taxonID	type	description	language	source
561187CDFFDC32589BD648DBFB25177F.taxon	description	(Figures 1 – 26)	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFDC32589BD648DBFB25177F.taxon	type_taxon	Type species: Formosargus kerteszi James, 1939, by monotypy.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFDC32589BD648DBFB25177F.taxon	diagnosis	Diagnosis	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFDC32589BD648DBFB25177F.taxon	description	Redescription Male. Length (mm): body, 6.0 – 6.5; wing, 5.5 – 6.0. Head (Figures 3 – 6). Typical of sargines, higher than wide and concave in dorsal view; eyes dichoptic, with very few sparse and minute setae on the eyes, and ommatidia uniform in size, except for a roughly circular area with slightly bigger ommatidia frontally. Frons, frontal callus, and face width nearly one-fifth of head or less, margins parallel-sided. Occiput black. Vertex area (from the posterior ocellus to the margin of the occiput) wide, mostly white to yellowish, except for a thin dark brown line on each side laterally; pilosity often yellowish, short to medium in length. Ocellar tubercle dark brown to black at centre, yellowish or brownish externally, reduced in size, often as wide as vertex or shorter; ocelli in the form of equilateral triangle; pilosity often yellowish, denser than in vertex, short to medium in length. Upper frons (black area between anterior ocellus and white lower frons) often wider than long or at most slightly longer and at least twice shorter than lower frons (whitish part above antenna); medial area of upper frons yellowish or brownish, at least twice as slender as the lateral area; black lateral area width varying from short to as wide as ocellar tubercle width, with short and dense yellowish to darkish pilosity; shape of anterior margin of upper frons at edge with lower frons: inverse-U-shaped (e. g. Figure 5 (a )), straight to nearly straight (Figure 6 (d ))) or U-shaped, with median triangular extension slightly (Figure 6 (e )) to strongly (Figure 5 (f )) pointing towards antennal insertion. Frontal callus (= upper part of the lower frons) not distinguishable from basal half of lower frons in colour; latter with short whitish pilosity near antenna insertion. Antennae inserted at basal third of head; scape slightly longer than pedicel; mostly yellowish to reddish brown, except for a dorsoapical arista-like terminal flagellomere often dark brown, longer than remaining antenna, with a few setae at base, but often with only two; basal four flagellomeres compact, with short, relatively dense, yellowish hair-like setae; fourth flagellomere rounded or subtriangular, as long as each previous flagellomere separately (Figure 2 (a, b )) or much longer (Figure 2 (c )). Face white to yellow, much shorter than lower frons, pilosity short, yellowish. Proboscis whitish to reddish yellow, labellum well developed; maxillary palpus short, twosegmented, distal segment enlarged and shorter than basal segment. Thorax (Figure 7). Scutum mostly pale yellow in background, but reddish in some species; with a varied pattern of dark brown markings, ranging from a single complete medial band over scutum (e. g. Figure 7 (a )), with or without additional anterior and posterior lateral bands (e. g. Figure 7 (h )), to a completely dark scutum (Figure 7 (i )); scutellum often marked with dark brown, at least in the centre. Pleuron pale yellow to reddish yellow in background, with anepisternum, katepisternum, laterotergite, and mediotergite often marked with dark brown; pilosity white to yellow. Legs varying from whitish to reddish yellow, with at least the last three to four hind tarsomeres often dark brown; virtually unmodified, except for hind leg with first tarsomere much longer than tibia, and a modified structure at apex of mid first tarsomere, often hook-like or coiled, unique for each species (Figure 8). Wing (Figure 9). Wing membrane hyaline, with brownish veins; only microtrichia over membrane, except on alula, basal area of cup and br, bm, and cua cells anteriorly. C ending at wing tip. R 2 + 3 originating slightly before or at level of r – m crossvein; basal third of R 2 + 3 often curved, apical two-thirds straight, parallel to R 1 towards wing margin. R 4 weakly curved at base, arising from R 5 with almost right angle, nearly straight on apical twothirds. R 5 slightly curved posteriorly, ending posterior to wing tip. M fading on nearly entire extension between bm and br cells, except on its extreme apical portion; distal branch of M 1 + 2 longer than proximal branch of M 1 + 2; m – m as long as short branch of M 3; M 1 and M 3 weakly developed compared to M 2, often discernible proximally or as a fold; M 4 connected to discal cell; r – m as long as R 4; m – cu crossvein connected to M 3 + 4 fork. Medial branches often not reaching wing margin. Cu vein basal to true cubital fork only setose on its basal half. Alula narrow, without microtrichia. Halter pale yellow to yellow. Abdomen. Pale yellow to reddish yellow in background, with at least a pair of roughly subrectangular dark brown spots on tergite 1, and tergites 2 – 5 mostly yellowish, with a pair of proximal or medial dark brown spots to nearly entirely dark; more than twice longer than broad, wider at about fourth segment; first segment slightly shorter than remaining segments, parallel-sided; divergent margins on segments 2 – 4, convergent margins on segment 5; pilosity short and yellowish. Terminalia (Figures 10 – 14). Genital capsule subrectangular, slightly taller than wide; distal margin weakly developed dorsally, exceeding ventral margin height; ventral margin of synsternite clearly U-shaped, often with projections medially; synsternite with a pair of well-developed inner projections, ventral to phallus, curved at base, parallel on distal two-thirds. Gonocoxal apodeme very reduced; gonocoxal bridge placed at distal margin of genitalia. Phallus trifid, with very shortened lobes; parameral sheath roughly elliptical or subrectangular, nearly entirely encapsulating the phallus; both very reduced in size, placed at distal half of genitalia. Gonostylus wide, varied in shape, with expansions or bilobed. Epandrium wider than long. Proctiger (tergite 10) well developed, as long as cercus, triangular; cercus slender, rodshaped. Female. Similar to male, except as follows: Length (mm): body, 5.0 – 10.0; wing, 5.0 – 9.0. Head (Figures 3 – 6). Frons, frontal callus and face wider than one-fifth of head. Face much shorter than lower frons or at most as long as. Thorax (Figure 7 (b, e – g )). Scutal bands or maculae over scutum often slender and less intense in colour. Abdomen. Mostly pale in colouration, varying from completely yellow to well maculated with dark brown on each visible tergite at centre or distally; spots of tergites 2 – 5 often weakly or not connected medially; clearly oval, with divergent margins on segments 1 – 2, parallel margins on segment 3, convergent margins on segments 4 – 5. Terminalia (Figures 15 – 17). Typical of sargines, mostly differing in the shape of genital fork. Tergite 8 rectangular, nearly twice longer than wide; sternite 8 wider than tergite 8, subtriangular on its distal fourth. Tergite 9 sclerite strongly chitinised laterally. Genital fork (sternite 9) often wider posteriorly, with anterior half gradually (Figure 17 (a )) or only basal third abruptly (Figure 17 (c )) narrowing into an often straight anterior end; posterior bridge from nearly straight (Figure 17 (k, l )) to weakly bilobed (e. g. Figure 17 (i, j )); posterolateral process curved at basal third, parallel on apical two-thirds; genital opening reduced, from elliptical to nearly circular-shaped, displaced posteriorly on the genital fork. Tergite 10 well developed, triangular. Cercus two-segmented; first segment of cercus nearly twice as long as second segment.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFDC32589BD648DBFB25177F.taxon	distribution	Geographic distribution Oriental Region: Taiwan (Pingtung; Orchid Island), Philippines (Benguet; Eastern Samar: Calicoan Island; Laguna; Palawan), Thailand (Trang), Malaysia (Pahang; Sabah), Indonesia (North Maluku: Island of Mangole; West Java); Australian Region: Indonesia (West Papua), Papua New Guinea (Morobe); Australia (Queensland) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFDC32589BD648DBFB25177F.taxon	discussion	Comments The Malagasy Gongrosargus Lindner, 1959 is the only other genus of Sarginae that has the M 4 vein branching off from the discal cell in the wing, with m – cu crossvein connected to the fork of M 3 + 4 (Hauser et al. 2017 b, p. 965, fig. 182). These, however, are very large flies and distinct due to the presence of more than five antennal flagellomeres (see Hauser et al. 2017 b, p. 957, fig. 130), while all remaining genera of Sarginae have only five. Therefore, this may indicate that the genus does not belong to the Sarginae, but may be instead more closely related to the Chrysochlorinae and Hermetiinae. Adisoemarto (1974) compared his new genus Amsaria with the genera Sargus and Ptecticus and pointed out it could be distinguished from the latter two by the shape of the antenna, wing veins and thorax colouration. This is in part correct. Formosargus does not have a thumb-like projection of the pedicel towards the inner face of the flagellar complex as seen in Ptecticus (e. g. Rozkošný and Kovac 1994, p. 78, fig. 16); and in some species, including that described by Adisoemarto, the fourth flagellomere alone can be as broad as the remaining flagellomeres (see his p. 70, fig. 4), a feature that excludes it from Sargus as well. In respect of the scutum colour, the presence of a single medial dark band is indeed rather unique for most species of the genus, but some Formosargus species have also additional lateral markings, bands, or a completely dark scutum. In the Oriental Region, a three-banded scutum can be seen in Ptecticus, for example in P. proximus Rozkošný and Kovac, 1996. Therefore, this feature, the colouration of the thorax, is also not a good choice for genus identification. Adisoemarto, however, was not aware of James’s description of Formosargus (James 1939), nor of his second species described in 1969, in which he stated that the genus could be distinguished by the m – cu connected to the M 3 + 4 fork and by veins M 1 and M 3 mostly fading. After examining photographs of the female holotype of Amsaria sagittocera (Figure 24) housed in the MZB collection and the illustrations in Adisoemarto (1974), it is clear to us that it belongs to Formosargus, and therefore, Amsaria is proposed as a new synonym of Formosargus. The male genitalia, illustrated for the first the time in this paper, is rather unique in the Sarginae, distinct from the genitalia of the other genera worldwide. In some Sarginae, there is a medial line, sometimes only visible as a subtle scar, over the ventral face of the synsternite, basal to the gonostylus insertion, with different degrees of sclerotisation that may be evidence of the fusion between the gonocoxites. This condition can be seen in Acrochaeta and some Merosargus, where the medial line can be further modified into a highly sclerotised projection internally (e. g. Fachin and Amorim 2015, p. 91, fig. 335). In the three genera, this medial line on the synsternite is present, although it is not well marked externally in Formosargus as it can be in Acrochaeta and Merosargus species. The medial line is, then, in Acrochaeta, Formosargus, and in some Merosargus, connected to the inner walls of the synsternite (e. g. Figure 10 (b )); Fachin and Amorim 2015, p. 19, fig. 40 and p. 90, fig. 331 )). Additionally, Formosargus shares with these two Neotropical genera the presence of an inner projection in the synsternite, but the condition is slightly different. The inner projection itself in Formosargus is rather long, parallel and well defined, sometimes with strong setae distally, and free of an innermost connection with the ventral or inner margins of the gonocoxite (Figure 11), while in some Merosargus and in some members of Acrochaeta (e. g. A. fasciata Wiedemann, 1830; see Fachin and Amorim 2015, p. 19, fig. 40), the inner projection is wide and simple, with no signal of reduction around the gonostylus, and in the other members of Acrochaeta (e. g. A. flaveola Bigot, 1879; see Fachin and Amorim 2015, p. 53, fig. 61), the inner projection is still platelike but reduced laterally and around the gonostylus. In Formosargus, both the phallus and the parameral sheath are highly reduced and displaced distally in the genitalia (see e. g. Figure 12 (a )), which may be related to the reduction of the gonocoxal apodeme as well, recognisable only by an enlarging at the centre of the gonocoxal bridge (see e. g. Figure 11 (a), see black arrow). These features have also not been found in other sargines and may be apomorphies for Formosargus as well. Another important finding is the presence of a modified structure at the base of the first tarsomere in the male mid leg (Figure 8), which is, besides the hind tibial flange of Ptecticus kovaci Hauser and Rozkošný, 2007, the only case of a modified leg within Sarginae. Males are indeed rarer than females and only James (1939, 1969) had males to examine, but he did not notice the structure in the male mid leg. In the species of which we have males, the feature is unique for each species, which indicates it may be associated with some sort of courtship.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	description	(Figures 3 (a), 5 (a – c), 7 (a – b), 8 (a – b), 15 (a), 17 (a – b), 18 (a), 21 – 22)	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	materials_examined	Type locality: Taiwan, Kankau (Koshun). Holotype: ♂ [DEI]; paratypes: 1 ♀, [DEI], 1 ♀, [WSU]; allotype: ♀ [DEI].	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	diagnosis	Diagnosis Upper frons short, wider than long; in males, anterior margin of upper frons inverse-Ushaped at the edge with lower frons (Figure 3 (a )). Scutal medial band narrower on anterior half, wider posteriorly (Figure 7 (a, b )); anepisternum with at least one dark spot. Abdomen yellowish, only males with a transverse dark brown band distally on tergite 1 (Figure 21).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	materials_examined	Material examined Type material. HOLOTYPE (Figures 7 (a), 21 (a – f )), ♂ labelled: ‘ 7. XI. [printed on white paper] ’; ‘ Kankau (Koshun) / Formosa / H. Sauter VI 1912 [printed on white paper] ’; ‘ Coll. DEI / Eberswalde [printed on white paper] ’; ‘ N. G. near / Ptecticus n. sp. [handwritten on white paper] / Det. by T. Shiraki / TAIHOKU, IV, 193 [printed on white paper] ’; ‘ Holotypus [printed on red paper] ’; ‘ Formosargus / kerteszi / ♂ [handwritten on red paper] / James [printed on red paper] ’ (DEI). PARATYPE (Figure 22 (a – d )), ♀ labelled: ‘ Kankau (Form.) / H. Sauter IX. 12 [1912] [printed on white paper] ’; ‘ Kertész det. [printed on white paper] ’; ‘ Formosargus / kerteszi / ♀ [handwritten on red paper] James / PARATYPE [printed on red paper] ’ (WSU). ALLOTYPE (Figure 22 (e – h )), ♀ labelled: ‘ Kankau (Formosa) / H. Sauter VII. 1912 [printed on white paper] ’; ‘ Formosargus / kerteszi / ♀ [handwritten on red paper] James / ALLOTYPE [printed on red paper] ’; ‘ Kertész det. [printed on white paper] ’; ‘ ALLOTYPUS [printed on red paper] ’ (DEI). Additional material. 1 ♀ (genitalia dissected and stored in microvial with glycerine), PHILIPPINES, [Eastern Samar], Calicoan PI [Calicoan Island], 10 August 1945, F. F. Bibby, no. 397, fermenting banana plant (USNM). 1 ♂, PHILIPPINES, Laguna, Calamba, Putting Lupa, 30 November 1976, kaingin, Malaise trap, UPLBMNH, DIP- 002490 (UPLB).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	description	Redescription. Female. Length (n = 1): body, 5.5 mm; wing, 5.0 mm. Head (Figures 3 (a), 5 (b, c )). Vertex area longer than ocellar tubercle, less than twice. Upper frons shortened, twice wider than long, at least twice shorter than lower frons; black lateral area nearly as wide as ocellar tubercle width, with short yellowish pilosity; upper frons anterior margin weakly convex. Fourth flagellomere rounded, as wide as each previous flagellomere alone or slightly wider. Face twice shorter than lower frons. Thorax (Figure 7 (a )). Medial dark brown band narrower on anterior part of scutum, wider on posterior half, at about twice its anterior width; one-third of scutellum dark brown; pleuron pale yellow, with only small dark brown macula on anepisternum. Tarsomeres yellowish. Wing. R 2 + 3 originating slightly before r – m, curved on basal third and nearly straight on apical two-thirds. Abdomen. Entirely pale yellow to yellow, without dark markings. Terminalia (Figures 15 (a), 17 (a, b )). Genital fork widest medially; anterior half clearly triangular; posterior bridge bilobed; posterolateral process wider on approximate basal half, narrowing into a rounded end; genital opening roughly oval.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	distribution	Geographic distribution. Oriental Region. Taiwan (Pingtung), Philippines (Eastern Samar: Calicoan Island; Laguna) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC9325E9B824982FE9A100B.taxon	discussion	Comments James’s description of Formosargus kerteszi is based on one male and three females (allotype and two paratypes). The additional male specimen herein presented from the Philippines conforms with the male holotype because both have a pair of diffuse black markings on the anepisternum, and a black mediotergite (Figure 21 (a )). The females from the same site as the male holotype and one additional female from the Philippines (Calicoan Island), however, differ from them by the presence of a single and small spot on the anespisternum and, strangely, their mediotergites are completely yellow. The two Australian species, F. lineatus and F. melanogrammus, also have a yellow mediotergite and similar scutal medial band to those of the females of F. kerteszi (see Figure 18 (a and d) for comparison), but, in contrast, both anepisterna are entirely yellow. Formosargus mangoleensis sp. nov., from Indonesia, is the only species sharing with the females of F. kerteszi one small spot on the anepisternum and a yellow mediotergite (Figure 20 (a )). It is interesting that they also have a rather similar scutal medial band, although in F. mangoleensis sp. nov. it is broader and does not reach the extreme anterior margin of the scutum (Figure 7 (g )). The fact is that these females of F. kerteszi tentatively do not fit in any other species and we do not have enough evidence to propose them as a separate species as well. Our data shows that each species has a distinct genital fork (Figure 17); however, only the female from Calicoan Island has the genital fork illustrated, so a possible confirmation of these females as a separate species from the males would demand the examination of the genital fork of the allotype and paratypes of F. kerteszi, which was not performed in this study. The holotype male genitalia is currently lost. We only have an additional male of F. kerteszi, which is without the terminal part of the abdomen including the genitalia, so it could not be illustrated and described. On the other hand, one available photograph of the male genitalia of the holotype (Figure 21 (e )) shows that F. kerteszi is similar to F. lineatus, particularly due to the presence of long inner projections of the synsternite (Figure 11 (a )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	description	(Figures 2 (a, b), 3 (b, c), 5 (d, e), 7 (c), 8 (c – e), 9 (a), 10 (a, b), 11, 15 (b), 17 (c, d), 18 (b – e), 23)	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	materials_examined	Type locality: Indonesia, Irian Jaya, Alkmaar. Holotype: ♀ [ZMAN].	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	diagnosis	Diagnosis Species very similar to F. kerteszi, but differs from it by pleuron completely yellowish in both sexes, without dark markings over anespisternum and mediotergite. Upper frons shortened as in other species, but roughly four times shorter than lower frons (Figure 5 (d, e )). Scutal band clearly more homogeneous in width, only slightly wider about at level of transverse suture (Figure 7 (c )), while in F. kerteszi, it is wider on posterior half (Figure 7 (a )). Abdomen in males only with a pair of dark brown spots on tergite 1 (Figure 18 (b )); completely yellowish in females as in F. kerteszi, while in F. melanogrammus, it has spots on tergites 2 – 5.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	materials_examined	Material examined Type material. HOLOTYPE (Figure 23 (a – c )), 1 ♀, INDONESIA, Irian Jaya [West Papua Province], Alkmaar (ZMAN). Additional material. 1 ♂ (left antenna slide-mounted; genitalia dissected and stored in microvial with glycerine), INDONESIA, [West Papua Province] Neth. Ind. - American, New Guinea Exped. [Expedition], Araucaria Camp 800 m, [- 3.5 ° S 139.183333 ° E], 09 March 1939, L. J. Toxopeus (RMNH). 1 ♀, N. Dutch New Guinea [= Indonesia], Waigeu [West Papua Province, Waigeo], Camp. Nok., 2500 ft. [feet], April 1938, L. E. Cheesman, B. M. 1938 – 593 (NHMUK). 2 ♀ (1 ♀ with left antenna and wing slide-mounted, genitalia dissected and stored in microvial with glycerine), PAPUA NEW GUINEA, Bulolo [Morobe Province], Rifle range LA, 23 February 1983, sp. no. 2014, Pandanus sp., H. R. Roberts, C. IE A 14858 (NHMUK). 1 ♀, Morobe Province, Tekadu, - 7.633333 ° S 146.566667 ° E, 09 February 2000, T. A Sear and Binatung, brigade MT (UCD).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	description	Redescription Male. Length (n = 1): body, 6.5 mm; wing, 6.0 mm. Head (Figure 3 (b), 5 (d )). Vertex area slightly longer than ocellar tubercle. Upper frons very reduced, twice wider than long, roughly four times shorter than lower frons; black lateral area of upper frons much narrower, short yellowish pilosity; anterior margin of upper frons inverse-U-shaped, pointing towards ocellar tubercle. Fourth flagellomere rounded, as wide as each previous flagellomere alone or slightly wider (Figure 2 (a )). Face three times shorter than lower frons. Thorax (Figure 7 (c )). Medial scutal band dark brown, homogeneous in width along extension, with a subtle enlarging at level of transverse suture; pleuron entirely yellow. Mid leg with a curled modified structure at apex of first tarsomere (Figure 8 (c – e )); tarsomeres yellowish. Wing (Figure 9 (a )). R 2 + 3 originating slightly before r – m, weakly sinuous. Abdomen. Mostly yellowish in background, with a pair of subrectangular dark brown maculae on tergite 1. Terminalia (Figures 10 (a, b), 11). Ventral margin of genitalia with a median bilobed and acuminate projection; synsternite with a pair of well-developed and long inner projections, curved at base, parallel on distal two-thirds, with strong bristles on last third, slightly exceeding distal margin of genitalia. Phallus exceeding gonocoxal bridge distally; parameral sheath as long as gonostylus, subrectangular. Gonostylus wide in lateral view, bilobed. Female. Similar to male, except as follows: Length (n = 4): body, 5.5 – 9.0 mm; wing, 5.5 – 8.0 mm. Head (Figures 3 (c), 5 (e )). Lower frons and face white. Thorax. Medial band narrower than in males (Figure 18 (d )). Abdomen. Homogeneously yellow, without dark markings on tergites. Terminalia (Figures 15 (b), 17 (c, d )). Genital fork equally wide on distal two-thirds, lateral margins projected backwards; basal third abruptly narrowing into a very thin, roughly straight and pointed anterior end; posterior bridge bilobed; posterolateral nearly parallel, with basal two-thirds enlarged, gradually narrowing into a rounded apex.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	distribution	Geographic distribution Australian Region. Indonesia (West Papua), Papua New Guinea (Morobe) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFCF32529BA84E8EFC8113BE.taxon	discussion	Comments Formosargus lineatus was originally described in Chrysochlora based on a female and only reported again by Lindner (1957) from a male from Indonesia (West Java) collected in 1939, which is here examined. The species was recently transferred to Formosargus (Lessard et al. 2020, referred to as Formosargus lineata), but images of the holotype are only now provided with the redescription of the species.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	description	(Figures 3 (d), 5 (f), 7 (d), 8 (f – h), 12, 18 (f), 19 (a), 24)	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	materials_examined	Type locality: Indonesia, West Java, Tjibodas, Mount Gede. Holotype: ♀ [MZB].	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	diagnosis	Diagnosis Vertex as broad as ocellar tubercle (Figure 3 (d )); anterior margin of upper frons with median triangular extension strongly pointing towards antennal insertion (Figure 5 (f )). Scutellum mostly dark brown (Figure 7 (d )); last three hind tarsomeres brownish (Figure 19 (a )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	materials_examined	Material examined Type material. HOLOTYPE (Figure 24 (a – e )), 1 ♀, INDONESIA, West Java, Tjibodas, Mount Gede (MZB). The holotype is without right wing, and fore and hind right legs (except for coxa and trochanter). Additional material. 1 ♂ (genitalia dissected and stored in microvial with glycerine), INDONESIA, Java [West Java], 15 km NW Bandung, 25 August 1991, Keith Snyder (LACM).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	description	Redescription Male. Length (n = 1): body, 6.0 mm; wing, 5.5 mm. Head (Figure 3 (d), 5 (f )). Vertex area nearly twice shorter than ocellar tubercle size. Upper frons nearly as long as wide, slightly shorter than lower frons; black lateral area of upper frons nearly as broad as ocellar tubercle width, with short yellowish pilosity; anterior margin of upper frons with a median triangular extension strongly pointing towards antennal insertion. Fourth flagellomere dark brown, pointed, as wide as previous flagellomeres combined. Face almost twice shorter than lower frons. Thorax (Figure 7 (d )). Scutum mostly dark, with only a pair of thin brownish anterior lateral lines lateral to medial band and yellow at level of transverse suture; only the medial band extends over transverse suture, reaching apex of scutellum; posterior scutal lateral bands fused to median posterior band, U-shaped; pleuron reddish yellow, with anepisternum and laterotergite marked with dark brown proximally, mediotergite dark brown. Last three hind tarsomeres brownish; mid leg with a hook-like modified structure at base of first tarsomere (Figure 8 (f – h )). Wing. R 2 + 3 originating at r – m level, weakly sinuous towards wing margin. Abdomen. Yellow in background, with a dark brown transverse band over distal half of tergite 1; tergites 2 – 5 with a pair of subrectangular dark brown spots medially; spots of tergite 3 reduced. Terminalia (Figure 12). Outer margin of genitalia projected; ventral margin of genitalia with a median rectangular plate-like projection; synsternite with a pair of short inner projections, gradually narrowing towards apex, no setae apically. Phallus not exceeding distal margin of gonocoxal apodeme; parameral sheath shorter than gonostylus, wider on distal half. Gonostylus bilobed, much wider on distal half. Female. Only known from the original description and from the photographs of the holotype provided here (Figure 24).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	distribution	Geographic distribution Oriental Region. Indonesia (West Java) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFC3326B9B1A4C39FC4E1770.taxon	discussion	Comments In the original description of Formosargus sagittocerus comb. nov., Adisoemarto (1974) mentioned brownish markings at least on the dorsocentral region of the scutum – likely referring to the medial band, on the base of the scutellum, and on the mediotergite. From his illustration (Adisoemarto 1974, p. 70, fig. 3) and holotype images (Figure 24), additional markings, although unclear, can be noted on the anterior and posterior lateral margins of the scutum, that may indicate a similar colour pattern to that seen in the male here presented. The LACM male is indeed darker than the female in the scutum (Figure 7 (d )) and in the medial area of the upper frons (Figure 5 (f )) as well. However, some level of sexual dimorphism is observed in the genus regarding the colour intensity of the body, in which the females are usually lighter than males in the head and thorax, often with smaller dark markings, but darker, with broader markings, in the abdomen (see Figure 20 (b and d) for comparison). The record of the male, again in Indonesia (West Java), less than 100 km west of the collection site of the female holotype (Figure 26), is additional evidence to sustain this specimen in the species. It is worth noting that the antenna of the holotype of F. sagittocerus illustrated by Adisoemarto (1974, p. 70, fig. 2) shows seven flagellomeres, but it probably has five, as in nearly all Sarginae. The conical shape of the flagellum in the illustration is the result of an enlargement of the fourth flagellomere in both sexes of some species of Formosargus. This also is present in F. trivittatus sp. nov., F. variegatus and F. woodleyi sp. nov.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFA326E9B93497DFD851503.taxon	description	(Figure 25)	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFA326E9B93497DFD851503.taxon	materials_examined	Type locality: Philippines, Palawan, Mantalingajan Range, Pinigisan, 600 m. Holotype: ♂ [UZMC], Allotype: ♀ [UZMC].	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFA326E9B93497DFD851503.taxon	diagnosis	Diagnosis Both scutal medial band and a pair of anterior lateral dark brown maculae discontinued at transverse suture; posterior part of scutum with a U-shaped dark brown macula; scutellum completely yellowish (Figure 25); anepisternum widely dark brown, laterotergite and mediotergite dark brown. All hind tarsomeres yellowish.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFA326E9B93497DFD851503.taxon	materials_examined	Material examined Type material. HOLOTYPE (Figure 25 (a – c )), ♂ labelled: ‘ Philippines, Palawan / Mantalingajan / Pinigisan 600 meter / 24 September 1961 / Noona Dan Exp. 61 – 62 [printed on white paper] ’; ‘ Caught in / malaise-traps / inside forest [printed on white paper] ’; ‘ zmuc / 00026038 [printed on white paper] ’; ‘ Formosargus / variegatus / ♂ James / HOLOTYPE [handwritten on red paper] ’ (UZMC). The holotype has the last four mid tarsomeres missing.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFA326E9B93497DFD851503.taxon	distribution	Geographic distribution Oriental Region. Philippines (Palawan) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFA326E9B93497DFD851503.taxon	discussion	Comments Formosargus variegatus has a scutum colour pattern similar to that seen in F. trivittatus sp. nov., since both species have at least three dark bands over the anterior part of the scutum, with yellow between the bands and the end of each of posterior lateral band connected to the medial band posteriorly. In F. variegatus, however, the medial band is completely interrupted at the transverse suture and the posterior lateral bands are strongly fused medially in the form of a semi-ellipse (Figure 25), while in F. trivittatus sp. nov., the medial band is narrowed at the level of the transverse suture and the posterior lateral bands are weakly connected to it, forming a triangular macula (Figure 7 (h )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	description	(Figures 1, 4 (a), 6 (a), 7 (e), 9 (b), 15 (c), 17 (e – f), 19 (b – c ))	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	diagnosis	Diagnosis (female) Dark brown pilosity on vertex area and ocellar tubercle (Figure 6 (a )); upper frons as wide as long, longer than frontal callus and lower frons combined. Scutal medial band broad, roughly as wide as one-third of scutum (Figure 7 (e )), only narrowing at level of transverse suture; mediotergite dark. Abdominal tergites marked with dark brown, a transverse proximal band on tergite 1 and subrectangular spots, narrowed medially, on tergites 2 – 5 (Figure 19 (b )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	materials_examined	Material examined HOLOTYPE, 1 ♀ (with puparium pinned beneath label), TAIWAN, Orchid Island, Chungui Bridge, 22.0 ° N 121.57 ° E, 20. iv – 01. v. 1997, V. Berezovskiy, coll. (LACM). PARATYPES, 3 ♀ (1 ♀ with left wing slide-mounted, genitalia dissected and stored in microvial with glycerine) (each puparium pinned beneath label), same data as holotype (LACM). Two females have an additional label mentioning that each adult emerged from the puparium on 21. v. 1997. Additional material. 1 ♂, TAIWAN, Orchid Island, Lanyu Weather Station, - 22.037306 ° S 121.558333 ° E [Note: male specimen not collected; only known from photograph (Figure 1)].	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	description	Description Female. Length (n = 4): body mm, 5.0 – 6.0; wing mm, 5.0. Head (Figures 4 (a), 6 (a )). Vertex area as long as ocellar tubercle, with medium dark brown pilosity. Upper frons as wide as long, slightly longer than lower frons; black lateral area of upper frons nearly as wide as ocellar tubercle width, with short dark brown pilosity; anterior margin of upper frons with median triangular extension slightly pointing towards antennal insertion. Fourth flagellomere rounded, as wide as each previous flagellomere. Face slightly shorter than lower frons. Thorax (Figure 7 (e )). Medial dark brown band over scutum broad, nearly as wide as one-third of scutum width, homogeneous in width along its entire extension, except for a narrowing at level of transverse suture; pleuron mostly yellowish, with only a small dark brown macula on anepisternum, mediotergite dark brown. Wing (Figure 9 (b )). R 2 + 3 originating at r – m level, weakly curved on basal third, nearly straight on apical two-thirds. Abdomen. Tergite 1 with a dark brown transverse band proximally; tergites 2 – 5 with a pair of dark brown subrectangular spots; only spots of tergite 4 connected medially. Terminalia (Figures 15 (c), 17 (e – f )). Genital fork equally wide on distal two-thirds, with a subtle widening at level of posterolateral process insertion; posterior, lateral and anterior margins projecting backwards; basal one-third abruptly narrowing into a wide and rounded anterior end; posterior bridge bilobed; posterolateral process wider on basal one-third, narrowing from midway to apex, nearly parallel on distal half; genital opening elliptical. Male Only known from a photograph (Figure 1).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	etymology	Etymology The specific epithet is a patronym dedicated to Vladimir V. Berezovskiy, a senior hymenopterist technician at the Entomology Research Museum in Riverside, USA, who first found the larva and identified the specimens in the LACM collection as belonging to Formosargus.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	distribution	Geographic distribution Oriental Region. Taiwan (Orchid Island) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFFE32609B824DBFFE2F109D.taxon	discussion	Comments This new species is the second species from Taiwan, but it is only found on Orchid Island, while Formosargus kerteszi is reported from the southern tip of the main island, which is the type locality, and now also in the Philippines (Figure 26). The male of F. berezovskiyi sp. nov., only known from a photograph (Figure 1), is very similar in the colour pattern to the male of F. kerteszi, with a single medial scutal band, but one important difference, also present in the females, is the shape and width of the band. In F. berezovskiyi sp. nov. it is clearly narrowed at the level of the transverse suture and wider than in other species, as much as one-third the width of the scutum (Figures 1, 7 (e )), while in F. kerteszi, the scutal band is more linear and thinner (Figures 7 (a, b )). Besides these features, females of F. berezovskiyi sp. nov. can be separated from all the other one-striped females by the presence of a black mediotergite and abdominal tergites strongly marked with dark brown (Figure 19 (b )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	description	(Figures 4 (b), 6 (b), 7 (f), 15 (d), 17 (g – h), 19 (d – e ))	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	diagnosis	Diagnosis (female) Species distinctly reddish in colour. Medial band on scutum slightly wider on posterior half (Figure 7 (f )), with a narrowing at level of the transverse suture and yellow one-third of scutum as wide as in F. berezovskiyi sp. nov., but differs from it by a pair of anterior lateral dark brown maculae over notopleuron (Figure 19 (d )). Abdominal tergites with subtle brownish markings, mostly conforming with the background colour (Figure 19 (e )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	materials_examined	Material examined HOLOTYPE, 1 ♀ (genitalia dissected and stored in microvial with glycerine), MALAYSIA, B. [British] N. [Northern] Borneo, [Sabah], Mt. [Mount] Kinabalu, Kamborangah, 7000 ft. [feet], 18 April 1929, [coll. H. M. Pendlebury] (NHMUK).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	description	Description Female. Length (n = 1): body, 10.0 mm; wing, 9.0 mm. Head (Figures 4 (b), 6 (b )). Vertex area twice longer than ocellar tubercle. Upper frons as wide as long, twice longer than lower frons; upper frons black lateral area broader than ocellar tubercle width, with short dark brownish pilosity; anterior margin of upper frons with a median triangular projection pointing towards antennal insertion. Fourth flagellomere rounded, as wide as each previous flagellomere alone or slightly wider. Face nearly as long as lower frons. Thorax (Figure 7 (f )). Medial scutal band dark brown, nearly as wide as one-third of scutum, with a subtle narrowing at level of transverse suture, wider posteriorly, weakly discontinued at base of scutellum; a subrectangular dark brown macula over notopleuron area, extending over anepisternum proximally; laterotergite reddish yellow, mediotergite with only a small dark brown mark. Basal half of hind femur dark brown. Wing. R 2 + 3 arising slightly before r – m, weakly curved towards R 1 on basal third, nearly straight on apical two-thirds. M 1 and M 3 less faint. Halter mostly reddish yellow, distal half of knob dark brown. Abdomen. Reddish, with a central transverse brownish band, nearly covering entire surface of each tergites 1 – 5 (Figure 19 (d )), conforming in colour with the background; pilosity reddish yellow. Terminalia (Figures 15 (d), 17 (g – h )). Genital fork wider medially at level of posterolateral process insertion; anterior half subtriangular, narrowing into a relatively slender and rounded anterior end; posterior bridge weakly bilobed; posterolateral process enlarged and rounded at base, much slenderer on next one-third, enlarging on last third towards apex, distal half nearly parallel; genital opening reduced, roughly oval. Male Unknown.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	etymology	Etymology The specific epithet, borneensis, refers to the collection site of the holotype of the new species, the island of Borneo.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	distribution	Geographic distribution Oriental Region. Malaysia (Sabah) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF132619BF84917FDEE1501.taxon	discussion	Comments This species is part of a group of species, including F. kerteszi, F. lineatus, F. berezovskiyi sp. nov., F. mangoleensis sp. nov. and F. melanogrammus, that share the presence of a single medial dark brown band on the scutum (see Figure 7 (a – c, e, g )) and a rounded fourth flagellomere (see Figure 2 (a, b )). Formosargus borneensis sp. nov., however, has a pair of additional subrectangular dark markings laterally on the anterior part of the scutum, mostly over the notopleuron (Figure 19 (e); see couplets 1 and 2 in the key). This feature indeed has similarities to the condition seen in the three-striped species, for example to that of F. trivittatus sp. nov. (Figure 20 (c )). The difference is that in F. trivittatus sp. nov. and in the other species related to it, the lateral macula is not only over the notopleuron but also extends over the anterior part of the scutum (see Figure 7 (h )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	description	(Figures 4 (c), 6 (c), 7 (g), 16 (a), 17 (i – j), 19 (f), 20 (a ))	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	diagnosis	Diagnosis (female) Medial band on scutum narrow, not reaching the extreme anterior end of scutum (Figure 7 (g )), wider posteriorly. Vertex at least twice the length of the ocellar tubercle (Figure 4 (c )); upper frons shortened, nearly three times shorter than lower frons (Figure 6 (c )). Mediotergite completely yellow; anepisternum with a small subrectangular dark spot (Figure 20 (a )). Abdomen entirely pale yellow.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	materials_examined	Material examined HOLOTYPE, 1 ♀ (genitalia dissected and stored in microvial with glycerine), INDONESIA, [North Maluku Province], Sula Isl. [Island] Mangole, 2 km W. Mandafuhi Camp, Mal. [Malaise] trap 19, c 70 m, 07 – 24 March 1995, C. v. Achterberg and Y. Yasir (RMNH). PARATYPE, 1 ♀, INDONESIA, [North Maluku Province], Mangole, Mandafuhi Camp, 05 March 1995, along road, C. v. Achterberg, RMNH 95 ’ (RMNH).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	description	Description Female. Length (n = 2): body, 10.0 mm; wing, 9.0 mm. Head (Figures 4 (c), 6 (c )). Vertex area twice longer than ocellar tubercle. Upper frons shortened, twice wider than long, nearly three times shorter lower frons; upper frons black lateral area nearly as broad as ocellar tubercle width, with short brownish pilosity; anterior margin nearly straight at the edge with lower frons, only medially pointing towards antennal insertion. Fourth flagellomere rounded, as wide as each previous flagellomere alone or slightly wider. Face three times shorter than lower frons. Thorax (Figure 7 (g )). Medial dark brown band covering less than one-third of scutum width, not reaching extreme anterior end of scutum, wider on posterior half and scutellum; pleuron pale yellow, with only a small subrectangular dark brown spot on anepisternum. Wing. R 2 + 3 originating slightly before r – m, weakly curved on basal third, straight on apical two-thirds. Abdomen. Completely yellow; pilosity yellow. Terminalia (Figures 16 (a), 17 (i, j )). Genital fork wider at posterior half at level of posterior bridge; anterior half gradually narrowing into a long and curved anterior end; posterior bridge bilobed; posterolateral process parallel, slightly wider at base, gradually narrowing towards a rounded apex; genital opening roughly circular. Male Unknown.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	etymology	Etymology The specific epithet, mangoleensis, refers to the collection site of the holotype of the new species, the island of Mangole, in the Sula Islands, Indonesia.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	distribution	Geographic distribution Oriental Region. Indonesia (North Maluku: Mangole Island) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF332639B8A4DBFFDF4113C.taxon	discussion	Comments This species, as mentioned, shares with the females of F. kerteszi a small dark spot on the anepisternum and mediotergite completely yellow. However, it is distinguished from F. kerteszi by a vertex area twice the length of the ocellar tubercle (Figure 6 (c )); also, in F. mangoleensis sp. nov., the upper frons is relatively shorter than that in F. kerteszi (see Figures 5 (b, c) and 6 (c) for comparison) and it has a medial band slightly wider, that does not reach the extreme apex of the anterior part of scutum (see Figures 7 (b) and 7 (g) for comparison). Besides these features, F. mangoleensis sp. nov. is so far the only species reported within the limits of Wallacea (Figure 26) – the area between Wallace’s Line and Lydekker’s Line – while F. kerteszi is found more to the north, beyond Wallace’s Line. The nearest species in geographical proximity to F. mangolensis sp. nov. is a record of F. lineatus from the Island of Waigeo, Indonesia, near the north-west boundary of Lydekker’s Line, which is in the Australian Region.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	description	(Figures 2 (c), 4 (d – e), 6 (d – e), 7 (h), 10 (c – e), 13, 16 (b), 17 (k – l), 20 (b – d ))	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	diagnosis	Diagnosis Scutum with three well-marked dark brown bands (Figure 7 (h )); medial band narrowing at level of transverse suture, not discontinued as in F. variegatus, with median macula triangular on posterior part of scutum; end of posterior lateral bands weakly or not connected to medial macula; as in F. variegatus, scutellum yellowish, at most dark brown at extreme base; anterior lateral band extending widely over anepisternum, laterotergite at centre and mediotergite dark brown. Male abdominal tergites irregularly marked with dark brown, tergite 1 with a transverse dark brown band medially, tergite 3 faintly dark brown at centre, and tergites 4 – 5 with a pair of subrectangular dark brown spots (Figure 20 (b )); in females, transverse dark brown bands over all tergites, spots medially discontinued on tergites 2 – 3 and 5 (Figure 20 (d )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	materials_examined	Material examined HOLOTYPE, 1 ♂ (left antenna slide-mounted; genitalia dissected and stored in microvial with glycerine), MALAYSIA, Malay Penins, Pahang, F. M. S., Fraser’s Hill, 4200 ft. [feet], 22 May 1932 (NHMUK). PARATYPE, 1 ♀ (genitalia dissected and stored in microvial with glycerine), THAILAND, Trang, Khao Pu-Khao Ya NP [National Park], - 7.550633 ° S 99.789483 ° E, 75 m, Malaise trap 20 – 23 January 2006, M. Sharkey leg. T 1958 (LACM).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	description	Description Male. Length (n = 1): body, 6.0 mm; wing, 6.0 mm. Head (Figures 4 (d), 6 (d )). Vertex area as long as ocellar tubercle. Upper frons almost twice wider than long, twice shorter than lower frons; upper frons black lateral area nearly as broad as ocellar tubercle width, short yellowish pilosity; anterior margin of upper frons mostly straight, with a median triangular projection weakly pointing towards antennal insertion. Fourth flagellomere dark brown, subtriangular, as wide as previous flagellomeres combined (Figure 2 (c )). Face almost three times shorter than lower frons. Thorax (Figure 7 (h )). Three longitudinal dark brown bands over scutum; medial band wider, not completely interrupted at level of transverse suture, expanding posteriorly into a triangular macula; anterior lateral band wide, extending over notopleuron and anepisternum; each posterior lateral band with its apical end connected to median posterior triangular macula; laterotergite at centre and mediotergite dark brown. Legs with basal half of hind tibia and all three last tarsomeres brown, second hind tarsomere white; mid leg with a small, pointed structure at base of first tarsomere. Wing. R 2 + 3 originating slightly beyond r – m, weakly sinuous towards wing margin. Abdomen. Reddish yellow; tergite 1 with a dark brown transverse band medially; tergite 3 with a pair of diffuse dark brown markings medially; tergites 4 – 5 with a pair of subrectangular dark brown markings. Terminalia (Figures 10 (c – e), 13). Outer margin of genitalia slightly prominent; ventral margin of genitalia with a median subrectangular projection; synsternite with a pair of short inner projections basally connected to the subrectangular projection, gradually narrowing towards apex, with regular setae on last third. Median lobes of phallus wider distally; parameral sheath shorter than gonostylus, roughly elliptical. Gonostylus roughly concave, wide in lateral view, with distal half much wider than basal half, dorsal margins projected upwards. Female. Similar to male, except as follows: Length (n = 1): body, 5.5 – 9.0 mm; wing, 5.5 – 8.0 mm. Head (Figures 4 (e), 6 (e )). Upper frons as long as lower frons. Fourth flagellomere yellowish. Thorax. Posterior lateral scutal bands not connected laterally to median posterior triangular macula. Abdomen. All abdominal tergites with transverse dark brown spots; tergite 1 with a band on proximal half; spots on tergites 2 – 3, 5 not connected medially; short darkish pilosity over dark markings. Terminalia (Figures 16 (b), 17 (k – l )). Genital fork wider at distal one-third, gradually narrowing towards a relatively wide, straight and rounded anterior end; posterior bridge nearly straight; posterolateral process curved on basal one-third, parallel on distal two-thirds, slightly pointing on last third; genital opening oval.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	etymology	Etymology The specific epithet is masculine and comes from the Latin word tri, meaning three, and vittae, for stripe or band, in reference to the three well-marked horizontal bands over the scutum of this species.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	distribution	Geographic distribution Oriental Region. Malaysia (Pahang), Thailand (Trang) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF232649BF34FB4FD7515F3.taxon	discussion	Comments Formosargus trivittatus sp. nov. is the only species of the genus reported from the continental portion of Southeast Asia, with one record from Malaysia (male) and one from Thailand (female). All the other species in the area have been found on islands (Taiwan and in the insular area of Southeast Asia).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	description	(Figures 4 (f), 6 (f), 7 (i), 8 (i – j), 9 (c), 10 (f), 14, 20 (e – f ))	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	diagnosis	Diagnosis (male) Darkest species of the genus. Scutum and abdomen widely marked with dark brown with coppery to bluish reflections (Figure 20 (e, f )). Ocellar tubercle much longer than vertex (Figure 4 (f )); frons, frontal callus, and face narrower than seen in other species, less than one-sixth of head width (Figure 6 (f )); upper frons longer than wide, as long as lower frons; anterior margin straight (Figure 6 (f )). M 1 and M 3 normally sclerotised (Figure 9 (c )).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	materials_examined	Material examined HOLOTYPE, 1 ♂ (left wing slide-mounted; genitalia dissected and stored in microvial with glycerine), PHILIPPINES, Luzon [Island] P. I., [Benguet], Zool. Park. Baguio 4500 ’, 02 – 04 January 1966, D. R. Davis (USNM).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	description	Description Male. Length (n = 1): body, 6.0 mm; wing, 6.0 mm. Head (Figure 4 (f), 6 (f )). Vertex area much shorter than ocellar tubercle. Upper frons longer than wide, nearly as long as lower frons; upper frons black lateral area width shorter than half of ocellar tubercle width, with short dark brown pilosity; anterior margin of upper frons straight at the edge with lower frons. Fourth flagellomere dark brown, pointed apically, as long as previous flagellomeres combined. Face slightly shorter than lower frons. Thorax (Figure 7 (i )). Scutum with a bluish dark macula covering almost entire surface, except for a thin line on transverse suture and postalar callus yellowish; scutellum mostly dark brown with weak copper shine, distal and lateral margins pale yellow; pleuron yellow in the background with a wide dark brown macula over anepistenum; laterotergite and mediotergite dark brown. Basal third of mid and hind femora, last four fore and mid tarsomeres, and last three hind tarsomeres dark brown; mid legs with a pointed structure at base of tibia and a long seta at base of first tarsomere (Figure 8 (i, j )). Wing (Figure 9 (c )). R 2 + 3 originating at r – m, weakly sinuous towards wing margin. M 1 and M 3 completely discernible, but weaker than M 2. Abdomen. Mostly dark, with coppery to bluish metallic reflections dorsally; yellowish only between tergites 1 and 2 medially, at centre from extreme posterior margin of tergite 2 to extreme anterior margin of tergite 4 and from extreme posterior margin of tergite 4 to basal half of tergite 5, and in each lateral corner between tergites 2 and 3, 3 and 4, and 4 and 5. Terminalia (Figures 10 (f), 14). Ventral margin of genitalia strongly U-shaped at centre, without median projection; synsternite with a pair of long and well-developed inner projections, with almost right angle at base, curved on next one-third, and straight on last third, with strong bristles apically, slightly exceeding distal margin of genitalia. Lobes of phallus enlarged on distal half; parameral sheath shorter than gonostylus, wider at base. Gonostylus wide on basal one-third, with a long spatulate expansion. Female Unknown.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	etymology	Etymology The specific epithet is a patronym honouring Norman E. Woodley, a great expert in Diptera with major contributions to the systematics and taxonomy of Stratiomyidae. He recognised this new species as belonging to the genus Formosargus and made it available to us.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	distribution	Geographic distribution Oriental Region. Philippines (Benguet) (Figure 26).	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
561187CDFFF432669BEE4DBFFE3C106C.taxon	discussion	Comments Formosargus woodleyi sp. nov. can be easily distinguished from others by a body mostly dark, nearly completely dark on the scutum, with bluish reflections over it, especially in the scutum and abdomen (Figure 20 (e, f )). Besides these features, this is the only species of the genus that has the upper frons clearly longer than wide, also slenderer compared to other males (Figure 6 (f )), also it has a much bigger ocellar tubercle, which is wider than the vertex (Figure 4 (f )), and M 1 and M 3 veins completely discernible (Figure 9 (c )). The remaining species are predominantly yellow or reddish, at most only considerably marked with black over the scutum, but never with a metallic shine.	en	Fachin, Diego Aguilar, Hauser, Martin (2022): Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym. Journal of Natural History 56 (37 - 40): 1513-1562, DOI: 10.1080/00222933.2022.2117107, URL: http://dx.doi.org/10.1080/00222933.2022.2117107
