taxonID	type	description	language	source
03B587806F47365CFF00CF4298142AE6.taxon	materials_examined	Type species: Scirus (Dactyloscirus) eupaloides Berlese, 1916, by original designation.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F47365CFF00CF4298142AE6.taxon	diagnosis	Diagnosis. Generally pale yellow to light brown or orange. Pedipalps 5 - segmented, with at least the terminal segment extending beyond subcapitulum; hyaline bulbous-tipped apophyses usually present on inner margin of telofemur and on joint of genu and tibiotarsus; the latter may be elongated, pointed or blunt apically, or less frequently inconspicuous or absent. Pedipalpal basifemora and telofemora with spine-like dorsal seta. Idiosoma covered dorsally with large propodosomal and variously developed hysterosomal shields in females, and more extensive dorsal shield covering almost entire idiosoma in males. Two pairs of long and plumose trichobothria (vi and sce) and 2 pairs of short and smooth tactile setae (ve and sci) on propodosomal shield, and 7 pairs on hysterosoma — c 1, d 1, e 1, f 1 and h 1 medially, and c 2 and h 2 laterally, setae h 2 often displaced ventrally on tapered posterior portion. Two pairs of cupules present: im dorsally near seta e 1, and ih ventrally near seta h 2. Ventrally on idiosoma, coxal plates I – II and III – IV contiguous, each usually bearing 3 tactile setae; genital plates large, each with 4 genital setae and 2 papillae; anal plates less distinct and striated like surrounding soft cuticle, with 1 pair of anal setae (ps 1) arising on the plate, and the 2 nd pair (ps 2) located outside it (thus the term para-anal also applied to it), together with the ventrally displaced seta h 2; additional 4 – 7 pairs of hysterogastral setae also present between posterior coxal and genital shields. Legs relatively short, usually not exceeding ¾ of the total body length; each leg ends in a pair of claws and 4 - rayed empodium; tarsi constricted apically forming conspicuous tarsal lobes that are completely separated from distal end of tarsi II – IV, and incompletely separated on tarsus I. Attenuate solenidia usually present on genua, tibiae and tarsi; famulus present on tarsus I, basally enlarged and usually ends in three tiny prongs; tibia IV always smooth trichobothrium. One of the dorsal tactile setae on basifemora III, IV and telofemora III, IV enlarged, long and spiculiform.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F473658FF00CA889B742953.taxon	description	(Figs. 1 – 3)	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F473658FF00CA889B742953.taxon	diagnosis	Diagnosis. In both sexes, pedipalps thin, curved and abruptly tapered from near base, 1 spls each on basi- and telofemur, 1 bulbous apophysis on telofemur, and 1 bulbous apophysis on joint of genu and tibiotarsus. Median shield absent in females, but a pair of lateral shields present on hysterosoma; male idiosoma strongly sclerotized, dorsal shield covering up to setae e 1. In both sexes, hysterosomal setae c 1, c 2, d 1 and e 1 minute (7 – 10), f 1 and h 1 2.25 – 2.5 times as long as the aforementioned setae. Genital setae 4 pairs: g 4 longest, 1.5 times longer than g 2, g 3, and 3 times than g 1 in females; all pairs subequally short in males. Chaetotaxy of basal segments of legs I – IV in both sexes: coxae 3 - 3 - 3 - 3, basifemora 5 - 5 - 3 - 2, and telofemora 5 - 5 - 4 - 4 sts.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F473658FF00CA889B742953.taxon	description	Description. FEMALE. Relatively large, with light brown idiosoma and slightly darker gnathosoma and legs; total body length 851 [851 – 1051 (945.2, n = 15)]; width 399 [339 – 532 (455.0, n = 14)]; Gnathosoma (Fig. 1 a): Relatively long and slender, 279 [266 – 306 (290.0, n = 15)] long, about ½ of total body length, 0.49 [0.36 – 0.55 (0.44, n = 15)], surface reticulate; hypostome gradually tapered as a cone; with 4 pairs of hypostomal setae and 2 pairs of adoral setae; hg 1 31, hg 2 31, hg 3 73, hg 4 24. Chelicerae 252 [252 – 265 (260.0, n = 15)] long, about ¼ of total body length, basal half inflated to about 4 times its distal width, surface reticulate, and with 1 subapical seta (Fig. 1 b). Pedipalps thin, curved and abruptly tapered from near base; 372 [372 – 412 (385.7, n = 14] long, about ½ of total body length, 0.44 [0.36 – 0.45 (0.41, n = 14)], extended beyond hypostome by half its length (Fig. 1 a); 5 - segmented and end in a small claw, surfaces strongly reticulate. Chaetotaxy of pedipalpal segments: trochanter 0; basifemur 1 spls, 27 long, arising dorsosubapically on outer margin; telofemur 1 spls, similar in length (27) to that located on basifemur, and 1 bulbous apophysis on outer margin, 17 long and with hyaline tip; genu 4 sts, 1 apophysis on joint of genu and tibiotarsus, 28 long, distally bulbous and hyaline; tibiotarsus 3 sts, 1 minute rodlike tubercle, and 1 tsl, basal sts on inner margin long (24) and fine, terminal solenidion tsl about as long, remaining setae minute. Idiosoma: Diamond-shaped, with prominent shoulders and sharply tapered posteriorly; 572 [559 – 771 (657.0, n = 15)] long. Dorsum (Fig. 2 a) with a transversely trapezoidal propodosomal shield and a pair of small lateral hysterosomal shields; median hysterosomal shield absent; surfaces of shields reticulate; lateral shields 54 × 7, more than 5 times as long as seta c 1 and about 2 / 3 of setal distance c 1 - c 2. Propodosomal shield bearing plumose trichobothria vi and sce, and 2 pairs of sts, ve and sci; vi 258 and sce 330 long and almost reaching anus; ve and sci minute (7 – 10), as long as the anteromedian hysterosomals; setal distances: vi – vi 30, vi – ve 78, vi – sce 97, ve – ve 136, ve – sce 23, sce – sce 140, sci – sci 46, sci – sce 51. Hysterosoma striate, striae densely laden with spine-like papillae, those between median setae coarser than lateral ones; complemented with 7 pairs of setae — c 1, c 2, d 1, e 1 and h 2 which minute (7 – 10), f 1 and h 1 which subequal (20 – 24) and 2.25 to 2.5 times longer than the former setae; setal distances: c 1 – c 1 58, c 1 – c 2 62, c 2 – c 2 168, d 1 – d 1 62, e 1 – e 1 58, f 1 – f 1 53, h 1 – h 1 41, h 2 – h 2 74. Cupule im rounded, mid-lateral to and approximately equidistant from setae e 1 and f 1. Venter (Fig. 2 b). Coxae I, II and III, IV contiguous, surfaces of coxal plates faintly reticulate; areas in intercoxal and opisthosomal regions that are not covered with ventral plates striate-papillate; with 4 pairs of hysterogastral setae arising between coxal plates III + IV, and 1 pair aggenital (or paragenital) setae lateral to genital shields. Genital plates faintly reticulate, each with 4 setae and 2 oval and unequal papillae; seta g 4 longest (51), 1.5 times longer than g 2 and g 3 (34), and 3 times than g 1 (17). Anal region with 2 pairs of anal setae: ps 1 on striated anal shield, and ps 2 on soft cuticle. Cupule ih conspicuous, slit-like, arising between ps 2 and h 2. Legs: Long and rather stout, lengths: I 466, II 385, III 492, IV 545; all leg segments strongly reticulate; tarsi ending in a pair of conspicuous lobes, and a pair of smooth claws and 4 - rayed empodium between the lobes; tarsal lobes II – IV separated from tarsus, those of leg I continuous with distal margin of tarsus. Chaetotaxy of leg segments (Fig. 3 a – d): coxae I – IV — 3 - 3 - 3 - 3 sts; trochantera I – IV — 1 - 1 - 2 - 1 sts; basifemora I – IV — 5 - 5 - 3 - 2 sts; telofemora I – IV — 5 - 5 - 4 - 4 sts; genu I — 4 sts, 4 asl, 1 mst in duplex with one of asl; genu II — 5 sts, 2 asl; genu III — 5 sts, 1 asl; genu IV — 5 sts, 2 asl; tibia I — 4 sts, 1 asl, 1 bsl, 1 mst in duplex with the asl; tbia II — 5 sts, 1 bsl; tibia III — 5 sts, 1 bsl; tibia IV — 4 sts, 1 T; tarsus I — 24 sts, 1 fam, 3 bsl, 1 mst in triplex with fam and one of the bsl, 1 dtsl; tarsus II — 22 sts, 1 bsl, 1 dtsl; tarsus III — 22 sts, 1 dtsl; tarsus IV — 20 sts, 1 dtsl. Famulus on tarsus I shorter (44) and about ¾ the length (58) of associated bsl; prongs at tip of famulus not observed. One of dorsal sts on basifemora III, IV and telofemora III, IV, noticeably thicker and longer than all others, spiculiform (macrosetae in: Shiba, 1986). MALE. Total body length 598 [545 – 598 (571.8, n = 2)], about half as long (0.58 – 0.6) as that of female; width 279 [239 – 279 (25.9, n = 2)]. Gnathosoma: Length 199 – 213 (206.2, n = 2), about 1 / 3 (0.36, n = 2) the total body length, hypostomal setae as in female. Pedipalps (Fig. 1 c) as in female, 213 [200 – 213 (206.2, n = 2)] long, about as long as gnathosoma and 1 / 3 of total body length; relatively more sclerotized and with more pronounced reticulations than in female. Chaetotaxy of pedipalpal segments as in female, including shapes, relative lengths and location of apophyses. Chelicera shaped as in female, 160 – 170 (165.5, n = 2) long. Idiosoma: Length 346 – 385 (365.0, n = 2) long. Dorsum (Fig. 1 d) almost entirely sclerotized, covered with a single dorsal shield bearing propodosomal and most of hysterosomal setae except f 1, h 1 and h 2 (Fig. 1 e); setae f 1 and h 1 about 10 long, only slightly longer than any of c 1 to e 1 and c 2 (7), h 2 minute, relative distances between dorsal setae as in female. Trichobothrium sce 228, slightly longer than vi (204), and not reaching anus. Venter as in female but more sclerotized, coxal plates I + II covering almost entire propodosomal venter and almost contiguous at midline, III + IV more narrowly separated on each side; opisthosoma much shorter, striate-papillate; coxal setae 3 - 3 - 3 - 3, and 3 pairs of hysterograstral setae arising on soft cuticle between coxal plates III + IV. Genital plates (Fig. 1 d) small, with 4 pairs of subequal short setae, and 2 pairs of unequal, oval papillae. Two pairs of setae in anal region: ps 1 on anal plate, and paranal ps 2 on soft, striated cuticle. Cupule ih conspicuous, slit-like, between ps 2 and h 2. Legs: Chaetotaxy of leg segments: coxae I – IV — 3 - 3 - 3 - 3 sts; trochantera I – IV — 1 - 1 - 2 - 1 sts; basifemora I – IV — 5 - 5 - 3 - 2 sts; telofemora I – IV — 5 - 5 - 4 - 4 sts; genu I — 4 sts, 4 asl, 1 mst in duplex with one of the asl; genu II — 5 sts, 1 asl; genu III — 5 sts, 1 asl; genu IV — 5 sts, 2 asl; tibia I — 4 sts, 1 asl, 1 bsl; tibia II — 5 sts, 1 bsl; tibia III — 5 sts, 1 bsl; tibia IV — 4 sts, 1 T; tarsus I — 20 sts, 1 fam, 3 bsl, 1 mst in triplex with fam and one of the bsl, 1 dtsl; tarsus II — 19 sts, 1 bsl, 1 dtsl, tarsus III — 18 sts, 1 dtsl; tarsus IV — 17 sts, 1 dtsl. Length of famulus (44) on tarsus I slightly over ¾ the length of associated asl (51); prongs at tip of famulus not observed. One dorsal sts on basifemora III, IV and telofemora III, IV long, thick and spiculiform as in female.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F473658FF00CA889B742953.taxon	materials_examined	Type material. Holotype female: unknown village in Langkat Regency (3.86540 o, 98.3088 oE), North Sumatra Province, INDONESIA, 10 Jan. 2014, collected by I. L. Lit, Jr., et al., caught by pitfall trap in corn field (sample LKT 2 - P 4 - R 1 - PF 2). Paratypes: 1 female, same data as for holotype but from a different sample; 11 females and 3 males, same data as for holotype but collected on 15 Dec. 2014 from 12 different samples; 1 male, same locality as for holotype, collected from soil in corn field, date unknown; 1 male, near Pioneer Learning Center, Tanjung Bintang District (5.3839 oN, 105.3834 oE), South Lampung Regency, Lampung Province, INDONESIA, collected from soil in corn field, date unknown. Additional material. From unknown village in Langkat Regency (3.86540 o, 98.3088 oE), North Sumatra Province, all collected on 15 Dec. 1963 by I. L. Lit, Jr. et al., by pitfall traps in corn field: 4 females and 1 nymph, from 5 different samples; 1 female, collected 10 Oct. 2014 from a different sample; 1 nymph, same data as preceding but date of collection unknown, from soil in corn field. From near Pioneer Learning Center, Tanjung Bintang District (5.3839 oN, 105.3834 oE), South Lampung Regency, Lampung Province, all collected by I. L. Lit, Jr.: 1 female and 1 male, date unknown, from 2 different soil samples; 1 female, date unknown, caught by pitfall trap; 1 female, date unknown, caught by pitfall trap outside experimental plots.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F473658FF00CA889B742953.taxon	etymology	Etymology. The specific epithet refers to the type locality, Sumatra Island.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F473658FF00CA889B742953.taxon	discussion	Remarks. The new species, Dactyloscirus sumatranus sp. nov. closely resembles D. dolichosetosus Den Heyer, 1979 from South Africa in sharing the following features in females: (1) the median shield is absent and a pair of lateral hysterosomal shields is present and longer than the adjacent setae c 1 and c 2; (2) bulbous apophyses are present on pedipalpal telofemur and on the joint of pedipalpal genu and tibiotarsus; (3) the posterior trichobothria sce on the propodosomal shield are very long and reaching the anus; and (4) the same number of femoral setae are present, namely, 5 - 5 - 3 - 2 on basifemora and 5 - 5 - 4 - 4 on telofemora. It differs from the latter species in the following characterstics: (1) the basal half of chelicera is strongly inflated, about four times wider than the distal part (vs. slender and gradually tapered towards the tip of claw in D. dolichosetosus); (2) the length of dorsolateral hysterosomal shields is about 2 / 3 the setal distance c 1 – c 2 (vs. equal); (3) hysterosomal setae f 1 and h 1 are at least twice as long as the anterior pairs c 1 – e 1 (vs. only slightly longer); and (4) the length of famulus on tarsus I is only ¾ the closely associated asl (vs. longer).	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F423655FF00CEF29A272873.taxon	description	(Figs. 4 – 7)	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F423655FF00CEF29A272873.taxon	diagnosis	Diagnosis. In both sexes, pedipalps short and thick, about ¼ of body length, 5 - segmented, with tibiotarsus almost straight and gradually tapered to the tip of claw; basifemur with 1 thick spls, telofemur with 1 thick spls and bulbous apophysis. Apophysis on joint of pedipalpal genu and tibiotarsus present, short and bulbous in female, not observed in male. Female idiosoma with a large propodosomal shield, without median but with a pair of small lateral shields on hysterosoma, and all dorsal setae arising from soft cuticle of hysterosoma; male idiosoma more extensively sclerotized, and bear setae c 1 – e 1 and c 2 on the dorsal shield. In both sexes, hysterosomal setae c 1, c 2, d 1, e 1 and h 2 minute (7 – 10), f 1 and h 1 2.25 times as long as the aforementioned pairs; g 4 longest of genital setae. Chaetotaxy of basal leg segments in both sexes: coxae 3 - 3 - 3 - 3, basifemora 5 - 5 - 3 - 2, telofemora 5 - 5 - 4 - 4.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F423655FF00CEF29A272873.taxon	description	Description. FEMALE. Moderately large and rather stout, with light brown idiosoma and slightly darker gnathosoma and legs; total body length 738 [738 – 745 (741.5, n = 2)]; width 332 [333 – 359 (345.5, n = 2)]. Gnathosoma (Fig. 4 a): Length 232 [226 – 232 (229, n = 2)], about 1 / 3 the total body length; hypostome stoutly conical, with 4 pairs of hypostomal setae of which hg 3 longest, and 2 pairs of adoral setae of which anterior pair slightly longer than posterior one. Chelicerae (Fig. 4 b) 190 [187 – 190 (188.5, n = 2)] long, about ¼ the total body length, basally inflated, basal width six times exceeding distal width, surface reticulate, and with 1 subapical seta. Pedipalps (Fig. 4 a) rather short, 226 [199 – 226 (212.5, n = 2)] long, about ¼ of total body length [0.27 – 0.31 (0.29)], and with tibiotarsus and distal part of genu extending beyond hypostome tip; 5 - segmented and ending in small claw, tibiotarsus thick, almost straight and gradually tapered distally; surfaces of all segments minutely reticulate. Chaetotaxy of pedipalpal segments: trochanter 0; basifemur 1 spls, 17 long, arising dorsosubapically on outer margin; telofemur 1 spls, similar in length and location as the former, and 1 short bulbous apophysis (17) with hyaline tip ventrally near inner margin; genu 4 sts and 1 small ovate apophysis (9) arising distally on inner surface; tibiotarsus 3 sts and 1 rod-like tubercle, basal sts on inner margin long and fine (17), terminal solenidion tsl about as long as wide, remaining setae minute. Idiosoma: Gradually tapered posteriorly from shoulders, 505 [505 – 519 (n = 2)] long. Dorsum (Fig. 5 a) with a transversely trapezoidal propodosomal shield and a pair of inconspicuous lateral hysterosomal shields; surfaces of shields reticulate; median hysterosomal shield absent; lateral shields 37 × 3, five times as long as seta c 1, and about ½ setal distance c 1 – c 2. Propodosomal shield bearing trichobothria vi and sce and 2 pairs of sts (ve and sci); vi 178, sce 221 long; ve and sci minute like anteromedian hysterosomals, 7 – 10; distances between setae: vi – vi 16, vi – ve 56, vi – sce 72, ve – ve 108, ve – sce 14, sce – sce 114, sci – sci 50, sci – sce 34. Soft cuticle of hysterosoma striate-papillate, striae between median setae coarser than lateral ones which fade out to thickened lateral areas of idiosoma; complemented with 7 pairs of setae — c 1, c 2, d 1, e 1 and h 2 which are minute (7 – 10), and f 1 and h 1 which are about twice longer (20) than the former setae; distances between setae: c 1 – c 1 50, c 1 – c 2 46, c 2 – c 2 140, d 1 – d 1 48, e 1 – e 1 44, f 1 – f 1 40, h 1 – h 1 36, and h 2 – h 2 72. Cupule im oval, mid-lateral to and slightly closer to e 1 than f 1. Venter (Fig. 5 b). Coxae I, II and III, IV contiguous, surfaces of coxal plates coarsely papillate to minutely reticulate; intercoxal and opisthosomal cuticle striate-papillate; with 4 pairs hysterogastral and 1 pair of aggenital (or paragenital) setae arising on hysterosomal area between coxal plates III + IV and genital plates. Genital plates minutely reticulate, each plate with 4 setae and two oval, subequal papillae; seta g 4 longest (34), g 2 24, g 3 17 and g 1 shortest (14). Two pairs of setae in anal region, ps 1 on anal plate, and paranal ps 2 on soft cuticle surrounding the plate, the latter 3 times longer than former. Cupule ih conspicuous, rounded, arising close to seta ps 2. Legs: Rather long and stout, lengths: I 372, II 293, III 359, IV 412, the last about half the length of body; all leg segments strongly reticulate; tarsi ending in a pair of conspicuous lobes, a pair of smooth claws and 4 - rayed empodium between lobes; tarsal lobes II – IV separated from tarsus, those of I continue with distal end of tarsus. Chaetotaxy of leg segments (Fig. 6 a – d): coxae I – IV — 3 - 3 - 3 - 3 sts; trochantera I – IV — 1 - 2 - 1 sts; basifemora I – IV — 5 - 5 - 3 - 2 sts; telofemora I – IV — 5 - 5 - 4 - 4 sts; genu I — 4 sts, 4 asl, and 1 mst in duplex with one of the asl; genu II — 5 sts, 1 asl; genu III — 5 sts, 1 asl; genu IV — 5 sts, 2 asl; tibia I — 5 sts, 1 asl, 1 mst in duplex with the asl; tibia II — 5 sts, 1 bsl; tibia III — 5 sts, 1 bsl; tibia IV — 4 sts, 1 T; tarsus I — 23 sts, 1 fam, 2 bsl, 1 asl, 1 mst in triplex with fam and the distal bsl, 1 dtsl; tarsus II — 22 sts, 1 bsl, 1 dtsl; tarsus III — 21 sts, 1 dtsl; tarsus IV — 21 sts, 1 dtsl. Famulus on tarsus I 24 long, almost ¾ the length of associated asl (34), prongs on its tip not observed. One of dorsal sts on basifemora III, IV and telofemora III, IV spiculiform, noticeably thicker and longer than all others. MALE. Smaller than female, total body length 558 – 567 (562.5, n = 2), width 226 – 253 (239.5, n = 2). Gnathosoma: Hypostome as in female, with 4 pairs of hypostomal and 2 pairs of adoral setae, hg 3 longest (Fig. 4 c). Chelicerae thin, 150 (n = 1), about as long as pedipalp, gradually tapered from distal third, basally reticulate, cheliceral seta present (Fig. 4 d). Pedipalps short and stout like that of female, 146 (n = 1), about ¼ of total body length; 5 - segmented, end in a small claw (Fig. 4 c). Chaetotaxy of palp segments: trochanter 0; basifemur 1 spls, 20; telofemur 1 spls, 20, and 1 short bulbous apophysis, length × width, 7 × 7, arising ventrally on inner margin; genu 4 sts, without apophysis; tibiotarsus 3 short sts, 1 minute tubercle and long tsl. Idiosoma: Length 359 – 394 (376.5, n = 2). Dorsal shield large, covering dorsum up to setae e 1, surface minutely reticulate (Fig. 6 a). Trichobothria vi and sce 204 and 221 long, respectively, plumose, and not reaching anus; setae ve, sci, c 1, c 2, d 1 and e 1 arising on dorsal shield, all short, 7 – 8 long; f 1 and h 1 arising on soft cuticle of opisthosoma, only slightly longer (10) than anterior pairs of hysteronotal setae situated on shield; relative distances between dorsal setae as in female. Venter (Fig. 6 b) similar to that of female but opisthosoma much shorter and tapered, genito-anal area more sclerotized, and anal setae ps 1 and ps 2 arising dorsally, along with genital setae g 3 and g 4. Three pairs of hysterogastral setae present on soft intercoxal area between coxal plates III + IV. Genital plates strongly sclerotized and bearing genital setae g 1 and g 2 ventrally, 14 and 17 long, respectively; setae g 3 and g 4 arising dorsally, both 10 long. Legs (Fig. 7 a – d): relatively short and stout, lengths: I 282, II 238, III 255, IV 299. Chaetotaxy of I – IV leg segments: basal segments from coxa to telofemur as in female; genu I — 4 sts, 3 asl; genu II — 6 sts, 1 asl; genu III — 5 sts, 1 asl; genu IV — 5 sts, 1 asl; tibia I — 4 sts, 1 asl and 1 mst in duplex, 1 bsl; tibia II — 5 sts, 1 bsl; tibia III — 5 sts, 1 bsl; tibia IV — 4 sts, 1 T; tarsus I — 20 sts, 1 fam, 2 bsl, 1 mst in triplex with fam and the distal bsl, 1 dtsl; tarsus II — 20 sts, 1 bsl, 1 dtsl; tarsus III — 19 sts, 1 dtsl; tarsus IV — 17 sts, 1 dtsl. Relative lengths of famulus and associated bsl as in female, prongs at its tip not observed. Long spiculiform setae present on basifemora and telofemora III and IV as in female.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F423655FF00CEF29A272873.taxon	materials_examined	Material. Holotype female: unknown village in Langkat Regency (3.86540 o, 98.3088 oE), North Sumatra Province, INDONESIA, 15 Dec. 2013, collected by I. L. Lit, Jr., et al., caught by pitfall trap in cornfield (sample LKT 1 - P 5 - R 1 - PF- 1). Paratypes: 1 female, same data as for holotype but from sample taken on 10 Jan. 2014; 1 male, same data as for holotype but collected from a different sample; and 1 male, same locality data as for holotype, date unknown.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F423655FF00CEF29A272873.taxon	etymology	Etymology. The specific epithet refers to the type habitat, in corn (Zea mays) fields, and is a combination of the Indonesian (Bahasa) words “ ladang ” (field) and “ jagung ” (corn); it is treated as a Latinized noun in apposition.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F423655FF00CEF29A272873.taxon	discussion	Remarks. The female of Dactyloscirus ladangjagung sp. nov. resembles that of D. hoffmannae Swift, 1996 from Hawaii in having the following characterstics: (1) the pedipalpal tibiotarsus is thick and gradually tapered distally; (2) the apophysis on pedipalpal genu is small and narrowly bulbous; (3) the median dorsal hysterosomal shield is absent; and (4) a pair of lateral hysterosomal shields is present. The females of the two species are separated by the number of setae (sts) on some leg segments, viz., 5 - 5 - 3 - 2 on basifemora in D. ladangjagung (vs. 5 - 5 - 3 - 1 in D. hoffmannae), and on tibiae 5 - 5 - 5 - 4 (vs. 4 - 5 - 4 - 2). In addition, seta g 4 is the longest of all genital setae in the female of D. ladangjagung while g 3 is longest in D. hoffmannae.	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
03B587806F4E3653FF00C83E9CA92A22.taxon	description	(Females)	en	Corpuz-Raros, Leonila A., Naredo, Jeremy C. B., Lit, Ireneo L., Caasi-Lit, Merdelyn T. (2022): Two new species of the genus Dactyloscirus (Acari: Prostigmata: Bdelloidea Cunaxidae) from Sumatra, Indonesia, with a key to known species. Zootaxa 5214 (1): 89-103, DOI: https://doi.org/10.11646/zootaxa.5214.1.3
