identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F171B72AFFD0456FFCAAFCF54390FB7F.text	F171B72AFFD0456FFCAAFCF54390FB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ligabuesaurus BONAPARTE, GONZALEZ RIGA & APESTEGUIA 2006	<div><p>LIGABUESAURUS BONAPARTE, GONZÁLEZ RIGA &amp; APESTEGUÍA, 2006</p> <p>Etymology</p> <p>The generic name was defined by Bonaparte et al. (2006) in honour of Italian philanthropist Dr Giancarlo Ligabue, with the Greek suffix - σαῦρος (sauros), lizard or reptile.</p> <p>Diagnosis</p> <p>As for the species.</p></div> 	https://treatment.plazi.org/id/F171B72AFFD0456FFCAAFCF54390FB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bellardini, Flavio;Coria, Rodolfo A;Pino, Diego A;Windholz, Guillermo J;Baiano, Mattia A;Martinelli, Augustin G	Bellardini, Flavio, Coria, Rodolfo A, Pino, Diego A, Windholz, Guillermo J, Baiano, Mattia A, Martinelli, Augustin G (2022): Osteology and phylogenetic relationships of Ligabuesaurus leanzai (Dinosauria: Sauropoda) from the Early Cretaceous of the Neuquén Basin, Patagonia, Argentina. Zoological Journal of the Linnean Society 196 (4): 1333-1393, DOI: 10.1093/zoolinnean/zlac003, URL: https://academic.oup.com/zoolinnean/article/196/4/1333/6553819
F171B72AFFD04546FCAFFB1E4417FD21.text	F171B72AFFD04546FCAFFB1E4417FD21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ligabuesaurus leanzai BONAPARTE, GONZALEZ RIGA & APESTEGUIA 2006	<div><p>LIGABUESAURUS LEANZAI BONAPARTE, GONZÁLEZ RIGA &amp; APESTEGUÍA, 2006</p> <p>Type species and etymology</p> <p>The name of the type species was erected in honour of geologist Dr Héctor Leanza, who reported about the presence of fossils at Cerro de los Leones, Picún Leufú, Neuquén Province, Argentina.</p> <p>Holotype</p> <p>MCF-PVPH-233 (Fig. 4A): a single, large-sized, incomplete and disarticulated sauropod specimen represented by ten maxillary teeth (MCF-PVPH-233/ 01), a posterior cervical vertebra (MCF-PVPH-233/02), an anterior dorsal vertebra (MCF-PVPH- 233 / 03), two articulated midposterior dorsal vertebrae (MCF-PVPH-233/04 and MCF-PVPH- 233 / 05), two articulated posterior dorsal vertebrae (MCF-PVPH-233/06 and MCF-PVPH-233/07), both scapulae (MCF-PVPH-233/08 and MCF-PVPH-233/09), a left humerus (MCF-PVPH-233/10), a proximal and distal epiphysis of the right humerus (MCF-PVPH-233/11 and MCF-PVPH-233/12), a right metacarpal II (MCF-PVPH- 233/13), a right metacarpal III (MCF-PVPH- 233/ 14), a distal epiphysis of the left metacarpal II (MCF-PVPH-233/15), a distal epiphysis of the left metacarpal IV (MCF-PVPH-233/16), a right femur (MCF-PVPH-233/17), a right tibia (MCF-PVPH-233/18), a right fibula (MCF-PVPH-233/19), a right astragalus (MCF-PVPH-233/20) and a nearly complete and articulated right pes, with five metatarsals and three phalanges (MCF-PVPH-233/21–MCF-PVPH-233/28).</p> <p>Referred specimens</p> <p>MCF-PVPH-261 (Fig. 4A): several postcranial elements from the type quarry no. 4 of Ligabuesaurus, consisting of a mid-cervical vertebra (MCF-PVPH-261/16), two posterior cervical vertebrae (MCF-PVPH-261/01 and MCF-PVPH-261/02), an anterior caudal vertebra (MCF-PVPH-261/15), an incomplete dorsal rib (MCF-PVPH-261/17), both coracoids (MCF-PVPH-261/05 and MCF-PVPH-261/06), a distal half of left radius(?) (MCF-PVPH-261/07), a partial left ilium (MCF-PVPH-261/08), both pubes (MCF-PVPH-261/09–MCF-PVPH-261/11), a left femur (MCF-PVPH-261/12), a proximal epiphysis of the left tibia (MCF-PVPH-261/13) and a proximal epiphysis of the left fibula (MCF-PVPH-261/14).</p> <p>MCF-PVPH-228 and MCF-PVPH-908 (Fig. 4B), a single, large-sized and incomplete sauropod specimen from quarry no. 3, represented by the following associated bones: two articulated posterior cervical vertebrae (MCF-PVPH- 228 / 01 and MCF-PVPH-261/02), an anterior dorsal vertebra (MCF-PVPH-908), two articulated mid-posterior dorsal vertebrae (MCF-PVPH-228/03 and MCF-PVPH-228/04), six incomplete dorsal ribs (MCF-PVPH-228/05–MCF-PVPH-261/10) and a right scapula (MCF-PVPH-228/11).</p> <p>MCF-PVPH-744 (Fig. 4B), one isolated, almost complete tooth.</p> <p>See the Supporting Information (Table S1 and Section 1.1.2 ‘Comments on referred specimens of Ligabuesaurus ’) for considerations about the composition of the type material of Ligabuesaurus.</p> <p>Locality and horizon</p> <p>The fossil remains of Ligabuesaurus come from the Cerro de los Leones locality, a hill located ~ 10 km to the southwest of Picún Leufú city, southern Neuquén Province, Patagonia, Argentina (Fig. 1A, B). The fluvial deposits outcropping in this area were referred to the lower section of the Cullin Grande Member (Martinelli et al., 2007), the upper member of the Lohan Cura Formation (Bajada del Agrio Group, Lower Cretaceous, Albian). The type quarry (no. 4) was opened in the fossiliferous level no. 2 (sensu Martinelli et al., 2007) in the southern flank of the Cerro de los Leones (Fig. 1C) and 40 m to the east of quarry no. 3, where part of the referred specimen was found (Supporting Information, Table S1). The sauropod remains were found in laminate mudstones with interbedded fine- to very fine-grained sandstones. These fluvial deposits were dated as Albian and are considered to have been formed in a distal floodplain in semi-arid climatic conditions (Martinelli et al., 2007).</p> <p>Comments on original diagnosis</p> <p>In the original description of Ligabuesaurus, Bonaparte et al. (2006) identified four autapomorphies. The first three are listed below with the numbers (1), (2) and (3). The fourth autapomorphy, listed by Bonaparte et al. (2006) as (4) rudimentary prespinal lamina (prsl) on the posterior cervical and anterior dorsal vertebrae, is not used here for the following reasons: in the posterior cervical vertebra MCF-PVPH-233/02 the prsl is not present (Fig. 5L), whereas in the anterior dorsal vertebra MCF-PVPH-233/03 there is a reduced lamina on the dorsalmost portion of the anterior face of the neural spine (Fig. 5M). However, in the anterior dorsal vertebra MCF-PVPH-908 the prsl is not rudimentary but represented by a narrow and prominent lamina, well developed from the base to the apex of the neural spine (Fig. 5N). Therefore, we consider that the rudimentary prsl on the posterior cervical and anterior dorsal vertebrae is not a pertinent autapomorphy for Ligabuesaurus and exclude it from the diagnosis.</p> <p>Revised diagnosis</p> <p>Ligabuesaurus leanzai is characterized by the following autapomorphies: (1) laminar and anteroposteriorly compressed neural spines on posterior cervical and anterior dorsal vertebrae that are rhomboid in shape and wider than the vertebral centra; (2) spinoprezygapophyseal laminae in posterior cervical vertebrae forked to form two pairs of laminae: the medial pair unites them towards the top of the neural spine, and the lateral pair form the lateral border of the neural spine; (3) posterior cervical and anterior dorsal vertebrae with low neural arch pedicels, less than onethird of the height of the anterior articular surface; (4) humeral head expanded posteriorly (D’Emic, 2012); (5) quadrangular ventral half of the coracoid in lateral view (Fig. 5A); (6) fossae on proximoventral faces of metatarsals II and III (D’Emic, 2012); and (7) deep pit on ventrodistal face of pedal phalanx II-1 (modified from D’Emic, 2012). With regard to (4), (6) and (7), in the extended contribution on the early evolution of Titanosauriformes, D’Emic (2012; appendix 4) also provided a diagnosis for Ligabuesaurus, identifying five autapomorphies, some of which are not included in the diagnosis to represent morphological features with a wide distribution within Sauropoda. In this sense, the distal scapular blade with rounded dorsal expansion (autapomorphy 1; D’Emic, 2012) is a condition that Ligabuesaurus shares with several Titanosauriformes (e.g. Brachiosaurus Riggs, 1903, Brontomerus Taylor, Wedel &amp; Cifelli, 2011, Euhelopus Romer, 1956, Giraffatitan Paul, 1988, Rukwatitan Gorscak et al., 2014), hence it is excluded from the diagnosis. Likewise, the gracile humerus of Ligabuesaurus (autapomorphy 3; D’Emic, 2012) is a plesiomorphic condition that is also present in several sauropods, such as Alamosaurus Gilmore, 1922, Chubutisaurus Del Corro, 1975, Europasaurus Mateus et al. in Sander et al., 2006, Giraffatitan, Patagosaurus Bonaparte, 1979, Rinconsaurus Calvo &amp; González Riga, 2003 and Wintonotitan Hocknull et al., 2009. In contrast, we agree with D’Emic that the deep pit on the ventrodistal face of the pedal phalanx represents an autapomorphy of Ligabuesaurus (autapomorphy 5; D’Emic, 2012), but we have reconsidered the pedal element (MCF-PVPH-233/28) as a phalanx II-1 and not a I-1 (contra D’Emic, 2012). However, also in the phalanx I-1 (MCF-PVPH-233/26) there is a ventrodistal vascular foramen, but it is small and poorly preserved, hence it is not included in the present diagnosis.</p> </div>	https://treatment.plazi.org/id/F171B72AFFD04546FCAFFB1E4417FD21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bellardini, Flavio;Coria, Rodolfo A;Pino, Diego A;Windholz, Guillermo J;Baiano, Mattia A;Martinelli, Augustin G	Bellardini, Flavio, Coria, Rodolfo A, Pino, Diego A, Windholz, Guillermo J, Baiano, Mattia A, Martinelli, Augustin G (2022): Osteology and phylogenetic relationships of Ligabuesaurus leanzai (Dinosauria: Sauropoda) from the Early Cretaceous of the Neuquén Basin, Patagonia, Argentina. Zoological Journal of the Linnean Society 196 (4): 1333-1393, DOI: 10.1093/zoolinnean/zlac003, URL: https://academic.oup.com/zoolinnean/article/196/4/1333/6553819
