberg 387 / 1.-9.8.13 / Bürgers S.G.” (ZMHB). Paratypes: 2 females: “D. N. Guinea / Mäanderberg / 1.-10.VIII.13 / Kais. Augustafl. Exp. / Bürgers S.G.” (ZMHB); 1 female: same data, but 19.-31.VII.13 (ZMHB).</p> <p>Description of the holotype. Total body length 8.5 mm, greatest width 4.2 mm. Dark reddish-brown, glossy, forebody and fore legs darker than rest of body; antennae and palpi reddish; setation yellowish.Antennae 9-segmented, club 3-lamellate, as long as funicle. Labroclypeus broadly concave anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views; anterolateral angles protruding, sharpened, reflexed; canthus angulate. Dorsal surface of labroclypeus densely punctured, punctures deep, annular to hexagonal or slightly oblong, increasing in size from anterior margin of labroclypeus to frons; setae erect, much exceeding diameter of corresponding puncture in length, longer on frons than on labroclypeus. Underside of labroclypeus densely setose. Frons flat. Punctures on dorsal head and pronotum with dense microscopical velvety pubescence; intervening spaces narrow, in part ridge-like, smooth and glossy. Pronotum transverse, with anterior margin concave and weakly bisinuate, anterolateral angles strongly protruding, acute. Sides of pronotum more convex anteriad than posteriad, widest at middle, slightly constricted postmedially, denticulate all along (about 18 denticles); basal margin evenly convex; posterolateral angles obtuse. Punctures on pronotum almost equal to those on frons in size, annular or hexagonal to oblong, setigerous, with setae erect, much exceeding their diameter in length, almost equal to only slightly shorter than diameter of compound eye; the similarly long setae present between every two lateral denticles. Hypomeron separated from prosternum by rather high emarginate carina bearing long setae; antennal pockets moderately deep. Punctures of hypomeron shallow, annular, setigerous, somewhat more densely arranged anteriorly than posteriorly; setae not suprassing diameter of punctures; intervening spaces smooth and glossy. Scutellar shield apically pointed, rather densely punctured; setae adpressed, hardly surpassing diameter of punctures; intervening spaces microreticulate, matt. Elytra slightly widened posteriad, broadly rounded apically, with very indistinct traces of longitudinal carinae. Punctures of elytral disc long and narrow, incision-shaped; intervening spaces in part microreticulate and opaque, in part smooth and glossy; setae erect, much exceeding length of corresponding punctures. Pygidium weakly convex on disc and slightly impressed along sides, densely punctured with annular punctures becoming smaller toward apex; intervening spaces very narrow, smooth and glossy; setae erect, much exceeding diameter of punctures in length. Thorax and abdominal ventrites densely punctured, with punctures annular, larger in size toward mid-ventral line, setigerous; intervening spaces smaller than diameter of punctures, smooth and glossy; setae semi-erect, becoming longer toward mid-ventral line, those on abdominal ventrites longer than on thorax, up to twice longer than diameter of corresponding puncture. Ventral profile of abdomen weakly concave. Protibia (Fig. 5B) widened distally, with delicate longitudinal carina dorsally, with two long and sharp apical teeth and very vague basal tooth; protibial spur straight, pointed. Metatibial spurs very slightly unequal in size, not sharpened at tip. All tarsomeres ordinary. Tarsal claws with pulvilli. All legs with long setae. Aedeagus and spiculum gastrale, as on Figs. 5C–F; parameres with lateral margins sinuous in dorsal view, lateral sides with bipartite longitudinal impression; spiculum gastrale strongly bent.</p> <p>Females. Total body length 8.5–10.2 mm, greatest width 4.2–4.8 mm. From reddish- to blackish-brown. Labroclypeus shallowly and broadly emarginated anteriorly, its lateral sides bisinuate to almost regularly convex from anterolateral angles to canthus; anterolateral angles usually less protruding than in male and rounded. Antennal club as long as in male. Sides of pronotum denticulate, with 17–20 denticles. Ventral profile of abdomen weakly concave to almost straight. Distalmost tooth on protibia longer than in male (Fig. 5G). Metatibial spurs also longer, with tips blunt, almost truncate.</p> <p>Differential diagnosis. This species shows no external differences from M. esau Heller, 1914 and has a very similar aedeagus; however, in contrast to the latter, the lateral impression of parameres is bipartite, and lateral margins of parameres are distinctly angulate in the dorsal view in the new species (Figs. 5C–D; Telnov 2020: figs. 614–616). The spiculum gastrale is also different in shape (Figs. 5E–F; Telnov 2020: fig. 561).</p> <p>Etymology. The species name (meaning hedgehog-like in Latin) is given in reference to the long setation of this new species.</p></div> 	https://treatment.plazi.org/id/038387EAFFBEAC6A12F11A59FA43FECC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFB0AC6B12F11EE3FBFFFEB4.text	038387EAFFB0AC6B12F11EE3FBFFFEB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius interruptocarinulatus Heller 1914	<div><p>Maechidius interruptocarinulatus Heller, 1914</p> <p>Fig. 6</p> <p>Additional material examined. 1 male (Fig. 6A): Cyclopengebg. / Lager I 800m 8-9.28 / E. Mayr S.G. (ZMHB); 2 males: D. N. Guinea / Kaiserin Augustaflüss-Expedition / Bürgers S.G. (ZMHB); 1 female: D. N. Guinea 98 / Kais. Augustafl Exp. VII.12 / Bürgers S.G. (ZMHB).</p> <p>Description of the males. Total body length 7.5–8.0 mm, greatest width 3.7–4.2 mm. Same characters as described for females by Telnov (2020: 58) with the following additions. Antennal club slightly longer than in female. Anterolateral angles of labroclypeus long and pointed, strongly protruding and distinctly reflexed, anterior margin of clypeus between them regularly concave. Lateral margins of pronotum from broadly and regularly rounded to slightly more sinuous posteriad. Intervening spaces between pronotal punctures very narrow and ridgelike, somewhat variable in arrangement, giving an allusion of fingerprints or of more regular longitudinally arranged ridges. Teeth on protibia narrower and more conspicuously inclined laterad than in female. Protibial spur hooked at tip (Fig. 6B). Tarsal claws with pulvilli. Aedeagus and spiculum gastrale, as on Figs. 6C–F; distal tips of parameres spatulate, curved at right angle.</p> <p>Remarks. The additional specimens show no valuable differences from the female holotype of that species from the Etna Bay (Naturhistorische Sammlungen Dresden, examined). This species is widely distributed throughout the western part of New Guinea and Waigeo Island and was already reported for the Cyclops Mountains by Telnov (2020).</p></div> 	https://treatment.plazi.org/id/038387EAFFB0AC6B12F11EE3FBFFFEB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFB1AC6912F11AB3F9FEFAA4.text	038387EAFFB1AC6912F11AB3F9FEFAA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius macrosoma Prokofiev 2022	<div><p>Maechidius macrosoma Prokofiev, new species</p> <p>Fig. 7</p> <p>Type material. Holotype, male (Fig. 7A): “D. N. Guinea 201 / Etappenberg 850 m 2.-8.xi.12 / Kais. Augustafl. Exp. / Bürgers S.G.” (ZMHB).</p> <p>Description of the holotype. Total body length 10.5 mm, greatest width 5.7 mm. Dark reddish-brown, forebody darker, glossy; elytra alutaceus; antennae and palpi rusty-brown, much lighter than body; setation pale. Antennae 9-segmented, club 3-lamellate. Labroclypeus broadly concave anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views; anterolateral angles short but conspicuous, pointed, laterally directed, separated from lateral sides of labroclypeus by distinct incision, reflexed; canthus broadly rounded. Dorsal surface of labroclypeus punctured and setose similarly to frons, but punctures smaller and setae shorter. Underside of labroclypeus rather densely setose. Frons slightly impressed inward of eye, deeply and densely punctured; punctures large, annular, setigerous; intervening spaces much narrower than diameter of punctures. Setae erect, moderately long, ×1.5–2.0 diameter of puncture; a few longer and stiffer setae along inner margin of eye. Pronotum transverse, with anterior margin concave and bisinuate; anterolateral angles strongly protruding, acute, surface of pronotum at their base weakly impressed. Pronotum widest at middle, with sides weakly convex and smoothly converging anteriad and shallowly but distinctly concave posteriad; basal margin smoothly and regularly convex; posterolateral angles straight. Lateral margins of pronotum sharply denticulate, with 20 blunt denticles; a moderately short seta (of same length as setae on disc) present between every two crenulae. Punctures and setation of pronotum similar to those on head, but puncturation denser and intervening spaces very narrow, ridge-like, and setae adpressed. Most punctures on forebody with dense microscopical velvety pubescence. Hypomeron separated from prosternum by moderately high straight carina bearing moderately long setae; antennal pockets weak. Punctures of hypomeron much shallower and smaller than on disc of pronotum, but setae of same length, adpressed; intervening spaces variable, microreticulate and matt anteriorly and laterally, becoming smooth and glossy near forecoxae. Scutellar shield apically rounded, with moderately deep annular punctures bearing long (×2–3 diameters of corresponding puncture) adpressed setae; intervening spaces somewhat variable, distinctly microreticulate, matt. Elytra weakly widened posteriad, subtruncate apically, lacking longitudinal carinae. Punctures of elytral disc narrow, incision-shaped; intervening spaces commensurable with length of punctures, somewhat wrinkled and distinctly microreticulate, silky; setae semi-erect, ×1.5–2 length of corresponding punctures. Pygidium with weak impressions along midline and at basilateral corners; punctures large, annular, moderately shallow; intervening spaces distinctly smaller than diameter of punctures, microreticulate, matt; setae semi-erect, ×2.0–2.5 diameter of corresponding puncture, becoming slightly longer toward apex. Thorax with moderately large annular punctures becoming denser laterally, disc of metasternum impunctate along midline; intervening spaces on mesosternum distinctly microreticulate and matt, on metasternum very finely microreticulate to smooth, shining; setae adpressed, ×2.0–2.5 diameter of corresponding puncture. Abdominal ventrites densely but slightly irregularly punctured, punctures similar to those on metasternum, setae longer; intervening spaces distinctly microreticulate, silky. Ventral profile of abdomen weakly concave. Protibia (Fig. 7B) widened distally, with two longitudinal carinae dorsally and two short triangular apical teeth diverging at nearly right angle; basal tooth untraceable; protibial spur absent. Metatibial spurs pointed, straight; lower spur slightly longer and broader than upper one.Tarsomeres not elongated; three basal metatarsomeres bearing a band of brush-like setae along their ventrolateral margins (Fig. 7C). Tarsal claws with pulvilli. Aedeagus and spiculum gastrale, as on Figs. 7D–G; parameres symmetrical, each bearing a short anteriorly directed process at base; spiculum gastrale strongly bent.</p> <p>Female unknown.</p> <p>Differential diagnosis. The new species is keyed to M. hirtipes Arrow, 1941 or M. subcostatus Heller, 1895 in Telnov (2020) (the key is ambiguous for separation of these species), but aedeagus of both compared species is very different (Telnov 2020: figs. 620–622, 724–725); furthermore, the new species lacks the longitudinal carinae on elytral disc (Figs. 7A, 8). In the new species, the long brush-like setae are present on the first three metatarsomeres, while in the compared species those are confined to the first metatarsomere only. The aedeagus of the new species is most similar to that in M. brocki Telnov, 2020 and M. penicilliger, new species, but both the compared species differ in the absence of the anteriorly directed ventral process at the base of parameres, and both are very different externally, particularly in the shape of the male labroclypeus, in the pronotal sides more excavate at their bases, in the presence of the longitudinal elytral carinae and the male’s protibial spur, and in the longer setation of the body. Maechidius macrosoma, new species can be further distinguished from M. brocki in the bidentate male protibia with more divaricate apical teeth and in the presence of brush-like setae along the ventral margin of first three basal metatarsomeres, and from M. penicilliger, new species in the distinctly microreticulate (vs. glossy) elytra and in the absence of small protuberances with thickened setae on the two anteriormost abdominal ventrites. The body size of the holotype is larger than in any compared species, but a single specimen known does not allow to exclude the possible variations.</p> <p>Etymology. The species name (meaning large-bodied in Greek) reflects the larger size of the holotype and only known specimen in comparison with similar species.</p></div> 	https://treatment.plazi.org/id/038387EAFFB1AC6912F11AB3F9FEFAA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFB3AC6F12F11A0BF8BAFD2C.text	038387EAFFB3AC6F12F11A0BF8BAFD2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius penicilliger Prokofiev 2022	<div><p>Maechidius penicilliger Prokofiev, new species</p> <p>Fig. 9</p> <p>Type material. Holotype, male (Fig. 9A): D. N. Guinea / Bürgers S.G. (ZMHB).</p> <p>Description of the holotype. Total body length 7.5 mm, greatest width 4.2 mm. Dark reddish-brown, glossy; elytra slightly more reddish than the rest of body; antennae, palpi and tarsi reddish; setation pale. Antennae 9- segmented, club 3-lamellate, long (as long as funicle). Labroclypeus broadly concave anteriorly, its lateral margins strongly bisinuate in both dorsal and lateral views; anterolateral angles small and pointed, only slightly reflexed; canthus rounded, with strongly convex outer edge. Dorsal surface of head coarsely punctured, with deep annular or hexagonal punctures becoming more oblong posteriad; bottom of punctures with dense microscopical velvety pubescence; intervening spaces very narrow; setae (largely worn) very short, strongly adpressed, not surpassing diameter of corresponding puncture, except few longer semi-erect setae along inner margin of eye and anterior margin of labroclypeus. Underside of labroclypeus rather densely setose. Frons flat. Pronotum transverse, with anterior margin concave and slightly bisinuate; anterolateral angles strongly protruding, acute, surface of pronotum at their base weakly impressed. Pronotum widest at middle, with sides convex and smoothly converging anteriad and moderately deeply excavated in front of posterolateral angles; basal margin convex; posterolateral angles pronounced and somewhat attenuated, acute. Lateral margins of pronotum denticulate, with 9 broad-based tuberculate denticles before laterobasal excavation and about 5 smaller crenulations along the latter; a very short seta (often worn) present between every two tubercles. Pronotal punctures large and coarse, oval to elongate-oval but becoming annular or hexagonal close to lateral and anterior margins, with dense microscopical velvety pubescence; intervening spaces very narrow, ridge-like; setae minute, rising from anterior margin of puncture, shorter than corresponding puncture (setae largely worn and only few of them observable). Hypomeron separated from prosternum by high, sharp, slightly arcuate carina bearing moderately long setae; antennal pockets deep; punctures large, annular, with short adpressed setae not surpassing diameter of puncture; intervening spaces generally much smaller than diameter of punctures, distinctly microreticulate, matt. Scutellar shield apically pointed, with fine irregular puncturation; setae probably worn. Elytra weakly widened posteriad, subtruncate apically, with four longitudinal carinae (two between sutural joint and humeral umbo, one extending from humerus and disappearing in posterior half of elytron, and one passing along lateral margin to apical umbo) and elevated sutural margin. Punctures of elytral disc narrow, incisionshaped; intervening spaces generally smaller than length of punctures, microreticulate in part but generally glossy; setae adpressed, not or only slightly surpassing length of corresponding punctures. Pygidium flat, with large, annular, moderately shallow punctures; intervening spaces very narrow, distinctly microreticulate, matt; setae semi-erect, equal to slightly exceeding diameter of corresponding puncture. Thorax and abdominal ventrites with moderatelysized (smaller on abdomen) shallow annular punctures; intervening spaces narrow, distinctly microreticulate and matt; disc of metasternum impunctate and smooth along mid-line; setae semi-erect, those on thorax short, not or only slightly surpassing diameter of corresponding puncture, but somewhat longer on disc of metasternum; abdominal ventrites with mixed short and long setae arranged in transverse rows (setae becoming longer medially and on more posterior ventrites). Ventral profile of abdomen distinctly concave. First two abdominal ventrites with a slight protuberance at midline bearing somewhat stiffened setae (less expressed on second ventrite) (Fig. 9B). Protibia only slightly widened distally, with two longitudinal carinae dorsally and two apical teeth directed forward and laterally; basal tooth untraceable (Fig. 9C). Protibial spur present, straight. Metatibial spurs pointed, with lower spur curved and 1.5 times longer than upper one. Tarsomeres not elongated; underside of three basal metatarsomeres with a brush of long setae (Fig. 9D). Tarsal claws with pulvilli. Aedeagus and spiculum gastrale, as on Figs. 9E–H; parameres symmetrical, with tips spatulate, regularly rounded ventrally; spiculum gastrale with shaft strongly bent.</p> <p>Female unknown.</p> <p>Differential diagnosis. The new species is very similar to M. brocki Telnov, 2020, both externally and in aedeagus, differs mainly in the lateral margins of pronotum less excavate at base, in the presence of the long brushlike setae on male’s first three metatarsomeres (absent in M. brocki) and in the shape of parameres having more symmetrically angulated tips less attenuated in the lateral view (Figs. 9E–H; Telnov 2020: figs. 601–603).</p> <p>Etymology. The species is named from the Latin words “peniculus” (brush) and “gero” (to carry), due to the characteristic setation of the hind tarsomeres.</p> <p>Remarks. Although not precisely stated on the label, this species was apparently collected on Huon Peninsula, as the other specimens from the Bürgers’ samplings housed in ZMHB.</p></div> 	https://treatment.plazi.org/id/038387EAFFB3AC6F12F11A0BF8BAFD2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFB5AC6F12F11D9EFA91FBE7.text	038387EAFFB5AC6F12F11D9EFA91FBE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius seriepunctatus Moser 1920	<div><p>Maechidius seriepunctatus Moser, 1920</p> <p>Fig. 10</p> <p>Additional material examined. 1 male (Fig. 10A): “Neu-Britannien / 20. [handwritten] Ralum [printed] 11.96 [handwritten] / E. Dahl S.” (ZMHB).</p> <p>Remarks. This specimen shows no differences from the lectotype described by Telnov (2020) and personally examined by myself. I did not study the genitalia of the lectotype and Telnov (2020) gave no description or figures of them. The aedeagus of a new specimen (Figs. 10B–C) is most similar to that in M. skalei Telnov, 2020 and M. woodlarkianus Heller, 1914, differing mainly by the tips of parameres more attenuated and curved in profile (cf. Telnov 2020: figs. 712–714, 742–744); however, both the compared species are very different externally. Spiculum gastrale was missing.</p> </div>	https://treatment.plazi.org/id/038387EAFFB5AC6F12F11D9EFA91FBE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFB6AC6D12F11FC2F953FF7D.text	038387EAFFB6AC6D12F11FC2F953FF7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidius telnovi Prokofiev 2022	<div><p>Maechidius telnovi Prokofiev, new species</p> <p>Fig. 11</p> <p>Type material. Holotype, male (Fig. 11A–B): “ New Guinea / Pratt ” (BMNH).</p> <p>Description of the holotype. Total body length 8.5 mm, greatest width 4.8 mm. Black, alutaceous; legs dark reddish-brown to blackish, with tarsi and forelegs more reddish than middle and hind tibiae and femora; antennae and palpi reddish-brown; setation pale. Antennae 9-segmented, club 3-lamellate, scapus strongly inflated at distal extremity. Labroclypeus broadly concave anteriorly, its lateral margins weakly sinuous in both dorsal and lateral views; anterolateral angles short and broad, moderately protruding, reflexed at distal margin only; canthus rounded. Punctures on dorsal surface of labroclypeus and frons annular, gradually increasing in size and density from anterior margin of labroclypeus toward frons, with dense microscopical velvety pubescence; intervening spaces very narrow, convex, very finely microreticulate. Frons slightly convex. Pronotum transverse, with anterior margin concave and only slightly bisinuate, and anterolateral angles strongly protruding, acute. Pronotum widest at middle, its sides weakly convex and smoothly converging anteriad before mid-length, weakly concave behind mid-length; basal margin regularly convex; posterolateral angles obtuse. Lateral margins of pronotum distinctly crenulate, with 24 crenulae better expressed in posterior two-thirds of lateral margin; a short seta present between every two crenulae. Pronotal punctures large and deep, ovate, with dense microscopical velvety pubescence; intervening spaces narrow, convex, glossy in part, forming an allusion of wavy longitudinal ridges. Hypomeron separated from prosternum by flange-like carina bearing long and sparse setae; antennal pockets very deep. Punctures of hypomeron moderately large and not densely distributed, annular, with setae not surpassing diameter of puncture; intervening spaces variable, matt, with very indistinct microsculpture. Scutellar shield apically rounded; puncturation similar to that on pronotum, but punctures smaller and intervening spaces less convex, matt. Elytra moderately widened posteriad, broadly rounded apically, with conspicuous apical declivity; apical umbones transversely extended, not callose. Base of elytra weakly impressed close to margin. Punctures of elytra round to ovate or slightly hexagonal, deep, with dense microscopical velvety pubescence, becoming smaller toward sutural joint and side-margins; intervening spaces very narrow, convex, forming an allusion of longitudinal carinae (two of them being most expressed on each elytron between scutellar shield and humeral umbo), very finely microreticulate, matt. Pygidium flat, with dense elongate-oval puncturation bearing dense microscopical velvety pubescence and short adpressed to semierect setae slightly or not surpassing length of punctures. Thorax rather densely punctured, with punctures round, moderately-sized (about twice smaller than those on elytral disc), bearing dense microscopical velvety pubescence and very short adpressed setae, slightly or not surpassing diameter of puncture; intervening spaces slightly variable but mainly smaller than diameter of punctures, somewhat convex and finely microreticulate. Abdomen concave in profile, with last two visible ventrites somewhat bulging ventrally (Fig. 11B); all ventrites lacking protuberances or patches of setae along their middle; puncturation similar to that on thorax but setae somewhat longer, especially on posteriormost visible ventrite. Protibia widened distally, bearing a pair of closely sitting submarginal longitudinal carinae; with two poorly developed apical teeth, basal tooth and protibial spur absent (Fig. 11C). Metatibial spurs straight and of equal size, their tips not acute. All tarsomeres short, ordinary. Tarsal claws with pulvilli. Aedeagus and spiculum gastrale, as on Figs. 11D–F; parameres symmetrical, distinctly clavate at tips, with short unpaired ventral tooth at base; spiculum gastrale fan-shaped, with anterior margin weakly concave.</p> <p>Female unknown.</p> <p>Differential diagnosis. This new species appears to be most similar to M. muticus Arrow, 1941 (Fig. 12). The aedeagi in both species are similar in bauplan; however, the new species can be distinguished by the more clavate tips of the parameres and distinctly toothed ventral collar at the base of the parameres (cf. Telnov 2020: figs. 673– 675). The shape of the spiculum gastrale is also distinctive (cf. Telnov 2020: fig. 555). Externally, the new species differs from M. muticus by larger punctures on pronotum and elytra, more convex intervening spaces bearing sparse setae, matt (vs. glossy) integument, less sharpened anterolateral angles of labroclypeus, weakly bisinuate anterior margin of pronotum (vs. strongly bisinuate, clearly convex at middle in M. muticus), absence of protibial spur and abdomen strongly concave in profile (vs. almost straight).</p> <p>Orientation of intervening spaces between pronotal punctures in the new species may allude a fingerprint-like pattern postulated for M. caperatus Telnov, 2020 and M. sturnus Arrow, 1941, although the course of ridges is much less sinuous than in these species (Telnov 2020: figs. 30, 87) and no fingerprint pattern is observed on elytra. Furthermore, both aforementioned species show no other resemblances with M. telnovi, new species in external features or genitalia.</p> <p>Etymology. This species is dedicated to Dmitry Telnov (BMNH), who prepared the recent comprehensive treatment of this genus (Telnov 2020).</p></div> 	https://treatment.plazi.org/id/038387EAFFB6AC6D12F11FC2F953FF7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
038387EAFFA8AC7312F11B2AFE8AFC4B.text	038387EAFFA8AC7312F11B2AFE8AFC4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maechidiini Burmeister 1855	<div><p>Emendations to Telnov’s (2020) key to the maechidiine species from the Indo-Australian transitional zone and Bali</p> <p>4. Elytra without tracks of longitudinal carinae; intervening spaces of elytra not transversely wrinkled................................................................................................ M. paralellicollis Moser, 1920</p> <p>– Elytra with vague tracks of longitudinal carinae; intervening spaces of elytra transversely wrinkled................... 4A</p> <p>4A. Punctures of head and pronotum ovoid, with fur-like micropubescence; about 16 denticles along lateral sides of pronotum....................................................................... M. astrolabius Prokofiev, new species</p> <p>– Punctures of head and pronotum annular, lacking micropubescence; 19–25 crenulae along lateral sides of pronotum..... 4B</p> <p>4B. 19–20 crenulae along lateral sides of pronotum........................................ M. humeralis Heller, 1914</p> <p>– 24–25 crenulae along lateral sides of pronotum............................. M. acutus (Narakusumo &amp; Balke, 2019)</p> <p>***</p> <p>9. Hypomeron separated from prosternum by low straight carina................................................ 9A</p> <p>– Hypomeron almost fused to prosternum............................................... M. rugicollis Moser, 1920</p> <p>9A. Aedeagus as on Figs. 5C–D................................................ M. echinoides Prokofiev, new species</p> <p>– Aedeagus as in Telnov (2020: figs. 614–616)............................................... M. esau Heller, 1914</p> <p>***</p> <p>12. Underside of male’s basal metatarsomere with a long setae which are not brush-like arranged and rather suberect; aedeagus as in Telnov (2020: 709 – 711)........................................................... M. similis Telnov, 2020</p> <p>– Underside or margins of male’s basal metatarsomere with a brush of long (sub)erect setae.......................... 13</p> <p>13. Abdominal ventrites medially with golden (sub)erect setae; intervening spaces of elytra densely microreticulate............................................................................................ M. tarsalis Arrow, 1941</p> <p>– Abdominal ventrites without golden (sub)erect setae (small and poorly delimited patches of thickened setae can present medially on anteriormost two abdominal ventrites, then intervening spaces of elytra glossy).............................. 13A</p> <p>13A. Elytra without longitudinal carinae; intervening spaces of elytra distinctly microreticulate, forming a silky appearance; aedeagus as on Figs. 7D–E....................................................... M. macrosoma Prokofiev, new species</p> <p>– Elytra with more or less expressed longitudinal carinae; intervening spaces of elytra glossy to wrinkled in part, not silky in appearance....................................................................................... 13B</p> <p>13B. Somewhat enlarged setae in irregular longitudinal rows on elytra; ventrolateral margins of male’s first metatarsomere with a brush of very long setae (nearly as long as tarsomere) (Fig. 8A); aedeagus as in Telnov (2020: 620–622)................................................................................................. M. hirtipes Arrow, 1941</p> <p>– No enlarged setae in longitudinal row on elytra; ventrolateral margins of male’s first to third metatarsomeres with a brush of long setae (but distinctly shorter than the length of tarsomere) (Fig. 9D); aedeagus of different shape................. 13C</p> <p>13С. Longitudinal carinae on elytral disc incomplete, not extending past mid-length of elytron (Fig. 8B); no medial tubercles or patches of setae on anteriormost two abdominal ventrites; aedeagus as in Telnov (2020: 724–725)................................................................................................... M. subcostatus Heller, 1895</p> <p>– Longitudinal carinae on elytra complete; small patches of stiffened yellowish setae on weak protuberance medially on anteriormost two abdominal ventrites (Fig. 9B); aedeagus as in Figs. 9E–F......... M. penicilliger Prokofiev, new species</p> <p>***</p> <p>15. Body with solid coating of microscopical velvety pubescence; tips of parameres crossing each other........................................................................................... M. addarcis Prokofiev, new species</p> <p>– No solid coating of microscopical velvety pubescence on body; tips of parameres not crossing each other ….......... 15A</p> <p>15A. See couplet 15 in Telnov (2020).</p> <p>***</p> <p>20. Parameres slender, narrowed toward apex................................................................ 21</p> <p>– Parameres broad................................................................................... 20A</p> <p>20A. Elytral puncturation not forming the irregularly paired rows of punctures; aedeagus as in Figs. 3C–D........................................................................................ M. bombycinus Prokofiev, new species</p> <p>– Elytral puncturation forming the irregularly paired rows of punctures; aedeagus as in Telnov (2020: figs. 679–680).................................................................................. M. nepenthephilus Telnov, 2020</p> <p>***</p> <p>56. Species from Sulawesi................................................................................ 57</p> <p>– Species from New Guinea............................................................................ 56A</p> <p>56A. Body matt; abdomen strongly concave in profile; protibial spur absent in male (Fig. 11C); aedeagus as in Figs. 11D–E.............................................................................. M. telnovi Prokofiev, new species</p> <p>– Body glossy; abdomen almost straight in profile; protibial spur present in male (Fig. 12C); aedeagus as in Telnov (2020: figs. 673–675)........................................................................ M. muticus Arrow, 1941</p></div> 	https://treatment.plazi.org/id/038387EAFFA8AC7312F11B2AFE8AFC4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prokofiev, Artem M.	Prokofiev, Artem M. (2022): New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 5205 (5): 445-462, DOI: https://doi.org/10.11646/zootaxa.5205.5.3
