taxonID	type	description	language	source
03E48797FFE5FF94FF214EDFC1C4FF4C.taxon	type_taxon	Type species. Pilumnus (Parapilumnus) quadridentatus De Man, 1895, by original designation. Other included species. Aniptumnus bijoyi sp. nov., A. nefissurus (Garth & Kim, 1983), A. neolaevis (Deb, 1995) comb. nov., and A. vietnamicus Ng & Clark, 2008.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE5FF94FF214EDFC1C4FF4C.taxon	diagnosis	Diagnosis. Adult: Carapace transversely subhexagonal, broader than long; anterolateral margins divided into 4 teeth posterior to exorbital angle, first low, lobiform, second to fourth more projecting. Male thoracic sternum moderately broad, smooth; sternite 2 clearly separated from sternite 3 by transverse suture; small section of sternite 8 sometimes exposed when pleon locked onto thoracic sternum. Chelipeds distinctly unequal. Ambulatory legs without dactylo-propodal lock; posterodistal margin of P 5 basis-ischium, with several large, closely packed spines or conical tubercles; posterior margin of P 5 merus with spines and / or conical tubercles proximally. G 1 sinuous, slender; G 2 short, about one-fifth length of G 1. Zoea I: Rostral spine much shorter than antennal protopods; antennae extremely long, much longer than carapace; furcae of telson prominently elongate, inner margin armed with spines.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE5FF94FF214EDFC1C4FF4C.taxon	discussion	Remarks. Aniptumnus Ng, 2002, has primarily been diagnosed by the condition of the fourth ambulatory leg (P 5) where the basis-ischium has a comb-like postero-distal margin due to the peculiar placement of some spines or conical tubercles, and the posterior margin of the merus is armed with spines and / or conical tubercles proximally (Ng 2002; Ng & Clark 2008). Subsequent to the establishment of the genus, Ng & Clark (2008) added two more species, A. nefissurus (Garth & Kim 1983) and A. vietnamicus Ng & Clark, 2008. Ghory et al. (2013) considered Pilumnopeus riui Takeda, 2001, described from Hainan Island, China, as a junior synonym of A. quadridentatus. Trivedi et al. (2021) reported A. quadridentatus in the Indian Ocean (West Bengal, India) by virtue of their considering it to be a senior synonym of H. neolaevis Deb, 1985. The identity of H. neolaevis is further discussed herein. Finally, Widyastuti & Rahayu (2022) recently reported a range extension for A. quadridentatus, with a new record from Timika, Papua, Indonesia. It is important to note, however, that A. nefissurus is atypical in apparently lacking the comb-like posterodistal margin on the P 5 basis-ischium, and both A. nefissurus and A. vietnamicus have the distal tip of their G 1 tapering to a point (but with varying degrees of curvature), which is typical of many pilumnines, whereas in the type species and A. bijoyi sp. nov., the distal tip of the G 1 has an apical lobe that is bluntly round and scoop-shaped. The position of A. nefissurus and A. vietnamicus in Aniptumnus will have to be evaluated further. Also, given that only A. quadridentatus has the first zoea morphology known, it will be critical to also determine the zoeal morphology of the other species of Aniptumnus as well. The systematic position of Aniptumnus within Pilumnidae / Pilumninae is also an interesting issue to resolve. It is, thus far, the only pilumnid reported to have such long antennae on the first zoea, although another pilumnid genus, Leelumnus Mendoza & Ng, 2011, also has a similar feature (Mendoza unpubl. data). In addition, both Aniptumnus and Leelumnus appear to favor estuarine areas and are a common component of fouling communities there (Mendoza & Ng 2011). Another genus, Cryptopilumnus Hsueh, Huang & Ng, 2009, has a similar granulation pattern on the P 5 basis-ischium and merus, although it can be separated from Aniptumnus by the presence of a dactylo-propodal lock on the ambulatory legs, the complete fusion of thoracic sternites 1 – 3, and the predisposition to live in holes and crevices within rocks (Hsueh et al. 2009), whereas Aniptumnus has no dactylo-propodal lock on the ambulatory legs, has fused thoracic sternites 1 and 2 but with sternite 3 separated from sternite 2 by a distinct suture, and it has not been recorded to live within rocks. Aniptumnus has also been compared to Latopilumnus Türkay & Schuhmacher, 1985, primarily due to similarities in the general form of the carapace and pereopods, although in the case of Latopilumnus, the spines and conical tubercles on the P 5 basis-ischium and merus are absent and the first zoea, which has short antennae, is typical of the family (Ng & Clark 2008). These genera will have to be included in future phylogenetic studies concerning Aniptumnus.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF94FF214E23C75EFA6E.taxon	materials_examined	Type locality. Matla River estuary, West Bengal, India.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF94FF214E23C75EFA6E.taxon	diagnosis	Diagnosis. Carapace transversely subhexagonal, broader than long; anterolateral margins divided into 4 teeth posterior to exorbital angle, first low, lobiform, second to fourth more projecting. Male thoracic sternum with small section of sternite 8 exposed when pleon locked onto thoracic sternum. Pleonal somite 2 much narrower than somite 1. Chelipeds distinctly unequal, minor chela distinctly more granulate, setose than major chela. Posterodistal margin of P 5 basis-ischium, with several closely packed conical tubercles; posteroproximal margin of P 5 merus with few large, conical tubercles proximally gradually being replaced by smaller granules distally until proximal two-thirds of merus. G 1 sinuous, distal tip bluntly round, without scoop-shaped apical lobe.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF94FF214E23C75EFA6E.taxon	discussion	Remarks. Heteropanope neolaevis was originally described by Deb (1995: 220) based on several specimens (“ 50 examples ”) collected from the Matla River, in the Gangetic Delta, West Bengal, India. The identity and taxonomy of this species has been extensively discussed by Deb (1999), Ng et al. (2018), and Trivedi et al. (2021). As Deb (1985) did not indicate a holotype, all of her specimens were syntypes. As such, to stabilize the taxonomy, Trivedi et al. (2021) designated one of the surviving male syntypes deposited in the collection of the Zoological Survey of India (ZSI) in Kolkata, India, as the lectotype and four other male syntypes as paralectotypes (ZSI-C 1503 / 2). They provided detailed photographs and drawings of the lectotype as well (Trivedi et al. 2021: figs. 7, 8, 10 G – I). They further remarked that the type material of H. neolaevis agreed well with the descriptions and figures of Aniptumnus quadridentatus (De Man, 1895), citing the similar morphology of the carapace, pereopods, and male thoracic sternite 8, and concluded that the two nominal taxa were conspecific. They also noted, however, that while the G 1 of H. neolaevis (Trivedi et al. 2021: fig. 10 G, H) closely resembled that of A. quadridentatus, the distal tip of the former is “ more rounded and less produced, but this can be explained by variation ” (Trivedi et al. 2021: 14). Comparison with the illustrations of the lectotype of A. quadridentatus and actual specimens (N = 44) from Singapore and Malaysia, however, has revealed that the G 1 of adult males always has a short but distinct, scoopshaped apical lobe that is somewhat perpendicular to the long axis (Fig. 5 F – I; cf. Ng 2002: fig. 1 F, G), and none had the morphology of H. neolaevis, wherein the well-produced, distal scoop-shaped apical lobe of the G 1 is absent (cf. Deb 1999: fig. 4; Trivedi et al. 2021: fig. 10 G – I). Furthermore, the lectotype of H. neolaevis has a narrower pleonal somite 2, and a much larger section of sternite 8 exposed (Trivedi et al. 2021: fig. 8 D) whereas in A. quadridentatus pleonal somite 2 is wider and sternite 8 is either minimally exposed (Fig. 5 E) or not at all (cf. Takeda 2001: fig. 3 E). As such, we consider H. neolaevis to be a valid species of Aniptumnus distinct from A. quadridentatus.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF9FFF214944C62CFACC.taxon	description	Figs. 1 – 4	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF9FFF214944C62CFACC.taxon	materials_examined	Type material. Holotype: male, 10.7 by 7.1 mm (CUSAT / SBN / HRJ / 1), Marine Science Boat Jetty, School of Marine Sciences, Cochin University of Science and Technology, Kochi, Kerala, SW India, 9 ° 57 ′ 50.8104 ″ N, 76 ° 16 ′ 56.7834 ″ E, coll. P. H. Praved & N. R. Hershey, 29 Sep. 2021 (fixed in 5 % formalin). Paratypes: 6 males, 8.1 by 5.9 mm to 10.5 by 7.1 mm, 4 females, 7.6 by 5.6 mm to 8.7 by 5.7 mm (CUSAT / SBN / HRJ / 2 – 11), same data as holotype; 1 male, 10.1 by 6.7 mm, 1 female, 8.2 by 5.1 mm (ZRC 2022.0137), same data as holotype; 1 male, 5.0 by 3.2 mm (ZRC 2019.1091), intertidal rocky area with algae, just outside School of Marine Sciences, Cochin University of Science and Technology, Kochi, Kerala, SW India, coll. N. R. Hershey, Feb. 2019. Comparative material. Aniptumnus quadridentatus (De Man, 1895). Singapore: 30 males, 5.5 by 3.9 mm to 13.9 by 9.5 mm, 18 females, 5.6 by 4.0 mm to 12.3 by 8.0 mm, 5 ovig. females, 8.2 by 5.8 mm to 10.4 by 7.0 mm (ZRC 2018.0639), off Pulau Ubin, floating fish farm (kelong FC 12 E), among clumps of Perna viridis attached to submerged ropes, coll. JCE Mendoza, I Iesa & MD Safaruan, 7 Nov. 2017; 1 female, 14.0 by 8.9 mm, 1 ovig. female, 10.7 by 7.3 mm (ZRC 2011. 0683), Changi Point, on Paku Buoy (fouling), coll. SLM Teo, FS Tan, SC Lee & JL Ong, 2010; 6 males, 5.8 by 4.0 mm to 11.1 by 7.4 mm, 3 females, 6.1 by 4.2 mm to 7.7 by 5.1 mm (ZRC 2021.0421), off Pulau Ubin, floating fish farm (kelong E 63 C), coll. BY Lee, 3 Sep. 2020. Malaysia: 8 males, 9.5 by 6.5 mm to 11.0 by 7.3 mm, 3 females, 8.9 by 5.6 mm to 9.7 by 6.2 mm, 2 ovig. females, 8.6 by 5.8 mm, 9.3 by 6.0 mm (ZRC 1992.10966 – 10978), Perak, Lumut, coll. H. Singh, 1989. Aniptumnus vietnamicus Ng & Clark, 2008. Vietnam: Holotype, male, 8.3 by 6.1 mm (ZRC 1970.1.10.1), Nhatrang Bay, coll. R. Serène, 10 Apr. 1966; paratype, female, 9.6 by 6.6 mm (ZRC 1970.1.10.2), same data as holotype. Type locality. India: Kochi, Kerala; Kollam-Kottapuram National Waterway No. 3, near Marine Science Boat Jetty, CUSAT-School of Marine Sciences.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF9FFF214944C62CFACC.taxon	diagnosis	Diagnosis. Carapace subhexagonal, broader than long (CW / CL: 1.4 – 1.5); dorsal surface finely granular with small patches of granules on gastric, branchial regions, sparsely setose with tufts of longer setae on front and on granular patches of gastric, branchial regions. Anterolateral, posterolateral margins subequal in length. First tooth of carapace exorbital margin lobiform, nearly flat, second tooth broadly triangular, denticulate, third and fourth teeth acutely triangular, slightly curved. Orbits without fissures, notch at junction of supra- and infraorbital margins absent. Third maxilliped merus distolateral angle produced, rounded. Chelipeds unequal; major chela external surface sparsely setose, finely granular with scattered larger granules; minor chela more densely setose, with rows of sharp conical granules. P 5 basis-ischium with conical tubercles on disto-posterior margin forming comb-like structure; merus posterior margin with sharp conical tubercles proximally. G 1 slender, sinuous, distal tip with short, narrow, scoop-shaped apical lobe. G 2 small, one-fifth length of G 1, terminal, subterminal segments uniformly curving mesially. Penis emerging from anterior tip of coxosternal condyle of P 5 coxa.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF9FFF214944C62CFACC.taxon	description	Description of male holotype. Carapace (Figs. 1, 2 A) transversely subhexagonal, broader than long (CW / CL = 1.5), dorsal regions poorly defined, dorsal surface convex longitudinally and transversely, generally smooth with patches of fine granules on gastric, branchial regions, sparsely setose, most setae aggregated around granular areas. Front moderately projecting, slightly deflexed ventrally, bilobed, lobes divided by median notch, frontal margins uniformly, moderately convex without any distinct lateral lobe or tooth, lined with small granules, with poorly defined row of long setae submarginally. Supraorbital margin concave in dorsal view, granular, without fissures; junction with infraorbital margin without notch. Anterolateral and posterolateral margins similar in length; anterolateral margin granular, sparsely setose, divided into 4 teeth; exorbital angle low, not prominently projecting, separated from first tooth by shallow concavity; first tooth wide, flat or weakly convex lobe, separated from second tooth by small notch; second tooth broadly subtriangular, posterior margin much longer than anterior, apex directed toward anterior; third, fourth teeth similar, acute, apices pointed obliquely; posterolateral margin nearly straight, finely granulate, convergent posteriorly; posterior margin slightly convex, with raised row of fine round granules. Suborbital, sub-branchial and pterygostomial regions finely granular. Orbits (Fig. 2 C) transversely ovate; eyes large, filling entire orbital cavity, ocular peduncle stout, unarmed, corneas pigmented. Infraorbital margin granular, without fissures, mesial suborbital tooth absent. Antennular fossae subrectangular, antennules robust, folding transversely and obliquely. Basal antennal article short, subrectangular, filling orbital hiatus; flagellum long, exceeding orbit. Proepistome narrow, smooth. Central region of posterior border of epistome projecting medially, notched laterally. Endostome with prominent oblique ridge on either side. Third maxillipeds (mxp 3; Fig. 2 B) finely granular externally, sparsely setose; palps thick, subcylindrical; merus subquadrate, distolateral angle projecting, rounded; ischium subrectangular, about 1.5 times as long as merus, with submedian sulcus, submarginal row of long setae mesially; basis subtriangular, separated from ischium by clear suture; exopod elongate, not tapering, distal tip reaching to three-quarters lateral margin of merus, flagellum long, well developed. Male thoracic sternum (Fig. 2 D, E) moderately broad, surface smooth, punctate. Sternites 1, 2 completely fused, forming broad triangle abutting ischia of third maxillipeds. Sternites 2, 3 separated by distinct suture (suture 2 / 3); sternites 3, 4 almost completely fused, demarcated only laterally by notches that continue mesially as shallow groove. Sutures 4 / 5, 5 / 6 interrupted medially. Sutures 6 / 7, 7 / 8 complete. Sternopleonal cavity (Fig. 2 E) deep, anterior tip on sternite 4, just reaching imaginary transverse line formed by coxo-sternal condyles of P 1 coxae. Tubercle of sternopleonal lock (press-button) anteriorly positioned on sternite 5, closer to suture with sternite 4. Median line complete at level of sternite 7, 8 only; short portion visible on posterior part of sternite 4 within sternopleonal cavity. Small, median, triangular decalcified area at junction between sternite 6, 7. Sternite 8 (Fig. 2 G) not visible when pleon folded against thoracic sternum, positioned beneath the posterolateral corner of the pleonal somite 2. Chelipeds (P 1; Figs. 1, 3 A, B) distinctly heterochelous. Anterior margins of basis-ischium, merus lined with subconical granules. External surface of the carpus granular, inner distal angle projecting as sharp tooth bordered with large tubercles. External surface of major palm (Fig. 3 B) finely granular, with rows of larger, widely spaced granules near upper margin, also proximally. Fingers thick, noticeably shorter than palm, cutting margins with small teeth; fixed finger with short longitudinal groove on the external surface; dactylus longer, slightly curved, upper margin granular proximally. Minor palm (Fig. 3 A) smaller, external surface setose, with large, sharp, conical granules, sometimes arranged in rows along long axis of palm; fingers slender, shorter than palm, cutting margins minutely dentate; fixed finger slightly deflexed; dactylus with longitudinal ridge on external surface, upper margin granular proximally. Ambulatory legs (P 2 – P 5; Figs. 1, 3 C) moderately long, slender, setose, broad surfaces finely granular; P 3 longest, merus-to-dactylus length 1.02 times CW), P 5 shortest, merus-to-dactylus length 0.78 times CW. Basisischium unarmed except in P 5, where posterodistal margin armed with prominent spines, forming comb-like structure. Merus subrectangular, elongate, anterior margin slightly convex, finely granulate, subdistal tooth present but effaced, posterior margin, slightly convex, finely granulate except in P 5, which has several spines and conical tubercles proximally, decreasing in size until mid-length of merus. Carpus subrectangular, widening distally, anterior margin convex, posterior margin straight, dorsal surface without granular ridge. Propodus subrectangular, anterior, posterior margins slightly convex. Dactylus mostly straight, tapering distally, ending in curved chitinous claw; dactylo-propodal lock absent. Male pleon (Fig. 2 D, F, G) smooth, with all somites and telson freely articulating, lateral margin concave. Somite 1 much wider than long, anterior margin deeply concave, posterior margin slightly sinuous, lateral margins straight. Somite 2 much wider than long, anterior margin slightly concave, posterior margin strongly convex, lateral margins rounded. Somite 3 widest, transversely subhexagonal, lateral margins strongly convex. Somites 4 – 6 trapezoidal, gradually decreasing in width, with somite 6 narrowest, lateral margins straight to slightly concave. Telson triangular with rounded apex, lateral margins convex, median length 0.83 times basal width. G 1 (Fig. 3 D, E, G, H) moderately long, slender, sinuous; margins lined with simple or plumose setae; distal part curved, ending in small scoop-shaped apical lobe, with stiff, simple spiniform subdistal setae; G 2 (Fig. 3 F) short, about one-fifth length of G 1, terminal, subterminal segments uniformly curving mesially. Penis (Fig. 2 E) emerging from anterior portion of coxosternal condyle of P 5 coxa. Female. The female morphology is similar to the male in all non-sexual aspects. Thoracic sternum relatively wider than that of male. Pleon (Fig. 4 A) much wider, lateral margin convex, setose, external surfaces smooth; somites and pleon freely articulating; somite 3 widest. Telson large, subtriangular, apex rounded, lateral margins convex, median length subequal to basal width. Vulvae (Fig. 4 B) mesially on thoracic sternite 6, abutting suture with sternite 5; semicircular in outline. Live colouration. Fresh specimens are usually covered with a copious amount of sediment and debris when collected such that they need to be gently brushed clean to expose the natural coloration (Fig. 1). The dorsal carapace and pereopods are generally tan or pale brown with patches and scattered spots of dark purple that are sometimes surrounded by lighter areas of purple. The setae covering them are similarly tan. The thoracic sternum, pleon, and ventral and internal surfaces are creamy white. The fingers of the chelae are brown with pale tips.	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF9FFF214944C62CFACC.taxon	etymology	Etymology. The species is named in honor of Prof. S. Bijoy Nandan, Dean, Faculty of Marine Sciences, Cochin University of Science and Technology, for his significant contributions in the field of marine biology and wetland ecology in India. Habitat and ecology. Aniptumnus bijoyi sp. nov. was found in larger numbers on encrusting communities such as clumps of mussels (Mytella strigata and Perna virdis), submerged structures, and roots of decaying aquatic weeds (Eichornia crasipes) in the School of Marine Science Jetty. Their occurrence was also noticed in the other parts of the Cochin Estuary. The species is usually abundant during the pre-monsoon period (January – May) in sites with a salinity range of 10 – 28 ppt and depth range of 1 – 6 m. The habitat is muddy to clayey-silty, the pH of the sediment generally ranged from 7.5 – 8, and the average redox potential from - 135 to - 275 mV. The total carbon of the sediment ranged from 30 – 35 g / kg and the total organic carbon ranged from 1.9 – 2.5 %. Aniptumnus bijoyi sp. nov. was observed to occur together with the brachyuran crabs Thranita crenata (Rüppell, 1830), Neorhynchoplax alcocki (Kemp, 1917), and N. demeloi (Kemp, 1917).	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
03E48797FFE6FF9FFF214944C62CFACC.taxon	discussion	Remarks. Aniptumnus bijoyi sp. nov. is morphologically most similar to A. quadridentatus (De Man, 1895) and A. neolaevis (Deb, 1985) comb. nov. in the general form of the carapace, thoracic sternum, male pleon, and pereopods (cf. Ng 2002; Trivedi et al. 2021; Widyastuti & Rahayu 2022). The new species is easily distinguished from A. nefissurus and A. vietnamicus by the general form of the carapace, by its less granular major chela and by the G 1 morphology (cf. Garth & Kim 1983: fig. 11; Ng & Clark 2008: figs. 19 – 21). Aniptumnus bijoyi sp. nov. can most easily be distinguished from A. neolaevis in the following features: the tuberculation on the posteroproximal margin of the P 5 merus is much more prominent, with the conical tubercles being larger (Fig. 3 C) (vs. tuberculation, while present, is much reduced, tubercles are smaller in A. neolaevis, cf. Trivedi et al. 2021: fig. 8 A); pleonal somite 2 of the male is considerably wider, and consequently, thoracic sternite 8 is completely concealed when the pleon is folded and locked onto the thoracic sternum (Fig. 2 E) (vs. pleonal somite 2 much narrower, thoracic sternite 8 exposed in A. neolaevis; cf. Trivedi et al. 2021: fig. 8 D); and the distal tip of the G 1 has a short, projecting, scoop-shaped apical lobe (Fig. 3 D, E, G, H) (vs. apical lobe absent in H. neolaevis; cf. Deb 1999: fig. 4; Trivedi et al. 2021: fig. 10 G, H). Aniptumnus bijoyi sp. nov. differs from A. quadridentatus in the following morphological features: the carapace outline is generally wider and shorter, with anterolateral and posterolateral margins nearly identical in length, and the posterolateral margins slightly convex (Fig. 2 A) (vs. carapace outline narrower and longer, with posterolateral margins longer than anterolateral margins, and posterolateral margins straight in A. quadridentatus; Fig. 5 A; cf. De Man 1897: pl. 13 fig. 6; Takeda 2001: figs. 2, 3 A; Widyastuti & Rahayu 2022: fig. 3 A); on the carapace anterolateral margin, the first tooth after the exorbital angle is much reduced, more like a flat lobe (Fig. 2 A) (vs. first tooth lobiform and low but with distinct, denticulate convex protuberance in A. quadridentatus; Fig. 5 A; cf. Takeda 2001: fig. 3 B; Ng 2002: fig. 1 A); the dorsal surface of the carapace is widely convex transversely in the frontal view (Fig. 2 C) (vs. dorsal surface more acutely convex transversely in frontal view in A. quadridentatus; Fig. 5 C; cf. Widyastuti & Rahayu 2022: fig. 3 B); there is no notch on the orbit at the junction of the supraorbital and infraorbital margins (Fig. 2 C) (vs. present in A. quadridentatus; Fig. 5 C); the distolateral angle of mxp 3 merus is more rounded and projecting (Fig. 2 B) (vs. less projecting in A. quadridentatus; Fig. 5 B; cf. Ng 2002: fig. 1 B; Widyastuti & Rahayu 2022: fig. 3 C); in the male, thoracic sternite 8 not visible when the pleon is folded and locked onto the thoracic sternum (Fig. 2 G) (vs. sternite 8, sometimes exposed, small but distinct, positioned in between the lateral margin of the pleonal somite 2 and the mesial margin of the coxo-sternal condyle of the P 5 coxa in A. quadridentatus; Fig. 5 E); and the distal tip of the G 1 has a narrower apical lobe (Fig. 3 D, E, G, H) (vs. apical lobe wider in A. quadridentatus; Fig. 5 F – I; cf. Takeda 2001: fig. 3 H, I; Ng 2002: fig. 1 F, G).	en	Hari, P. Praved, Hershey, N. Regina, Mendoza, Jose Christopher E. (2022): On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala. Zootaxa 5214 (2): 261-272, DOI: https://doi.org/10.11646/zootaxa.5214.2.6
