taxonID	type	description	language	source
03B187913A4CFFF9FF3081A68BF2FC20.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Austria (Schrank 1781). Geographical distribution: Holarctic (Davids et al. 2006), remarkably Europe (GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in Vila Nova de Cerveira, and Valença (São Pedro da Torre) municipalities.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	materials_examined	Material examined: 1 undetermined sex adult; Vila Nova de Cerveira, Portugal; 41 º 56 ′ 12.768 ″ N, 8 º 45 ′ 9.108 ″ W; October 2019; coll. by C. Antunes and E. Martins; catalogue number NatMIP - CAOr- 0003. 1 undetermined sex adult; São Pedro da Torre, Portugal; 41 º 59 ′ 27 ″ N, 8 º 40 ′ 50.76 ″ W; 11 May 2020; coll by C. Antunes and E. Martins; catalogue number NatMIP-CAOr- 0005. Ecological notes: In lakes, ponds and slow-moving waters, as well as present in duckweed Lemna spp. (Davids et al. 2006); found also in the main river stretch and in the guts of European flounder P. flesus.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021), and also represents the first occurrence for Northern Europe. Hydrozetes lemnae (Coggi, 1897) Synonym: Notaspis lemnae Coggi, 1897 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	materials_examined	Type material: Possibly not deposited at scientific collection. Type locality: Italy, in the area around Bologna, Italy (Coggi 1897). Geographical distribution: Semi-cosmopolitan (Palaearctic, Nearctic, Ethiopic, Eastern, Australian, and Neotropical geographical zones) (Subías 2018).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in the tributary section “ Rio Mouro, Ponte do Curto ” (Monção municipality, 42 º 4 ′ 29.27 ″ N, 8 º 23 ′ 39.12 ″ W).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	materials_examined	Material examined: 1 undetermined sex adult; Ponte do Curto, Portugal, 42 º 4 ′ 29.27 ″ N, 8 º 23 ′ 39.12 ″ W; September 2020; coll. by Aquamuseu Team; NatMIP-CAOr- 0011. Ecological notes: Freshwater, feeding on submerged plant fragments (Pérez-Íñigo 1997). Present in macrophytes (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4CFFF9FF3081A68BF2FC20.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4DFFF9FF3082EB89A3F880.taxon	materials_examined	Type material: Syntypes: Possibly IFREMER 39042 (Greenland), IFREMER 34702 (Portugal, Azores), IFREMER 38960 and 20172 (Spain, Bay of Biscay), IFREMER 21648 (Angola) (GBIF 2022). Type locality: Deep Atlantic basins: Greenland, Labrador Basin, 58 ° 47 ′ 7 N, 52 ° 56 ′ 5 W; Portugal, Azores Sill, 36 ° 48 ′ 5 N, 27 ° 06 ′ 0 W; Spain, Bay of Biscay, 44 ° 06 ′ 3 N, 4 ° 22 ′ 4 W; northern edge of Angola Basin, 9 ° 12 ′ S, 10 ° 29 ′ E (Bartsch 1978). Geographical distribution: Atlantic Ocean (Bartsch 1978; GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4DFFF9FF3082EB89A3F880.taxon	distribution	Distribution in Portugal: Azores Archipelago (Bartsch 1978). Ecological notes: Typical marine benthic, found from 2006 to 4223 m depth, associated with poriferans and hydrozoans (Bartsch 1978).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4DFFFEFF30807F8D61FF50.taxon	materials_examined	Type material: Holotype: (Bartsch 1996), possibly IFREMER 58497 (GBIF 2022). Type locality: Portugal: Azores Triple Junction, 38 º 20.35 ′ N, 30 º 40.15 ′ W (Bartsch 1996). Geographical distribution: Portugal: Azores Triple Junction (Bartsch 1996; GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4DFFFEFF30807F8D61FF50.taxon	distribution	Distribution in Portugal: Azores Triple Junction (Bartsch 1996; GBIF 2022). Ecological notes: Typical benthic marine, found from 540 to 919 m depth, at hydrothermal vent associated with hydrozoans, gorgonians, polychaetes, molluscs, brachiopods, amphipods and crinoids (Bartsch 1996).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF3086228DFEFDB8.taxon	materials_examined	Type material: Possible IFREMER 25084 (GBIF 2022). Type locality: Portugal: Azores Archipelago (Bartsch 1988). Geographical distribution: Portugal: Azores Archipelago (Bartsch 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF3086228DFEFDB8.taxon	distribution	Distribution in Portugal: Azores Archipelago (Bartsch 1988). Ecological notes: Depth range 1505 - 1540 m, marine benthic, found with foraminiferans and pteropods (Bartsch 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF30859A8D46FB94.taxon	materials_examined	Type material: Holotype: MNHN 63 B 8 (Bartsch 1994); Paratype: One female: MNHN 63 B 7 (Bartsch 1994). Type locality: Portugal: Azores Archipelago: Lucky Strike hydrothermal field, 37 ° 20 ′ N, 32 ° 17 ′ W (Bartsch 1994). Geographical distribution: Portugal: Azores Archipelago: Lucky Strike hydrothermal field (Bartsch 1994; GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF30859A8D46FB94.taxon	distribution	Distribution in Portugal: Azores Archipelago: Lucky Strike hydrothermal field (Bartsch 1994, 1996; GBIF 2022). Ecological notes: Depth 1636 m, marine benthic (Bartsch 1994), common on rocky formations, associated with poriferans, hydrozoans, nematodes, polychaetes, molluscs, crustaceans and pycnogonids (Bartsch 1996).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF3083E6892BF98C.taxon	materials_examined	Type material: Holotype deposited at the MNHN (Bartsch 1996); Paratype: One female (Bartsch 1996), IFREMER 58496 (GBIF 2022). Type locality: Portugal: Azores Archipelago: Vent Field Menez-Gwen, Azores Triple Junction, 37 º 50.46 ′ N, 32 º 31.35 ′ W (Bartsch 1996; GBIF 2022). Geographical distribution: Portugal: Azores Archipelago (Bartsch 1996; GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF3083E6892BF98C.taxon	distribution	Distribution in Portugal: Azores Archipelago: Vent Field Menez-Gwen, Azores Triple Junction (Bartsch 1996). Ecological notes: Typical marine benthic, found from depth 840 to 870 m, hydrothermal vent associated with hydrozoans, polychaetes, molluscs and crustaceans; legs fouled with black particles and filamentous bacteria (Bartsch 1996).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF3081EE8BC1F858.taxon	materials_examined	Type material: Holotype: MNHN 63 B 5 (Bartsch 1994). Type locality: Portugal: Azores Archipelago, Lucky Strike hydrothermal field, Azores Triple Junction, 37 ° 20 ′ N, 32 ° 17 ′ W (Bartsch 1994; GBIF 2022). Geographical distribution: Portugal: Azores Archipelago: Lucky Strike field, Azores Triple Junction (Bartsch 1994; GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4AFFFEFF3081EE8BC1F858.taxon	distribution	Distribution in Portugal: Azores Archipelago: Lucky Strike field, Azores Triple Junction (Bartsch 1994). Ecological notes: Marine benthic, deep sea at 1636 m (Bartsch 1994).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFFFF3086FB8968FC68.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Denmark (probably surroundings of Copenhagen) or Norway (Müller 1776). Geographical distribution: Western Palaearctic (Davids et al. 2006).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFFFF3086FB8968FC68.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral, Alcobaça municipality; South: Alentejo, Guadiana River-between Serpa and Beja municipalities (Lundblad 1956; Valdecasas 1988). Ecological notes: In oligotrophic and euritrophic slow-moving waters and sensitive to high electrolyte concentrations (absent in brackish water) (Davids et al. 2006), present in dead twigs, plants, such as grasses, Alisma spp., Myriophyllum spp. and Scirpus spp. (Lundblad 1956); Larval insects hosts: parasitic of 26 different water beetle species from families Dytiscidae, Haliplidae, Noteridae, Lacophilidae, and Hydrophilidae (Davids et al. 2006).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFFFF308383888DF91F.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Europe (Linnaeus 1758). Geographical distribution: Holarctic (Davids et al. 2006).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFFFF308383888DF91F.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, Serra da Estrela-Lagoa da Torre (Davids et al. 2005), Vilar Formoso- Côa River (Lundblad 1956); South: Alentejo (Valdecasas 1988), in Guadiana River-between Serpa and Beja municipalities (Lundblad 1956). Ecological notes: Freshwater (Linnaeus 1758), in slow-flowing and stagnant waters (Pešić 1999), often on riparian vegetation, decaying wood and leaf litter; Larval insects hosts: adults and larvae of Heteroptera and adults of Odonata (Davids et al. 2006). Found also in clear waters, present in Myriophyllum spp. and Juncus spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFCFF30808F8BA2FD48.taxon	materials_examined	Type material: Paratypes deposited in SMF, catalogue numbers 43764 - 0506 - VIE / 111, 43765 - 0519 - VIE / 111, 43784 - 0520 - VIE / 112 (GBIF 2022). Type locality: Germany: Bremen, close to peat channel (GBIF 2022; Koenike 1903). Geographical distribution: Palaearctic (Davids et al. 2006).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFCFF30808F8BA2FD48.taxon	distribution	Distribution in Portugal: South: Alentejo (Valdecasas 1988), 10 km north of Alcácer do Sal (Lundblad 1956). Ecological notes: Freshwater, with some saltwater influence, in slow-moving streams and standing waters, present in some plants including grasses (Lundblad 1956); Larval insects hosts: Dytiscidae, coleopterans (Davids et al. 2006). Hydrachna rubicunda Viets, 1930	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFCFF30808F8BA2FD48.taxon	materials_examined	Type material: Holotype deposited in SMF, catalogue number 43790 - 3819 - VIE / 112 (GBIF 2022). Type locality: Spain (GBIF 2022): Zaragoza, Balsa de la Torre Choa (Viets 1930). Geographical distribution: Iberian Peninsula (Lundblad 1956; Valdecasas 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFCFF30808F8BA2FD48.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, at standing and temporary waters, including rice field channels; present in aquatic plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A4BFFFCFF30808F8BA2FD48.taxon	discussion	Remarks: Possible synonym to Hydrachna globosa (Davids et al. 2006).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A48FFFCFF3084BA8A24F9D4.taxon	materials_examined	Type material: Neotype deposited in NRM, catalogue number NHRS-GULI 000050084 (GBIF 2022; NRM 2022). Type locality: Denmark: Gribsö (GBIF 2022; NRM 2022), based on neotype. Geographical distribution: Europe (Więcek et al. 2020).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A48FFFCFF3084BA8A24F9D4.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas, and Guadiana River-between Serpa and Beja municipalities (Lundblad 1956). Ecological notes: Freshwater environments with low alkalinity; feeds of dead aquatic insects and the adult stage individuals have a preference for Chironomidae (Diptera) egg masses (Di Sabatino et al. 2010); present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp., and Scirpus spp. (Lundblad 1956); Adapted to acidic marshes environments (Więcek et al. 2013).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A48FFFCFF3084BA8A24F9D4.taxon	discussion	Remarks: The old records of Hydrodroma despiciens constitute a species complex that has been demonstrated by morphological taxonomy and supported molecular data (Gerecke 2017; Więcek et al. 2020). Therefore, it is likely that the Lundblad records for Portugal may be Hydrodroma pilosa but this has not yet been confirmed (Pers. comm., H. Smit). We point out that there is a male in the NRM collection, Lundblad No. Präp. 4242 from the Rio Guadiana near Serpa (Pers. comm., G. Lindberg)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A48FFFDFF3080368C98FE28.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: France: probably Montpellier (Dugès 1834). Geographical distribution: Southwestern Palaearctic in Netherlands and England, and around the Mediterranean for Marocco, Italy, Spain, France, and Syria, (Dugès 1834; GBIF 2022; Gerecke 1991; Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A48FFFDFF3080368C98FE28.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, at standing and temporary waters; present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956); Larvae tend to attach to the abdomen of hydrophilid beetles, such as Laccobius moraguesi Régimbart, 1898, Hemisphaera guignoti Schäfer, 1975 and Hydrochus grandicollis Kiesenwetter, 1870 (Di Sabatino et al. 2010), as well as parasitic on odonates insects (Dugès 1834).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF3085C28D7CFB6C.taxon	materials_examined	Type material: Holotype (Schaub 1888), possibly not deposited in any scientific collection. Type locality: Europe (Schaub 1888), probably North Atlantic. Geographical distribution: Palaearctic (GBIF 2022; Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF3085C28D7CFB6C.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, at standing and temporary waters; present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956); Larval insect hosts: Ephydridae dipterans (Di Sabatino et al. 2010). Hydryphantes (Hydryphantes) placationis Thon, 1899	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF3085C28D7CFB6C.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Germany: Böhmen (Thon 1899). Geographical distribution: Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF3085C28D7CFB6C.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, in slow-flowing waters (Di Sabatino et al. 2010), standing and temporary waters; present in waters plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF308287891AF8AC.taxon	description	Synonym: Thyas thori Walter, 1907 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF308287891AF8AC.taxon	materials_examined	Type material: Deposited at NMHB, Holotype one male catalogue number NMB-ACWA 800, Paratypes one male 802, one female 801, two deuteronymph 803 and 804 (Pers. comm, U. Wüest). Type locality: Streams close to the Rhine River, between Germany-Münstertal and Switzerland-Birsfelden. Geographical distribution: Central Europe, Mediterranean.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFFDFF308287891AF8AC.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: Freshwater, at high hydrodynamic sections of streams, up to 2300 m altitude (Di Sabatino et al. 2010; Walter 1907).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFF2FF3080D38B8CFE0C.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Germany: Elbe River (Rathen), and Scheidebach River (Mittlere Erzgebirge, Annaberg-Buchholz (Piersig 1898). Geographical distribution: Western Palaearctic (Valdecasas 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A49FFF2FF3080D38B8CFE0C.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: In streams; Larval insect hosts: Chironomidae dipterans.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30856E8BD9FCBC.taxon	description	Synonym: Thyopsis cancellata maderensis Lundblad, 1941	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30856E8BD9FCBC.taxon	materials_examined	Type material: Holotype one male NHRS-GULI 000089980, and one nymph type NHRS-GULI 000089981, deposited at NRM (Pers. comm., G. Lindberg). Type locality: Portugal: Madeira Island (Lundblad 1941 b). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30856E8BD9FCBC.taxon	distribution	Distribution in Portugal: Madeira Island (Lundblad 1941 b). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30849E8ABCF83C.taxon	materials_examined	Type material: Holotype (Michael 1895), catalogue numbers (no differentiation by gender) deposited in NHMUK: 6549752 and 6550259 (GBIF 2022). Type locality: England: Cornwall, Land’s End (Michael 1895). Geographical distribution: England (Di Sabatino et al. 2010); France (Lundblad 1956); Iberian Peninsula (Borges et al. 2008; Valdecasas 1988): Portugal (Estremadura and Madeira Island); Spain (Cadiz, Gerona, Madrid); Mediterranean (Di Sabatino et al. 2010), Türkiye (Erman et al. 2010), Iran (Pešić et al. 2004).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30849E8ABCF83C.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace (Lundblad 1956); Madeira Island (Borges et al. 2008). Ecological notes: Freshwater, slow and mid streams, thin water layers (Pešić et al. 2004), with some saltwater influence (Michael 1895) and vicinity of the marine environment. Trichothyas (Lundbladia) rutae (Lundblad, 1941) Synonyms: Lundbladia rutae Lundblad, 1941 (original designation); Trichothyas (Lundbladia) petrophila rutae (Lundblad, 1941)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30849E8ABCF83C.taxon	materials_examined	Type material: Holotype one female (Lundblad 1941 b); Syntypes deposited in NRM: Male individuals, catalogue number NHRS-GULI 000006564 and NHRS-GULI 000006565; Females, NHRS-GULI 000006566 and NHRSGULI 000006567 (GBIF 2022). Type locality: Portugal: Madeira Island, in Caramujo (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Lundblad 1941 b); Spain: Canary Islands (Di Sabatino et al. 2010; Valdecasas 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A46FFF2FF30849E8ABCF83C.taxon	distribution	Distribution in Portugal: Madeira Island (Hughes et al. 1998; Lundblad 1941 b), in Caramujo (GBIF 2022). Ecological notes: Freshwater (Hughes et al. 1998)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3085B78A77F813.taxon	materials_examined	Type material: Deposited in NRM: Holotype one female NHRS-GULI 000089974 (Lundblad no. 4207), Allotype male NHRS-GULI 000089975 (Pers. comm., G, Lindberg). Type locality: Spain: Cádiz (Valdecasas 1988), in Algeciras, Andalucía (Lundblad 1954); and Cangas de Onís, Asturias (Lundblad 1956). Geographical distribution: Portugal: Beira Alta (Lundblad 1956); Spain: Cádiz, Asturias-Oviedo (Valdecasas 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3085B78A77F813.taxon	distribution	Distribution in Portugal: Centre: Santa Comba Dão, Beira Alta (Lundblad 1956). Ecological notes: Warm fast-flowing water streams, present in moss in a waterfall (Lundblad 1956). Protzia curvipalpis Lundblad, 1956	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3085B78A77F813.taxon	materials_examined	Type material: Deposited in NRM: Holotype Lundblad No. Präp. 4416; Paratype Lundblad No. Präp. 4417 and 4421 (Lundblad 1956). Type locality: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Geographical distribution: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3085B78A77F813.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: Cold freshwater, found in cascade staircase with plenty of moss, together with platyhelminth Crenobia alpina (Lundblad 1956). Protzia octopora Lundblad, 1954	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3085B78A77F813.taxon	materials_examined	Type material: Deposited in NRM: Holotype one male, Lundblad No. Präp. 4418; Paratype: Lundblad No. Präp. 4420 (Lundblad 1956). Type locality: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Geographical distribution: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3085B78A77F813.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: Cold freshwater, found in cascade staircase with plenty of moss, together with platyhelminth Crenobia alpina (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3087938B53FD9C.taxon	description	Synonyms: Thyas barbigera Viets, 1908; Todothyas incerta (Lundblad, 1941) (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3087938B53FD9C.taxon	materials_examined	Type material: Holotype deposited at SMNH (Di Sabatino et al. 2009) Type locality: Portugal, Madeira Island (Di Sabatino et al. 2009) Geographical distribution: Holarctic distribution (Di Sabatino et al. 2009)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3087938B53FD9C.taxon	distribution	Distribution in Portugal: Azores, Madeira (Bica da Cana) (Di Sabatino et al. 2009) Ecological notes: Larval host on culicid midges with high dispersal capacity (Di Sabatino et al. 2009).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A47FFF3FF3087938B53FD9C.taxon	discussion	Remarks: This species is synonymous with T. barbigera (Di Sabatino et al. 2009) since Lundblad only relied on a deuteronymph to describe the species knowing its similarity.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF3086238BF2FCBC.taxon	materials_examined	Type material: Lectotype and paralectotypes deposited at SMF, catalogue number SMF 2799 length (Gerecke 2009). Type locality: Germany: between Falkau and Saig (Schwoerbel 1957). Geographical distribution: South central Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF3086238BF2FCBC.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in front of “ Morraceira das Varandas ” Island.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF3086238BF2FCBC.taxon	materials_examined	Material examined: 0 / 1 / 0; Morraceira das Varandas, Portugal; 41 º 54 ′ 36.7 ″ N, 8 º 49 ′ 4.49 ″ W; 24 September 2020; coll. by Aquamuseu Team; catalogue number NatMIP-CATr- 0006. Ecological notes: Interstitial on springs and streams (Di Sabatino et al. 2010). In Minho River, one individual was collected in the main sector, on sediment type ‘ moderately sorted medium sand’ (Cantallo et al. 2021), in which the logarithmic main grain size value was 1.848.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF3086238BF2FCBC.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30842B8C49FB6F.taxon	discussion	Synonyms: Lebertia (Lebertia) africana Walter, 1926; Lebertia (Lebertia) affinis Sokolow, 1927; Lebertia (Lebertia) tenuistrata Viets, 1937; Lebertia (Lebertia) elongata Besseling, 1957	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30842B8C49FB6F.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Norway (Thor 1899). Geographical distribution: Western Palaearctic (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30842B8C49FB6F.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: Mainly occurring in streams and oligotrophic lakes, but even on mosses and under stones; parasitic larvae of insect-chironomids (Di Sabatino et al. 2010). In Portugal, this species was found on a cold cascade staircase, together with platyhelminth Crenobia alpina (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30825B8BBDF8C8.taxon	materials_examined	Type material: Deposited in NRM: Holotype Lundblad No. Präp. 4449; Paratype: Lundblad No. Präp. 4450 (Lundblad 1956). Type locality: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Geographical distribution: Iberian Peninsula (Valdecasas 1988), French Pyrenees (Angelier et al. 1985).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30825B8BBDF8C8.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), inValença municipality-São Pedro da Torre; Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30825B8BBDF8C8.taxon	materials_examined	Material examined: 1 / 1 / 0; São Pedro da Torre, Portugal; 41 º 59 ′ 42.05 ″ N, 8 º 40 ′ 27.94 ″ W; 24 September 2020; coll by Aquamuseu Team; catalogue number NatMIP-CATr- 0002. Ecological notes: Cold freshwater, found in cascade staircase with plenty of moss, together with platyhelminth Crenobia alpina (Lundblad 1956); Possibly rhithrobiont-like (on river and tributaries habitats) species (Di Sabatino et al. 2010). In Minho River, two individuals were collected in the main sector, on sediment type ‘ moderately sorted coarse sand’ (Cantallo et al. 2021), in which the logarithmic main grain size value was 0.633.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF0FF30825B8BBDF8C8.taxon	discussion	Remarks: First record mentioned for Minho River (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF1FF3080378BF2FE28.taxon	discussion	Synonyms: Lebertia (Neolebertia) sublitoralis Walter, 1922; Lebertia (Lebertia) shadini Sokolow 1940	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF1FF3080378BF2FE28.taxon	materials_examined	Type material: D eposited at Koenike’s collection, catalogue number Ppt. 1112 (Koenike 1911). Type locality: Germany: Delme River, close to Grossherzogtum Oldenburg, Delmenhorst (Koenike 1911). Geographical distribution: Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF1FF3080378BF2FE28.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in Vila Nova de Cerveira municipality.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF1FF3080378BF2FE28.taxon	materials_examined	Material examined: 0 / 1 / 0; Vila Nova de Cerveira, Portugal; 41 º 56 ′ 12.768 ″ N, 8 º 45 ′ 9.108 ″ W; October 2019; coll. by C. Antunes and E. Martins; catalogue number NatMIP-CATr- 0001. Ecological notes: On the sandy bottom of low land streams and lakes (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A44FFF1FF3080378BF2FE28.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF308696893BFC20.taxon	discussion	Synonym: Lebertia (Mixolebertia) schmidtii Thor, 1911	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF308696893BFC20.taxon	materials_examined	Type material: Holotype (Koenike 1902), possibly not deposited in any scientific collection. Type locality: Ireland: Dartrey (Koenike 1902). Geographical distribution: North Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF308696893BFC20.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in Valença municipality-São Pedro da Torre.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF308696893BFC20.taxon	materials_examined	Material examined: 0 / 0 / 1; Pedro da Torre, Portugal; 41 º 59 ′ 42.05 ″ N, 8 º 40 ′ 27.94 ″ W; 24 September 2020; coll by Aquamuseu Team; catalogue number NatMIP-CATr- 0007. Ecological notes: In standing waters (Di Sabatino et al. 2010). In Minho River, one individual was collected in the main sector, on sediment type ‘ moderately sorted coarse sand’ (Cantallo et al. 2021), in which logarithmic main grain size value was 0.633.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF308696893BFC20.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021), and also represents the first occurrence for Southern Europe.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF30849E8C24FAB4.taxon	discussion	Synonym: Hexalebertia madericola Lundblad, 1942 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF30849E8C24FAB4.taxon	materials_examined	Type material: Deposited in NRM: Male individuals, catalogue numbers NHRS-GULI 000006556 and NHRSGULI 000006557; Females, NHRS-GULI 000006558 and NHRS-GULI 000006559 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1942), in Rabaçal (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF30849E8C24FAB4.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1942), Levada do Risco, Lomo de Mouro (Gerecke 2009), Levada Faja do Rodrigues, Pico Ruivo do Paul da Serra, Bica da Cana (García-Jiménez et al. 2017), Rabaçal (GBIF 2022). Ecological notes: Freshwater environment (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF3082128C24F91C.taxon	distribution	Synonym: Hexalebertia maderigena Lundblad, 1942 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF1FF3082128C24F91C.taxon	materials_examined	Type material: Deposited in NRM: Male individuals, catalogue numbers NHRS-GULI 000006560 and NHRSGULI 000006561; Females, NHRS-GULI 000006562 and NHRS-GULI 000006563 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1942), in Risco waterfall (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1942), in Risco waterfall (GBIF 2022). Ecological notes: Freshwater environment (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF6FF30818A8BF2FE98.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Italy (Maglio 1924). Geographical distribution: Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF6FF30818A8BF2FE98.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in Valença municipality-São Pedro da Torre.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF6FF30818A8BF2FE98.taxon	materials_examined	Material examined: 0 / 1 / 0; São Pedro da Torre, Portugal; 41 º 59 ′ 42.05 ″ N, 8 º 40 ′ 27.94 ″ W; 24 September 2020; coll. by Aquamuseu Team; catalogue number NatMIP-CATr- 0004. Ecological notes: On lakes and rivers (Di Sabatino et al. 2010). In Minho River, one individual was collected in the main sector, on sediment type ‘ moderately sorted coarse sand’ (Cantallo et al. 2021), in which the logarithmic main grain size value was 0.633.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A45FFF6FF30818A8BF2FE98.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF6FF3086068AD5FC20.taxon	discussion	Synonyms: Lebertia porosa obscura Thor, 1900; Lebertia vigintimaculata Thor, 1900; Lebertia porosa britannica Thor, 1906; Lebertia porosa dorsalis Thor, 1906; Lebertia porosa italica Thor, 1906; Lebertia pachydermis Koenike, 1908; Lebertia lacertosa Koenike, 1911; Lebertia seclusa Koenike, 1914; Lebertia rivalis Koenike, 1918; Lebertia violacea Viets, 1921; Lebertia curvata Viets, 1925; Lebertia leioderma Viets, 1925; Lebertia gibbosa Lundblad, 1926; Lebertia porosa distans Thor 1926; Lebertia riabuschinskii Thor, 1926; Lebertia caucasica Sokolow, 1927; Lebertia jenissejensis Sokolow, 1930; Lebertia valenciana Viets, 1930; Lebertia pachypalpis Sokolow, 1934; Lebertia stackelbergi Sokolow, 1934; Lebertia stackelbergi saxicola Sokolow, 1936; Lebertia caucasica cubanica Sokolow, 1940; Lebertia validipes Lundblad, 1942; Lebertia crassipalpis Halbert, 1944; Lebertia rotunda Halbert, 1944; Lebertia hibernica Viets, 1949; Lebertia heterotricha Láska, 1951.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF6FF3086068AD5FC20.taxon	materials_examined	Type material: Paratypes: deposited in SMF, catalogue numbers 45001 - 1691 - VIE / 174 and 45002 - 1693 - VIE / 174 (GBIF 2022). Type locality: Norway: Lister-spring close to Vanse Kirche (Thor 1900). Geographical distribution: Holarctic (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF6FF3086068AD5FC20.taxon	distribution	Distribution in Portugal: Madeira, In Moinhos-Ribeiro Frio (GBIF 2022). Ecological notes: Freshwater, springs, flowing and standing water bodies; also found in eutrophic waters (Van Der Hammen & Smit 1996; Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	materials_examined	Type material: Holotype Lundblad No. Präp. 4457, deposited in NRM (Lundblad 1956). Type locality: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956). Geographical distribution: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956). Ecological notes: Freshwater, at slow-moderate flowing streams, remarkably on a sandy substrate; present in Carex spp. plants (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	discussion	Remarks: Since Frontipoda Koenike, 1891 is a synonym with Oxus Kramer, 1877 (Besch 1964), we recommend Frontipoda hastata Lundblad, 1954 as a junior synonym with Oxus hastata (Lundblad, 1954). Oxus lusitanicus Lundblad, 1954	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	materials_examined	Type material: Holotype (Lundblad 1954), Lundblad No. Präp. 4456, deposited in NRM (Lundblad 1956). Type locality: Portugal: Beira Alta (Valdecasas 1988), Côa River (Lundblad 1954), near 10 km to Vilar Formoso (Lundblad 1956). Geographical distribution: Portugal: Beira Alta (Valdecasas 1988), Côa River (Lundblad 1954), near 10 km to Vilar Formoso (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), Côa River (Lundblad 1954), near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: Freshwater, at the riverine clear water, present in Myriophyllum spp. and Juncus spp. (Lundblad 1956). Oxus oblongus Kramer, 1879	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	materials_examined	Type material: Holotype (Kramer 1879), possibly not deposited in any scientific collection. Type locality: Germany: Schleusingen (Kramer 1879). Geographical distribution: Germany (Kramer 1879),	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral, in Alcabaça municipality; South: Estremadura, in Sintra-Monserrate Park and Palace (Lundblad 1956; Valdecasas 1988). Ecological notes: Freshwater, at temporary pond waters and slow-moderate motion streams (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A42FFF7FF3083CB892CFE7D.taxon	discussion	Remarks: When Lundblad described the specimens he found in Portugal, Spain and France he emphasized that they could be aberrant forms, similar to Oxus oblongus (Lundblad 1956). Later Oxus oblongus was mentioned as a possible synonym of Oxus strigatus (Di Sabatino et al. 2010; Smit & Gerecke 2010) but this has not been fully clarified either.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	materials_examined	Type material: Holotype (Piersig 1896), possibly not deposited in any scientific collection. Type locality: Germany: Drebabach and Plothenbach streams, Thüringen (Piersig 1896). Geographical distribution: Western Palaearctic (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: Freshwater, mainly found in fast-flowing rivers (Stryjecki et al. 2018) and organic pollutiontolerant species; Larval insect hosts: Chironomidae dipterans (Di Sabatino et al. 2010). Sperchon (Hispidosperchon) setiger Thor, 1898	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	materials_examined	Type material: Paratype: Deposited in SMF, catalogue number 47270 - 1552 - VIE / 289 (GBIF 2022). Type locality: Norway (GBIF 2022): Roslandsaa, Jaederen (Thor 1898 b). Geographical distribution: Western Palaearctic, remarkably Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), Alcobaça municipality (Lundblad 1956). Ecological notes: Mainly found on streams with a reduced flow; Larval insect hosts: Simuliidae and Chironomidae dipterans (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	discussion	Remarks: The species Sperchon meridianus Angelier, 1954 was considered a synonym of S. setiger, which was recently verified as valid species based on molecular data (Pešić & Smit 2022). Sperchon (Sperchon) brevirostris Koenike, 1895	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	materials_examined	Type material: Holotype (Koenike 1895 b), possible not deposited in any scientific collection. Paratype: Individual deposited in SMF, catalogue number 47069 - 1559 - VIE / 279 (GBIF 2022). Type locality: Portugal: Azores Archipelago (Koenike 1895 b). Geographical distribution: Europe (GBIF 2022; Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	distribution	Distribution in Portugal: Azores Archipelago (Koenike 1895 b), including Terceira Island, and São Miguel Island (Lagoa do Congro, Lagoa das Furnas, Lagoa do Fogo) (Barrois 1896); Madeira Island (Borges et al. 2008; Hughes et al. 1998). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998), in flowing tributaries and standing water bodies (Barrois 1896; Di Sabatino et al. 2010); Stenothermic species (Pešić 1999). Sperchon (Sperchon) glandulosus Koenike, 1886	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	materials_examined	Type material: Deposited in SMF: Holotype (Koenike 1886), catalogue number 47174 - 2188 - VIE / 284; Paratype, catalogue number 47171 - 1562 - VIE / 284 (GBIF 2022). Type locality: Germany: Bremen (Koenike 1886). Geographical distribution: Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	distribution	Distribution in Portugal: Azores Archipelago: Faial, São Miguel and Terceira Islands (Barrois 1887). Ecological notes: Freshwater, in tributaries and eventually lakes near to coast; Larval insects hosts: Chironomidae dipterans (Di Sabatino et al. 2010). Sperchon (Sperchon) thienemanni Koenike, 1907 Synonym: Sperchon rugosus Koenike, 1911	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	materials_examined	Type material: Possibly not deposited in any scientific collection. Type locality: Germany, in Jasmund Peninsula-Rügen Island (Koenike 1907). Geographical distribution: Europe (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), tributary section “ Rio Mouro, Ponte do Curto ” (Monção municipality, 42 º 4 ′ 29.27 ″ N, 8 º 23 ′ 39.12 ″ W).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	materials_examined	Material examined: 1 undetermined sex adult; Ponte do Curto, Portugal; 42 º 4 ′ 29.27 ″ N, 8 º 23 ′ 39.12 ″ W; 24 September 2020; coll by Aquamuseu Team; catalogue number NatMIP-CATr- 0014. Ecological notes: Freshwater, common in rheohelocrenes springs, with crenophilous-type habit; larvae parasites of insect-chironomids (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A43FFF4FF3084648BF2FC9F.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A40FFF4FF3084FF8903FA24.taxon	materials_examined	Type material: Possible type material: ZMB / Arach 285343 (GBIF 2022). Type locality: Germany, in Brandenburg (Protz 1896). Geographical distribution: Holarctic (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A40FFF4FF3084FF8903FA24.taxon	distribution	Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in the tributary section “ Rio Mouro, Ponte do Curto ” (Monção municipality, 42 º 4 ′ 29.27 ″ N, 8 º 23 ′ 39.12 ″ W); Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A40FFF4FF3084FF8903FA24.taxon	materials_examined	Material examined: 1 undetermined sex adult; Ponte do Curto, Portugal; 42 º 4 ′ 29.27 ″ N; 24 September 2020; coll. by Aquamuseu Team; catalogue number NatMIP-CATr- 0012. Ecological notes: On muddy sediment in streams, feeds on chironomid larvae (Di Sabatino et al. 2010). Present in macrophytes (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A40FFF4FF3084FF8903FA24.taxon	discussion	Remarks: First record mentioned for Minho River (Cantallo et al. 2021), and also represents the first occurrence for North Portugal.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A40FFF4FF3081CF888DF83B.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Germany, in Lake Issinger Weiher, close to Regensburg municipality (Koch 1837). Geographical distribution: Palaearctic (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A40FFF4FF3081CF888DF83B.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Colares, Sintra municipality (Lundblad 1956). Ecological notes: In cold lakes and shaded tributaries (Di Sabatino et al. 2010), covered with Azolla spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFF5FF3086238D03FBB0.taxon	description	Synonyms: Atractides lusitanicus Lundblad, 1941 (original designation); Torrenticola (Monatractides) lusitanica Lundblad, 1956	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFF5FF3086238D03FBB0.taxon	materials_examined	Type material: Holotype (Lundblad 1941 a), possibly not deposited in any scientific collection. Type locality: Portugal: Convent of the Capuchos, Sintra municipality (Lundblad 1941 a). Geographical distribution: Iberian Peninsula (Valdecasas 1988), Mediterranean and Iran (Erman et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFF5FF3086238D03FBB0.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Convent of the Capuchos, Sintra municipality (Lundblad 1941 a). Ecological notes: Freshwater, mainly small streams, with dead leaves, twigs and little moss in a deeply cut ravine in the coniferous forest (Lundblad 1956). Monatractides (Monatractides) stenostomus (Viets, 1930) Synonyms: Atractides stenostomus Viets, 1930 (original designation); Torrenticola (Monatractides) stadleri var. stenostoma (Viets, 1930); Torrenticola (Monatractides) stenostoma (Viets, 1930)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFF5FF3086238D03FBB0.taxon	materials_examined	Type material: Holotype (Viets 1930), with catalogue number SMF 47795 at SMF (Pers. comm., R. Gerecke). Type locality: Spain: Madrid (Valdecasas 1988), in Manzanares River, Charca de Marmoto (Viets 1930). Geographical distribution: France and Iberian Peninsula (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFF5FF3086238D03FBB0.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1956). Ecological notes: Freshwater, in tributaries (Lundblad 1956) with slow-moderate and warm streams, on a sandystone substrate (Di Sabatino et al. 2010), as well as on sandy bottom and present in aquatic plants (Viets 1930).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Possibly not deposited in any scientific collection (Martin 2009). Type locality: French Pyrenees (Angelier 1949; Valdecasas 1988). Geographical distribution: French Pyrenees (Angelier 1949; Martin 2009; Valdecasas 1988) and Portugal (Cantallo et al. 2021). Distribution in Portugal: North: International Minho River (Cantallo et al. 2021), in Vila Nova de Cerveira municipality-Lovelhe, close to Marina da Lenta. Material examined: 1 / 0 / 0; Lovelhe; 41 º 57 ′ 2.39 ″ N, 8 º 44 ′ 51.5 ″ W; 24 September 2020; coll. by Aquamuseu Team; catalogue number NatMIP-CATr- 0008. Ecological notes: In low-order streams in mountain areas (Di Sabatino et al. 2010); In Minho River, one individual was collected in the main sector, on sediment type ‘ moderately sorted coarse sand’ (Cantallo et al. 2021), in which logarithmic main grain size value was 0.575.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021). Torrenticola (Torrenticola) affinis (Lundblad, 1941) Synonym: Atractides affinis Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Syntypes deposited in NRM: Males individuals, catalogue numbers NHRS-GULI 000006514 and NHRS-GULI 000006515; Females, NHRS-GULI 000006516 and NHRS-GULI 000006517 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Caramujo (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) anomala (Koch, 1837) Synonyms: Atractides anomalus Koch, 1837 (original designation); Torrenticola longirostris Szalay, 1933; Torrenticola piriformis Láska, 1953 Type material: Unknown, possibly lost. Type locality: Germany (Koch 1837). Geographical distribution: Holarctic (Erman et al. 2010; Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	distribution	Distribution in Portugal: North: International Minho River Basin (Cantallo et al. 2021), in Mouro River: Monção municipality-Ponte do Curto. Geographical distribution: Holarctic (Erman et al. 2010; Di Sabatino et al. 2010). Distribution in Portugal: North: International Minho River Basin (Cantallo et al. 2021), in Mouro River: Monção municipality-Ponte do Curto.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Material examined: 1 undetermined sex adult; Ponte do Curto, Portugal; 42 º 4 ′ 29.27 ″ N, 8 º 23 ′ 39.12 ″ W; 24 September 2020; coll by Aquamuseu Team; catalogue number NatMIP-CATr- 0009. Ecological notes: In middle-order streams (Erman et al. 2010), with the ability to withstand a wide range of temperatures — eurythermal species (Di Sabatino et al. 2010).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	discussion	Remarks: First record mentioned for Portugal (Cantallo et al. 2021). Torrenticola (Torrenticola) crassa (Lundblad, 1941) Synonym: Atractides crassus Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Holotype one male NHRS-GULI 000089982, and one female Allotype NHRS-GULI 000089983, deposited at NRM (Pers. comm., G. Lindberg). Type locality: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) crassirostris (Lundblad, 1941) Synonym: Atractides crassirostris Lundblad, 1941 (original designation) Type material: Syntypes deposited in NRM: Males individuals, catalogue numbers NHRS-GULI 000006510 and NHRS-GULI 000006511; Females, NHRS-GULI 000006512 and NHRS-GULI 000006513 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Caramujo (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Distribution in Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) elliptiformis (Lundblad, 1941) Synonym: Atractides elliptiformis Lundblad, 1941 (original designation) Type material: Syntypes deposited in NRM: Males individuals, catalogue numbers NHRS-GULI 000006526 and NHRS-GULI 000006527; Females, NHRS-GULI 000006528 and NHRS-GULI 000006529 (GBIF 2022). Type locality: Portugal: Madeira Island, in Vinte e Cinco Fontes, Levada (GBIF 2022; Lundblad 1941 b). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998), in Vinte e Cinco Fontes, Levada (GBIF 2022). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) insulicola (Lundblad, 1941) Synonym: Atractides insulicola Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Holotype and paratypes deposited in NRM: Males individuals, catalogue numbers NHRS-GULI 000006501 and NHRS-GULI 000006502; Females, NHRS-GULI 000006503 and NHRS-GULI 000006504 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Caramujo (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Distribution in Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) mandibularis (Lundblad, 1941) Synonym: Atractides mandibularis Lundblad, 1941 (original designation) Type material: Holotype one male NHRS-GULI 000097846, and one female Allotype NHRS-GULI 000097847, deposited at NRM (Pers. comm., G. Lindberg). Type locality: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) nesiotes (Lundblad, 1941) Synonym: Atractides nesiotes Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Holotype and paratypes deposited in NRM: Males individuals, catalogue numbers NHRSGULI 000006522 and NHRS-GULI 000006523; Females, NHRS-GULI 000006524 and NHRS-GULI 000006525 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Vinte e Cinco Fontes, Levada (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) pharyngealis (Lundblad, 1941) Synonym: Atractides pharyngealis Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Holotype (Lundblad 1941 b); Holotype and paratypes deposited in NRM: Male individuals, catalogue numbers NHRS-GULI 000006505 and NHRS-GULI 000006506; Females, NHRS-GULI 000006507, NHRS-GULI 000006508 and NHRS-GULI 000006509 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Ribeiro do Inferno (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b), in Ribeiro do Inferno (GBIF 2022). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) rotunda (Lundblad, 1941) Synonym: Atractides rotundus Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	materials_examined	Type material: Holotype and paratypes deposited in NRM, Male individuals: catalogue numbers NHRSGULI 000006518 and NHRS-GULI 000006519; Females: NHRS-GULI 000006520 and NHRS-GULI 000006521 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), Caramujo (GBIF 2022). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; GBIF 2022; Hughes et al. 1998). Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Torrenticola (Torrenticola) tenuipalpis (Lundblad, 1956) — nov. stat. Synonym: Torrenticola (Torrenticola) amplexa var. tenuipalpis Lundblad, 1956 (original designation) Type material: Holotype deposited in NRM, Lundblad No. Präp. 4470 (Lundblad 1956). Type locality: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1956). Geographical distribution: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1956). Ecological notes: Freshwater, at slow-moderate flowing streams, remarkably on a sandy substrate; present in Carex spp. plants (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A41FFEBFF3083888CF6F852.taxon	discussion	Remarks: We propose to higher the ranking of Torrenticola (Torrenticola) amplexa to that of a full species given the elongated P. IV. A record from North America (Quaglia & Conroy 1984) needs revision.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE8FF3086FA8A77FCBC.taxon	materials_examined	Type material: Holotype (Lundblad 1954), deposited at NRM, catalogue number NHRS-GULI 000002776 (GBIF 2022). Type locality: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada (Lundblad 1956), Buçaco mountain (GBIF 2022). Geographical distribution: Portugal: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada (Lundblad 1956), Buçaco mountain (GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE8FF3086FA8A77FCBC.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada (Lundblad 1956), Buçaco mountain (GBIF 2022). Ecological notes: Cold freshwater, found on cascade staircase with plenty of moss, together with platyhelminth Crenobia alpina (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE8FF30849E8931FB40.taxon	materials_examined	Type material: Holotype (Lundblad 1954) deposited in NRM, Lundblad No. Präp. 4514 (Lundblad 1956). Type locality: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956). Geographical distribution: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE8FF30849E8931FB40.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956). Ecological notes: Freshwater, in slow-moderate flowing streams, on a sandy substrate; present in Carex spp. plants (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE8FF3082128AA9F938.taxon	materials_examined	Type material: Females (Piersig 1903), possibly not deposited in any scientific collection. Type locality: Germany: Bohemian Forest, Mauth; Kleine Mittweidastream, Ore Mountains, near Nitzschhammer (Piersig 1903). Geographical distribution: Central and Southeastern Europe (Gerecke et al. 2016; Szalay 1970).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE8FF3082128AA9F938.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa CombaDão (Lundblad 1956). Ecological notes: The species lives at slow-moderate flowing streams, on sandy substrate, present in Carex spp. plants (Lundblad 1956), in the plains (e. g. lakes), pools of low mountain ranges, as well as in the interstitial and groundwaters (Gerecke et al. 2016; Szalay 1970).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE9FF30801B8CF9FEE0.taxon	materials_examined	Type locality: Tunisia: Oued Djoumine stream, Mateur (Walter 1928). Geographical distribution: Southwestern Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5CFFE9FF30801B8CF9FEE0.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), Côa River, near 10 km to VilarFormoso (Lundblad 1956). Ecological notes: Freshwater, lentic waters (e. g. pools) and interstitial habitats (Gerecke et al. 2016), as well as at riverine clear water, present in plants Myriophyllum spp. and Juncus spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFE9FF3086B38AD1FCBC.taxon	materials_examined	Type material: Holotype (Thor 1898 a), possibly not deposited in any scientific collection. Type locality: Denmark: Kristiania, nowadays Oslo (Thor 1898 a). Geographical distribution: Holarctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFE9FF3086B38AD1FCBC.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: Present on low-order small rivers, including stream rheocrenes; Larval insect hosts: Chironomidae dipterans (Gerecke et al. 2016). In Portugal, this species was found on a cold cascade staircase, together with platyhelminth Crenobia alpina (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	description	Synonym: Maderomegapus hystricipes Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Holotype one male NHRS-GULI 000089969 and one female Allotype NHRS-GULI 000089970 deposited at NRM (Pers. comm., G. Lindberg). Type locality: Portugal: Madeira Island (Lundblad 1941 b) Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998). Atractides (Atractides) insulanus (Lundblad, 1941) Synonym: Megapus insulanus Lundblad, 1941 (original designation) Type material: Deposited in NRM: Holotype one male (Gerecke 2003; Lundblad 1941 b), catalogue number NHRS-GULI 000006542 (GBIF 2022); Paratype: (Gerecke 2003), catalogue number NHRS-GULI 000006544 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Ribeira da Janela waterfall (Gerecke 2003). Geographical distribution: Portugal: Madeira Island (GBIF 2022; Gerecke 2003).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b), in Ribeira da Janela waterfall (Gerecke 2003). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998), springs (rhithrobiontic habit), in thin layers of flowing water (Gerecke 2003). Atractides macaronensis (Lundblad, 1941) Synonym: Megapus macaronensis Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Deposited in NRM: Holotype one male (Lundblad 1941 b), catalogue number NHRS-GULI 000006536 (GBIF 2022); Paratype catalogue number NHRS-GULI 000006537 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Vinte e Cinco Fontes, Levada (GBIF 2022; Gerecke 2003). Geographical distribution: Portugal: Madeira Island (GBIF 2022; Gerecke 2003).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998), in Vinte e Cinco Fontes, Levada (GBIF 2022; Gerecke 2003). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998), rhithrobiont (river and tributary environment), in thin layers of flowing water, preference at rocky habitats (Gerecke 2003). Atractides maderensis (Lundblad, 1941) Synonyms: Megapus maderensis Lundblad, 1941 (original designation	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Deposited in NRM: Holotype one male (Gerecke 2003; Lundblad 1941 b), catalogue number NHRSGULI 000006546 (GBIF 2022); Paratype: (Gerecke 2003), catalogue number NHRS-GULI 000006547 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Ribeira da Janela, Risco; Vinte e Cinco Fontes, Levada (GBIF 2022; Gerecke 2003). Geographical distribution: Portugal: Madeira Island (GBIF 2022; Gerecke 2003).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b), in Ribeira da Janela, Risco; Vinte e Cinco Fontes, Levada (GBIF 2022; Gerecke 2003). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998), found in springs and river habitats (rhithrobiontic species) (Gerecke 2003). Atractides (Polymegapus) rutae (Lundblad, 1941) Synonyms: Megapus rutae Lundblad, 1941 (original designation); Megapus (Allomegapus) rutae Lundblad, 1941	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Deposited in NRM: Holotype one male (Lundblad 1941 b), catalogue number NHRS-GULI 000006538 (GBIF 2022); Paratypes catalogue number NHRS-GULI 000006539 (GBIF 2022); one female catalogue number NHRS-GULI 000006540 (GBIF 2022). Type locality: Portugal: Madeira Island (Lundblad 1941 b), in Rabaçal (GBIF 2022; Gerecke 2003). Geographical distribution: Portugal: Madeira Island (GBIF 2022; Gerecke 2003; Lundblad 1941 b); Spain: Canary Islands, in Tenerife (Lundblad 1962; Valdecasas 1988).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: Madeira Island (Lundblad 1941 b), in Rabaçal (GBIF 2022; Gerecke 2003). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998), springs, and thin layers of flowing water (Gerecke 2003). Atractides (Atractides) clavipes Lundblad, 1954	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Holotype (Lundblad 1954), Lundblad No. Präp. 4500, deposited in NRM (Lundblad 1956). Type locality: Portugal: Sintra-Monserrate Park and Palace (Lundblad 1954, 1956). Geographical distribution: Germany, France, Iberian Peninsula and Italy (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra, Monserrate Park and Palace (Lundblad 1956). Ecological notes: Crenobiont species: at springs habitats (Gerecke et al. 2016) and small streams, present in plant Alsophila spp. (Lundblad 1956). Atractides (Atractides) nodipalpis Thor, 1899 Synonyms: Megapus nodipalpis Thor, 1899 (original designation); Megapus tenuitarsus Komárek, 1921; Megapus nodipalpis parviporus Besseling, 1933; Atractides moravicus Láska, 1952; Atractides limnicola Schwoerbel, 1959; Atractides microcavaticus Schwoerbel, 1961; Atractides schlienzi Schwoerbel, 1961	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Paratypes 3 specimen deposited in SMF, catalogue numbers 42618 - 1516 - VIE / 053, 42619 - 1517 - VIE / 053, 42620 - 1518 - VIE / 053 (GBIF 2022). Type locality: Norway: Gudbrandsdalen (GBIF 2022; Gerecke 2003; Lundblad 1956). Geographical distribution: Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1956). Ecological notes: Warm fast-flowing streams, present in moss in a waterfall (Lundblad 1956). Organic pollutiontolerant species with rhithrobiont freshwater preferences (river and tributaries environment); Larval insect hosts: Chironomidae dipterans (Gerecke et al. 2016). Atractides (Tympanomegapus) ibericus Lundblad, 1956 Synonym: Atractides lacustris ibericus Lundblad, 1956 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	materials_examined	Type material: Holotype one male Lundblad No. Präp. 4551, deposited in NRM (Lundblad 1956). Type locality: Portugal: Santa Comba Dão, Beira Alta (Lundblad 1956). Geographical distribution: Portugal: Santa Comba Dão, Beira Alta (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5DFFEFFF3083568931FE0C.taxon	distribution	Distribution in Portugal: Centre: Santa Comba Dão, Beira Alta (Lundblad 1956). Ecological notes: Freshwater, in slow-moderate motion streams, on a sandy substrate; present in Carex spp. plants (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	materials_examined	Type material: One adult male deposited in NTNU-VM, catalogue number VP-Hyd 712 (Pešić et al. 2017) Type locality: Norway: Heltneelva (GBIF 2022; Ström 1768) Geographical distribution: Palaearctic (GBIF 2022; Ström 1768)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1956). Ecological notes: Freshwater rhithrobiont-like (on river and tributaries habitats); Organic pollution tolerant-species with preferences for streams and rheocrenes in lowland and low mountain ranges; Larval insect hosts: Chironomidae dipterans (Gerecke et al. 2016). Found also on a sandy substrate; present in Carex spp. plants (Lundblad 1956) Hygrobates (Hygrobates) longipalpis (Hermann, 1804) Synonym: Hydrarachna longipalpis Hermann, 1804 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	materials_examined	Type material: Neotype designated by Pešić et al. 2019 RMNH (Pešić et al. 2019). Type locality: Netherlands: Gelderland (GBIF 2022). Geographical distribution: Holarctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral (Valdecasas 1988), in Fonte Fria, Mealhada, Buçaco mountain (Lundblad 1956). Ecological notes: In standing or slow-flowing waters (Gerecke et al. 2016; Lundblad 1956); Larval insect hosts: Chaoboridae and Chironomidae dipterans (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	discussion	Remarks: Given the recent re-establishment of the species Hygrobates prosiliens Koenike, 1915 this observation requires further analysis, as the first inhabits in standing water while H. longipalpis has a preference for slowflowing water (Pešić et al. 2019). Hygrobates (Hygrobates) longiporus Thor, 1898 Synonym: Hygrobates falcilaminatus Walter, 1926	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	materials_examined	Type material: Possibly not deposited in any scientific collection. Type locality: Norway (Thor 1898 b). Geographical distribution: Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	distribution	Distribution in Portugal: Centre: Beira Litoral, in Alcabaça municipality; and Beira Alta, in Santa Comba Dão; and Côa River, near 10 km to Vilar Formoso (Lundblad 1956; Valdecasas 1988). Ecological notes: In standing freshwaters, such as lakes (Gerecke et al. 2016), clear water, on sandy and muddy bottoms, with scattered leaves, dead twigs, plants, e. g. grasses, Carex spp. and Alisma spp. (Lundblad 1956). Hygrobates (Hygrobates) setosus Besseling, 1942 Synonym: Hygrobates nigromaculatus setosus Besseling, 1942 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	materials_examined	Type material: Female specimens in the “ Collectie-Romijn ” (Besseling 1942), nowadays deposited in RMNH registration number ZMA. ACA. P. 35358 and ZMA. ACA. P. 35360 (GBIF 2022). Type locality: Netherlands: Dorpsbeekte Epe (Besseling 1942; GBIF 2022). Geographical distribution: Palaearctic (Gerecke et al. 2016; Smit & van der Hammen 2000).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	distribution	Distribution in Portugal: North: International Minho River Basin: Monção (tributaries, “ Rios Gadanha e Mouro ”) and Valença (main river stretch) municipalities (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	materials_examined	Material examined: 2 / 0 / 0; Pinheiros, Portugal; 42 º 2 ′ 14.7 ″ N, 8 º 30 ′ 6,9 ″ W; 25 September 2020; coll. by Aquamuseu Team; catalogue number NatMIP-CATr- 0013. 0 / 1 / 0; Valença, Portugal; 42 º 2 ′ 57.01 ″ N, 8 º 37 ′ 54.74 ″ W; 24 September 2020; coll. By Aquamuseu Team; catalogue number NatMIP-CATr- 0015. 0 / 1 / 0; Ponte de Mouro; 42 ° 04 ′ 30.0 ″ N, 8 ° 23 ′ 38.5 ″ W; 24 September 2020; coll. by Aquamuseu Team; catalogue number NatMIP-CATr- 0016. Ecological notes: In freshwater tributaries pools; with parasitic larvae of insect-dipterans (Gerecke et al. 2016). In the present study, this species was found in association with macrophytes (“ Rio Gadanha ”). One specimen was found in a rock environment close to the spring (“ Rio Mouro ”). One individual was collected in the main sector from the Minho River, on sediment type ‘ moderately sorted coarse sand’, in which the logarithmic main grain size value was 0.345 (Cantallo et al. 2021).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	discussion	Remarks: This species was reported for Portugal by Cantallo et al. (2021), which also represents the first occurrence for Southern Europe. Hygrobates (Hygrobates) trigonicus Koenike, 1895 Synonym: Hygrobates properus Láska, 1954	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	materials_examined	Type material: Holotype (Koenike 1895 a), possibly not deposited in any scientific collection. Type locality: Poland: Słupice (Koenike 1895 a). Geographical distribution: Western Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A5BFFECFF30855B8C23FC68.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Santa Comba Dão, and Côa River, near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: Freshwater, rhithrobiont-like species, found in river and tributary environments (Gerecke et al. 2016), in clear water, on a sandy substrate; present in plants, such as Carex spp., Juncus spp., Myriophyllum spp. (Lundblad 1956); Larval insect hosts: Chironomidae dipterans (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFECFF3082458C67F909.taxon	materials_examined	Type material: Holotype one male catalogue number NMB-ACWA 2713 deposited at NHMB (Pers. comm, U. Wüest) Type locality: Algeria: Jijel Province, Oued Nil (Walter 1925). Geographical distribution: European (France, Iberian Peninsula and Italy) and North African Mediterranean area (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFECFF3082458C67F909.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (in Santa Comba Dão, and Côa River, near 10 km to Vilar Formoso); South: Alentejo (in Ribeira de Odivelas, Ribeira do Roxo, and Guadiana River-between Serpa and Beja municipalities), Estremadura (in Colares-Sintra municipality) (Lundblad 1956; Valdecasas 1988). Ecological notes: Freshwater, at standing and slow-flowing freshwater ecosystems (Gerecke et al. 2016), shaded and clear tributaries, on a sandy bottom; present in plants, such as grasses Azolla spp., Batrachium spp., Carex spp., Juncus spp., Myriophyllum spp., Nymphea spp., Scirpus spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Holotype one female (Viets 1918), Syntypes deposited in SMF, catalogue numbers 45385 - 2009 - VIE / 193, 45386 - 2010 - VIE / 193, 45387 - 2011 - VIE / 193 (GBIF 2022). Type locality: Spain (GBIF 2022): Valencia (Valdecasas 1988; Viets 1918). Geographical distribution: Iberian Peninsula, Canary Islands, South France, Italy and North Africa (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace (Lundblad 1956). Ecological notes: Freshwater, at temporary and small permanent waters (Gerecke et al. 2016; Viets 1918). Limnesia (Limnesia) atlantica Lundblad, 1941	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Holotype one male NHRS-GULI 000089977, one female Allotype NHRS-GULI 000089978, and one nymph type NHRS-GULI 000089979 deposited at NRM (Pers. comm., G. Lindberg). Type locality: Portugal: Madeira Island (Lundblad 1941 b). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998; Lundblad 1941 b).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998). Ecological notes: Freshwater environment (Borges et al. 2008; Hughes et al. 1998). Limnesia (Limnesia) fulgida Koch, 1836	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Unknown, possibly lost. Geographical distribution: Holarctic (Gerecke et al. 2016; Smit & Van der Hammen 2000).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Gerecke et al. 2016; Smit & Van der Hammen 2000). Ecological notes: At standing freshwaters; larvae parasites of Chironomidae and Sphaeroceridae dipterans (Gerecke et al. 2016); present in plants, such as Batrachium spp., Myriophyllum spp., and Nymphea spp. (Lundblad 1956). Limnesia (Limnesia) iberica Lundblad, 1954	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Holotype one male deposited in NRM, Lundblad No. Präp. 4478 (Lundblad 1954); Paratype: One female, Lundblad No. Präp. 4479 (Lundblad 1956). Type locality: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956). Geographical distribution: Portugal: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1954, 1956). Ecological notes: Freshwater, at slow-moderate motion streams, on a sandy substrate; present in Carex spp. plants (Lundblad 1956). Limnesia (Limnesia) maculata (Müller, 1776) Synonym: Hydrachna maculata Müller, 1776 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Denmark or Norway (Müller 1776). Geographical distribution: Holarctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, at standing and temporary waters; present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956); Deutonymph larvae parasites of insect-chironomids Trichoptera and Cladocera larvae (Böttger 1969; Gerecke et al. 2016). Limnesia (Limnesia) manubriata Walter, 1928	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Syntypes deposited at NHMB two male catalogue numbers NMB-ACWA 2815 and 2816, two female 2813 and 2814 and a deuteronymph 2812. Type locality: Algeria, in Kabylie, Agoulmine Temjout; and Tunisia, in Ajim, Djerba Island (Walter 1928). Geographical distribution: Portugal and North of Africa (Lundblad 1956; Valdecasas 1988; Walter 1928).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace (Lundblad 1956). Ecological notes: Freshwater, at standing environments (Lundblad 1956). Limnesia (Limnesia) walteri Migot, 1926	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	materials_examined	Type material: Syntypes: Deposited in SMF, catalogue numbers 45725 - 3914 - VIE / 210, 45724 - 3905 - VIE / 210, and 45723 - 3901 - VIE / 210 (GBIF 2022). Type locality: France: Banyuls-sur-Mer, Barrage de la Baillaury (Migot 1926), Spain (GBIF 2022). Geographical Distribution: Russia, Southwestern Europe from Portugal to Greece, and North Africa, including the Sahara desert (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A58FFE2FF3081F58B46FEC4.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Santa Comba Dão, and Côa River, near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: In standing freshwaters, characteristic of remnant pools in streams (Gerecke et al. 2016), and can be found at an altitude of 50 m (Migot 1926); clear water, on a sandy bottom, present in plants, e. g. grasses, Carex spp., Juncus spp. and Myriophyllum spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE2FF3085138A5BFB82.taxon	materials_examined	Type material: Deposited in NHMB: Holotype one male, catalogue number 4458; Paratype: One male, catalogue number 4459 (GBIF 2022; Plazi. org 2022). Type locality: Switzerland: Ticino (GBIF 2022). Geographical distribution: Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE2FF3085138A5BFB82.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Côa River, near 10 km to Vilar Formoso (Lundblad 1956); South: Estremadura (Valdecasas 1988), in Colares, Sintra municipality (Lundblad 1956). Ecological notes: Occurs in eutrophic, oligotrophic and dystrophic lakes and slow-running streams, from coast to a depth of 50 m (Gerecke et al. 2016); clean waters, covered with Azolla spp.; and present in Juncus spp. and Myriophyllum spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE2FF3083998CA5F9B1.taxon	description	Synonyms: Nesaea pusilla Neumann, 1875 (original designation); Piona tuberifera Viets, 1910; Piona rotunda disjuncta Viets, 1930	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE2FF3083998CA5F9B1.taxon	materials_examined	Type material: Possible type material of an adult specimen deposited in NRM, catalogue number NHRSGULI 000079078 (GBIF 2022). Type locality: Sweden: Öland (GBIF 2022). Geographical distribution: Holarctic (Gerecke et al. 2016; Plazi. org 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE2FF3083998CA5F9B1.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Slightly brackish to freshwater, at standing / lentic and temporary waters (Gerecke et al. 2016); present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE3FF3081B08AC4FE70.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Northern Europe (Hermann 1804). Geographical distribution: Holarctic, remarkably Europe (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE3FF3081B08AC4FE70.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace (Lundblad 1956). Ecological notes: Freshwater (Lundblad 1956), at pools of streams and small habitats with a short temporary water cover; Larval insects hosts: Chironomidae dipterans (Gerecke et al. 2016). Tiphys torris (Müller, 1776) Synonym: Hydrachna torris Müller, 1776 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE3FF3081B08AC4FE70.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Denmark (probably surroundings of Copenhagen) or Norway (Müller 1776). Geographical distribution: Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A56FFE3FF3081B08AC4FE70.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace (Lundblad 1956). Ecological notes: In lentic freshwater ecosystems and small tributaries, including eutrophic, oligotrophic and dystrophic environments (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A57FFE3FF3084538D53F8C8.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Germany (Pešić et al. 2007; Piersig 1894). Geographical distribution: Palaearctic (Gerecke et al. 2016; Pešić et al. 2007).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A57FFE3FF3084538D53F8C8.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace; Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Lenitobiont freshwater, in standing and temporary waters, including pools, lakes, ponds and ditches (Pešić et al. 2007); present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956); Larval insect hosts: Chironomidae (Gerecke et al. 2016). Neumania (Neumania) uncinata Walter, 1927 Synonym: Neumania atlantida Lundblad, 1941	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A57FFE3FF3084538D53F8C8.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Southeastern France, in Provence (Walter & Motas 1927). Geographical distribution: Western Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A57FFE3FF3084538D53F8C8.taxon	distribution	Distribution in Portugal: South: Estremadura (Valdecasas 1988), in Sintra-Monserrate Park and Palace (Lundblad 1956); Madeira Island (Lundblad 1941 b). Ecological notes: Freshwater: springs, in flowing and standing water bodies, with preference for sandy or gravel substrate (Esen et al. 2011; Gerecke et al. 2016; Pešić et al. 2007). Neumania (Soarella) papillosa (Soar, 1902) Synonym: Ecpolus papillosus Soar, 1902 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A57FFE3FF3084538D53F8C8.taxon	materials_examined	Type material: Holotype (Soar 1902), probably not deposited in any scientific collection. Type locality: Britain (Soar 1902). Geographical distribution: Europe (Gerecke et al. 2016; Pešić et al. 2007).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A57FFE3FF3084538D53F8C8.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Santa Comba Dão, and Côa River, near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: Freshwater, common in eutrophic to dystrophic standing waters (Gerecke et al. 2016), in clear water, on a sandy substrate; present in plants, such as Carex spp., Juncus spp., Myriophyllum spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE1FF3081EF8CA0FF34.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Denmark (probably surroundings of Copenhagen) or Norway (Müller 1776). Geographical distribution: Western Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE1FF3081EF8CA0FF34.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Côa River, near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: In fresh-oligohaline and eutrophic slow-flowing waters, but with a preference for large water bodies; Larval insects hosts: Chaoboridae (Diptera), Chironomidae (Diptera) and Hydrophilus spp. larvae (Coleoptera) (Gerecke et al. 2016); present in Juncus spp., Myriophyllum spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE0FF3087938988FAFF.taxon	materials_examined	Type material: Unknown, possibly lost. Type locality: Denmark (probably surroundings of Copenhagen) or Norway (Müller 1776). Geographical distribution: Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE0FF3087938988FAFF.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão, and Côa River, near 10 km to Vilar Formoso; South: Estremadura, in Colares-Sintra municipality; Alentejo, in Ribeira de Odivelas; Guadiana River-between Serpa and Beja municipalities (Lundblad 1956; Valdecasas 1988). Ecological notes: In standing and slow-moving freshwater ecosystems, with ectoparasitic larvae of insectchironomids and on larval Trichoptera (Gerecke et al. 2016), in clear water, on a sandy substrate; present in plants, such as Azolla spp. Batrachium spp., Carex spp., Juncus spp., Myriophyllum spp., Nymphea spp., and Scirpus spp. (Lundblad 1956). Unionicola parvula Lundblad, 1954 Synonym: Unionicola (Pentatax) parvula Lundblad, 1954 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE0FF3087938988FAFF.taxon	materials_examined	Type material: Deposited in NRM: Holotype one female (Lundblad 1954), Lundblad No. Präp. 4120; Paratype: One male Lundblad No. Präp. 4381 (Lundblad 1956). Type locality: France: Béziers (Lundblad 1954, 1956). Geographical distribution: France and Iberian Peninsula (Gerecke et al. 2016; Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE0FF3087938988FAFF.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Santa Comba Dão (Lundblad 1956). Ecological notes: Freshwater, at slow-moderate flowing streams (Gerecke et al. 2016), on sandy substrate; present in Carex spp. plants (Lundblad 1956). Unionicola hankoi Szalay, 1927	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE0FF3087938988FAFF.taxon	materials_examined	Type material: Paratype: One male deposited in NRM, Lundblad No. Präp. 4501 (Lundblad 1956). Type locality: Hungary: Balatonsee (Lundblad 1956; Szalay 1927). Geographical distribution: Eastern and Southern Europe (Gerecke et al. 2016; Tuzovskij & Semenchenko 2015).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A54FFE0FF3087938988FAFF.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Côa River, near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: In lentic and lotic freshwater ecosystems (Erman et al. 2010); Found in the mantle cavity (females) or the branchiae (deutonymph forms) of mollusc-unionids (Gerecke et al. 2016); present in Myriophyllum spp. and Juncus spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE1FF3087938BD9FE70.taxon	distribution	Synonym: Arrenurus (Megaluracarus) autochthonus Lundblad, 1941 (original designation)	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE1FF3087938BD9FE70.taxon	materials_examined	Type material: Holotype one male NHRS-GULI 000089971, Allotype female NHRS-GULI 000089972, nymph type NHRS-GULI 000089973 deposited at NRM (Pers. comm., G. Lindberg). Type locality: Portugal: Madeira Island (Lundblad 1941 b). Geographical distribution: Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE1FF3087938BD9FE70.taxon	distribution	Distribution in Portugal: Madeira Island (Borges et al. 2008; Hughes et al. 1998). Ecological notes: Freshwater (Borges et al. 2008; Hughes et al. 1998).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE1FF30856F8931FC0C.taxon	distribution	Synonyms: Arrenurus hofsteni Walter, 1910; Arrenurus sinuator commutatus Viets, 1930	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE1FF30856F8931FC0C.taxon	materials_examined	Type material: Holotype (Barrois & Moniez 1887), possibly not deposited in any scientific collection. Type locality: Northern France, in Pont-à-Vendin (Barrois & Moniez 1887). Geographical distribution: Western Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE1FF30856F8931FC0C.taxon	distribution	Distribution in Portugal: Centre: Beira Alta (Valdecasas 1988), in Côa River, near 10 km to Vilar Formoso (Lundblad 1956). Ecological notes: In fresh to oligohaline waters, being a eutrophication-tolerant species. Larval insect hosts: Diptera Chaoboridae, Chironomidae and Culicidae (Gerecke et al. 2016); present in Myriophyllum spp. and Juncus spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE6FF3084F48A6CFE5D.taxon	materials_examined	Type material: One male paratype, catalogue number 42261 - 1418 - VIE / 035 (GBIF 2022). Type locality: United Kingdom (GBIF 2022). Geographical distribution: Western Palaearctic (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE6FF3084F48A6CFE5D.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, in slow-moving streams, ponds and lakes; Larval insects hosts: Zygoptera odonates (Gerecke et al. 2016); present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956). Arrenurus (Arrenurus) securifer Viets, 1930	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE6FF3084F48A6CFE5D.taxon	materials_examined	Type material: Paratypes: One male and one deposited in SMF, catalogue numbers 42372 - 3916 - VIE / 040 and one male and one female 42373 - 3917 - VIE / 040 (AQUILA 2022). Type locality: Spain: Madrid (Valdecasas 1988), in Manzanares River, Charca de Marmoto (Viets 1930). Geographical distribution: Southern France and Iberian Peninsula (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE6FF3084F48A6CFE5D.taxon	distribution	Distribution in Portugal: Centre: Beira Alta, in Côa River, near 10 km to Vilar Formoso; South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, in streams, ponds and lakes (Gerecke et al. 2016), as well as on sandy bottoms (Viets 1930); present in plants, such as Batrachium spp., Juncus spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956). Arrenurus (Arrenurus) szalayi Lundblad, 1954	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE6FF3084F48A6CFE5D.taxon	materials_examined	Type material: Deposited in NRM: Holotype one male Lundblad No. Präp. 4403 (Lundblad 1954); one female Lundblad No. Präp. 4405 (Lundblad 1956). Type locality: Portugal: Alentejo, in Ribeira de Odivelas (Lundblad 1954, 1956). Geographical distribution: Iberian Peninsula (Camacho & Valdecasas 1988; Lundblad 1956; Valdecasas 1988). Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1954, 1956). Ecological notes: Freshwater, in permanent ponds with grasses and algae (Camacho & Valdecasas 1988). Arrenurus (Megaluracarus) globator (Müller, 1776) Synonym: Hydrachna globator Müller, 1776 (original designation) Type material: Unknown, possibly lost. Type locality: Denmark (probably surroundings of Copenhagen) or Norway (Müller 1776). Geographical distribution: Palaearctic: Europe, Türkiye, Iran, Altai, Japan (Gerecke et al. 2016).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A55FFE6FF3084F48A6CFE5D.taxon	distribution	Distribution in Portugal: South: Alentejo, in Ribeira de Odivelas (Lundblad 1956). Ecological notes: Freshwater, with a preference for slow-moving streams, lakes and ponds; partially eutrophicationtolerant species; Larval insects hosts: Culicidae (Diptera), Chironomidae (Diptera), Dytiscidae (Coleoptera), Gyrinidae (Coleoptera) (Gerecke et al. 2016); present in plants, such as Batrachium spp., Myriophyllum spp., Nymphea spp. (Lundblad 1956).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A52FFE6FF30848B8BD9FACF.taxon	materials_examined	Type material: Holotype: One specimen deposited at Instituto Sperimentale per la Zoologia Agraria (Castilho et al. 2016; GBIF 2022). Type locality: Italy: Lido de Venezia (Canestrini & Canestrini 1881; Castilho et al. 2016; GBIF 2022). Geographical distribution: Western Europe (GBIF 2022).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A52FFE6FF30848B8BD9FACF.taxon	distribution	Distribution in Portugal: North: Leça da Palmeira, Cabo do Mundo, Vila Praia da Âncora, and Moledo beaches (Costa-Garcia 2018). Ecological notes: Marine, intertidal, barnacle epibiont (Costa-Garcia 2018; Halbert 1920).	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
03B187913A52FFE6FF30848B8BD9FACF.taxon	discussion	Remarks: This species is considered marine, however, its status as truly aquatic should be furtherly checked that it is an active species in all its life stages in water, as well as its breeding.	en	Cantallo, Harold, Gomes, Nuno, Antunes, Carlos, Martins, Duarte, Costa, Dimítri De Araújo (2022): Checklist of water mites from mainland Portugal and its archipelagos. Zootaxa 5213 (4): 336-370, DOI: https://doi.org/10.11646/zootaxa.5213.4.2
