taxonID	type	description	language	source
7D518786FFA47118FF7026ED7F2AF2E2.taxon	description	(Figs 1 – 4) http: // zoobank. org / urn: lsid: zoobank. org: act: 7316 C 9 ED-A 5 DA- 4843 - 9 EC 3 - 354 D 88 F 9 C 5 AD	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA47118FF7026ED7F2AF2E2.taxon	materials_examined	Type material. Holotype, female: VIETNAM, Ninh Binh Province, Cuc Phuong National Park, 390 m, 20 ° 21 ′ 03 ″ N 105 ° 35 ′ 36 ″ E (12 - 09), SD Gaimari, M Hauser, Pham HT, 24 – 28. III. 2012, ex yellow pan trap (CDFA). Paratype, female, as holotype except: MSUC _ ARC _ 314157 (1, MSUC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA47118FF7026ED7F2AF2E2.taxon	diagnosis	Diagnosis. This species is distinguished by the unique elytral sculpturing in which the striae are deeply impressed on the elytral disc and declivity forming deep strial furrows and interstrial ridges, and elytral disc with a weak transverse saddle-like depression. Similar species. Anisandrus sinivali Smith, Beaver & Cognato, 2020 a, A. eggersi (Beeson, 1930)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA47118FF7026ED7F2AF2E2.taxon	description	Description (female). 3.55 mm long (mean = 3.55 mm; n = 2); 2.22 × as long as wide. Body black, legs and antennae dark brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, reticulate, moderately punctate; punctures each bearing a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, slightly impressed. Antennal scape long and slender, longer than club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face. Pronotum 0.94 × as long as wide. In dorsal view rounded, type 1, sides convex, rounded anteriorly; anterior margin with a row of five large serrations. In lateral view short and tall, type 3, disc as long as anterior slope, summit at midpoint. Anterior slope with densely spaced, large coarse asperities, becoming lower and more strongly transverse towards summit. Disc subshiny, impunctate, feebly asperate, basal and lateral areas densely finely punctate, each puncture bearing moderate, erect, hair-like setae, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Mycangial tuft present along basal margin, tuft densely setose, approximately the width of scutellum. Scutellum broad, large, linguiform, flush with elytra, flat, shiny. Elytra: 1.28 × as long as wide, 1.37 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 1 / 2, then broadly rounded to apex; surface shining. Disc with a weak medial transverse saddle-like depression, striae strongly impressed from basal fifth, with large, deep subcontiguous punctures, glabrous; interstriae costate, punctate, punctures uniseriate, setose, setae 2 – 3 × width of interstriae 2, erect hair-like. Declivity occupying ~ 2 / 5 elytra, evenly rounded, declivital face appearing bisulcate, deeply impressed between striae 1 and 2; striae deeply impressed, strial punctures larger and deeper than those of disc; interstriae impunctate, tuberculate and setose along interstriae 1, 3, 4, setae 2 – 4 × width of interstriae 2, erect, hair-like, interstriae 2 narrower than interstriae 3 at midpoint of declivity, declivital interstriae 1 – 3 armed with a blunt tubercle at summit, interstriae 3 armed by one or two denticles near declivital summit ventrad to tubercle on interstriae 2. Posterolateral margin costate to interstriae 5, unarmed by granules. Legs: procoxae contiguous. Protibiae obliquely triangular, broadest at apical 1 / 3; posterior face smooth; apical 1 / 2 of outer margin with seven or eight very large socketed denticles, their length much longer than basal width. Meso- and metatibiae flattened; outer margins evenly rounded with nine or ten and ten or 11 large socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA47118FF7026ED7F2AF2E2.taxon	distribution	Distribution. Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA47118FF7026ED7F2AF2E2.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA47118FF7026ED7F2AF2E2.taxon	etymology	Etymology. L. proscissus = plowed up. In reference to the deep furrows on elytral declivity. A participle of the verb proscindo.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	description	(Figs 5 – 8) http: // zoobank. org / urn: lsid: zoobank. org: act: 9 DDB 74 AA- 4 C 67 - 47 E 9 - A 190 - 34 B 555962 B 28	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	materials_examined	Type material. Holotype, female: NEPAL, Ghanpokhara, 5,500 – 7,000 ft, 2. v. 1954, J. Quinlan, NHMUK 014591805 (NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	diagnosis	Diagnosis. This species can be distinguished by the mesonotal mycangial tuft the length of the scutellum; elytral disc flat; declivital interstriae punctate; posterolateral margin feebly carinate to interstriae 5; declivity feebly bisulcate, margins ornamented by a minute tubercle at the base of interstriae 2 and three minute tubercles on the basal quarter of interstriae 3; and declivital interstriae bearing fine erect hair-like setae. Similar species. Anisandrus feronia Smith, Beaver & Cognato, 2020 a, A. mussooriensis (Eggers, 1930)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	description	Description (female). 3.25 mm long (n = 1); 2.41 × as long as wide. Body bicolored: pronotum red, elytra dark brown, legs and antennae yellow-brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, alutaceous, subshiny, punctate; punctures small, shallow, setose; punctures each bearing a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, slightly impressed. Antennal scape long and slender, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 as long as pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face. Pronotum 0.96 × as long as wide. In dorsal view rounded, type 1, sides convex, rounded anteriorly; anterior margin with a row of five serrations. In lateral view short and tall, type 3, disc shorter than anterior slope, summit at basal 2 / 5. Anterior slope with densely spaced, large coarse asperities, becoming lower and more strongly transverse towards summit. Disc subshiny with dense, fine punctures bearing moderate, semi-erect hair-like setae, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded. Mycangial tuft present along basal margin, tuft moderately setose, approximately the width of scutellum. Scutellum broad, large, linguiform, flush with elytra, flat, shiny. Elytra: 1.44 × as long as wide, 1.50 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 3 / 4, then narrowly rounded to apex; surface opalescent. Disc weakly convex, striae not impressed, with small, deep subcontiguous punctures, setose, setae as long as two punctures, recumbent, hair-like; interstriae flat, unarmed by granules, punctate, punctures strongly confused, setose, setae 1 – 1.5 × width of interstriae 2, erect hair-like. Declivity occupying ~ 3 / 8 elytra, steeply rounded, declivital face feebly bisulcate, interstriae 1 and 3 convex, depressed between striae 1 and 2; striae not impressed, strial punctures much larger and deeper than those of disc, and bearing setae as described for disc; interstriae minutely uniseriate punctate, setae 1 – 2 × width of interstriae 2, erect, hair-like, interstriae 2 about as wide as interstriae 3 at midpoint of declivity, declivital margins ornamented by a minute tubercle at the base of interstriae 2 and three minute tubercles on the basal quarter of interstriae 3. Posterolateral margin carinate to interstriae 5. Legs: procoxae contiguous. Protibiae obliquely triangular, broadest at apical 1 / 3; posterior face smooth; apical 1 / 2 of outer margin with six very large socketed denticles, their length much longer than basal width. Meso- and metatibiae flattened; outer margins evenly rounded with nine very large socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	distribution	Distribution. Nepal.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA4711EFF70213D7FC8F489.taxon	etymology	Etymology. L. simplex = simple. In reference to the unadorned elytral declivity, which is atypical for Anisandrus. An invariable adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	description	(Figs 9 – 12) http: // zoobank. org / urn: lsid: zoobank. org: act: 061 FF 3 E 7 - 6 B 54 - 44 BF- 91 E 1 - A 961 A 95 A 6 E 7 E	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	materials_examined	Type material. Holotype, female: BRUNEI [DARUSSALAM], Kuala Belalong FSC, E 116.7 ', N 4.34 ', Dipterocarp forest, Dipterocarpus beccarii, Aerial FIT 4, 270 m alt., 24. v. [19] 91, N. Mawdsley, NM 169, NHMUK 014591833 (NHMUK). Paratypes, female, as holotype except: Aerial FIT 2, 270 m alt., 4. vii. [19] 91, NM 222, NHMUK 01838 (1, NHMUK); as previous except: Aerial FIT 4, 270 m alt., 17. vii. [19] 91, NM 242 (1, RABC); as previous except: Aerial FIT 5, 255 m alt., 7. vi. [19] 91, NM 179, MSUC _ ARC _ 314153 (1, MSUC); as previous except: 18. vii. [19] 91, NM 240, NHMUK 01840 (1, NHMUK); as previous except: Aerial Malaise 6, 275 m alt., 3. vii. [19] 91, NM 213 (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	diagnosis	Diagnosis. This species is distinguished by the protibiae posterior faces inflated, granulate; antennal club wider than long; pronotum lateral margin oblique; pronotum anterior margin without serrations; posterolateral carina acute, carinate; declivital interstriae 1 strongly elevated on apical half, denticulate on apical half; and discal interstriae 1 – 6 ending in small triangular teeth projecting over the declivity. The species is closely related to A. suturalis (Eggers, 1936), but can be distinguished by the whole of the elytral disc and declivity shining, not matte posteriorly; the posterior part of the elytral disc with the striae more deeply grooved, and interstriae 1 – 6 ending in small triangular teeth projecting over the declivity; sutural interstriae without denticles on upper half of declivity, remaining declivital interstriae without granules and long hairs. Similar species. Arixyleborus suturalis, A. minor (Eggers, 1940)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	description	Description (female). 1.5 – 1.7 mm long (mean = 1.6 mm; n = 4); 2.5 – 2.91 × as long as wide. Pronotum dark brown, legs and antennae yellow-brown, elytra bicolored: basal half of disc brown, posterior half of disc and declivity reddish brown. Head: epistoma entire, transverse, lined with a row of hair-like setae. Frons slightly convex from epistoma to upper level of eyes; surface alutaceous, dull, impunctate; punctures above epistoma large, coarse. Eyes deeply emarginated above level of antennal insertion, upper portion of eyes smaller than lower part. Submentum flat, deeply impressed below genae, shaped as a large triangle. Antennal scape short and thick, about as long as club. Antennal funicle 4 - segmented, segments equal in size. Pedicel as long as funicle. Club wider than long, obliquely truncate, type 2 with segment 1 covering basal half of anterior face and most of posterior face, its margin completely costate, segment 2 narrow, pubescent with corneous part, visible on anterior face only. Pronotum 1.17 – 1.33 × as long as wide. In dorsal view very elongate and rounded frontally, type 9, sides parallel on basal 2 / 3; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 8, disc flat, summit at apical 1 / 3. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, alutaceous, sparsely finely punctate. Lateral margins obliquely costate. Base transverse, posterior angles strongly oblique. Scutellum moderately sized, ‘ u’ shaped, flat, flush with elytra. Elytra: 1.42 × as long as wide, 1.33 – 1.48 × as long as pronotum, shining. Base feebly concave, humeral angles rounded. Sides straight from base to apical 3 / 4; lateral profile of declivity obliquely truncate; apex angulate, entire. Disc occupying 3 / 4 of declivity; anterior half of disc with interstriae smooth, nearly glabrous, finely, sparsely punctate; posterior half with striae more deeply grooved, and interstriae 1 – 6 ending in small triangular teeth projecting over the declivity. Declivital interstriae 1 strongly elevated on apical half, denticulate; interstriae impunctate; strial punctures very large, glabrous; dorso- and lateral margins ornamented with long incurved setae arising from each tooth projecting over the declivity and on the interstriae. Posterolateral declivital margin acute, carinate, laterally extending to interstriae 7. Legs: Procoxae contiguous. Protibiae slender, broadest at apical third, posterior face inflated, tuberculate; five large socketed denticles present on outer margin of apical half. Meso- and metatibiae with evenly rounded outer margins, flattened, posterior faces unarmed, seven small socketed denticles present on outer margin.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	distribution	Distribution. Brunei Darussalam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA2711EFF7027147A23F381.taxon	etymology	Etymology. In reference to the collection locality, Kuala Belalong. An adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	description	(Figs 13 – 16) http: // zoobank. org / urn: lsid: zoobank. org: act: 9 AA 798 BE- 1 BE 1 - 4 F 9 D- 9439 - 7 BD 1 A 42 AF 277	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	materials_examined	Type material. Holotype, female: INDONESIA, Sulawesi Utara, Dumoga-Bone N. P., Plot A, ca 200 m, lowland forest, flight intercept trap, NHMUK 014591832 (NHMUK). Paratypes, female, as holotype: (1, RABC), MSUC _ ARC _ 314134, MSUC _ ARC _ 314135 (2, MSUC); as previous except: NHMUK 014591841 (1, NHMUK); as previous except: Rothamsted light trap, site 2, 220 m, H. Barlow, February 1985, NHMUK 014591842 (1, NHMUK); as previous except: April 1985, NHMUK 014591843 (1, NHMUK); as previous except: November 1985, NHMUK 014591844 (1, NHMUK); as previous except: G. Mogongonipa summit, 1008 m, at light, November 1985 (1, RABC); as previous except Plt C, ca 400 m, lowland forest, Feb – April 1985, pitfall trap, NHMUK 014591845 (1, NHMUK); as previous except: 200 – 300 m, October 1985, litter, NHMUK 014591846 (1, NHMUK); Central Sulawesi, Lore Lindu N. P., Marena forest, alt. m 600, 15. xii. 1985, J. Krikken & J. van Tol (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	diagnosis	Diagnosis. This species is distinguished by its very stout form and elytra that are laterally constricted at both the basal and apical tenth. Similar species. Beaverium obstipus (Schedl, 1935)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	description	Description (female). 4.15 – 4.6 mm long (mean = 4.32 mm; n = 5); 1.91 – 2.19 × as long as wide. Body, legs and antennae ferruginous. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface shining, punctate, alutaceous. Eyes moderately emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrow, triangular, weakly impressed. Antennal scape long and slender, as long as club. Pedicel narrower than scape, shorter than funicle. Antennal funicle 4 - segmented, segment 1 shorter than pedicel. Club as broad as tall, circular, club type 4, flattened, segment 1 corneous, transverse on anterior face, occupying about 2 / 5 of club; segment 2 narrow, corneous; segments 1 – 3 visible on posterior face. Pronotum 0.9 – 1.05 × as long as wide. In dorsal view rounded, type 1, sides parallel on basal 2 / 3; anterior margin with a short continuous recurved carina. In lateral view robust and subquadrate, type 5, disc flat, summit at basal third. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, coarsely, densely punctate. Lateral margins obliquely costate. Base transverse, posterior angles strongly oblique. Scutellum moderate, broadly linguiform, flush with elytra, flat, shiny. Elytra: 1.05 – 1.19 × as long as wide, 1.18 – 1.33 × as long as pronotum, shining. Base transverse, humeral angles rounded, sides convex with constrictions at basal and apical tenth, then narrowly rounded to apex; lateral profile of declivity rounded, apex posterolaterally broadened, entire. Disc convex, appearing humped, subshiny; striae not impressed, with large, shallow punctures separated by 0.5 – 3 diameters of a puncture, glabrous; interstriae flat, punctures half size of strial punctures, confused, each with a long, erect seta (mostly abraded). Declivity occupying ~ 1 / 2 of elytra, gradual, face flat, smooth, subshiny, glabrous; strial and interstrial punctures strongly confused, indistinguishable, striae not impressed, punctures larger than those of discal. Posterolateral margin elevated, costate to interstriae 5. Legs: procoxae contiguous; prosternal coxal piece tall and pointed. Protibiae obliquely triangular broadest at apical 1 / 4; posterior face smooth; apical 1 / 2 of outer margin with 8 – 10 moderate socketed denticles, each approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded with 13 or 14 and 11 or 12 large socketed denticles, respectively, their length longer than basal width.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	distribution	Distribution. Indonesia (Sulawesi).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA3711CFF7024727AFCF489.taxon	etymology	Etymology. L. brevis = short, caudatus = tailed, referring to the steep elytral declivity.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	description	(Figs 17 – 20) http: // zoobank. org / urn: lsid: zoobank. org: act: DC 32 B 1 DE-A 19 F- 44 F 6 - 829 B- 75936 E 221 D 4 F	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	materials_examined	Type material. Holotype, female: INDIA, Koppa, Quard Hitlow Estate, 30. x. [19] 90, borer on Grevillea robusta, S. D. Samuel coll., 12 IIE 21466, NHMUK 014591743 (NHMUK). Paratype, female, as holotype except: MSUC _ ARC _ 314154 (1, MSUC)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	diagnosis	Diagnosis. This species is distinguished by the unique pronotal apex which is produced ventrad to form a quadrate process bearing four large teeth and the area dorsad of process appearing bisulcate. Similar species. Cnestus nitidipennis (Schedl, 1951), C. protensus (Eggers, 1930)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	description	Description (female). 4.2 mm long (mean = 4.2 mm; n = 2); 1.91 × as long as wide. Head red, frons, epistoma black, pronotum red with black anterior quadrate process, elytra black, each elytron with a large transparent area on the declivity, antennae and legs light brown. Head: mandibles enlarged (in lateral view protruding forward at 90 ° to the plane of the frons, dorsoventrally deeper than normal; in anterior view, with an upwardly directed, smooth, rounded process on the dorsal side); epistoma moderately and broadly emarginate with a row of hair-like setae. Frons transversely impressed from epistoma to upper level of eyes; median carina absent; surface setose, shagreened, tuberculate, moderately covered with erect hair-like setae, tubercles larger, denser above eyes. Eyes very shallowly emarginate just above antennal insertion, upper part much smaller than lower part. Antennal scape long and slender, longer than club. Pedicel as wide as scape, shorter than funicle. Funicle 3 - segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face. Pronotum 0.95 × as long as wide. In dorsal view conical, type 6, sides parallel in basal 1 / 2, rounded anteriorly; anterior margin produced ventrad to form a quadrate process bearing four large teeth, area dorsad of process appearing bisulcate. In lateral view uniformly rounded without distinct summit, type 1, disc flat. Anterior slope with densely spaced, coarse and large asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, erect hairlike setae, some longer hair-like setae at anterior and lateral margins. Disc shining, alutaceous with dense, coarse punctures, glabrous. Lateral margins carinate. Base carinate, medially emarginate, posterior angles broadly rounded. Mycangial tuft absent. Scutellum large, linguiform, flush with elytra, flat, shining. Elytra: 0.95 × as long as wide, as long as pronotum, glabrous. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 1 / 2, then broadly rounded to apex. Strial punctures minute; interstriae impunctate. Declivity strongly rounded, smooth, transversely impressed before apex; six striae present. Posterolateral margin carinate to interstriae 3. Legs: procoxae contiguous. Protibiae very slender with three large, narrow denticles on outer margin; posterior face smooth. Meso- and metatibiae flattened, outer margins evenly rounded with five and 12, large similarly sized socketed denticles, their length longer than basal width.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	distribution	Distribution. India (Karnataka).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	biology_ecology	Host plants. The type series was collected from Grevillea robusta (Proteaceae).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA0711CFF70244A7DA9F3FE.taxon	etymology	Etymology. L. luculentus = brilliant, excellent. A variable adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	description	(Figs 21 – 24) http: // zoobank. org / urn: lsid: zoobank. org: act: 17 DCDCE 9 - DF 6 C- 4 B 6 F- 8 B 94 - F 57908 A 9 DE 74	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	materials_examined	Type material. Holotype, female: VIETNAM, Lao Cai Prov., Hoang Lien N. P., 22.35 N, 103.77 E, 1500 m, 17. v. 2019, VN 159, SM Smith, AI Cognato, ex 3 – 5 cm branch, SAX 397 [DNA voucher], MSUC _ ARC _ 314124 (MSUC). GenBank. CAD: OP 607183 and COI: OP 617754.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	diagnosis	Diagnosis. This species is distinguished by the short, steep declivity that is ~ 1 / 5 of total elytral length, armed with large distinct tubercles on interstriae 1 and 3, interstriae 2 minutely granulate on basal third; moderate body length; posterolateral margins rounded; declivital interstriae 1 setae in three confused rows, interstriae 2 setae uniseriate; declivity strongly shagreened; strial and interstrial punctures small, shallow, strongly confused and indistinguishable. Cyclorhipidion achlys shares a similar declivital setal pattern and size as C. pelliculosum (Eichhoff, 1878), but can be distinguished by the strial punctures which are large, distinct and seriate. Similar species. Cyclorhipidion pelliculosum	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	description	Description (female). 3.4 mm long (n = 1); 3.09 × as long as wide. Body dark reddish brown, antennae and legs brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface subshining, impunctate, alutaceous, finely rugose. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrow, triangular, deeply impressed. Antennal scape short and thick, as long as club. Pedicel narrower than scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying about 2 / 5 of club; segment 2 narrow, corneous; segments 1 ‒ 2 present on posterior face. Pronotum 1.09 × as long as wide. In dorsal view subquadrate, sides convex, type 3, narrowly rounded anteriorly; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 7, disc flat, summit at apical 2 / 5. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc dull, alutaceous, densely, finely punctate, finely setose, setae recumbent, hair-like. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Scutellum moderate, broad, linguiform, flush with elytra, flat, shining. Elytra: 2.0 × as long as wide, 1.83 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 9 / 10, then strongly rounded to apex. Disc flat, shining, striae and interstriae moderately setose, setae long, erect, hair-like, striae not impressed, punctures minute, confused, spaced by 4 ‒ 6 diameters of a puncture; interstriae not impressed, minutely punctate, punctures equal to strial punctures, strongly confused, separated by 3 ‒ 6 diameters of a puncture. Declivity occupying 1 / 5 of elytra, very steeply rounded, declivital face convex, strongly shagreened, distinctly separated from smooth, shining disc; striae 1 ‒ 3 equidistant, strial and interstrial punctures small, shallow, strongly confused, indistinguishable, strial punctures as large as those of disc; interstriae 1 and 2 flat, interstriae 3 convex, interstriae 1 bearing three rows of confused setae on declivital face, interstriae 2 bearing one row of uniseriate setae, setae long, semi-erect hair-like; interstriae 1 and 3 armed with a row of moderate to large distinct tubercles, interstriae 2 minutely granulate on basal third. Posterolateral margin rounded, granulate, extending to interstriae 7. Legs: procoxae contiguous. Protibiae semi-circular with evenly rounded outer edge, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with eight moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded each with 14, and 12, small to moderate socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	distribution	Distribution. Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA17112FF7024017C71F539.taxon	etymology	Etymology. G. achlys = mist in eyes before death. Achlys is the Greek Goddess of death-mist, and the personification of misery and suffering. This species represents the numerous nearly indistinguishable pseudocryptic Cyclorhipidion species that dwell in the misty mountain tops of Southeast Asia and perpetuate misery among taxonomists. A noun in apposition.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	description	(Figs 25 – 28) http: // zoobank. org / urn: lsid: zoobank. org: pub: 78 CCF 021 - B 47 F- 48 D 6 - 8914 - BF 9 AD 4 A 33 C 79	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	materials_examined	Type material. Holotype, female: INDONESIA, Sulawesi Utara, Dumoga Bone N. P., ‘ Clarke’ Camp lower montane forest, 1140 m, at light, March 1985, NHMUK 014591834 (NHMUK). Paratypes, female, as holotype except: G. Mogongonipa summit, 1008 m, at light, May 1985, MSUC _ ARC _ 314151 (1, MSUC); as previous except: November 1985 (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	diagnosis	Diagnosis. This species can be distinguished by the posterior face of the protibiae inflated and denticulate, almost appearing to have a double row of denticles on outer margin; declivital interstriae 1 – 4 bearing a regular row of conical denticles or tubercles; and declivital striae weakly impressed. Cyclorhipidion conidentatus is most similar to C. sus (Schedl, 1973) and C. multipunctatum (Browne, 1980) which both have the posterior face of the protibiae inflated and denticulate and a type 4 antennal club. C. conidentatus can be distinguished from both by the presence of conical denticles or tubercles along the entire length of declivital interstriae 2 that are as large as those of other interstriae. Similar species. Cyclorhipidion sus, C. multipunctatum	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	description	Description (female). 3.1 – 3.5 mm long (mean = 3.37 mm; n = 3); 2.38 – 2.50 × as long as wide. Body dark reddish brown, the elytra darker especially on the declivity, vestiture fairly long and dense. Head: epistoma entire, transverse, lined with a row of hair-like setae. Frons broadly convex, slightly transversely depressed between the eyes, weakly shining, finely papillate on lower part above epistoma, the papillae with short, fine, erect hairs, traces of a fine median carina, upper part with punctures larger, sparser and impressed; vertex impunctate. Eyes deeply emarginate above level of antennal insertion, upper portion of eyes smaller than lower part. Submentum narrowly triangular, deeply impressed. Scape short and thick, approximately as long as club. Pedicel as long as funicle. Antennal funicle 4 - segmented, segment 1 as long as pedicel. Club as broad as tall, circular, club type 4, flattened, segments 1 and 2 visible on posterior face. Pronotum 0.96 – 1.07 × as long as wide. In dorsal view long and rounded frontally, type 7, sides evenly rounded, widest in the middle, rounded anteriorly; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 7, disc flat, summit just in front of middle. Anterior slope strongly convex, very densely covered with small, granulate asperities, decreasing in size towards the summit, posterior part of disc subnitid, finely densely punctate, the pubescence fairly long, inclined towards the summit. Scutellum linguiform, fairly large, shining, impunctate. Elytra: 1.42 × as long as wide, 1.33 – 1.48 × as long as pronotum and slightly wider. Base transverse, edge oblique, humeral angles rounded, sides parallel in basal 2 / 5, then slightly narrowed, rather abruptly incurved apically, with a constriction just before apex. Declivity beginning after basal half of elytra, convex, its slope gradually increasing towards apex. Basal part of disc flat, shining, striae and interstriae finely densely, irregularly, punctate, strial and interstrial punctures indistinguishable, strongly confused, setose, punctures with fairly long, inclined hairs; posterior part of disc and declivity matte, striae 1 and 2 sulcate on upper two-thirds (weakly so in both paratypes), strial punctures clear, larger and much shallower than those on interstriae, closely placed, weakly impressed; interstriae much wider than striae, each with a regular row of conical denticles or tubercles (more strongly developed in the larger specimens), extending almost to beginning of declivity on interstriae 1 and 2, progressively less close on the outer interstriae, sutural interstriae widened and outwardly convex on apical third of declivity, all interstriae with several irregular rows of minute punctures bearing long semi-erect hairs. Posterolateral margin rounded. Legs: procoxae contiguous. Protibiae semi-circular with evenly rounded outer margin, broadest at apical 1 / 3; eight moderate denticles on outer margin; posterior face weakly inflated, denticulate near outer margin. Meso- and metatibiae flattened, outer margins evenly rounded with 15 and 14 large similarly sized socketed denticles, respectively, their length longer than basal width.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	distribution	Distribution. Indonesia (Sulawesi).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAE7113FF7027837A95F489.taxon	etymology	Etymology. L. conus = cone, dentatus = toothed, referring to the conical denticles on the elytra.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	description	(Figs 29 – 32) http: // zoobank. org / urn: lsid: zoobank. org: act: A 396202 C- 2677 - 481 C- 939 D- 75 F 51 D 99 BDA 7	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	materials_examined	Type material. Holotype, female: THAILAND, Chiang Mai, Omkoi District, Omkoi Wildlife Sanctuary, 28. VI. 2013, C. Bateman, ex Shorea sp., MSUC _ ARC _ 314126 (MSUC). Paratypes, as holotype, MSUC _ ARC _ 314139 (1, MSUC); SAX 404, MSUC _ ARC _ 314140 [DNA voucher] (1, MSUC). GenBank. CAD: OP 650089 and COI: OP 644414.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	diagnosis	Diagnosis. This species is distinguished by the elytral disc with a shallow median saddle-like impression; declivity very steep; declivital posterolateral margin carinate to interstriae 5; elytral apex bearing three large triangular spines at each interstriae 2, 3 and 4 that are at least the width of an interstriae, medial pair largest and decreasing in size laterally; declivital face unarmed by tubercles; declivital interstriae impunctate and alutaceous. Similar species. Cyclorhipidion miyazakiense (Murayama, 1936), C. hsuae Lin & Smith, 2022, and C. wui Lin & Smith, 2022	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	description	Description (female). 2.6 – 2.8 mm long (mean = 2.7 mm; n = 3); 2.45 – 2.55 × as long as wide. Body, antennae, and legs ferruginous. Elytra slightly darker than rest of body. Head: epistoma entire, transverse, with a row of hairlike setae. Frons weakly convex to upper level of eyes; surface subshining, impunctate, alutaceous, finely rugose; median line feebly elevated. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrow, triangular, deeply impressed. Antennal scape regularly thick, as long as club. Pedicel narrower than scape, shorter than funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 4; segment 1 corneous, sinuate on anterior face, occupying about 1 / 5 of club; segment 2 narrow, soft; segments 1 ‒ 3 present on posterior face. Pronotum 1.09 – 1.18 × as long as wide. In dorsal view elongate, rounded frontally, type 7, sides parallel on basal 2 / 3; anterior margin without serrations. In lateral view elongate with disc slightly longer than anterior slope, type 7, disc flat, summit just anteriad of midpoint. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshining, alutaceous, densely, finely punctate, finely setose, setae short, erect, hair-like. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Scutellum moderate, broad, linguiform, flush with elytra, flat, shining. Elytra: 1.33 – 1.36 × as long as wide, 1.15 – 1.25 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 5 / 8, then broadly rounded to apex. Disc with a shallow median saddle-like impression, shining, striae and interstriae densely setose, setae about as long as interstriae 2, semi-erect, hair-like, striae not impressed, punctures large, uniseriate, spaced by one diameter of a puncture; interstriae not impressed, minutely punctate, punctures minute, strongly confused, separated by 2 ‒ 3 diameters of a puncture. Declivity occupying 1 / 4 of elytra, steeply rounded, declivital face flat, unarmed alutaceous, dull, separation between the smooth, shining disc and dull declivity distinct; three striae present, striae 1 ‒ 3 equidistant, striae distinctly impressed, punctures very large, shallow, subcontiguous, shagreened, much larger than on disc; interstriae flat, impunctate, setose, setae erect, shorter than interstriae 1 width; interstriae 1 and 2 bearing two rows of confused erect setae on declivital face; elytral apex bearing three large triangular spines at interstriae 2, 3 and 4 that are at least the width of an interstriae, medial pair largest and decreasing in size laterally. Posterolateral margin carinate, extending to interstriae 5. Legs: procoxae contiguous; prosternal coxal piece tall and pointed. Protibiae semi-circular with evenly rounded outer edge, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with seven or eight moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded each with ten or 11 and 9 – 11 moderate socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	distribution	Distribution. Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	biology_ecology	Host plants. The type series was collected from Shorea (Dipterocarpaceae).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAF7113FF7027147F13F3E5.taxon	etymology	Etymology. L. gladi - = sword, - gerum = carrier. In reference to the wide and long denticles on the elytral apex. A variable adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	description	(Figs 33 – 36) http: // zoobank. org / urn: lsid: zoobank. org: act: 934 B 4 C 11 - 70 DB- 40 D 8 - 91 E 6 - CC 492 C 8 BCC 99	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	materials_examined	Type material. Holotype, female: VIETNAM, Lao Cai Prov., Hoang Lien N. P., 22.35 N, 103.77 E, 1500 – 2000 m, 21. v. 2019, VN 198, SM Smith, AI Cognato, ex 2 cm branch, SAX 396 [DNA voucher], MSUC _ ARC _ 314128 (MSUC). GenBank. CAD: OP 607182 and COI: OP 617753.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	diagnosis	Diagnosis. This species is distinguished by the moderate size; declivity obliquely truncate, strongly shagreened and dull; pronotum longer than wide, subquadrate from dorsal view (type 3); and declivital interstrial punctures replaced by confused tubercles. Cyclorhipidion lapilliferum is very similar to C. circumcisum (Sampson, 1921) from which it can be distinguished by the smaller size (3.4 mm vs 4.0 mm) and more elongate form (3.09 × vs 2.67 × as long as wide). Similar species. C. circumcisum, C. muticum Smith, Beaver & Cognato, 2020 a	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	description	Description (female). 3.4 mm long (n = 1); 3.09 × as long as wide. Body, legs and antennae ferruginous. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface shagreened, impunctate, alutaceous, moderately rugose. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~ 1 / 3; segment 2 narrow, corneous; segments 1 ‒ 3 present on posterior face. Pronotum 1.18 × as long as wide. In dorsal view subquadrate, sides convex, type 3, narrowly rounded anteriorly; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 7, disc flat, summit at apical 1 / 3. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc shagreened, alutaceous, densely, finely punctate, setose, setae dense, long, fine hair-like. Lateral margins obliquely costate. Base transverse, posterior angles narrowly rounded. Scutellum large, broad, linguiform, shiny, flush with elytra, flat. Elytra: 1.91 × as long as wide, 1.62 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal ~ 4 / 5, then sharply angulate to apex. Disc ascending posteriorly, shiny, basal 1 / 4 shagreened, impunctate; striae and interstriae not impressed, sparsely setose, setae short erect, hairlike, striae seriate, punctures small, shallow; interstrial punctures confused, minute, punctures strongly confused, separated by 3 ‒ 5 diameters of a puncture. Declivity occupying 1 / 3 of elytra, obliquely truncate, face weakly convex, strongly shagreened, dull, densely setose; six striae present, striae weakly impressed, punctures difficult to discern, minute, shallow, shagreened, slightly larger than on disc, striae 2 equidistant between 1 and 3; interstriae weakly convex, interstriae lightly setose, setae long, very fine, erect hair-like; interstriae impunctate, densely covered by confused tubercules, tubercles numerous, small to minute. Posterolateral margin costate, tuberculate, extending laterally to interstriae 7, setose, setae long, fine, erect, hair-like. Legs: procoxae contiguous. Protibiae semi-circular with evenly rounded outer edge, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with nine moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded with 12 and 14 moderate socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	distribution	Distribution. Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAC7111FF7026A57F68F04D.taxon	etymology	Etymology. L. lapilliferum = having pebbles. In reference to the granules on the elytral declivity. A variable adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	description	(Figs 37 – 40) http: // zoobank. org / urn: lsid: zoobank. org: act: 621995 C 4 - 9 DA 1 - 4 B 17 - 8 B 98 - 96 A 776625 C 56	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	materials_examined	Type material. Holotype, female: NEPAL, Kathmandu Dist., Godwari, 5600 ’, 25. v. 1983, M. J. D. Brendell, under bark of dead trees, NHMUK 014591823 (NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	diagnosis	Diagnosis. This species is distinguished by moderate size; declivital slope gradual; separation between the smooth shiny elytral disc and shagreened declivity gradual, not sharply distinct; declivital striae weakly impressed, strial punctures small, indistinct; declivital interstriae 1 – 3 armed with a row of few and sparse seriate granules; and elytral apex and posterolateral margin granulate. This species is superficially similar to species in the C. pelliculosum species group (Smith et al. 2020 a) which includes C. achlys, C. bodoanum (Reitter, 1913), C. inarmatum (Eggers, 1923), and C. tenuigraphum (Schedl, 1953 a) among others. C. nepalense can be distinguished by the following combination of characters (C. nepalense given first): declivital interstriae 2 feebly granulate vs unarmed (rarely granulate in some C. pelliculosum and C. tenuigraphum); the more gradual slope of the elytral declivity that occupies 1 / 3 of the elytra vs steep declivity occupying 1 / 4 of elytra; elytral disc with confused strial and interstrial punctures that are the same size vs strial punctures distinct, seriate and interstrial punctures confused. Similar species. Cyclorhipidion petrosum Smith, Beaver & Cognato, 2020 a	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	description	Description (female). 2.7 mm long (n = 1); 3.0 × as long as wide. Body, legs and antennae light brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface subshiny, punctate, alutaceous, rugose. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrow, triangular, deeply impressed. Antennal scape short and thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~ 2 / 5; segment 2 narrow, corneous; segments 1 and 2 present on posterior face. Pronotum 1.22 × as long as wide. In dorsal view long and rounded frontally, type 7, sides parallel in basal 3 / 4, rounded anteriorly; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 7, disc flat, summit at apical 2 / 5. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, alutaceous, densely, finely punctate, finely setose, setae short, erect, hair-like. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Scutellum large, broad, linguiform, shiny, flush with elytra, flat. Elytra: 1.78 × as long as wide, 1.45 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallelsided in basal 3 / 4, then broadly rounded to apex. Disc flat, shiny, striae and interstriae densely setose, setae long, semi-recumbent, hair-like; striae and interstriae not impressed, minutely punctate, punctures strongly confused, separated by 2 ‒ 4 diameters of a puncture. Declivity occupying ~ 1 / 3 of elytra, declivital slope gradual, rounded, shagreened, separation between the smooth shiny disc and shagreened declivity gradual, not sharply distinct; three striae present, striae 2 equidistant between 1 and 3, striae weakly impressed, punctures moderate, much larger than on disc; interstriae feebly convex, interstriae setose, setae moderate, long, semi-erect hair-like; interstriae minutely punctate, interstriae 1 – 3 armed with a row of few and sparse seriate granules. Posterolateral margin rounded, granulate. Legs: procoxae contiguous; prosternal coxal piece tall and pointed. Protibiae semi-circular with evenly rounded outer edge, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with seven moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded with ten moderate socketed denticles.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	distribution	Distribution. Nepal.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAD7116FF7022507D8FF7DD.taxon	etymology	Etymology. In reference to the collection locality, Nepal. An adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	description	(Figs 41 – 44) http: // zoobank. org / urn: lsid: zoobank. org: act: 33 F 3 D 649 - FE 24 - 4 F 82 - 9 BE 6 - 9166 B 4 CD 7040	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	materials_examined	Type material. Holotype, female: VIETNAM, Lao Cai Prov., Hoang Lien N. P., 22.35 N, 103.77 E, 1500 – 2000 m, 21. v. 2019, VN 198, SM Smith, AI Cognato, ex 2 cm branch, SAX 392 [DNA voucher], MSUC _ ARC _ 314131 (MSUC). Paratype, female: as holotype except: SAX 393 [DNA voucher], MSUC _ ARC _ 314132 (1, MSUC). GenBank. CAD: OP 607178; OP 607179 and COI: OP 617749; OP 617750.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	diagnosis	Diagnosis. This species is distinguished by the short, steep declivity that is ~ 1 / 5 of total elytral length, armed with moderate tubercles on interstriae 1 and 3, interstriae 2 always unarmed; moderate body length; posterolateral margins rounded; declivital interstriae 1 setae in two confused rows, interstriae 2 setae uniseriate; and the declivity strongly shagreened, strial punctures shallow, obscure. The species is most similar to C. inarmatum and C. cognatoi Lin & Smith 2022 with which it shares similar declivital setal pattern and body size but both of which have a strongly shining elytral declivity. C. taedulum can be distinguished from both by the elytral length / width ratio and elytral / pronotal length / width ratio. In C. taedulum the elytra is 1.89 × as long as wide and 1.7 × as long as pronotum. In C. cognatoi the elytra is 1.67 – 1.79 × as long as wide and 1.46 – 1.58 × as long as pronotum and in C. inarmatum the elytra is 1.62 – 1.70 × as long as wide and 1.31 – 1.41 × as long as pronotum. Similar species. This species closely resembles C. inarmatum, C. bodoanum, C. cognatoi all of which have two rows of confused setae on declivital interstriae 1 and one row of setae on declivital interstriae 2.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	description	Description (female). 2.7 mm long (mean = 2.7 mm; n = 2); 3.0 × as long as wide. Body light brown, antennae and legs light brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface subshining, impunctate, alutaceous, finely rugose; median line feebly elevated. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrow, triangular, deeply impressed. Antennal scape short and thick, as long as club. Pedicel narrower than scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying about 2 / 5 of club; segment 2 narrow, soft; segments 1 ‒ 2 present on posterior face. Pronotum 1.11 × as long as wide. In dorsal view very elongate and rounded frontally, type 9, sides parallel on basal 3 / 4; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 8, disc flat, summit at apical 1 / 3. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshining, alutaceous, densely, finely punctate, finely setose, setae short, erect, hair-like. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Scutellum moderate, broad, linguiform, flush with elytra, flat, shining. Elytra: 1.89 × as long as wide, 1.7 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 4 / 5, then broadly rounded to apex. Disc flat, shining, striae and interstriae moderately setose, setae long, semi-recumbent, hair-like; striae not impressed, punctures small, uniseriate, spaced by 1.5 diameters of a puncture; interstriae not impressed, minutely punctate, punctures much smaller than strial punctures, strongly confused, separated by more than 2 ‒ 5 diameters of a puncture. Declivity occupying 1 / 5 of elytra, steeply rounded, declivital face convex, strongly shagreened, separation between the smooth, shining disc and shagreened declivity distinct; three striae present, striae not impressed, punctures large, shallow, subcontiguous, shagreened, much larger than on disc; interstriae flat, minutely punctate, interstriae 1 bearing two rows of confused setae on declivital face, interstriae 2 being one row of uniseriate setae, setae long, semi-erect; interstriae 1 and 3 with three tubercles along their length, interstriae 2 unarmed. Posterolateral margin rounded, granulate, extending to interstriae 7. Legs: procoxae contiguous; prosternal coxal piece tall and pointed. Protibiae semi-circular with evenly rounded outer edge, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with seven or eight moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded each with ten and ten or 11 moderate socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	distribution	Distribution. Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAA7117FF7025E07CB4F7F9.taxon	etymology	Etymology. L. taedulus = boring. The hackneyed appearance of this species is a recurring pattern among pseudocryptic Cyclorhipidion. A variable adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	description	(Figs 45 – 48) http: // zoobank. org / urn: lsid: zoobank. org: act: 25 B 61327 - 5 FE 5 - 45 A 7 - 90 B 8 - AF 88 DA 3 E 7 F 37	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	materials_examined	Type material. Holotype, female: VIETNAM, Lao Cai Prov., Hoang Lien N. P., 22.35 N, 103.77 E, 1500 – 2000 m, 19. v. 2019, VN 170, SM Smith, AI Cognato, ex 5 cm branch, MSUC _ ARC _ 314130 (MSUC). Paratypes, as holotype MSUC _ ARC _ 314141, MSUC _ ARC _ 314142, (2, MSUC), NHMUK 014591848 (1, NHMUK); as holotype except: 17. v. 2019, VN 159, ex 3 – 5 cm branch, MSUC _ ARC _ 314143 (1, MSUC; 1, NMNH); VIETNAM: Lao Cai Prov., 17. v. 2019, VN 156, ex 4 cm branch, SAX 394 [DNA voucher], MSUC _ ARC _ 314144 (1, MSUC). GenBank. CAD: OP 607180 and COI: OP 617751.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	diagnosis	Diagnosis. This species is distinguished by the moderate size and comparatively stout form; declivity obliquely truncate with rounded margins; pronotum about as long as wide and subquadrate from dorsal view, type 3; declivital interstrial punctures replaced by sparse, small, confused tubercles; declivital strial punctures moderate, distinct; declivital face appearing convex and strongly shiny; and declivital striae feebly impressed and interstriae inflated on apical 1 / 2. This species most closely resembles C. amputatum Smith, Beaver & Cognato, 2020 a from which it can be distinguished by the following combination of characters (C. titorum given first): smaller size and stouter form 4.2 – 4.6 mm long, 2.47 – 2.71 × as long as wide vs 5.0 mm long, 3.13 × as long as wide; declivital striae feebly impressed vs distinctly impressed; declivity densely setose, interstriae 1 and 2 with three confused rows of long dense setae vs declivity lightly setose, interstriae 1 and 2 with two confused rows of short setae. Similar species. Cyclorhipidion amputatum, C. muticum Smith, Beaver & Cognato, 2020, C. circumcisum	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	description	Description (female). 4.2 – 4.6 mm long (mean = 4.44 mm; n = 5); 2.47 – 2.71 × as long as wide. Body dark brown, antennae and legs light brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface shiny, punctate, alutaceous, rugose. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum broadly triangular, deeply impressed. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~ 1 / 3; segment 2 narrow, corneous; segments 1 and 2 present on posterior face. Pronotum 0.94 – 1.0 × as long as wide. In dorsal view subquadrate and, sides convex, type 3, narrowly rounded anteriorly; anterior margin without serrations. In lateral view tall, type 2, disc flat, summit at anterior ~ 2 / 5. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, alutaceous, densely, finely punctate, finely setose, setae short, erect, hair-like. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Scutellum large, broad, linguiform, shiny, flush with elytra, flat. Elytra: 1.12 – 1.71 × as long as wide, 1.12 – 1.81 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 9 / 10, then sharply angulate to apex. Disc flat, shiny, striae and interstriae densely setose, setae long, semi-recumbent, hair-like, striae and interstriae strongly confused, indistinguishable; striae and interstriae not impressed, minutely punctate, punctures strongly confused, separated by 2 ‒ 5 diameters of a puncture. Declivity occupying ~ 1 / 3 of elytra, obliquely truncate, face convex, strongly shiny; five striae present, striae feebly impressed, striae 2 equidistant between 1 and 3, strial punctures moderate, distinct, subcontiguous to separated by one diameter of a puncture, subshiny, much larger than on disc; interstriae inflated on apical 1 / 2 of declivity, interstriae setose, setae dense, long, semi-erect hair-like, interstriae 1 and 2 with three confused rows; interstriae impunctate, coarsely tuberculate, tubercles sparse, small, strongly confused, variably sized. Posterolateral margin forming a costa extending laterally to interstriae 7; costa tuberculate, setose, setae long, erect, fine, hair-like. Legs: procoxae contiguous; prosternal coxal piece tall and pointed. Protibiae semi-circular with evenly rounded outer margin, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with ten or 11 moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded with 15 or 16 moderate socketed denticles.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	distribution	Distribution. Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFAB7114FF7025C47A2DF04D.taxon	etymology	Etymology. L. patronym for the Titus family. Given that xyleborines farm fungi as a family, it is appropriate to honor the Titus family farmers. Rebecca, Paul, and Rose have provided SMS and AIC (and hundreds of others in Central Michigan) with sustainable foods (including mushrooms) for over 15 years.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	description	(Figs 49 – 52) http: // zoobank. org / urn: lsid: zoobank. org: act: 5090 DCC 4 - 5358 - 461 E- 9 DCD- 59 F 2 D 7 B 1 A 55 D	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	materials_examined	Type material. Holotype, female: JAPAN, G. Lewis, NHMUK 014591822 (NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	diagnosis	Diagnosis. This species is distinguished by its stout form; declivital posterolateral margin costate and granulate; pronotum appearing subquadrate when viewed dorsally (type 3); protibiae outer margins obliquely triangular bearing five short obtuse denticles; declivital strial punctures approximately the same size on disc and declivity; declivity gradually rounded; and declivital surface opalescent. This species is very similar to E. praevius (Blandford, 1894) and can be distinguished by the following combination of characters (E. alastos given first): stouter form, 2.56 vs 2.72 – 2.92 × as long as wide; stout elytra 1.44 vs 1.6 – 1.83 × as long as wide; declivital surface opalescent vs strongly shining; declivital interstriae minutely granulate vs distinctly denticulate. Similar species. E. praevius, E. andamanensis (Blandford, 1896), Xyleborus aquilus Blandford, 1894	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	description	Description (female). 3.2 mm long (n = 1); 2.56 × as long as wide. Body ferruginous. Legs and antennae light brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons flattened to upper level of eyes; surface alutaceous sparsely, finely punctate, setose; punctures each a long, erect hair-like seta. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Antennal scape long and slender, longer than club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 longer than pedicel. Club approximately circular, flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~ 1 / 3; segment 2 narrow, soft; segments 1 ‒ 3 present on posterior face. Pronotum 1.12 × as long as wide. In dorsal view subquadrate with prominent anterolateral corners, type 3, sides weakly convex; anterior margin without serrations. In lateral view tall, type 2, disc flat, summit at midpoint. Anterior slope with densely spaced, broad asperities, becoming lower and more strongly transverse towards summit. Disc strongly shiny with sparse, minute punctures, some longer hairlike setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded, almost quadrate. Scutellum small, linguiform, flush with elytra, flat, shiny. Elytra: 1.44 × as long as wide, 1.43 × as long as pronotum. Base transverse, edge oblique and unarmed by granules, humeral angles rounded, parallel-sided in basal 3 / 4, then angulate to apex. Disc convex, shiny, striae not impressed, with large, shallow punctures separated by less than one diameter of a puncture, glabrous; interstriae flat, very sparsely finely punctate, punctures half size of strial punctures, each with a long, erect seta (mostly abraded). Declivity occupying approximately half of elytra, gradual, face flat, opalescent; striae impressed, strial punctures as large as those of disc; interstriae impunctate, interstriae uniformly minutely granulate along the entire length. Posterolateral margin elevated, carinate and granulate to interstriae 7. Legs: procoxae contiguous. Protibiae obliquely triangular, posterior face smooth; apical 1 / 3 of outer margin with five moderate, spaced socketed denticles, their length as long as basal width. Mesotibiae flattened; outer margin evenly rounded with eight moderate socketed denticles.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	distribution	Distribution. Japan.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFA97115FF7026A57AE2F255.taxon	etymology	Etymology. G. alastos = unforgettable. The specimen resided at least 140 years in the NHMUK before its recognition as a new species and its description. Adjective applied as a noun in apposition.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	description	(Figs 53 – 56) http: // zoobank. org / urn: lsid: zoobank. org: act: FB 14 CA 12 - 8 DFE- 45 C 1 - B 89 A- 3115003 FA 64 D	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	materials_examined	Type material. Holotype, female: PAPUA NEW GUINEA, Mt. Wilhelm, - 5.720874, 145.2695, 1200 m. a. s. l., 1 – 3. xi. 2012, P 1643, plot 20, FIT-MW 1200 - T- 4 / 8 - d 08, Philip, Alois, Novotny, Leponce, ex flight intercept trap (MNHN). Paratypes, female, as holotype except: 27 – 29. x. 2012, P 1537, plot 7, FIT-MW 1200 - G- 2 / 8 - d 08 (1, MNHN), 29 – 31. x. 2012, P 1554, plot 9, FIT-MW 1200 - I- 3 / 8 - d 05, MSUC _ ARC _ 314145 (1, MSUC); - 5.73196, 145.2522, 700 m. a. s. l., 9 – 11. xi. 2012, P 1209, plot 14, FIT-MW 700 - N- 8 / 8 - d 16, Keltim, Uma, Novotny, Leponce, ex flight intercept trap, NHMUK 014591847 (1, NHMUK), 30. x – 1. xi. 2012, P 1212, plot 15, FIT-MW 700 - O- 3 / 8 - d 06 (1, ANIC); - 5.720874, 145.2695, 1200 m. a. s. l., 1 – 3. xi. 2012, P 1571, plot 12, FIT-MW 1200 - L- 4 / 8 - d 08, Philip, Alois, Novotny, Leponce, ex flight intercept trap, MSUC _ ARC _ 314146 (1, MSUC), 9 – 11. xi. 2012, P 1607, plot 15, FIT-MW 1200 - 0 - 8 / 8 - d 16 (1, BPBM), 3 – 5. xi. 2012, P 1620, plot 17, FIT-MW 1200 - Q- 5 / 8 - d 10 (1, MNHN), 9 – 11. xi. 2012, P 1631, plot 18, FIT-MW 1200 - R- 8 / 8 - d 16 (1, MNHN), 1 – 3. xi. 2012, P 1579, plot 12, FIT-MW 1200 - L- 4 / 8 - d 08, MSUC _ ARC _ 314152 (1, MSUC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	diagnosis	Diagnosis. This species is distinguished by the uniquely sculptured elytral declivity with the declivital face sulcate and encircled by sinuate lateral margins and declivity covered in star-shaped scales. Similar species. Leptoxyleborus sordicauda (Motschulsky, 1863)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	description	Description (female). 2.8 – 3.0 mm long (mean = 2.9 mm; n = 3); 2.23 – 2.31 × as long as wide. Body brown, legs and antennae yellow brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons flattened to upper level of eyes; median area with an impunctate, strongly shining kite-shaped area, lateral areas above epistoma impressed, densely punctate with large deep punctures, punctures each a long, erect hair-like seta. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Antennal scape long and slender, longer than club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 longer than pedicel. Club approximately circular, flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~ 1 / 3; segment 2 narrow, soft; segments 1 and 2 present on posterior face. Pronotum 0.92 – 0.96 × as long as wide. In dorsal view conical and elongate, type 5, sides weakly convex; anterior margin with a row of ten serrations. In lateral view tall, type 2, disc flat, summit at basal 2 / 5. Anterior slope with densely spaced, broad asperities, becoming lower and more strongly transverse towards summit. Disc strongly shiny with sparse, minute punctures, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles obliquely rounded. Scutellum large, linguiform, flush with elytra, flat, shiny. Elytra: 1.31 – 1.38 × as long as wide, 1.42 – 1.5 × as long as pronotum. Base transverse, edge oblique and unarmed by granules, humeral angles rounded, parallel-sided in basal 8 / 9, then acutely angulate to apex, appearing laterally broadened. Disc very short, 1 / 5 of total elytral length, flat, shiny; striae not impressed, with very large, deep punctures separated by less than one diameter of a puncture, glabrous; interstriae flat, very sparsely finely punctate, punctures minute, each with a long, recumbent hair-like seta, as long as the distance between punctures. Declivity deeply concave, laterally margined, costate from suture to striae 4, then acutely carinate and appearing sinuate in lateral view from interstriae 4 to sutural apex; concavity with strial and punctures as large as those of disc and densely covered in a dense layer of minute star-like scales. Legs: procoxae contiguous, elongate, longer than basal width; prosternal coxal piece tall and pointed. Protibiae distinctly triangular, broadest at apical 1 / 4; posterior face smooth; apical 1 / 3 of outer margin with six or seven very large socketed denticles, their length much longer than basal width. Meso- and metatibiae broad, flattened outer margin evenly rounded; outer margin of mesotibiae with 11 – 13 very large socketed denticles; metatibiae with evenly rounded with 10 – 12 moderate socketed denticles.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	distribution	Distribution. Papua New Guinea.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB6710BFF7026A57A05F53B.taxon	etymology	Etymology. L. regina = queen. In reference to the sinuate lateral margins of the elytral declivity which is reminiscent of a “ conch ” collared robe worn by Queen Elizabeth I. Noun in apposition.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	description	(Figs 57 – 60) http: // zoobank. org / urn: lsid: zoobank. org: act: 024 CA 706 - 3 D 66 - 4 B 20 - 9 CA 6 - 5 E 5 B 543 A 4 DE 2	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	materials_examined	Type material. Holotype, female: MALAYSIA, Sabah, Rhino Ridge, Danum Valley, 11. xi. 2006, J. Hulcr, ex Burseraceae, SAX 503 [DNA voucher] (NMNH). GenBank. CAD: OP 607245.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	diagnosis	Diagnosis. This species is distinguished by the elytral disc broadly, deeply transversely impressed with a saddlelike depression. Similar species. Tricosa cattienensis Cognato, Smith & Beaver, 2020 a, T. indochinensis Cognato, Smith & Beaver, 2020 a.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	description	Description (female). 3.1 mm long (n = 1); 2.81 mm × as long as wide. Head and pronotum ferruginous, elytra dark brown, legs and antennae light brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface shagreened, dull, punctate; punctures above epistoma small, fine, shallow, punctures larger, deeper on frons. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum slightly impressed below genae, narrowly triangular. Scape regularly thick, shorter than club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 as long as pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying basal ~ 1 / 3; segment 2 narrow, corneous; segments 1 and 2 present on posterior face. Pronotum 1.09 × as long as wide. In dorsal view elongate and parallel sided, type 7, sides parallel in basal 2 / 3, rounded anteriorly; anterior margin with a row of six serrations. In lateral view elongate, disc longer than anterior slope, type 7, summit low, on apical 2 / 5. Anterior slope with densely spaced narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, erect hair-like setae. Disc subshiny, alutaceous, finely punctate, finely setose, setae short, erect, hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded. Scutellum small, triangular, flat, flush with elytra. Elytra: 1.73 × as long as wide, 1.58 × as long as pronotum. Base transverse, margins oblique; humeral angles rounded; attenuate at apical half, apex broadly rounded. Disc broadly, deeply transversely impressed with a saddle-like depression, surface smooth, shining, unarmed; striae seriate punctate, punctures separated by less than one diameter of a puncture, glabrous; interstrial minutely uniseriate punctate, widely spaced, each with a long, erect seta 1.5 × as long as interstriae 2. Declivity occupying ~ 2 / 5 of elytra, gradual, face flat, opalescent; striae impressed, strial punctures as large as those of disc; interstriae impunctate, interstriae tubercles arranged in an alternating pattern of a small denticle followed by a large denticle, with the apices curved ventrad; interstrial setae as long or twice as long as those of disc. Posterolateral margins rounded and granulate. Legs: procoxae contiguous, prosternal posterocoxal piece conical, slightly inflated. Protibiae distinctly triangular, broadest at apical half, posterior face flat, unarmed; five large denticles present on outer margin of apical third. Meso- and metatibiae obliquely triangular, flattened, posterior face unarmed; metatibiae bearing eight moderate socketed denticles on outer margin.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	distribution	Distribution. Malaysia (Sabah).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	biology_ecology	Host plants. The holotype was collected from Burseraceae.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB77108FF70278B7B41F51D.taxon	etymology	Etymology. G. hipparion = little horse. In reference to the saddle-shaped elytra. Noun in apposition.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	description	(Figs 61 – 64) http: // zoobank. org / urn: lsid: zoobank. org: act: 1122 E 2 F 2 - 0 E 6 A- 4 E 0 E- 9 D 89 - E 7320313921 A	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	materials_examined	Type material. Holotype, female: THAILAND, Chiang Mai, Doi Pui, 18 – 22. x. 2004, TH 2, W. Puranasakul, ex ethanol trap, MSUC _ ARC _ 314133 (MSUC). Paratypes, as holotype except: xi. 2004, S. A. Dole, THA 149, MSUC _ ARC _ 314137 (1, MSUC), i. 2005, S. A. Dole, THA 193, MSUC _ ARC _ 314138 (1, MSUC). GenBank. CAD: OP 607261 and COI: OP 617825.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	diagnosis	Diagnosis. This species is distinguished by the posterior margin of elytra broadly rounded; declivital face with interstriae 1 and 2 unarmed by tubercles; declivital face moderately sulcate, striae punctate; lateral declivital margins elevated, bearing 6 – 8 pairs of small to moderate, narrow, sharply pointed and recurved spines (often asymmetric) on basal 2 / 3 of declivity; and sulcate area of declivity shagreened, subshining. Similar species. Xyleborinus dumosus sp. nov., Xyleborinus echinopterus Smith, Beaver & Cognato, 2020, X. octiesdentatus (Murayama, 1931), X. kwangruengensis Park & Smith, 2020 (in Park et al. 2020)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	description	Description (female). 2.5 mm long (mean = 2.5 mm; n = 3); 2.78 × as long as wide. Body, legs, and antennae uniformly brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, alutaceous, dull, densely, coarsely punctate, setose; punctures each a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, deeply impressed. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face. Pronotum 1.11 × as long as wide. In dorsal view basic and parallel-sided type 2, sides parallel in basal 2 / 3, rounded anteriorly; anterior margin without serrations. In lateral view elongate, disc longer than anterior slope, type 7, summit low, on apical 2 / 5. Anterior slope with densely spaced narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, alutaceous, finely punctate, finely setose, setae short, erect, hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded. Scutellum minute, conical, disconnected from elytra, surrounded by dense mycangial tuft of setae. Elytra: 1.67 × as long as wide, elytra 1.50 × as long as pronotum. Base transverse, medially emarginate near scutellum and mycangial tuft, edge oblique, humeral angles angulate, parallel-sided in basal 3 / 4, then broadly rounded to apex. Disc occupying basal 2 / 3, smooth, shiny, glabrous, unarmed; striae not impressed, glabrous, with moderate punctures separated by 1 – 2 diameters of a puncture; interstriae flat, sparsely finely biseriate punctate, punctures 1 / 2 as large as those of striae, each bearing erect hair-like setae, setae approximately as long as width of interstriae 2. Declivital face steeply rounded, moderately sulcate between interstriae 3, shagreened, subshiny, striae and interstriae moderately setose, setae long, semi-erect hair-like, as long as the distance between suture and interstriae 3 on declivital margins and semi-recumbent and no longer than an interstrial width on declivital face; three striae present, striae 1 and 2 subcontiguous, confused; strial punctures larger, deeper than on disc; interstriae 1 and 2 flat, armed by two and three denticles, respectively on declivital base, unarmed on face; interstriae 2 very narrow; interstriae 3 forming declivital margin, distinctly elevated, 6 – 8 pairs of small to moderate, narrow sharply pointed and recurved spines (often asymmetric) on basal 2 / 3. Posterolateral margin rounded, denticulate from interstriae 3 – 7. Legs: procoxae contiguous, prosternal coxal piece inflated. Protibiae obliquely triangular, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with seven moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae flattened, outer margin evenly rounded with eight moderate socketed denticles.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	distribution	Distribution. Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB47108FF7027B77F78F3AD.taxon	etymology	Etymology. G. acanthos - = thorn, - pteron = wing. In reference to the spines on the elytral declivity. A noun in apposition.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	description	(Figs 65 – 68) http: // zoobank. org / urn: lsid: zoobank. org: act: 5183 F 61 A-E 30 B- 4213 - 8 D 9 E-DA 2418 C 755 D 4	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	materials_examined	Type material. Holotype, female: VIETNAM, Lao Cai Prov., Hoang Lien N. P., 22.35 N, 103.77 E, 1500 – 2000 m, 19. v. 2019, VN 177, SM Smith, AI Cognato, ex 3 – 5 cm branch, SAX 434 [DNA voucher], MSUC _ ARC _ 314129 (MSUC). GenBank. CAD: OP 607184 and COI: OP 617755.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	diagnosis	Diagnosis. This species is distinguished by the posterior margin of elytra broadly rounded; declivital face with interstriae 1 and 2 unarmed by tubercles; declivital face moderately sulcate, striae punctate; lateral declivital margins feebly elevated, bearing 6 – 8 pairs of minute to small, narrow, sharply pointed and recurved spines (often asymmetric) on basal 2 / 3 of declivity; and sulcate area of declivity shagreened, dull. Similar species. Xyleborinus acanthopteron sp. nov., X. echinopterus, X. octiesdentatus, X. kwangruengensis	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	description	Description (female). 2.9 mm long (n = 1); 2.9 × as long as wide. Body uniformly dark brown, legs and antennae yellow brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, alutaceous, dull, densely, coarsely punctate, setose; punctures each a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, deeply impressed. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, as long as funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face. Pronotum 1.11 × as long as wide. In dorsal view basic and parallel-sided, type 2, sides parallel in basal 2 / 3, rounded anteriorly; anterior margin without serrations. In lateral view elongate, disc longer than anterior slope, type 7, summit low, on apical 2 / 5. Anterior slope with densely spaced narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, alutaceous, finely punctate, finely setose, setae short, erect, hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded. Scutellum minute, conical, disconnected from elytra, surrounded by dense mycangial tuft of setae. Elytra: 1.9 × as long as wide, elytra 1.9 × as long as pronotum. Base transverse, medially emarginate near scutellum and mycangial tuft, edge oblique, humeral angles angulate, parallel-sided in basal 3 / 4, then broadly rounded to apex. Disc occupying basal 2 / 3, smooth, shiny, glabrous, unarmed; striae not impressed, glabrous, with moderate punctures separated by 1 – 2 diameters of a puncture; interstriae flat, sparsely finely uniseriate punctate, punctures 1 / 3 as large as those of striae, each bearing an erect hair-like seta, setae approximately as long as width of interstriae 2. Declivital face steeply rounded, moderately sulcate between interstriae 3, shagreened, dull, striae and interstriae feebly setose, setae long, semi-erect hair-like, as long as the distance between suture and interstriae 3 on declivital margins and semi-erect and no longer than an interstriae 2 width on declivital face; three striae present, striae 1 and 2 in irregular rows; strial punctures as larger, deeper than on disc; interstriae 1 and 2 flat, armed by two and three denticles, respectively on declivital base, unarmed on face; interstriae 2 punctate, punctures as described for striae; interstriae 3 forming declivital margin, feebly elevated, 6 – 8 pairs of minute to small, narrow, sharply pointed and recurved spines (often asymmetric) on basal 2 / 3. Posterolateral margin rounded, spinose from interstriae 3 – 7. Legs: procoxae contiguous, prosternal coxal piece inflated. Protibiae obliquely triangular, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with seven moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae flattened, outer margin evenly rounded with eight or nine and nine moderate socketed denticles.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	distribution	Distribution. Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB57109FF7026A57F68F359.taxon	etymology	Etymology. L. dumosus = covered with thornbushes. In reference to the spiny elytral declivity. A variable adjective.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	description	(Figs 69 – 72) http: // zoobank. org / urn: lsid: zoobank. org: act: CB 76 DB 7 C- 26 ED- 4674 - 8985 - E 8377 F 10 C 5 C 0	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	materials_examined	Type material. Holotype, female: JAPAN, Kagoshima Pref., Tarumizu Oonohara, Broadleaf forest, 425 m, 5 November 2001, Yoshikazu Sato coll, ex ETOH-baited trap (NUFM). Paratypes, as holotype except: 17 July 2000 (1, NMNH); 1 Aug 2000, MSUC _ ARC _ 314127 (1, MSUC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	diagnosis	Diagnosis. This species is distinguished by the entire length of interstriae 2 unarmed by tubercles; elytra attenuate from basal half, apex acute; declivital interstriae flat, interstriae 2 not impressed; and denticles on interstriae 1 and 3 moderate. Similar species. Xyleborinus perpusillus (Eggers, 1927)	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	description	Description (female). 2.2 mm long (mean = 2.2 mm; n = 3); 3.38 × as long as wide. Body red brown. Legs and antennae yellow brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, alutaceous, subshiny, finely, moderately punctate, setose; punctures each with a long, erect hairlike seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, deeply impressed. Antennal scape short and thick, shorter than club. Pedicel as wide as scape, shorter than funicle. Funicle 4 - segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate, type 1; segment 1 corneous, occupying basal 3 / 5 and encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face. Pronotum 1.23 × as long as wide. In dorsal view long and rounded frontally, type 7, sides parallel in basal 3 / 4, rounded anteriorly; anterior margin without serrations. In lateral view elongate, disc much longer than anterior slope, type 7, summit prominent, on anterior 2 / 5. Anterior slope with densely spaced narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc shiny, alutaceous, finely punctate, finely setose, setae short, erect, hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded. Scutellum minute, conical, disconnected from elytra, surrounded by dense mycangial tuft of setae. Elytra: 2.15 × as long as wide, 1.75 × as long as pronotum. Base transverse, medially emarginate near scutellum and mycangial tuft, edge oblique, humeral angles rounded, attenuate from basal half, apex acute. Disc occupying basal ~ 2 / 3, smooth, shiny, unarmed; striae not impressed, glabrous, with small punctures separated by 1 – 2 diameters of a puncture; interstriae flat, sparsely finely uniseriate punctate, punctures 1 / 3 those of striae, each bearing erect hair-like setae, setae approximately as long as width of interstriae 2. Declivital face strongly shagreened, gradually sloping, three striae present, striae 1 and 2 subcontiguous, strial punctures minute, shallow, glabrous; interstriae impunctate, denticulate, setose, setae uniseriate, variably sized, setae between denticles from as long as a denticle, setae as base of a denticle 2 × as long as a denticle; interstriae 1 with a row of with six or seven small to moderate denticles along its length; interstriae 2 flat, extremely narrow, nearly obsolete, unarmed along its entire length; interstriae 3 with a row of 4 – 6 small to moderate denticles along its length, their apices acutely pointed, spine-like. Posterolateral margin rounded, denticulate from interstriae 3 – 5. Legs: procoxae contiguous. Protibiae obliquely triangular, broadest at apical 1 / 3; posterior face smooth; apical 1 / 3 of outer margin with five moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae flattened; outer margin evenly rounded with eight and seven moderate socketed denticles, respectively.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	distribution	Distribution. Japan (Kyushu).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	biology_ecology	Host plants. Unknown.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB2710FFF7026A57B55F661.taxon	etymology	Etymology. Named in honor of the Japanese scolytid taxonomist Akira Nobuchi. Noun in apposition.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7024F97CDDF72A.taxon	discussion	New record. INDIA: Sikkim, Gopaldhara, Rungbong Vall., H. Stevens (2, NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7024F97CDDF72A.taxon	distribution	Distribution. Bangladesh, Bhutan, China (Chongqing, Guangxi, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang), India (Sikkim *, Uttarakhand, Uttar Pradesh, West Bengal), Laos, East Malaysia, Myanmar, Nepal, Taiwan, Thailand, Vietnam. Established in the USA.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7025917A40F012.taxon	discussion	New records. CHINA: Yunnan, Xishuangbanna, Mengla, Menglun, Trop. Bot. Gdn, 21.929 N, 101.253 E, 600 m, XTBG 600 - 3 FI, 2. ii. 2018, L. Z. Meng (1, RABC); as previous except: Bubeng, 21.610 N, 101.582 E, 717 m, FIT, BB 600 - 3 FI, 3. iv. 2018 (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7025917A40F012.taxon	distribution	Distribution. China * (Yunnan), India (Tamil Nadu, West Bengal), East Malaysia.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7022497DA7F15E.taxon	distribution	Distribution. China (Fujian, Yunnan *), East & West Malaysia, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF70231D7C26F21A.taxon	discussion	New record. BURMA [= MYANMAR]: N. Toungoo, Pyonchaung Res., 14. XI. 1921, B. M. Bhatia ex Albizia lebbek (felled tree) (1, NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF70231D7C26F21A.taxon	distribution	Distribution. China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan), Japan, Myanmar *, Taiwan, Vietnam; imported and established in USA.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF70231D7C26F21A.taxon	discussion	Remarks. Previously recorded from Pistacia (Anacardiaceae), Ilex (Aquifoliaceae), Quercus sp. (Fagaceae), Cinnamomum (Lauraceae), Broussonetia (Moraceace), and Ligustrum (Oleaceae) (Smith et al. 2020 a). Albizia lebbek (Fabaceae) is a new record.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7020417BFEF31E.taxon	distribution	Distribution. China (Fujian, Jiangxi, Yunnan *), India (Assam, West Bengal), Myanmar, Nepal, Taiwan, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB3710FFF7021427AC4F3AA.taxon	distribution	Distribution. China * (Yunnan), Indonesia (Java, Sumatra), East & West Malaysia, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7026A47CB1F540.taxon	distribution	Distribution. China * (Yunnan), Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7027787C00F58C.taxon	distribution	Distribution. China * (Yunnan), Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7027CC7A99F6D8.taxon	distribution	Distribution. Bhutan, China * (Yunnan), India (West Bengal), Myanmar, Nepal, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7024807AC6F764.taxon	distribution	Distribution. China (Sichuan, Yunnan *), India (Uttarakhand, West Bengal), Nepal, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7025547D25F7B0.taxon	discussion	New record. CHINA, Yunnan, Honghe, Hekou, Dajianshan, 22.908 N, 103.697 E, 2111 m, FIT, DJS 12, 01. v. 2018, L. Z. Meng (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7025547D25F7B0.taxon	distribution	Distribution. China * (Yunnan), India (Meghalaya), Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7022287C0AF0B4.taxon	distribution	Distribution. China (Fujian, Guangdong, Guangxi, Jiangxi, Yunnan *), India (Nicobar Is, West Bengal), Indonesia (Bali, Batoe Is, Java, Maluku, Sulawesi, Sumatra), Laos, East & West Malaysia, New Guinea, Philippines, Solomon Islands, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7023247C88F1C0.taxon	distribution	Distribution. China (Chongqing, Fujian *, Sichuan).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7023F87DE6F2E0.taxon	discussion	New records. CHINA: Fujian, Putian City, Xianyou Co., Jiuli Lake, 25.454 N, 118.819 E, 410 m, ex Castanopsis chinensis, 6. viii. 2020, Zhang, L., Cao Y-F. (1, RABC). VIETNAM: Tam Dao N. P., 13. III. 2016, M. Jusino, in flight, UFFE 11620 (1, MSUC; 1, UFFE).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7023F87DE6F2E0.taxon	distribution	Distribution. China (Fujian *, Guangxi, Hainan), Thailand, Vietnam *.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7023F87DE6F2E0.taxon	discussion	Remarks. The species was previously recorded only from Liquidambar formosana (Altingiaceae) (Smith et al. 2020 a). Castanopsis chinensis (Fagaceae) is a new host record.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710CFF7020D87AD3F32C.taxon	distribution	Distribution. ‘ Borneo’, China (Fujian, Yunnan *), Indonesia (Sumatra), East & West Malaysia.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710DFF7021AC7F6EF541.taxon	discussion	New records. CHINA: Yunnan, Wenshan, Maguan, Gulingqin, 22.732 N, 103.996 E, 582 m, FIT, GLQ 570 - 5 FI, 21. ix. 2018, L. Z. Meng (1, RABC); Honghe, Gejiu, Lvshuihe, 23.029 N, 103.406 E, 555 m, FIT, LSH 52, 02. v. 2018, L. Z. Meng (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB0710DFF7021AC7F6EF541.taxon	distribution	Distribution. China (Hong Kong, Yunnan *), India (West Bengal), Indonesia (Java), Myanmar, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7027787ADBF58D.taxon	distribution	Distribution. Bangladesh, China (Guizhou, Yunnan *), India (Sikkim, West Bengal), Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7027CC7D87F6D9.taxon	distribution	Distribution. Brunei Darussalam, China * (Yunnan), East & West Malaysia.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7024807C33F765.taxon	distribution	Distribution. China * (Yunnan), Taiwan.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7025547DE6F0FD.taxon	distribution	Distribution. United States (California).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7025547DE6F0FD.taxon	discussion	Discussion. Wood (1975) described this species collected from Palo Alto, (Santa Clara Co.), California in 1944 and an additional paratype from Knight’s Landing (Yolo Co.) in 1949. This species was previously known from the type series (Smith & Cognato 2022). This specimen from Calaveras County is the only known nontype specimen. This species is undoubtedly Asian but has yet to be reported outside of California. Smith & Cognato (2022) provide further discussion on the history of this species.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7022FC7B58F109.taxon	distribution	Distribution. China * (Yunnan), Indonesia (Java, Sumatra), East & West Malaysia, Philippines, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7023B07C1AF255.taxon	distribution	Distribution. Bhutan, China (Yunnan), India (Himachal Pradesh, West Bengal), Indonesia (Sumatra), Laos, Myanmar, Nepal *, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7020047A39F2E1.taxon	distribution	Distribution. China (Fujian, Guangxi, Sichuan, Yunnan *), Japan, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB1710DFF7020D87D59F32D.taxon	distribution	Distribution. China * (Yunnan), India (Arunachal Pradesh).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB17102FF7021A57DE6F5F1.taxon	description	GenBank. COI: OP 208788 and CAD: OP 219692.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB17102FF7021A57DE6F5F1.taxon	distribution	Distribution. China * (Yunnan), United States (Florida, Georgia, South Carolina) (Smith & Cognato 2022).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB17102FF7021A57DE6F5F1.taxon	discussion	Remarks. This species was recently described from an adventive population in the United States. Subsequently, a specimen was discovered in the UFFE cryo-collection of scolytines. Identification of the specimen was confirmed with phylogenetic analysis which placed it sister to the US C. nemesis specimens with 100 % bootstrap support (Fig. 73). The Chinese specimen has 6.1 % COI and 0.78 % CAD sequence difference as compared to the US specimens. This amount of difference is less than 10 % COI and 2 % CAD differences which are suggested thresholds for xyleborine species limits (Cognato et al. 2020 b).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7027E87CB1F63D.taxon	distribution	Distribution. China * (Yunnan), Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7024BC7D19F765.taxon	discussion	New records. CHINA: Fujian, Wuyishan City, Wuyi Mtn Nat. Res., Guadun Vill., 27.743 N, 117.634 E, 1220 m, ex Castanopsis fargesii, 10. vii. 2018, Lai, S-C., Zhang, L., Lv Jia, Hu, K-P. (2, RABC); Yunnan, Xishuangbanna, Mengla, Bubeng, 21.615 N, 101.579 E, 860 m, FIT, BB 800 - 2 FI, 24. iv. 2018, L. Z. Meng (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7024BC7D19F765.taxon	distribution	Distribution. China * (Fujian, Yunnan), Laos, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7025547B22F005.taxon	distribution	Distribution. Brunei Darussalam, China * (Yunnan), Laos *, East & West Malaysia, Thailand, Vietnam (Smith et al. 2020 ab, as Cyclorhipidion pruinosulum (Browne )).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7025547B22F005.taxon	discussion	Remarks. Xyleborus pruinosulus Browne, 1979, was synonymised with this species by Smith et al. (2022).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7022B47B7FF16D.taxon	discussion	New record. LAOS: Champasak Prov., 15 ° 02 ′ N 106 ° 31 - 34 ′ E, Ban Nong Panouan env., 770 - 800 m, 10 - 17. vi. 2010, M. Geiser, D. Hauck, ex coffee plantations, secondary degraded primary forest, hand collecting, sweeping, beating, coll. At flowering trees and in small water puddles, light trapping (1, NHB). Distribution. ‘ Borneo’, India (Andaman Islands), Indonesia (Sumatra), Laos *, East & West Malaysia.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF70236C7C45F255.taxon	distribution	Distribution. Bhutan, China (Yunnan), Indonesia (Java, Sumatra, Sulawesi), Laos, East & West Malaysia, Nepal *, Philippines, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7020057B27F2E1.taxon	distribution	Distribution. China (Chongqing, Guizhou), Laos *, East & West Malaysia, New Guinea, Taiwan, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBE7102FF7020D97CD4F3E5.taxon	distribution	Distribution. Bangladesh, China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Jiangxi, Yunnan), India (Andaman Is, Arunachal Pradesh, Assam, Tamil Nadu, Uttarakhand, West Bengal), Indonesia (Java), Japan (Ryukyu Is), East & West Malaysia, Nepal *, New Guinea, South Korea, Sri Lanka, Taiwan, Thailand, Vietnam. Imported to Africa (South Africa, Tanzania).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF7026A47CD7F540.taxon	distribution	Distribution. Bhutan, China (Yunnan), India (Arunachal Pradesh, Assam, Meghalaya, Sikkim *, West Bengal), Laos, Myanmar, Taiwan, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF7027787F71F644.taxon	distribution	Distribution. Recorded in Southeast Asia from China * (Yunnan), India (Kerala), Laos, Myanmar, Taiwan, Thailand, Vietnam. It also occurs in Malaysia and the Philippines, through Indonesia to New Guinea and the Solomon Islands.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF7024747F5DF6AC.taxon	distribution	Distribution. China (Jiangxi, Yunnan *), Indonesia (Sumatra), Japan, East & West Malaysia, Solomon Islands, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF70252C7F40F04C.taxon	distribution	Distribution. China (Guangxi, Jiangxi, Yunnan *), India (Andaman Is, West Bengal), Indonesia (Java, Maluku, Sumatra), Japan, Laos *, East & West Malaysia, Myanmar, New Guinea, Philippines, Taiwan, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF70220C7A92F108.taxon	discussion	New records. CHINA: Fujian, Longyan City, Wuping Co., Liangye Mtn Nat. Res., 25.142 N, 116.135 E, 403 m, ex Sapium discolor, 29. viii. 2019, Zhang, L., Liu, Z-H. (1, RABC); as previous except: ex Liquidambar formosana (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF70220C7A92F108.taxon	distribution	Distribution. Bangladesh, China (Fujian *, Guizhou, Yunnan), India (Arunachal Pradesh, Assam, West Bengal), Laos, West Malaysia, Sri Lanka, Taiwan, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF70220C7A92F108.taxon	discussion	Remarks. The species is polyphagous (Maiti & Saha 2004), but has not previously been recorded from Triadica cochinchinensis (= Sapium discolor) (Euphorbiaceae) or Liquidambar formosana (Altingiaceae).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF7023B07D55F20C.taxon	discussion	New records. CHINA: Guangxi, Shiwandashan, 15. iv. 2015, You Li, ex Fagaceae, UFFE 19151 (3, UFFE). INDIA: Assam, Pawai Tea Est., 6. xii. 1983, 932 / 6, C. I. E. A. 15663, ex tea (1, NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF7023B07D55F20C.taxon	distribution	Distribution. Bangladesh, China (Guangxi *, Yunnan), India (Assam *, West Bengal), Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF7023B07D55F20C.taxon	discussion	Remarks. Previously recorded only from Chrysophyllum (Sapotaceae) and Cinnamomum (Lauraceae). Fagaceae and tea (Camellia sinensis) (Theaceae) are new records.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7100FF70204C7A38F310.taxon	distribution	Distribution. China (Chongqing, Sichuan, Yunnan *), Indonesia (Sumatra), Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7101FF7021487C4AF541.taxon	discussion	New records. CHINA: Guangxi region, Miaoershan, S slope, 20 - 27 Jun 1997, 800 - 1300 m, Bolm (1, NHB). LAOS: Bokeo prov., 5 km W Ban Toup, Bokeo Nature Reserve, 500 - 700 m, 20 ° 27 - 28 ′ N, 100 ° 24 ′ E, 4. - 18. v. 2011, NHB Basel, Laos 2011 expedition, M. Brancucci, M. Geiser, D. Hauck, Z. Kraus, A. Phantala, E. Vongphachan (1, NHB); Phongsaly prov., PHONGSALY env., 21 ° 41 ′ N, 102 ° 06 ′ E, ~ 1500 m, 6. - 17 v. 2004, Vit. Kubáň (1, NHB).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBC7101FF7021487C4AF541.taxon	distribution	Distribution. China (Fujian, Guangdong, Guangxi *, Hong Kong, Hunan, Jiangxi, Yunnan, Zhejiang), Japan, Laos *, South Korea, Taiwan, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7027797D6DF5A9.taxon	discussion	New records. CHINA: Fujian, Sanming City, Taining Co., Emei Mtn Nat. Res., 27.006 N, 117.071 E, 1430 m, 12. viii. 2020, Zhang, L., Cao, Y-F. (2, RABC); Yunnan, Xishuangbanna Tropical Botanical Garden, 20. vii. 2014, C. Bateman, UFFE 11682 (1, UFFE).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7027797D6DF5A9.taxon	distribution	Distribution. China * (Fujian, Yunnan), Japan, Taiwan.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7024117A1FF7DD.taxon	discussion	New records. CHINA: Fujian, Longyan City, Wuping Co., Liangye Mtn Nat. Res., 25.142 N, 116.135 E, 403 m, ex Cinnamomum camphora, 31. viii. 2019, Zhang, L., Liu, Z-H. (2, RABC); Guizhou, Guiyang, Guiyang Forest Park, 29. ix. 2015, You Li, ex Liquidambar formosana, UFFE 11945 (1, UFFE); Hainan: Qiongzhong, Jinping, 27. x. 2016, You Li, ex Litchi, UFFE 16583 (1, UFFE); Yunnan, Honghe, Gejiu, Lvshuihe, 23.030 N, 103.405 E, 538 m, FIT, LSH 45, 29. v. 2018, L. Z. Meng (1, RABC); Yunnan, Wenshan, Maguan, Gulingqin, 22.732 N, 103.996 E, 582 m, FIT, GLQ 59, 18. vi. 2018, L. Z. Meng (1, RABC).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7024117A1FF7DD.taxon	distribution	Distribution. China (Fujian *, Guizhou *, Hainan *, Hong Kong, Jiangxi, Yunnan *).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7024117A1FF7DD.taxon	discussion	Remarks. Previously recorded only from Fagaceae (Smith et al. 2020 a). Liquidambar formosana (Altingiaceae), Cinnamomum camphora (Lauraceae), and Litchi sp. (Sapindaceae) are new records.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF70259D7FF1F005.taxon	distribution	Distribution. ‘ Borneo’, Brunei Darussalam, China * (Yunnan), Indonesia (Java, Sulawesi), East & West Malaysia, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7022B47F40F16D.taxon	distribution	Distribution. ‘ Borneo’, Brunei Darussalam, China * (Yunnan), Indonesia (Sumatra), East Malaysia, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF70236B7F9DF27B.taxon	distribution	Distribution. China * (Yunnan), Indonesia (Maluku), Laos, East & West Malaysia, Myanmar, New Guinea, Sri Lanka, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7020627D85F28A.taxon	distribution	Distribution. China * (Yunnan), Indonesia (Java), West Malaysia, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7021367A8EF3AA.taxon	discussion	New record. INDIA: [West] Bengal: Matelli T. E. [Tea Estate] Northl, 14. XI. [19] 84, No. 970.6, on Albizia lebbeck, C. I. E. A. 1663 (1, NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7021367A8EF3AA.taxon	distribution	Distribution. Brunei Darussalam, India * (West Bengal), Indonesia (Java, Sulawesi), Japan, East & West Malaysia, New Guinea, Philippines, Sri Lanka, Taiwan, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBD7101FF7021367A8EF3AA.taxon	discussion	Remarks. Previously recorded from Buchanania, Mangifera (Anacardiaceae), Swietenia (Meliaceae), and Castanopsis (Fagaceae) (Smith et al. 2020 a). Albizia lebbeck (Fabaceae) is a new host record.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7026A47CC9F51C.taxon	discussion	New records. [MYANMAR]: N. Toungoo, Saing Yane Res., San Thwin coll., 28. II. [19] 31, ex. Dipterocarpus turbinatus (3, NHMUK), 17. III. [19] 31 (1, NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7026A47CC9F51C.taxon	distribution	Distribution. Cambodia, India (Andaman Is, Assam), Indonesia (Sumatra), East & West Malaysia, Myanmar *, Philippines, Sri Lanka, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF70275C7C1CF5A8.taxon	distribution	Distribution. China * (Yunnan), Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7024107FECF6AC.taxon	discussion	New record. CHINA: Guangdong, Quangzhou, South China Agricultural University, 28. v. 2017, You Li, ex Elaeocarpus, UFFE 17028 (1, UFFE).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7024107FECF6AC.taxon	distribution	Distribution. China (Fujian, Guangdong *, Jiangxi, Yunnan).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7024107FECF6AC.taxon	discussion	Remarks. Previously recorded from Fagaceae and Schima superba (Theaceae). Elaeocarpus (Elaeocarpaceae) is a new host record.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF70252C7A4FF794.taxon	discussion	New records. CHINA: Yunnan, Honghe, Hekou, Dajianshan, 22.909 N, 103.697 E, 2130 m, FIT, DJS 43, 12. v. 2018, L. Z. Meng (1, RABC). LAOS: NE, Hua Phan prov., 20 ° 12 ′ N, 104 ° 01 ′ E, PHU PHAN Mt., ~ 1750 m, 17. v. - 3. vi. 2007, Vit. Kubáň (1, NHB). Distribution. China (Sichuan, Yunnan *), India (Assam), Laos *, Nepal, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7025C47FC2F0B4.taxon	discussion	New records. CHINA: Yunnan, Honghe, Hekou, Dajianshan, 22.908 N, 103.697 E, 2111 m, FIT, DJS 10 - 3, 12. v. 2018, L. Z. Meng (1, RABC). [MYANMAR]: [Kachin State], Myitkyina, Hpemaw, 24. xi. [19] 30, Po Yone, F. R. S. No. 500, ex Juglans regia (3, NHMUK).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7025C47FC2F0B4.taxon	distribution	Distribution. China (Chongqing, Guizhou, Sichuan, Yunnan *), Myanmar *, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7025C47FC2F0B4.taxon	discussion	Remarks. Previously recorded from Pinus armandi (Pinaceae) (Smith et al. 2020 a). Juglans regia (Juglandaceae) is a new host record.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7023247CD2F19C.taxon	discussion	New records. CHINA: Yunnan, Honghe, Hekou, Dajianshan, 22.908 N, 103.697 E, 2136 m, FIT, DJS 33, 12. v. 2018, L. Z. Meng (3, RABC). LAOS: NE, Hua Phan prov., 20 ° 12 ′ N, 104 ° 01 ′ E, PHU PHAN Mt., ~ 1750 m, 17. v. - 3. vi. 2007, Vit. Kubáň (16, NHB).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7023247CD2F19C.taxon	distribution	Distribution. China * (Yunnan), Laos *, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF7023DC7FCAF2BC.taxon	distribution	Distribution. Bangladesh, China (Fujian, Guangdong, Guangxi, Hong Kong, Hunan, Jiangxi, Sichuan, Yunnan *), India (Assam, West Bengal), Japan, Myanmar, South Korea, Taiwan, Thailand, Vietnam. Imported to and established in USA.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF70213C7F9CF3E4.taxon	discussion	New records. LAOS: NE, Hua Phan prov., 20 ° 12 ′ N, 104 ° 01 ′ E, PHU PHAN Mt., ~ 1750 m, 17. v. - 3. vi. 2007, Vit. Kubáň (5, NHB); as previous except: 20 ° 13 ′ 09 - 19 ″ N 103 ° 59 ′ 54 ″ E, 1480 - 1550 m, PHOU PANE Mt., 9. - 16. vi. 2009, David Hauck (2, NHB).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBA7106FF70213C7F9CF3E4.taxon	distribution	Distribution. Laos *, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7026A47D1EF5F0.taxon	discussion	New record. CHINA: Hainan: Qiongzhong, Wanling, 26. x. 2016, You Li, ex Acacia mangium, UFFE 16949 (1, UFFE).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7026A47D1EF5F0.taxon	distribution	Distribution. Throughout tropical parts of the Afrotropical, Australian, Oriental regions and Costa Rica (Kirkendall & Jordal 2006); recorded in Southeast Asia from Bangladesh, Bhutan, Cambodia, China (Guangxi, Hainan *, Hong Kong, Shanxi, Yunnan), India (Andaman Is, Assam, Chhattisgarh, Haryana, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Nicobar Is, Tamil Nadu, Uttarakhand, Uttar Pradesh, West Bengal), Laos, Myanmar, Nepal, Taiwan, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7027E87C1CF63C.taxon	distribution	Distribution. China * (Yunnan), Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7024BC7C53F764.taxon	discussion	New records. LAOS: NE, Hua Phan prov., 20 ° 12 ′ N, 104 ° 01 ′ E, PHU PHAN Mt., ~ 1750 m, 17. v. - 3. vi. 2007, Vit. Kubáň (2, NHB). Xieng Khouang prov., 19 ° 37 - 8 ′ N, 103 ° 20 ′ E, 30 km NE Phonsavan: Ban Na Lam → Phou Sane Mt., 1300 - 1500 m, 10. - 30. v. 2009, M. Brancucci (1, NHB).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7024BC7C53F764.taxon	description	Distribution. China (Hainan), Laos *.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7025547A82F7B0.taxon	distribution	Distribution. China (Guangdong, Guizhou *, Hong Kong), Japan, South Korea, Taiwan, Vietnam *.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7022287C8EF151.taxon	distribution	Distribution. In temperate and tropical regions around the world. Within Southeast Asia it is recorded from China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang), India (Karnataka, Kerala, Tamil Nadu), ‘ Indochina’, Laos *, South Korea, Taiwan, Thailand, Vietnam. Established in the Neotropics, USA and Europe.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7023087A05F19C.taxon	distribution	Distribution. China (Yunnan), Indonesia (Java, Moluccas, Sulawesi), Laos *, Thailand.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7023DC7C3AF374.taxon	distribution	Distribution. From the Indian subcontinent and southern China through Southeast Asia and Indonesia west of Wallace’s line, Philippines, Taiwan. Presumably imported to Australia. Within Southeast Asia it is recorded from China (Chongqing, Fujian, Guangdong, Guizhou *, Hainan, Hong Kong, Jiangxi, Sichuan, Yunnan), India (Andaman Is, Assam, Karnataka, Meghalaya, Sikkim, Tamil Nadu, Uttarakhand, West Bengal), Laos, Myanmar, Taiwan, Thailand, Vietnam. Also recorded from Australia, Japan (Ryukyu Is.), Indonesia, West Malaysia, New Guinea, Philippines, and Sri Lanka.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFBB7107FF7021647ABAF380.taxon	distribution	Distribution. China (Hainan, Hong Kong, Yunnan), Indonesia (Java), Laos *, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB87104FF7026A47B73F51C.taxon	discussion	New records. LAOS: NE, Hua Phan prov., 20 ° 12 ′ N, 104 ° 01 ′ E, PHU PHAN Mt., ~ 1750 m, 17. v. - 3. vi. 2007, Vit. Kubáň (3, NHB). Phongsaly prov., PHONGSALY env., 21 ° 41 - 2 ′ N, 102 ° 06 - 8 ′ E, ~ 1500 m, 28. v. - 20. vi. 2003, Vit. Kubáň (15, NHB).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB87104FF7026A47B73F51C.taxon	distribution	Distribution. China (Fujian, Jiangxi, Xizang), India (West Bengal), Laos *, Taiwan, Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB87104FF70275C7DF5F5A8.taxon	distribution	Distribution. China * (Fujian), India (Meghalaya), Myanmar, Nepal.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB87104FF7024107D01F6F4.taxon	distribution	Distribution. China (Hong Kong, Yunnan *), Thailand, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB87104FF7024E47D3FF7DC.taxon	discussion	New records. CHINA: Guizhou, Huaxi, Guiyanag, SLP, 15. vi. 2018, Shang Tian, ex trap, UFFE 22240 (1, UFFE); as previous except: 12. viii. 2018, UFFE 22249 (1, UFFE); as previous except: 6. viii. 2018, Xinyao Gu, UFFE 22259 (4, UFFE).	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
7D518786FFB87104FF7024E47D3FF7DC.taxon	distribution	Distribution. China (Guizhou *, Yunnan), Myanmar, Vietnam.	en	Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong, Cognato, Anthony I. (2022): New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa 5209 (1): 1-33, DOI: 10.11646/zootaxa.5209.1.1
