taxonID	type	description	language	source
C909A4C764EF59999A3119E7EB8BF7B6.taxon	description	Figs 4, 5 Suggested Common Name: Tham Than Lot Bent-toed Gecko	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
C909A4C764EF59999A3119E7EB8BF7B6.taxon	diagnosis	Diagnosis. Cyrtodactylus fluvicavus sp. nov. can be separated from all other species of the Cyrtodactylus brevipalmatus group by the combination of having 11 - 13 supralabials, 9 or 10 infralabials, 26 - 30 paravertebral tubercles, 14 - 18 rows of longitudinally arranged tubercles, 30 - 39 transverse rows of ventrals, 154 - 175 longitudinal rows of ventrals, 9 - 11 expanded subdigital lamellae on the fourth toe, 10 - 13 unexpanded subdigital lamellae on the fourth toe, 19 - 22 total subdigital lamellae on the fourth toe; 7 - 9 expanded subdigital lamellae on the fourth finger, 9 - 11 unexpanded subdigital lamellae on the fourth finger, 17 - 19 total subdigital lamellae on the fourth finger; 9 - 12 total enlarged femoral scales, 8 - 11 total femoral pores in males; 14 or 15 precloacal pores in males; 14 or 15 enlarged precloacals; enlarged femorals and enlarged precloacals not continuous; proximal femorals less than one-half the size of the distal femorals; small tubercles on forelimbs and flanks; small dorsolateral caudal tubercles and narrow ventrolateral caudal fringe; ventrolateral caudal fringe composed scales of different size; tail circular in cross-section; slightly enlarged unpaired medial subcaudals not posteromedially furrowed; maximum SVL 78.2 mm; three dark transverse body bands (Tables 4 - 6).	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
C909A4C764EF59999A3119E7EB8BF7B6.taxon	description	Description of holotype (Fig. 4). Adult male SVL 72.5 mm; head moderate in length (HL / SVL 0.28), width (HW / HL 0.70), depth (HD / HL 0.42), distinct from neck, triangular in dorsal profile; lores concave slightly anteriorly, weakly inflated posteriorly; prefrontal region concave; canthus rostralis rounded; snout elongate (ES / HL 0.42), rounded in dorsal profile; eye large (ED / HL 0.25); ear opening obliquely elongate, small; eye to ear distance greater than diameter of eye; rostral rectangular, divided by a deep furrow, bordered posteriorly by large left and right supranasals and one small azygous internasal, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two smaller postnasals, bordered ventrally by first supralabial; 12 R / 12 L rectangular supralabials, second through seventh supralabials nearly same size as first, then tapering abruptly below eye; 10 R / 10 L infralabials tapering smoothly to just below and slightly past posterior margin of eye; scales of rostrum and lores flat to domed, larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small tubercles; superciliaries subrectangular, largest dorsally; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapezoidal postmentals contacting medially for 50 % of their length posterior to mental; one row of slightly enlarged, elongate sublabials extending posteriorly to sixth (L) and fifth (R) infralabial; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales. Body relatively short (AG / SVL 0.46) with well-defined ventrolateral folds; dorsal scales small, granular interspersed with larger, conical, semi-regularly arranged, weakly keeled tubercles; tubercles extend from occipital region onto base of tail and slightly beyond as paravertebral rows; smaller tubercles extend anteriorly onto nape and occiput, diminishing in size anteriorly; approximately 17 longitudinal rows of tubercles at midbody; approximately 30 paravertebral tubercles; small tubercles on flanks; 34 longitudinal rows of flat, imbricate, ventral scales much larger than dorsal scales; 155 transverse rows of ventral scales; 15 large, pore-bearing, precloacal scales; no deep precloacal groove or depression; and two rows of post-precloacal scales on midline. Forelimbs moderate in stature, relatively short (ForL / SVL 0.14); granular scales of forearm slightly larger than those on body, interspersed with large tubercles; palmar scales rounded, slightly raised; digits well-developed, relatively short, inflected at basal interphalangeal joints; digits narrower distal to inflections; subdigital lamellae wide, transversely expanded proximal to joint inflections, narrower transverse lamellae distal to joint inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; 8 R / 8 L expanded and 10 R / 10 L unexpanded lamellae beneath the fourth finger; hind limbs larger and thicker than forelimbs, moderate in length (TibL / SVL 0.16), covered dorsally by granular scales interspersed with moderately sized, conical tubercles dorsally and posteriorly and anteriorly by flat, slightly larger, subimbricate scales; ventral scales of thigh flat, subimbricate, larger than dorsals; subtibial scales flat, imbricate; one row of 5 R / 6 L enlarged pore-bearing femoral scales not continuous with enlarged pore-bearing precloacal scales, terminating distally at knee; proximal femoral scales smaller than distal femorals, the former forming an abrupt union with much smaller, rounded, ventral scales of posteroventral margin of thigh; plantar scales flat; digits relatively long, well-developed, inflected at basal interphalangeal joints; 9 R / 9 L wide, transversely expanded subdigital lamellae on fourth toe proximal to joint inflection extending onto sole, and 11 R / 11 L unexpanded lamellae beneath the fourth toe; and claws well-developed, claw base sheathed by a dorsal and ventral scale. Tail original, 97.6 mm long (TL / SVL 1.34), 5.2 mm in width at base, tapering to a point; sub-circular or nearly round in cross-section; dorsal scales flat, square bearing tubercles forming paravertebral rows and small tubercles forming a dorsolateral longitudinal row; slightly larger, posteriorly directed, semi-spinose tubercles forming narrow but distinct ventrolateral caudal fringe; larger scales of ventrolateral fringe occur at regular intervals; medial subcaudals slightly enlarged but not paired, distinctly enlarged single medial subcaudals absent; subcaudals, larger than dorsal caudals; base of tail bearing hemipenial swellings; 3 R / 2 L conical postcloacal tubercles at base of hemipenial swellings; and postcloacal scales flat, imbricate.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
C909A4C764EF59999A3119E7EB8BF7B6.taxon	distribution	Distribution. Cyrtodactylus fluvicavus sp. nov. is currently known from the type locality at Tham Than Lot Noi-Tham Than Lot Yai Nature Trail in Chaloem Rattanakosin National Park, Si Sawat District, Kanchanaburi Province, western Thailand (Fig. 1).	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
C909A4C764EF59999A3119E7EB8BF7B6.taxon	etymology	Etymology. The specific epithet fluvicavus comes from the Latin fluvius, meaning stream, river, or flow and the Latin cavus, meaning hollow or hole and refers to a landmark cave in the Chaloem Rattanakosin National Park which has a stream that flows through it.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
7E67190E06B0564CB4C8E5A323705A73.taxon	description	Fig. 10 Suggested Common Name: Ko Chang Bent-toed Gecko	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
7E67190E06B0564CB4C8E5A323705A73.taxon	materials_examined	Additional material examined. Cyrtodactylus cf. kochangensis sp. nov. adult male THNHM 01667 from Khlong Naka Wildlife Sanctuary, Suk Samran District, Ranong Province, Thailand (~ N 9.4596, E 98.5044, elevation unknown), collected by Yodchaiy Chuaynkern between 28 December 2000 and 2 January 2001.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
7E67190E06B0564CB4C8E5A323705A73.taxon	diagnosis	Diagnosis (based on the holotype). Cyrtodactylus kochangensis sp. nov. can be separated from all other species of the Cyrtodactylus brevipalmatus group by the combination of having 12 or 13 supralabials, nine infralabials, 34 paravertebral tubercles, 14 rows of longitudinally arranged tubercles, 35 transverse rows of ventrals, 172 longitudinal rows of ventrals, 8 or 9 expanded subdigital lamellae on the fourth toe, 11 or 12 unexpanded subdigital lamellae on the fourth toe, 19 - 21 total subdigital lamellae on the fourth toe; eight expanded subdigital lamellae on the fourth finger, ten unexpanded subdigital lamellae on the fourth finger, 18 total subdigital lamellae on the fourth finger; 12 total enlarged femoral scales; 12 enlarged precloacal scales; enlarged femoral and enlarged precloacal scales not continuous; proximal femoral scales less than one-half the size of the distal femorals; small tubercles on forelimbs and flanks; large dorsolateral caudal tubercles and a wide ventrolateral caudal fringe not composed homogeneous scales; tail square in cross-section; slightly enlarged paired medial subcaudals not posteromedially furrowed; maximum SVL 60.1 mm; five dark transverse body bands (Tables 4, 5).	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
7E67190E06B0564CB4C8E5A323705A73.taxon	description	Description of holotype (Fig. 10). Adult female SVL 60.1 mm; head moderate in length (HL / SVL 0.29), width (HW / HL 0.67), depth (HD / HL 0.38), distinct from neck, triangular in dorsal profile; lores flat anteriorly, weakly inflated posteriorly; prefrontal region slightly concave; canthus rostralis rounded; snout elongate (ES / HL 0.40), rounded in dorsal profile; eye large (ED / HL 0.24); ear opening subcircular, small; eye to ear distance greater than diameter of eye; rostral rectangular, furrowed dorsally, bordered posteriorly by large left and right supranasals and one slightly smaller azygous internasal, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two slightly smaller postnasals, bordered ventrally by first supralabial; 12 R / 13 L rectangular supralabials tapering smoothly to below eye; 9 R / 9 L infralabials tapering smoothly to below eye; scales of rostrum and lores domed, slightly larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small tubercles; superciliaries subrectangular, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapazoidal postmentals contacting medially for approximately 40 % of their length posterior to mental; one row of six (R, L) slightly enlarged sublabials extending posteriorly fifth infralabials, subsequent sublabials much smaller; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales. Body relatively short (AG / SVL 0.48) with well-defined ventrolateral folds; dorsal scales small, granular, interspersed with larger conical, semi-regularly arranged, weakly keeled tubercles; tubercles extend from occipital region onto base of tail and slightly beyond as paravertebral rows; tubercles of nape and occiput smaller than those on body; approximately 14 longitudinal rows of tubercles at midbody; approximately 34 paravertebral tubercles; tubercles on upper flanks smaller than those on dorsum; approximately 35 longitudinal rows of flat, imbricate, ventral scales much larger than dorsal scales; approximately 172 transverse rows of ventral scales; no pore-bearing, precloacal scales; 12 enlarged precloacal scales; no deep precloacal groove or depression; and approximately three rows of post-precloacal scales on midline. Forelimbs moderate in stature, relatively short (ForL / SVL 0.13); granular scales of forearm slightly larger than those on body, interspersed with tubercles; palmar scales rounded, slightly raised; digits well-developed, relatively short, inflected at basal interphalangeal joints; digits narrower distal to inflections; subdigital lamellae wide, transversely expanded proximal to joint inflections, narrower transverse lamellae distal to joint inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; 8 R / 8 L expanded and 10 R / 10 L unexpanded lamellae beneath the fourth finger; hind limbs larger and thicker than forelimbs, moderate in length (TibL / SVL 0.14), covered dorsally by granular scales interspersed with moderately sized, conical tubercles dorsally and posteriorly and anteriorly by flat, slightly larger, subimbricate scales; ventral scales of thigh flat, subimbricate, larger than dorsals; subtibial scales flat, imbricate; no pore-bearing femoral scales; 6 R / 6 L enlarged femoral scales; enlarged femoral scales not contiguous with enlarged precloacal scales, terminating distally at knee; proximal femoral scales smaller than distal femorals, the former forming an abrupt union with much smaller, rounded, ventral scales of posteroventral margin of thigh; plantar scales flat; digits relatively long, well-developed, inflected at basal interphalangeal joints; 9 R / 8 L wide, transversely expanded subdigital lamellae on the fourth toe proximal to joint inflection that extend onto sole, and 12 R / 11 L unexpanded lamellae beneath the fourth toe; and claws well-developed, sheathed by a dorsal and ventral scale at base. Tail original (but in two pieces), long 80.3 mm (TL / SVL 1.34), 4.0 mm in width at base, tapering to a point; nearly square in cross-section; dorsal scales flat, square bearing tubercles forming paravertebral rows and large tubercles forming a dorsolateral longitudinal row; slightly larger, posteriorly directed, semi-spinose tubercles forming wide distinct ventrolateral caudal fringe; scales of ventrolateral fringe generally interspersed at regular intervals with larger spinose scales; medial subcaudal scales paired, slightly enlarged; subcaudals, larger than dorsal caudal scales; base of tail bearing hemipenial swellings; one conical postcloacal tubercle at base of hemipenial swellings; and postcloacal scales flat, imbricate.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
7E67190E06B0564CB4C8E5A323705A73.taxon	distribution	Distribution. Cyrtodactylus kochangensis sp. nov. is currently known only from the type locality at Ko Chang Island, Ko Phayam Subdistrict, Mueang Ranong District, Ranong Province, Thailand. The additional population of pending species status occurs in the Khlong Naka Wildlife Sanctuary, Suk Samran District Ranong Province.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
7E67190E06B0564CB4C8E5A323705A73.taxon	etymology	Etymology. The specific epithet kochangensis is in reference to the type locality, Ko Chang, Ranong Province, Thailand	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
2BBE9CD258DC5F5B991D225EF3E44746.taxon	description	Figs 7, 8 Suggested Common Name: Pa La-U Bent-toed Gecko	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
2BBE9CD258DC5F5B991D225EF3E44746.taxon	diagnosis	Diagnosis. Cyrtodactylus rivularis sp. nov. can be separated from all other species of the Cyrtodactylus brevipalmatus group by the combination of having 12 or 13 supralabials, 9 - 11 infralabials, 33 or 34 paravertebral tubercles, 18 - 20 rows of longitudinally arranged tubercles, 34 - 37 transverse rows of ventrals, 160 - 166 longitudinal rows of ventrals, nine expanded subdigital lamellae on the fourth toe, 12 or 13 unexpanded subdigital lamellae on the fourth toe, 21 or 22 total subdigital lamellae on the fourth toe; eight expanded subdigital lamellae on the fourth finger, 10 - 12 unexpanded subdigital lamellae on the fourth finger, 18 - 20 total subdigital lamellae on the fourth finger; 14 - 16 total enlarged femoral scales; 15 enlarged precloacals; enlarged femorals and enlarged precloacals not continuous, and lacking pores; proximal femorals less than one-half the size of the distal femorals; small tubercles on forelimbs and flanks; large dorsolateral caudal tubercles and wide ventrolateral caudal fringe; ventrolateral caudal fringe composed generally homogeneous scales; tail square in cross-section; single enlarged unpaired medial subcaudals not posteromedially furrowed; maximum SVL 73.9 mm; three or four dark transverse body bands (Tables 4, 5).	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
2BBE9CD258DC5F5B991D225EF3E44746.taxon	description	Description of holotype (Fig. 7). Adult female SVL 73.9 mm; head moderate in length (HL / SVL 0.27), width (HW / HL 0.73), depth (HD / HL 0.40), distinct from neck, triangular in dorsal profile; lores concave slightly anteriorly, weakly inflated posteriorly; prefrontal region concave; canthus rostralis rounded; snout elongate (ES / HL 0.41), rounded in dorsal profile; eye large (ED / HL 0.29); ear opening horizontally elongate, small; eye to ear distance greater than diameter of eye; rostral rectangular, divided dorsally by a deep furrow, bordered posteriorly by large left and right supranasals and one slightly smaller azygous internasal, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two smaller postnasals, bordered ventrally by first supralabial; 13 R / 12 L rectangular supralabials, first two largest, then tapering abruptly below eye; 11 R / 10 L infralabials tapering smoothly to just below eye and then more rapidly beyond posterior margin of eye; scales of rostrum and lores flat to domed, larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small tubercles; superciliaries subrectangular, largest anteriorly; mental triangular, bordered laterally by first infralabials, posteriorly by large left and right elongate postmentals contacting medially for approximately 40 % of their length posterior to mental; one row of two (R) and four (L) slightly enlarged sublabials extending posteriorly to third (L) and second (R) infralabial, subsequent sublabials much smaller; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales. Body relatively short (AG / SVL 0.47) with well-defined ventrolateral folds; dorsal scales small, granular, interspersed with larger conical, semi-regularly arranged, weakly keeled tubercles; tubercles extend from occipital region onto base of tail and slightly beyond as paravertebral rows; tubercles of nape and occiput small; approximately 20 longitudinal rows of tubercles at midbody; approximately 34 paravertebral tubercles; tubercles on flanks nearly same size as those on dorsum; 34 longitudinal rows of flat, imbricate, ventral scales much larger than dorsal scales; 160 transverse rows of ventral scales; no pore-bearing, precloacal scales; 15 enlarged precloacal scales; no deep precloacal groove or depression; and three rows of post-precloacal scales on midline. Forelimbs moderate in stature, relatively short (ForL / SVL 0.13); granular scales of forearm slightly larger than those on body, interspersed with tubercles; palmar scales rounded, slightly raised; digits well-developed, relatively short, inflected at basal interphalangeal joints; digits narrower distal to inflections; subdigital lamellae wide, transversely expanded proximal to joint inflections, narrower transverse lamellae distal to joint inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; 8 R / 8 L expanded and 11 R / 11 L unexpanded lamellae beneath the fourth finger; hind limbs larger and thicker than forelimbs, moderate in length (TibL / SVL 0.15), covered dorsally by granular scales interspersed with moderately sized, conical tubercles dorsally and posteriorly and anteriorly by flat, slightly larger, subimbricate scales; ventral scales of thigh flat, subimbricate, larger than dorsals; subtibial scales flat, imbricate; no pore-bearing femoral scales; 8 R / 8 L enlarged femoral scales; enlarged femoral scales not contiguous with enlarged precloacal scales, terminating distally at knee; proximal femoral scales smaller than distal femoral scales, the former forming an abrupt union with much smaller, rounded, ventral scales of posteroventral margin of thigh; plantar scales flat; digits relatively long, well-developed, inflected at basal interphalangeal joints; 9 R / 9 L wide, transversely expanded subdigital lamellae on fourth toe proximal to joint inflection that extend onto sole, and 13 R / 13 L unexpanded lamellae beneath the fourth toe; and claws well-developed, sheathed by a dorsal and ventral scale at base. Posterior one-half of tail regenerated, tail long 91.5 mm (TL / SVL 1.24), 4.8 mm in width at base, tapering to a point; nearly square in cross-section; dorsal scales flat, square bearing large tubercles forming a discontinuous dorsolateral longitudinal row; slightly larger, posteriorly directed, semi-spinose tubercles forming large distinct ventrolateral caudal fringe; scales of ventrolateral fringe generally homogeneous; single medial subcaudals enlarged but not paired; subcaudal scales, larger than dorsal caudal scales; base of tail bearing hemipenial swellings; 2 R / 2 L conical postcloacal tubercles at base of hemipenial swellings; and postcloacal scales flat, imbricate.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
2BBE9CD258DC5F5B991D225EF3E44746.taxon	distribution	Distribution. Cyrtodactylus rivularis sp. nov. is currently known from the type locality at Pa La-U Waterfall, Kaeng Krachan National Park, Huai Sat Yai Subdistrict, Hua Hin District Prachuap Khiri Khan Province, Thailand.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
2BBE9CD258DC5F5B991D225EF3E44746.taxon	etymology	Etymology. The specific epithet rivularis is derived from the Latin rivus, meaning stream, brook, or creek refers to rocky brook or stream habitat of the new species.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
333ED315A45555789D5E3A3BC20D8AEB.taxon	description	Fig. 12 Suggested Common Name: Uthai Thani Bent-toed Gecko	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
333ED315A45555789D5E3A3BC20D8AEB.taxon	diagnosis	Diagnosis. Cyrtodactylus uthaiensis sp. nov. can be separated from all other species of the Cyrtodactylus brevipalmatus group by the combination of having 13 - 15 supralabials, 10 - 11 infralabials, 33 paravertebral tubercles, 17 rows of longitudinally arranged tubercles, 36 transverse rows of ventrals, 159 longitudinal rows of ventrals, eight expanded subdigital lamellae on the fourth toe, 12 unexpanded subdigital lamellae on the fourth toe, 20 total subdigital lamellae on the fourth toe; seven expanded subdigital lamellae on the fourth finger, 11 unexpanded subdigital lamellae on the fourth finger, 18 total subdigital lamellae on the fourth finger; 16 total enlarged femoral scales, 12 total femoral pores; 14 enlarged pore-bearing precloacals; enlarged femorals and enlarged precloacals not continuous; proximal femorals less than one-half the size of the distal femorals; small tubercles on forelimbs and flanks; large dorsolateral caudal tubercles and wide ventrolateral caudal fringe; ventrolateral caudal fringe composed scales of different size; tail circular in cross-section; slightly enlarged medial subcaudals intermittent, medially furrowed, posteriorly emarginated; maximum SVL 58.1 mm; and six dark transverse body bands (Tables 5, 6).	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
333ED315A45555789D5E3A3BC20D8AEB.taxon	description	Description of holotype (Fig. 12). Adult male SVL 58.1 mm; head moderate in length (HL / SVL 0.28), width (HW / HL 0.68), depth (HD / HL 0.39), distinct from neck, triangular in dorsal profile; lores concave slightly anteriorly, weakly inflated posteriorly; prefrontal region slightly concave; canthus rostralis rounded; snout elongate (ES / HL 0.40), rounded in dorsal profile; eye large (ED / HL 0.29); ear opening elliptical, small; eye to ear distance greater than diameter of eye; rostral rectangular, dorsally furrowed, bordered posteriorly by large left and right supranasals, bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by large supranasal, posteriorly by two smaller postnasals, ventrally by first supralabial; 13 R / 15 L rectangular supralabials tapering smoothly to posterior margin of eye; 10 R / 11 L infralabials tapering smoothly to posterior margin of eye; scales of rostrum and lores flat to domed, slightly larger than granular scales on top of head and occiput; scales of occiput intermixed with distinct, small tubercles; superciliaries subrectangular, largest dorsally and anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by large left and right trapezoidal postmentals contacting medially for approximately 40 % of their length posterior to mental; one row of slightly enlarged, elongate sublabials extending posteriorly to fifth (L) and seventh (R) infralabial; gular and throat scales small, granular, grading posteriorly into slightly larger, flatter, smooth, imbricate, pectoral and ventral scales. Body relatively short (AG / SVL 0.46) with well-defined ventrolateral folds; dorsal scales small, granular, interspersed with larger, conical, semi-regularly arranged, weakly keeled tubercles; tubercles extend from occipital region onto base of tail and slightly beyond as paravertebral rows; smaller tubercles extend anteriorly onto nape and occiput, diminishing in size anteriorly; approximately 17 longitudinal rows of tubercles at midbody; approximately 33 paravertebral tubercles; small tubercles on flanks; 36 longitudinal rows of flat, imbricate, ventral scales much larger than dorsal scales; 159 transverse rows of ventral scales; 16 total large femoral scales; 12 total femoral pores; 14 enlarged pore-bearing precloacals; no deep precloacal groove or depression; and two rows of post-precloacal scales on midline. Forelimbs moderate in stature, relatively short (ForL / SVL 0.14); granular scales of forearm slightly larger than those on body, interspersed with large tubercles; palmar scales rounded, slightly raised; digits well-developed, relatively short, inflected at basal interphalangeal joints; digits narrower distal to inflections; subdigital lamellae wide, transversely expanded proximal to joint inflections, narrower transverse lamellae distal to joint inflections; claws well-developed, claw base sheathed by a dorsal and ventral scale; 7 R / 7 L expanded and 11 R / 11 L unexpanded lamellae beneath the fourth finger; hind limbs larger and thicker than forelimbs, moderate in length (TibL / SVL 0.14), covered dorsally by granular scales interspersed with moderately sized, conical tubercles dorsally and posteriorly and anteriorly by flat, slightly larger, subimbricate scales; ventral scales of thigh flat, subimbricate, larger than dorsals; subtibial scales flat, imbricate; one row of 6 R / 6 L of enlarged pore-bearing femoral scales not continuous with enlarged pore bearing precloacal scales, terminating distally at knee; 8 R / 8 L enlarged femoral scales; proximal femoral scales smaller than distal femorals, the former forming an abrupt union with much smaller, rounded, ventral scales of posteroventral margin of thigh; plantar scales flat; digits relatively long, well-developed, inflected at basal interphalangeal joints; 8 R / (broken) L wide, transversely expanded subdigital lamellae on fourth toe proximal to joint inflection that extend onto sole, 12 R / (broken) L unexpanded lamellae beneath first toe; and claws well-developed, sheathed by a dorsal and ventral scale at base. Tail original, long 76.7 mm (TL / SVL 1.32), 4.1 mm in width at base, tapering to a point; sub-circular or nearly round in cross-section; dorsal scales flat, square bearing tubercles forming paravertebral rows and large tubercles forming a dorsolateral longitudinal row; slightly larger, posteriorly directed, semi-spinose tubercles forming small but distinct ventrolateral caudal fringe; larger scales of ventrolateral fringe occur at regular intervals; slightly enlarged medial subcaudals intermittent, medially furrowed, posteriorly; single enlarged medial subcaudals absent; subcaudal scales, larger than dorsal caudal scales; base of tail bearing hemipenial swellings; 3 R / 3 L conical postcloacal tubercles at base of hemipenial swellings; and postcloacal scales flat, imbricate.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
333ED315A45555789D5E3A3BC20D8AEB.taxon	distribution	Distribution. Cyrtodactylus uthaiensis sp. nov. is currently known from the type locality at Thung Na Ngam Subdistrict, Lan Sak District, Uthai Thani Province, Thailand.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
333ED315A45555789D5E3A3BC20D8AEB.taxon	etymology	Etymology. The specific epithet uthaiensis refers to the type locality, Uthai Thani Province, Thailand.	en	Grismer, L. Lee, Aowphol, Anchalee, Yodthong, Siriporn, Ampai, Natee, Termprayoon, Korkhwan, Aksornneam, Akrachai, Rujirawan, Attapol (2022): Integrative taxonomy delimits and diagnoses cryptic arboreal species of the Cyrtodactylus brevipalmatus group (Squamata, Gekkonidae) with descriptions of four new species from Thailand. ZooKeys 1129: 109-162, DOI: http://dx.doi.org/10.3897/zookeys.1129.90535, URL: http://dx.doi.org/10.3897/zookeys.1129.90535
