taxonID	type	description	language	source
5F6B87EEFF8F9C0B1CF9FDD35CE5FA43.taxon	type_taxon	Type: Retiboletus ornatipes (Peck) Binder & Bresinsky, Feddes Repert 113 (1 – 2): 37 (2002).	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0B1CF9FDD35CE5FA43.taxon	description	Description: Basidiomata small to medium-sized. Pileus subhemispherical to broadly convex, sometimes applanate; surface dry, velutinous, tomentose to matted tomentose, brown, yellow-brown, yellowish brown, grayish brown, gray, blackish gray, or black, sometimes with an olive, grayish green or blackish green tinge; margin always incurved; context yellow, pale yellow to light yellow, or white to grayish white or pallid gray to cream, unchanging or staining yellow, yellowish brown, pale brown, brown, blackish brown, or ferrugineous when injured. Hymenophore poroid, adnate or slightly depressed around apex of stipe; pores angular, grayish white, pale yellow, yellow to light yellow, unchanging or usually staining yellowish brown, brown to dark brown or pale orange when injured; tubes grayish white or yellowish to light yellow, unchanging or usually staining yellowish brown, pale brown, brown to dark brown when injured. Stipe central, subcylindric, solid, usually flexuous; surface dry, yellowish, yellow to brownish yellow or grayish white, gray, blackish yellow to blackish, always with yellow tinge downwards; surface always covered with concolorous or much darker reticulum; context yellow to bright yellow, or white to cream or grayish on the upper part, but with distinct yellow tinge downwards, unchanging or staining vivid yellow to yellowish brown or pale brown, brown to blackish brown or indistinct blue when injured; annulus absent. Basal mycelium white or yellow. Odor indistinct. Basidiospores smooth, subfusiform to ellipsoid. Cheilo- and pleurocystidia abundant, fusiform or subfusiform to fusoid-ventricose. Pileipellis subrepent to trichodermium composed of vertically arranged or interwoven thin-walled hyphae. Clamp connections absent in all tissues.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0B1CF9FDD35CE5FA43.taxon	biology_ecology	Ecology and distribution: Known from northern Central America and East Asia, associated with species of the families Fagaceae and Pinaceae.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0B1CF9FDD35CE5FA43.taxon	discussion	Notes: The genus Retiboletus can be easily distinguish from other genera in Boletaceae by the grayish black or yellow tinged pileus, the concolorous stipes covered with reticulum or sometimes granular-like squamules, the yellow to bright yellow or white to grayish or cream hymenophore, and white or yellow basal mycelium. Species of the genus Boletus share the reticulate stipe with Retiboletus, but the surface of the immature poroid hymenophore of Boletus is covered with a layer of tangled white hyphae (Cui et al. 2016). The genus Retiboletus receives high bootstrap support values as a monophyly in present and former studies. Species of this genus cluster into two major clades corresponding to two subgenera, i. e., R. subg. Nigroretiboletorum and R. subg. Retiboletus	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0A1CF9F9EC5D31FE61.taxon	type_taxon	Type: Retiboletus ornatipes (Peck) Binder & Bresinsky, Feddes Repert 113 (1 – 2): 37 (2002).	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0A1CF9F9EC5D31FE61.taxon	diagnosis	Diagnosis: Retiboletus subg. Retiboletus differs from R. subg. Nigroretiboletorum by its yellow to light yellow hymenophore and context, which is unchanging or staining yellow to yellowish brown when injured.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0A1CF9F9EC5D31FE61.taxon	description	Description: Basidiomata small to large-sized. Pileus subhemispherical to broadly convex, sometimes applanate; surface dry, velutinous to tomentose to matted tomentose, yellowish brown, gray brown to brown or yellow, sometimes with olive tinge; margin incurved; context pale yellow to light yellow, unchanging or staining yellow to yellowish brown in color when injured. Hymenophore poroid, adnate or slightly depressed around apex of stipe; pores angular pale yellow, yellow, to light yellow, usually changing yellowish brown to brown or pale orange when injured; tubes yellowish to light yellow, usually changing yellowish brown, pale brown when injured. Stipe central, subcylindric, solid, usually flexuous; surface dry, yellowish, yellow to brownish yellow, prominently and coarsely reticulate nearly to base or the upper; reticulum brownish black or yellow to light yellow; context yellow, unchanging or changing vivid yellow or yellowish brown in color when injured; annulus absent. Basal mycelium yellow. Odor indistinct. Basidiospores smooth, subfusiform to ellipsoid; Cheilo- and pleurocystidia abundant, fusiform or subfusiform; Pileipellis a trichoderm composed of vertically or interwoven thin-walled hyphae; Stipitipellis hymeniform, composed of thin-walled hyphae with narrowly or broadly clavate, subfusiform or fusiform terminal cells. Clamp connections absent in all tissues.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0A1CF9F9EC5D31FE61.taxon	biology_ecology	Ecology and distribution: Known from northern Central America and East Asia, associated with species of the family Fagaceae and the family Pinaceae.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8F9C0A1CF9F9EC5D31FE61.taxon	discussion	Currently known species: Retiboletus brevibasidiatus Raspé & Chuankid (2021: 297), R. flavoniger (Halling, G. M. Muell. & L. D. Gómez) Binder & Halling (2002: 30), R. kauffmanii, R. ornatipes, R. retipes, R. sinensis N. K. Zeng & Zhu L. Yang (2016: 363).	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C0A1CF9FDC25C3AFA54.taxon	etymology	Etymology: the epithet refers to species of this subgenus with gray, grayish-brown or black colored pileus.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C0A1CF9FDC25C3AFA54.taxon	type_taxon	Type: Retiboletus griseus (Frost in Peck) Binder & Bresinsky, Feddes Repert 113 (1 – 2): 37 (2002).	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C0A1CF9FDC25C3AFA54.taxon	diagnosis	Diagnosis: Retiboletus subg. Nigroretiboletorum differs from R. subg. Retiboletus by its white, grayish white or pallid gray to cream context, unchanging or staining pale brown, brown, blackish brown, or ferrugineous when injured, and the white to grayish white or grayish pink to dark purple hymenophore, which is unchanging or staining pale brownish, brown, dark brown or ferrugineous when injured.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C0A1CF9FDC25C3AFA54.taxon	description	Description: Basidiomata small to medium-sized. Pileus subhemispherical to convex, sometimes applanate; surface dry, densely subtomentose, grayish brown, pale brown to blackish brown or grayish white to blackish; margin incurved; context white, grayish white or pallid gray to cream, unchanging or staining pale brown, brown, blackish brown, or ferrugineous when injured. Hymenophore poroid, adnate or slightly depressed around apex of stipe; pores angular to roundish, white to grayish white or grayish pink to dark purple, unchanging or staining brownish, brown, dark brown or ferrugineous when injured; tubes grayish white or gray-white when young, then gray, grayish yellow, or lilac, unchanging or staining pale brown, brown, or dark brown when injured. Stipe central, subcylindrical, solid, usually flexuous; surface dry, grayish white, gray to blackish, but with yellow to gray-yellow tinge downwards; surface often covered with concolorous or much darker reticulum, sometimes without indistinct reticulum; context white, cream to grayish on the upper part, but yellowish to cream downwards, unchanging or staining pale brown, brown, blackish brown, ferrugineous or indistinct blue when injured; annulus absent. Basal mycelium white or cream. Odor indistinct. Basidiospores smooth, subfusiform to ellipsoid. Cheilo- and pleurocystidia subfusiform or fusiform, abundant. Pileipellis a trichoderm composed of more or less vertically arranged, or sometimes slightly interwoven hyphae. Clamp connections absent in all tissues.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C0A1CF9FDC25C3AFA54.taxon	biology_ecology	Ecology and distribution: Known from North and Central America and East Asia, associated with plants of the families Fagaceae and Pinaceae.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C0A1CF9FDC25C3AFA54.taxon	discussion	Currently known species: Retiboletus ater, R. brunneolus, R. fuscus, R. griseus, R. nigerrimus (R. Heim) Manfr. Binder & Bresinsky, R. nigrogriseus, R. pseudogriseus, R. sinogriseus, R. vinaceipes B. Ortiz, Lodge & T. J. Baroni, R. zhangfeii and the newly described species R. cyanescens.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C081CF9F9DE5C72FF65.taxon	etymology	Etymology: cyanescens referring to the context in stipe staining indistinct blue when injured.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C081CF9F9DE5C72FF65.taxon	materials_examined	Type: China. Yunnan Province: Anning County, alt. 2050 m, 15 July 2021, Y. C. Li 2967 (holotype: KUN-HKAS 122939).	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C081CF9F9DE5C72FF65.taxon	diagnosis	Diagnosis: Retiboletus cyanescens can be distinguished from the other species in Retiboletus by the combination of the gray to blackish pileus, the white, grayish white to yellowish gray stipe which is yellowish to brownish yellow downwards, the entirely reticulate stipe surface, the cream basal mycelium, the bluish discoloration in the middle part of the stipe when injured, the palisadoderm pileipellis, and the subtropical to temperate distribution.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C081CF9F9DE5C72FF65.taxon	description	Description: Basidiomata small to medium-sized. Pileus 3 – 10 cm in diameter, subhemispherical to applanate, surface dry, finely subtomentose, dark brown (5 F 8), brown (5 E 8), black (4 F 2), or brown-gray (4 C 2) in the center and brownish (5 B 4 - 5) or grayish (4 B 2) towards margin. Context whitish (4 A 1) to cream (4 A 3), without color change when injured. Hymenophore adnate to slightly decurrent, or sometimes depressed around apex of stipe when mature; surface initially whitish (4 A 1) and then dirty white or grayish (4 B 1); pores angular to roundish, 0.5 – 1.5 mm wide; tubes up to 4 – 6 mm in length, concolorous or a little paler than hymenophoral surface, without color change when injured or sometimes with rust brownish spots when touched. Stipe 3 – 7 × 0.3 – 0.7 cm, clavate to subcylindrical, flexuous, solid; surface grayish white (4 B 2) to pale range (5 A 3), covered with blackish to dark brown reticulum, without color change when hurt; context white (4 A 1) to cream (4 A 3) at upper part and cream to yellowish (4 A 2) downwards, staining indistinct blue in the middle part when injured; basal mycelium cream (4 A 3) without color change when injured. Taste and odor mild. Basidia 28 – 40 × 9 – 12 μm, clavate, thin-walled, 4 - spored, hyaline to yellowish in KOH. Basidiospores [100 / 5 / 5] (9.5) 10 – 13 (14.5) × 4 – 5 μm, [Q = (2.11) 2.30 – 3.13 (3.25), Q m = 2.60 ± 0.18], sub- fusiform and inequilateral with slightly suprahilar depression in side view, elongate fusoid or narrowly oblong in ventral view, slightly thick-walled (up to 0.5 μm), brownish or pale brown in KOH, olive-brown to brown in Melzer’s reagent, smooth. Hymenophoral trama boletoid; hyphae cylindrical, 3 – 14 μm wide, hyaline to yellowish in KOH, yellowish to yellow in Melzer’s reagent. Cheilocystidia 27 – 37 × 5 – 7 μm, fusiform or subfusiform, thin-walled, brownish to pale yellow-brown in KOH; surface without encrustations. Pleurocystidia relatively big, 44 – 65 × 6 – 10 μm, fusiform or subfusiform, thin- to slightly thickwalled (up to 0.5 μm), concolorous with cheilocystidia; surface without encrustations. Caulocystidia forming the reticulum over the stipe surface, morphologically similar to cheilo- and pleurocystidia. Pileipellis a trichoderm about 150 μm thick, composed of more or less vertically arranged to slightly interwoven, brown to dark brown hyphae, 6 – 16 μm wide; terminal cells 21 – 38 × 6 – 16 μm, narrowly clavate to subcylindrical or subfusiform, sometimes narrowly mucronate, rostrate, thin-walled, yellowish brown to brownish in KOH, yellow-brown to brown in Melzer’s reagent. Pileal trama composed of thin-walled hyphae, 3 – 9 μm wide, hyaline to yellowish in KOH, yellowish to yellow in Melzer’s reagent. Clamp connections absent in all tissues.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C081CF9F9DE5C72FF65.taxon	biology_ecology	Habitat, ecology and distribution: Solitary on the ground in subtropical to temperate forests dominated by plants of the family Fagaceae; currently known from southwestern China.	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
5F6B87EEFF8E9C081CF9F9DE5C72FF65.taxon	materials_examined	Additional specimens examined: China. Yunnan Province: Kunming City, Anning County, alt. 2050 m, 15 July 2021, Y. C. Li 2975 and Y. C. Li 2980 (KUN-HKAS 122940 and KUN-HKAS 122941, respectively); Lijiang County, Wenbi Mountain, alt. 2400 m, 14 July 2019, C. Huang 013 (KUN-HKAS 106692); Kunming City, Kunming Institute of Botany, alt. 1980 m, Z. L. Yang 5698 (KUN-HKAS 79637).	en	Li, Jin, Wang, Zhen, Liu, En-De, Yang, Zhu L., Li, Yan-Chun (2022): Morphological and molecular data reveal Retiboletus cyanescens sp. nov. and the new subgenus Nigroretiboletorum (Boletaceae). Phytotaxa 572 (3): 232-242, DOI: 10.11646/phytotaxa.572.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.572.3.2
