taxonID	type	description	language	source
03FAAD53FFABFFC3FF045AF4E63F924D.taxon	description	(Figs 1 A, 2)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFABFFC3FF045AF4E63F924D.taxon	materials_examined	Additional specimens examined. EAST MALAYSIA • 1 ♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87554, E 117.94135, 54.5 ± 6.1 m. a. s. l.; 11 May 2022, 20 h 58; on the leaf litter; coll. M. K. Tan & T. Robillard; SBH. 22.19 (FRC) • 1 ♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87416, E 117.93786, 89.2 ± 6.8 m. a. s. l.; 18 May 2022, 20 h 07; on the leaf litter; coll. M. K. Tan & T. Robillard; SBH. 22.152 (MNHN) • 1 ♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87328, E 117.93749, 95.1 ± 5.6 m. a. s. l.; 18 May 2022, 20 h 57; on the leaf litter; coll. M. K. Tan & T. Robillard; SBH. 22.153 (FRC) • 1 ♂; Sabah State, Sepagaya Waterfall, near Lahad Datu; N 4.98354, E 118.14498, 182.4 ± 7.4 m. a. s. l.; 13 May 2022, 11 h 39; on the leaf litter; coll. M. K. Tan & T. Robillard; SBH. 22.50 (ZRC) • 1 ♂, 1 ♀; Sabah State, Mount Silam, lowland forest; N 4.97589, E 118.19060, 341 m. a. s. l.; 13 May 2022, 20 h; on the leaf litter; coll. M. K. Tan & T. Robillard; TR 22 - 10 (MNHN)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFABFFC3FF045AF4E63F924D.taxon	distribution	Distribution. Borneo, Sabah State: Sepilok, Mount Silam [new locality record], Sepagaya near Lahad Datu [new locality record]	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFABFFC3FF045AF4E63F924D.taxon	materials_examined	Type locality. EAST MALAYSIA, Sabah State, Sepilok Calling song (1 ♂, in captivity, 26.4 ° C) (Fig. 2). The calling song generally consists of a sequence of echemes which can be highly variable in duration (8 – 26 s). The echeme sequence typically begins with the echemes more erratic in terms of number of syllables, syllable durations and intervals. Each echeme is made up of 4 – 10 syllables (= pulses), and has an average duration of 140.9 ± 65.3 ms (78.0 – 249.1 ms). Consecutive echemes have an average interval of 371.6 ± 86.3 ms (285.6 – 474.3 ms). The average syllable duration is 14.1 ± 1.5 ms (11.9 – 15.6 ms) and the average interval between consecutive syllables is 7.2 ± 1.6 ms (5.4 – 9.7 ms). The dominant frequency is 8.34 kHz. Compared to the continuous trill of Cycloptiloides timah Ingrisch, 2006 from Singapore (see Tan et al., under review), the calling song of C. bimaculatus also differs by a distinctly shorter syllable duration (14.1 ms in C. bimaculatus vs. 25.7 ms in C. timah) and by the intervals between consecutive syllables (7.2 ms vs. 104.5 ms). The dominant frequency is also distinctly lower (8.3 kHz vs. 9.1 kHz). It should be noted that the average temperature of the recordings for the two species are different (26.5 ° C in C. bimaculatus vs. 30.5 ° C in C. timah), which may have contributed partly to the differences in the call parameters between the two species.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAEFFC2FF045FBFE69891A1.taxon	description	(Fig. 3)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAEFFC2FF045FBFE69891A1.taxon	materials_examined	Additional specimens examined. BRUNEI DARUSSALAM • 1 ♂; Temburong District, Kuala Belalong Field Studies Centre, along Sungei Mata Ikan, N 4.54742, E 115.15715, 73.8 ± 27.9 m. a. s. l.; 22 September 2016, 22 h 33; on a foliage of a tree after rain; coll. M. K. Tan; KB. 16.4 (ZRC) • 1 ♂; Temburong District, Kuala Belalong Field Studies Centre, along Sungei Mata Ikan, N 4.54733, E 115.15720, 113.4 ± 14.2 m. a. s. l.; 23 September 2016, 19 h 00; on a foliage of a tree after rain; coll. M. K. Tan; KB. 16.15 (ZRC)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAEFFC2FF045FBFE69891A1.taxon	discussion	Remarks. When Ectatoderus nigrofasciatus Tan et al., 2021 was first described, the basal part of the male genitalia of the holotype was not examined (Tan et al., 2021: Fig. 7). The apical part of the male genitalia, previously thought to be the complete genitalia, was considered to resemble that of Ectatoderus argentatus Ingrisch, 2006 from Thailand. However, after examining more specimens collected from Brunei Darussalam, it was found that the basal half of the male genitalia of E. nigrofasciatus has in fact the medial valve curved to a spiral. This character is more similar to that in Ectatoderus angusticollis Chopard, 1969 from Singapore (Fig. 3). Below, the diagnosis is revised accordingly. Revised diagnosis. The species is characterised by the scapus being dorsally black, but ventrally and laterally brown, and the antennae being entirely black basally; the head has a distinct black band behind eyes; and the male phallic complex is very large and forming a spiral. The new species is most similar and closest in distribution to E. angusticollis. In both species, the pronotum is more strongly widening toward apex, and the TIs and TIIs are provided with black dorsal stripes. The male phallic complex is very similar to that of E. angusticollis, including the spiral-shaped medial valve and the shape of the ventro-external sclerite. The species differs from E. angusticollis by the male phallic complex being exceptionally larger despite of having very similar body size (Fig. 3); the scapus and antennae being black (instead of testaceous); and the head being provided with distinct black and white bands behind eyes. The new species is also similar to E. argentatus having the pronotum strongly widening apically and the frons dark; it differs by the scapus and basal antennal segments being darkened instead of yellowish brown, and by the male phallic complex forming a spiral at base instead of being merely curved.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAEFFC2FF045FBFE69891A1.taxon	distribution	Distribution. Borneo, BRUNEI DARUSSALAM: Belait and Temburong [new locality record] Districts	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAEFFC2FF045FBFE69891A1.taxon	materials_examined	Type locality. BRUNEI DARUSSALAM, Belait District, near Andulau Forest Reserve	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAFFFCFFF0459A1E1E69315.taxon	description	(Figs 1 B, 4, 5)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAFFFCFFF0459A1E1E69315.taxon	discussion	Remarks. The specimen collected from Sabah exhibits some differences with O. pullus collected from Belait (Brunei Darussalam) and Singapore: legs with more whitish marmoration, supra-anal plate whitish when alive. However, we could not identify clear-cut differences in the genitalia, including both the sclerotized median valve and the membranous lateral valves between the specimen from Sabah and those from Brunei Darussalam (Fig. 5). As Tan et al. (2021) have also suggested that there might be a polymorphic species and that more evidence was needed (e. g., DNA or acoustics) to better understand the species boundaries. At the moment, the external morphology and genitalia morphology are not congruent between specimens from Brunei Darussalam, Singapore and Sabah; we also do not have acoustic data for the Brunei population; and no molecular data are available. Furthermore, the specimen from Sabah also shares some similarities (but also with clear differences) in the external morphology of Ornebius minusculus (Chopard, 1929) from Sipura Island (Sumatra) (http: // coldb. mnhn. fr / catalognumber / mnhn / eo / ensif 4531), although the genitalia for this species has not been examined. As such, without more specimens from Sabah to examine variations, we refrained from describing it as a new species, tentatively identified it as O. pullus and preferred to revise its taxonomy when the chance arises.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAFFFCFFF0459A1E1E69315.taxon	distribution	Distribution. Brunei Darussalam (Belait and Brunei-Muara Districts), East Malaysia (Sabah State) [new locality record], Singapore.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFAFFFCFFF0459A1E1E69315.taxon	materials_examined	Type locality. BRUNEI DARUSSALAM, Brunei-Muara District, near bridge over river Sungai Lubang Barus on road coming from Tutong. Calling song (1 ♂, in captivity, 31.5 ° C) (Fig. 4). The calling song generally consists of a sequence of syllables (= pulses), which can be highly variable in duration (2 – 14 s or 4 – 25 syllables). The average syllable duration is 37.5 ± 5.0 ms (31.0 – 46.0 ms) and the average interval between consecutive syllables is 501.2 ± 20.4 ms (469.3 – 538.4 ms). The dominant frequency is 7.97 ± 0.13 kHz (7.78 – 8.16 kHz). Compared to the calling song of Ornebius pullus from Singapore (see Tan et al., under review), the calling song of this specimen from Sabah has a similar call structure (i. e., the calls of both species are made up of a sequence of syllables). The biggest difference is in the interval between consecutive syllables being distinctly longer (501.2 ms vs. 400.0 ms) than that from Singapore. The syllable is only marginally longer in duration on average (37.5 ms vs. 32.4 ms) and the dominant frequency is not different (8.0 kHz vs. 8.1 kHz).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	description	(Figs 1 C, 1 D, 6, 7) Ornebius sp. — Tan et al. under review [calling songs from Singapore] Specimens examined. Holotype: SINGAPORE • 1 ♂; Pasir Ris mangrove boardwalk; N 1.37805, E 103.95147; 19 March 2022, night time; on foliage of mangrove tree; coll. M. K. Tan; ZRC Paratypes: SINGAPORE • 1 ♂; Pasir Ris mangrove boardwalk; N 1.37805, E 103.95147; 2 May 2022, night time; on foliage of mangrove tree; coll. M. K. Tan; MNHN-EO-ENSIF 11343	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	diagnosis	Diagnosis. The new species has genitalia which appear most similar to those in the species group Ornebius (rufonigrus) Ingrisch, 1987, but differs by the dark-grey colouration; the maxillary palps being not so elongated (more similar to Ornebius alvarezi Tan, Ingrisch, Baroga-Barbecho & Yap, 2019 from the Philippines; elongated oblong in Ornebius dumoga Ingrisch, 2006 from Sulawesi and in Ornebius consternus Ingrisch, 2006 from Sulawesi), the FW colouration darker (orange in O. dumoga and in Ornebius rufonigrus Ingrisch, 1987 from Indochina and Malay Peninsula) and less covered by the pronotum (compared to O. consternus). The male genitalia of this new species differ from congeners from the species group by the internal sclerites at base being smaller instead of expanded. We also compared the new species with other species currently not placed in any species group (i. e., with unknown male genitalia). Our species resembles species with wholly dark / black or smoky FWs, such as Ornebius obscuripennis (Chopard, 1930) from Sarawak and Ornebius fuscipennis (Chopard, 1929) from Sumatra (including Padang, Pulau Sipura, Pulau Telo and Batu Islands). The new species differs from O. obscuripennis by the pronotal lateral lobe and lateral parts of the head being dark-grey (i. e., unicolorous with dorsal parts) instead of forming a yellow lateral band. It also differs from O. fuscipennis by much smaller body size (BL 8.4 vs. 10.5 mm; PL 2.8 vs. 4.5 mm; TL 2.5 vs. 2 mm; FIIIL 4.4 vs. 7 mm); by the FW more uniformly coloured instead of smoky with posterior margin blackish; by the apical segment of the maxillary palps longer than subapical and third; by the pronotal disc not covered with yellow-brown scales; and by the legs that are not unicolourous yellowish. Lastly, the new species also differs from a syntopic species, Ornebius tampines Tan & Robillard, 2012 by the overall body and FW colouration, the body shape is more typical of Ornebius, instead of slenderly built, the shape of the male genitalia and by the calling songs (see Tan et al., under review). It is also somewhat similar to a sympatric species, O. pullus by the wholly black FWs, but differs by the lack of white margins along the lateral lobe and the posterior margin of the dorsal disc of the pronotum, by the shape of male genitalia and by the calling songs (see Tan et al., under review)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	etymology	Etymology. The species name refers to the dark-grey colouration, resembling the grey-wolf, Canis lupus.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	description	Description. Small species; habitus of male as shown in Fig 1 C, 1 D. Dorsum of head flattened, covered with dark-grey scales (Figs. 6 A). Frontal rostrum 2.0 times wider than scapus (Fig. 6 A). Scapus dark-grey (Fig 6 A); basal antennal segments black, thereafter brown with some segments black (Figs 6 B, 6 C). Frons and mouthparts of dark colour (Fig. 6 B). Maxillary palps pale grey; with apical (= fifth) segment longer than subapical (= fourth) segment, third segment of subequal length as subapical segment; apical segment pyriform (Fig. 6 C). Lateral parts of head, including genae, of dark colour, without bands (Fig. 6 C). Pronotal disc brown and wholly covered with dark-grey scales, about 1.3 times longer than wide, with anterior margin narrow and straight; lateral margin widening slightly posteriorly; covering only base of mirror of FW; posterior margin convex (Fig. 6 D). Lateral lobe of pronotum also wholly covered with dark-grey scales (Figs 1 D, 6 C). TI with internal tympanum small and oval; without external tympanum. FIII 1.4 – 1.5 times longer than TIII; TIII 2.6 times longer than MTaIII. Femora generally pale, covered with marmoration of dark-grey scales except in basal parts. Tibiae and tarsi generally grey. Abdominal tergites covered with dark-grey scales; sternites pale. Abdominal apex darkened. Cercus covered with grey scales. Male. FW mostly infumated black, dorsal field darkest at posterior margin (Fig. 6 D); lateral field likewise in some areas with infumated white veins (Fig. 6 E). Supra-anal plate with last abdominal tergite and epiproct distinctly separated by a transverse suture. Last abdominal tergite transverse, posterior margin abruptly indented in middle; with oval pale spot in middle (not obvious in dry-pinned specimen) (Fig. 6 F vs. 6 G); posterior margin on either side of the indentation with two patches of short and strong setae. Supra-anal plate less transverse than last abdominal tergite, black coloured, apex subtruncate (Figs 6 F, 6 G). Subgenital plate black (Fig. 6 H). Paraproct process black, elongate, cylindrical and tapering into a subacute apex (Fig. 6 H). Genitalia as in Fig. 7. Central lobe of phallus with lateral valves (lv) elongate, apical area curved slightly ventrad in lateral view; tapering into obtuse apex, in apical area with inner margin curved in dorsal view; both sides together almost forming a dorsally and ventrally open tube. External sclerite (es) well developed, with external part more strongly sclerotized than inner part (in dorsal view). Medial valve (mv) membranous with base widened (in dorsal view) and upcurved (in lateral view). Internal sclerites (is) at base small but widened and curled, afterwards straight and elongate. Female. Unknown. Measurements (2 ♂, in mm). ♂ Holotype: BL = 8.4; PronL = 2.8; PronW = 2.2; FWL = 2.5; FWW = 2.3; FIIIL = 4.4; TIIIL = 3.1; MaTIIIL = 1.2. ♂ paratype: BL = 8.4; PronL = 2.8; PronW = 2.2; FWL = 2.5; FWW = 2.3; FIIIL = 4.4; TIIIL = 2.9; MaTIIIL = 1.1.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	biology_ecology	Ecology. This species is found among foliage of mangrove trees, occurring in syntopy with another mangrovespecialist, Ornebius tampines.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	distribution	Distribution. SINGAPORE	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
03FAAD53FFA2FFCDFF045E87E37095B1.taxon	materials_examined	Type locality. SINGAPORE, Pasir Ris mangrove Calling song. Described in Tan et al. (under review). The calling song of this species is similar to that of Ornebius rufonigrus Ingrisch, 1987 from the same species group with regard to echeme duration and dominant frequency, although the new species produces echeme-sequences composed of triple-chirps instead of double-chrips as in O. rufonigrus.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul, Robillard, Tony (2022): Taxonomy and bioacoustics of scaly crickets (Orthoptera, Mogoplistidae Mogoplistinae) from Borneo and Singapore. Zootaxa 5213 (2): 177-189, DOI: https://doi.org/10.11646/zootaxa.5213.2.6
