taxonID	type	description	language	source
03C087D47F36FFB3FF7EE1C2168D7EE7.taxon	materials_examined	Material examined. Phjŏngjang, 18 July 1981, 32 males; Sokam, 8 July 1981, 1 male; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F36FFB3FF7EE1C2168D7EE7.taxon	discussion	Remarks. A set of subtle morphological differences used in diagnostics, often overlapped by intraspecific variations, make the determination of adult males of Cladotanytarsus one of the most difficult within Tanytarsini and Chironomidae. Accurately mounted and illustrated specimens, their precise measurements and appropriate interpretation of diagnostic features are thus decisive. In fact, determination of Cladotanytarsus sinjongensis may be a challenge when based on the illustrated original description, although several crucial characters indicate that the specimens presently examined are conspecific with those described from South Korea (Ree & Kim 1988). The males from North and South Korea compared here [data on the South Korean material, with remarks given in square brackets] are approximately of the same size, with the wing 1.18 – 1.45 mm long [1.2 – 1.3 mm], AR 0.90 – 1.03 [0.91 – 1.05], frontal tubercles 14 – 18 μm long [15 – 16 μm], similar lengths of palpomeres, leg ratios and the same body colouration. The hypopygial anal point is more or less lanceolate or tongue-shaped (Fig. 2 A, B), 35 – 45 μm long [7 – 9 μm, inconsistency relative to the scaled fig. 4], the superior volsella is slender, narrowed at mid length, with the swollen and apically rounded distal part (Fig. 2 A, C) [probably distorted during mounting], the stem of the median volsella is slightly curved, ~ 40 μm long [35 – 38 μm], with branched lamellae (Fig. 2 A, D), as in all known Cladotanytarsus [the deep furcations overlooked].	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F36FFB0FF7EE07E102D7B4B.taxon	description	The genus Cladotanytarsus is divided into two subgenera: Cladotanytarsus s. str. Kieffer, 1921 and Lenziella Kieffer, 1922 (Giłka 2011 b, Giłka & Spies 2012, Puchalski & Giłka 2017). Since no character typical of Lenziella (a globular basal swelling in the inferior volsella nor lobes of the mid and hind leg tibiae apices) has been found in C. sinjongensis males, we place the species in the subgenus Cladotanytarsus s. str. The genus is represented by nearly 80 species known from all continents except Antarctica (Puchalski et al. 2018). From North Korea, apart from C. sinjongensis, now found and redescribed below, two Cladotanytarsus species have so far been recorded by Reiss (1980), but determined to the genus level, as Cladotanytarsus sp. K 1 and K 2.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F36FFB0FF7EE7EE16117DF3.taxon	description	Subtribe: Tanytarsina Zavřel, 1917	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F35FFB2FF7EE2C115DC7FB3.taxon	materials_examined	Material examined. Kŭmgang-san, 28 June – 2 July 1981, 1 male; Phjŏngjang, 18 July 1981, 3 males; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F35FFB2FF7EE2C115DC7FB3.taxon	discussion	Remarks. Micropsectra atrofasciata is an eurytopic and one of the most common species within the genus and the tribe in Europe (Giłka 2011 c), distributed across the Palaearctic region. The wide geographical range and distinct intraspecific variations in the body size and colouration, depending also on a generation (seasonal heteromorphism), as well as in the shape of some key structures of the hypopygium (cf. Giłka 2000, 2001 a, 2011 a; see also remarks to Micropsectra joganplumosa and Tanytarsus volgensis below), were probably the reason for multiple descriptions of this species under different names. A redefinition of M. atrofasciata by Stur and Ekrem (2006) includes eight junior synonyms. Here, we analyzed another name, Micropsectra tonewdeea Sasa et Tanaka, 2002, based on description of the adult male and photographs of the holotype displayed on the website of the NMNS (405: 070). A comparison of diagnostic characters shows that the type specimen described for this name falls within the wide range of morphological variability of M. atrofasciata. As far as we know, the name, treated as synonymous here, has not been used apart from the original description based on materials from locus typicus in Gunma Prefecture, Honshu (Sasa & Tanaka 2002).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F35FFB3FF7EE1F310BD784D.taxon	description	One of the largest genera in the tribe Tanytarsini. Nearly 150 species names have been ascribed to Micropsectra, although many of them still require a redefinition or generic transfer. The genus is divided into several species groups. In the material analysed we found three species of the atrofasciata - and notescens groups.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F34FFB2FF7EE59110787BBF.taxon	materials_examined	Material examined. Myohyang-san, 22 – 25 June 1981, 1 male, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F34FFB2FF7EE59110787BBF.taxon	discussion	Remarks. As in the previous species, males of Micropsectra joganplumosa are variable in the body colouration and main metric characters that probably resulted in description of the species under a series of names, i. e. Micropsectra tusimalemea, M. tusimamenea and M. tusimaquerea. The specimens analyzed (Sasa & Suzuki 1999) were originally found as belonging to the species closely related to each other and / or to several further Micropsectra described in the same paper. However, the slight differences (scutum colouration, AR values, and length of digitus) are here defined as intraspecific variations. In fact, the drawings presented in original descriptions for all these names do not reflect shapes of the most important diagnostic structures, especially when compared with photographs of their type specimens (cf. Sasa & Okazawa 1991, fig. 1.1 a – j + NMNS 217: 029 and Sasa & Suzuki 1999, figs 18, 19 and 23 + NMNS 372: 016, 372: 018, 354: 069). Some drawings presented under these names indicate a similar or identical shape of the same hypopygial structures (median and inferior volsellae); on the other hand, the same structure (anal point) nearly identical on photographs differs distinctly on the drawings. In case of such inconsistency, we assume that at least part of the drawings may have been mistakenly assigned to a specimen / name, thus the comparison of the diagnostic structures can only be based on photographs of the designated types. Despite of deformations of the holotypes on slides (hypopygia) some crucial characters indicate that they are conspecific. The characters best defining the species are: the shape of the anal point (Fig. 3 A, B), the digitus and superior volsella, typical of the Micropsectra notescens species group (Fig. 3 A, C), and the median volsella in shape of a hockey stick bearing numerous spoon-shaped lamellae (Fig. 3 D). The species is known from the west of Japan: Toyama Prefecture in Honshu (as M. joganplumosa) and Tsushima Island, where specimens were collected at one locality, in the same time, using the same method (as M. tusimalemea, M. tusimamenea and M. tusimaquerea). Now Micropsectra joganplumosa is also recorded from central part of North Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F34FFB5FF7EE18D10A57F5F.taxon	materials_examined	Material examined. Kŭmgang-san, 28 June – 2 July 1981, 3 males, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F34FFB5FF7EE18D10A57F5F.taxon	discussion	Remarks. The enormity of names coming from Japan pertain also to a series of Micropsectra (Sasa 1984, 1993; Sasa & Okazawa 1994; Sasa & Suzuki 1999; Sasa et al. 2001) of which specimens described under six names at least have to be compared with M. jokatertia. They are: M. chuzelonga Sasa, 1984, M. inamenea Sasa, Kitami et Suzuki, 2001, M. inaneoa Sasa, Kitami et Suzuki, 2001, M. johanaprima Sasa et Okazawa, 1994, M. tomoprima Sasa, 1993, and M. tusimaneoa Sasa et Suzuki, 1999. Based on more or less distinct characters given in original descriptions and observable on photographs of type specimens (cf. NMNS 051: 011, 398: 015, 402: 023, 250: 078, 244: 001, 356: 017, respectively to the above list of names), M. jokatertia (NMNS 352: 051) differs from them in having a slender gonostylus (Fig. 4 A vs. M. chuzelonga, M. inamenea), a longer and / or more slender anal point and its crest (Fig. 4 A vs. M. chuzelonga, M. inaneoa, M. johanaprima, M. tomoprima, M. tusimaneoa), a longer or more slender digitus (Figs 4 A & 5 G vs. M. chuzelonga, M. johanaprima, M. tusimaneoa), and a lower number of spoon-shaped lamellae of the median volsella (Figs 4 B & 5 C vs. M. inamenea). Consequently, we refrain from synonymization of M. jokatertia, although synonymy within the compared names (excl. M. jokatertia) is highly probable. Micropsectra jokatertia fits well the concept of the notescens group analysed on the basis of key structures of the hypopygium (Giłka 2001 b). When M. jokatertia is included to a comparison of several well-defined species, it places between M. rilensis Giłka, 2001 and M. recurvata Goetghebuer, 1928 in this series (Fig. 5).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE61E17277DF1.taxon	materials_examined	Material examined. Myohyang-san, 22 – 25 June 1981, 2 males, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE61E17277DF1.taxon	discussion	Remarks. An illustrated diagnostic description of the adult male, based on the material examined here, provided Giłka (2012). The species has been recently analysed also for intraspecific variations (Orel 2021), however, with data referred to the present material erroneously given to be from South Korea. To our knowledge, Neozavrelia fengchengensis has so far been recorded from the Northeast China (Liaoning Province), East Siberia (Chita Region), several other sites in the Russian Far East, and from North Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE5EF156E7C2B.taxon	description	Over 30 described species belong to Neozavrelia, although many of them still remain in wrong combinations, while some names require revision as potential synonyms. Two species have so far been recorded from North Korea (Giłka 2012).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE05017037B25.taxon	materials_examined	Material examined. Kŭmgang-san, 28 June – 2 July 1981, 1 male, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE05017037B25.taxon	discussion	Remarks. An illustrated redescription of the species with notes on systematics and possible synonymy have been based on the presently examined material (Giłka 2012).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE2E7169979F0.taxon	discussion	Remarks. The two specific names, including Paratanytarsus confusus, were alternatively mentioned by Reiss (1980), thus we assume that at least one species has been found in North Korea. Later synonymization of P. confusus with P. dissimilis enabled a clear definition of this species (Langton et al. 1988). Diagnostic characters for adult males of the two sibling species, P. dissimilis and P. inopertus, compared Giłka (2011 a) in the couplet 6 of the key for Paratanytarsus. Though both are eurytopic and widespread in the Holarctic (Giłka 2011 c), none has been confirmed in the presently examined material.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE1E0175F7831.taxon	description	Nearly 70 described species belong to Paratanytarsus, including several species still being incorrectly ascribed to other genera. At least three species have been recorded in North Korea; apart from the below species, Reiss (1980) recorded one more, determined as Paratanytarsus sp. K 1.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB9FF7EE13011377829.taxon	discussion	Remarks. The adult male of Paratanytarsus laccophilus is clearly defined and easily separable even from its closest relative, P. paralaccophilus Giłka et Paasivirta, 2008 known from Lapland (Giłka & Paasivirta 2008). In North Korea, the species was recorded by Reiss (1980) but not confirmed in the material presently analyzed.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB9FF7EE200164A7911.taxon	materials_examined	The genus includes nearly 100 species distributed worldwide. Some names, especially those coming from east and south of Asia, still require a revision (potential synonyms, incorrect combinations). In the material presently examined we found one species.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB8FF7EE30511737FB3.taxon	materials_examined	Material examined. Myohyang-san, 22 – 25 June 1981, 3 males, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB8FF7EE30511737FB3.taxon	discussion	Remarks. Neither the original description, based on simple schematic drawings, nor the photographs of the Rheotanytarsus tusimatfegeus holotype, not very successfully slide-mounted (NMNS 371: 001), allow for an unambiguous definition of this species when each of the characters' source taken separately, but both in combination indicate the species analyzed here. North Korean males are slightly smaller and have a proportionally lower AR than those from Japan (wing length 1.60 – 1.70 vs 1.80 – 1.96, AR ~ 0.65 vs 0.76 – 0.88), while other characters are conspecific. Rh. tusimatfegeus has so far been reported from the Sea of Japan region, in Tsushima (Sasa & Suzuki 1999) and most likely in Russia (Makarchenko et al. 2005). Now recorded for the first time in North Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE77A117B7A8B.taxon	materials_examined	Material examined. Phjŏngjang, 18 July 1981, 1 male, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE77A117B7A8B.taxon	discussion	Remarks. Reiss and Fittkau (1971) indicated this species as representative of the chinyensis group. The concept was recently refuted, and the former chinyensis group was proposed to be split into several unrelated clusters / groups (Lin et al. 2018 a, b). Alternatively, the group was postulated to be reduced. However, Tanytarsus chinyensis was not included in the analyses (op. cit.), thus according to this concept, the group name-bearing species is unplaced in any of the species groups proposed, apparently, except for the chinyensis group (see also remarks to T. tamadecimus below). In the material analysed, we found a single male collected in the Botanical Garden in Phjŏngjang.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE08D17E57BF3.taxon	discussion	Remarks. Tanytarsus heusdensis was ascribed to the chinyensis species group (Reiss & Fittkau 1971); recently proposed to be excluded into the separate heusdensis complex / group (Lin et al. 2018 a, b). The species was recorded from North Korea by Reiss (1980), but not confirmed in our material.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFBBFF7EE25016367EE7.taxon	materials_examined	Material examined. Kŭmgang-san, 28 June – 2 July 1981, 1 male; Myohyang-san, 22 – 25 June 1981, 2 males; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFBBFF7EE25016367EE7.taxon	discussion	Remarks. The North Korean specimens fit well the original description of Tanytarsus iriolemeus (Sasa & Suzuki 2000) except for one character crucial in species group concepts for Tanytarsus - the strong bristle-like median setae placed in a roundish field of the hypopygial anal tergite. This structure is omitted in the original description but present in the holotype specimen (slightly beyond the range of sharpness on the photo presented at NMNS website, 385: 035). This character combined with the anal point elongate, bearing a few spinulae (Fig. 6 A, B; cf. also Tanytarsus innarensis Brundin 1947), the superior volsella roundish, medially excavated, with a well-protruded posteromedian corner, the digitus long apically rounded (Fig. 6 C), and the shape of the median volsella (Fig. 6 D) may indicate that T. iriolemeus belongs to the verralli species group. In the analyses by Lin et al. (2018 b), T. iriolemeus falls in the tamakutibasi cluster that together with the aterrimus + chinyensis + curticornis groups is sister to a clade comprising the Holarctic excavatus, recurvatus, and verralli groups.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE59B103A7CD2.taxon	description	With nearly 400 described species, Tanytarsus is the largest genus in the tribe and the second most diverse genus of the family Chironomidae (cf. Dantas et al. 2022). At least 11 species of Tanytarsus have been recorded from North Korea. The review comprises 3 species found previously (Reiss 1980, apart form 6 species ascribed to Tanytarsus as “ sp. K 1 – 6 ”) and 8 species here recorded for the first time from North Korea (Table 2).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3DFFBBFF7EE52516407A2A.taxon	materials_examined	Material examined. Phjŏngjang, 13 June 1981, 2 males; Sokam, 8 July 1981, 3 males; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3DFFBBFF7EE52516407A2A.taxon	discussion	Remarks. A diagnostic character of the Tanytarsus occultus male is the hypopygial anal point, usually with a square or concave apex, which the shape, however, may be affected by strong intraspecific variations. T. occultus was described on the basis of a males’ series by Brundin (1949), who referred the description to his illustration of a specimen’s hypopygium presented two years earlier [Brundin 1947, originally erroneously as Tanytarsus holochlorus (= T. mendax Kieffer, 1925)]. The illustration was apparently based on a specimen with a hypopygial anal point of a structure rarely observed in T. occultus. That choice could lead to confusion until Reiss & Fittkau (1971) redescribed the male on the basis of Brundin’s series, indicating variations in the shape of the anal point, thus stabilizing the status of the species. Recently, one more close species was described, Tanytarsus latens Giłka, Paasivirta, Gadawski et Grabowski, 2018, in which the anal point is similar or nearly identical (variable) to that illustrated from the atypical specimen of T. occultus by Brundin (1947). In case of these two species, the anal point structure can be thus critically misleading in their delimitation. The characters best separating males of the two species is the shape / length of the superior volsella and the digitus, and the arrangement of anal tergite bands (cf. Giłka et al. 2018). T. occultus is one of the best-defined species in the genus in term of morphological variability and diagnostics supported by molecular analyses (Reiss & Fittkau 1971; Ekrem et al. 2003, Ekrem 2004; Giłka & Paasivirta 2007, Giłka et al. 2018). Only now recorded from North Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3DFFBBFF7EE01E16FD790D.taxon	materials_examined	Material examined. Kaesŏng, 16 July 1981, 4 males; Sokam, 8 July 1981, 3 males; leg. W. Krzemiński. Comparative material. Tanytarsus smolandicus: Sweden, Nedre Dalälven, 11 August 2003, 1 male, Storsjön, 18 July 2003, 20 males; leg. W. Giłka.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3DFFBBFF7EE01E16FD790D.taxon	discussion	Remarks. Ekrem (2002) redescribed the male of Tanytarsus oscillans based on the holotype specimen. In remarks on its close relatives, T. smolandicus Brundin, 1947 (terra typica in Sweden) and T. unagiseptimus Sasa, 1985 (Japan), selected diagnostic characters were indicated to separate these species. It seems, however, that these features (number of median setae, length of frontal tubercles, AR value, presence / absence of microtrichia between anal point crests and slight differences in shape of inferior volsella), in part at least, may be within the morphological variability of T. oscillans and / or T. smolandicus. According to the concept by Lin et. al. (2018 b), T. oscillans and T. unagiseptimus are sister species (not ascribed to any group analyzed) while T. smolandicus has not been included. Interestingly, a concept of possible synonymy between the abovementioned names has been raised recently, when DNA sequences from specimens sampled in Fennoscandia and Japan, determined as T. oscillans or T. unagiseptimus, have been compared that resulted in ~ 97 – 98 % of DNA sequence compatibility (Paasivirta, pers. comm.). Unfortunately, our specimens collected decades ago (kept in denatured ethanol or slide-mounted in Canada balsam) do not give a chance to be analyzed molecularly for supporting this concept.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3CFFBAFF7EE23211DC79B7.taxon	materials_examined	Material examined. Kaesŏng, 16 July 1981, 1 male; Phjŏngjang, 13 June 1981, 7 males, 18 July 1981, 1 male; Sariwŏn, 18 June 1981, 3 males; Sokam, 8 July 1981, 2 males; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3CFFBAFF7EE23211DC79B7.taxon	discussion	Remarks. Tanytarsus oyamai is probably one of the most common Tanytarsini in the southern part of the Sea of Japan basin; also frequent in North Korea - collected from half of the explored sites. The adult male is well-defined, thus only several illustrations are attached to show variations of the most important diagnostic structures (Fig. 6 E – H).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE4F217B57F60.taxon	materials_examined	Material examined. Phjŏngjang, 18 July 1981, 2 males, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE4F217B57F60.taxon	discussion	Remarks. Tanytarsus pallidicornis is one of the best-defined in the genus. This common eurytopic species is widely distributed in the Holarctic but only now recorded from North Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE5B517197D59.taxon	materials_examined	Material examined. Phjŏngjang, 18 July 1981, 9 males; Sokam, 8 July 1981, 2 males; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE5B517197D59.taxon	discussion	Remarks. A group membership of Tanytarsus takahashii, based on morphology, was discussed by Ekrem (2002) who indicated several possible placements for this species (the mendax, eminulus or lestagei group). Later, molecular analyses supported the concept of the lestagei group as the place for T. takahashii (Lin et al. 2018 b). The species has been also compared with the fossil species T. serafini Giłka, 2010, and suggested to belong to the serafini group (cf. Giłka 2010, pl. II, figs. 2 – 7 and Fig. 7 A – C).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE7FC14B778DB.taxon	materials_examined	Material examined. Hwangju, 18 – 19 June 1981, 2 males; Kŭmgang-san, 28 June – 2 July 1981, 3 males; Sokam, 8 July 1981, 1 male; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE7FC14B778DB.taxon	discussion	Remarks. The Korean specimens of Tanytarsus tamadecimus fit the original description, except for a couple of characters, possibly taken from specimens deformed before their illustrating. According to Sasa (1980), the digitus is “ composed of a long spatulated dorsal process and a ventral hook-like process connected with each other ”. In fact, at the Sasa’s illustrations the hook-like process looks so unusually (not known from any other species) as we assume it could be misidentified with a long darkly pigmented tubercle at the digitus base that was omitted in the original illustrations (cf. Sasa 1980, fig. 29 B, F, G and Fig. 8 B). Character / shape overinterpretation seems to pertain also to the anal point and median volsella (cf. Sasa 1980, fig. 29 D, E and Fig. 8 A, C), thus we attach illustrations of selected variations to supplement the description. Tanytarsus tamadecimus and T. chinyensis are treated here as close relatives (see also remarks to T. chinyensis). The concept of the chinyensis species group sensu Reiss and Fittkau (1971) was recently refuted and / or its species composition was suggested to be reduced (Lin et al. 2018 a, b); however, neither T. chinyensis nor T. tamadecimus have been included in the analyses. Consequently, we keep the chinyensis group, with the two species and T. cretensis Reiss, 1987, plus T. tamagotoi Sasa, 1983 and T. simantoseteus Sasa, Suzuki et Sakai, 1998, as it was suggested (op. cit.), plus a series of species by Sasa and co-authors: T. chuzesecundus Sasa, 1984, T. inawaijeus Sasa, Kitami et Suzuki, 2000, T. oyaberotundus Sasa, Kawai et Ueno, 1988 and T. tusimatkeleus Sasa et Suzuki, 1999. Males of all the species have the broad thumb-like or spatulate digitus and the horn-like posteromedian corner of the superior volsella.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBEFF7EE37D15657F5F.taxon	materials_examined	Material examined. Kŭmgang-san, 28 June – 2 July 1981, 3 males, leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBEFF7EE37D15657F5F.taxon	discussion	Remarks. Ekrem (2002) listed characters found as crucial in delimitation of Tanytarsus tamaundecimus on the basis of the original description. However, the type material has not been examined, neither photographs of the holotype are displayed at the NMNS website. The examined Korean specimens have been preliminarily confirmed as T. tamaundecimus (Ekrem, pers. comm.), thus several key structures and their variations are supplemented here (Fig. 8 D – F).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFBEFF7EE06F16747B3F.taxon	description	So far, over 20 specific names have been ascribed to Stempellina as the genus, subgenus of Tanytarsus or to other Tanytarsini generic names. According to current knowledge, the number of known species is lower due to systematic combinations other than original or doubtful status of some names (nomina dubia). Stempellina includes 15 described species: 2 extinct and 13 extant (Giłka 2005, Zakrzewska et al. 2020), thus its share in the tribe’s world extant fauna is less than 2 %. Stempellina are recorded in all continents except Antarctica, mostly in the Holarctic. In North Korea, one unknown species has been found.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFBEFF7EE5F615057D6F.taxon	materials_examined	Material examined. Hwangju, 18 – 19 June 1981, 7 males; Kaesŏng, 16 July 1981, 1 male; Phjŏngjang, 11 June 1981, 27 males; 13 June 1981, 28 males; 18 July 1981, 26 males; Sariwŏn, 18 June 1981, 5 males; Sokam, 8 July 1981, 29 males; leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFBEFF7EE5F615057D6F.taxon	discussion	Remarks. This eurytopic and widespread Holarctic species (cf. Giłka 2011 c) was found as the most frequent and abundant tanytarsine in North Korea - recorded from two sites by Reiss (1980, as T. fimbriatus) and now confirmed at further 5 localities (123 males = over 55 % of all examined specimens). Interestingly, an extremely wide range of the body size and colouration have been observed in the examined males, among which the biggest specimens, darker are nearly twice as long as the smallest ones, lighter (see also remarks to Micropsectra atrofasciata and M. joganplumosa).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFA0FF7EE117164A7B92.taxon	description	(Fig. 9 A – E)	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFA0FF7EE117164A7B92.taxon	materials_examined	Type material. Holotype, adult male: North Korea, Paektu Mountain, Samdžijŏn at Samji Lake (41 ° 50 ' N 128 ° 20 ' E; ~ 1400 m a. s. l.), 23 – 25.08.1992, sweep net, leg. A. Palaczyk.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFA0FF7EE117164A7B92.taxon	etymology	Derivatio nominis. The specific epithet commemorates Oktawiusz Wincenty Radoszkowski (1820 – 1895), the meritorious entomologist and the first Polish discoverer of insects from Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFA0FF7EE117164A7B92.taxon	diagnosis	Diagnosis. Antenna with 5 – 6 flagellomeres, borders between flagellomeres 5 – 9 indistinct, ultimate segments completely fused, plume consisted of short sparse setae. Frontal tubercles robust. Hypopygial median setae absent. Anal point short and apically swollen, anal point crests thin, spinulae absent. Superior volsella extensive, apex round, with small apical protuberance. Stem of median volsella straight, with setiform and long spindle – shaped lamellae. Description. Adult male (n = 1). Body size. Minute species, total length ~ 2 mm. Wing length 1010 μm. Colouration (in alcohol). Eyes, scutal stripes, postnotum and sternum light brown, remaining body parts pale yellowish, wing membrane transparent, veins a bit darker. Head. Eyes bare, reniform, broadly separated by frons. Antenna only with 5 – 6 well discernible flagellomeres, borders between flagellomeres 5 – 9 indistinct, ultimate segments completely fused, AR ~ 2.6 (when flagellum measured as 6 - segmented), ~ 2.0 (as 7 - segmented), ~ 1.3 (as 9 - segmented); plume weak, consisted of short and sparse setae (Fig. 9 A). Frontal tubercles robust, ~ 30 μm long and ~ 10 μm wide at base, conical, with apex rounded. Length of palpomeres 2 – 5 (μm): 32, 66, 72, 117. Clypeus with 16 setae. Thorax chaetotaxy. Setae weak, poorly observable; Ac not observed, likely absent, Dc 7 – 8, Pa 1, Scts 2 (pair of setae placed medially). Wing. Squama bare, anal lobe reduced, cell m 3 + 4, distal part of cells m 1 + 2 and r 4 + 5 with macrotrichia, cell r 4 + 5 with a single row of macrotrichia in 3 / 4 distal part; veins C, R, R 1, R 4 + 5 (distal 1 / 4 – 1 / 2 part), M 1 + 2 (distal 3 / 4 part), Cu 1 and false vein above M 3 + 4 always with macrotrichia; An and false vein under Cu 1 (distal part) usually with some macrotrichia, other veins bare; M 3 + 4 ending distinctly distal of R 4 + 5. Legs. Fore leg tibia with s-shaped spur (~ 5 μm long). Combs of mid and hind leg tibiae fine, each bearing spur: straight and equally long (~ 20 μm) on mid leg, and unequal on hind leg - longer spur (~ 40 μm) slender and evenly curved, shorter spur (~ 16 μm) straight; ta 1 of p 2 bearing 2 hook – shaped sensilla chaetica. For length of legs segments and legs ratios see Table 1. Hypopygium. Gonostylus (~ 75 μm) as long as gonocoxite. Anal tergite with bands V-shaped broadly separated and 2 – 3 posterolateral setae, median setae absent. The tergite tapering towards short and apically slightly swollen anal point, anal point crests thin, spinulae absent (Fig. 9 B, C). Superior volsella extensive, broadest at base, tapering towards round apex bearing small apical protuberance; 3 setae on anteromedian margin (Fig. 9 B, D, E). Stem of median volsella straight, ~ 30 μm long, with setiform and long spindle – shaped lamellae reaching tip of inferior volsella (Fig. 9 B, E). Inferior volsella slightly curved, parallel-sided, with distal part rounded (Fig. 9 B, E).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFA0FF7EE117164A7B92.taxon	discussion	Remarks. Except for a combination of hypopygial characters given in the diagnosis, the adult male of Stempellina radoszkowskii differs from all known Stempellina by lower number of antennal flagellomeres. This character was recently analysed in extant and fossil species of the genus (Zakrzewska et al. 2020), as well as in Stempellinella (Zakrzewska & Jankowska 2021). A distinct trend towards reduction of the number of flagellomeres (12 or less) in extant species relative to those of fossil is confirmed here.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F26FFA0FF7EE1BB114B787B.taxon	description	The small genus Zavrelia comprises 11 described and named species distributed in the northern hemisphere (Zorina 2008; Ekrem & Stur 2009; Kobayashi 2014). Only one species has so far recorded from North Korea.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F26FFA0FF7EE2F7149979BD.taxon	materials_examined	Material examined. Kŭmgang-san, 28 June – 02 July 1981, 2 males (paratypes), leg. W. Krzemiński.	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F26FFA0FF7EE2F7149979BD.taxon	discussion	Remarks. Detailed illustrated descriptions of adults and immature stages, with remarks on systematics and ecology of Zavrelia sinica gave Ekrem and Stur (2009). The species is known from two sites, in the Northeast China and North Korea, where just a couple of specimens, including three adult males have been collected (two examined here).	en	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
