identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
10F6926364BA5F4ABC8FE4177CB8C73B.text	10F6926364BA5F4ABC8FE4177CB8C73B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Centroptilum alamiae Kaltenbach, Vuataz & Gattolliat 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Centroptilum alamiae Kaltenbach, Vuataz &amp; Gattolliat sp. nov.</p>
            <p>Figs 4b, e, 5b, 6, 7, 8, 9</p>
            <p> Differential diagnosis to other species of  Centroptilum . </p>
            <p> Larva. Following combination of characters: A) labrum with anterior margin slightly concave; ratio width vs. length ca. 1.5  × (Fig. 7a); B) maxillary palp ca. 1.7  × as long as galea-lacinia, segment III apically rounded; segment III ca. 1.6  × as long as segment II (Fig. 7g); C) inner distal margin of labial palp segment III slightly concave (Fig. 7k); D) dorsal margin of fore femur with occasional short, spine-like setae; row of stout, pointed setae near margin (Fig. 8a); E) tarsus approx. as long as tibia (Fig. 8a); F) claw with two rows of denticles, each row with ca. 20 small to minute denticles (Fig. 8b); G) paraproct with 30-45 pointed spines, sometimes with split tips, few additional, submarginal spines (Fig. 8j). </p>
            <p>Description.</p>
            <p>Larva (Figs 4b, e, 5b, 7 - 9). Body length 5.6-7.0 mm. Caudalii broken. Antennae reaching apex of fore protoptera.</p>
            <p>Colouration (Fig. 9a-c). Head, thorax and abdomen dorsally brown, with dark grey-brown pattern as in Fig. 9a. Head, thorax and abdomen ventrally light brown, with dark grey-brown lateral marks on thorax (Fig. 9c). Legs light brown, femur distomedially slightly darker, tarsus basally and distally slightly darker, claw basally darker. Caudalii light brown, darker annulated.</p>
            <p> Labrum (Fig. 7a). Rectangular, width ca. 1.5  × maximum length. Distal margin with broad, angulated, medial emargination. Anterior margin slightly concave. Dorsal surface scattered with long, medium and short, simple setae; setae not arranged in a submarginal arc. Ventrally with marginal row of setae composed of anterolateral long, simple, pointed setae and medial long, apically blunt, pectinate setae; ventral surface with ca. nine short, stout setae near lateral and anterolateral margin. </p>
            <p>Right mandible (Fig. 7b, c). Incisor and kinetodontium separated. Incisor with three denticles; kinetodontium with two denticles. Prostheca stick-like, distally with three denticles. Margin between prostheca and mola almost straight, with two tufts of long setae. Tuft of setae at apex of mola present.</p>
            <p>Left mandible (Fig. 7d, e). Incisor and kinetodontium separated. Incisor with four denticles; kinetodontium with three denticles. Prostheca stick-like, distolaterally denticulate. Margin between prostheca and mola straight, with large brush-like tuft of long setae. Subtriangular process short, on level of area between prostheca and mola. Tuft of setae at apex of mola absent.</p>
            <p>Hypopharynx and superlinguae (Fig. 7f). Lingua as long as superlinguae. Lingua longer than broad; distal half laterally not expanded; distal margin with short, fine setae, tuft of stout setae short. Superlinguae distally rounded; lateral margins rounded; fine, short to long, simple setae along distal margin.</p>
            <p> Maxilla (Fig. 7g). Galea-lacinia ventrally with four or five simple, apical setae under canines. Canines long and slender. With three denti-setae, distal denti-seta canine-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one pectinate, spine-like seta and three simple, spine-like setae (dorsolateral insertions); and ca. six long setae, partly with bifurcated tips (bifurcation often difficult to see; ventrolateral insertions). Maxillary palp 3-segmented, ca. 1.7  × as long as length of galea-lacinia; palp segment III ca. 1.6  × length of segment II; setae on maxillary palp fine, simple, scattered over surface of segments I, II, and III; apex of last segment rounded. </p>
            <p>Labium (Fig. 7h-k). Glossa nearly as broad and slightly shorter than paraglossa; inner and outer margins with many short, spine-like setae; apex with two medium, robust setae; dorsal surface with long, fine, simple, scattered setae. Paraglossa curved inward; ventrally with many long setae along outer lateral and apical margin, and row of long, stout, pointed, simple setae along inner lateral margin; dorsal surface with long, fine, simple, scattered setae. Labial palp 3-segmented. Segment III nearly trapezoidal with rounded distal corners, distal margin slightly concave; outer lateral margin with short to medium, fine, simple setae, distal margin with short, spine-like and short, fine, simple setae; ventral surface with medium, fine, simple, scattered setae. Segment II with medium, fine, simple, scattered setae along outer lateral margin and on ventral surface; dorsally with seven or eight short, spine-like setae along distal margin. Segment I with medium, fine, simple setae scattered on ventral surface and on outer lateral margin.</p>
            <p>Hind protoptera well developed.</p>
            <p> Foreleg (Fig. 8a, b) very slender. Ratio of foreleg segments 1.6:1.0:1.0:0.4. Femur. Length ca. 5  × maximum width. Dorsal margin with occasional short, spine-like setae, row of short, pointed setae near margin. Apex slightly rounded. Short, stout, pointed setae scattered along ventral margin; femoral patch absent. Tibia. Dorsal margin bare. Ventral margin with row of short, curved, spine-like setae and some aditional stout, pointed setae along margin. Anterior surface scattered with short, stout, pointed, and partly serrate setae along ventral margin. Patellatibial suture present in basal 1/3 area. Tarsus. Dorsal margin bare. Ventral margin with dense row of short, curved, serrate, spine-like setae. Claw with two rows of 17-20 minute denticles each, in basal ca. 1/3 area, increasing in size distally; subapical setae absent. </p>
            <p>Terga (Figs 4b, e, 5b). Posterior margin of terga: I smooth, without spines; II-VI (VII) with small triangular spines; VII-IX with triangular, pointed spines.</p>
            <p>Sterna. Posterior margin of sterna I-VI smooth, without spines. Posterior margin of sterna VII-VIII with small, triangular spines.</p>
            <p>Tergalii (Figs 8c-i, 9d). Present on segments I-VII. Costal margins with minute denticles and short, fine, simple setae, anal margins almost smooth. Tracheae extending from main trunk to inner and outer margins. Tergalius I as long as length of segments II and III combined; tergalius IV as long as length of segments V and VI combined; tergalius VII as long as length of segments VIII and IX combined.</p>
            <p>Paraproct (Fig. 8j). With irregular row of 30-45 pointed marginal spines of different size, some with split tips, and few additional spines in second row. Cercotractor with minute, irregular, marginal spines.</p>
            <p>Caudalii (Fig. 8k). Spines at posterior margins of segments short triangular, pointed.</p>
            <p>Subimago. Judging from subimaginal tarsomeres developing under cuticle of last instar female larvae, all tarsomeres of all legs of female subimago have pointed microlepids on surface (see Kluge 2022).</p>
            <p>Imago. Unknown.</p>
            <p>Etymology.</p>
            <p>Dedicated to Prof. Majida El Alami, committed researcher on aquatic insects in Morocco, and collector of some of the specimens; in recognition of her substantial contribution to the knowledge of the systematics, ecology, and distribution of Moroccan mayflies.</p>
            <p>Biological aspects.</p>
            <p>The specimens were collected in calm edge waters, loose substrate, low to moderate current, high temperatures, and sites rich in filamentous algae and mosses (Fig. 6a, b; El Alami et al. 2022a).</p>
            <p>Distribution</p>
            <p>(Fig. 6e). Morocco.</p>
            <p>Type-material.</p>
            <p> Holotype. Morocco • larva; Oued  Kelâa , Akchour; 35°14'32"N, 05°10'10"W; 13.03.2021; leg. S. El Yaagoubi; on slide; GBIFCH00592619, GBIFCH00592620, GBIFCH00592621; MZL. Paratypes. Morocco • 6 larvae; same data as holotype; 2 on slides; GBIFCH00980875, GBIFCH00980876; 4 in alcohol; GBIFCH00975645, GBIFCH00975646; MZL • 7 larvae; Oued Jnane Niche (sup.); 16.03.2014; leg. M. El Alami; in alcohol; GBIFCH00975647; MZL • 12 larvae; Oued Jnane Niche (sup.); 17.05.2015; leg. M. El Alami; 1 on slide; 11 in alcohol; LESCB. </p>
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	https://treatment.plazi.org/id/10F6926364BA5F4ABC8FE4177CB8C73B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kaltenbach, Thomas;Vuataz, Laurent;Samraoui, Boudjema;El Yaagoubi, Sara;El Alami, Majida;Gattolliat, Jean-Luc	Kaltenbach, Thomas, Vuataz, Laurent, Samraoui, Boudjema, El Yaagoubi, Sara, El Alami, Majida, Gattolliat, Jean-Luc (2022): Two new species of Centroptilum Eaton, 1869 from North Africa (Ephemeroptera, Baetidae). ZooKeys 1131: 71-97, DOI: http://dx.doi.org/10.3897/zookeys.1131.91017, URL: http://dx.doi.org/10.3897/zookeys.1131.91017
05D708EB910D574698C7DA1192F054E8.text	05D708EB910D574698C7DA1192F054E8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Centroptilum samraouii Kaltenbach, Vuataz & Gattolliat 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Centroptilum samraouii Kaltenbach, Vuataz &amp; Gattolliat sp. nov.</p>
            <p>Figs 1, 2, 3, 4a, d, 5a, 6</p>
            <p> Differential diagnosis to other species of  Centroptilum . </p>
            <p> Larva. Following combination of characters: A) labrum with anterior margin nearly straight; ratio width vs. length ca. 1.6  × (Fig. 1a); B) maxillary palp ca. 1.9  × as long as galea-lacinia, segment III apically pointed; segment III ca. 1.3  × as long as segment II (Fig. 1g); C) inner distal margin of labial palp segment III concave (Fig. 1j); D) dorsal margin of fore femur with occasional short, spine-like setae (Fig. 2a); E) fore tarsus slightly longer than tibia (1.1  × ; Fig. 2a) F) claw with two rows of denticles, each with ca. 20 small to minute denticles (Fig. 2b); G) paraproct with 17-23 pointed spines, plus some additional submarginal spines (Fig. 2j). </p>
            <p>Description.</p>
            <p>Larva (Figs 1 - 3, 4a, d, 5a). Body length 3.8-4.2 mm. Cerci: ca. 2/3 of body length. Paracercus: nearly as long as cerci. Antennae reaching apex of fore protoptera.</p>
            <p>Colouration (Fig. 3a, b). Head, thorax and abdomen dorsally brown, with dark grey-brown pattern as in Fig. 3a. Head and thorax ventrally brown, with dark grey-brown lateral marks on thorax (Fig. 3b). Abdomen ventrally light brown. Legs light brown, apex of femur and claw darker. Caudalii ecru, brown annulated.</p>
            <p> Labrum (Fig. 1a). Rectangular, width ca. 1.6  × maximum length. Distal margin with broad, angulated, medial emargination. Anterior margin nearly straight. Dorsal surface scattered with long, medium and short, simple setae; setae not arranged in a submarginal arc. Ventrally with marginal row of setae composed of anterolateral long, simple, pointed setae and medial long, apically blunt, pectinate setae; ventral surface with ca. seven short, stout setae near lateral and anterolateral margin. </p>
            <p>Right mandible (Fig. 1b, c). Incisor and kinetodontium separated. Incisor with three denticles; kinetodontium with two denticles. Prostheca stick-like, distally with two denticles. Margin between prostheca and mola almost straight, with two tufts of long setae. Tuft of setae at apex of mola present.</p>
            <p>Left mandible (Fig. 1d, e). Incisor and kinetodontium separated. Incisor with four denticles; kinetodontium with three denticles. Prostheca stick-like, distally denticulate. Margin between prostheca and mola straight, with large brush-like tuft of long setae. Subtriangular process short, on level of area between prostheca and mola. Tuft of setae at apex of mola absent.</p>
            <p>Hypopharynx and superlinguae (Fig. 1f). Lingua as long as superlinguae. Lingua longer than broad; distal half laterally not expanded; distal margin with short, fine setae, tuft of stout setae short. Superlinguae distally rounded; lateral margins rounded; fine, short to long, simple setae along distal margin.</p>
            <p> Maxilla (Fig. 1g, h). Galea-lacinia ventrally with 3-5 simple, apical setae under canines. Canines long and slender. With three denti-setae, distal denti-seta canine-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one pectinate, spine-like seta and two simple, spine-like setae (dorsolateral insertions); and ca. eight long setae with bifurcated tips (bifurcation often difficult to see; ventrolateral insertions). Maxillary palp 3-segmented, ca. 1.9  × as long as length of galea-lacinia; palp segment III ca. 1.3  × length of segment II; setae on maxillary palp fine, simple, scattered over surface of segments I, II, and III; apex of last segment pointed. </p>
            <p>Labium (Fig. 1i, j). Glossa nearly as broad and slightly shorter than paraglossa; inner and outer margins with many short, spine-like setae; apex with two medium, robust setae; dorsal surface with long, fine, simple, scattered setae. Paraglossa curved inward; ventrally with many long setae along outer lateral and apical margin, and row of long, stout, pointed, simple setae along inner lateral margin; dorsal surface with long, fine, simple, scattered setae. Labial palp 3-segmented. Segment III nearly trapezoidal with rounded distal corners, distal margin concave; outer lateral margin with short to medium, fine, simple setae, distal margin with short, spine-like and short, fine, simple setae; ventral surface with medium, fine, simple, scattered setae. Segment II with medium, fine, simple, scattered setae along outer lateral margin and on ventral surface; dorsally with 5-7 short, spine-like setae along distal margin. Segment I with medium, fine, simple setae scattered on ventral surface.</p>
            <p>Hind protoptera well developed.</p>
            <p> Foreleg (Fig. 2a, b) very slender. Ratio of foreleg segments 1.6:1.0:1.1:0.4. Femur. Length ca. 5  × maximum width. Dorsal margin with occasional short, spine-like setae. Apex slightly rounded. Short, stout, pointed setae scattered along ventral margin; femoral patch absent. Tibia. Dorsal margin bare. Ventral margin with row of short, curved, spine-like setae and additional stout, pointed setae along margin. Anterior surface scattered with few stout, pointed, and partly serrate setae along ventral margin. Patellatibial suture present in basal  ¼ area. Tarsus. Dorsal margin bare. Ventral margin with dense row of short, curved, serrate, spine-like setae. Claw with two rows of 17-20 minute denticles each, in basal ca. 1/3 area, increasing in size distally; subapical setae absent. </p>
            <p>Terga (Figs 4a, d, 5a). Posterior margin of terga: I smooth, without spines; II with rudimentary spines; III with small, triangular spines; IV-IX with triangular spines.</p>
            <p>Sterna. Posterior margin of sterna I-VI smooth, without spines. Posterior margin of sterna VII-VIII with small, triangular spines.</p>
            <p>Tergalii (Figs 2c-i, 3c). Present on segments I-VII. Costal margins with minute denticles and short, fine, simple setae, anal margins almost smooth. Tracheae extending from main trunk to inner and outer margins. Tergalius I as long as length of segments II-IV combined; tergalius IV as long as length of segments V and VI combined; tergalius VII as long as length of segments VIII and IX combined.</p>
            <p>Paraproct (Fig. 2j). With 17-23 pointed marginal spines of different size, and some additional spines in second row. Cercotractor with minute, irregular, marginal spines.</p>
            <p>Caudalii (Fig. 2k). Spines at posterior margins of segments elongated triangular with long points.</p>
            <p>Subimago. Judging from subimaginal tarsomeres developing under cuticle of last instar female larvae, all tarsomeres of all legs of female subimago have pointed microlepids on surface (see Kluge 2022).</p>
            <p>Imago. Unknown.</p>
            <p>Etymology.</p>
            <p> Dedicated to Prof.  Boudjéma Samraoui, committed researcher on aquatic insects in Algeria, and collector of the new species; in recognition to his substantial contribution to the knowledge of the ecology and distribution of Algerian mayflies. </p>
            <p>Biological aspects.</p>
            <p> Centroptilum samraouii sp. nov. occupies the headwaters of steep, narrow and intermittent streams (Fig. 6c, d; Samraoui et al. 2021b, c). </p>
            <p>Distribution</p>
            <p>(Fig. 6e). Algeria.</p>
            <p>Type-material.</p>
            <p>Holotype. Algeria • larva; Guitna sup., Ghora; 36°36'42"N, 08°21'19"E; 22.01.2020; leg. B. Samraoui; on slides; GBIFCH00592552, GBIFCH00592551, GBIFCH00592622; MZL. Paratypes. Algeria • 2 larvae; Guitna sup., Ghora; 36°36'42"N, 08°21'19"E; 05.11.2019; leg. B. Samraoui; on slides; GBIFCH00895417, GBIFCH00895418; MZL • 3 larvae; Guitna sup.; 36°36'42"N, 08°21'19"E; 09.10.2019; leg. B. Samraoui; on slide; GBIFCH00592553; 2 in alcohol; GBIFCH00975620, GBIFCH00975623; MZL • larva; Louar inf., Ghora; 36°37'03"N, 08°22'49"E; 05.11.2019; leg. B. Samraoui; on slide; GBIFCH00592555; MZL • larva; Algeria; Guitna inf.; 07.11.2018; leg. B. Samraoui; in alcohol; GBIFCH00975621; MZL.</p>
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	https://treatment.plazi.org/id/05D708EB910D574698C7DA1192F054E8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kaltenbach, Thomas;Vuataz, Laurent;Samraoui, Boudjema;El Yaagoubi, Sara;El Alami, Majida;Gattolliat, Jean-Luc	Kaltenbach, Thomas, Vuataz, Laurent, Samraoui, Boudjema, El Yaagoubi, Sara, El Alami, Majida, Gattolliat, Jean-Luc (2022): Two new species of Centroptilum Eaton, 1869 from North Africa (Ephemeroptera, Baetidae). ZooKeys 1131: 71-97, DOI: http://dx.doi.org/10.3897/zookeys.1131.91017, URL: http://dx.doi.org/10.3897/zookeys.1131.91017
