taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318182/files/figure.png	https://doi.org/10.5281/zenodo.7318182	Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm.	Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318184/files/figure.png	https://doi.org/10.5281/zenodo.7318184	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318186/files/figure.png	https://doi.org/10.5281/zenodo.7318186	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318188/files/figure.png	https://doi.org/10.5281/zenodo.7318188	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318191/files/figure.png	https://doi.org/10.5281/zenodo.7318191	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318193/files/figure.png	https://doi.org/10.5281/zenodo.7318193	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318197/files/figure.png	https://doi.org/10.5281/zenodo.7318197	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318199/files/figure.png	https://doi.org/10.5281/zenodo.7318199	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318203/files/figure.png	https://doi.org/10.5281/zenodo.7318203	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318205/files/figure.png	https://doi.org/10.5281/zenodo.7318205	Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318207/files/figure.png	https://doi.org/10.5281/zenodo.7318207	Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318209/files/figure.png	https://doi.org/10.5281/zenodo.7318209	Figure 13. Male terminalia of Formosargus trivittatus sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	Figure 13. Male terminalia of Formosargus trivittatus sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318211/files/figure.png	https://doi.org/10.5281/zenodo.7318211	Figure 14. Male terminalia of Formosargus woodleyi sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; prct, proctiger.	Figure 14. Male terminalia of Formosargus woodleyi sp. nov., holotype. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; prct, proctiger.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318213/files/figure.png	https://doi.org/10.5281/zenodo.7318213	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318216/files/figure.png	https://doi.org/10.5281/zenodo.7318216	Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318220/files/figure.png	https://doi.org/10.5281/zenodo.7318220	Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process.	Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318222/files/figure.png	https://doi.org/10.5281/zenodo.7318222	Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal.	Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318226/files/figure.png	https://doi.org/10.5281/zenodo.7318226	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318230/files/figure.png	https://doi.org/10.5281/zenodo.7318230	Figure 21. Formosargus kerteszi, male,holotype [DEI]. (a) Habitus, lateral view. (b) Habitus, dorsal view. (c) Wing. (d) Antenna. (e) Male terminalia, ventral view. (f) Labels.	Figure 21. Formosargus kerteszi, male,holotype [DEI]. (a) Habitus, lateral view. (b) Habitus, dorsal view. (c) Wing. (d) Antenna. (e) Male terminalia, ventral view. (f) Labels.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318232/files/figure.png	https://doi.org/10.5281/zenodo.7318232	Figure 22. Formosargus kerteszi, female, paratype [WSU] and allotype [DEI]. (a–d). Paratype. (e–i). Allotype.	Figure 22. Formosargus kerteszi, female, paratype [WSU] and allotype [DEI]. (a–d). Paratype. (e–i). Allotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318234/files/figure.png	https://doi.org/10.5281/zenodo.7318234	Figure 23. Chrysochlora lineata, female, holotype [ZMAN]. (a) Habitus, dorsal view. (b) Habitus, lateral view. (c) Habitus, frontal view.	Figure 23. Chrysochlora lineata, female, holotype [ZMAN]. (a) Habitus, dorsal view. (b) Habitus, lateral view. (c) Habitus, frontal view.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318236/files/figure.png	https://doi.org/10.5281/zenodo.7318236	Figure 24. Amsaria sagittocera, female, holotype [MZB]. (a–b) Habitus, dorsal view. (c–d) Habitus, lateral view. (e) Head, frontal view.	Figure 24. Amsaria sagittocera, female, holotype [MZB]. (a–b) Habitus, dorsal view. (c–d) Habitus, lateral view. (e) Head, frontal view.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318238/files/figure.png	https://doi.org/10.5281/zenodo.7318238	Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen).	Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen).	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFDC32589BD648DBFB25177F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318186/files/figure.png	https://doi.org/10.5281/zenodo.7318186	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318230/files/figure.png	https://doi.org/10.5281/zenodo.7318230	Figure 21. Formosargus kerteszi, male,holotype [DEI]. (a) Habitus, lateral view. (b) Habitus, dorsal view. (c) Wing. (d) Antenna. (e) Male terminalia, ventral view. (f) Labels.	Figure 21. Formosargus kerteszi, male,holotype [DEI]. (a) Habitus, lateral view. (b) Habitus, dorsal view. (c) Wing. (d) Antenna. (e) Male terminalia, ventral view. (f) Labels.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318232/files/figure.png	https://doi.org/10.5281/zenodo.7318232	Figure 22. Formosargus kerteszi, female, paratype [WSU] and allotype [DEI]. (a–d). Paratype. (e–i). Allotype.	Figure 22. Formosargus kerteszi, female, paratype [WSU] and allotype [DEI]. (a–d). Paratype. (e–i). Allotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318213/files/figure.png	https://doi.org/10.5281/zenodo.7318213	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318222/files/figure.png	https://doi.org/10.5281/zenodo.7318222	Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal.	Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318220/files/figure.png	https://doi.org/10.5281/zenodo.7318220	Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process.	Figure 17. Genital fork (sternite 9) of Formosargus species. (a–b). F. kerteszi (a. dorsal view; b. ventral view). (c–d). F. lineatus (c. dorsal view; d. ventral view). (e–f) F. berezovskiyi sp. nov., holotype (e. dorsal view; f. ventral view). (g–h) F. borneensis sp. nov., holotype (g. dorsal view; h. ventral view). (i–j) F. mangoleensis sp. nov., holotype (i. dorsal view; j. ventral view). (k–l) F. trivittatus sp. nov., paratype (k. dorsal view;l. ventral view). Abbreviations: go, genital opening;pb, posterior bridge; pp, posterolateral process.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC9325E9B824982FE9A100B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318205/files/figure.png	https://doi.org/10.5281/zenodo.7318205	Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	Figure 11. Male terminalia of Formosargus lineatus. (a) Genitalia, dorsal view; arrow: reduced gonocoxal apodeme. (b) Genitalia, lateral view; arrow: inner projection of synsternite. (c) Genitalia, ventral view; arrow: medial line. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; gonst, gonostylus; ph, phallus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318184/files/figure.png	https://doi.org/10.5281/zenodo.7318184	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318191/files/figure.png	https://doi.org/10.5281/zenodo.7318191	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318222/files/figure.png	https://doi.org/10.5281/zenodo.7318222	Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal.	Figure 18. Habitus of Formosargus species. (a) F. kerteszi, female, dorsal. (b–e) F. lineatus (b–c. male, dorsal and lateral, respectively; d–e. female, dorsal and lateral, respectively). (f) F. sagittocerus, male, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318234/files/figure.png	https://doi.org/10.5281/zenodo.7318234	Figure 23. Chrysochlora lineata, female, holotype [ZMAN]. (a) Habitus, dorsal view. (b) Habitus, lateral view. (c) Habitus, frontal view.	Figure 23. Chrysochlora lineata, female, holotype [ZMAN]. (a) Habitus, dorsal view. (b) Habitus, lateral view. (c) Habitus, frontal view.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318186/files/figure.png	https://doi.org/10.5281/zenodo.7318186	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318197/files/figure.png	https://doi.org/10.5281/zenodo.7318197	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318199/files/figure.png	https://doi.org/10.5281/zenodo.7318199	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318203/files/figure.png	https://doi.org/10.5281/zenodo.7318203	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318213/files/figure.png	https://doi.org/10.5281/zenodo.7318213	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFCF32529BA84E8EFC8113BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318186/files/figure.png	https://doi.org/10.5281/zenodo.7318186	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	Figure 3. Heads of Formosargus species, dorsal view. (a) F. kerteszi, female. (b–c) F. lineatus (b. male; c. female). (d) F. sagittocerus, male.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318191/files/figure.png	https://doi.org/10.5281/zenodo.7318191	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318226/files/figure.png	https://doi.org/10.5281/zenodo.7318226	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318236/files/figure.png	https://doi.org/10.5281/zenodo.7318236	Figure 24. Amsaria sagittocera, female, holotype [MZB]. (a–b) Habitus, dorsal view. (c–d) Habitus, lateral view. (e) Head, frontal view.	Figure 24. Amsaria sagittocera, female, holotype [MZB]. (a–b) Habitus, dorsal view. (c–d) Habitus, lateral view. (e) Head, frontal view.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318197/files/figure.png	https://doi.org/10.5281/zenodo.7318197	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318207/files/figure.png	https://doi.org/10.5281/zenodo.7318207	Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	Figure 12. Male terminalia of Formosargus sagittocerus. (a) Genitalia,dorsal view;arrow:reduced gonocoxal apodeme. (b) Genitalia, lateral view. (c) Genitalia, ventral view. (d) Epandrium, proctiger and cerci, dorsal view. Abbreviations: cerc; cercus; epand, epandrium; goncx, gonocoxite; goncx brg, gonocoxal bridge; gonst, gonostylus; pm sh, parameral sheath; mps, median process of synsternite; prct, proctiger.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFC3326B9B1A4C39FC4E1770.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFA326E9B93497DFD851503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318238/files/figure.png	https://doi.org/10.5281/zenodo.7318238	Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen).	Figure 25. Formosargus variegatus, male, holotype [UZMC]. (a) Habitus, dorsal view.(b) Habitus, lateral view. Arrow points to a modified structure in the mid leg. (c) Labels (© Natural History Museum of Denmark, University of Copenhagen).	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFA326E9B93497DFD851503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFA326E9B93497DFD851503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318182/files/figure.png	https://doi.org/10.5281/zenodo.7318182	Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm.	Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318188/files/figure.png	https://doi.org/10.5281/zenodo.7318188	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318193/files/figure.png	https://doi.org/10.5281/zenodo.7318193	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318226/files/figure.png	https://doi.org/10.5281/zenodo.7318226	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318199/files/figure.png	https://doi.org/10.5281/zenodo.7318199	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318213/files/figure.png	https://doi.org/10.5281/zenodo.7318213	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFFE32609B824DBFFE2F109D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318188/files/figure.png	https://doi.org/10.5281/zenodo.7318188	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318226/files/figure.png	https://doi.org/10.5281/zenodo.7318226	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318213/files/figure.png	https://doi.org/10.5281/zenodo.7318213	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318184/files/figure.png	https://doi.org/10.5281/zenodo.7318184	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF132619BF84917FDEE1501.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318188/files/figure.png	https://doi.org/10.5281/zenodo.7318188	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318193/files/figure.png	https://doi.org/10.5281/zenodo.7318193	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318216/files/figure.png	https://doi.org/10.5281/zenodo.7318216	Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF332639B8A4DBFFDF4113C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318191/files/figure.png	https://doi.org/10.5281/zenodo.7318191	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 5. Heads of Formosargus species, frontal view. (a–c) F. kerteszi (a. male, Philippines; b. female, Allotype [DEI], Taiwan; c. female, Philippines). (d–e) F. lineatus (d. male; e. female). (f) F. sagittocerus, male. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318184/files/figure.png	https://doi.org/10.5281/zenodo.7318184	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318188/files/figure.png	https://doi.org/10.5281/zenodo.7318188	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318203/files/figure.png	https://doi.org/10.5281/zenodo.7318203	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318216/files/figure.png	https://doi.org/10.5281/zenodo.7318216	Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	Figure 16. Female terminalia of Formosargus species, dorsal view. (a) F. mangoleensis sp. nov, holotype. (b) F. trivittatus sp. nov., paratype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF232649BF34FB4FD7515F3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318188/files/figure.png	https://doi.org/10.5281/zenodo.7318188	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318228/files/figure.png	https://doi.org/10.5281/zenodo.7318228	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318193/files/figure.png	https://doi.org/10.5281/zenodo.7318193	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318199/files/figure.png	https://doi.org/10.5281/zenodo.7318199	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318195/files/figure.png	https://doi.org/10.5281/zenodo.7318195	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318197/files/figure.png	https://doi.org/10.5281/zenodo.7318197	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	Figure 8. Male mid legs of Formosargus species, male. (a–b) F. kerteszi. (c–e) F. lineatus. (f–h) F. sagittocerus. (i–j) F. woodleyi sp. nov., holotype. Arrow: modified structure at base of basitarsus. Abbreviations: fem, femur; tsm 1, first tarsomere.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318203/files/figure.png	https://doi.org/10.5281/zenodo.7318203	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	Figure 10. Male terminalia of Formosargus species (epandrium, proctiger, and cerci removed). (a–b) F. lineatus (a. dorsal view; b. lateral view). (c–e) F. trivittatus sp. nov., holotype (c. dorsal view; d. ventral view; e. lateral view). (f) F. woodleyi sp. nov., lateral view, holotype. Abbreviations: goncx apod, gonocoxal apodeme; goncx brg, gonocoxal bridge; gonst, gonostylus; ph, phallus; pm sh, parameral sheath. Scale bars: 0.1 mm.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
561187CDFFF432669BEE4DBFFE3C106C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7318240/files/figure.png	https://doi.org/10.5281/zenodo.7318240	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line.	2022-10-12	Fachin, Diego Aguilar;Hauser, Martin		Zenodo	biologists	Fachin, Diego Aguilar;Hauser, Martin			
