taxonID	type	description	language	source
515B87B6FF84FFCFFF63A9F2FCF8FD10.taxon	distribution	Distribution in Caucasus. Azerbaijan (Skalický 1998; Sazhnev 2018 b). Distribution. The species was described from China (Xinjiang) (Reitter 1887), distributed in Middle Asia (Kazakhstan, Tajikistan, Turkmenistan), Near East (Israel, Iran, Iraq) (Derwesh 1965; Mascagni 2016; Litovkin & Sazhnev 2018) and Caucasus (Azerbaijan) (Skalický 1998).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF84FFCFFF63AA92FE23F8BC.taxon	materials_examined	Material examined. 52 exs. GEORGIA. Mtskheta-Mtianeti Krai: Mtskheta Distr., Kura River opposite Dzegvi vill., 41 ° 51.199 ' N 44 ° 35.737 ' E, 10.08.2016, leg. D. M. Palatov, 2 ex. (IBIW). RUSSIA. Astrakan Oblast: Krasnoyarsky Distr., Dosang vill., 8.05.2009, leg. A. V. Kovalev, 4 exs. (ZISP); same place, bank Akhtuba River, 46 ° 54 ' 30.50 " N 47 ° 54 ' 25.50 " E, at light, 1 – 8.05.2011, leg. I. A. Zabaluev, 7 exs, (IBIW). Dagestan Republic: Kumtorkalinsky Distr., Kumtorkala vill., Sarykum barkhan, at light, 9.06.2013, leg. E. V. Ilyina, 1 ex. (IBIW); same place, Kumtorkala station, 31.07.2016, E. V. Ilyina, 1 ex. (IBIW), 19.05.2018, leg. E. V. Ilyina, 5 exs. (IBIW); same place, right bank Shura-Ozen River, near the railway bridge, night at light, 23.05.2019, leg. I. A. Zabaluev, 21 exs, (IBIW); same place, Kumtorkala station, 2 – 3.06.2019, leg. I. A. Zabaluev, 4 exs. (IBIW); Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E. V. Ilyina, 2 exs. (IBIW); Magaramkent Distr., Tagirkent-Kazmalar vill., 20.07.2014, leg. E. V. Ilyina, 2 exs. (IBIW); Magaramkent Distr., State Nature Zakaznik “ Samursky ”, 29 – 30.07.2015, leg. E. V. Ilyina, 1 ex. (IBIW); Nogaysky Distr., sands Karagaily-Kum, 44 ° 02 ' 15.4 " N 45 ° 38 ' 34.2 " E, 16.08.2016, leg M. V. Nabozhenko, S. V. Nabozhenko & G. M. Abdurakhmanov, 1 ex. (IBIW). Volgograd Oblast: Palassovsky Distr., Elton vill., at light, 21 – 28.05.2006, leg A. V. Matalin, 1 ex. (IBIW).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF84FFCFFF63AA92FE23F8BC.taxon	distribution	Distribution in European Russia and Caucasus. ST: AST VOL (Sazhnev 2016 c) CA: ADG DAG KAB (Tersky Distr.) KCH / STA (Podkumok River) (Motschulsky 1854; Jakobson 1913; Zaitsev 1946; Nikitsky 2010; Sazhnev 2017; Sazhnev & Ilyina 2017). Armenia: Elisabethpol, Governorate (Jacobson 1913; Mascagni 2016); Azerbaijan: Agdash (Geok Tepe) (Schneider & Leder 1877; Jacobson 1913; Zaitsev 1946; Sazhnev 2018 b); Georgia: Karayaz, Tbilisi (Zaitsev 1946, 1953; Mascagni 2016). Distribution. The species was described from Italy (Etruria), distributed in South Europe, Caucasus (Russian North Caucasus, Armenia, Azerbaijan, Georgia), Near East (Arab Emirates, Iran, Iraq, Lebanon, Saudi Arabia, Egypt (Sinai )), Middle Asia (Kazakhstan, Kirgizstan, Tajikistan, Turkmenistan and Uzbekistan), Pakistan and North Africa (Mascagni 2016; Sazhnev 2020). In Russia, this species occurs only in the extreme south of European part and North Caucasus (Sazhnev & Shapovalov 2016; Sazhnev 2016 b). The records for most part of Siberia (species is possible in the southwest West Siberia) and the Russian Far East are erroneous and require confirmation (Sazhnev 2020).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF84FFCFFF63AA92FE23F8BC.taxon	discussion	Remarks. The species recorded for Mtskheta-Mtianeti Krai of Georgia for the first time. The record from Khashuri (Mikhailovo) (Schneider & Leder 1877) on the opinion by F. A. Zaitsev (1953) is other species Augyles sericans (Kiesenwetter, 1843).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF84FFC8FF63AF3EFEC8F9E0.taxon	materials_examined	Material examined. 250 exs. RUSSIA. Astrakhan Oblast: Enotaevsky Distr., Zamyany vill., at light, 16.09.2007, leg. O. G. Brekhov, 30 exs. (IBIW). Ivanovo Oblast: Pestryakovsky Distr., Purezhka River, 56 ° 28 ' 30.3 " N 42 ° 38 ' 4.2 " E, at light, 24 – 25.06.2011, leg. D. V. Potanin, 2 exs. (IBIW). Kaliningrad Oblast: Nemansky Distr., 3 km E of Bolshoe Selo vill., Neman River, 55 ° 03 ' 07.3 " N 22 ° 09 ' 02.1 " E, 12.06.2019, leg. A. A. Przhiboro, 12 exs. (ZISP). Kirov Oblast: Kilzemsky Distr., Tautovo vill., Kilmez River, 7.06.2005, leg. E. A. Korobeynikova, 1 ex. (IBIW). Leningrad Oblast: Kingiseppsky Distr., Kingisepp (Yamburg), 6.07.1904, leg. Barovsky, 2 exs. (ZISP); Gatchinsky Distr., env. Malkovoe vill., Suyda River, 16.07.2018, leg. A. G. Koval, 3 exs. (VIZR). Lipetsk Oblast: Eletsky Distr., 30 km E from Elets, Don River left bank, on sand, 26.07.2004 leg. A. O. Bieńkowski, 1 ex. (PCB). Republic of Mari El: Tsaryovokokshaysky Uyezd, 12.07.1913, unknown collector, 1 ex. (ZISP). Moscow Oblast: Serpukhovsky Distr., 3 km W of Luzhi vill., 18.07.2011, leg. A. V. Matveev, 1 ex. (ZISP). Nizhniy Novgorod Oblast: Dzerzhinsky Distr., Gavrilovka vill. 6.07.2013, leg. D. V. Potanin, 8 exs. (IBIW). Perm Krai: Perm, Balmoshnaya River, soil traps, 17.09. – 5.10.2017, leg. V. O. Kozminykh, 1 ex. (IBIW). Rostov Oblast: Sholokhovsky Distr., Kruzhilinsky vill., at light, 26 – 30.06.2003, leg. O. V. Brekhov, 2 exs. (IBIW); Sholokhovsky Distr., Vyoshinskaya vill., at light, 3 – 4.07.2003, leg. O. V. Brekhov, 2 exs. (IBIW). Ryazan Oblast: Spassky Distr., Brykin Bor vill., bank of Pra River, 54 ° 42 ' 48.1 " N 40 ° 51 ' 10.8 " E, on sand, 28.06.2017, leg. A. S. Sazhnev, 2 exs. (IBIW). Saratov Oblast: Lysogorsky Distr., 3 km NE Semyonovka vill., bank of Medveditsa River, on sand, 25 – 28.06.2012, leg. A. S. Sazhnev, 13 exs. (IBIW); Voskresensky Distr., Sinodskoe vill., Tereshka River, 51 ° 58 ' 06.4 " N 46 ° 39 ' 44.5 " E, 15.07.2012, leg. N. V. Rodnev, 3 exs. (IBIW); Balashovsky Distr., Repnoe vill., pools, 29.05.2017, leg. A. N. Volodchenko, 1 ex. (IBIW). Republic of Tatarstan: Kazan, Gavrilova str., Kazanka River, on sand, 13.07.2010, leg. D. A. Klyomin, 1 ex. (IBIW). Vladimir Oblast: Vozhegodsky Distr., Grudino (Zarechie) vill., sandy bank of Vozhega River, 28.07.2016, leg. D. Vtorushin, 81 exs. (IBIW), same place, 31.07.2016, leg. D. Vtorushin, 55 exs. (IBIW). Volgograd Oblast: Uryupinsky Distr., 5 km W pf Mikhaylovskaya vill., bank of Medveditsa River, 21.06.2011, leg. A. V. Matveev, 1 ex (ZISP). Ilovlinsky Distr., Baybaev vill., at light, 23.05.2002, leg. O. G. Brekhov, 1 ex. (PCBr). Voronezh Oblast: Novousmansky Distr., 2 km W Maklok vill., bank of Usmanka River (env. Chistoe Lake), sand, 12.07.2016, leg. A. S. Sazhnev, 10 exs. (IBIW). Yaroslavl Oblast: Yaroslavsky Distr., Krasnye Tkachi settlement, bank of Kotorosl River, 5.06.1993, leg. D. V. Vlasov, 3 exs. (YMR); Nekouzsky Distr., Borok settlement, Radovsky Island, sandy bank of Rybinskoe Reservoir, 14.06.2016, leg. A. S. Sazhnev, 13 exs. (IBIW). Addition material. 10 exs. KAZAKHSTAN. Aktyubinsk Oblast: [Uralsk Oblast: Temirsky Ueazd], Chatarly River, 6.06.1908, leg. B. Uvarov & D. Borodin, 2 ex. (ZISP); same place, necropolis Daumshar, 9.06.1908, leg. B. Uvarov & D. Borodin, 3 exs. (ZISP); same place, Kokdzhida sands, 10.06.1908, leg. B. Uvarov & D. Borodin, 1 ex. (ZISP); same place, 17.06.1908, leg. B. Uvarov & D. Borodin, 4 ex. (ZISP).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF84FFC8FF63AF3EFEC8F9E0.taxon	distribution	Distribution in European Russia. NT: ARH SPB VLG CT: IVA KIR KOS KÖN MOS NNO * RZN TUL VLD YAR VL: CHV MAR MRD SAM TAT ST: AST LIP ROS SAR VOL * VRN UR: ORB PER (Lindemann 1871; Zaitzev 1907; Pomerantsev 1908; Jacobson 1913; Lebedev 1925; Gussakovsky 1926; Semyonov, 2009; Tsurikov 2009; Alekseev & Shapoval 2019; Litovkin 2012; Litovkin et al., 2013; Dedyukhin & Kholmogorova, 2014; Sazhnev 2014, 2016 c, 2018 c; Egorov 2019; Sazhnev & Kozminykh 2019; Sazhnev et al., 2018; Egorov et al., 2020) Distribution. The species was described from Austria, distributed in Europe, Near East (Israel, Iraq), Asian Turkey, and Middle Asia (Turkmenistan, Uzbekistan) (Mascagni 2016). Records from the south Siberia and the Russian Far East (Kirejtshuk 2001) should be attributed to other species of the Augyles cribratellus species group sensu Charpentier (1965) (Sazhnev 2018 c). The record of this species from Issyk-Kul hollow in Kyrgyzstan (Ovchinnikov 1996) require verification (Litovkin & Sazhnev 2018).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF84FFC8FF63AF3EFEC8F9E0.taxon	discussion	Remarks. The species recorded for two subjects of Russia (Nizhniy Novgorod and Volgograd Oblasts) for the first time.	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF83FFC9FF63AEFCFB64FB7C.taxon	materials_examined	Material examined. 36 exs. RUSSIA. Kaluga Oblast: Kirovsky Distr., Berezhki vill., 54 ° 07 ' 16.0 " N 34 ° 26 ' 29.0 " E, at light, 27 – 28.06.2016, leg. V. V. Perov, 3 exs. (IBIW). Komi Republic: Ust-Tsilemsky Distr., Ust-Tsilma vill., 29.08.1904, leg. Zhuravsky, 1 ex. (ZISP); Syktyvkar, Vezhdino vill., 20.07.1991, leg. A. Zheleznyakov, 3 exs. (MPSU). Krasnodar Krai: Novorossiysk, 18.08.1878 (var. basimaculatus Reitter, 1908), unknown collector, 1 ex. (ZISP). Leningrad Oblast: Kingiseppsky Distr., Romanovka vill., env. Kingisepp (Yamburg), 30 – 31.05.1904, leg. Barovsky, 4 exs. (ZISP). Moscow Oblast: [Klinsky uezd, Babkovo], 3.06.1905, D. S., 1 ex. (ZISP); 23 km W Zvenigorod, env. Glubokoe Lake, at light, 14.06.1998, leg. A. O. Bieńkowski, 2 exs. (PCB); 5 km N Zelenograd, Nikolskoe vill., at light, 23.06.2006, leg. A. O. Bieńkowski, 1 ex. (PCB). Nenets Autonomous Okrug: Gulf of Khaydypur, More-Yu River, 7 – 15.08.2015, leg. O. Makarova & N. Bizin, 2 exs. (MSPU). Nizhniy Novgorod Oblast: Arzamas Distr., Volchikha vill., [Gorbatovsky uezd], 4 – 9.05.1894, leg. Jacobson, 1 ex. (ZISP). Saint Petersburg: Lesnoy, 20.05.1862, leg. А. Moravits, 1 ex. (ZISP); env. Pavlovsk, 5.06.1906, leg. V. Barovsky, 2 exs. (ZISP). Saratov Oblast: Tatishchevo Distr., env. Storozhovka vill., Kurdyum River, 4.09.2014, leg. I. A. Zabaluev, 1 ex. (IBIW). Tambov Oblast: 20 km NNE Tambov, Zarechie vill., at light, 16.08.2008, leg. R. N. Ishin, 1 ex. (PCL). Tver Oblast: Bologoe, 26.06.1903, unknown collector, 1 ex. (ZISP). Ulyanovsk Oblast: Radishchevsky Distr., 6 km N Vyazovka vill., saline steppe, 3.05.2005, leg. A. Yu. Isaev, 1 ex. (ZISP); Radishchevsky Distr., 8 km S Vyazovka vill., 29.06.2008, leg. A. V. Kovalev, 1 ex. (ZISP). Vladmir Oblast: Murmansky Distr., Berdishchevo, unknown collector, 1 ex. (ZISP). Vologda Oblast: Vozhegodsky Distr., env. Gridino vill., sand bank Vozhega River, 31.07.2016, leg. D. Vtorushin, 1 ex. (IBIW). Yaroslavl Oblast: Yaroslavl, 10.05.1923, unknown collector, 1 ex. (ZMMU), same place, 14.05.1923, unknown collector, 1 ex. (ZMMU); Rostov Distr., Melenki station, 4.06.2006, leg. V. D. Titov, 1 ex. (YMR); Uglich Distr., env. Uglich, Uglich Reservoir, 23.05.1999, leg. Maslov, 1 ex. (YMR); Uglich Distr., Meteevo vill., Uleyma River, 14.06.1997, leg. D. V. Vlasov, 1 ex. (YMR); Yaroslavl Distr., 9.08.1926, leg. Jakovlev, 2 exs. (ZISP); Yaroslavl Distr., Berditsyno vill., 3.05.1989, leg. A. Jakovlev, 1 ex. (ZISP); Yaroslavl Distr., Belkino vill., Tallitsa River, 2.07.1999, leg. D. V. Vlasov, 1 ex. (YMR).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF83FFC9FF63AEFCFB64FB7C.taxon	distribution	Distribution in European Russia and Caucasus. NT: KAR NEN SPB VLG CT: KÖN KLG MOS NNO * NOV TAM * TVR * VLD YAR VL: TAT ULY ST: LIP SAR CA: KRD (Novorossiysk) UR: EKB KOM (Motschulsky 1854; Lindemann 1871; Yakovlev 1902; Zaitsev 1907, 1916, 1946; Jacobson 1913; Zaitsev 1915, 1915 a, 1930; Lebedev 1925; Bercio & Folwaczny 1979; Semyonov 2009; Tsurikov 2009; Litovkin et al. 2013; Sazhnev 2016 a, 2017, 2018; Sazhnev & Shapovalov 2016; Sazhnev et al. 2019; Sazhnev & Kozminykh 2019). Distribution. Holarctic species (Charpentier 1979; Mascagni 2016) with a disjunction of the range on the Black Sea coast of the North Caucasus (Zaitsev 1946). Species was described from Germany, distributed in North and Central Europe, Central Asia (northern Kazakhstan and Mongolia), West and East Siberia, Russian Far East, North America (Mascagni 2016; Sazhnev 2020).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF83FFC9FF63AEFCFB64FB7C.taxon	discussion	Remarks. The first record this species for Caucasus was make as Augyles seriepilosus Motschulsky, 1860 (Zaitsev 1916), which F. A. Zaitsev considered as probably synonym of Augyles intermedius species (Zaitsev 1946). But Augyles seriepilosus (species dubius) was described from East Siberia (Transbaikal Region) (Motschulsky 1860), specimen of « Augyles seriepilosus » with Motschulsky labels is in ZISP collection, and it possible refers to Asian species Augyles interspidulus (Charpentier, 1979). The species recorded for Nizhniy Novgorod, Tambov and Tver Oblasts for the first time.	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF82FFCAFF63AD7FFD0AFF40.taxon	materials_examined	Material examined. 22 exs. ABKHAZIA. Sukhumi, 30.03.1960 leg. Chernyshev, 1 ex. (IBIW); Bzyb’ River near Pitsunda, 43 ° 11 ' 34.4 " N 40 ° 17 ' 14.6 " E, leg. A. A. Prokin, 7 exs. (IBIW). AZERBAIJAN. Lankaran District: Avrora vill., Hirkan National Park, at light, 4.05.1980, leg. S. G. Korolev, 1 ex. (IBIW). RUSSIA. Astrakhan Oblast: Krasnoyarsky Distr., env, Dosang vill., at light, 8.05.2009, leg. A. V. Kovalev 4 exs. (IBIW, ZISP); same place, Akhtuba River, 46 ° 54 ' 30.50 " N 47 ° 54 ' 25.50 " E, at light, 1 – 8.05.2011, leg. I. A. Zabaluev, 2 exs. (IBIW). KarachayCherkess Republic: Urup Distr., Urup River near Pregradnoye vill., 43 ° 58 ' 25.2 " N 41 ° 12 ' 04.7 " E, 15.08.2016, leg. A. A. Prokin, 4 exs. (IBIW). Krasnodar Krai: 3 km N Aderbievka vill., W slope of Kotsekhur mountain range, shore of Aderba River, h ~ 160 m, 44 ˚ 37 ’ 52 " N 38 ˚ 06 ’ 06 " E, 3.05.2016, 1 ex, K. V. Makarov & O. L. Makarova leg. (IBIW). Republic of North Ossetia-Alania: Alagirsky Distr., Barzikau vill., shore of Fiagdon River, h ≈ 1200 m, 42 ° 50 ' 37 " N 44 ° 18 ' 44 " E, 11.05.2016, leg. A. S. Sazhnev, 1 ex. (IBIW). Saratov Oblast: [Saratov Uezd], on the clay bank, 2.06.1900, leg. A. I. Jakovlev, 1 ex. (ZISP). Ulyanovsk Oblast: Novospassky Distr., 3 km E of Marievka vill., 53 ° 08 ' N 48 ° 09 ' E, 114 m, 7.07.2009, leg. A. V. Kovalev, 1 ex. (IBIW). CRIMEA. Tauria, 14.06.1911, leg. V. Pliginskiy, 1 ex (ZISP).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF82FFCAFF63AD7FFD0AFF40.taxon	distribution	Distribution in European Russia and Caucasus. ST: AST SAR * VL: SAM ULY CA: KCH KRD NOS (Isaev et al. 2001; Sazhnev 2014, 2017; Sazhnev & Prokin 2022). Crimea (state with limited recognition); Abkhazia (state with limited recognition); Azerbaijan (Sazhnev 2018 b). Distribution. West-Palearctic species was described from France, distributed in Europe, Caucasus, Near East (Syria), Asian Turkey and North Africa (Mascagni 2016; Sazhnev 2020).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF82FFCAFF63AD7FFD0AFF40.taxon	discussion	Remarks. The species recorded for Saratov Oblast for the first time. This species was recorded from Crimea [Tauria] as Augyles obliteratus (Plyginskiy 1928).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63A95CFEFEFC75.taxon	materials_examined	Material examined. 4 exs. RUSSIA: Orenburg Oblast: Orenburg, bank of Ural River, Petropavovskaya Grove, at light, 15.07.2012, leg. A. S. Ukrainsky, 4 exs. (IBIW).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63A95CFEFEFC75.taxon	distribution	Distribution in European Russia and Caucasus: VL: SAM UR: ORB (Litovkin et al. 2013: as Augyles sp.; Sazhnev & Kozminykh 2019). Armenia (Mascagni 2016). Distribution. Species distributed in Europe (including southern Ural) Asian Turkey, Mongolia and North Africa (Mascagni 2016; Sazhnev & Kozminykh 2019).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63A95CFEFEFC75.taxon	discussion	Remarks. Comparison of examined specimens (as well as Mongolian specimens) with European material and types is required. The diagnosis of the taxon Augyles marmota and the list of its synonyms needs revision. It is possible that more than one species is hidden under this name. [Augyles (Augyles) nebulosus (Kuwert, 1890)]	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63A95CFEFEFC75.taxon	distribution	Distribution in Caucasus. Azerbaijan: Agdash (Aresh, Geok Tepe), Mingachevir (Zaitsev 1946; Sazhnev 2018 b). Distribution. The species was described from Turkmenistan (Kuwert, 1890), is distributed in Caucasus (Azerbaijan) and Middle Asia (Turkmenistan, Uzbekistan) (Mascagni 2016).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63A95CFEFEFC75.taxon	discussion	Remarks. There is no indication for Azerbaijan in the Palaearctic Catalogue (Mascagni 2016), it is necessary to add E: AB.	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63AC66FDC7F9E2.taxon	materials_examined	Material examined. 8 exs. CRIMEA. Donuzlav bay, Krasnoyarskoe vill., 24.07.2016, leg. V. V. Shaporinsky, 1 ex. (IBIW). RUSSIA. Astrakhan Oblast: Krasnoyarsky Distr., env, Dosang vill., Akhtuba River, 46 ° 54 ' 30.50 " N 47 ° 54 ' 25.50 " E, at light, 1 – 8.05.2011, leg. I. A. Zabaluev, 1 ex. (IBIW); same place, in sediments, 28.04.2014, leg. A. Zaitsev & M. Pilipenko, 1 ex. (ZMMU); Akhtubinsky Distr., Bolkhuny vill., bank of Akhtuba River, 17.07.2014, leg. E. A. Negrobova, 1 ex. (IBIW). Karachay-Cherkess Republic: Urup Distr., Urup River near Pregradnoye vill., 43 ° 58 ' 25.2 " N 41 ° 12 ' 04.7 " E, 15.08.2016, leg. A. A. Prokin, 1 ex. (IBIW); Dagestan Republic: Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E. V. Ilyina, 2 exs. (IBIW). Orenburg Oblast: Gaysiy Distr., Khmelevka vill., at light, 7.05.2012 leg. A. S. Ukrainsky, 1 ex. (IBIW); Orenburg, bank of Ural River, Petropavovskaya Grove, at light, 15.07.2012, leg. A. S. Ukrainsky, 1 ex. (IBIW).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63AC66FDC7F9E2.taxon	distribution	Distribution in European Russia and Caucasus. ST: AST CA: DAG KCH UR: ORB (Sazhnev 2017; Sazhnev & Ilyina 2017; Sazhnev & Kozminykh 2019) Georgia (Mascagni 2016). Distribution. The species was described from Iraq, is distributed in South Europe, Caucasus, Near East (Iraq, Iran, Syria, Yemen), Asian Turkey, Middle Asia (Afghanistan, Kazakhstan, Turkmenistan, Uzbekistan) and Mongolia (Mascagni 2016; Sazhnev 2020).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63AEF9FBE0F85E.taxon	materials_examined	Material examined. 2 exs. AZERBAIJAN. Lankaran District: Lenkoran, leg. Leder (Reitter), 1 ex., (ZISP); Masally Distr.: Goryachie vody, Vilyash-chay River, 20.08.1932, leg. Znoyko, 1 ex. (ZISP).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63AEF9FBE0F85E.taxon	distribution	Distribution in Caucasus. Azerbaijan (this paper). Distribution. The species was described from France, is distributed in Europe, Near East (Iran, Iraq), Caucasus (Azerbaijan), Asian Turkey and Middle Asia (Afghanistan, Turkmenistan, Uzbekistan) (Mascagni 2016).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF81FFCAFF63AEF9FBE0F85E.taxon	discussion	Remarks. The species recorded for Azerbaijan (add E: AB) for the first time.	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF80FFCBFF63A8D2FAD8FB7F.taxon	materials_examined	Material examined. 18 exs. AZERBAIJAN. Qabala District: Nidzh vill, shore of Turianchay River, 38 ° 34 ' 58.6 " N 48 ° 47 ' 38.2 " E, 6.06.2017, leg. A. A. Prokin, 1 ex. (IBIW); Lankaran District: Lenkoran, leg. Leder (Reitter), 2 exs., as Augyles scutellatus (ZISP); Alekseevka vill., Hirkan National Park, at light, 11.07.1978, leg. V. V. Belov, 1 ex. (IBIW), same place, 13.07.1978, leg. V. V. Belov, 1 ex. (IBIW), same place, 15.07.1978, leg. V. V. Belov, 1 ex. (IBIW). GEORGIA. Mtskheta-Mtianeti Krai: Mtskheta Distr., Kura River opposite Dzegvi vill., 41 ° 51.199 ' N 44 ° 35.737 ' E, 10.08.2016, leg. D. Palatov, 3 exs. (IBIW). Samtskhe – Javakheti Krai: Borjomi, 3.05.1911, unknown collector, 1 ex. (ZISP). RUSSIA. Dagestan Republic: Magaramkent Distr., State Nature Zakaznik “ Samursky ”, 15 – 17.05.2015, leg. E. V. Ilyina, 1 ex. (IBIW), 29 – 30.07.2015, leg. E. V. Ilyina, 1 ex. (IBIW); Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E. V. Ilyina, 2 exs. (IBIW); Magaramkent Distr., Tagirkent-Kazmalar vill., 20.07.2014, leg. E. V. Ilyina, 1 ex. (IBIW). Krasnodar Krai: Cauc. occ., Adler, 08, leg. Dr. Lgocki, 1 ex., as Augyles scutellatus (ZISP); Tuapse Distr., env. Tuapse, Agoy River, 44.15 ° N, 39.06 ° E, 20.08.2016, leg. A. A. Prokin, 1 ex. (IBIW); Sochi, Adlersky Distr., Adler, Mzymta River, 2 – 3.09.2004, leg. S. A. Svobodny, 2 exs. (IBIW). Orenburg Oblast: Gaysiy Distr., Khmelevka vill., at light, 8.05.2012 leg. A. S. Ukrainsky, 1 ex. (IBIW).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF80FFCBFF63A8D2FAD8FB7F.taxon	distribution	Distribution in European Russia and Caucasus. CT: RZN VL: MAR / TAT ST: AST CA: DAG KRD UR: ORB (Jacobson 1913; Zaitsev 1916; Lebedev 1925; Litovkin et al. 2013; Sazhnev 2014; Sazhnev & Shapovalov 2016; Sazhnev 2017; Sazhnev & Ilyina 2017; Sazhnev & Kozminykh 2019). Azerbaijan: Agdash (Geok Tepe) (Zaitsev 1946, 1953; Sazhnev 2018 b), Elisabethpol Governorate (including as Augyles scutellatus) (Jacobson 1913), Ganja (Kirovabad, as Augyles vitticollis) (Zaitsev 1946), Lankaran (as Augyles vitticollis) (Zaitsev 1946), Talysh (as Augyles scutellatus) (Jacobson 1913); Georgia: Borjomi (Zaitsev 1946, 1953), Khashuri (Mikhailovo) (Schneider & Leder 1877; Zaitsev 1946), Kvemo Kartli Krai (Bolnisi — former Ekaterinenpheld, including as Augyles vitticollis) (Zaitsev 1946), Matani (Zaitsev 1946, 1953), Tbilisi (Tiflis, including as Augyles scutellatus) (Jacobson 1913). Distribution. European species (described from Switzerland), distributed in Caucasus, Near East (Israel, Syria) (Mascagni, 2016; Sazhnev 2020).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF80FFCBFF63A8D2FAD8FB7F.taxon	discussion	Remarks. By F. A. Zaitsev (1946) typical material of Augyles scutellatus (Motschulsky, 1854) was studied and this taxon to Augyles sericans was synonymized. Since Augyles vitticollis (Reitter 1884) is junior synonym of Augyles scutellatus (Mascagni 2016), both taxons should be referred to synonyms of Augyles sericans.	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF80FFCBFF63AD7EFCC0F890.taxon	materials_examined	Material examined. 37 exs. AZERBAIJAN. Baku: Sangachal, shore of Caspian Sea, 40 ° 09 ' 39.53 " N 49 ° 28 ' 20.53 " E, 6.06.2013, leg. D. G. Kasatkin, 5 exs. (IBIW); Gobustal, 6.06.2013, leg. I. V. Shokhin & D. G. Kasatkin, 8 exs. (IBIW). RUSSIA. Dagestan Republic: Magaramkent Distr., State Nature Zakaznik “ Samursky ”, 15 – 17.05.2015, leg. E. V. Ilyina, 7 exs. (IBIW); Magaramkent Distr., estuary of Samur River, Caspian Sea coast, at light, 5.06.2013, leg. E. V. Ilyina, 7 exs. (IBIW); Magaramkent Distr., Gazardkam-Kazmalar vill., 19.07.2014, leg. E. V. Ilyina, 9 exs. (IBIW); Magaramkent Distr., Tagirkent-Kazmalar vill., 20.07.2014, leg. E. V. Ilyina, 1 ex. (IBIW).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
515B87B6FF80FFCBFF63AD7EFCC0F890.taxon	distribution	Distribution in Caucasus. CA: DAG (Sazhnev 2016 b, 2017; Sazhnev & Ilyina 2017). Azerbaijan: Baku (Sazhnev 2018 b); Georgia (Mascagni 2016). Distribution. The species was from Turkmenistan, distributed in Caucasus, Near East (Arab Emirates, Iran, Iraq, Israel, Syria, Saudi Arabia), Asian Turkey, Middle Asia (Kazakhstan, Kirgizstan, Turkmenistan and Uzbekistan) and North Africa (Algeria) (Mascagni 2016; Sazhnev 2020).	en	Sazhnev, Alexey S. (2022): Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries genus Augyles of European part of Russia and Caucasus Region. Zootaxa 5209 (5): 581-593, DOI: https://doi.org/10.11646/zootaxa.5209.5.5
