taxonID	type	description	language	source
7C6D8795FA64FFEEA2A2FAB8C93D20E5.taxon	type_taxon	Type species: Cassisotropis aciformis Taszakowski et al. sp. nov., here designated.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA64FFEEA2A2FAB8C93D20E5.taxon	diagnosis	Diagnosis. Cassisotropis is recognized within Cylapinae by the following set of features: head, pronotum, thoracic pleura, mesoscutum, scutellum and abdomen with net-like sculpturing created by microsetae (Figs 2 B – F); head and anterior part of pronotum with several long, erect, cylindrical, regularly distributed, apically serrate bristles (Figs 2 A – E); head with indistinct tubercles on vertex and frons (Figs 2 B – E); clypeus with moderately developed longitudinal keel medially (Fig. 2 C, D); pronotal collar separated from remainder of pronotum by weakly developed sulcus (Figs 1 A, B, Fig. 2 B); calli upraised and conical (Figs 1 C, D, Figs 2 A, B); posterior lobe with relatively small, weakly raised tubercles, each situated mediolaterally, bordering posterior margin (Fig. 2 B); metepisternal scent gland evaporative areas triangular (Fig. 2 F); embolium wide (Figs 1 A, B); tarsus two-segmented, tarsomere II not divided (Fig. 2 H); aedeagus with theca strongly sclerotized with sail-like process subapically (Fig. 3 C); sclerotized part of ductus seminis inside endosoma long, its apex reaching subapical portion of endosoma (Fig. 3 D); endosoma with long, needle-like spicule (Fig. 3 D); left paramere sensory lobe with strongly developed protuberance (Fig. 3 F); first and second valvulae (gonapophyses 8 and 9) sharply pointed, not serrate apically (Figs 4 H, I); first valvulae connected by membranous structure (Figs 4 A, G).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA64FFEEA2A2FAB8C93D20E5.taxon	description	Description. Male. Macropterous. TEXTURE AND VESTITURE. Dorsum covered with sparse, short, irregularly distributed setae (Figs 1 A, 2 B – D). Head, pronotum, thoracic pleura, mesoscutum, scutellum, and abdomen with net-like sculpturation based on microsetae (Figs 2 B – D). Head. Vertex, frons, and clypeus with several long, erect, cylindrical, regularly distributed, apically serrate bristles (Figs 2 A – D); antennal segment I almost glabrous on basal one-third, covered with sparse semirecumbent setae; antennal segment II covered with setae similar to those on segment I but denser; segments III and IV missing in holotype (Figs 2 A – D). Thorax. Pronotum. With four pairs of long, erect, cylindrical, regularly distributed, apically serrate bristles that are placed on pronotal calli, humeral angles, anterior portion of lateral margin and anterior part of pronotal calli (Figs 2 B, D). Thoracic pleura. Clothed with sparse, short setae. Legs. Coxae and femora covered with fine, recumbent, rather short setae; tibiae and tarsi with relatively dense, stiff, semirecumbent setae, tibiae also with small black spinules organized in several straight and regular rows (Figs 2 A, B). Abdomen. Clothed with relatively dense, moderately long semirecumbent setae (Fig. 2 A). STRUCTURE. Body elongate oval (Fig. 1 A). Head. Elongated horizontally (Figs 1 A, C); vertex and frons with indistinct shallow longitudinal sulcus medially; vertex not carinate posteriorly with two indistinct small contiguous tubercles on medial portion of lateral margin; frons with two pairs of indistinct small tubercles bordering sides of medial longitudinal furrow (Figs 2 B – D); clypeus with moderately developed medial rib along entire length, clypeal base situated above antennal insertions and ventral margin of eye (Figs 2 C, D); antennal insertion contiguous with sulcus between maxillary and mandibular plates (Figs 2 C, D); eye relatively large, reniform, its ventral margin reaching gula; mandibular plate without sulcus posteriorly (Fig 2 D); antennal segment I mostly cylindrical, somewhat narrowed basally (Fig. 2 D); segment II cylindrical, relatively stout, as thick as segment I (Fig. 2 A); labium thin and long, reaching beyond metacoxae; segment I reaching xyphus, subdivided near medial part (Fig. 2 A); segment II subdivided subapically. Thorax. Pronotum. Trapezoid, rather short; collar flattened, relatively broad, separated from remainder of pronotum by weakly developed, shallow sulcus; anterior lobe with small pit between calli; calli well developed, upraised, conical, sharply pointed; remainder of anterior lobe sloping toward lateral margin; lateral margin strongly carinate; humeral angle broad; posterior lobe with relatively small, weakly raised tubercles, each situated mediolaterally, bordering posterior margin (Fig. 2 B). Scutellum. Somewhat convex medially (Fig. 2 B). Thoracic pleura. Mesepimeral spiracle slit-like; scent gland evaporative area triangular, occupying entire ventral portion of metepisternum; metepimeron narrow, carinate posteriorly. Hemelytron. Claval commissure as long as scutellum; embolium broad; membrane with both cells well developed, major cell relatively short, its apex not reaching beyond apex of cuneus (Fig. 1 A). Legs. Tarsus two-segmented; tarsomere I about two times shorter than tarsomere II; tarsomere II not subdivided (Fig. 2 H); pretarsal structure with the well-developed middle row of tiles on the unguitractor plate; pretarsal claw with well-developed subapical claw tooth (Fig. 2 I). Abdomen. Genitalia. Pygophore trapezoid, its dorsal wall as long as ventral wall, aperture terminated posteriorly (Figs 2 G, 3 A, B); aedeagus elongated; theca strongly sclerotized with relatively large sail-like process placed at right hand side subapically; ductus seminis moderately long and thin; distal, sclerotized part of ductus seminis inside endosoma relatively thick, long, apex reaching subapical portion of endosoma, weakly curved, mostly cylindrical, broadened and irregularly shaped; secondary gonopore undifferentiated; endosoma weakly membranous with relatively long, needle-like sclerite originating from membranous oval lobe placed apically of endosoma (Figs 3 C, D); left paramere stout, C-shaped, apical process moderately long, straight, weakly tapering in lateral view, paramere body with small keel dorsally on area adjacent to apical process, sensory lobe with massive, long protuberance (Figs 3 E – G); right paramere relatively thin, C-shaped, apical process relatively long and thick in lateral view, sensory lobe weakly developed, round (Figs 3 H, I). Female. Similar to male in texture, vestiture, and structure (Figs 1 B, D). Head. Antennal segment II thinner than segment I, covered with sparse semirecumbent setae (Figs 1 B, D). Abdomen. Genitalia. Genital chamber (or bursa copulatrix) semicircular, mostly membranous, rather thin, not reaching laterally beyond rami of first valvulae (ra 1, g 8 ra), sclerotized rings large (sr), paired, each occupying lateral one-third of genital chamber, ovoid, weakly rimmed (Figs 4 A – C, E); lateral oviducts (odl) not contiguous with each other, their apical part reaching weakly beyond lateral margin of genital chamber, dorsal sac (ds) (sensu Pluot-Sigwalt & Matocq 2017) present, weakly developed, membranous, semiovoid, spermathecal gland (sgl) originating from posterior part of dorsal sac (Figs 4 B – D); posterior wall of genital chamber with interramal sclerite (irs) strongly sclerotized, triangular (Fig. 4 F); vestibulum without any sclerite around vulva; first and second valvulae (gonapophyses 8 and 9) (va 1, gp 8) with apex sharpened and not serrate (Figs 4 H, I); first valvulae (gonapophysis 8) connected by membrane (m) (Figs 4 A, G).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA64FFEEA2A2FAB8C93D20E5.taxon	etymology	Etymology. This genus is dedicated to Dr. Gerasimos Cassis in recognition of his great contribution to the study of the Heteroptera. The name is the combination of Cassis (Dr Gerasimos Cassis) and the Greek noun tropis, meaning keel (in reference to the genus Peritropis). The gender is feminine.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA64FFEEA2A2FAB8C93D20E5.taxon	discussion	Remarks. Cassisotropis and Infernotropis possess the characters presented by Gorczyca (2000) as diagnostic for Fulviini, i. e., the porrect head, antenna shorter than body length, with the segment II longest, labium thin, long, reaching at least beyond metafemora, forecoxae and forefemora enlarged. They also have a set of characters that are commonly found in the tribe in its current concept, such as: labial segments I and II subdivided; clypeal base situated above antennal insertions and ventral margin of eye; antennal insertion contiguous with sulcus between maxillary and mandibular plates; eye enlarged, reniform, its ventral margin reaching gula; mandibular plate without sulcus posteriorly; the genital capsule with the dorsal wall long, only weakly shorter than ventral wall, genital opening terminal in orientation; bursa copulatrix relatively thin, not reaching laterally beyond rami of the first valvulae (Wolski & Henry 2012, 2015; Wolski et al. 2017; Namyatova & Cassis 2019, 2021, 2022; Wolski 2021). Therefore, both genera are here included in the tribe Fulviini. Cassisotropis is similar and likely closely related to a group of fulviine genera having the dorsal surface verrucose or with net-like sculpturation based on tuberculate microsetae as noted among others in Peritropis Uhler, 1891 (Gorczyca & Wolski 2007: Fig. 9; Wolski & Henry 2012: Figs 27 – 32), species belonging to the anthocoroides group of the genus Fulvius Stål, 1862 (Pluot-Sigwalt & Chérot 2013), genera Euchilofulvius Poppius, 1909, Peritropisca Carvalho & Lorenzato, and Rewafulvius Carvalho, 1972 (Wolski & Gorczyca 2014: Figs 29 – 31), Sulawesifulvius Gorczyca, Chérot & Štys, 2004 (Wolski et al. 2017: Fig. 7), Ceratofulvius Reuter, 1902 (Namyatova & Cassis 2019: Fig. 18 J) or Schmitzofulvius Gorczyca, 1998 (Wolski, pers. obs.). Within these genera, Cassisotropis is most similar to Peritropisca Carvalho and Lorenzato, 1978 known from Papua New Guinea in both sharing following set of characters: vertex and frons with longitudinal medial sulcus (Figs 2 C, D; Wolski & Gorczyca 2014: Figs 23, 24), pronotal calli upraised, conical, and sharply pointed (Figs 2 A, B; Wolski & Gorczyca 2014: Figs 3, 4), posterior lobe of pronotum with two well-developed mediolateral keels (Fig. 2 B; Wolski & Gorczyca 2014), embolium wide (Figs 1 A, B), and tarsus two-segmented with the second tarsomere not subdivided. Cassisotropis differs from Peritropisca in the lack of the transverse furrow along the posterior margin of head, medial and longitudinal keel on the posterior lobe of pronotum (as in Wolski & Gorczyca 2014: Figs 23, 24), and the metathoracic scent gland evaporative area relatively broad, occupying ventral part of metepisternum (Fig. 2 F), whereas in Peritropisca it is narrow, restricted to posterior portion of metepisternum (Wolski & Gorczyca 2014: Fig. 33). Both genera can also be easily distinguished by the shape of the male genitalia (Fig. 3; Wolski & Gorczyca 2014: Figs 11 – 12). Among the Madagascan genera, Cassisotropis is most similar to Schmitzofulvius sharing (apart from the verrucose dorsal surface) the upraised, conical and sharply pointed calli (Figs 2 A, B; Gorczyca 1999). The new genus can be distinguished primarily by the embolium not distinctly narrowed basally and the lack of translucent patch on corium (Gorczyca 1999, 1998).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	description	(Figs. 1 – 4)	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	materials_examined	Material examined. Holotype (♂): ’ CE MADAGASCAR 2014 \ Toamasina pr.; Andisabe- \ Mantadia N. P. 961 m \ Analamazaotra forest \ 18 ° 56.324 ' S 48 ° 25.396 ' E \ 961 m; 9. – 12. ii., M. Trýzna lgt. ’; Paratype: (♀): same as holotype.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	diagnosis	Diagnosis. See the generic diagnosis.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	description	Description. Male. COLORATION. Dorsum yellow fuscous with black and brown areas (Fig. 1 A). Head. Contrastingly yellow; vertex with small reddish patch on medial part of posterior margin, with relatively large fuscous patch in middle of area adjacent to frons, and with two black patches, each contiguous with posterior part of eye; frons with small reddish patch on medial portion of area adjacent to vertex, with two black patches, each contiguous with medial part of eye and with two black patches places between antennal insertion and eye; mandibular plate with two reddish patches medially; maxillary plate and buccula yellow anteriorly, dark brown posteriorly; gula mostly yellowish with dark brown areas; antennal segment I yellow, tinged with red subapically; segment II yellow with relatively broad reddish annulation subapically and black fuscous narrow annulation apically; segments III and IV missing in holotype; labium dark brown weakly tinged with dark yellow (Figs 1 A, C). Thorax. Pronotum. Black with broad yellow fuscous band medially being paler on collar and anterior lobe; humeral angle yellow (Fig. 1 A). Mesoscutum and scutellum. Black; mesoscutum indistinctly tinged with yellow; scutellum with yellow spot apically (Fig. 1 A). Thoracic pleura. Pro- and mesopleuron black; mesepimeron weakly and narrowly tinged with yellow dorsally and ventrally; metepisternum fuscous anteriorly, dark yellow posteriorly, scent gland evaporative areas and peritreme yellow; metepimeron dark brown. Hemelytron. Yellow fuscous with brown and black areas; clavus, endocorium and cuneus fuscous, entirely mottled with yellow and with brown mottling medially; clavus with relatively large, yellow patch subapically; endocorium with blackish patch apically; cuneus with blackish tinge near medial part on inner margin; exocorium and embolium yellow, tinged with fuscous and brown; membrane with indistinct yellow tinges (Fig. 1 A). Legs. Coxae yellow; fore coxa weakly tinged with red; femora dark brown fuscous with two irregular annulations near apex, apical part weakly tinged with red; profemur weakly and narrowly tinged with yellow; meso- and metafemora with well-developed yellow tinge; tibiae and tarsi dark yellow (Figs 1 A, C). Abdomen. Reddish, tinged with fuscous, pygophore fuscous (Fig. 1 C). Female. Similar to male in coloration (Figs 1 B, D). Measurements. Male (n = 1) / Female (n = 1) Body. Length, clypeus – apex of membrane: 4.04 / 4.47; width: 1.72 / 1.80. Head. Length: 0.68 / 0.73; width, including compound eyes: 0.75 / 0.74; vertex width: 0.34 / 0.40; eye dorsal width: 0.21 / 0.17; height in lateral view: 0.41 / 0.42. Antenna. Length of segment I: 0.46 / 0.48; II: 1.27 / 1.27; III: – / 0.53; IV: – / –. Labium. Total length: – / –; length of segment I: 0.54 / 0.63; II: 0.66 / –; III: 0.58 / –; IV: 0.46 / –. Pronotum. Mesal pronotal length: 0.59 / 0.63; lateral pronotal margin length: 0.65 / 0.72; posterior pronotal maximal width (straight): 1.33 / 1.35. Scutellum. Anterior width: 0.66; length (mesoscutum excluded): 0.44 / 0.51, length (mesoscutum included): 0.78 / 0.81. Hemelytron. Claval commissure: 0.72 / 0.84; heme length: 3.15 / 3.31; corium length: 1.74 / 1.79; outer cuneal margin length (straight): 0.64 / 0.67; cuneus width: 0.70 / 0.66; cell length: 0.80 / 0.87; cell width: 0.33 / 0.41. Hind leg. Femur length: 1.36 / 1.43; tibia length: 1.98 / 2.04; tarsus length: 0.42 / 0.39; length of tarsal segment I: 0.19 / 0.19; II: 0.30 / 0.31.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	biology_ecology	Biology. Unknown; specimens were collected in rainforest.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	distribution	Distribution. Central west Madagascar (Fig. 9).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA66FFEAA2A2FAB8CF8223C1.taxon	etymology	Etymology. The specific epithet is derived from the Latin acus (needle) and forma (shape) and is used to denote endosoma with needle-like sclerite apically.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA6CFFE6A2A2F9EECF3625F1.taxon	type_taxon	Type species: Infernotropis madagascariensis Taszakowski et al. sp. nov., here designated.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA6CFFE6A2A2F9EECF3625F1.taxon	diagnosis	Diagnosis. Recognized by the following combination of characters: body large (> 6.20 mm); dorsum clothed with fine, adpressed, regularly distributed, moderately dense scale-like, silvery setae that are distinctly broadened toward apex (Figs 5, 6 A, D); antennal segment II stout, with basal half cylindrical and apical half strongly flattened dorsoventrally (Figs 5, 6 A), its outer surface in males with dense, long, erect and semirecumbent setae (Fig. 6 A); pronotum distinctly narrowing toward anterior part, bell-like (Figs 5 A, B); tarsus three-segmented; tarsomere I about as long as tarsomeres II and III combined (Fig. 6 E); sclerotized portion of ductus seminis inside endosoma with basal twothirds thin and cylindrical, apical one-third broadened, irregularly shaped with apico-lateral portions well expanded beyond middle of endosoma; apical part of endosoma with large, inflated, serrated lobe embraced basally by apicolateral parts of distal portion of ductus seminis; endosoma also with long, flattened sclerotized lobe originated on its base and terminated subapically (Figs 7 A – C); dorsal labiate plate (dlp) with large sclerite occupying most of it, laterally possessing rib-like projections directed dorsad (Figs 8 B, D, E); ventral labiate plate (vlp) with elongate sclerotization laterally (sv) (Figs 8 B – E).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA6CFFE6A2A2F9EECF3625F1.taxon	description	Description. Male. Macropterous. TEXTURE AND VESTITURE. Dorsum weakly rugose, clothed with fine, adpressed, regularly distributed, moderately dense scale-like, silvery setae that are distinctly broadened toward apex (Figs 5, 6 A, D). Head. Weakly rugose (Figs 6 B, C); antennal segment I covered with short and moderately dense, adpressed simple setae; segment II with inner surface clothed with setae similar to those present on segment I, outer surface with dense, long, erect and semirecumbent setae; segment III covered with relatively dense and long, erect and semirecumbent fine setae (Figs 5 A, C, 6 A) (segment IV missing in male specimen). Thorax. Thoracic pleura. Weakly rugose, covered with sparse, fine, recumbent setae (Fig. 6 A). Hemelytron. Clavus and endocorium with setae organized in regular rows along their veins; exocorium, embolium, and clavus with more irregularly distributed vestiture (Figs 5 A, C, 6 A). Legs. Coxae and femora covered with fine, recumbent, rather short setae; tibiae and tarsi with relatively dense, stiff, semirecumbent setae, tibiae also with small black spinules organized in several straight and regular rows (Figs 5 A, C, 6 A). Abdomen. Covered with relatively long and dense, recumbent and semirecumbent setae (Fig. 6 A). STRUCTURE. Body stout, elongate oval (Fig. 5 A). Head. Porrect, in anterior view about twice wider than high, frons forming obtuse angle with vertex; vertex with weakly developed, longitudinal sulcus medially; base of clypeus situated above antennal insertions and ventral margin of eye; antennal insertion contiguous with sulcus between maxillary and mandibular plates; eye relatively large, reniform, its ventral margin reaching gula; mandibular plate without sulcus posteriorly; antenna short, equal to median of body; segment I mostly cylindrical, somewhat narrowed basally; segment II stout, its basal half cylindrical, apical half strongly flattened dorso-ventrally; segment III thin, cylindrical; labium thin, reaching well beyond hind coxae; segment I equal to length of head, subdivided near apex (Figs 5 A, C, 6 A – C). Thorax. Pronotum. Distinctly narrowing toward anterior part, bell-like; collar present, convex, well separated from remainder of pronotum by deep sulcus; lateral margin weakly sinuate, almost straight, not carinate; posterior margin somewhat concave; humeral angle relatively broad. Mesoscutum and scutellum. Mesoscutum well exposed; scutellum weakly convex (Figs 5 A, C, 6 A). Thoracic pleura. Mesepimeral spiracle slit-like; scent gland evaporative area triangular, occupying entire ventral portion of metepisternum; peritreme moderately upraised (Fig. 6 A). Hemelytron. Claval commissure as long as scutellum; embolium narrow; membrane with both cells well developed; major cell relatively short, its apex weakly reaching beyond apex of cuneus (Figs 5 A, C). Legs. Tarsus three-segmented; tarsomere I about as long as tarsomeres II and III combined (Fig. 6 E); pretarsal claw with subapical tooth (Fig. 6 F). Abdomen. Genitalia. Pygophore with dorsal wall as long as ventral wall, its opening terminated posteriorly; aedeagus with theca moderately sclerotized; ductus seminis outside endosoma long and thin, sclerotized portion of ductus seminis inside endosoma with basal two-thirds thin and cylindrical, apical one-third broadened, irregularly shaped with apico-lateral portions well expanded beyond middle of endosoma; secondary gonopore undifferentiated; endosoma divided into vesica and conjunctiva; apical part of endosoma with large, inflated, serrated lobe embraced basally by apico-lateral parts of distal portion of ductus seminis; endosoma laterally with long, flattened sclerotized lobe originated on its base and terminated subapically (Figs 7 A – C); left paramere C-shaped; apical process relatively long; paramere body with long, stiff setae (Figs 7 D – F); right paramere with apical process short, broadened and irregularly shaped in lateral view; paramere body covered with long, stiff setae; sensory lobe somewhat protruding, round (Figs 7 G – I). Female. Similar to male in coloration, texture, vestiture, and structure. Body more oval than in male (Fig. 5 B, D). Head. Antennal segment II entirely covered with short, moderately dense, adpressed setae; segment IV longer than III, thin, cylindrical. Abdomen. Genitalia. Bursa copulatrix relatively broad, weakly extending laterally beyond rami of the first valvulae (ra 1) (Fig. 8 A); dorsad labiate plate (dlp) with large sclerite occupying most of it, laterally possessing rib-like projections directed dorsal (Figs 8 A – E); sclerotized ring (sr) situated laterally, thickrimmed, relatively large (Figs 8 A – C); lateral oviducts (odl) placed centrally on genital chamber, relatively thick and short; spermathecal gland (sgl) opens centrally in the genital chamber, between the lateral oviducts (Fig. 8 B); ventral labiate plate (vlp) with elongate sclerotization laterally (sv) (Figs 8 B – E); posterior wall of genital chamber with interramal lobe strongly sclerotized, triangular (Fig. 8 F); vestibulum without any sclerite around vulva (Fig 8 C); first and second valvulae (gonapophyses 8 and 9) (va 1) with apex sharpened and not serrate (Figs 8 G, H); first valvulae (gonapophyses 8) connected by membrane (Fig. 8 A).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA6CFFE6A2A2F9EECF3625F1.taxon	etymology	Etymology. This name is derived from the Latin adjective infernus meaning of the lower origin, infernal, of hell and is used to denote distinct, flame-colored patches of the hemelytron and apical portions of each femur found in the type species. The gender is feminine.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA6CFFE6A2A2F9EECF3625F1.taxon	discussion	Remarks. Infernotropis is a usual genus distinguished from other cylapines by the combination of diagnostic characters presented above. It is superficially similar to the Afrotropical genus Linnavuorifulvius Gorczyca & Wolski, 2016 in both sharing the stout, elongate-oval body and the dorsal surface weakly rugose. The new genus can be easily distinguished among others by the dorsum sprinkled with fine, adpressed, regularly distributed silvery setae, the antennal segment II flattened dorsoventrally on the apical half, and the shape of the male genitalia (Fig. 7; Gorczyca et al. 2016: Figs 1, 2 A – C).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	description	(Figs. 5 – 8)	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	materials_examined	Material examined. Holotype (♂): ’ West MADAGASCAR \ nr. KIRINDI N. P., Andranomena \ Spec. Res.; 27 m (Andranovory \ nampela); S 20 ° 08 ' 52 '' E 44 ° 30 ' 52 '' \ 1 – 4. ii. 2013; M. Trýzna lgt. ’; Paratypes: (5 ♀): same as holotype.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	diagnosis	Diagnosis. See the generic diagnosis.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	description	Description. Male. COLORATION. Dorsum dark brown to black with distinct yellowish and orange areas (Fig. 5 A, C). Head. Black; vertex with two yellow patches each contiguous with inner margin of eye; maxillary plate, clypeus, gula, and buccula weakly tinged with yellow; antennal segment I dark brown; segment II dark brown, fuscous subapically, medial portion with small yellow patch on inner surface, apex with yellow annulation; segments III and IV fuscous; labium fuscous (Figs 5 A, C). Thorax. Pronotum. Black with two relatively large yellow to contrastingly orange patches contiguous with medial part of posterior margin (Figs 5 A, C). Scutellum. Black (Fig. 5 A). Thoracic pleura. Dark brown to black, sometimes weakly tinged with yellow on metepisternum. Hemelytron. Dark brown to black with two contrastingly orange, contiguous, rather small patches, one situated on embolium apically and other localized on cuneus basally; membrane fuscous (Figs 5 A, C). Legs. Dark brown with yellow and orange areas; femora sometimes weakly tinged with yellow, always with narrow, orange annulation apically; tibiae with rather broad, yellow annulation apically; tarsi dark yellow (Figs 5 A, C). Abdomen. Dark brown to black (Fig. 5 C). Female. Similar to male in coloration (Figs 5 B, D). Measurements. Male (n = 1) / Female (n = 3) Body. Length, clypeus – apex of membrane: 6.20 / 6.27 – 6.68; width: 2.04 / 2.56 – 2.69. Head. Length, s: 0.90 / 1.10 – 1.15; width, including compound eyes: 1.10 / 1.10 – 1.17; vertex width: 0.29 / 0.39 – 0.40; eye dorsal width: 0.40 / 0.37 – 0.43; height in lateral view: 0.68 / 0.71 – 0.73. Antenna. Length of segment I: 0.66 / 0.77 – 0.81; II: 2.55 / 2.30 – 2.44; III: 0.70 / 0.63 – 0.66; IV: – / 0.98 – 1.07. Labium. Total length: 3.84 / –; length of segment I: 0.95 / 1.00 – 1.06; II: 1.31 / –; III: 0.98 / –; IV: 0.68 / –. Pronotum. Mesal pronotal length: 1.03 / 1.03 – 1.12; lateral pronotal margin length: 1.11 / 1.21 – 1.22; posterior pronotal maximal width (straight): 1.68 / 1.89 – 1.97. Scutellum. Anterior width: 0.66; length (mesoscutum excluded): 0.84 / 0.82 – 0.96, length (mesoscutum included): 1.18 / 1.23 – 1.39. Hemelytron. Claval commissure: 1.10 / 1.12 – 1.47; heme length: 4.45 / 4.31 – 4.78; corium length: 2.56 / 2.64 – 2.94; outer cuneal margin length (straight): 0.76 / 0.79 – 0.82; cuneus width: 0.92 / 1.08 – 1.13; cell length: 1.14 / 1.26 – 1.30; cell width: 0.44 / 0.42 – 0.53. Hind leg. Femur length: 2.03 / –; tibia length: 2.95 / 2.99 – 3.10; tarsus length: 0.65 / 0.64 – 0.65; length of tarsal segment I: 0.37 / 0.36 – 0.37; II: 0.22 / 0.21; III: 0.22 / 0.24 – 0.26.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	biology_ecology	Biology. Unknown	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	distribution	Distribution. Central east Madagascar (Fig. 9).	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	etymology	Etymology. It is named after the type locality, Madagascar.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
7C6D8795FA69FFE0A2A2FF38CF2427E8.taxon	discussion	Remarks. In his catalogue, Gorczyca (2006) listed eight Cylapinae species from Madagascar. In addition to the two newly described species, we found a specimen of Fulvius anthocoroides (Reuter, 1875) in the collection of the Moravian Museum in Brno bearing the following label data: “ West MADAGASCAR \ nr. KIRINDI N. P., Andranomena \ Spec. Res.; 27 m (Andranovory \ nampela); S 20 ° 08 ' 52 '' E 44 ° 30 ' 52 '' \ 1 – 4. ii. 2013; M. Trýzna lgt. ”. Currently, a total of 11 species (including eight endemic) are known from Madagascar.	en	Taszakowski, Artur, Masłowski, Adrian, Wolski, Andrzej, Gorczyca, Jacek (2022): Two new genera and species of the Fulviini (Hemiptera, Heteroptera, Miridae Cylapinae) from Madagascar, with a checklist of Madagascan cylapines. Zootaxa 5209 (1): 93-110, DOI: https://doi.org/10.11646/zootaxa.5209.1.5
