identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C087D47F36FFB3FF7EE1C2168D7EE7.text	03C087D47F36FFB3FF7EE1C2168D7EE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cladotanytarsus (Cladotanytarsus) sinjongensis Ree et Kim, 1988 Ree & Kim 1988	<div><p>Cladotanytarsus (Cladotanytarsus) sinjongensis Ree et Kim, 1988</p> <p>Cladotanytarsus sinjongensis Ree et Kim, 1988: 19 (male, fig. 4a–c; South Korea).</p> <p>(Fig. 2A–D)</p> <p>Material examined. Phjŏngjang, 18 July 1981, 32 males; Sokam, 8 July 1981, 1 male; leg. W. Krzemiński.</p> <p>Remarks. A set of subtle morphological differences used in diagnostics, often overlapped by intraspecific variations, make the determination of adult males of Cladotanytarsus one of the most difficult within Tanytarsini and Chironomidae. Accurately mounted and illustrated specimens, their precise measurements and appropriate interpretation of diagnostic features are thus decisive. In fact, determination of Cladotanytarsus sinjongensis may be a challenge when based on the illustrated original description, although several crucial characters indicate that the specimens presently examined are conspecific with those described from South Korea (Ree &amp; Kim 1988). The males from North and South Korea compared here [data on the South Korean material, with remarks given in square brackets] are approximately of the same size, with the wing 1.18–1.45 mm long [1.2–1.3 mm], AR 0.90–1.03 [0.91–1.05], frontal tubercles 14–18 μm long [15–16 μm], similar lengths of palpomeres, leg ratios and the same body colouration. The hypopygial anal point is more or less lanceolate or tongue-shaped (Fig. 2A, B), 35–45 μm long [7–9 μm, inconsistency relative to the scaled fig. 4], the superior volsella is slender, narrowed at mid length, with the swollen and apically rounded distal part (Fig. 2A, C) [probably distorted during mounting], the stem of the median volsella is slightly curved, ~40 μm long [35–38 μm], with branched lamellae (Fig. 2A, D), as in all known Cladotanytarsus [the deep furcations overlooked].</p> </div>	https://treatment.plazi.org/id/03C087D47F36FFB3FF7EE1C2168D7EE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F36FFB0FF7EE07E102D7B4B.text	03C087D47F36FFB0FF7EE07E102D7B4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cladotanytarsus Kieffer 1921	<div><p>Genus: Cladotanytarsus Kieffer, 1921</p> <p>The genus Cladotanytarsus is divided into two subgenera: Cladotanytarsus s. str. Kieffer, 1921 and Lenziella Kieffer, 1922 (Giłka 2011b, Giłka &amp; Spies 2012, Puchalski &amp; Giłka 2017). Since no character typical of Lenziella (a globular basal swelling in the inferior volsella nor lobes of the mid and hind leg tibiae apices) has been found in C. sinjongensis males, we place the species in the subgenus Cladotanytarsus s. str. The genus is represented by nearly 80 species known from all continents except Antarctica (Puchalski et al. 2018). From North Korea, apart from C. sinjongensis, now found and redescribed below, two Cladotanytarsus species have so far been recorded by Reiss (1980), but determined to the genus level, as Cladotanytarsus sp. K 1 and K2.</p> </div>	https://treatment.plazi.org/id/03C087D47F36FFB0FF7EE07E102D7B4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F36FFB0FF7EE7EE16117DF3.text	03C087D47F36FFB0FF7EE7EE16117DF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsini Zavrel 1917	<div><p>Tribe: Tanytarsini Zavřel, 1917</p> <p>Subtribe: Tanytarsina Zavřel, 1917</p></div> 	https://treatment.plazi.org/id/03C087D47F36FFB0FF7EE7EE16117DF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F35FFB2FF7EE2C115DC7FB3.text	03C087D47F35FFB2FF7EE2C115DC7FB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Micropsectra atrofasciata (Kieffer 1911) Stur & Ekrem 2006	<div><p>Micropsectra atrofasciata (Kieffer, 1911)</p> <p>Micropsectra atrofasciata (Kieffer, 1911): Stur &amp; Ekrem 2006: 182 (male, pupa, larva, figs 3, 7; synonyms; Palaearctic).</p> <p>Micropsectra tonewdeea Sasa et Tanaka, 2002: 34 (male, fig. 8a–m; Japan), syn. nov.</p> <p>Material examined. Kŭmgang-san, 28 June – 2 July 1981, 1 male; Phjŏngjang, 18 July 1981, 3 males; leg. W. Krzemiński.</p> <p>Remarks. Micropsectra atrofasciata is an eurytopic and one of the most common species within the genus and the tribe in Europe (Giłka 2011c), distributed across the Palaearctic region. The wide geographical range and distinct intraspecific variations in the body size and colouration, depending also on a generation (seasonal heteromorphism), as well as in the shape of some key structures of the hypopygium (cf. Giłka 2000, 2001a, 2011a; see also remarks to Micropsectra joganplumosa and Tanytarsus volgensis below), were probably the reason for multiple descriptions of this species under different names. A redefinition of M. atrofasciata by Stur and Ekrem (2006) includes eight junior synonyms. Here, we analyzed another name, Micropsectra tonewdeea Sasa et Tanaka, 2002, based on description of the adult male and photographs of the holotype displayed on the website of the NMNS (405:070). A comparison of diagnostic characters shows that the type specimen described for this name falls within the wide range of morphological variability of M. atrofasciata. As far as we know, the name, treated as synonymous here, has not been used apart from the original description based on materials from locus typicus in Gunma Prefecture, Honshu (Sasa &amp; Tanaka 2002).</p> </div>	https://treatment.plazi.org/id/03C087D47F35FFB2FF7EE2C115DC7FB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F35FFB3FF7EE1F310BD784D.text	03C087D47F35FFB3FF7EE1F310BD784D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Micropsectra Kieffer 1909	<div><p>Genus: Micropsectra Kieffer, 1909</p> <p>One of the largest genera in the tribe Tanytarsini. Nearly 150 species names have been ascribed to Micropsectra, although many of them still require a redefinition or generic transfer. The genus is divided into several species groups. In the material analysed we found three species of the atrofasciata - and notescens groups.</p> </div>	https://treatment.plazi.org/id/03C087D47F35FFB3FF7EE1F310BD784D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F34FFB2FF7EE59110787BBF.text	03C087D47F34FFB2FF7EE59110787BBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Micropsectra joganplumosa Sasa et Okazawa, 1991 Sasa & Okazawa 1991	<div><p>Micropsectra joganplumosa Sasa et Okazawa, 1991</p> <p>Micropsectra joganplumosa Sasa et Okazawa, 1991: 55 (male, fig. 1.1a–j; Japan).</p> <p>Micropsectra tusimalemea Sasa et Suzuki, 1999: 17 (male, fig. 18a–g; Japan), syn. nov.</p> <p>Micropsectra tusimamenea Sasa et Suzuki, 1999: 18 (male, fig. 19a–g; Japan), syn. nov.</p> <p>Micropsectra tusimaquerea Sasa et Suzuki, 1999: 21 (male, fig. 23a–g; Japan), syn. nov.</p> <p>(Fig. 3A–D)</p> <p>Material examined. Myohyang-san, 22–25 June 1981, 1 male, leg. W. Krzemiński.</p> <p>Remarks. As in the previous species, males of Micropsectra joganplumosa are variable in the body colouration and main metric characters that probably resulted in description of the species under a series of names, i.e. Micropsectra tusimalemea, M. tusimamenea and M. tusimaquerea. The specimens analyzed (Sasa &amp; Suzuki 1999) were originally found as belonging to the species closely related to each other and/or to several further Micropsectra described in the same paper. However, the slight differences (scutum colouration, AR values, and length of digitus) are here defined as intraspecific variations. In fact, the drawings presented in original descriptions for all these names do not reflect shapes of the most important diagnostic structures, especially when compared with photographs of their type specimens (cf. Sasa &amp; Okazawa 1991, fig. 1.1a–j + NMNS 217:029 and Sasa &amp; Suzuki 1999, figs 18, 19 and 23 + NMNS 372:016, 372:018, 354:069). Some drawings presented under these names indicate a similar or identical shape of the same hypopygial structures (median and inferior volsellae); on the other hand, the same structure (anal point) nearly identical on photographs differs distinctly on the drawings. In case of such inconsistency, we assume that at least part of the drawings may have been mistakenly assigned to a specimen/name, thus the comparison of the diagnostic structures can only be based on photographs of the designated types. Despite of deformations of the holotypes on slides (hypopygia) some crucial characters indicate that they are conspecific. The characters best defining the species are: the shape of the anal point (Fig. 3A, B), the digitus and superior volsella, typical of the Micropsectra notescens species group (Fig. 3A, C), and the median volsella in shape of a hockey stick bearing numerous spoon-shaped lamellae (Fig. 3D). The species is known from the west of Japan: Toyama Prefecture in Honshu (as M. joganplumosa) and Tsushima Island, where specimens were collected at one locality, in the same time, using the same method (as M. tusimalemea, M. tusimamenea and M. tusimaquerea). Now Micropsectra joganplumosa is also recorded from central part of North Korea.</p> </div>	https://treatment.plazi.org/id/03C087D47F34FFB2FF7EE59110787BBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F34FFB5FF7EE18D10A57F5F.text	03C087D47F34FFB5FF7EE18D10A57F5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Micropsectra jokatertia Sasa et Ogata, 1999 Sasa & Ogata 1999	<div><p>Micropsectra jokatertia Sasa et Ogata, 1999</p> <p>Micropsectra jokatertia Sasa et Ogata, 1999: 92 (male, fig. 4a–n; Japan).</p> <p>(Figs 4A, B; 5C, G)</p> <p>Material examined. Kŭmgang-san, 28 June – 2 July 1981, 3 males, leg. W. Krzemiński.</p> <p>Remarks. The enormity of names coming from Japan pertain also to a series of Micropsectra (Sasa 1984, 1993; Sasa &amp; Okazawa 1994; Sasa &amp; Suzuki 1999; Sasa et al. 2001) of which specimens described under six names at least have to be compared with M. jokatertia. They are: M. chuzelonga Sasa, 1984, M. inamenea Sasa, Kitami et Suzuki, 2001, M. inaneoa Sasa, Kitami et Suzuki, 2001, M. johanaprima Sasa et Okazawa, 1994, M. tomoprima Sasa, 1993, and M. tusimaneoa Sasa et Suzuki, 1999. Based on more or less distinct characters given in original descriptions and observable on photographs of type specimens (cf. NMNS 051:011, 398:015, 402:023, 250:078, 244:001, 356:017, respectively to the above list of names), M. jokatertia (NMNS 352:051) differs from them in having a slender gonostylus (Fig. 4A vs. M. chuzelonga, M. inamenea), a longer and/or more slender anal point and its crest (Fig. 4A vs. M. chuzelonga, M. inaneoa, M. johanaprima, M. tomoprima, M. tusimaneoa), a longer or more slender digitus (Figs 4A &amp; 5G vs. M. chuzelonga, M. johanaprima, M. tusimaneoa), and a lower number of spoon-shaped lamellae of the median volsella (Figs 4B &amp; 5C vs. M. inamenea). Consequently, we refrain from synonymization of M. jokatertia, although synonymy within the compared names (excl. M. jokatertia) is highly probable.</p> <p>Micropsectra jokatertia fits well the concept of the notescens group analysed on the basis of key structures of the hypopygium (Giłka 2001b). When M. jokatertia is included to a comparison of several well-defined species, it places between M. rilensis Giłka, 2001 and M. recurvata Goetghebuer, 1928 in this series (Fig. 5).</p> </div>	https://treatment.plazi.org/id/03C087D47F34FFB5FF7EE18D10A57F5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE61E17277DF1.text	03C087D47F33FFB5FF7EE61E17277DF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neozavrelia fengchengensis Wang et Wang 1996	<div><p>Neozavrelia fengchengensis Wang et Wang, 1996</p> <p>Neozavrelia fengchengensis Wang et Wang, 1996: 121 (male, fig. 1; China); Guo &amp; Wang 2005: 189 (male, in key; China); Makarchenko et al. 2005: 414 (Russia); Giłka 2012: 35 (male, figs 1–4; North Korea); Orel 2021: 254 (male, figs 1–8; Russia).</p> <p>Material examined. Myohyang-san, 22–25 June 1981, 2 males, leg. W. Krzemiński.</p> <p>Remarks. An illustrated diagnostic description of the adult male, based on the material examined here, provided Giłka (2012). The species has been recently analysed also for intraspecific variations (Orel 2021), however, with data referred to the present material erroneously given to be from South Korea. To our knowledge, Neozavrelia fengchengensis has so far been recorded from the Northeast China (Liaoning Province), East Siberia (Chita Region), several other sites in the Russian Far East, and from North Korea.</p> </div>	https://treatment.plazi.org/id/03C087D47F33FFB5FF7EE61E17277DF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE5EF156E7C2B.text	03C087D47F33FFB5FF7EE5EF156E7C2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neozavrelia Goetghebuer	<div><p>Genus: Neozavrelia Goetghebuer in Goetghebuer et Thienemann, 1941</p> <p>Over 30 described species belong to Neozavrelia, although many of them still remain in wrong combinations, while some names require revision as potential synonyms. Two species have so far been recorded from North Korea (Giłka 2012).</p> </div>	https://treatment.plazi.org/id/03C087D47F33FFB5FF7EE5EF156E7C2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE05017037B25.text	03C087D47F33FFB5FF7EE05017037B25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neozavrelia tamanona (Sasa 1980) Gilka 2012	<div><p>Neozavrelia tamanona (Sasa, 1980)</p> <p>Tanytarsus tamanonus Sasa, 1980: 25 (male, figs 26a–e, 27a–h; Japan).</p> <p>Neozavrelia tamanona (Sasa, 1980): Sasa &amp; Kawai 1987: 30 (combination; Japan); Guo &amp; Wang 2005: 195 (male, figs 10–11; China); Giłka 2012: 36 (male, figs 5–6; North Korea).</p> <p>Material examined. Kŭmgang-san, 28 June – 2 July 1981, 1 male, leg. W. Krzemiński.</p> <p>Remarks. An illustrated redescription of the species with notes on systematics and possible synonymy have been based on the presently examined material (Giłka 2012).</p></div> 	https://treatment.plazi.org/id/03C087D47F33FFB5FF7EE05017037B25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE2E7169979F0.text	03C087D47F33FFB5FF7EE2E7169979F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paratanytarsus dissimilis (Johannsen 1905) Palmen 1960	<div><p>Paratanytarsus dissimilis (Johannsen, 1905) or P. inopertus (Walker, 1856)</p> <p>Paratanytarsus confusus Palmen or inopertus (Walk.): Reiss 1980: 149 (North Korea).</p> <p>Paratanytarsus dissimilis (Johannsen, 1905): Langton et al. 1988: 319 (= confusus Palmén, 1960). Paratanytarsus dissimilis (Johannsen, 1905) &amp; Paratanytarsus inopertus (Walker, 1856): Giłka 2011a: 39, 40 (males in key, figs 122–131).</p> <p>Remarks. The two specific names, including Paratanytarsus confusus, were alternatively mentioned by Reiss (1980), thus we assume that at least one species has been found in North Korea. Later synonymization of P. confusus with P. dissimilis enabled a clear definition of this species (Langton et al. 1988). Diagnostic characters for adult males of the two sibling species, P. dissimilis and P. inopertus, compared Giłka (2011a) in the couplet 6 of the key for Paratanytarsus. Though both are eurytopic and widespread in the Holarctic (Giłka 2011c), none has been confirmed in the presently examined material.</p> </div>	https://treatment.plazi.org/id/03C087D47F33FFB5FF7EE2E7169979F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F33FFB5FF7EE1E0175F7831.text	03C087D47F33FFB5FF7EE1E0175F7831.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paratanytarsus Thienemann et Bause	<div><p>Genus: Paratanytarsus Thienemann et Bause in Bause, 1913</p> <p>Nearly 70 described species belong to Paratanytarsus, including several species still being incorrectly ascribed to other genera. At least three species have been recorded in North Korea; apart from the below species, Reiss (1980) recorded one more, determined as Paratanytarsus sp. K 1.</p> </div>	https://treatment.plazi.org/id/03C087D47F33FFB5FF7EE1E0175F7831	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB9FF7EE13011377829.text	03C087D47F3FFFB9FF7EE13011377829.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paratanytarsus laccophilus (Edwards 1929) Reiss & Sawedal (1981) Reiss 1980	<div><p>Paratanytarsus laccophilus (Edwards, 1929)</p> <p>Tanytarsus (Lundstroemia) laccophilus Edwards, 1929: 409 (male, fig. 14c; UK).</p> <p>Paratanytarsus laccophilus (Edwards, 1929): Reiss 1980: 149 (North Korea); Reiss &amp; Säwedal 1981: 101 (male, pupa, figs, in keys).</p> <p>Remarks. The adult male of Paratanytarsus laccophilus is clearly defined and easily separable even from its closest relative, P. paralaccophilus Giłka et Paasivirta, 2008 known from Lapland (Giłka &amp; Paasivirta 2008). In North Korea, the species was recorded by Reiss (1980) but not confirmed in the material presently analyzed.</p> </div>	https://treatment.plazi.org/id/03C087D47F3FFFB9FF7EE13011377829	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB9FF7EE200164A7911.text	03C087D47F3FFFB9FF7EE200164A7911.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rheotanytarsus Thienemann et Bause	<div><p>Genus: Rheotanytarsus Thienemann et Bause in Bause, 1913</p> <p>The genus includes nearly 100 species distributed worldwide. Some names, especially those coming from east and south of Asia, still require a revision (potential synonyms, incorrect combinations). In the material presently examined we found one species.</p></div> 	https://treatment.plazi.org/id/03C087D47F3FFFB9FF7EE200164A7911	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3FFFB8FF7EE30511737FB3.text	03C087D47F3FFFB8FF7EE30511737FB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rheotanytarsus tusimatfegeus (Sasa et Suzuki 1999)	<div><p>Rheotanytarsus tusimatfegeus (Sasa et Suzuki, 1999)</p> <p>Tanytarsus tusimatfegeus Sasa et Suzuki, 1999: 27 (male, fig. 31a–e; Japan).</p> <p>R.? tusimatfegeus Sasa et Suzuki, 1999: Makarchenko et al. 2005: 415 (Russia).</p> <p>Material examined. Myohyang-san, 22–25 June 1981, 3 males, leg. W. Krzemiński.</p> <p>Remarks. Neither the original description, based on simple schematic drawings, nor the photographs of the Rheotanytarsus tusimatfegeus holotype, not very successfully slide-mounted (NMNS 371:001), allow for an unambiguous definition of this species when each of the characters' source taken separately, but both in combination indicate the species analyzed here. North Korean males are slightly smaller and have a proportionally lower AR than those from Japan (wing length 1.60–1.70 vs 1.80–1.96, AR ~ 0.65 vs 0.76–0.88), while other characters are conspecific. Rh. tusimatfegeus has so far been reported from the Sea of Japan region, in Tsushima (Sasa &amp; Suzuki 1999) and most likely in Russia (Makarchenko et al. 2005). Now recorded for the first time in North Korea.</p> </div>	https://treatment.plazi.org/id/03C087D47F3FFFB8FF7EE30511737FB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE77A117B7A8B.text	03C087D47F3EFFB8FF7EE77A117B7A8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus chinyensis Goetghebuer 1934	<div><p>Tanytarsus chinyensis Goetghebuer, 1934</p> <p>Tanytarsus chinyensis Goetghebuer: Reiss &amp; Fittkau 1971: 99, 169 (male, fig. 6, systematics); Giłka &amp; Paasivirta 2009: 32, 40 (male, fig. 28, systematics); Lin et al. 2018a: 330, 387 (systematics); Lin et al. 2018b: 10, 11 (systematics).</p> <p>Material examined. Phjŏngjang, 18 July 1981, 1 male, leg. W. Krzemiński.</p> <p>Remarks. Reiss and Fittkau (1971) indicated this species as representative of the chinyensis group. The concept was recently refuted, and the former chinyensis group was proposed to be split into several unrelated clusters/groups (Lin et al. 2018a, b). Alternatively, the group was postulated to be reduced. However, Tanytarsus chinyensis was not included in the analyses (op. cit.), thus according to this concept, the group name-bearing species is unplaced in any of the species groups proposed, apparently, except for the chinyensis group (see also remarks to T. tamadecimus below). In the material analysed, we found a single male collected in the Botanical Garden in Phjŏngjang.</p> </div>	https://treatment.plazi.org/id/03C087D47F3EFFB8FF7EE77A117B7A8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE08D17E57BF3.text	03C087D47F3EFFB8FF7EE08D17E57BF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus heusdensis Goetghebuer 1923	<div><p>Tanytarsus heusdensis Goetghebuer, 1923</p> <p>Tanytarsus heusdensis Goetghebuer: Reiss &amp; Fittkau 1971: 101, 170 (male, figs 8, 9a–c, systematics); Reiss 1980: 149 (North Korea); Giłka &amp; Paasivirta 2009: 40 (male, fig. 30, systematics); Lin et al. 2018a: 375 (male, fig. 29a–d, systematics); Lin et al. 2018b: 10, 11 (fig. 1, systematics).</p> <p>Remarks. Tanytarsus heusdensis was ascribed to the chinyensis species group (Reiss &amp; Fittkau 1971); recently proposed to be excluded into the separate heusdensis complex/group (Lin et al. 2018a, b). The species was recorded from North Korea by Reiss (1980), but not confirmed in our material.</p> </div>	https://treatment.plazi.org/id/03C087D47F3EFFB8FF7EE08D17E57BF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFBBFF7EE25016367EE7.text	03C087D47F3EFFBBFF7EE25016367EE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus iriolemeus Sasa et Suzuki, 2000 Sasa & Suzuki 2000	<div><p>Tanytarsus iriolemeus Sasa et Suzuki, 2000</p> <p>Tanytarsus iriolemeus Sasa et Suzuki, 2000: 20 (male, fig. 21 a–n; Japan).</p> <p>(Fig. 6A–D)</p> <p>Material examined. Kŭmgang-san, 28 June – 2 July 1981, 1 male; Myohyang-san, 22–25 June 1981, 2 males; leg. W. Krzemiński.</p> <p>Remarks. The North Korean specimens fit well the original description of Tanytarsus iriolemeus (Sasa &amp; Suzuki 2000) except for one character crucial in species group concepts for Tanytarsus - the strong bristle-like median setae placed in a roundish field of the hypopygial anal tergite. This structure is omitted in the original description but present in the holotype specimen (slightly beyond the range of sharpness on the photo presented at NMNS website, 385:035). This character combined with the anal point elongate, bearing a few spinulae (Fig. 6A, B; cf. also Tanytarsus innarensis Brundin 1947), the superior volsella roundish, medially excavated, with a well-protruded posteromedian corner, the digitus long apically rounded (Fig. 6C), and the shape of the median volsella (Fig. 6D) may indicate that T. iriolemeus belongs to the verralli species group. In the analyses by Lin et al. (2018b), T. iriolemeus falls in the tamakutibasi cluster that together with the aterrimus + chinyensis + curticornis groups is sister to a clade comprising the Holarctic excavatus, recurvatus, and verralli groups.</p> </div>	https://treatment.plazi.org/id/03C087D47F3EFFBBFF7EE25016367EE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3EFFB8FF7EE59B103A7CD2.text	03C087D47F3EFFB8FF7EE59B103A7CD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus van der Wulp 1874	<div><p>Genus: Tanytarsus van der Wulp, 1874</p> <p>With nearly 400 described species, Tanytarsus is the largest genus in the tribe and the second most diverse genus of the family Chironomidae (cf. Dantas et al. 2022). At least 11 species of Tanytarsus have been recorded from North Korea. The review comprises 3 species found previously (Reiss 1980, apart form 6 species ascribed to Tanytarsus as “sp. K1–6”) and 8 species here recorded for the first time from North Korea (Table 2).</p> </div>	https://treatment.plazi.org/id/03C087D47F3EFFB8FF7EE59B103A7CD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3DFFBBFF7EE52516407A2A.text	03C087D47F3DFFBBFF7EE52516407A2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus occultus Brundin 1949	<div><p>Tanytarsus occultus Brundin, 1949</p> <p>Tanytarsus holochlorus Edw.: Brundin 1947: 67 (male, fig. 100; Sweden).</p> <p>Tanytarsus occultus Brundin, 1949: 847 (male, Sweden); Reiss &amp; Fittkau 1971: 89, 123 (male, fig. 35a–c, in key); Ekrem et al. 2003: 315 (male, pupa, figs 248–250; Europe, North America); Ekrem 2004: 137 (pupa, figs 183–187); Giłka &amp; Paasivirta 2007: 107– 110 (male, figs 2–4, 13, 16; Fennoscandia); Giłka et al. 2018: 573–576 (male, diagnostics, barcoding, phylogeny).</p> <p>Material examined. Phjŏngjang, 13 June 1981, 2 males; Sokam, 8 July 1981, 3 males; leg. W. Krzemiński.</p> <p>Remarks. A diagnostic character of the Tanytarsus occultus male is the hypopygial anal point, usually with a square or concave apex, which the shape, however, may be affected by strong intraspecific variations. T. occultus was described on the basis of a males’ series by Brundin (1949), who referred the description to his illustration of a specimen’s hypopygium presented two years earlier [Brundin 1947, originally erroneously as Tanytarsus holochlorus (= T. mendax Kieffer, 1925)]. The illustration was apparently based on a specimen with a hypopygial anal point of a structure rarely observed in T. occultus. That choice could lead to confusion until Reiss &amp; Fittkau (1971) redescribed the male on the basis of Brundin’s series, indicating variations in the shape of the anal point, thus stabilizing the status of the species. Recently, one more close species was described, Tanytarsus latens Giłka, Paasivirta, Gadawski et Grabowski, 2018, in which the anal point is similar or nearly identical (variable) to that illustrated from the atypical specimen of T. occultus by Brundin (1947). In case of these two species, the anal point structure can be thus critically misleading in their delimitation. The characters best separating males of the two species is the shape/length of the superior volsella and the digitus, and the arrangement of anal tergite bands (cf. Giłka et al. 2018). T. occultus is one of the best-defined species in the genus in term of morphological variability and diagnostics supported by molecular analyses (Reiss &amp; Fittkau 1971; Ekrem et al. 2003, Ekrem 2004; Giłka &amp; Paasivirta 2007, Giłka et al. 2018). Only now recorded from North Korea.</p> </div>	https://treatment.plazi.org/id/03C087D47F3DFFBBFF7EE52516407A2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3DFFBBFF7EE01E16FD790D.text	03C087D47F3DFFBBFF7EE01E16FD790D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus oscillans Johannsen 1932	<div><p>Tanytarsus oscillans Johannsen, 1932</p> <p>Tanytarsus oscillans Johannsen, 1932: Ekrem 2002: 24 (male, figs 34, 35; Indonesia; = Tanytarsus cultellus Chaudhuri et Datta, 1992, = Tanytarsus sibafegeus Sasa, Sumita et Suzuki, 1999).</p> <p>Material examined. Kaesŏng, 16 July 1981, 4 males; Sokam, 8 July 1981, 3 males; leg. W. Krzemiński. Comparative material. Tanytarsus smolandicus: Sweden, Nedre Dalälven, 11 August 2003, 1 male, Storsjön, 18 July 2003, 20 males; leg. W. Giłka.</p> <p>Remarks. Ekrem (2002) redescribed the male of Tanytarsus oscillans based on the holotype specimen. In remarks on its close relatives, T. smolandicus Brundin, 1947 (terra typica in Sweden) and T. unagiseptimus Sasa, 1985 (Japan), selected diagnostic characters were indicated to separate these species. It seems, however, that these features (number of median setae, length of frontal tubercles, AR value, presence/absence of microtrichia between anal point crests and slight differences in shape of inferior volsella), in part at least, may be within the morphological variability of T. oscillans and/or T. smolandicus. According to the concept by Lin et. al. (2018b), T. oscillans and T. unagiseptimus are sister species (not ascribed to any group analyzed) while T. smolandicus has not been included. Interestingly, a concept of possible synonymy between the abovementioned names has been raised recently, when DNA sequences from specimens sampled in Fennoscandia and Japan, determined as T. oscillans or T. unagiseptimus, have been compared that resulted in ~97–98% of DNA sequence compatibility (Paasivirta, pers. comm.). Unfortunately, our specimens collected decades ago (kept in denatured ethanol or slide-mounted in Canada balsam) do not give a chance to be analyzed molecularly for supporting this concept.</p> </div>	https://treatment.plazi.org/id/03C087D47F3DFFBBFF7EE01E16FD790D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F3CFFBAFF7EE23211DC79B7.text	03C087D47F3CFFBAFF7EE23211DC79B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus oyamai Sasa 1979	<div><p>Tanytarsus oyamai Sasa, 1979</p> <p>Tanytarsus oyamai Sasa, 1979: 3 (male, figs 5–7; Japan); Ekrem 2002: 26 (male, pupa, larva, figs 32, 33, 63, 70, 78, 89, 94, 99, 104, 108 + references therein; Japan).</p> <p>(Fig. 6E–H)</p> <p>Material examined. Kaesŏng, 16 July 1981, 1 male; Phjŏngjang, 13 June 1981, 7 males, 18 July 1981, 1 male; Sariwŏn, 18 June 1981, 3 males; Sokam, 8 July 1981, 2 males; leg. W. Krzemiński.</p> <p>Remarks. Tanytarsus oyamai is probably one of the most common Tanytarsini in the southern part of the Sea of Japan basin; also frequent in North Korea - collected from half of the explored sites. The adult male is well-defined, thus only several illustrations are attached to show variations of the most important diagnostic structures (Fig. 6E–H).</p> </div>	https://treatment.plazi.org/id/03C087D47F3CFFBAFF7EE23211DC79B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE4F217B57F60.text	03C087D47F39FFBFFF7EE4F217B57F60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus pallidicornis (Walker 1856) Reiss & Fittkau 1971	<div><p>Tanytarsus pallidicornis (Walker, 1856)</p> <p>Tanytarsus pallidicornis (Walker, 1856): Reiss &amp; Fittkau 1971: 140, 193 (male, figs 58, 59, systematics); Giłka 2011a: 79 (male, diagnostics, figs 349–352; Poland).</p> <p>Material examined. Phjŏngjang, 18 July 1981, 2 males, leg. W. Krzemiński.</p> <p>Remarks. Tanytarsus pallidicornis is one of the best-defined in the genus. This common eurytopic species is widely distributed in the Holarctic but only now recorded from North Korea.</p> </div>	https://treatment.plazi.org/id/03C087D47F39FFBFFF7EE4F217B57F60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE5B517197D59.text	03C087D47F39FFBFFF7EE5B517197D59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus takahashii Kawai et Sasa, 1985 Kawai & Sasa 1985	<div><p>Tanytarsus takahashii Kawai et Sasa, 1985</p> <p>Tanytarsus takahashii Kawai et Sasa, 1985: 22 (male, fig. 8; Japan); Ekrem 2002: 28 (male, figs 47, 48; Japan + references therein).</p> <p>(Fig. 7A–C)</p> <p>Material examined. Phjŏngjang, 18 July 1981, 9 males; Sokam, 8 July 1981, 2 males; leg. W. Krzemiński.</p> <p>Remarks. A group membership of Tanytarsus takahashii, based on morphology, was discussed by Ekrem (2002) who indicated several possible placements for this species (the mendax, eminulus or lestagei group). Later, molecular analyses supported the concept of the lestagei group as the place for T. takahashii (Lin et al. 2018b). The species has been also compared with the fossil species T. serafini Giłka, 2010, and suggested to belong to the serafini group (cf. Giłka 2010, pl. II, figs. 2–7 and Fig. 7A–C).</p> </div>	https://treatment.plazi.org/id/03C087D47F39FFBFFF7EE5B517197D59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBFFF7EE7FC14B778DB.text	03C087D47F39FFBFFF7EE7FC14B778DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus tamadecimus Sasa, 1980 Sasa 1980	<div><p>Tanytarsus tamadecimus Sasa, 1980</p> <p>Tanytarsus tamadecimus Sasa, 1980: 26 (male, figs 28, 29; Japan).</p> <p>(Fig. 8A–C)</p> <p>Material examined. Hwangju, 18–19 June 1981, 2 males; Kŭmgang-san, 28 June – 2 July 1981, 3 males; Sokam, 8 July 1981, 1 male; leg. W. Krzemiński.</p> <p>Remarks. The Korean specimens of Tanytarsus tamadecimus fit the original description, except for a couple of characters, possibly taken from specimens deformed before their illustrating. According to Sasa (1980), the digitus is “composed of a long spatulated dorsal process and a ventral hook-like process connected with each other”. In fact, at the Sasa’s illustrations the hook-like process looks so unusually (not known from any other species) as we assume it could be misidentified with a long darkly pigmented tubercle at the digitus base that was omitted in the original illustrations (cf. Sasa 1980, fig. 29B, F, G and Fig. 8B). Character/shape overinterpretation seems to pertain also to the anal point and median volsella (cf. Sasa 1980, fig. 29D, E and Fig. 8A, C), thus we attach illustrations of selected variations to supplement the description.</p> <p>Tanytarsus tamadecimus and T. chinyensis are treated here as close relatives (see also remarks to T. chinyensis). The concept of the chinyensis species group sensu Reiss and Fittkau (1971) was recently refuted and/or its species composition was suggested to be reduced (Lin et al. 2018a, b); however, neither T. chinyensis nor T. tamadecimus have been included in the analyses. Consequently, we keep the chinyensis group, with the two species and T. cretensis Reiss, 1987, plus T. tamagotoi Sasa, 1983 and T. simantoseteus Sasa, Suzuki et Sakai, 1998, as it was suggested (op. cit.), plus a series of species by Sasa and co-authors: T. chuzesecundus Sasa, 1984, T. inawaijeus Sasa, Kitami et Suzuki, 2000, T. oyaberotundus Sasa, Kawai et Ueno, 1988 and T. tusimatkeleus Sasa et Suzuki, 1999. Males of all the species have the broad thumb-like or spatulate digitus and the horn-like posteromedian corner of the superior volsella.</p> </div>	https://treatment.plazi.org/id/03C087D47F39FFBFFF7EE7FC14B778DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F39FFBEFF7EE37D15657F5F.text	03C087D47F39FFBEFF7EE37D15657F5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus tamaundecimus Sasa, 1980 Sasa 1980	<div><p>Tanytarsus tamaundecimus Sasa, 1980</p> <p>Tanytarsus tamaundecimus Sasa, 1980: 27 (male, female, pupa, figs 30–32; Japan); Ekrem 2002: 28 (diagnostics). (Fig. 8D–F)</p> <p>Material examined. Kŭmgang-san, 28 June – 2 July 1981, 3 males, leg. W. Krzemiński.</p> <p>Remarks. Ekrem (2002) listed characters found as crucial in delimitation of Tanytarsus tamaundecimus on the basis of the original description. However, the type material has not been examined, neither photographs of the holotype are displayed at the NMNS website. The examined Korean specimens have been preliminarily confirmed as T. tamaundecimus (Ekrem, pers. comm.), thus several key structures and their variations are supplemented here (Fig. 8D–F).</p> </div>	https://treatment.plazi.org/id/03C087D47F39FFBEFF7EE37D15657F5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFBEFF7EE06F16747B3F.text	03C087D47F38FFBEFF7EE06F16747B3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stempellina Thienemann et Bause	<div><p>Genus: Stempellina Thienemann et Bause in Bause, 1913</p> <p>So far, over 20 specific names have been ascribed to Stempellina as the genus, subgenus of Tanytarsus or to other Tanytarsini generic names. According to current knowledge, the number of known species is lower due to systematic combinations other than original or doubtful status of some names (nomina dubia). Stempellina includes 15 described species: 2 extinct and 13 extant (Giłka 2005, Zakrzewska et al. 2020), thus its share in the tribe’s world extant fauna is less than 2%. Stempellina are recorded in all continents except Antarctica, mostly in the Holarctic. In North Korea, one unknown species has been found.</p> </div>	https://treatment.plazi.org/id/03C087D47F38FFBEFF7EE06F16747B3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFBEFF7EE5F615057D6F.text	03C087D47F38FFBEFF7EE5F615057D6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytarsus volgensis Miseiko 1967	<div><p>Tanytarsus volgensis Miseiko, 1967</p> <p>Tanytarsus fimbriatus Reiss et Fittkau, 1971: 118 (male, figs 29, 30); Reiss 1980: 149 (North Korea).</p> <p>Tanytarsus volgensis Miseiko, 1967: Ekrem 2004: 139 (male, figs 194–202; = fimbriatus Reiss et Fittkau, 1971); Giłka &amp; Paasivirta 2007: 107–109 (male, figs 7–9, 12, 14).</p> <p>Material examined. Hwangju, 18–19 June 1981, 7 males; Kaesŏng, 16 July 1981, 1 male; Phjŏngjang, 11 June 1981, 27 males; 13 June 1981, 28 males; 18 July 1981, 26 males; Sariwŏn, 18 June 1981, 5 males; Sokam, 8 July 1981, 29 males; leg. W. Krzemiński.</p> <p>Remarks. This eurytopic and widespread Holarctic species (cf. Giłka 2011c) was found as the most frequent and abundant tanytarsine in North Korea - recorded from two sites by Reiss (1980, as T. fimbriatus) and now confirmed at further 5 localities (123 males = over 55% of all examined specimens). Interestingly, an extremely wide range of the body size and colouration have been observed in the examined males, among which the biggest specimens, darker are nearly twice as long as the smallest ones, lighter (see also remarks to Micropsectra atrofasciata and M. joganplumosa).</p> </div>	https://treatment.plazi.org/id/03C087D47F38FFBEFF7EE5F615057D6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F38FFA0FF7EE117164A7B92.text	03C087D47F38FFA0FF7EE117164A7B92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stempellina radoszkowskii Giłka & Gadawski 2022	<div><p>Stempellina radoszkowskii sp. nov.</p> <p>http://zoobank.org/ urn:lsid:zoobank.org:act: F90C1ED3-A009-432A-94B6-BD34FE2AA118</p> <p>(Fig. 9A–E)</p> <p>Type material. Holotype, adult male: North Korea, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.33333&amp;materialsCitation.latitude=41.833332" title="Search Plazi for locations around (long 128.33333/lat 41.833332)">Paektu Mountain</a>, Samdžijŏn at Samji Lake (41°50'N 128°20'E; ~ 1400 m a.s.l.), 23– 25.08.1992, sweep net, leg. A. Palaczyk.</p> <p>Derivatio nominis. The specific epithet commemorates Oktawiusz Wincenty Radoszkowski (1820–1895), the meritorious entomologist and the first Polish discoverer of insects from Korea.</p> <p>Diagnosis. Antenna with 5–6 flagellomeres, borders between flagellomeres 5–9 indistinct, ultimate segments completely fused, plume consisted of short sparse setae. Frontal tubercles robust. Hypopygial median setae absent. Anal point short and apically swollen, anal point crests thin, spinulae absent. Superior volsella extensive, apex round, with small apical protuberance. Stem of median volsella straight, with setiform and long spindle–shaped lamellae.</p> <p>Description. Adult male (n = 1).</p> <p>Body size. Minute species, total length ~ 2 mm. Wing length 1010 μm.</p> <p>Colouration (in alcohol). Eyes, scutal stripes, postnotum and sternum light brown, remaining body parts pale yellowish, wing membrane transparent, veins a bit darker.</p> <p>Head. Eyes bare, reniform, broadly separated by frons. Antenna only with 5–6 well discernible flagellomeres, borders between flagellomeres 5–9 indistinct, ultimate segments completely fused, AR ~2.6 (when flagellum measured as 6-segmented), ~2.0 (as 7-segmented), ~1.3 (as 9-segmented); plume weak, consisted of short and sparse setae (Fig. 9A). Frontal tubercles robust, ~30 μm long and ~10 μm wide at base, conical, with apex rounded. Length of palpomeres 2–5 (μm): 32, 66, 72, 117. Clypeus with 16 setae.</p> <p>Thorax chaetotaxy. Setae weak, poorly observable; Ac not observed, likely absent, Dc 7–8, Pa 1, Scts 2 (pair of setae placed medially).</p> <p>Wing. Squama bare, anal lobe reduced, cell m 3+4, distal part of cells m 1+2 and r 4+5 with macrotrichia, cell r 4+5 with a single row of macrotrichia in 3/4 distal part; veins C, R, R 1, R 4+5 (distal 1/4–1/2 part), M 1+2 (distal 3/4 part), Cu 1 and false vein above M 3+4 always with macrotrichia; An and false vein under Cu 1 (distal part) usually with some macrotrichia, other veins bare; M 3+4 ending distinctly distal of R 4+5.</p> <p>Legs. Fore leg tibia with s-shaped spur (~5 μm long). Combs of mid and hind leg tibiae fine, each bearing spur: straight and equally long (~20 μm) on mid leg, and unequal on hind leg - longer spur (~40 μm) slender and evenly curved, shorter spur (~16 μm) straight; ta 1 of p 2 bearing 2 hook–shaped sensilla chaetica. For length of legs segments and legs ratios see Table 1.</p> <p>Hypopygium. Gonostylus (~75 μm) as long as gonocoxite. Anal tergite with bands V-shaped broadly separated and 2–3 posterolateral setae, median setae absent. The tergite tapering towards short and apically slightly swollen anal point, anal point crests thin, spinulae absent (Fig. 9B, C). Superior volsella extensive, broadest at base, tapering towards round apex bearing small apical protuberance; 3 setae on anteromedian margin (Fig. 9B, D, E). Stem of median volsella straight, ~30 μm long, with setiform and long spindle–shaped lamellae reaching tip of inferior volsella (Fig. 9B, E). Inferior volsella slightly curved, parallel-sided, with distal part rounded (Fig. 9B, E).</p> <p>Remarks. Except for a combination of hypopygial characters given in the diagnosis, the adult male of Stempellina radoszkowskii differs from all known Stempellina by lower number of antennal flagellomeres. This character was recently analysed in extant and fossil species of the genus (Zakrzewska et al. 2020), as well as in Stempellinella (Zakrzewska &amp; Jankowska 2021). A distinct trend towards reduction of the number of flagellomeres (12 or less) in extant species relative to those of fossil is confirmed here.</p> </div>	https://treatment.plazi.org/id/03C087D47F38FFA0FF7EE117164A7B92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F26FFA0FF7EE1BB114B787B.text	03C087D47F26FFA0FF7EE1BB114B787B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zavrelia Kieffer, Thienemann et Bause 1913	<div><p>Genus: Zavrelia Kieffer, Thienemann et Bause in Bause, 1913</p> <p>The small genus Zavrelia comprises 11 described and named species distributed in the northern hemisphere (Zorina 2008; Ekrem &amp; Stur 2009; Kobayashi 2014). Only one species has so far recorded from North Korea.</p> </div>	https://treatment.plazi.org/id/03C087D47F26FFA0FF7EE1BB114B787B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
03C087D47F26FFA0FF7EE2F7149979BD.text	03C087D47F26FFA0FF7EE2F7149979BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zavrelia sinica Ekrem et Stur, 2009 Ekrem & Stur 2009	<div><p>Zavrelia sinica Ekrem et Stur, 2009</p> <p>Zavrelia sinica Ekrem et Stur, 2009: 137 (male, female, pupa, larva, figs 10–11; China, North Korea).</p> <p>Material examined. Kŭmgang-san, 28 June – 02 July 1981, 2 males (paratypes), leg. W. Krzemiński.</p> <p>Remarks. Detailed illustrated descriptions of adults and immature stages, with remarks on systematics and ecology of Zavrelia sinica gave Ekrem and Stur (2009). The species is known from two sites, in the Northeast China and North Korea, where just a couple of specimens, including three adult males have been collected (two examined here).</p> </div>	https://treatment.plazi.org/id/03C087D47F26FFA0FF7EE2F7149979BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Giłka, Wojciech;Gadawski, Piotr	Giłka, Wojciech, Gadawski, Piotr (2022): Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea. Zootaxa 5214 (2): 151-175, DOI: https://doi.org/10.11646/zootaxa.5214.2.1
