taxonID	type	description	language	source
03FFB25377777A6FDD90FE106DAF57E1.taxon	diagnosis	Diagnosis. Body medium-sized (4.50 – 7.20 mm); pronotum moderately narrow, usually sharply narrowed posteriad toward subparallel-sided mediobasal portions; elytra moderately short, slightly broader than long, slightly or significantly broadened posteriad; aedeagus long, with moderately small basal part, gradually narrowed toward subacute or rounded apex, with a pair of sclerotized dorsal lobes in apical portion, without (G. (s. str.) zwickianus) or with two narrow and elongate middle structures, and with moderately short flagellum.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6FDD90FE106DAF57E1.taxon	discussion	Species included. G. (s. str.) bodemeyeri Bernhauer, 1902, G. (s. str.) championi Cameron, 1924, G. (s. str.) zwickianus Fagel, 1976.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6FDD90FE106DAF57E1.taxon	distribution	Distribution. Species of the bodemeyeri group are distributed in the Palaearctic Region, and known from Causasus, Turkey, Iran and the Himalayan Region. Bionomics. The species of the bodemeyeri group inhabit banks of rivers, streams and lakes, and can be found in gravel, under stones, etc. Specimens were collected at elevations from 180 to 3800 m a. s. l. from April to October.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6FDD90FE106DAF57E1.taxon	discussion	Remarks. This group included G. (s. str.) zwickianus and G. (s. str.) bodemeyeri, which were included by Bordoni (1984) to major and rousi groups respectively. Based on the shape of the parameres with wide apical portions and weakly sclerotized internal sac and narrow flagellum, the bodemeyeri group is similar to the convexicollis group of the nominal subgenus Geodromicus, but differs by the presence of paired sclerotized structures in apical and middle portions (except G. (s. str.) zwickianus) of the median lobe. Key to bodemeyeri group	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6BDD90F97F6CDE5462.taxon	description	(Figs. 1, 7 – 12, 25)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6BDD90F97F6CDE5462.taxon	materials_examined	Type material examined. Holotype of Geodromicus bodemeyeri Bernhauer, 1902, ♂: ‘ Asia minor | Bulghar Maaden | v. Bodemeyer’ <printed>, ‘ unbekannt’ <handwritten>, ‘ Bodemeyeri | Brnh Type. ’ <handwritten>, ‘ Chicago NHMus | M. Bernhauer | Collection’ <printed>, ‘ HOLOTYPE | teste D. J. Clarke 2014 | GDI Imaging Project’ <violet, printed>, ‘ PHOTOGRAPHED | Kelsey Keaton 2014 | Emu Catalog’ <blue, printed>, ‘ FMNHINS | 2819535 | FIELD MUSEUM’ <printed, with barcode on left side of the label> (FMNH). The photographs of the habitus and type labels of the holotype are available in Arthropod Collections Database of FMNH (last access: 02.11.2022). Additional material examined. RUSSIA: KABARDINO-BALKARIA: 1 ♂: ‘ Caucasus, Naltshik, rip-fl. [?], ix. [19] 60 ’ (ZMUC); NORTH OSSETIA-ALANIA: 1 ♂: Alagir, riverbank of Ardon River. 650 m a. s. l. 06.06.1984. S. K. Alexeev leg. (cSh); TURKEY: IZMIR: 1 ♂: ‘ ♂ ’, ‘ Anamas Gbg. ’, ‘ Kl [ein]. - As [ien]. ’, ‘ Pisidischer’, ‘ Taurus’, ‘ Weirather’, ‘ Inssbruck’, ‘ bodemeyeri Bernh. ’, ‘ ex coll. Scheerpeltz’, ‘ Geodromicus bodemeyeri B. det. A. Bordoni 1983 extraxit et delin. ’ (NMW); BURSA: 2 ♀♀: Uludað. 31.07.1988. M. Jäch leg. (NMW); BOLU: 1 ♀: Abant. 1400 m a. s. l. 17.07.1971. A. Vigna leg. (cB); NIÐDE: 1 ♂: ‘ Asia Minor Bulghar [Bolghar Dagh] Maaden v. Bodemeyer’, ‘ Bodemeyeri Bh. det. Bernh. ’ (FMNH); GüMüşHANE: 1 ♀: Kösedagi Pass. Stream, 2 – 3 m wide, granite. 29.05.1989. M. Jäch leg. (NMW); ERZINZAN: 2 ♂♂, 1 ♀: 10 km SE Refahiye. Small creeks, volcanic rock, with minor calcareous rocks. 10.06.1989. M. Jäch leg. (NMW); ERZURUM: 1 ♂: Tortum-Narman. Small creek at Kirecli Pass, ca. 2200 m a. s. l., as well as small spring-fed creek E of the pass at 2300 m a. s. l. 09.06.1989. M. Jäch leg. (NMW); KARS: 1 ♂: Sarikamis. Small streams flowing through meadows, 2 – 5 m wide, ca. 10 km S Sarikamis, ca. 2000 m, volcanic; also small artificial forest pond in pine forest. 08.06.1989. M. Jäch leg. (NMW); 1 ♂: Kagizman. 08.06.1989. S. Schödl leg. (NMW).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6BDD90F97F6CDE5462.taxon	description	Redescription. Measurements (n = 14): HW: 0.92 – 1.07; HL: 0.59 – 0.72; OL: 0.27 – 0.32; TL: 0.15 – 0.19; AL (averaged): 3.40; PL: 0.94 – 1.25; PWmax: 1.12 – 1.47; PWmin: 0.83 – 1.02; ESL: 1.68 – 1.92; EW: 1.69 – 2.17; MTbL (averaged): 1.35; MTrL (averaged): 0.66 (MTrL 1 – 4: 0.34; MTrL 5: 0.32); AW: 1.61 – 2.42; AedL: 1.12 – 1.18; BL: 5.20 – 7.00 (holotype of G. (s. str.) bodemeyeri: 5.50). Habitus as in Fig. 1. Body brown to reddish-brown, with paler elytra and abdomen (holotype paler, with darker head and yellowish large spots on each elytron); mouthparts, antennae and legs yellow-brown to brown (apical segment of maxillary palpi usually yellow, some specimens with yellowish antennomeres 1 – 2 or 1 – 3, femuri and tarsi). Body glossy; forebody without microsculpture, except neck with very dense isodiametric meshes and abdominal tergites with dense transverse or isodiametric microreticulation (some specimens with fine, dense, transverse or isodiametric meshes between ocelli). Pubescence of forebody white or yellow, very dense, moderately long, semierect, longer on apical portion of head; abdomen with dense, decumbent pubescence, slightly shorter and finer than that on forebody. Head 1.4 – 1.5 times as broad as long, slightly convex, more elevated between ocelli and eyes, frontal portion with slightly convex supra-antennal elevations, with relatively deep, wide and subtriangular anterio-median depression, narrowed and extended basad to level of anterior margins of eyes, sometimes connected with interocellar depression; temples moderately convex and long, slightly less than twice as long as longitudinal length of eyes; interocellar depression subrectangular, moderately large and deep, separated from infraorbital portions by distinctly convergent latero-anteriad narrow and deep anteocellar foveae (grooves in front of ocelli), reaching level of apical third of eyes. Eyes moderately large and convex. Ocelli small, distance between ocelli about as long as distance between ocellus and posterior margin of eye, or slightly shorter. Punctation irregular, moderately fine, dense or sparse, sometimes denser in middle and / or infraorbital portions. Maxillary palpomere 3 about as long as preceding segment, significantly broadened apicad; apical palpomere distinctly shorter than 3, from middle gradually narrowed toward subacute apex. Antenna long, reaching apical third or middle of elytra when reclined; basal antennomere robust and long, about three times as long as broad, antennomere 2 distinctly narrower and about twice shorter than basal antennomere, 3 about as broad as and 1.3 – 1.6 times as long as 2, 4 slightly shorter than 3, 5 – 8 distinctly longer than 4, 9 – 10 slightly broader and longer than 8, apical antennomere 1.2 – 1.4 times as long as 10, from apical third gradually narrowed toward subacute apex. Pronotum convex, slightly broader than long, from widest anterior third gradually narrowed toward widely rounded anterior and sharply narrowed toward subacute hind angles; short laterobasal margins somewhat subparallel; middle portion without longitudinal depression, or with traces of it in middle in some specimens, with shallow or deep transverse mediobasal depression; anterior margin slightly concave, straight or rounded, about as that or slightly shorter than straight posterior margin. Punctation regular, very dense, slightly larger and deeper than that in middle of head, with interspaces between punctures in middle about as long diameters of two – four nearest punctures, sparser in medioapical portion in some specimens, mediobasal portion without punctation. Scutellum large, without punctation, with fine transverse meshes or without them, with elongate subtrangular apex. Elytra convex, slightly broader than long, slightly broadened posteriad, 1.5 – 1.7 times as long as pronotum; lateral margins narrowly flattened and slightly reflexed in latero-apical portions; hind margins straight or slightly rounded. Punctation variable, about as that on pronotum, slightly denser and larger or finer and sparser, usually finer in middle along suture. Legs slender; protarsi about 1.3 times as long as protibia; meso- and metatarsi distinctly longer, about 1.2 times as long as tibia; apical metatarsomere about as long as preceding four segments. Abdomen slightly narrower, as broad as or slightly broader than elytra, with two large, transverse tomentose spots in middle of abdominal tergite IV and narrow palisade fringe on apical margin of abdominal tergite VII. Male. Pronotum usually wide, strongly convex. Profemuri wide or very wide in some specimens; protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII straight or sligthly concave. Apical margin of abdominal sternite VIII widely concave. Aedeagus with moderately small basal part, gradually narrowed toward wide median lobe, and from apical third of median lobe slightly (Figs. 7, 9 – 11) or sharply (Fig. 12) narrowed toward small rounded apex; parameres broadened toward moderately wide apical portions, exceeding apex of median lobe or slightly shorter (Fig.), with four moderately long apical setae; median lobe with two sclerotized, elongate and concave dorsal lobes in apical portion and two narrow and very long, sclerotized, parallel structures in middle portion; internal sac with narrow and moderately long flagellum, stretching from basal portion toward median sclerotized structures, with two indistinct and very fine fields of thorns in basal or middle portion (Figs.). Lateral aspect of the aedeagus as in Fig. 8. Female. Pronotum moderately narrow, less convex. Profemuri and protarsomeres 1 – 4 narrow. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII straight or rounded. Comparative notes. Based on the general shape of the body and the aedeagus, G. (s. str.) bodemeyeri is similar to G. (s. str.) zwickianus, from which it can be distinguished by the slightly narrower pronotum and the presence of the sclerotized parallel structures in the middle of the median lobe.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6BDD90F97F6CDE5462.taxon	distribution	Distribution. Geodromicus (s. str.) bodemeyeri is distributed in the Caucasus and Turkey (Fig. 25). Bionomics. Specimens were collected at elevations from 650 to 2300 m a. s. l. and some specimens were taken from under stones and gravel along river banks. One specimen (“ G. convexus ”) was collected near shore of the lake (Khnzorian 1962).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377777A6BDD90F97F6CDE5462.taxon	discussion	Remarks. Geodromicus bodemeyeri based on the holotype was originally described from “ Kleinasien, Bulghar Maaden ”. Bordoni (1984) provided morphological features of G. (s. str.) bodemeyeri, including sketchy figure of the aedeagus (Bordoni (1984): fig. X. 1) and include it to the rousi group (Shavrin 2021). Geodromicus convexus based on three males was originally described from Armenia (Sevan Lake and Megri, Araks River) and Azerbaijan (Nüsnüs, Ordubadskiy District). I`m not studied the type material, but the studied male from Naltshik (Kabardino-Balkaria) is conspecific with types and it was compared with them by A. Solodovnikov in 2000. The aedeagus (Fig. 11) and the description of this species also corresponds with the external morphological features of G. (s. str.) bodemeyeri. Geodromicus rivularis was originally described based on male and female from Araks River, Armenia (the same locality together with G. convexus). Iablokoff-Khnzorian (1989) provided the figure of the aedeagus of the holotype (Iablokoff-Khnzorian (1989): fig. 2), which very similar in the shape with the aedeagus studied by me in specimen from North Ossetia-Alania (Fig. 12). In the same article, Iablokoff-Khnzorian (1989) synonymized G. convexus with G. rivularis. This synonymyzation was not reported in catalogues (e. g. Herman (2001), Schülke & Smetana (2015 )). Morphological features provided in the original description of G. rivularis correspond with external and internal morphological features of G. (s. str.) bodemeyeri. Thus, I synonymyzed G. rivularis with it.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377737A64DD90F9286C795102.taxon	description	(Figs. 2, 13 – 15, 27)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377737A64DD90F9286C795102.taxon	materials_examined	Type material examined. Lectotype (here designated) of Geodromicus championi Cameron, 1924, ♀ (Fig. 2): ‘ SYN- | TYPE’ <printed label with blue margins>, ‘ Type | H. T. ’ <round label with red margin>, ‘ Chakrata Dist. | Sanj Khud 6500`’ <printed>, ‘ Dr. Cameron. | 27. V. [19] 22. ’ <printed>, ‘ Geodromicus | championi | Cam. ’ <handwritten>, ‘ M. Cameron | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ LECTOTYPE | Geodromicus | championi | Cameron, 1924 | Shavrin A. V. des. 2016 ’ <red, printed> (BMNH). Paralectotypes: 2 ♂♂ (one male dissected, other male damaged), 4 ♀♀ (one female without antennomeres 3 – 6): ‘ Chakrata Dist. | Sainj Khud 6500`’ <printed>, ‘ SYN- | TYPE’ <printed label with blue frame>, ‘ Dr. Cameron | 27. V. [19] 22. <printed>, ‘ G. championi Cam. ’ <handwritten>, ‘ M. Cameron | Bequest. | B. M. 1955 - 147. ’ <printed> (BMNH). Paralectotype, ♀: ‘ Chakrata Dist. Sanj Khud 6500 ’, ‘ Dr. Cameron 27. V. [19] 22. ’, ‘ Geodromicus championi Cam. ’ (NMPC). Holotype of Geodromicus nepalicus Coiffait, 1977, ♂ (dissected): ‘ Ufer des Tila Khola | b. Jumla, 26.9. - 4.10.72 ’ <printed>, ‘ Gebiet von Jumla | Westnepal, lg. H. Franz’ <printed>, ‘ Geodromicus | spec. | det. H. Franz [printed] ’ <handwritten>, ‘ TYPE’ <red, printed>, ‘ Geodromicus | nepalicus | H. COIFFAIT [printed] det. 19 [printed] 76 ’ <handwritten> (NMW). Paratype, 1 ♀: ‘ Sinja Khola zw. Chauta u. Neurigad’ <printed>, ‘ Paratype’ <red>, ‘ Geodromicus nepalicus H. Coiffait 1976 ’ <handwritten> (MHNH). Additional material examined. PAKISTAN: 1 ♂: Chitral Bumburet. 2200 – 2350 m a. s. l. 24.05.1983. C. Besuchet & I. Löbl (MHNG); NEPAL: KARNALI: 1 ♂, 2 ♀♀: Gothichaur, 29 ° 12 ′ 10 ′ N 82 ° 18 ′ 56 ′′ E. 2800 m a. s. l., river bank. 08.06.1997. H. F. Grill leg. (NME); 1 ♂: Jumla, 12 km E Jumla, Jharjwala. 2500 m a. s. l. 01.05.1995. A. Weigel leg. (cSh); 2 ♀♀: Jumla, Tila River, NN. 2300 m a. s. l. 29.04.1995. M. Hartmann & A. Weigel leg. (cSh, NME); 1 ♂: same data. J. Weipert leg. (NME); GANDAKI: 1 ♂, 1 ♀: SW Dhaulagiri, E Dhorpatan, 28 ° 30 ′ 08 ′′ N 83 ° 07 ′ 45 ′′ E. 3000 m a. s. l. 22.09.2012. J. Schmidt leg. (NME).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377737A64DD90F9286C795102.taxon	description	Redescription. Measurements (n = 15): HW: 1.06 – 1.20; HL: 0.64 – 0.80; OL: 0.32 – 0.40; TL: 0.10 – 0.12; AL (averaged): 3.54; PL: 1.04 – 1.40; PWmax: 1.43 – 1.80; PWmin: 0.96 – 1.25; ESL: 1.91 – 2.35; EW: 2.14 – 2.55; MTbL (averaged): 0.68; MTrL (averaged): 0.32 (MTrL 1 – 4: 0.17; MTrL 5: 0.15); AW: 1.95 – 2.45; AedL: 0.82 – 1.00; BL: 5.80 – 7.20 (lectotype of G. championi: 7.20; holotype of G. nepalicus: 6.20). Habitus as in Fig. 2. Body reddish-brown to black (some specimens with paler elytra); antennae brown (some specimens with paler antennomeres 1 – 2); mouthparts and legs yellow-brown to black. Forebody without microsculpture except some specimens with traces of indistinct transverse microreticulation in apical portion of head, pronotum with indistinct transverse meshes (some specimens) and abdominal tergites with very dense transverse microsculpture. Head 1.5 – 1.6 times as broad as long, frontal portion with strongly convex supra-antennal elevations, with deep and wide median depression, gradually narrowed and extended basad; temples moderately short, distinctly concaved between posterior margin of eye and smooth postocular ridge (indistinct in some specimens); interocellar depression very deep, moderately narrow or wide; anteocellar foveae deep, reaching level of middle or apical third of eyes. Distance between ocelli 1.2 – 1.4 times as long as distance between ocellus and posterior margin of eye. Punctation moderately large and dense, sometimes finer and sparser in middle, and larger and deeper on infraorbital portions. Antenna very long, usually exceeding middle of elytra when reclined; antennomere 2 1.3 times as long as basal antennomere, 3 distinctly longer than 2, 4 – 9 slightly shorter than 3, 10 slightly shorter than 9, apical antennomere 1.2 – 1.3 times as long as 10. Pronotum slightly transverse, 1.4 – 1.5 times as broad as long, from widest anterior third strongly narrowed apicad toward subparallel and moderately long laterobasal margins; middle portion without or with indistinct longitudinal and transverse, moderately deep mediobasal depressions. Punctation very dense, distinctly larger and deeper than that in middle of head, sparser in middle and mediobasal portions; mediobasal portion of some specimens sometimes without punctation. Elytra slightly or strongly broadened posteriad, 1.6 – 1.8 times as long as pronotum. Punctation very dense, about as that on pronotum or distinctly denser, with interspaces between punctures as long as diameter of one – two nearest punctures, sometimes denser and coarser in parascutellar portion and along suture. Metatibia twice as long as metatarsus. Abdomen slightly narrower than elytra, with two moderately small transverse tomentose spots in middle of abdominal tergite IV. Male. Protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII straight or slightly concave. Apical margin of abdominal sternite VIII widely concave. Aedeagus with small basal part, gradually narrowed apicad and broadened in about middle of wide median lobe; apical third of median lobe strongly narrowed toward apical portion and from apical portion gradually narrowed toward subacute apex; parameres broadened in apical portion, with four moderately long apical setae; median lobe with two very long sclerotized structures in middle; internal sac with narrow, with very long flagellum, from basal portion stretching apicad between parallel median structures (Figs. 13, 15). Lateral aspect of the aedeagus as in Fig. 14. Female. Protarsomeres 1 – 4 narrow. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded. Comparative notes. Geodromicus (s. str.) championi can be distinguished from other species of the bodemeyeri group by the darker coloration of the body, more transverse pronotum, the shape of the apical portion of the aedeagus and longer flagellum.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377737A64DD90F9286C795102.taxon	distribution	Distribution. Geodromicus (s. str.) championi is widely distributed in the Himalayan Region and known from Pakistan, India (Uttarakhand), and Nepal (Fig. 27). Bionomics. Specimens were collected at elevations from 1980 to 3000 m a. s. l. and were taken from under stones and gravel along river banks.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377737A64DD90F9286C795102.taxon	discussion	Remarks. Based on the unspecified number of syntypes, G. (s. str.) championi was originally described from “ Chakrata District; Sanj Khud and Bina, Gad, 6500 to 8000 feet above the sea ” (India, Uttarakhand). The specimen with the best preservation and additional label “ Type H. T. ” was designated as the lectotype in order to fix the identity of the name. Geodromicus (s. str.) nepalicus was described based on the holotype from “ Rive du Tila Khola, près Jumla, Népal occidental … ”. During the study of the holotype, I have not found sufficient external and internal morphological differences between it and other studied specimens of G. championi. Thus, it was synonymized with the latter species. Geodromicus (s. str.) championi is here recorded from Pakistan for the first time.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777C7A61DD90FC886D5155DA.taxon	description	(Figs. 3 – 6, 19 – 25)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777C7A61DD90FC886D5155DA.taxon	materials_examined	Type material examined. Holotype of Geodromicus zwickianus Fagel, 1976, ♂ (dissected): ‘ SO-ANATOLIEN | Ost-Taurus bei | Sürgü | 17.5.1970 Zwick’ <printed>, ‘ det. P. Zwick | Geodromicus [handwritten] ’ <printed>, ‘ G. Fagel det., 197 [printed] 2 | Geodromicus | zwickianus | n. sp. ’ <printed>, ‘ TYPE’ <pink, printed label with black margins>, ‘ Holotypus ♂ | Geodromicus | zwickianus | Fagel | V. Gusarov rev. 1993 ’ <red, handwritten>, ‘ Geodromicus ♂ | zwickianus Fagel | V. I. Gusarov det. 199 [printed] 3 ’ <handwritten>, ‘ Geodromicus | zwickianus Fagel, 1976 | Shavrin A. V. det. 2019 ’ <printed> (MHNG). Holotype of Geodromicus iranicus Coiffait, 1981, ♂ (Fig. 3; dissected): ‘ IRAN, Schalous Pass | 2600 – 2900 m, Elbursgeb. | 2. IX. 1960 | leg. J. Klapperich’ <printed>, ‘ TYPE’ <red, printed>, ‘ Geodromicus | iranicus | Coiffait’ <handwritten>, ‘ Geodromicus | bodemeyeri | Bernhauer, 1902 ’ (MHNG). Paratypes: 2 ♀♀ [one paratype without left antennomeres 2 – 11]: ‘ IRAN, Schalous Pass | 2600 – 2900 m, Elbursgeb. | 2. IX. 1960 | leg. J. Klapperich’ <printed>, ‘ PARATYPE’ <red, printed>, ‘ Geodromicus | iranicus | Coiffait’ <handwritten in blue> (MHNG); 1 ♂, 1 ♀: same data as the previous (MHNH). Additional material examined. TURKEY: HAKKâRI: 5 ♂♂, 3 ♀♀: NE of Yüksekova. 2100 m. 20.08.1970. Exp. Nat. Mus. Praha (NMPC, cSh); ANTALYA: 1 ♂: Gömbe b. Elmali, 36 ° 33 ′ N 29 ° 37 ′ E. 1800 m a. s. l., river bank. 07.06.2006. W. Marggi leg. (cSch); IRAN: ARDABIL: 1 ♂, 1 ♀: Meshkinshahr, Lahrud, Shabil, 38 ° 19 ′ 43.8 ′′ N 47 ° 50 ′ 40.4 ′′ E. 2660 m a. s. l. 05.08.2005. S. Serri & J. Frisch leg. (MNHUB); 1 ♀: Nir, Gor Gor fall, 38 ° 10 ′ 13.8 ′′ N 47 ° 54 ′ 10.5 ′′ E. 2440 m a. s. l. 06.08.2005. S. Serri & J. Frisch leg. (MNHUB); 2 ♂♂, 5 ♀♀: N of Nir, Alvares, Kuh-e Sabalan, 38 ° 12 ′ 07 ′′ N 47 ° 52 ′ 57 ′′ E. 2960 m a. s. l. 06.08.2005. J. Frisch leg. (MNHUB); 1 ♂, 1 ♀: S Lahrud, Gholorsuie, Kuh-e Sabalan, 38 ° 19 ′ 36 ′′ N 47 ° 51 ′ 24 ″ E. 2530 m a. s. l. 05.08.2005. J. Frisch & S. Serri leg. (MNHUB); ÂZARBâIJâN-E GHARBÎ: 1 ♂: 15 km S Mešginšahr, 38 ° 15 ′ N 47 ° 45 ′ E. 2900 m a. s. l. 20.06.2000. E. P. Hajdaj leg. (cSh); 1 ♂, 1 ♀: Khoyu – Qotur road: 2 km W Qotur, Qotur River, 38 ° 28 ′ 58 ″ N 44 ° 22 ′ 37 ″ E. 1950 m a. s. l. 29.08.2008. J. Frisch & S. Serri leg. (MNHUB); 1 ♀: Khoy-Siyah Shesmeh road, 9 km W Zar Abad, 38 ° 47 ′ 15 ″ N 44 ° 32 ′ 08 ″ E. 1970 m a. s. l. 30.08.2008. J. Frisch & S. Serri leg. (MNHUB); ÂZARBâIJâN-E SHARQI: 1 ♂, 1 ♀: S Tabriz, Kandovan, Kuh-e Sahand, 37 ° 47 ′ 29 ″ N 46 ° 15 ′ 02 ″ E. 2250 m a. s. l. 08.08.2005. J. Frisch leg. (MNHUB); 1 ♀: Shargi, 10 km N Kaleyher. 17 – 18.05.2007. A. Anicthchenko leg. (cSh); GILAN: 1 ♀: S Astara, 5 km W Lavandvil: Koteh Komeh, 38 ° 18 ′ 26 ″ N 48 ° 47 ′ 14 ″ E. 180 m a. s. l. 10.10.2011. J. Frisch leg. (MNHUB); HAMADAN: 1 ♀: Hamedan sur Hamedan, 34 ° 44 ′ N 48 ° 27 ′ E. 2600 m a. s. l. 11.09.1975. A. Senglet leg. (MHNG); MAZANDARAN: 1 ♂, 1 ♀: W Polour, Demavend (N). 2400 m a. s. l. 23.06.1998. Heinz leg. (cSh); 1 ♂: C Elburz, Kandovan Pass, 36 ° 09 ′ 10 ″ N 51 ° 18 ′ 99 ″ E. 3000 m a. s. l. 31.05.2008. J. Weipert leg. (NME); 3 ♂♂, 1 ♀: C Elburz, Kandovan Tunnel, 2670 m, 36 ° 09 ′ 57 ″ N, 51 ° 19 ′ 18 ″ E, 31.05.2008, leg. A. Skale (NME, cSh); TEHRAN: 1 ♂: Elburz Mts., Kandovan Pass, N Tajrish, 36 ° 09 ′ 35.3 ″ N 51 ° 17 ′ 50.5 ″ E. 2850 m, brook bank, under stones, in gravel and alluvial soil. 10.06.2014. D. Wrase & Laser (cSch); 2 ♂♂, 3 ♀♀: after Kandovan tunnel, 36 ° 06 ′ 16.8 ″ N 51 ° 18 ′ 55.8 ″ E. 2575 m a. s. l. 24.07.2005. S. Serri & J. Frisch (cSh, MNHUB); 7 ♂♂, 5 ♀♀: Kandovan Val. 2545 m a. s. l. 10 – 11.08.1970. Exp. Nat. Mus. Praha (cSh, NMPC); 2 ♂♂, 3 ♀♀: C Elburz, Darband Sar Valley. 2500 – 3000 m a. s. l. 16.07.1970. Exp. Nat. Mus. Praha (NMPC); 4 ♂♂, 3 ♀♀: Kuhha-ye Tu-Chal. 2000 – 2500 m a. s. l. 18 – 19.07.1970. Exp. Nat. Mus. Praha (cSh, NMPC); 1 ♂: M. Elburs, Gadjereh. 2600 m a. s. l. 19.08.1966. V. Sbordoni leg. (cB); 1 ♂: Firouzkuh, Tange Vashi, Savashi waterfall, 35 ° 52 ′ 07.2 ″ N 52 ° 43 ′ 46.5 ″ E. 2250 m a. s. l. 28.06.2006. Alipanah leg. (MNHUB); 1 ♂: C Elburz, S Polur, 35 ° 47 ′ 50 ″ N 52 ° 02 ′ 90 ″ E. 2570 m a. s. l., riverbank. 29.05.2008. J. Weipert (NME); 2 ♂♂: C Elburz, Shemshak, Shemshak River. 2350 m a. s. l. 26.05.2008. A. Skale leg. (cSh, NME); SEMNAN: 1 ♂: Shahmirzad, Chashm vill., 35 ° 51 ′ 17.3 ″ N 53 ° 17 ′ 31.9 ″ E. 2045 m a. s. l. 22.05.2006. S. Serri & J. Frisch (MNHUB); KERMAN: 2 ♂♂: Qohrud Mts., 15 km E Korin. 3500 m a. s. l. 05.05.2008. A. Anichtchenko leg. (cSh); 5 ♂♂, 7 ♀♀: Küh-e Hezâr, above Âbshâr-e Râyen water fall, SW Râyen, 29 ° 32 ′ 49.6 ′ N 57 ° 17 ′ 56.5 ″ E. 05.06. 2014. 2920 m a. s. l., high plaine brook bank, in gravel detritus sifted. D. Wrase leg. (cSh, cSch, cZ, NME); OSTâN-E KOHGÎLűYE VA BűYER-AHMAD: 5 ♂♂, 5 ♀♀: Yasuj, Zagros Mts., Si Sakht Pass. 3800 m a. s. l. 04 – 05.06.2008. A. Anichtchenko’ (BMNH, cSh); 1 ♀: Yasuj, NW Shiraz, vill. Kakan, 30 ° 40 ′ N 51 ° 43 ′ E. 13.06.1999. E. & P. Hajdaj leg. (cSh).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777C7A61DD90FC886D5155DA.taxon	description	Redescription. Measurements (n = 100): HW: 0.86 – 1.21; HL: 0.61 – 0.81; OL: 0.22 – 0.30; TL: 0.12 – 0.20; AL (averaged): 3.42; PL: 0.85 – 1.34; PWmax: 1.14 – 1.56; PWmin: 0.73 – 1.18; ESL: 1.31 – 1.99; EW: 1.56 – 2.12; MTbL (averaged): 1.20; MTrL (averaged): 0.56 (MTrL 1 – 4: 0.29; MTrL 5: 0.27); AW: 1.63 – 2.02; AedL: 0.72 – 1.28; BL: 4.50 – 6.90 (holotype of G. zwickianus: 6.30; holotype of G. iranicus: 6.35). Habitus as in Figs. 3 – 6. Body yellow-brown to black (some specimens with darker head, yellow-brown or reddish (Fig. 5) pronotum, sometimes with paler middle portion of elytra and paratergites, and sometimes with indistinct reddish spot on each elytron (Fig. 6 )); mouthparts, antennae and legs yellow-brown to brown (apical segment of maxillary palpi usually paler, some specimens with yellowish antennomeres 1 – 2 or 1 – 3 and tarsi). Forebody without microsculpture, except for neck with very dense and coarse isodiametric sculpture, medioapical portion of pronotum in some specimens with traces of indistinct meshes and abdominal tergites with very dense transverse or isodiametric microreticulation. Head 1.4 times as broad as long, frontal portion with slightly or strongly convex supra-antennal elevations, with deep and wide subtriangular or rectangular anterio-median depression; temples 1.5 – 1.8 times as long as longitudinal length of eyes; interocellar depression moderately narrow and deep, separated from infraorbital portions by deep anteocellar foveae (indistinct in some specimens) reaching about level of middle or apical third of eyes. Distance between ocelli slightly shorter or distinctly longer than distance between ocellus and posterior margin of eye. Punctation irregular and fine, denser in middle, sparser and sometimes slightly larger on infraorbital portions. Antennomere 2 1.4 – 1.7 times as long as basal antennomere, 3 distinctly longer than 2, 4 – 5 or 4 – 6 slightly shorter than 3, 7 – 10 broader and slightly shorter or longer than 6. Pronotum 1.2 – 1.3 times as broad as long, from widest anterior third sharply narrowed toward moderately short subparallel laterobasal margins; middle portion without longitudinal depression, with shallow and sometimes indistinct or with moderately wide and deep transverse mediobasal depression; anterior margin slightly concave or rounded, distinctly narrower than straight posterior margin. Punctation dense, about as that or slightly larger and deeper than that on head, sparser and sometimes finer in medioapical (some specimens) or mediobasal portion (sometimes mediobasal portion in front of mediobasal depression without punctures). Elytra 1.4 – 1.5 times as long as pronotum, slightly or widely broadened posteriad. Punctation about as that on pronotum, but distinctly sparser and sometimes finer in middle or along suture and usually denser and coarser in parascutellar portion (some specimens with slightly larger and deeper punctation than that on pronotum). Metatibia about twice as long as metatarsus. Abdomen narrower than elytra, with distinct, wide and large, transverse or oval tomentose spots in middle of abdominal tergite IV. Male. Pronotum wide and convex. Profemuri and protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII straight, rounded or sligthly concave. Apical margin of abdominal sternite VIII concave. Median lobe moderately wide, gradually or strongly narrowed toward subacute apex; parameres significantly broadened toward apical portions, exceeding apex of median lobe (Figs. 16, 18, 23) or slightly shorter (Figs. 19 – 20, 22 – 24), with four moderately long apical setae; apical portion of median lobe with two narrow sclerotized structures; internal sac weakly sclerotized, with narrow and short or elongate flagellum (Figs. 16 – 24). Lateral portion of the aedeagus as in Figs. 17, 21. Female. Pronotum moderately narrow, slightly convex. Profemuri and protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VIII and sternite VIII straight or rounded. Comparative notes. Based on the general shape of the body and the aedeagus, G. (s. str.) zwickianus is similar to G. (s. str.) bodemeyeri, from which it can be distinguished by the slightly broader pronotum and the abscence of the sclerotized parallel structures in the middle of the median lobe.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777C7A61DD90FC886D5155DA.taxon	distribution	Distribution. Geodromicus (s. str.) zwickianus is distributed in Turkey and Iran (Fig. 25). Bionomics. Specimens were collected at elevations from 180 to 3800 m a. s. l. and were taken from under stones and gravel along river banks, and sometimes sifted from alluvial soils and gravel detritus. The locality of G. (s. str.) zwickianus in Qohrud Mts. as in Fig. 26.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777C7A61DD90FC886D5155DA.taxon	discussion	Remarks. Geodromicus (s. str.) zwickianus based on the holotype was originally described from “ S. O. Anatolien: Ost Taurus bei Sürgü … ”. Geodromicus iranicus based on eight specimens was originally described from “ Iran … dans le Massif de l`Elbrouz ”. During the study of the types and additional material from Turkey and Iran, I have not found sufficient morphological differences between types and other studied specimens. Thus, I synonymized G. iranicus with G. (s. str.) zwickianus. The studied specimens from Turkey are usually with slightly darker body. The studied specimens from Zagros Mts. have slightly shorter and narrower pronotum with slightly larger punctation than that in other studied specimens. Despite that, shapes of aedeagi are slightly vary and have similar internal structures (Figs. 16, 18 – 20, 22 – 24).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A63DD90FF796DAF5476.taxon	diagnosis	Diagnosis. Body medium-sized (4.25 – 7.90 mm); pronotum convex, moderately narrow or distinctly transverse, from widest anterior third gradually or sharply narrowed toward subparallel-sided short or elongate laterobasal portion; elytra slightly longer than broad (G. (s. str.) malcolmi) or distinctly broader than long, slightly or significantly broadened posteriad; aedeagus narrow or wide, with parameres slightly or significantly broadened anteriad, with weakly sclerotized internal sac and very narrow short or long flagellum.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A63DD90FF796DAF5476.taxon	discussion	Species included. G. (s. str.) convexicollis Luze, 1903, G. (s. str.) kashmirensis Cameron, 1930, G. (s. str.) lebedevianus Roubal, 1929, G. (s. str.) malcolmi sp. n., G. (s. str.) macrothorax Kashcheev, 1999, G. (s. str.) medvedevi Shavrin, 2009.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A63DD90FF796DAF5476.taxon	distribution	Distribution. The species of the bodemeyeri group are distributed in the Palaearctic Region, and known from the Central Asia and the Himalayan Region.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A63DD90FF796DAF5476.taxon	discussion	Bionomics. The species of the convexicollis group inhabit banks of rivers and streams, and can be found in gravel, under stones, mosses, drift, etc. Specimens were collected at elevations from 700 to 4200 m a. s. l. from April to October. Remarks. Previously, Bordoni (1984) included Geodromicus hauserianus (synonym of G. (s. str.) convexicollis) to the rousi group (Shavrin 2021). Based on the general shape of the apical portions of the parameres and weakly sclerotized internal sac with narrow flagellum, the convexicollis group is similar to the bodemeyeri group, from which it can be distinguished by the absence of paired sclerotized structures in apical and middle portions of the median lobe. Key to convexicollis group	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A78DD90F91F6AB85680.taxon	description	(Figs. 28 – 39)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A78DD90F91F6AB85680.taxon	materials_examined	Type material examined: Holotype of Geodromicus convexicollis Luze, 1903, ♂: ‘ globulicollis Zett. | Taschkent, | Akunin’ <handwritten>, ‘ convexicollis | Luze’ <handwritteb>, ‘ 205 ’ <printed>, ‘ c. Eppelsh. | Steind. d. ’ <printed>, ‘ Co- | Typus’ <pink, printed>, ‘ Holotypus | Geodromicus | convexicollis Luze, 1903 | Zerche 1991 ’ <printed> (NMW.) Holotype of Geodromicus (s. str.) hauserianus Bordoni, 1984, ♂ (Fig. 29; dissected): ‘ OST-BUCHARA | Tschitschantan | Nufswald F. Hauser 1898. ’ <printed>, ‘ HOLOTYPUS’ <red, printed>, ‘ Geodromicus (s. str.) | hauserianus n. sp. | Det. A. Bordoni 19 [printed] 83 | holotypus ♂ ’ <handwritten> (NMW). Paratypes: 2 ♀♀ (on one pin): ‘ OST-BUCHARA | Tschitschantan | Nufswald F. Hauser 1898. ’ <printed>, ‘ PARATYPUS’ <red, printed; 2 labels>, ‘ Geodromicus (s. str.) | hauserianus n. sp. | Det. A. Bordoni 19 [printed] 83 | paratypi ♀♀ ’ <handwritten> (NMW). Holotype of Geodromicus cylindricus Tronquet, 1981, ♂ (aedeagus and apical abdominal segment are glued on the same card with the specimen): ‘ Band I Amir 2600 m 14. VIII. [19] 75 ’ <handwritten>, ‘ Afg [h] an [istan]. G. M [eurgues]. et O. L [edoux]. ’ <handwritten>, ‘ Museum Paris Coll J. Jarrige’ <handwritten>, ‘ TYPE’ <red, printed>, ‘ Geodromicus | cylindricus n. sp’ <handwritten> (MHNH). Additional material examined. KAZAKHSTAN: EAST KAZAKSTAN REGION: 1 ♂, 2 ♀♀: Tyshkan tract, 6 km E Sarybel. 1200 m a. s. l. 12.05.1985. V. G. Shilenkov leg. (cSh, ZMUC); 4 ♀♀: Tyshkan-tau Mts., Tyshkan R., 10 km N Sarybel. 2000 m a. s. l .. 19.05.1995 (cZ); 1 ♂, 1 ♀: same data, 44 ° 30 N, 80 ° 04 E. 1800 m a. s. l. 28.07.2002. Müller-Motzfeld leg. (cA); 1 ♂, 1 ♀: Tarbagatay, below Pik Sarymobe, Kalymuvat. 1250 m a. s. l. 05.08.2001. Müller-Motzfeld leg. (cA); 3 ♂♂, 2 ♀♀: Tarbagatay, Bazar R. 22.08.1988. V. Kashcheev leg. (ZIN); 3 ♂♂, 6 ♀♀: same data. 21.08.1988 (cSh, ZIN); 7 ♀♀: Altai, Katon-Karaga, Sarymsakty R. 15 – 19.07.1989. V. Kashcheev leg. (ZIN); 2 ♂♂, 1 ♀: Altai, Tauchilik and Karakoba rivers. 11.08.1989. V. Kashcheev leg. (cSh); 4 ♂♂: same data. 16.08.1988 (BMNH, cSh, ZIN); 1 ♀: Chingistay [Shyngystay], 49 ° 11 ′ 33.0 ″ N 85 ° 57 ″ 06.8 ′ E′. 896 m a. s. l. V. Kashcheev leg. (cSh); 1 ♂, 1 ♀: Sorvenok R. near Aksubas Mt. 12 – 14.08.1989. V. Kashcheev leg. (ZIN); 1 ♂, 1 ♀: Sarym-Sakty Mts., Ozernaya R., valley. 2300 m a. s. l. 15 – 20.07.2007. A. Plutenko leg. (cSch); 1 ♂, 1 ♀: southern slope of Kurchum Mts., 10 km N Urunkhayka. 1500 m a. s. l., near water. 28.07.1988. I. I. Kabak leg. (ZIN); TURKISTAN: 1 ♂: 15 km E Kaskasu. Meadow-steppe zone, stream, in mosses. 12.07.1983. B. V. Iskakov leg. (ZIN); 1 ♂: 13 km SE Kaskasu. Subalpine zone, stream, in mosses. 11.07.1983. B. V. Iskakov leg. (ZIN); 1 ♂: Ugam Mts., Sary-Aygyr gorge, 42 ° 11 ′ 08 ″ N 70 ° 19 ′ 59 ″ E. 07.08.2012. S. V. Kolov leg. (cKh); 1 ♂: Aksu-Zhabagly Nature Reserve, Zhabagly R., Kshi-Kaindy. 16.06.2011. V. Kashcheev leg. (ZIN); 2 ♂♂, 1 ♀: same data, Koksay R. 02.07.2001 (cSh); 1 ex.: Bolshoy Baldabrek R. pass, above Darbal. 27.04.1967. E. L. Guryeva leg. (ZIN); JAMBYL: 22 ♂♂, 29 ♀♀: Kyrgyz Mnt., Merke R, Merke. 12.06.1988. V. Kashcheev leg. (cSh, ZIN); 3 ♂♂, 5 ♀♀: same data. 30.06.2011 (cSh); 2 ♂♂, 7 ♀♀: Bukhtarma R., Dzhambul. 10.08.1988. V. Kashcheev leg. (ZIN); 2 ♂♂: same R., near Archaty. 12.08.1988. V. Kashcheev leg. (cSh); 2 ♀♀: same R., Chingistay [Shyngystay]. 30.08.2010. V. Kashcheev leg. (ZIN); 4 ♂♂, 14 ♀♀: Chylik R., Sarybastau Pass. 15.06.1988. V. Kashcheev leg. (ZIN); 1 ♂, 1 ♀: Karatau Mt. Zhamantas R., 43 ° 52 ′ 05 ″ N 68 ° 11 ′ 55 ″ E. 731 m a. s. l. 10.05.2010. A. V. Ivanov leg. (cSh); 1 ♀: Karatau. 16.05.1974. S. Yu. Gryuntal leg. (ZMM); 1 ♀: Kungey Mnt., Kolbastau Pass. 27.07.1988. V. Kashcheev leg. (ZIN); JETISU: 20 ♂♂, 19 ♀♀: Dzhungar Alatau, Usek R. 10.07.2005. V. Kashcheev leg. (cSh, ZIN); 15 ♂♂, 35 ♀♀, 6 exs.: same data, Kesken-Terek R. 18 – 28.08.1988 (cSh, ZIN); 5 ♂♂, 2 ♀♀: same data, Kokjota Mnt., Topolevka. 27.06.2006 (ZIN); 4 ♂♂, 10 ♀♀: same data, Argynkaty R. 31.08.2006 (BMNH, cSh, ZIN); 6 ♂♂, 1 ♀: same data, Kapal R. 1620 m a. s. l. 02.08.1984. V. Kashcheev leg. (cSh, ZIN); 4 ♀♀: same data, Tentek R., Kokzhar. 25.08.2006 (BMNH, ZIN); 4 ♂♂, 1 ♀: same data, Burkhan R. 12.08.2004 (cSh); 1 ♂: same data, Oisaz-Chizhe R. 06.07.1990 (cSh); 1 ♀: same data, foothills of Berbokan Mts., environs of Uspenovka, arm of Orto-Tentek R. t = 14 ° C, pH = 5.4. 13.08.1991. A. Shatrovsky leg. (cA); ALMATY: 57 ♂♂, 66 ♀♀: Ketmen Mnt., Dolaity R. 09 – 15.07.1988. V. Kashcheev leg. (cSh, ZIN); 2 exs.: same Mnt., Bolshoy Ketmen. 16.06.1988. В. V. Kashcheev leg. (ZIN); 12 ♂♂, 4 ♀♀: same data. 18.07.1988 (cSh, ZIN); 4 ♂♂, 2 ♀♀: same data, Bolshoy Ketmen Pass. 20.07.2009 (cSh, ZIN); 1 ♂: S Ketmen Pass, 43 ° 20 ′ N, 80 ° 19 ′ E. 2750 m a. s. l. 31.07. Müller-Motzfeld leg. (cA); 7 ♂♂, 13 ♀♀: Ketmen Mnt., Bolshoy Kyrgisal. 10.06.1988. V. Kashcheev leg. (ZIN); 19 ♂♂, 13 ♀♀: same data. 13.07.2009 (ZIN); 4 ♂♂, 6 ♀♀: same Mnt., Shalkydysu R. 17.07.1988. V. Kashcheev (cSh, ZIN); 1 ♂: same Mnt., Avat R. 14.07.2008. V. Kashcheev leg. (ZIN); 6 ♂♂, 16 ♀♀: Ile Karatau, Zhynishke R. 20.06.1988. V. Kashcheev (cSh, ZIN); 13 ♂♂, 23 ♀♀: same data. 22.08.1988 (BMNH, cSh, ZIN); 1 ♀: Chilik R., Bartogay reservoir, 43 ° 19 ′ 21 ″ N 78 ° 29 ′ 57 ″ E. 06.09.2012. Kolov leg (cKh); 32 ♂♂, 46 ♀♀: Kyrgyzsay Pass, 43 ° 18 ′ 40.3 ′ N′ 079 ° 32 ′ 26.9 ′ E. 12.07.2009. V. Kashcheev leg. (ZIN, cSh); 1 ♂, 18 exs.: Kungey, Chilik R., Sarybastau Pass. 12 – 15.06.1988. V. Kashcheev leg. (ZIN); 1 ♂: Ile Alatau, Batan R., Chin-Turgen. 25.06.1988. V. Kashcheev leg. (ZIN); 2 ♀♀: same data, Big Almaty Lake. 28.06.1991. V. Kashcheev leg. (ZIN); 11 exs.: Zhinishke, Zhinishke R. 17.06.1988. V. Kashcheev leg. (ZIN); 2 exs.: same data, Koksay Pass. 20.06.1988 (ZIN); 4 exs.: same data, Assy Pass. 24.06.1988 (ZIN); 1 ♀: Kungey Alatau, near Alma-Ata, 77 ° 00 ′ N 43 ° 10 ′ E. 2500 – 3500 m a. s. l. 07.1990. J. Kolibáč leg. (NHMB); 1 ♀: SW environs of Saryzhaz, 89 ° 42 ′ 48 ″ N 54 ° 79 ′ 30 ″ E. 2000 – 2300 m a. s. l. D. Král leg. (NHMB); 2 exs.: Burakozhyr. 31.08.1988. V. Kashcheev leg. (ZIN); 1 ex.: 73 км S Glinkovo, Boralday R. Near water. 24.05.1986. B. V. Iskakov leg. (ZIN); UZBEKISTAN: NAVOI: 3 exs.: ‘ T [u] rk [e] st [an]. Mnt. Nurata Uchum Glazunow 1892 ’ (ZIN); TASHKENT: 2 ♂♂, 1 ♀: ‘ Ouzbekistan 8 – 68 Mts Tschimgan 1500 m H. C. ’ (MHNH); 1 ♀: Bolshoy Chimgan. 05.07.1981. K. Majer leg. (MNHUB); 1 ex.: Chimgan, 12 Klyuchey. 20.07.1924. Yu. Martynova leg. (ZIN); 11 exs.: Chimgan. 30.07.1920. I. Ivanov leg. (ZIN); 1 ex.: Uganskiy Mts., 20 km N Khumsan. 16.05.1966. V. A. Zaslavskiy leg. (ZIN); 1 ex.: same data, valley of Ugan River. 17.05.1963. E. L. Guryeva leg. (ZIN); 2 exs.: Tashkent. 17.07.1925. Petrova leg. (ZIN); 1 ♂: ‘ Zangi Bazar Süd- Turkestan K. Küchler S. G. 8. V. 1913 ’, ‘ Bodemeyeri Bernh. ’, ‘ Bernhauer determ. ’, ‘ Geodromicus Bodemeyeri cotyp. Bernh. ’, ‘ Frankfurt / Main Senckenberg- Museum’ (SF); 1 ex.: Angren R. 23.09.1963. O. L. Kryzhanovsky (ZIN); 5 ♂♂, 11 ♀♀: Chatkal Mt. range, Okhangaron District, 6 km NW Irtash, Iertash [Yertoshoy] River (right confluent of Okhangaron River), 41 ° 10 ′ 40.7 ″ N 70 ° 16 ′ 17.9 ″. 2400 m a. s. l. 05 – 06.08.2021. A. V. Shavrin leg. (BMNH, cSh); 3 ♂♂, 1 ♀: same data, 41 ° 10 ′ 46 ″ N 70 ° 17 ′ 28.7 ″ E. 1900 m a. s. l. 03.08.2021 (cSh); 1 ♂: same data, Aksay R., snow spot. 41 ° 30 ′ 46.0 ″ N 70 ° 02 ′ 32.9 ″ E. 2200 m a. s. l .. 09 – 10.08.2021. A. Anicthchenko leg. (cSh); 2 ♂♂, 1 ♀: same data, 41 ° 10 ′ 24.3 ″ N 70 ° 16 ′ 06.5 ″. 2500 m a. s. l. 05 – 06.08.2021. A. Shavrin leg. (cSh); 1 ♂: Sublikay. 24.07.2017. Yu. G. Arzanov leg. (cSh); FERGANA: 1 ♀: Pamir Alai, Chamsaabad. 1800 m a. s. l. 14.07.1984. D. Wrase leg. (cSch); SYRDARYA: 1 ex.: Chimchanka R. 22.07.1924. O. Martynova leg. (ZIN); SAMARKAND: 2 ♀♀: Aman Kutan, environs of Samarkand. 1200 m a. s. l. 16 – 18.05.1974. A. Pfeffer leg. (NMPC); KASHKADARVA: 1 ♂: Yakkabagskiy District, Kalta-Kol. 07.09.1993. 1400 m a. s. l. K. Kolesnichenko leg. (cSh); KYRGYZSTAN: TALAS: 1 ♀: Kyrgyzskiy Mts., Kegety Valley. 2800 m a. s. l. 03.06.1994. W. Dolin leg. (cSch); 1 ♂: ‘ Talass-Tal. Fischer’ (SNMB); CHUI: 1 ♀: Kyrgyzskiy Mts., 5 km S Bishkek Orto-San. 01.10.1992. S. V. Ovchinnikov leg. (cSh); 3 ♂♂, 2 ♀♀: Sosnovka, Kara-Balta R. Stones near river, Semenovia transiliensis. 18.05.1943. K. V. Arnoldi leg. (cSh, ZMM); 1 ♂: Kungey- Alatoo Mts., upper course of Tschon-Kemin R. ca. 42 ° 47 ′ N 76 ° 19 ′ E. 2200 – 2500 m a. s. l. 08.1999. Hetzel leg. (cF); 1 ♀: Kara-Balta. 27.05.1995. S. Ovchinnikov leg. (cSh); YSSIK-KOL: 3 ♂♂, 4 ♀♀: Terskey-Alatoo Mts., valley of Chon-Kyzyl-Suu R., 2650 m a. s. l., under stones near water. 01.07.1988. V. Yanushev leg. (cSh, ZMM); 1 ex.: same Mts. and R., 2500 – 2700 m a. s. l. 02.07.1959. Zaslavskiy leg. (ZIN); 1 ♂, 1 ♀: Terskey-Alatau, Ottuk valley, 42 ° 21 N 79 ° 03 E. 3000 m a. s. l. 25.07.2000. Müller-Motzfeld leg. (cA); 1 ♂: Chichkan Valley near pass, 42 ° 11 ′ N, 72 ° 58 ′ E. 2370 m a. s. l. 31.07.1999. Müller-Motzfeld leg. (cA); 3 ♂♂, 1 ♀: ‘ Turkestan Issyk-Kul V. M. Duchor’ (MZHF); NARYN: 2 ♂♂, 4 ♀♀: Inner Tian Shan, 8 – 14 km NE Son-Kul Lake, ca. 41 ° 52 ′ N 75 ° 21 ′ E. 3100 – 3600 m a. s. l. 08.1999. Hetzel leg. (cF); 1 ♀: Ala Arteche. 2000 m a. s. l. 09.07.1984. D. Wrase leg. (cSch); 1 ♂, 1 ♀: At-Bashi. 22 – 24.08.1995. S. Saluk leg. (cSh); 1 ♂, 3 ♀♀: Son-Köl region, 41 ° 43 ′ 53 N, 75 ° 05 ′ 03 E. 3000 m a. s. l. 14.07.2000. Müller-Motzfeld leg. (cA); JALAL-ABAD: 3 exs.: Fergana mt. Range, Arstanbab. 28.05.1961. L. Medvedev leg. (ZIN); 2 exs.: same data. 2600 m. 27 – 28.05.1961. E. L. Guryeva leg. (ZIN); 1 ♂, 2 ♀♀: Chatkal Mts., Arkit. 1400 m a. s. l. 14.09.1945. K. Arnoldi leg. (ZMM); OSH: 1 ♂, 1 ♀: West Tian Shan, Yarodar. Stream, litter and under stones. 24 – 25.06.1983. K. Eskov leg. (ZMM); 2 ♂♂, 3 ♀♀: Lailak Dist., Sarken-Gush rivers. 2100 m a. s. l. 9.6.1996. A. Putschkov leg. (cSch, cSh); 1 ♀: Zaalayskiy Mts., Bardabo outpost. 3200 – 3400 m a. s. l. 19 – 20.07.1996. V. A. Mikhailov leg. (cA); 1 ♂, 1 ♀: Alai Valley, Kelte-Bulak R. near Unkur. 3100 m a. s. l. 22.07.1999. Müller-Motzfeld leg. (cA); 2 ♂♂, 1 ♀: Alai, White Mts. near Kara Kintyk, 39 ° 42 ′ N, 73 ° 27 ′ E. 3200 m a. s. l. 21.07,1999. Schubert leg. (cA); 1 ♀: Alai valley, Belye Gory, Kysyl R. 21.07.1999. Müller-Motzfeld leg. (cA); 1 ex.: northern slopes of Alai Mts. Under stones. 01.08.1938. M. Chirkun leg. (ZIN); 2 exs.: ‘ Alai. 4. IV. 1899 ’ (ZIN); TAJIKISTAN: SUGHD: 6 ♂♂, 3 ♀♀: ‘ Seravschan Dschai-See’ (NMW, MHNH); 3 exs.: ‘ Sevarschan Lac. Dschai Glasunow 1892 ’ (ZIN); 1 ♀: ‘ Turkestan Dshai-See Rutschin Pass Glasunov’ (BMNH); 1 ♀: ‘ Seravshan Turkestan’ (MHNH); 1 ♂: ‘ Seravschan Turkestan’, ‘ asiaticus Solsk. ’ (IRSNB); 1 ♂: ‘ Turkestan Seravschan’ (BMNH); 1 ♀: ‘ Turkestan Seravschan Kulikulan L. ’ (NMPC); 1 ♀: ‘ Turkestan Lac. Kulikulan Glasunov 1892 ’ (NMPC); 2 ♀♀: ‘ Seravschan Kulikulan S’, ‘ Turkestan’ (MZHF, NMPC); 2 ♂♂, 1 ♀: ‘ Seravshan Kuli Kulansee’, ‘ im 250 km de Tashkent’ (MHNH); 1 ♂, 2 ♀♀, 4 exs.: ‘ Turkestan Lac. Kulikulan Glasunov 1892 ’ (NMW, ZIN); 2 ♂, 1 ♀: Zeravshan Mts., environs of Kulikulan. 25.08.1940 (ZMUC); 1 ♂: ‘ Co-Type’, ‘ ♂ ’, ‘ Seravschan Kulikulan — S, Turkestan, convexicollis’, ‘ ex coll. Luze’, ‘ COTYPUS Geodromicus convexicollis Luze. ’, ‘ M. Cameron Bequest. B. M. 1955 - 147. ’ (BMNH); 1 ♀: ‘ ♀ ’, ‘ Seravchan Kulikulan. S’, ‘ Turkestan’, ‘ Glasunow’, ‘ convexicollis’, ‘ COTYPUS Geodromicus convexicollis G. Luze’, ‘ Geodromicus convexicollis Luze’ (NMPC); 1 ♀: ‘ ♀ ’, ‘ TURKESTAN LAC. KULIKULAN Glasunov 1982 ’, ‘ COTYPUS Geodromicus convexicollis Luze’, ‘ Geodromicus convexicollis 3273 Luze’, ‘ Collection Arthur Spälti’ (MHNG); 1 ♀: ‘ Turkestan Lac. Kulikulan Glasunov 1892 ’, ‘ COTYPUS Geodromicus convexicollis Luze’, ‘ Sammlung Dr. J. B. Jörger Masans-Chur 1957 ’ (NHMB); 2 exs.: ‘ Seravschan Boschara Glasunow 1892 ’ (ZIN); 6 ♂♂, 3 ♀♀: Seravshan, Kette Masor. 2300 m a. s. l. 20.05.1984. Mikhailov leg. (NHMB); 1 ♂, 1 ♀: Seravshan Mts., Zavron Valley. 2100 – 3000 m a. s. l. 12 – 13.07.1990. M. Schulke & D. Wrase leg. (cSch); 1 ♂: same Mts., north slope. Marguzorskiye lakes. 24.05.1984. V. Mikhailov leg. (cSch); 1 ♀: same Mts., north slope, gorge, near Mazor. 02.06.1984. V. Mikhailov leg. (cSch) 2 exs.: Iskanderdarya near sources. 12.08.1947. Kirichenko leg. (ZIN); 1 ♂: Syr-Darja, Kanibadam. 22.04. (ZMM); 1 ♂: ‘ OST-BUCHARA Tschitschantan [Vorukh]. Coll. Hauser 1898 ’ (FMNH); 1 ♂: Turkesrtan Mts., north slope of Shakhristan Pass. 28.06.1983. S. Alekseev leg. (cR); NOHIYAHOI TBEI JUMHURÎ: 1 ♂: Gissar Mts., Kondara. 1200 – 1300 m a. s. l. 20 – 23.05.1988. V. G. Shilenkov leg. (ZMUC); 1 ex.: same data. 14.09.1937. Gussakovskiy leg. (ZIN); 2 ♂♂: same Mts., Ziddi. 2400 – 2700 m a. s. l. 19.06.1982. Jelinek leg. (cZ, NMPC); 1 ♂, 1 ♀: Hissar-Alai, Adjuk-Cleft nr. Warsob. 01 – 03.07.1990. M. Schülke leg. (cSch); 2 ♀♀: southern slope of Petr I Mts., Sangvor Reserve, Lyuli-Kharvi. 23.08.1975 (cSh); KHATLON: 1 ♀: ‘ Ost-Buchara Kuljab’ (NMW); VILOYATI MUKHTORI KűHISTONI: 1 ♂: Vanchskiy District, valley of Banch R. near environs of Banchskiy forestry. 17.05.1982. V. Mikhailov leg. (cSch); 1 ♀: Stchugnanskiy Mts., environs of botanical garden. 2200 – 2300 m a. s. l .. 05.05.1978. V. Mikhailov leg. (cSch) 1 ex.: Dalvaz, Kala-i-Khum. 1820 m a. s. l .. 05 – 12.07.1989. Grabichevskiy leg. (ZIN); AFGHANISTAN: BADAKSHAN: 1 ♂, 1 ♀: Badakshan Mts., Shiwa Lake. 2800 m a. s. l. 20.07.2008. C. Reitter leg. (cF); LOCALITY NOT SPECIFIED: 1 ♀: ‘ Ala-Tau. Syr. D [arya]. 6.08. ’ ‘ A. LEBEDEV’, ‘ bodemeyeri Bh. det. Bernh. ’, ‘ planicollis Brnh. Typus unic. ’, ‘ Chicago NHMus M. Bernhauer Collection’ (FMNH); 1 ex.: ‘ Alatau Geb. E. Fischer’ (ZIN).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A78DD90F91F6AB85680.taxon	description	Redescription. Measurements (n = 500): HW: 0.65 – 0.93; HL: 0.55 – 0.67; OL: 0.27 – 0.30; TL: 0.10 – 0.20; AL: 3.13 – 3.55; PL: 0.85 – 1.18; PWmax: 1.04 – 1.68; PWmin: 0.73 – 0.93; ESL: 1.49 – 1.98; EW: 1.63 – 2.26; MTbL (averaged): 1.30; MTrL (averaged): 0.62 (MTrL 1 – 4: 0.35; MTrL 5: 0.27); AW: 1.49 – 2.20; AedL: 0.97 – 1.30; BL: 4.30 – 7.20 (holotype of G. convexicollis: 5.00; holotype of G. hauserianus: 6.60; holotype of G. cylindricus: 6.00). Habitus as in Fig. 28 – 30. Body brown to reddish-brown or black, sometimes with paler (yellowish or reddish) elytra and paratergites of abdomen; mouthparts, antennae and legs yellow-brown to brown (antennae in some specimens yellowish, or with paler antennomeres 1 – 3, femori (except apical portions) and tarsi). Head with traces of shallow transverse meshes in apical portion and / or isodiametric or subdiagonal meshes on infraorbital portions; neck with dense and coarse isodiametric micropunctation; medioapical portion of pronotum in some specimens with very indistinct transverse microreticulation; abdomen with very dense transverse or isodiametric microsculpture. Head 1.1 – 1.3 times as broad as long, with slightly or strongly convex supra-antennal elevations, with moderately deep and wide anterio-median depression; temples moderately convex, 1.5 times to slightly more than twice as long as longitudinal length of eyes, gradually rounded or sometimes with subacute angle between posterior margin of eye and distinct postocular ridge in some specimens; interocellar depression subrectangular or subtrapezoidal, large and deep, separated from infraorbital portions by moderately deep and long foveae, reaching level of middle length or apical third of eyes. Ocelli small or moderately large, distance between ocelli about as long as or slightly longer than distance between ocellus and posterior margin of eye. Punctation variable, fine or moderately large and deep, sparse or dense, usually denser and deeper in middle and sometimes finer and sparser on infraorbital portion. Antennomere 3 slightly longer and narrower than antennomere 2, 4 – 5 slightly narrower and shorter than 3, 6 – 8 slightly longer than 5, 9 – 10 slightly longer than 8, apical antennomre 1.3 – 1.7 times as long as 10. Pronotum transverse, 1.4 – 1.8 times as broad as long, from widest middle or anterior third sharply narrowed toward short or moderately long subparallel mediolateral portions; middle portion without longitudinal depression, with transverse, shallow or moderately deep mediobasal depression. Punctation variable. moderately sparse, about as that or slightly larger than that on middle portion of head, or distinctly denser, larger and deeper, usually sparser in / or middle and mediobasal portions (some specimens without punctures in front of mediobasal depression). Elytra slightly broader than long and slightly or significantly broadened posteriad, 1.6 – 1.7 times as long as pronotum. Punctation variable, about as that on pronotum or distinctly denser, larger and deeper, usually denser and coarser in parascutellar portion, and finer and sparser along suture. Metatibia about twice as long as metatarsus. Abdomen distincly narrower than elytra, with two large and transverse tomentose spots in middle of abdominal tergite IV (sometimes indistinct in some specimens). Male. Pronotum wide, usually strongly convex. Profemuri and protarsomeres 1 – 4 distinctly widened. Apical margin of abdominal tergite VIII straight, rounded or slightly concave. Apical margin of abdominal sternite VIII concave. Median lobe very wide, from apical third strongly narrowed toward rounded or subacute apex; parameres slightly broadened apicad, reaching or exceeding apex of median lobe, with four short apical setae; internal sac weakly sclerotized, without visible structures except narrow and moderately short flagellum in middle or basal portion (Figs. 31, 33 – 36). Lateral aspect of the aedeagus as in Fig. 32. Female. Pronotum moderately small, somewhat narrow, less convex. Profemuri and protarsomeres 1 – 4 narrow. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded. Comparative notes. Based on the coloration, body length and the general shape of the body, G. (s. str.) convexicollis is similar to Central Asian G. (s. str.) kashmirensis, from which it can be distinguished by slightly more transverse pronotum and moderately narrow apical portions of the parameres. From all species of the convexicollis group it differs by the very wide median lobe.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A78DD90F91F6AB85680.taxon	distribution	Distribution. Geodromicus (s. str.) convexicollis is widely distributed in the Middle Asia, and known from Kazakhstan, Uzbekistan, Kyrgyzstan, Tajikistan and north-eastern Afghanistan (Fig. 37). Bionomics. Specimens were collected at elevations from 700 to 3600 m a. s. l. and were taken from under stones near streams and rivers, or were sifted from the gravel, soil, drift and mosses. Some specimens were collected from under stones near snow spots. It was collected together with specimens of G. (s. str.) macrothorax in several localities of south-eastern Kasakhstan (Saryhastau Gorge, Kokjota Mnt., and Dzhungar Alatau). The localities in Chatkal Mts. (Uzbekistan) as in Figs. 38 – 39.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253777B7A78DD90F91F6AB85680.taxon	discussion	Remarks. Geodromicus convexicollis Luze, 1903 was originally described based on one specimen (“ Nach einem männlichen Exemplare beschrieben. ”) from Tashkent, Uzbekistan. Zerche (1992) studied the holotype and redescribed the species. Bordoni (1984) based on the holotype and three paratypes described G. hauserianus from “ Ost. Buchara, Tschiftschantan … ”. It was synonymized by Zerche (1991) with G. (s. str.) convexicollis without the study of the type specimens. I studied the type specimens in NMW (see above) and confirmed this synonymization here. Geodromicus cylindricus Tronquet, 1981 was originally described from “ Afghanistan: Baudi-I-Amir ”. I studied the holotype and found that this specimen is conspecific with other specimens of G. (s. str.) convexicollis. Sketchy aedeagus provided by Tronquet (1981) is also similar to other studied specimens of G. convexicollis. Thus, I synonymized it with G. (s. str.) convexicollis.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377607A7ADD90FB0B68C355CA.taxon	description	(Figs. 40 – 43, 46 – 51)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377607A7ADD90FB0B68C355CA.taxon	materials_examined	Type material examined. Lectotype (here designated) of Geodromicus kashmirensis Cameron, 1930, ♂ (Fig. 20; dissected): ‘ Type’ <round label with red margin, printed>, ‘ Ferozepur Nala 6500 | Gulmarg, Kashmir. | B. M. Bhatia. | 2. [handwritten] VI. 1928. ’ <printed>, ‘ Under | wet stones’ <printed>, ‘ 311 ’ <handwritten>, ‘ M. Cameron | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ G. | kashmiricus | TYPE Cam’ <handwritten>, ‘ LECTOTYPE | Geodromicus | kashmirensis Cameron, 1930 | Shavrin A. V. des. 2022 ’ <red, printed>, ‘ Geodromicus (s. str.) | kashmirensis Cameron, 1930 | Shavrin A. V. det. 2022 ’ <printed> (BMNH). Lectotype (here designated) of Geodromicus similis Cameron, 1930, ♀ (Fig. 41; right antennomeres 9 – 11 glued on the same card with the specimen): ‘ Type’ <round label with red margin, printed>, ‘ Ferozepur Nala 6500 | Gulmarg, Kashmir. B. M. Bhatia. | 2. [handwritten, in black] VI. 1928. ’ <printed>, ‘ Under | wet stones’ <printed>, ‘ 313 <handwritten>, ‘ Geodromicus | similis | TYPE [in red, underlined] Cam. ’ <handwritten>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ LECTOTYPE | Geodromicus | similis Cameron, 1930 | Shavrin A. V. des. 2022 ’ <red, printed>, ‘ Geodromicus (s. str.) | kashmirensis Cameron, 1930 | Shavrin A. V. det. 2022 ’ <printed> (BMNH). Holotype of Geodromicus consors Cameron, 1930, ♀ (Fig. 42): ‘ Type’ <round label with red margin, printed>, ‘ Kashmir | Gulmarg | vi-vii- 31 | Dr. Cameron’ <printed>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ G. | consors | TYPE [red, underlined] Cam. ’ <handwritten>, Geodromicus (s. str.) | kashmirensis Cameron, 1930 | Shavrin A. V. det. 2022 ’ <printed> (BMNH). Holotype of Geodromicus (Geodromicus s. str.) afghanicus Scheerpeltz, 1961, ♀: ‘ Panjao 24 [handwritten] / 7 [handwritten] 1948 | Afghanistan N. Haarløv | St. 85. [handwritten] ’ <printed>, ‘ 3. Danske Exp. III | Centralasien’ <printed>, ‘ ♀ ’ <handwritten>, ‘ TYPUS | Geodromicus [handwritten] | afghanicus [handwritten] | O. Scheerpeltz. ’ <red, printed>, ‘ Geodromicus | afghanicus | nov. spec. | det. Scheerpeltz, 19 [printed] 58 ’ <handwritten>, ‘ ZMUC | 00523323 ’ <printed>, ‘ Holotypus ♀ | Geodromicus | afghanicus | Scheerpeltz | V. Gusarov rev. 1993 ’ <red, handwritten>, ‘ Geodromicus ♀ | afghanicus Scheerp. | V. I. Gusarov det. 1993 [printed] ’ <handwritten> (ZMUC). Holotype of Geodromicus karakorus Coiffait, 1977, ♂ (Fig. 43; dissected): ‘ Nangmah-Tal | Karakorum, 1.8. [19] 70 | lg. Koblmüller’ <handwritten>, ‘ Geodromicus | spec. | det. H. Franz [printed] ’ <handwritten>, ‘ TYPE’ <red, printed>, ‘ Geodromicus | karakorus | H. COIFFAIT det. 19 [printed] 76 ’ <handwritten>, ‘ Geodromicus | kashmirensis Cameron, 1930 | Shavrin A. V. det. 2022 ’ <printed> (NMW). Additional material examined. AFGHANISTAN: 1 ♂: ‘ Kot Gai 2500 m 21. VIII. [19] 74 ’, ‘ Palktia Afghan [istan]. ’, ‘ Ft. de C ... [unreadable] G. M [eurgues]. et G. L [edoux] ’, ‘ MUSEUM PARIS | COLL. J. JARRIGE’, ‘ Geodromicus afghanensis Coiffait’ (MHNH); 1 ♀: ‘ St. 280 ’, ‘ Voyage en Afghanistan K. Lindberg’, ‘ Bend-Amir 1.8. [19] 57 ’, ‘ … [unreadable] Steien. ’ ‘ Geodromicus afghanicus Scheerp. ’, ‘ coll. Scheerpeltz’ (NHMW); 1 ♂: ‘ M. Daniel lgt by a trap on a lorry’, ‘ Kabul — 12. VI. Tangi Gar [19] 67. AFGHANISTAN’ (NMPC); 1 ♂: ‘ COTYPUS Geodromicus afghanicus O. Scheerpeltz’, ‘ ♂ ’, ‘ J. Klapperich Pagmangebirge 2300 m, 30.5. [19] 52 O. Afghanistan’, ‘ MUSEUM KOENIG BONN’, ‘ Geodromicus afghanicus H. COIFFAIT det. 1981 ’ (MHNH); 1 ♀: ‘ ♀ ’, ‘ Afghanistan Badagsthan’, ‘ COTYPUS Geodromcius afghanicus O. Scheerpeltz’, ‘ Geodromicus afghanicus | 3278 B n. sp. ’, ‘ Collection Arthur Spälti’ (MHNG); 1 ♂ [additional printed label: ‘ m’], 1 [additional printed label: ‘ f’]: ‘ Paratypus’, ‘ COTYPUS Geodromicus afghanicus O. Scheerpeltz’, ‘ J. Klapperich Pagmangebirge [Pagman] 2400 m, 6. 7. [19] 52 O. Afghanistan’, ‘ H. Coiffait det. 1981 ’, ‘ MUSEUM KOENIG BONN’, ‘ Geodromicus afghanicus Scheerp., 1960 V. I. Gusarov det 1993 ’ (ZFMK); PAKISTAN: 1 ♂, 1 ♀: Khyber Pakhtunkhwa, Great Karakorum Mts., to west from Chikar. 3200 – 3500 m a. s. l .. 01 – 10.08.2009. V. Gurko leg. (cSch); 1 ♂: Gilgit-Baltistan, Hunza-Nagar District, W Chikar. 3200 – 3500 m a. s. l. 01 – 10.08.2009. V. Gurko leg. (cSh); INDIA: KASHMIR: 13 ♂♂, 24 ♀♀, 1 spec.: ‘ Kashmir Gulmarg vi-vii- 31 Dr. Cameron’, ‘ M. Cameron. Bequest. B. M. 1955 - 147. ’ (BMNH); 3 ♀♀ (one specimen with additional label: ‘ consors Cam’): ‘ Paratype’, ‘ Kashmir Gulmarg vi-vii- 31 Dr. Cameron’, ‘ M. Cameron. Bequest. B. M. 1955 - 147. ’ (BMNH); 2 ♂♂, 2 ♀♀: ‘ Gulmarg Kashmir 6 / [19] 31. Cameron’, ‘ similis Cam. auct. det. ’, ‘ W. Steel coll. B. M. 1969 - 552 ’; 1 ♂, 4 ♀♀: ‘ Kashmir Gulmarg 8,000 – 9,000 ft. vi-vii- 1931 Dr. Cameron’, ‘ W. Steel coll. B. M. 1969 - 552 ’ (BMNH); 2 ♀♀: ‘ Kashmir Khelanmarg 10,000 ft. vi-vii- 1931 Dr. Cameron’ (BMNH); 1 ♀: ‘ Kashmir Lianmarg 10,000 ft. vi-vii- 1931 Dr. Cameron’ (BMNH); 2 ♂♂, 1 ♀, 3 exs.: Tangmarg Pir Panjal. 2600 m a. s. l. 21 – 25.05.1976. Martens & Schawaller leg. (MHNG, MHNH); 2 ♂♂: ‘ Kashmir’ (BMNH); LADAKH: 1 ♂, 1 ♀: Chemre. 06.1995. Sauer leg. (cSch); 3 ♂♂ (specimens from FMNH with additional label: ‘ amnicola Cam. det. Bernhauer Standinger don. ’), 4 ♀♀: ‘ Gya-Ladak Taglang-Pass Himalaya mont. ’ (FMNH, NMPC, SNSD).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377607A7ADD90FB0B68C355CA.taxon	description	Redescription. Measurements (n = 50): HW: 0.90 – 1.18; HL: 0.50 – 0.79; OL: 0.26 – 0.30; TL: 0.10 – 0.15; AL (averaged): 3.42; PL: 0.80 – 1.45; PWmax: 1.07 – 1.60; PWmin: 0.72 – 1.10; ESL: 1.71 – 2.25; EW: 1.82 – 2.25; MTbL (averaged): 1.55; MTrL (averaged): 0.60 (MTrL 1 – 4: 0.40; MTrL 5: 0.20); AW: 1.70 – 2.10; AedL: 1.05 – 2.05; BL: 4.25 – 7.20 (holotype of G. kashmirensis: 7.00; holotype of G. karakorus: 5.20; holotype of G. consors: 6.80; lectotype of G. similis: 5.00). Habitus as in Figs. 40 – 43. Body reddish-brown to dark-brown, sometimes with darker head, paler middle portion of elytra and yellowish intersegmental membranes of abdomen and / or paratergites in some specimens; mouthparts, antennae and legs yellow-brown to brown (some specimens with darker antennomeres 1 – 2 or 1 – 3 and tibiae). Head without or with distinct shallow and transverse microreticulation in middle; neck with very dense rugose and isodiametric microsculpture; pronotum without or with very indistinct transverse meshes in medioapical portion. Head 1.4 – 1.8 times as broad as long, with slightly or distinctly elevated infraorbital portions, frontal portion with relatively strongly convex supra-antennal elevations, with very deep, wide and subtriangular anterio-median depression, temples wide and convex, or distinctly narrowed posteriad behind posterior margins of eyes, about twice as long as longitudinal length of eyes; interocellar depression variable, deep and wide, or slightly impressed and narrow, separated from infraorbital portions by moderately deep, long anteocellar foveae (indistinct in some specimens), reaching level of middle or apical third of eyes. Ocelli small or moderately large, distance between ocelli 1.3 – 1.7 times as long as distance between ocellus and posterior margin of eye. Punctation irregular, moderately sparse and fine: denser, slightly larger and deeper in middle and infraorbital portion, or finer and sparser in middle and larger and denser on infraorbital portions. Antenna reaching middle length or apical third of elytra when reclined; antennomere 3 1.3 – 1.4 times as long as 2, antennomere 4 slightly broader and distinctly shorter than 3, antennomeres 5 – 10 indistinctly longer than 4 (sometimes antennomeres 8 – 10 slightly broader than 7). Pronotum 1.3 – 1.4 times as broad as long, from widest middle or medio-apical third strongly narrowed toward moderately long subparallel laterobasal margins, with shallow or moderately deep transverse mediobasal depression. Punctation dense, larger and deeper than that in middle portion of head, sometimes finer and sparser in mediobasal portion (some specimens with small oval impunctate portion in front of mediobasal depression). Elytra 1.4 times to twice as broad as long, slightly or widely broadened posteriad, 1.5 times to twice as long as pronotum. Punctation dense, about as that on pronotum or slightly larger and deeper, denser and finer in parascutellar portion and sometimes sparser and finer along suture. Metatibia more than twice as long as metatarsus. Abdomen distinctly narrower than elytra, with two very large and transverse tomentose spots in middle of abdominal tergite IV. Male. Pronotum large, very wide and convex. Profemuri and protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII straight or slightly concave. Apical margin of abdominal sternite VIII concave. Aedagus with moderately wide basal part, gradually narrowed toward rounded or subacute apex; aedeagus widely broadened apicad, reaching or slightly exceeding apex of median lobe, with four moderately short apical setae; internal sac with very narrow and moderately short flagellum in basal or middle portion (Figs. 46 – 49). Lateral aspect of the aedeagus as in Fig. 50. Female. Pronotum small, moderately narrow, less convex. Profemuri and protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VII and sternite VIII rounded. Comparative notes. Based on the coloration, body length and the general shape of the body, G. (s. str.) kashmirensis is similar to G. (s. str.) convexicollis, from which it can be distinguished by the less transverse pronotum, distinctly narrower median lobe and broader apical portions of the parameres.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377607A7ADD90FB0B68C355CA.taxon	distribution	Distribution. Geodromicus (s. str.) kashmirensis is known from several localities of Afghanistan, Pakistan and India (Ladakh, Kashmir) (Fig. 51). Bionomics. Specimens were collected at elevations from 1900 to 3500 m a. s. l. and were taken from under stones near streams and rivers.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377607A7ADD90FB0B68C355CA.taxon	discussion	Remarks. Geodromicus kashmirensis was originally described based on unspecified number of syntypes from “ Gulmarg, Ferozepur Nala, alt. 6500 feet … ”. I designate here the studied specimen from BMNH as the lectotype in order to fix the ideltity of the name. Geodromicus similis was originally described based on unspecified number of syntypes from the same locality as that in G. kashmirensis. I designate here this specimen (female) as the lectotype in order to fix the identity of the name. Based on the morphological features, it conspecific with the lectotype of G. kashmirensis and other studied specimens of this species. Thus, I synonymized G. similis with G. (s. str.) kashmirensis. Geodromicus consors was originally described based on the holotype (“ Type in my collection ”) from “ Kashmir: Gulmarg ”. I studied this type and found that it is conspecific with the lectotype of G. kashmirensis and other studied specimens of G. (s. str.) kashmirensis. Consequently, G. consors is synonymized with it. Geodromicus (Geodromicus s. str.) afghanicus was described based on the female from “ Panjao in Afghanistan ... ”. Tronquet (1981) cited it as “ G. afghanensis Coiffait (sous presse) ” from “ Khot Gai, massif du Tarte Soleiman, au Sud de Djalalabad, 2 500 m … ” and provided figures of the habitus (Tronquet 1981: fig. 3) and the aedeagus (Tronquet 1981: figs. 4: A – B). Based on the study of the holotype and additional material, I found that G. afghanicus is morphologically conspecific with G. (s. str.) kashmirensis. In consequence, G. afghanicus is synonymized with it. Geodromicus karakorus was originally described based on the holotype and paratype from “ Vallée Nang Mah, Karakorum … ”. Based on the study of the holotype, I found that this species based on the external and internal morphological features is conspecific with G. (s. str.) kashmirensis. Thus, I synonymyzed G. karakorus with this species.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776D7A74DD90FF796A73500E.taxon	description	(Figs. 44 – 45, 52 – 53, 63)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776D7A74DD90FF796A73500E.taxon	materials_examined	Type material examined. Neotype (here designated) of Geodromicus lebedevianus Roubal, 1929, ♀ (right antennomeres 9 – 11 missing): ‘ Kirghizia — E Talasskij | Alatau, Tschitschkan, | Itagar river near Dangi | 42 ° 09 ′ 37 N, 72 ° 51 ′ 04 E | 2000 m, 19. VII. 1998. | leg. Müller-Motzfeld’ <printed>, ‘ NEOTYPE | Geodromicus | lebedevianus Roubal, 1929 | Shavrin A. V. des. 2022 ’ <red, printed> (MNHUB). Material examined: KYRGYZSTAN: TALAS: 1 ♂, 2 ♀♀: Tian-Shan slope of Kyrgyzskyi Mts., Kara-Balta. 17.05.1990. S. V. Ovchinnikov leg. (cSh); YSSIK-KOL: 2 ♂♂, 1 ♀: Chichkan Valley, 42 ° 07 ′ N 72 ° 48 ′ E. 1850 m a. s. l. 30. VII. 1999. Müller-Motzfeld leg. (cA, cSch); 1 ♂: same data. 14.08.2003. T. Kölkebeck leg. (cSch).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776D7A74DD90FF796A73500E.taxon	description	Redescription. Measurements (n = 8): HW: 0.84 – 0.87; HL: 0.56 – 0.64; OL: 0.25 – 0.27; TL: 0.10 – 0.12; AL (averaged): 3.03; PL: 0.78 – 0.87; PWmax: 0.98 – 1.05; PWmin: 0.74 – 0.80; ESL: 1.54 – 1.65; EW: 1.72 – 1.85; MTbL (averaged): 1.27; MTrL (averaged): 0.56 (MTrL 1 – 4: 0.31; MTrL 5: 0.25); AW: 1.56 – 1.86; AedL: 0.82 – 0.87; BL: 4.72 – 6.05 (neotype: 4.90). Habitus as in Figs. 44 – 45. Body yellow-brown to dark-brown or black, with yellow-brown middle portions of elytra, sometimes forming very long and wide pale macula on each elytron; mouthparts, antennae and legs yellow to yellowish-brown (some specimens with paler antennomeres 1 – 2 or 1 – 3). Forebody without microsculpture except of head with very fine traces in frontal portion (some specimens) and coarse, dense and transverse meshes between ocelli; abdominal tergites with dense transverse microreticulation, usually finer on tergites VI – VII. Head 1.3 – 1.5 times as broad as long, with moderately deep and wide anterio-median depression; temples more than twice shorter than longitudial length of eyes, gradually narrowed toward neck; interocellar depression rectangular, narrow or moderately wide and deep, separated from infraorbital portions by narrow and deep anteocellar foveae, reaching level of middle or apical third of eyes. Ocelli large, distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Punctation fine and sparse, sometimes denser in middle. Antenna reaching middle length of elytra when reclined; antennomere 3 indistinctly about as long as 2, 4 slightly shorter than 3, 5 – 9 slightly longer than 4, 10 slightly shorter than 9, apical antennomere 1.3 – 1.6 times as long as 10. Pronotum small, slightly convex, 1.2 times as broad as long, from widest anterior third strongly narrowed basad toward long subparallel mediolateral portions; mediobasal portion with transverse and moderately deep depression. Punctation dense or sparse (some specimens), distinctly larger and deeper than that on head, finer in middle and / or mediobasal portions (some specimens with narrow impunctated portion in front of mediobasal depression). Elytra 1.1 times as broad as long, slightly or widely broadened posteriad, 1.8 – 1.9 times as long as pronotum. Punctation sparser or denser than that on pronotum, but distinctly larger and deeper, sometimes coarser and denser in parascutellar portion and denser or sparser and somewhat finer along suture. Metatibia slightly more than twice as long as metatarsi. Male. Profemuri slightly widened; protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII rounded or truncate. Apical margin of abdominal sternite VIII concave. Female. Profemuri and protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VIII and abdominal sternite VIII straight or rounded. Aedeagus with moderately wide basal portion, gradually narrowed toward rounded apex; parameres strongly broadened toward apical portions, distinctly exceeding apex of median lobe, with three moderately short apical and two – three preapical setae; internal sac weakly sclerotized, with very long and narrow flagellum, slightly widened in basal portion (Fig. 52). Lateral aspect of the aedeagus as in Fig. 53. Comparative notes. Based on the shape of the moderately narrow pronotum, G. (s. str.) lebedevianus is somewhat similar to G. (s. str.) medvedevi, from which it can be distinguished by the more sharply narrowed laterobasal portions of the pronotum, coloration of the elytra, and longer and broader parameres.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776D7A74DD90FF796A73500E.taxon	distribution	Distribution. Geodromicus (s. str.) lebedevianus is known from several localities of mountain regions of Kyrgyzstan (Fig. 63). Bionomics. Specimens were collected at elevations from 1850 to 2000 m a. s. l The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776D7A74DD90FF796A73500E.taxon	discussion	Remarks. Geodromicus lebedevianus was originally described based on the unspecified number of syntypes from “ Russisch-Asien: Talass-Tal … ”. The original description provided morphological features corresponding with some species of the convexicollis group, e. g. G. convexicollis. The studied specimens above have similar morphological features as that in the original description such as coloration of the body, small and narrow pronotum (“ Halsschild etwa so lang wie breit … ”), etc. Roubal compared it with several species of Geodromicus, including G. (s. str.) convexicollis, from which he distinguished it by “ durch die ganz andere Position der Ozellen, die Halsschildfurche etc. ”. The type material was looked by me in the collection of SNMB in 2015, but was not found except the box with the bottom label “ lebedevianus Roubal ” and one hole under it. Apparently, the type (types?) was lost. Thus, I designate a neotype here in the interest of the stability of nomenclature. I have chosen a specimen for it from the same mountain range (Talas Alatau) from which G. lebedevianus was originally described. The record of G. (s. str.) lebedivianus from Aksu-Zhabagly Nature Reserve by Kashcheev & Ishkov (2001) require confirmation. I identified only specimens of G. (s. str.) convexicollis during the study of Kashcheev`s material from this Nature Reserve (see above).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776E7A70DD90FB626AFF5476.taxon	description	(Figs. 54 – 59, 63)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776E7A70DD90FB626AFF5476.taxon	materials_examined	Material examined: KAZAKHSTAN: EAST KAZAKHSTAN: 1 ♂: 30 km S Panfilova, Sarybel. Under stones near the river. 27.07.1984. B. V. Iskakov leg. (ZIN); JAMBYL: 2 ♂♂: Chylik R., Sarybastau Gorge. 15.06.1988. V. Kashcheev leg. [paratypes?] (ZIN); 3 ♂♂: Dzhungar, Kokjota Mt., Topolevka. 27.06.2006. V. Kashcheev leg. (BMNH, cSh); 2 ♂♂: same data, Usek R. 10.07.2005 (cSh); 4 ♂♂, 2 ♀♀: same data, Argynkaty R. 31.08.2006 (cSh, ZIN); 1 ♂, 1 ♀: same data, Keskenterek R. 25.06.1998 (cSh); 7 ♂♂, 3 ♀♀: same data, Tentek R., Kokzhar. 25.08.2006 (cSh, MHNG, ZIN); ALMATY: 1 ♂, 1 ♀: Chelkydysu, 43 ° 13 ′ 02.4 ″ N 80 ° 17 ′ 04.7 ′ E′. 2345 m a. s. l. 21.07.2009. V. Kashcheev leg. (cSh); 6 ♂♂, 1 ♀: Ile Karatau, Zhynishke R. 22.06.1988. V. Kashcheev leg. (cSh); 1 ♂, 1 ♀: Prokhladnaya R. 2000 m a. s. l. 01.09.2008. V. Kashcheev leg. (ZIN); 7 ♂♂, 10 ♀♀: Ketmen Mts .., Shalkydysu R. 17.07.1988. V. Kashcheev leg. (BMNH, NHMB, cSh, ZIN); 1 ♂, 1 ♀: Baynakol R., near Karatogan. 11.09.1988. V. Kashcheev leg. (cSh); KYRGYZSTAN: 2 ♂♂: Issyk-Kul region, Kungey Alatau, Cholpon-Ata District, Grigoryevka, 42 °. 65 ′ N 77 ° 47 ′ E. 2200 m a. s. l. 09.06.2006. R. Ruta leg. (cRR).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776E7A70DD90FB626AFF5476.taxon	description	Redescription. Measurements (n = 50): HW: 0.86 – 1.25; HL: 0.57 – 0.76; OL: 0.27 – 0.32; TL: 0.12 – 0.15; AL (averaged): 3.78; PL: 0.88 – 1.61; PWmax: 0.78 – 1.86; PWmin: 0.68 – 1.21; ESL: 1.46 – 2.29; EW: 1.58 – 2.36; MTbL (averaged): 1.37; MTrL (averaged): 0.70 (MTrL 1 – 4: 0.40; MTrL 5: 0.30); AW: 1.21 – 2.14; AedL: 0.90 – 1.00; BL: 4.85 – 7.90. Habitus as in Figs. 57 – 59. Body yellow-brown to black, usually with paler elytra and sometimes paratergites of abdomen; mouthparts, antennae and legs yellow-brown to brown, usually with darker protibia and antennomeres 2 – 11 or 3 – 11. Head without or with fine transverse or isodiametric microsculpture, more distinct in middle and / or infraorbital portions; pronotum with very fine transverse microreticulation or without them; abdominal tergites with very dense transverse microreticulation, finer on abdominal tergites VI – VII. Head 1.5 – 1.6 times as broad as long, slightly or moderately strongly convex, with very deep and wide subtriangular or subrectangular anterio-median depression; temples convex, less than twice as long as longitudinal length of eyes; interocellar depression deep, subrectangular and moderately wide, separated from infraorbital portions by moderately deep anteocellar foveae, reaching level of apical third of eyes (some specimens without distinct foveae). Punctation sparse and fine, slightly larger, deeper and denser on infraorbital portions. Antenna reaching apical third or middle of elytra when reclined; antennomere 2 short and narrow, 3 distinctly longer than 2, 4 slightly broader and distinctly shorter than 3, 5 – 8 slightly longer than 4, 9 – 10 slightly broader than 8, apical antennomere 1.3 – 1.5 times as long as 10. Pronotum very convex, from middle or anterior third gradually or sharply narrowed toward short subparallel laterobasal margins in front of subacute hind angles; middle portion without or with very indistinct and narrow longitudinal depression, with shallow and weakly defined or with moderately deep and transverse mediobasal depression. Punctation distinctly denser, larger and deeper than that on head, with interspaces between punctures in middle about as long as one – four nearest punctures, sparser and finer in medioapical and mediobasal portions (some specimens with small oval impunctated area in front of mediobasal depression). Elytra slightly broader than long, slightly or widely broadened posteriad, 1.4 – 1.6 times as long as pronotum. Punctation denser, larger, deeper and sometimes coarser than that on pronotum, finer and denser in parascutellar portion and sometimes larger and denser along suture. Metatibia about twice as long as metarsi. Abdomen distinctly narrower than elytra. Male. Pronotum very wide and convex, sometimes significantly enlarged in some specimens (Fig. 58). Profemuri and protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII truncate or rounded. Apical margin of abdominal sternite VIII concave. Aedeagus slender, narrow, with moderately narrow basal portion, gradually narrowed toward rounded or subacute apex; parameres very wide, distinctly broadened toward apical portions, significantly exceeding apex of median lobe, with four long apical and three to four subapical setae; internal sac weakly sclerotized, with narrow and long flagellum (Figs. 54, 56). Lateral aspect of the aedeagus as in Fig. 55. Female. Pronotum moderately small, less convex. Profemuri and protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Geodromicus (s. str.) macrothorax differs from the remaining species of the convexicollis group by the shape of very wide and convex pronotum, sometimes with the presence of an indistinct longitudinal depression, and the different shape of the aedeagus with narrow median lobe and very long and wide parameres, significantly exceeding apex of the median lobe, with four long apical and three to four subapical setae.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776E7A70DD90FB626AFF5476.taxon	distribution	Distribution. Geodromicus (s. str.) macrothorax is distributed in the Middle Asia, and known from several localities in Kazakhstan and Kyrgyzstan (Fig. 63). Bionomics. Specimens were collected at elevations from 2000 to 2340 m a. s. l. and were taken from under stones near streams and rivers. It was collected together with specimens of G. (s. str.) convexicollis in some localities of south-eastern Kazakhstan (see above).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253776E7A70DD90FB626AFF5476.taxon	discussion	Remarks. Geodromicus macrothorax was originally described based on the holotype amd seven paratypes from “ Kungey-Alatau, upper flow of Chylik River, Sarybastau Gorge ”. According Kashcheev (1999), all types were deposited in ZIN, the collection of the Institute of Zoology in Astana, and the private collection of the author. During the study of collection of V. A. Kashcheev in ZIN, I have not found the type specimens which have been labeled as “ holotype ” or “ paratype ”. Two studied males from Chylik River (see the Material section) apparently refer to the paratypes of the private collection of V. Kashcheev (1953 – 2012) deposited to ZIN in 2014. It is quite possible that type specimens can be found in this museum later.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377687A4DDD90F9416BFC53BA.taxon	description	(Figs. 27, 60, 64 – 65)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377687A4DDD90F9416BFC53BA.taxon	materials_examined	Type material examined. Holotype, ♂ (Fig. 60; dissected): ‘ Arni Gad, | Mussoorie. ’ <yellow, printed>, ‘ Dr. Cameron | 13. IV. 22 ’ <printed>, ‘ Lesteva | torrentum | Cam. ’ <handwritten>, ‘ HOLOTYPE | Geodromicus (s. str.) | malcolmi sp. n. | Shavrin A. V. des. 2022 ’ <red, printed> (NMPC).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377687A4DDD90F9416BFC53BA.taxon	description	Description. Measurements (holotype): HW: 0.97; HL: 0.60; OL: 0.25; TL: 0.15; AL: 3.30; PL: 0.85; PWmax: 1.08; PWmin: 0.95; ESL: 1.87; EW: 1.82; MTbL: 1.25; MTrL: 0.57 (MTrL 1 – 4: 0.32; MTrL 5: 0.25); AW: 1.65; AedL: 0.94; BL: 4.90. Habitus as in Fig. 60. Body dark-brown, with slightly paler abdomen; mouthparts, antennae and legs brown (tarsi slightly paler). Forebody without microsculpture except head with traces of shallow transverse microreticulation in frontal portion and moderately coarse meshes between ocelli. Head 1.6 times as broad as long, with distinctly convex infraorbital portions and supra-antennal elevations, with deep and wide anterio-median depression, distinctly narrowed basad toward wide and deep interocellar depression; temples convex, 1.6 times as long as longitudinal length of eyes; interocellar depression separated from infraorbital portions by deep and long anteocellar foveae, reaching level of apical third of eyes. Ocelli large, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Punctation moderately sparse and fine, finer in middle and slightly larger, deeper and sparser in infraorbital portions. Antennomere 3 distinctly longer than antennomere 2, 4 distinctly shorter than 3, 5 – 10 slightly longer than 4, apical antennomere 1.3 times as long as 10. Pronotum moderately small, convex, 1.2 times as broad as long, from widest anterior third gradually narrowed toward moderately long subparallel mediolateral portions; mediobasal portion with deep, transverse depression. Punctation dense, moderately large and deep, sparser in medial portion in front of mediobasal depression. Elytra slightly longer than broad, slightly broadened posteriad, slightly more than twice as long as pronotum. Punctation dense, slightly larger and deeper than that on pronotum, coarser in parascutellar and finer and sparser in medioapical portions. Metatibia slightly more than twice as long as metatarsus. Male. Apical margin of abdominal tergite VIII and abdominal sternite VIII concave. Aedeagus with very wide basal portion, sharply narrowed toward narrow median lobe, from apical third gradually narrowed toward subacute apex; parameres wide, slightly widened apicad and slightly exceeding apex of median lobe, with four apical moderately long setae; internal sac without visible structures except short flagellum in middle portion (Fig. 64). Lateral aspect of the aedeagus as in Fig. 65. Female unknown. Comparative notes. Geodromicus (s. str.) malcolmi differs from the remaining species of the convexicollis group by the shape of the elytra slightly longer than broad and twice longer than pronotum, as well as by the different shape of the aedeagus.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377687A4DDD90F9416BFC53BA.taxon	distribution	Distribution. The new species is known only from the type locality in Uttarakhand, India (Fig. 27). Bionomics. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377687A4DDD90F9416BFC53BA.taxon	etymology	Etymology. The species is named in the memory of famous coleopterologist Malcolm Cameron (1873 – 1954), collector of the type specimen.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377687A4DDD90F9416BFC53BA.taxon	discussion	Remarks. Specimen apparently was collected together with Lesteva torrentum Cameron, 1924 from ‘ Mussorie District, 6500 to 7000 feet above the sea’ (Cameron 1924; Shavrin 2015).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4DDD90FE106B495140.taxon	description	(Fig. 61)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4DDD90FE106B495140.taxon	discussion	Remarks. Geodromicus medvedevi was originally described based on the holotype (Fig. 61) and three paratypes from “ NW Pakistan … 50 km ENE Chitral Shandur pass. Ca. 4200 m … ” (types in NMW) and was compared with G. cylindricus. Based on the narrow pronotum, it is similar to G. lebedevianus from Kyrgyzstan, from which it can be distinguished by the lateral margins of the pronotum gradually narrowed basad, and the parameres slightly broadened apicad and slightly exceeding apex of the median lobe.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4EDD90FC1F69605196.taxon	description	(Figs. 62, 66 – 70)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4EDD90FC1F69605196.taxon	materials_examined	Type material examined. Holotype of Geodromicus affinis Cameron, 1934, ♂ (Fig. 62; dissected; left antennomeres 8 – 11 missing): ‘ Type’ <round label with red margin>, ‘ N. India [underlined by yellow] | KASHMIR [handwrutten] ’ <printed>, ‘ Y. N. I. E | K 83 Kargil — | 24 May 1912 Wet | mossy stoned by spring, Geodromicus | affinis | TYPE [underlined] Cam’ <handwritten> (BMNH). Holotype of Geodromicus torrentis Puthz, 1973, ♂ (dissected): ‘ WEST-PAKISTAN | SWAT-REGION’ <handwritten>, ‘ Guhral spring | H 3 F | 17.7. [19] 68 ’ <handwritten>, ‘ HOLOTYPUS | ♂ [handwritten] ’ <red, printed>, ‘ Geodromicus | torrentis | nov. spec. | det. V. Puthz 19 [printed] 72 ’ <handwritten> (MHNG). Additional material examined. PAKISTAN: KHYBER PAKHTUNKHWA: 2 ♂♂: Swat, Utrar. 2200 – 2800 m a. s. l. 12 – 14.07.1982. Erber & Heinz leg. (MHNG); INDIA: KASHMIR: 1 ♂: Sonamarg. 13.07.1981. G. Ledoux leg. (cSh).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4EDD90FC1F69605196.taxon	description	Redescription. Measurements (n = 5): HW: 0.97 – 1.10; HL: 0.67 – 0.70; OL: 0.27 – 0.30; TL: 0.15 – 0.17; AL (averaged): 4.16; PL: 0.80 – 0.99; PWmax: 1.20 – 1.50; PWmin: 1.01 – 1.15; ESL: 1.92 – 2.12; EW: 2.00 – 2.26; MTbL (averaged): 1.40; MTrL (averaged): 0.65 (MTrL 1 – 4: 0.40; MTrL 5: 0.25); AW: 1.52 – 2.11; AedL: 0.50 – 1.02; BL: 5.25 – 6.70 (holotype of G. affinis: 6.50; holotype of G. torrentis: 6.70). Habitus as in Fig. 62. Body reddish-brown to black, with slightly paler abdomen; legs brown; mouthparts, antennae and tarsi yellow to dark-brown. Body without microsculpture except neck with very dense isodiametric meshes and abdominal tergites with fine and transverse microreticulation. Head 1.4 – 1.5 times as broad as long, with strongly convex supra-antennal elevations and infraorbital portions, with very deep and wide subrectangular anterio-median and moderately deep subtriangular interocellar depressions; temples slightly less than twice as long as longitudinal length of eyes; interocellar depression separated from infraorbital portions by strongly convergent latero-anteriad moderately narrow and deep anteocellar foveae. Ocelli small or moderately large, distance between ocelli slightly broader than distance between ocellus and posterior margin of eye. Punctation moderately fine and dense, slightly larger and deeper on infraorbital portions. Antenna reaching apical third of elytra when reclined; antennomere 2 about 1.3 times as long as basal antennomere, 3 1.3 – 1.4 times as long as 2, 4 – 5 slightly shorter than 3, 6 – 9 slightly longer and broader than 5, 10 slightly shorter than 9, apical antennomere 1.3 – 1.4 times as long as 10. Pronotum transverse, 1.5 times as broad as long, from widest anterior portion gradually or sharply narrowed toward moderately long and subparallel laterobasal margins; hind angles subacute or somewhat rounded; middle portion without or with indistinct longitudinal depression or with traces of it in medioapical portion; mediobasal depression very deep, transverse. Punctation moderately dense, distinctly larger and deeper than that on head, slightly denser in medioapical and sparser in mediobasal portions, sometimes with impunctate oval portion in front of mediobasal depression. Scutellum without punctures and microsculpture. Elytra distinctly broader than long, strongly broadened posteriad, more than twice as long as pronotum; hind margins broadly rounded. Punctation about as that on pronotum, or slightly larger and deeper, dense, finer along suture and in apical portions, finer or coarser in parascutellar area. Metatibia more than twice as long as metatarsus. Abdomen distinctly narrower than elytra, with very wide, transverse tomentose spots in middle of abdominal tergite IV. Male. Apical margin of abdominal tergite VIII truncate or rounded. Apical margin of abdominal sternite VIII concave. Aedeagus with moderately small and wide basal part, gradually narrowed toward narrow median lobe and rounded apex; parameres with broadened apical portions, distinctly exceeding apex of median lobe, with four (Figs. 66 – 67, 69) to six (Puthz 1973: figs. 1 – 2) moderately long apical setae; internal sac with short and wide strongly sclerotized flagellum, from widest middle portion gradually narrowed apically (Figs. x, x) and with short narrow basal part curved in basal portion. Lateral aspect of the aedeagus as in Fig. 68. Female unknown. Comparative notes. Based on the slender shape of the aedeagus and the presence of sclerotized wide flagellum, it is similar to species of the plagiatus group (Shavrin 2022), but it differs from them by the presence of an additional narrow basal part of the flagellum. Based on the length of the body, coloration, distance between ocelli and broadened parameres, it is somewhat similar to Eastern Palaearctic G. (s. str.) paramerus Shavrin 2000, known from Siberia and China (Shavrin 2000, 2022; Cheng et al. 2020), from which it can be distinguished by the more transverse pronotum, longer elytra, and by the narrower apical portions of the median lobe and the parameres.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4EDD90FC1F69605196.taxon	distribution	Distribution. The species is known from several localities in north-eastern Pakistan and India (Kashmir) (Fig. 70). Bionomics. Two males from Utrar (Pakistan, Swat) were collected at elevations from 2200 to 2800 m a. s. l. The holotype of G. affinis was collected in wet mosses. The detailed bionomical data are unknown	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377557A4EDD90FC1F69605196.taxon	discussion	Remarks. Geodromicus affinis Cameron, 1934 based on the holotype was originally described from “ Indian Tibet: Kargil ”. Puthz (1973) studied the type and provided the figure of the aedeagus (Puthz 1973: fig. 2). Geodromicus torrentis Puthz, 1973 based on the holotype was descibed from “ Pakistan: Swat Region: Guhral ”. I not found essential external and internal morphological differences between this specimen and additional material from Pakistan and Kashmir (see above), except the smaller length of the aedeagus (Puthz 1973: fig. 1). The length of the aedeagus in some species of Geodromicus can signigificantly vary. Thus, G. torrentis is synonymized with G. (s. str.) affinis.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377567A4BDD90FC3C6CFE5742.taxon	description	(Figs. 70 – 73, 78 – 79)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377567A4BDD90FC3C6CFE5742.taxon	materials_examined	Type material examined. Holotype of Geodromicus alticola Cameron, 1945, ♂ (Figs. 71, 73): ‘ Type’ <round label with red margin, printed>, ‘ Kashmir | Lianmarg | 10,000 ft. | vi-vii- 1931 | Dr. Cameron’ <printed>, ‘ G. | alticola | TYPE [red] Cam’ <handwritten>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ Geodromicus | alticola Cameron [handwritten] | Shavrin A. det. 2016 ’ <printed> (BMNH). Holotype of Geodromicus peraffinis Cameron, 1945, ♀ (Fig. 72): ‘ Type’ <round label with red margin, printed>, ‘ Kashmir | Lianmarg | 10,000 ft. | vi-vii- 1931 | Dr. Cameron’ <printed>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ G. | peraffinis | TYPE [red] Cam’ <handwritten>, ‘ Geodromicus | alticola Cameron [handwritten] | Shavrin A. det. 2016 ’ <printed> (BMNH). Holotype of Geodromicus brancuccii Coiffait, 1984, ♂: ‘ Pakistan 1979 | W. Wittmer’ <printed>, ‘ Sharan 1. - 2.7. | 2400 – 2700 m’ <printed>, ‘ TYPE’ <red, printed>, ‘ Geodromicus | brancucii | H. COIFFAIT [printed] det. 19 [printed] 83 ’ <handwritten> ‘ Geodromicus | brancuccii | Coiffait, 1984 | Shavrin A. V. det. 2015 [printed] <handwritten> (NHMB). Additional material examined. PAKISTAN: KHYBER PAKHTUNKHWA: 2 ♀♀: Swat, Kalâm, Falakser Glacier, Matiltan. 2500 m a. s. l. 19.08.1987. Heinz leg. (cSh); 1 ♂: Kaghan Valley, high meadows N Sharan. 2900 – 3100 m a. s. l. 27 – 29.07.2001. Heinz leg. (NME); AZAD JAMMU AND KASHMIR: 1 ♂: NW Junkar. 3500 m a. s. l. 25.07.2003 (cSch); INDIA: KASHMIR: 1 ♀: Margan Pass. 3000 m a. s. l. 20.07.1980. H. Aspöck & C. Rausch leg. (NHMB); 1 ♂, 2 ♀♀: ‘ Kashmir Lianmarg 10,000 ft. vi-vii- 1931 Dr. Cameron’ (BMNH).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377567A4BDD90FC3C6CFE5742.taxon	description	Redescription. Measurements (n = 12): HW: 1.00 – 1.10; HL: 0.73 – 0.85; OL: 0.25 – 0.27; TL: 0.17 – 0.22; AL: 3.67 – 4.15; PL: 1.01 – 1.25; PWmax: 1.25 – 1.45; PWmin: 0.85 – 0.97; ESL: 1.55 – 1.95; EW: 1.97 – 2.35; MTbL (averaged): 1.70; MTrL (averaged): 0.62 (MTrL 1 – 4: 0.32; MTrL 5: 0.30); AW: 1.87 – 2.17; AedL: 1.25 – 1.34; BL: 5.60 – 6.95 (holotype of G. alticola: 6.75; holotype of G. peraffinis: 6.95; holotype of G. brancuccii: 6.00). Habitus as in Figs. 71 – 72. Body reddish-brown to dark-brown, sometimes with slightly paler elytra and abdomen; mouthparts, legs and antennae yellow-brown to brown (antennomeres 4 – 11 sometimes paler; apical maxillary palpomere usually yellow). Clypeus with very fine, transverse microreticulation; latero-apical portions between supra-antennal elevations and apical margin of eye with dense and coarse meshes between punctures; portion between ocelli and neck with very dense and coarse isodiametric microsculpture; medioapical portion with indistinct transverse meshes in some specimens; scutellum with very fine transverse microsculpture in middle, indistinct in some specimens; abdominal tergites with fine and dense transverse microreticulation. Head 1.2 – 1.3 times as broad as long, with strongly convex supra-antennal elevations and slightly elevated infraorbital portions, with deep and wide, subtriangular anterio-median depression; temples slightly shorter than longitudinal length of eyes; interocellar depression subrectangular, moderately narrow and deep, separated from infraorbital portions by moderately narrow and deep anteocellar foveae, reaching level of apical third of eyes, or slightly longer. Eyes large, very convex, strongly protruded laterad. Ocelli moderately large, distance between ocelli about as long as distance between ocellus and posterior margin of eye, or slightly broader. Punctation irregular, moderately fine and sparse or somewhat large and deep, finer and sparser in middle or in infraorbital portions. Antenna reaching apical third of elytra when reclined; antennomere 2 narrow, 1.6 – 1.8 times as long as basal antennomere; 3 about as broad and 1.3 – 1.4 times as long as 2, 4 slightly shorter than 3, 5 – 8 slightly longer than 4, 9 – 10 slightly broader in their apical portions than 8, apical antennomere slightly longer than 10. Pronotum 1.1 – 1.2 times as broad as long, from widest anterior third strongly narrowed posterio-laterad toward acute apex; mediolateral margins slightly concaved in some specimens; laterobasal margins moderately long, subparallel; middle portion without or with indistinct longitudinal depression (one specimen), with shallow or moderately deep transverse mediobasal depression; anterior margin straight or slightly concave, about as broad as or slightly broader than straight or broadely concave posterior margin; lateral margins widely impressed and slightly reflexed; median part of pronotum separated from lateral flattened margins by slightly elevated and long ridge, starting from latero-apical and extending to laterobasal portions (Fig. 73). Punctation slightly larger and deeper than that on pronotum, dense or sparse, finer and sparser in mediobasal portion in front of mediobasal depression. Elytra somewhat flattened, strongly broader than long and strongly broadened posteriad, 1.4 – 1.5 times as long as pronotum; hind margins rounded. Punctation variable, from sparse and moderately fine (about as that on pronotum) or distinctly larger, deeper, and significantly denser than that on pronotum. Metatibia significantly more than twice as long as metatarsus. Abdomen slightly narrow than elytra, with a pair of wide, transverse tomentose spots in middle of abdominal tergite IV. Male. Profemuri and protarsomeres 1 – 4 wide. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII concave. Aedeagus with moderately wide basal part, gradually narrowed toward moderately wide preapical portion, with subacute apex; parameres narrow, distinctly shorter than apex of median lobe, with four short apical setae; preapical portion of median lobe with a pair of long sclerotized structures; internal sac weakly sclerotized, long, with strongly sclerotized, wide, moderately long flagellum, spirally folded in basal portion (Fig. 78). Lateral aspect of the aedeagus as in Fig. 79. Female. Profemuri and protarsomeres 1 – 4 narrow. Apical margins of abdominal tergite VIII and sternite VIII truncate or rounded. Comparative notes. Geodromicus (s. str.) alticola differs from other species of the genus by the unusual shape of lateral portions of the pronotum with the presence of long lateral ridge between it and the median portion, and by details of the external and internal morphology of the aedeagus.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377567A4BDD90FC3C6CFE5742.taxon	distribution	Distribution. The species is known from several localities in north-eastern Pakistan and India (Kashmir) (Fig. 70). Bionomics. Specimens were collected from 2400 to 3500 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377567A4BDD90FC3C6CFE5742.taxon	discussion	Remarks. Geodromicus alticola was originally described based on the holotype (“ Type in my collection ”) from “ Kashmir: Lianmarg … ”. Geodromicus peraffinis was originally described based on the holotype (female) from the same locality with G. alticola. It was compared by Cameron with G. alticola (“ Extremely like alticola Cam., from which it only differs in the shorter, slightly transverse thorax .. the puncturation scarcely differing from that of alticola … ”). I have not found external morphological differences between this type and other studied specimens of G. alticola. Proportions of the pronotum and elytra, as well as the punctation of the forebody vary in this species. Thus, I synonymyzed G. peraffinis with G. alticola. Geodromicus brancuccii was originally described based on the holotype from “ Pakistan, Sharan … ”. I have not found sufficient morphological differences between it and other studied specimens of G. alticiola. Consequently, G. brancuccii is synonymized with it. Besides that, the original description and the general shape of the aedeagus (Coiffait 1984: figs. 37 – 38) of G. pakistanus Coiffait, 1984, originally described from “ Pakistan, Murree-Abbottabad … ” are also similar to that of studied specimens of G. (s. str.) alticola. But the study of the holotype of this species is required.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377537A4ADD90FA486B4D55AE.taxon	description	(Figs. 70, 74, 80 – 81)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377537A4ADD90FA486B4D55AE.taxon	materials_examined	Type material examined. Lectotype (here designated) of Geodromicus curvipes Cameron, 1924, ♂ (Fig. 74): ‘ Type | H. T. ’ <round label with red margin, printed>, ‘ Gahan 7000`| Simla Hills. | Dr. Cameron | IX. 1921. ’ <printed>, ‘ Geodromicus | curvipes | Cam’ <handwritten>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ LECTOTYPE | Geodromicus | curvipes Cameron, 1924 | Shavrin A. V. des. 2022 ’ <red, printed>, ‘ Geodromicus (s. str.) | curvipes Cameron, 1924 | Shavrin A. det. 2022 ’ <printed> (BMNH). Paralectotype, ♂ (dissected): ‘ Gahan 7000`| Simla Hills’ <printed>, ‘ Dr. Cameron | IX. 1921. ’ <printed>, ‘ Geodromicus | curvipes | Cam’ <handwritten>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ Geodromicus (s. str.) | curvipes Cameron, 1924 | Shavrin A. det. 2022 ’ <printed> (BMNH).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377537A4ADD90FA486B4D55AE.taxon	description	Redescription. Measurements (n = 2): HW: 0.90 – 0.95; HL: 0.62; OL: 0.25; TL: 0.17 – 0.19; AL (lectotype): 2.50; PL: 1.11 – 1.12; PWmax: 1.12; PWmin: 0.78 – 0.85; ESL: 1.36 – 1.42; EW: 1.49 – 1.54; MTbL (lectotype): 0.90; MTrL (lectotype): 0.57 (MTrL 1 – 4: 0.30; MTrL 5: 0.27); AW: 1.48 – 1.55; AedL: 0.87; BL: 4.80 – 4.95 (lectotype). Habitus as in Fig. 74. Body, mouthparts, antennae and legs dark-brown; tarsi yellow-brown. Head with fine microsculpture: transverse in clypeus and isodiametric in middle portion; neck with very dense isodiametric meshes; pronotum with very fine isodiametric microreticulation, indistinct in mediobasal portion; scutellum with fine transverse microsculpture; abdomen with dense and transverse meshes. Pubescence of forebody very fine, semierect, moderately short and sparse, longer in apical portion of head; abdomen with moderately dense and very fine, decumbent pubescence. Head 1.4 – 1.5 times as broad as long, with distinctly convex supra-antennal elevations, middle portion between anterior margin of eyes and infraorbital portions; anterio-median depression subrectangular, wide and moderately deep; temples very convex, 1.3 – 1.4 times as long as longitudinal length of eyes; interocellar depression narrow, subtriangular and moderately deep, with narrow and deep anteocellar foveae, reaching level of apical third of eyes. Eyes large, strongly protruded laterad. Ocelli large, distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Punctation moderately large and deep, sparser and finer in middle portion. Maxillary palpomere 3 slightly broadened apically; apical palpomere slightly shorter than 3, from middle gradually rounded toward rounded apex. Antenna reaching middle of elytra when reclined; antennomere 2 about 1.3 times as long as basal antennomere, 3 distinctly longer than 2, 4 shorter than 3, 5 – 8 slightly longer and broader than 4, 9 – 10 slightly broader than 8, apical antennomere 1.3 times as long as 10. Pronotum convex, about as broad as long, from widest middle strongly narrowed anteriad toward rounded apical angles and gradually narrowed toward base; narrowest basal part of pronotum slightly elongate, indistinctly concave laterally and with slightly divergent lateral subacute hind angles; middle portion with long and moderately wide longitudinal depression (lectotype with depression reaching middle length of pronotum; longitudinal depression of paralectotype reaching mediobasal portion of pronotum); mediobasal depression transverse and moderately deep; apical portion of pronotum slightly protruded apicad, with anterior margin widely concave, about as broad as slightly concave basal margin, with bordered and slightly reflexed edges, larger in latero-apical portions; lateral margins of pronotum narrowly bordered. Punctation about as that in middle portion of head, but slightly denser, significantly sparser and finer in mediobasal portion in front of mediobasal depression Elytra convex, depressed in middle, slightly broader than long and slightly broadened posteriad, 1.2 times as long as pronotum; hind margins widely rounded. Punctation denser, coarser, larger and deeper than that on pronotum, with distance between punctures in middle as long as diameters of one – two nearest punctures, finer along suture (paralectotype) and / or apical portions. Metatibia about 1.5 times as long as metatarsus. Abdomen about as broad as elytra, with two large and transverse tomentose spots in middle of abdominal tergite IV and moderately wide palisade fringe on apical margin of abdominal tergite VII. Male. Profemuri very wide; inner margins of protibia widely curved; protarsomeres 1 – 4 moderately wide. Apical margin of abdominal tergite VIII truncate or slightly concave. Apical margin of abdominal sternite VIII widely concave. Aedeagus with wide basal part, significantly narrowed toward median lobe; median lobe with elongate and moderately wide apical portion, from apical third gradually narrowed toward rounded apex; parameres narrow and short, not reaching apex of median lobe, with three moderately short apical and two subapical setae; internal sac with moderately short and wide field of small thorns, with elongate, sclerotized, apical structures and very long, narrow flagellum (Fig. 80). Lateral aspect of the aedeagus as in Fig. 81. Female unknown. Comparative notes. Based on the general shape and the coloration of the body with similar shape of the pronotum, the presence of microsculpture and longitudinal depression on the pronotum, the similar shape of the protibia in males and the general shape of the aedeagus, G. (s. str.) curvipes is similar to the Himalayan G. (s. str.) subsimilis (see below). From this species it can be distinguished by the smaller body, slightly narrower pronotum, finer microsculpture on the pronotum, broader apical portion of the median lobe and different internal morphology of the aedeagus, with the presence of the long flagellum (missing in G. (s. str.) subsimilis).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377537A4ADD90FA486B4D55AE.taxon	distribution	Distribution. The species is known from the type localitiy in Himachal Pradesh, India (Fig. 70). Bionomics. Specimens were collected from 2100 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB25377537A4ADD90FA486B4D55AE.taxon	discussion	Remarks. Geodromicus curvipes was originally described based on unspecified number of specimens from “ Simla Hills; Gahan, 7000 feet above the sea ”. The male in better condition and with additional label ‘ Type H. T. ’ is designated as the lectotype in order to fix the identity of the name.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A45DD90FF796B0454C6.taxon	description	(Figs. 70, 75)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A45DD90FF796B0454C6.taxon	materials_examined	Type material examined. Holotype of Geodromicus ruficornis Cameron, 1941, ♀ (Fig. 75): ‘ Type’ <round label with red margin, printed>, ‘ Kashmir | Gulmarg | vi-vii- 31 | Dr. Cameron’ <printed>, ‘ G. ruficornis | TYPE [red] curvipes Cam’ <handwritten>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ Geodromicus | ruficornis Cameron, 1941 | Shavrin A. det. 2022 ’ <printed> (BMNH). Additional material examined. INDIA: KASHMIR: 1 ♀: ‘ Kashmir Gulmarg vi-vii- 31 Dr. Cameron’ (BMNH).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A45DD90FF796B0454C6.taxon	description	Redescription. Measurements (n = 2): HW: 0.76 – 0.82; HL: 0.58 – 0.60; OL: 0.20; TL: 0.13 – 0.15; AL (holotype): 0.40; PL: 0.72 – 0.74; PWmax: 0.87 – 0.90; PWmin: 0.67 – 0.72; ESL: 1.13 – 1.31; EW: 1.48 – 1.65; MTbL (holotype): 0.82; MTrL (holotype): 0.42 (MTrL 1 – 4: 0.22; MTrL 5: 0.20); AW: 1.57 – 1.60; BL: 4.60 – 4.75 (holotype). Habitus as in Fig. 75. Body reddish-brown, with slightly paler elytra; mouthparts, antennae and legs yellowbrown; apical palpomere of maxillary palpi and tarsi yellow. Head with fine transverse microsculpture, indistinct in middle and somewhat coarser on infraorbital portions, portion between ocelli and neck with dense isodiametric microreticulation; medioapical portion of pronotum with indistinct transverse meshes (indistinct in holotype). Pubescence of forebody moderately dense, yellow, semierect, longer on head; abdomen with very dense, decumbent pubescence. Head 1.3 times as broad as long, with slightly convex supra-antennal elevations, middle portion between apical margins of eyes and infraorbital portions, with deep and wide subrectangular anterio-median depression; temples convex, distinctly shorter than longitudinal length of eyes; interocellar depression moderately large and deep, subtrapezoidal, with deep anteocellar foveae, reaching level of apical third of eyes. Eyes moderately small, strongly protruded laterad. Ocelli small, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Punctation moderately fine, slightly larger in middle and denser on infraorbital portions. Apical palpomere about as long as palpomere 3, from middle gradually narrowed toward subacute apex. Antenna reaching middle of elytra when reclined; antennomere 2 about 1.3 times as long as basal antennomere, 3 distinctly longer than 2, 4 – 8 slightly shorter than 3, 9 – 10 indistinctly broader than 8, apical antennomere about 1.4 times as long as 10. Pronotum small, slightly convex, from widest anterior third strongly narrowed posteriad toward obtuse hind angles; laterobasal portions moderately short, slightly concave; middle portion without longitudinal depression; mediobasal depression shallow, transverse; anterior margin rounded, about as long as slightly concave posterior margin. Punctation moderately sparse, slightly larger and deeper than that on head, sparser and finer in mediobasal portion. Elytra strongly broader than long and significantly broadened posteriad, 1.5 – 1.7 times as long as pronotum, reaching basal margin of abdominal tergite IV; hind margins widely rounded. Punctation larger, deeper and denser than that on pronotum (coarser and larger in holotype). Metatibia about twice as long as metatarsus. Abdominal tergite IV with two moderately small and round tomentose spots. Male unknown. Female. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Geodromicus ruficornis can be distinguished from other Himalayan species of the genus by shapes of small pronotum and the moderately short elytra strongly broadened posteriad. Currently, it is impossible to find relative species due the missing of males for this species.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A45DD90FF796B0454C6.taxon	distribution	Distribution. The species is known from the type localitiy in Kashmir, India (Fig. 70). Bionomics. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A44DD90F8F66AEC55CA.taxon	description	(Figs. 70, 76, 82 – 83)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A44DD90F8F66AEC55CA.taxon	materials_examined	Type material examined. Holotype of Geodromicus subsimilis Cameron, 1941, ♂: ‘ Type’ <printed label with red margin>, ‘ SYN- | TYPE’ <printed label with blue margin>, ‘ Ghum dist. | Mangpo | v- 31 | Dr. Cameron’ <printed>, ‘ G. | subsimilis | TYPE [red] Cam. ’ <handwritten>, ‘ M. Cameron. | Bequest. | B. M. 1955 - 147. ’ <printed>, ‘ Geodromicus | subsimilis Cam. [handwritten] | Shavrin A. det. 2016 ’ <printed> (BMNH). Additional material examined. INDIA: WEST BENGAL: 1 ♂: ‘ Ghum. dist. Mangpo v- [19] 31 Dr. Cameron’, ‘ M. Cameron. Bequest. B. M. 1955 - 147. ’, ‘ subsimilis Cam. ’, ‘ SYNTYPE’ (BMNH); 2 ♂♂: ‘ Ghum district vvi- [19] 31 Dr. Cameron’, ‘ stream moss’, ‘ W. Steel coll. B. M. 1969 - 552. ’, ‘ SYNTYPE’ (BMNH); 1 ♂: ‘ Ghum dist. Tiger Hill 8,500 – 10,000 ft. v-vi- 1931 Dr. Cameron’, ‘ Stream moss’, ‘ M. Cameron. Bequest. B. M. 1955 - 147. ’, ‘ SYNTYPE’ (BMNH); NEPAL: 1 ♀: Mechi / Taplejung, 24 km NE Taplejung, 1550 m, Sekathum, Camp, 27 ° 32 ’ 10 ″ N 87 ° 48 ’ 29 ″ E. 06.05.2003. A. Weigel leg. (NME).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A44DD90F8F66AEC55CA.taxon	description	Redescription. Measurements (n = 6): HW: 0.91 – 0.97; HL: 0.66 – 0.70; OL: 0.25; TL: 0.20 – 0.22; AL (holotype): 3.08; PL: 1.16 – 1.25; PWmax: 1.20 – 1.35; PWmin: 0.84 – 0.90; ESL: 1.38 – 1.70; EW: 1.60 – 1.85; MTbL (holotype): 1.37; MTrL (holotype): 0.51 (MTrL 1 – 4: 0.26; MTrL 5: 0.25); AW: 1.45 – 1.67; AedL: 0.60 – 0.80; BL: 5.15 – 6.45 (holotype). Habitus as in Fig. 76. In general appearance it similar in habitus with G. (s. str.) curvipes. Body, mouthparts, antennae and legs brown to dark-brown (legs in some specimens slightly paler); tarsi yellow-brown. Head with distinct microsculpture: finer and transverse on clypeus, coarse, transverse and very dense in middle portion; pronotum with distinct and dense, transverse meshes; scutellum with fine transverse microsculpture; abdomen with fine and dense, transverse microreticulation. Head 1.3 times as broad as long, with deep and wide, subtriangular anterio-median depression; temples convex, slightly less than longitudinal length of eyes; interocellar depression wide and deep, subrectangular or subtriangular. Punctation about as that on head, but denser. Apical maxillary palpomere distinctly shorter than preceding palpomere, from middle strongly narrowed toward subacute apex. Pronotum very convex, slightly broader than long, from widest middle gradually narrowed both anteriad and posteriad; narrowest basal part of pronotum moderately short, subparallel; hind angles subacute; middle portion with long, moderately wide and deep longitudinal depression, deeper in middle and sometimes indistinct in medioapical and mediobasal portions; mediobasal depression transverse, shallow or indistinct, or moderately deep; apical portion of pronotum distinctly protruded apicad, with wide and slightly concave anterior margin and with distinctly bordered apical margins. Punctation very dense, about as that on head, sparser and finer in mediobasal portion in front of mediobasal depression. Elytra 1.1 – 1.3 times as long as pronotum. Punctation very dense, slightly larger and deeper than that on pronotum. Metatibia twice as long as metatarsus. Abdomen slightly narrower than elytra. Male. Profemuri and protarsomeres 1 – 4 wide; inner margins of protibia significantly and widely curved. Pronotum somewhat elongate and more convex. Apical margins of abdominal tergite VIII and sternite VIII slightly concave. Aedeagus with moderately small and narrow basal part, gradually narrowed toward median lobe; median lobe with very narrow apical portion, from apical third gradually narrowed toward subacute apex; parameres narrow, slightly broadened apically, with three moderately long apical setae; internal sac short and wide, with paired narrow elongate sclerotized structures in basal, and very long four sclerotized structures in apical portion; flagellum missing (Fig. 82). Lateral aspect of the aedeagus as in Fig. 83. Female. Profemuri and protarsomeres 1 – 4 narrow; protibia without modifications. Pronotum moderately short, slightly convex. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Based on the general shape and the coloration of the body with the similar shape of the pronotum, the presence of microsculpture and longitudinal depression on the pronotum, the shape of the curved protibia in males, and the general shape of the aedeagus, G. (s. str.) subsimilis is similar to Indian G. (s. str.) curvipes (see above). From this species it can be distinguished by the longer body, slightly broader pronotum, coarser microsculpture on the pronotum, narrower apical portion of the median lobe and different internal morphology of the aedeagus.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A44DD90F8F66AEC55CA.taxon	distribution	Distribution. The species is known from three locations in Nepal and West Bengal, India (Fig. 70). Bionomics. Specimens were collected from 1550 to 3050 m a. s. l. The detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775D7A44DD90F8F66AEC55CA.taxon	discussion	Remarks. Geodromicus subsimilis was originally described based on the holotype (“ Type in my collection ”) from “ Ghum district: Mangpu: Tiger Hill, 8000 – 10,000 feet ”. It is recorded here from Nepal for the first time.	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FF316C355018.taxon	materials_examined	Material examined. RUSSIA: MARITIME PROVINCE: 3 ♂♂: Tigrovyi. 25 – 27.07.1990. S. Becvar leg. (MHNG); 1 ♀: environs of Vladivostok, Sedanka. 28.07.1992. S. Beloborodov leg. (NHMB).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FF316C355018.taxon	discussion	Remarks. Geodromicus (s. str.) beibienkoi is widely distributed in the Eastern Palaearctic Region and known from Russia (Siberia, Russian Far East), Korea and China (Cheng et al. 2020).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FDBB6A5551BE.taxon	description	Materialexamined. JAPAN: FUKUSHIMAPREFECTURE: 1 ♀: Niwasaka. 30.04.1990 (MHNG); NAGANOPREFECTURE: 1 ♂: ‘ Yatsugadake Py, SHINANO by M. Suzuki. 20. Aug. 1918 ’ (ZIN); KYOTO PREFECTURE: 1 ♂: ‘ Mt Kibune near KYOTO by M. Suzuki. 27. Jul. 1919 ’ (ZIN).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FDBB6A5551BE.taxon	discussion	Remarks. Geodromicus (s. str.) lestevoides is known from the Russian Far East, Korea, Japan and north-western China (Shavrin 2018, Cheng et al. 2020).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FC1E6D6656CA.taxon	materials_examined	Material examined. CHINA: GANSU: 1 ♂: Dalija Shan, 46 km W Linxia, 2980 m a. s. l., near small river in a pasture formation under quite close to the water, from soaking wet moss. 10.07.1994. A. Smetana leg. (MHNG).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FC1E6D6656CA.taxon	discussion	Remarks. The species is known from Russia (Eastern Siberia, Buryatia) and China (Cheng et al. 2020).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FACA6C495443.taxon	materials_examined	Material examined. RUSSIA: MARITIME PROVINCE: 1 ♀: 30 km E Ussuriysk, Kamenushka, 20 – 25.06.1990. S. Kasantsev leg. (NHMB).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90FACA6C495443.taxon	discussion	Remarks. Geodromicus (s. str.) sibiricus is widely distributed in the Eastern Palaearctic Region: from Baikal Lake to Maritime province of Russia, northwest to China and Korea (Cheng et a l. 2020).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90F97569A255E3.taxon	description	(Fig. 77)	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90F97569A255E3.taxon	materials_examined	Material examined. CHINA: SICHUAN: 25 ♂♂, 22 ♀♀: Gongga Shan, Hailuogou, above Camp 3, 29 ° 35 ′ N 102 ° 00 ′ E. 2800 – 3300 m. 06 – 08.07.1998. J. Farkač leg. (NHMB); 1 ♂, 1 ♀: Gonggashan-Hailuogou, 29 ° 35 ′ N 102 ° 00 ′ E. 03 – 06.07.1994. 2900 – 3200 m a. s. l. J. Farkač & D. Král leg. (NHMB).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
03FFB253775E7A46DD90F97569A255E3.taxon	discussion	Remarks. Habitus as in Fig. 77. Geodromicus (s. str.) smetanai is known from several locations of Sichuan, China (Shavrin 2018).	en	Shavrin, Alexey V. (2022): The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa 5213 (5): 451-496, DOI: https://doi.org/10.11646/zootaxa.5213.5.1
