taxonID	type	description	language	source
A5415E5FF410FFDA9849FE09FE74FC9D.taxon	description	Figs 4, 6 A – B	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9849FE09FE74FC9D.taxon	distribution	Distribution Dominican Republic.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9817FCB5FC5CFBD3.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9817FCB5FC5CFBD3.taxon	distribution	Distribution Bolivia; Costa Rica; Ecuador; Nicaragua; Panama; Suriname.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9855FB72FEC0FA28.taxon	description	Figs 4, 47 – 55, Supp. file 2 B, Supp. file 3 (Figs 1 A – 4 H)	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9855FB72FEC0FA28.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9872FA38FEC0F961.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDA9872FA38FEC0F961.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDB9BC0F8E4FECFFEE3.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF410FFDB9BC0F8E4FECFFEE3.taxon	distribution	Distribution Cuba.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB982BFE82FECFFDD5.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB982BFE82FECFFDD5.taxon	distribution	Distribution Cuba.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB9826FD74FECFFCCF.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB9826FD74FECFFCCF.taxon	distribution	Distribution Cuba.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB9BCFFC5EFECFFB21.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB9BCFFC5EFECFFB21.taxon	distribution	Distribution Cuba.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB985AFB40FEC0FA1B.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB985AFB40FEC0FA1B.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB9842FA2BFEC0F90D.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB9842FA2BFEC0F90D.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB986CF91DFED7F867.taxon	description	Figs 4, 6 C – D	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF411FFDB986CF91DFED7F867.taxon	distribution	Distribution Bolivia.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC9872FEF2FF37FE4A.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC9872FEF2FF37FE4A.taxon	distribution	Distribution Peru.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC986AFDE4FF37FCBC.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC986AFDE4FF37FCBC.taxon	distribution	Distribution Peru.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC9847FCCEFEEFFB91.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC9847FCCEFEEFFB91.taxon	distribution	Distribution Jamaica.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC9800FBB0FED2FAEE.taxon	description	Fig. 4	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC9800FBB0FED2FAEE.taxon	distribution	Distribution Mexico.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC986CFA7EFF37F9C1.taxon	description	Figs 4, 6 E – F	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC986CFA7EFF37F9C1.taxon	distribution	Distribution Peru.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC987FF960FEC0F83A.taxon	description	Fig. 5, 56 – 64	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF416FFDC987FF960FEC0F83A.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9863FEF3FED2FDBE.taxon	description	Fig. 5	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9863FEF3FED2FDBE.taxon	distribution	Distribution Mexico.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9875FDD1FF37FCBE.taxon	description	Fig. 5	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9875FDD1FF37FCBE.taxon	distribution	Distribution Peru.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9870FCD1FEC0FBB8.taxon	description	Figs 4, 6 G – H	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9870FCD1FEC0FBB8.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9874FBC8FEC0FAAB.taxon	description	Figs 4, 6 I – J	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9874FBC8FEC0FAAB.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD986BFAC3FEC0F9AA.taxon	description	Figs 4, 6 K – L	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD986BFAC3FEC0F9AA.taxon	distribution	Distribution Brazil.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9872F9BAFEF5F89C.taxon	description	Figs 5, 6 M – N	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDD9872F9BAFEF5F89C.taxon	distribution	Distribution Colombia.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDE9870F8ACFED2FEE3.taxon	description	Fig. 5	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF417FFDE9870F8ACFED2FEE3.taxon	distribution	Distribution Mexico.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE981EFE82FDD5FD39.taxon	description	Fig. 5	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE981EFE82FDD5FD39.taxon	distribution	Distribution Guatemala; Mexico; Panama.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE9870FD48FB6AFB51.taxon	description	Figs 5, 6 O – P	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE9870FD48FB6AFB51.taxon	distribution	Distribution Bolivia.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE9870FD48FB6AFB51.taxon	discussion	Remarks We received photos of the dorsal view and labels of the type (MNHN), but the specimen shown in the photos does not match the original description. The pronotum of this specimen is entirely reddish brown and 3 mm long, while in the description Pic (1931) wrote “ capite antice et thorace lateraliter rufis (…) Long. 4 mill. (…) Voisin de anale Reitt. ”. The specimen on the photo better fits the description of C. collare, by its coloration (pronotum reddish brown) and length of 3 mm. We could not locate the syntypes of C. collare. The illustration of C. testaceicorne (Fig. 5) provided here was based on the original description and not on the specimen shown in the available photos (Fig. 6 O).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE9816FAF1FDEAF9D3.taxon	description	Fig. 5	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFDE9816FAF1FDEAF9D3.taxon	distribution	Distribution Mexico. Descriptions of new species	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	description	urn: lsid: zoobank. org: act: ABFE 4 E 1 F-DB 31 - 46 C 1 - BDC 7 - BDD 1 C 4 FF 81 E 3 Figs 4, 7 – 13, 46; Supp. file 1 A	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	diagnosis	Diagnosis TL: 3.00 – 3.28 mm in males and 3.32 – 3.40 mm in females. Entirely black, with lateral areas of some ventral sclerites lighter (Fig. 7 A – C). Eyes long (Fig. 7 D). Hypomeron and metaventrite smooth. Metanepisternum and metepimeron with imbricate microsculpture. Tergite VIII of males coarsely punctate (Fig. 11 D). Aedeagus strongly sclerotized (Fig. 12 A – C); apex and parameres short (Fig. 12 A). Distal gonocoxites of females straight and thick (Fig. 13 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	etymology	Etymology In homage to the entomologist Julien Achard (1881 – 1925) for his significant contribution on the Neotropical Scaphidiinae.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	materials_examined	Material examined Holotype BRAZIL • ♂; Minas Gerais, Viçosa, EPTEA Mata do Paraíso; 19 Nov. 2019; LabCol leg.; “ Fungo 27 \\ Em Leucoagaricus sp. \\ Cyparium achardi von Groll & Lopes-Andrade HOLOTYPUS ” [red paper]; CELC. (Supp. file 1 A). Paratypes BRAZIL • 3 ♂♂, 1 ♀ (1 ♂ entirely dissected, preserved in glycerin); same collection data as for holotype; “ T. do Pesquisador ”; 8 Nov. 2016; C. Lopes-Andrade et al. leg.; “ \\ Em Marasmiellus volvatus ”; CELC • 2 ♀♀ (1 ♀, abdomen dissected, preserved in glycerin); same collection data as for holotype; 14 Nov. 2019; “ Fungo 22 \\ Em Marasmiellus sp. ”; CELC • 1 ♀; same collection data as for holotype; “ Fungo 26 \\ Em Hygrocybe sp. ”; CELC • 1 ♂; same collection data as for holotype; “ Fungo 28 \\ Em Leucocoprinus ianthinus ”; CELC • 1 ♂; same collection data as for holotype; “ Fungo 12 \\ Em Marasmius haematocephalus ”; CELC • 1 ♀; same collection data as for holotype; 26 Nov. 2019; LabCol leg.; “ Fungo 53 \\ Em Leucocoprinus sp. ”; CELC • 1 spec.; Viçosa, Recanto das Cigarras, Mata da Biol.; 20 Nov. 2019; LabCol leg; “ Fungo 19 \\ Em Leucocoprinus cepistipes ”; CELC. All paratypes additionally labelled “ Cyparium achardi von Groll & Lopes-Andrade PARATYPUS [yellow paper] ”.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	description	Description MEASUREMENTS (holotype, in mm). TL 3.12, PL 1.31, PA 0.80, PB 1.90, EW 1.20, EL 2.17, IS 0.18, HW 0.80. COLORATION. Black; iridescent (Fig. 7 A – C). Clypeus yellowish-brown; mouth parts yellowish (Fig. 7 D); antennomeres I – VI and apical half of XI yellow; VII – X dark yellow (Fig. 7 D – F). Hypomeron, epipleuron, and metepimeron brown to black, lighter laterally (Fig. 7 B). Thorax and abdomen brown to black, lighter laterally (Fig. 7 B). Coxae and femora light to chestnut brown, lighter posteriorly in some specimens; tibiae light to dark brown; tarsi yellow (Fig. 7 B – C). HEAD. Punctation dense, fine (Fig. 7 D). Eyes tapered apically (Fig. 7 D). Labrum transverse; lateral areas rounded, smoothly delimited apically; central margin slightly curved; sclerotized portion rounded with a row of short setae; lateral setae slightly exceeding margins of labrum; slightly porose centrally (Fig. 7 G). Mandibles curved, subapical serrations on left mandible conspicuous (Fig. 7 H – I). Maxillae with palpomere II thickened; galea and lacinia densely pubescent, narrow (Fig. 7 J). Mentum with sides distinctly angulated, curved apically (Fig. 8 A). Setae of labial palpomere II exceeding palpomere III; palpomere III thick, with long apical setae (Fig. 8 A). Hypopharynx with sclerotized plate cup-shaped (Fig. 8 A – B). Postgena with strigulate microsculpture; gular pores sparse; gula small and rounded at anterior region (Fig. 7 C). Antennae variable in size between males and females (Fig. 7 E – F). PROTHORAX. Pronotum smooth, shining; punctation somewhat dense, fine; pubescence short, fine (Fig. 8 D – E); somewhat squared, lateral areas curved, forming an obtuse angle at lateral areas of posterior margin (Fig. 8 E). Hypomeron smooth (Fig. 8 F). Notosternal suture straight, inward-directed (Fig. 8 F). Profurca elongated, exceeding half of foramen (Fig. 8 G). Prosternal process acute apically (Fig. 9 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl, 2005) almost straight (Fig. 9 B). Scutellum rounded posteriorly (Fig. 9 B). Anterior phragma wide, slightly curved (Fig. 9 C). Mesanepisternum smooth (Fig. 9 D). Procoxal rest rounded, curved posteriorly (Fig. 9 D). Mesoventral line rounded; median line moderately curved; area between median line and mesocoxal line longer laterally (Fig. 9 D). Mesoventral process large and straight at base in lateral view (Fig. 9 E). METATHORAX. Metanotum with wide alacrista, rounded and somewhat turned to the posterior end; scutoscutellar suture slightly straight; median membranous area wide and very short (Fig. 9 F). Metaventrite smooth; punctation sparse, fine; some specimens with faint lateral imbricate microsculpture (Fig. 9 D). Mesocoxal line forming an open angle next to each coxal cavity and finely punctate under coxal cavity (Fig. 9 D). Metanepisternum and metepimeron (Figs 7 B – C, 9 D) with imbricate microsculpture, but almost inconspicuous in some specimens. Intercoxal plates smooth (Fig. 7 B). Metendosternite with straight furcal arms; ‘ stalk ridge’ not exceeding half of stalk length (Fig. 10 A); ventral longitudinal flange flat in lateral view (Fig. 10 B). WINGS. Elytra slightly longer than wide, partially covering tergite VI (Fig. 7 A); basal and lateral lines punctate (Fig. 8 D); sutural line dashed; adsutural area with a row of short setae (Fig. 7 A); six rows of coarse punctures (not including sutural line), but row 5 and 6 somewhat intermixed (Figs 7 A, 10 C); apical coarse punctation moderately sparse; apical serrations moderately large, well visible (Fig. 10 D); pubescence short, fine, denser on disc. Epipleuron with a row of punctures (poorly marked in some specimens). Hind wings fully developed (Fig. 10 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Pro- and mesofemora somewhat fusiform (Fig. 10 F – G); punctation moderately sparse and coarse. Metafemora straight, finely punctate (Fig. 10 H). Mesotibiae densely spinose, spines fine (Fig. 10 G). Metatibiae sparsely spinose, spines fine (Fig. 10 H). ABDOMEN. Tergites VI – VIII with imbricate microsculpture (Fig. 11 A). Tergite VII trapezoidal; punctation dense, coarse; pubescence sparse, fine (Fig. 11 A). Ventrite 1 sparsely and coarsely punctate; pubescence sparse, fine; strigulate microsculpture (Fig. 11 B – C). Metacoxal lines finely punctate. Ventrites 2 – 5 sparsely and finely punctate; pubescence sparse, fine; imbricate microsculpture anteriorly and with strigulate microsculpture middle-posteriorly. Males MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.18 (0.06), 0.11 (0.05), 0.10 (0.04), 0.09 (0.04), 0.10 (0.05), 0.07 (0.06), 0.09 (0.10), 0.09 (0.11), 0.10 (0.12), 0.10 (0.13), 0.17 (0.14); (n = 5, including the holotype; in mm): TL 3.00 – 3.28 (mean = 3.14, standard deviation ± 0.10), PL 1.20 – 1.31 (1.25 ± 0.04), PA 0.80 – 0.97 (0.89 ± 0.06), PB 1.72 – 2.07 (1.93 ± 0.13), SL 0.13 – 0.17 (0.14 ± 0.01), SW 0.15 – 0.20 (0.18 ± 0.02), EI 1.72 – 1.92 (1.81 ± 0.07), EL 2.07 – 2.32 (2.18 ± 0.09), EW 1.07 – 1.25 (1.19 ± 0.07), EH 0.70 – 0.80 (0.75 ± 0.05), HW 0.74 – 0.83 (0.78 ± 0.03), IS 0.15 – 0.24 (0.17 ± 0.03), WA 0.14 – 0.17 (0.16 ± 0.01), MC 0.72 – 0.92 (0.81 ± 0.08), MB 0.42 – 0.50 (0.46 ± 0.04), VL 0.62 – 0.70 (0.64 ± 0.03). Antennae almost reaching level of mesanepisternum, smaller than in females (Fig. 7 E). Pro- and mesotarsomeres I – III enlarged, with tenent setae (Fig. 10 F). Tergite VIII triangular; punctation dense, coarse, subglabrous (Fig. 11 D). Tergite IX with rectangular ventral struts (Fig. 11 E, F). Sternite VIII rectangular, longish; finely punctate and with strigulate microsculpture (Fig. 11 G). Sternite IX thicker posteriorly and triangular anteriorly (Fig. 11 F). Aedeagus strongly sclerotized, enlarged and darker basely and narrowed apically, apex of median lobe short (Fig. 12 A, B); openings in dorsal view somewhat enlarged, forming an acute angle (Fig. 12 C); internal sac with irregular sclerites, with two hooks (Fig. 12 D, E), parameres short, somewhat enlarged apically (Fig. 12 A). Females MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.20 (0.06), 0.13 (0.05), 0.11 (0.04), 0.12 (0.05), 0.11 (0.06), 0.08 (0.07), 0.11 (0.11), 0.10 (0.13), 0.11 (0.14), 0.10 (0.16), 0.16 (0.15); (n = 4, paratype; in mm): TL 3.32 – 3.40 (mean = 3.37, standard deviation ± 0.04), PL 1.23 – 1.39 (1.33 ± 0.07), PA 0.97 – 1.05 (1.01 ± 0.04), PB 2.05 – 2.17 (2.10 ± 0.05), SL 0.18 – 0.21 (0.19 ± 0.01), SW 0.17 – 0.20 (0.19 ± 0.01), EI 1.90 – 2.00 (1.94 ± 0.05), EL 2.27 – 2.40 (2.35 ± 0.05), EW 1.25 – 1.27 (1.25 ± 0.01), EH 0.80 – 1.00 (0.88 ± 0.09), HW 0.81 – 0.84 (0.84 ± 0.02), IS 0.16 – 0.25 (0.20 ± 0.04), WA 0.15 – 0.21 (0.18 ± 0.02), MC 0.87 – 0.95 (0.90 ± 0.03), MB 0.42 – 0.52 (0.46 ± 0.04), VL 0.67 – 0.75 (0.71 ± 0.03). Antennae longer than in males, surpassing mesanepisternum (Fig. 7 F). Tergite VIII triangular, rounded posteriorly; punctation dense, coarse (inconspicuous in some specimens); subglabrous (Fig. 13 A). Sternite VIII rectangular with strigulate microsculpture (Fig. 13 B). Vagina and bursa copulatrix membranous; vaginal plate with an apical sclerite and a pair of spermatheca at base of oviduct (Fig. 13 C – D). Distal gonocoxites parallel, straight and thick; gonostyli larger at apex (Figs 13 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	biology_ecology	Host fungi Adults were collected from Marasmiellus volvatus Singer (Agaricales, Omphalotaceae) (1 record, 4 individuals), Marasmiellus sp. Murrill (1, 2), Marasmius haematocephalus (Mont.) Fr. (Agaricales, Marasmiaceae) (1, 1), Leucocoprinus ianthinus (Sacc.) P. Mohr (Agaricales, Agaricaceae) (1, 1), L. cepistipes (Sowerby) Pat. (1, 1), Leucoprinus sp. (1, 1), Hygrocybe sp. (Agaricales, Hygrophoraceae) (1, 1), Leucoagaricus sp. (Agaricales, Agaricaceae) (1, 1).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	discussion	Remarks Similar to C. pygidiale, which occurs in Jataí, Goiás, Brazil (1174 km from the collection site of C. achardi sp. nov.), but they differ in the humeral and apical region of the elytra, and in the scutellar shield (blackish in C. achardi and reddish in C. pygidiale); elytral punctation is also different: rows 1 – 3 are coarser in C. pygidiale and the remainder are coarser in C. achardi; furthermore, rows 1 – 3 are somewhat outward directed, while they are inward directed in C. pygidiale.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF414FFC8985BF980FD6EFA25.taxon	distribution	Distribution Known only from Mata da Biologia, campus of the Universidade Federal de Viçosa, Viçosa, state of Minas Gerais, Southeast Brazil (Fig. 46).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	description	urn: lsid: zoobank. org: act: 8 A 653 AB 2 - 6 DEF- 4 FA 6 - A 885 - 438 BDAF 11 D 5 F Figs 4, 14 – 21, 46; Supp. file 1 B, Supp. file 2 A	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	diagnosis	Diagnosis TL: 2.07 – 2.25 mm in males and 2.10 – 2.35 mm in females. Brown (Fig. 14 A). Hypomeron, metaventrite and intercoxal plates with strigulate microsculpture. Metaventrite coarsely punctate above intercoxal plates (Figs 14 B, 17 D). Tergite VIII in males triangular (Fig. 19 D). Openings of aedeagus in dorsal view forming an acute angle (Fig. 20 C); internal sac with drop-like sclerites (Fig. 20 D – E). Tergite VIII of females with a posterior invagination (Fig. 21 A).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	etymology	Etymology In homage to Dr Richard A. B. Leschen (New Zealand Arthropod Collection) for his great contributions to the systematics of Coleoptera, especially regarding Scaphidiinae.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	materials_examined	Material examined Holotype BRAZIL • ♂; Minas Gerais, Viçosa, EPTEA Mata do Paraíso; 19 Nov. 2019; LabCol leg.; “ Fungo 30 \\ em Agaricus sylvaticus \\ Cyparium lescheni von Groll & Lopes-Andrade HOLOTYPUS ” [red paper]; CELC. (Supp. file 1 B) Paratypes BRAZIL • 1 ♂, 4 ♀♀; same collection data as for holotype, “ T. dos Gigantes ”; 15 Feb. 2015; S. Aloquio, A. Orsetti and M. Bento leg.; CELC • 1 ♀; same collection data as for holotype, “ T. da Madeira ”; 27 Feb. 2015; I. S. C. Pecci-Maddalena et al. leg.; CELC • 1 ♂; same collection data as for holotype; 13 Mar. 2015; S. Aloquio, A. Orsetti, C. Lopes-Andrade and M. Bento leg.; CELC • 1 ♂; same collection data as for holotype; 9 Nov. 2016; I. Pecci-Maddalena and C. Lopes-Andrade leg.; “ \\ ex Psathyrella candolleana ”; CELC • 5 ♂♂, 7 ♀♀ (2 ♂♂, 1 ♀, entirely dissected, preserved in glycerin; 1 ♂, 2 ♀♀, abdomen dissected, preserved in glycerin); same collection data as for holotype; CELC • 1 ♂; 1 ♀; same collection data as for holotype; CAMB • 1 ♂; 1 ♀; same collection data as for holotype; CERPE • 2 ♂♂; 1 ♀, 1 spec.; same collection data as for holotype; 21 Nov. 2019; LabCol leg.; “ Fungo 06 \\ Em Agaricus dulcidulus e Leucocoprinus brebissoni ”; CELC • 2 ♀♀; Viçosa, Recanto das Cigarras, Mata da Biol.; 20 Nov. 2019; LabCol leg.; “ Fungo 29 \\ Em Entoloma (Inocephalus) sp. ”; CELC • 8 ♂♂, 3 ♀♀ (1 ♂, entirely dissected, preserved in glycerin; 2 ♂♂, 1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; “ Fungo 08; CELC \\ Em Psathyrella sp. ”; CELC • 6 ♂♂, 11 ♀♀, 1 spec.; Viçosa, Mata da Biologia; 15 Oct. 2021; E. von Groll and A. Orsetti leg.; “ Fungo 20 \\ Em Agaricus sp. ”; CELC. All paratypes additionally labelled “ Cyparium lescheni von Groll & Lopes-Andrade PARATYPUS [yellow paper] ”.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	description	Description MEASUREMENTS (holotype, in mm). TL 2.32, PL 0.76, PA 0.70, PB 1.40, EW 0.86, EL 1.62, IS 0.20, HW 0.62. COLORATION. Iridescent. Brown; antennomeres I – VI, clypeus, mouthparts, tarsi yellow (Figs 14 A, 15 A). Variation: few paratypes with pronotum and hypomeron reddish brown (Fig. 14 D – F); scutellum reddish brown or black (Fig. 14 D); meso- and metathorax reddish brown, each sclerite lighter laterally (Fig. 14 E); elytra blackish (Fig. 14 D); epipleuron dark ochre; coxae and trochanters dark ochre; femora dark ochre, apex lighter; tibiae dark ochre, lighter anteriorly and posteriorly; tarsi yellow (Fig. 14 E); tergite VI and VII blackish; tergite VIII yellow; ventrite 1 dark ochre, 2 – 4 dark yellow, 5 and 6 yellow (Fig. 14 E). HEAD. Punctation dense, fine (Fig. 15 A). Eyes long (Fig. 15 A). Labrum sub-rectangular, lateral margins smoothly rounded, not well delimited apically; central margin sub-straight; sclerotized portion inwardly curved; lateral setae considerably extending from the margins of the labrum; densely porose centrally (Fig. 15 D). Mandibles strongly curved; subapical serrations on left mandible conspicuous (Fig. 15 E – F). Maxillary palps, galea and lacinia moderate pubescent, elongated (Fig. 15 G). Mentum straight apically and rounded laterally; sides well delimited (Fig. 16 A). Setae of labial palpomere II extending palpomere III; palpomere III longish with long apical setae (Fig. 16 A). Hypopharynx with narrow and rounded sclerotized-plate (Fig. 16 A – B). Post gena microsculptured with transversal lines; gular pores sparse; gula long and sharp (Fig. 16 C). Antennal club very distinct; antennomere XI longish, hexagonal, more or less rounded apically; remarkably different between sexes (Fig. 15 B – C). PROTHORAX. Pronotum smooth, shining; punctuation dense, fine; pubescence short, fine (Fig. 16 D – E); transverse, sub-straight laterally, forming an obtuse angle at lateral areas of posterior margin (Fig. 16 E). Hypomeron with strigulate microsculpture. Notosternal suture straight, inward directed (Fig. 16 F). Profurca somewhat thick and elongated, slightly extending half length of foramen (Fig. 16 G). Prosternal process acuminated (Fig. 17 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl 2005) strongly wavy (Fig. 17 B). Anterior phragma large and straight (Fig. 17 C). Mesanepisternum with strigulate microsculpture. Procoxal rests triangular; slightly curved posteriorly (Fig. 17 D). Mesoventral lines wavy and obtuse; median lines moderately wavy; area between median and mesoventral lines thin (Fig. 17 D). Mesoventral process long, slightly curved at top and base, forming a not well marked ridge (Fig. 17 E). METATHORAX. Metanotum with alacrista somewhat enlarged anteriorly and turned to anterior portion; scutoscutellar suture very strongly curved; median membranous area remarkably wide, and short (Fig. 17 F). Metaventrite with strigulate microsculpture; punctuation sparse, fine; coarsely punctate above intercoxal plates (Fig. 17 D). Mesocoxal line forming a smooth angle between coxal cavities (Fig. 17 D). Metanepisternum and metepimeron with imbricate microsculpture. Intercoxal plates with strigulate microsculpture. Metendosternite with curved furcal arms; ‘ stalk ridge’ exceeding half length of stalk (Fig. 18 A); ventral longitudinal flange long and narrow in lateral view (Fig. 18 B). WINGS. Elytra slightly wider than long, partially covering tergite VI (Fig 14 A); basal and lateral lines punctate (Fig. 16 D); sutural lines dashed; adsutural area with a row of setae; six rows of coarse punctures (not including sutural line) (Figs 14 A, D, 18 C); apical coarse punctation sparse; apical fine punctation dense (Fig. 18 D); apical serrations almost inconspicuous (Fig. 18 D); pubescence short and fine. Epipleuron with a line of coarse punctures. Hind wings fully developed (Fig. 18 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Femora strongly fusiform (Fig. 18 F – H). Profemora with inconspicuous punctuation; mesofemora with sparse and coarse punctuation; metafemora with fine and shallow punctuation. Mesotibiae densely spinose, spines fine; metatibiae sparsely spinose, spines fine. ABDOMEN. Tergite VI and VII with imbricate microsculpture (Fig. 19 A). Tergite VII trapezoidal (Fig. 19 A), triangular in some paratypes with tergite VIII not exposed; punctation sparse, fine; pubescence sparse, fine. Ventrites 1 – 5 sparsely and finely punctate; pubescence sparse, fine; with strigulate microsculpture (Fig. 19 B – C). Metacoxal lines finely punctate. Males MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.14 (0.05), 0.10 (0.04), 0.08 (0.03), 0.07 (0.04), 0.06 (0.04), 0.06 (0.05), 0.09 (0.08), 0.08 (0.08), 0.09 (0.10), 0.09 (0.11), 0.14 (0.10); (n = 8, including the holotype, unless otherwise specified; in mm): TL 2.07 – 2.32 (mean = 2.18, standard deviation ± 0.08), PL 0.74 – 0.80 (0.78 ± 0.02), PA 0.66 – 0.70 (0.69 ± 0.02), PB 1.30 – 1.40 (1.36 ± 0.04), SL (n = 7) 0.07 – 0.15 (0.11 ± 0.02), SW (n = 7) 0.09 – 0.14 (0.11 ± 0.02), EI 1.24 – 1.36 (1.30 ± 0.04), EL 1.44 – 1.62 (1.53 ± 0.08), EW 0.76 – 0.86 (0.81 ± 0.03), EH 0.50 – 0.62 (0.55 ± 0.04), HW 0.59 – 0.65 (0.62 ± 0.02), IS 0.15 – 0.20 (0.17 ± 0.02), WA 0.12 – 0.19 (0.14 ± 0.02), MC 0.55 – 0.64 (0.60 ± 0.03), MB 0.23 – 0.30 (0.27 ± 0.02), VL 0.40 – 0.49 (0.45 ± 0.03). Antennae shorter than in females; club less distinct than in females (Fig. 15 B). Pro- and mesotarsomeres I – III enlarged, with tenet setae (Fig. 18 F – G). Tergite VIII heptagonal, acuminated posteriorly; punctuation inconspicuous; subglabrous (Fig. 19 D). Tergite IX with rectangular ventral struts (Fig. 19 E – F). Sternite VIII rectangular, with a projection (Fig. 19 G). Sternite IX straight, constricted centrally and thicker apically (Fig. 19 F). Aedeagus sclerotized, apex of median lobe short; openings in dorsal view forming an acute angle (Fig. 20 A – C); internal sac with weak irregular sclerites, with drop-like sclerites (Fig. 20 D – E); parameres short, enlarged apically in lateral view (Fig. 20 A). Females MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.14 (0.05), 0.11 (0.05), 0.11 (0.04), 0.06 (0.04), 0.06 (0.05), 0.05 (0.06), 0.10 (0.11), 0.09 (0.13), 0.07 (0.13), 0.08 (0.14), 0.12 (0.12); (n = 7, paratypes; in mm): TL 2.10 – 2.35 (mean = 2.26, standard deviation ± 0.09), PL 0.76 – 0.86 (0.81 ± 0.04), PA 0.66 – 0.74 (0.71 ± 0.02), PB 1.34 – 1.50 (1.43 ± 0.06), SL 0.09 – 0.13 (0.11 ± 0.01), SW 0.12 – 0.14 (0.13 ± 0.01), EI 1.30 – 1.40 (1.35 ± 0.04), EL 1.48 – 1.66 (1.58 ± 0.06), EW 0.76 – 0.84 (0.81 ± 0.03), EH 0.52 – 0.65 (0.60 ± 0.04), HW 0.61 – 0.66 (0.63 ± 0.02), IS 0.18 – 0.20 (0.18 ± 0.01), WA 0.13 – 0.15 (0.14 ± 0.00), MC 0.57 – 0.65 (0.62 ± 0.03), MB 0.25 – 0.32 (0.30 ± 0.02), VL 0.43 – 0.52 (0.50 ± 0.03). Antennae larger than in males; antennal club more distinct (Fig. 15 C). Tergite VIII hexagonal, with a posterior invagination; punctuation inconspicuous; subglabrous (Fig. 21 A). Sternite VIII rectangular with a projection (Fig. 21 B). Vagina membranous; bursa copulatrix with sclerites on wall (Fig. 21 C, Supp. file 2 A). Vaginal plate with an apical sclerite, more or less developed (Fig. 21 C – D, Supp. file 2 A). Two filiform spermatheca (Supp. file 2 A). Distal gonocoxites straight, thin (Fig. 21 C – E); gonostyli long, somewhat thick, and parallel (Fig. 21 C – E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	biology_ecology	Host fungi Adults were collected from Psathyrella candolleana (Fr.) Maire (Agaricales, Psathyrellaceae) (1 record, 1 individual), Psathyrella sp. (1, 11), Agaricus sp. (1, 18; von Groll et al. 2021: figs 1 – 4), A. dulcidulus Schulzer (1, 4), A. sylvaticus Schaeff. (1, 20) and Entoloma (Inocephalus) sp. (Agaricales, Entolomataceae) (1, 2).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	discussion	Remarks Measurements similar to those of C. newtoni, except for the comparatively longer antennae. It was found co-occurring with C. newtoni in the same host fungi. However, they differ at first view by the coloration: C. lescheni sp. nov. is brown with few individuals of the same coloration as C. newtoni (pronotum reddish brown and elytra black); a few individuals of C. newtoni are brown, but darker than C. lescheni. They also differ by the microsculpture of the metaventrite: present in C. lescheni and absent in C. newtoni. Tergite VIII in males is triangular in C. lescheni and straight apically in C. newtoni. Parameres in C. newtoni are longer and the sclerite of the internal sac is also different. Tergite VIII in females has an invagination in C. lescheni, while in C. newtoni it is entire.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF402FFF39857FA44FC95FE49.taxon	distribution	Distribution Known only from Mata da Biologia and Mata do Paraíso, campus of the Universidade Federal de Viçosa, Viçosa, state of Minas Gerais, Southeast Brazil (Fig. 46).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	description	urn: lsid: zoobank. org: act: 88 FF 6 EF 0 - 5 AED- 4505 - 9 E 56 - 2200 ED 4 ABEEF Figs 4, 22 – 29, 46; Supp. file 1 C	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	diagnosis	Diagnosis TL: 2.53 – 2.78 mm in males and 2.34 – 2.68 mm in females. Pronotum, hypomeron and scutellum reddish brown (Fig. 22 A). Elytra black, anterior region reddish brown (Fig. 22 A). Hypomeron and mesanepisternum with strigulate microsculpture. Metaventrite smooth, coarsely punctate above intercoxal plates (Fig. 25 D). Metanepisternum and metepimeron with imbricate microsculpture. Aedeagus apex long, parameres longish, weak sclerites in internal sac (Fig. 28 A – E). Distal gonocoxites straight and slender (Fig. 29 E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	etymology	Etymology In homage to Dr Ivan Löbl (Muséum d’histoire naturelle, Genève, CH), for his remarkable contributions to the systematics of Scaphidiinae, which are the greatest source of information for our ongoing works on these beetles.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	materials_examined	Material examined Holotype BRAZIL • ♂; Minas Gerais, Viçosa, EPTEA Mata do Paraíso; 12 Nov. 2019; LabCol leg.; “ \\ Em Xylodon flaviporus \\ Cyparium loebli von Groll & Lopes-Andrade HOLOTYPUS ” [red paper]; CELC (Supp. file 1 C). Paratypes BRAZIL • 1 ♂, 1 ♀ (same pin); same collection data as for holotype; 8 Dec. 2014; I. S. C. Pecci-Maddalena leg.; CELC • 1 ♂, 2 ♀ (1 ♀, abdomen and head dissected, stored in glycerin); same collection data as for holotype, “ Trilha dos Gigantes ”; 15 Feb. 2015; S. Aloquio, A. Orsetti and M. Bento leg.; CELC • 2 specs; same collection data as for holotype; 19 Feb. 2015; FIT; A. Orsetti, S. Aloquio and M. Bento leg.; CELC • 1 ♂; same collection data as for holotype; S. Aloquio, A. Orsetti and M. Bento leg.; CELC • 6 ♂♂, 2 ♀♀ (1 ♂ entirely dissected, preserved in glycerin); same collection data as for holotype; “ T. da Madeira ”; 27 Feb. 2015; I. S. C. Pecci-Maddalena et al. leg.; CELC • 1 ♂, 1 ♀; same collection data as for holotype; CAMB • 1 ♀ (head dissected, preserved in glycerin); same collection data as for holotype; 13 Mar. 2015; S. Aloquio, A. Orsetti, C. Lopes-Andrade and M. Bento leg.; CELC • 1 ♂; same collection data as for holotype; 20 Mar. 2015; A. Orsetti and I. Gonçalves leg.; CELC • 6 ♂♂, 2 ♀♀ (2 ♂♂ dissected, preserved in glycerin); same collection data as for holotype; 9 Nov. 2016; I. Pecci-Maddalena and C. Lopes-Andrade leg.; “ \\ ex Psathyrella candolleana ”; CELC • 1 ♂, 1 ♀; same collection data as for holotype; CERPE • 2 ♂♂, 5 ♀♀ (1 ♀, abdomen dissected, preserved in glycerin); same collection data as for holotype; 19 Nov. 2019; LabCol leg.; “ \\ em Agaricus sylvaticus ”; CELC. All paratypes additionally labelled “ Cyparium loebli von Groll & Lopes-Andrade PARATYPUS [yellow paper] ”.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	description	Description MEASUREMENTS (holotype, in mm). TL 2.59, PL 1.00, PA 0.80, PB 1.72, EW 1.06, EL 1.86, IS 0.22, HW 0.72. COLORATION. Black, iridescent (Fig. 22 A – C). Frons dark brown; clypeus yellowish-brown; mouthparts yellow (Fig. 23 A); antennomeres I – VI and apex of XI yellow; VII – basal part of XI darker (Fig. 23 A – C). Pronotum, hypomeron and scutellum reddish brown (Fig. 22 A). Elytra black, anterior region reddish brown (Fig. 22 A). Meso- and metathorax in ventral view brown to dark brown. Procoxae dark ochre; meso- and metacoxae brownish red; femora dark brown, apex light brown; tibiae dark ochre, base and apex lighter; tarsi yellow (Fig. 22 B – C). Ventrite 1 dark brown to brown; each next segment lighter; ventrites 5 and 6 yellowish. Variation: some paratypes with pronotum and base of elytra ochreous (Fig. 22 D – F). HEAD. Punctation dense, fine (Fig. 23 A). Eyes wide and rounded (Fig. 23 A). Labrum rectangular, lateral margins rounded, well delimited apically; central margins straight and wide; sclerotized portion inwardly curved; lateral setae well exceeding margins of labrum; densely porose centrally (Fig. 23 D). Mandibles strongly curved and somewhat long apically; subapical serrations on left mandible conspicuous (Fig. 23 E – F). Maxillae with palpomere III short; galea with a row of lateral setae; galea densely pubescent, lacinia moderately pubescent, wider than galea (Fig. 23 G). Mentum forming a straight projection apically (Fig. 24 A). Setae of labial palpomere II far exceeding palpomere III; palpomere III longish, with short apical setae (Fig. 24 A – B). Post-gena microsculptured with close transversal lines; gula densely porose, but limited to central region; gula long and narrow (Fig. 24 C). Antennal club well distinct; antennomere XI pentagonal, rounded apically (straight in some individuals); no notable difference between sexes (Fig. 23 B – C). PROTHORAX. Pronotum smooth, shining; punctuation dense, fine; pubescence short, fine (Fig. 24 D – E); transverse, sub-straight laterally, forming an obtuse angle at lateral areas of posterior margin (Fig. 24 E). Hypomeron with strigulate microsculpture. Notosternal suture straight, slightly inward directed (Fig. 24 F). Profurca thin, only reaching half length of foramen (Fig. 24 G). Prosternal process rounded (Fig. 25 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl 2005) wavy (Fig. 25 B). Scutellum tapering posteriorly (Fig. 25 B). Anterior phragma large and obtuse (Fig. 25 C). Mesanepisternum with strigulate microsculpture. Procoxal rests triangular, strongly curved posteriorly (Fig. 25 D). Mesoventral and median lines wavy; area between median and mesocoxal lines not specially enlarged (Fig. 25 D). Process of metaventrite curved at top and straight at base, forming a ridge (Fig. 25 E). METATHORAX. Metanotum with alacrista enlarged anteriorly and turned to sides; scutoscutellar suture curved; median membranous area wide and short (Fig. 25 F). Metaventrite smooth, punctuation sparse and fine; coarsely punctate above intercoxal plates (Figs 22 B, 25 D). Mesocoxal line forming an angle between coxal cavities, and finely punctate under coxal cavities (Fig. 25 D). Metanepisternum and metepimeron with imbricate microsculpture. Intercoxal plates smooth. Metendosternite with arms almost straight; ‘ stalk ridge’ exceeding half length of stalk (Fig. 26 A); ventral longitudinal flange curved in lateral view (Fig. 26 B). WINGS. Elytra slightly wider than longer; partially covering tergite VI (Fig. 26 A); basal and lateral lines punctate (Fig. 24 D); sutural line dashed; adsutural area with a row of setae; six rows of coarse punctures (not including sutural line), but rows 5 and 6 somewhat intermixed (Figs 22 A, 26 C); apical coarse punctation sparse; apical serrations small, sparse; pubescence short and fine (Fig. 26 D). Epipleuron with diffuse, coarse, and close punctures. Hind wings fully developed (Fig. 26 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Femora somewhat fusiform (Fig. 26 F – H). Pro- and mesofemora sparsely and coarsely punctate; metafemora with shallow and fine punctation. Mesotibia densely spinose, spines fine (Fig. 26 G). Metatibiae sparsely spinose, spine fine (Fig. 26 H). ABDOMEN. Tergites VI – VIII with imbricate microsculpture (Fig. 27 A). Tergite VII triangular; punctation sparse, coarse; pubescence sparse, fine (Fig. 27 A). Ventrites (Fig. 27 B) sparsely pubescent; with strigulate microsculpture. Ventrite 1 sparsely and coarsely punctate (Fig. 27 C). Ventrites 2 – 5 moderately sparse and finely punctate. Metacoxal lines finely punctate. Males MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.15 (0.06), 0.11 (0.05), 0.10 (0.04), 0.08 (0.04), 0.08 (0.05), 0.06 (0.06), 0.09 (0.11), 0.08 (0.12), 0.08 (0.13), 0.08 (0.14), 0.13 (0.14); (n = 7, including the holotype; in mm): TL 2.53 – 2.78 (mean = 2.65, standard deviation ± 0.08), PL 1.00 – 1.08 (1.02 ± 0.03), PA 0.80 – 0.84 (0.82 ± 0.02), PB 1.60 – 1.80 (1.70 ± 0.06), SL 0.13 – 0.17 (0.15 ± 0.02), SW 0.13 – 0.16 (0.14 ± 0.01), EI 1.48 – 1.62 (1.52 ± 0.05), EL 1.74 – 1.90 (1.82 ± 0.05), EW 0.92 – 1.06 (0.98 ± 0.05), EH 0.43 – 0.71 (0.63 ± 0.10), HW 0.70 – 0.74 (0.72 ± 0.01), IS 0.20 – 0.23 (0.21 ± 0.01), WA 0.16 – 0.19 (0.17 ± 0.01), MC 0.68 – 0.74 (0.70 ± 0.02), MB 0.26 – 0.36 (0.32 ± 0.04), VL 0.52 – 0.56 (0.54 ± 0.02). Pro- and mesotarsomeres I – III enlarged, with tenet setae (Fig. 26 F – G). Tergite VIII heptagonal, acuminate posteriorly; punctuation fine, almost inconspicuous; subglabrous (Fig. 27 D). Tergite IX with more or less bent ventral struts (Fig. 27 E – F). Sternite VIII rectangular, with a small projection (Fig. 27 G). Sternite IX triangular at ends and centrally constricted (Fig. 27 F). Aedeagus sclerotized, enlarged at base, apex of median lobe long (Fig. 28 A – C); openings in dorsal view long and somewhat enlarged, forming an acute angle (Fig. 28 C); internal sac with weak irregular sclerites, with two hooks (Fig. 28 D – E); parameres thin, longish (Fig. 28 A – B). Females MEASUREMENTS (n = 1, paratype). Antennomeres (length (width )): 0.16 (0.06), 0.10 (0.05), 0.09 (0.04), 0.07 (0.04), 0.07 (0.05), 0.05 (0.06), 0.08 (0.11), 0.08 (0.13), 0.08 (0.15), 0.08 (0.15), 0.14 (0.14); (n = 7, paratypes; in mm): TL 2.34 – 2.68 (mean = 2.57, standard deviation ± 0.12), PL 0.86 – 1.02 (0.96 ± 0.05), PA 0.76 – 0.86 (0.82 ± 0.04), PB 1.50 – 1.78 (1.68 ± 0.09), SL 0.13 – 0.17 (0.15 ± 0.01), SW 0.13 – 0.17 (0.14 ± 0.01), EI 1.40 – 1.64 (1.52 ± 0.08), EL 1.64 – 1.98 (1.83 ± 0.11), EW 0.86 – 1.06 (0.95 ± 0.07), EH 0.65 – 0.75 (0.69 ± 0.03), HW 0.66 – 0.74 (0.70 ± 0.02), IS 0.20 – 0.25 (0.23 ± 0.02), WA 0.17 – 0.19 (0.18 ± 0.01), MC 0.60 – 0.76 (0.71 ± 0.06), MB 0.29 – 0.36 (0.32 ± 0.03), VL 0.46 – 0.58 (0.54 ± 0.04). Tergite VIII triangular; punctation inconspicuous; subglabrous (Fig. 29 A). Sternite VIII rectangular with a distinct thin projection (Fig. 29 B). Vagina and bursa copulatrix membranous without sclerites (Fig. 29 C). Vaginal plate with an apical T-shaped sclerite (Fig. 29 D). Spermatheca not detected. Distal gonocoxites straight, slender (Fig. 29 C, E); gonostyli parallel, long, and slender (Fig. 29 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	biology_ecology	Host fungi Adults were collected from Psathyrella candolleana (1 record, 10 individuals), Xylodon flaviporus (Berk. & M. A. Curtis ex Cooke) Riebesehl & E. Langer (Hymenochaetales, Schizoporaceae) (1, 1) and Agaricus sylvaticus (1, 7).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	discussion	Remarks Similar to C. lescheni sp. nov., especially the teneral specimens, in the strigulate microsculpture of the hypomeron and metaventrite, reddish brown pronotum, and the size of the antennae, but differ in the comparatively larger body length and in the reddish brown anterior region of the elytra.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF439FFFD98A8FDD9FC95FBE9.taxon	distribution	Distribution Known only from Mata da Biologia and Mata do Paraíso, campus of the Universidade Federal de Viçosa, Viçosa, state of Minas Gerais, Southeast Brazil (Fig. 46).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	description	urn: lsid: zoobank. org: act: 8 E 4 DB 834 - 61 A 4 - 47 B 2 - B 9 B 3 - 2 C 0606708142 Figs 4, 30 – 37, 46; Supp. file 1 D	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	diagnosis	Diagnosis TL: 2.00 – 2.22 mm in males and 2.00 – 2.30 mm in females. Pronotum reddish brown (Fig. 32 D). Elytra black (Fig. 30 A). Hypomeron with strigulate microsculpture. Metaventrite smooth; coarsely punctate above intercoxal plates (Figs 30 B, 33 D). Intercoxal plates smooth. Apex of tergite VIII in males straight (Fig. 35 D). Aedeagus openings in dorsal view narrow, forming an obtuse angle (Fig. 36 C). Internal sac with a plate-like sclerite (Fig. 36 D). Tergite VIII in females without apical invagination (Fig. 37 A).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	etymology	Etymology In homage to Dr Alfred F. Newton (Field Museum, Chicago, USA) for his significant contribution to the systematics of Staphylinidae and more specifically of Scaphidiinae.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	materials_examined	Material examined Holotype BRAZIL • ♂; Minas Gerais, Viçosa, EPTEA Mata do Paraíso; 19 Nov. 2019; LabCol leg.; “ \\ em Agaricus sylvaticus \\ Cyparium newtoni von Groll & Lopes-Andrade HOLOTYPUS ” [red paper]; CELC (Supp. file 1 D). Paratypes BRAZIL • 1 ♀; same collection data as for holotype; 19 Feb. 2015; S. Aloquio, A. Orsetti and M. Bento leg.; “ \\ Trilha caminho das águas embaixo de casca de tronco caído ”; CELC • 1 ♀; same collection data as for holotype, “ T. dos Gigantes ”; 15 Feb. 2015; S. Aloquio, A. Orsetti and M. Bento leg.; CELC • 3 ♂♂, 1 ♀, 2 specs; same collection data as for holotype, “ Trilha da Madeira ”; 27 Feb. 2015; I. S. C. PecciMaddalena et al. leg.; CELC • 1 ♂, 1 ♀; same collection data as for holotype; CAMB • 2 ♂♂, 1 ♀; same collection data as for holotype; 13 Mar. 2015; S. Aloquio, A. Orsetti, C. Lopes-Andrade and M. Bento leg.; CELC • 5 ♂♂, 4 ♀♀ (2 ♂♂ dissected, preserved in glycerin); same collection data as for holotype; 9 Nov. 2016; I. Pecci-Maddalena and C. Lopes-Andrade leg.; “ \\ ex. Psathyrella candolleana ”; CELC • 2 ♂♂, 2 ♀♀ (1 ♂ dissected, preserved in glycerin); same collection data as for holotype; 21 Nov. 2019; LabCol leg.; “ \\ Em Agaricus dulcidulus e Leucocoprinus brebissoni ”; CELC • 1 ♂, 2 ♀♀, 1 spec. (1 ♂, abdomen dissected, preserved in glycerin); same collection data as for holotype; “ \\ Em Agaricus sylvaticus ”; CELC • 1 ♂, 1 ♀ (1 ♂, abdomen dissected, preserved in glycerin); Viçosa, Recanto das Cigarras, Mata da Biol.; 20 Nov. 2019; LabCol leg.; “ Fungo 29 \\ Em Entoloma (Inocephalus) sp. ”; CELC • 2 ♂♂, 3 ♀♀, 1 spec. (1 ♂, entirely dissected, preserved in glycerin; 1 ♂, 1 ♀ (abdomen dissected, preserved in glycerin); same collection data as for preceding; “ Fungo 08 \\ Em Psathyrella sp. ”; CELC • 1 ♂, 3 ♀♀; Viçosa, Mata da Biologia; 15 Oct. 2021; E. von Groll and A. Orsetti leg.; “ Fungo 20 \\ Em Agaricus sp. ”; CELC. All paratypes additionally labelled “ Cyparium newtoni von Groll & Lopes-Andrade PARATYPUS [yellow paper] ”.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	description	Description MEASUREMENTS (holotype, in mm). TL 2.00, PL 0.72, PA 0.60, PB 1.30, EW 0.74, EL 1.44, IS 0.17, HW 0.56. COLORATION. Frons dark brown; clypeus and mouthparts yellowish brown (Fig. 30 D); antennomeres I – VI yellowish brown; VII – XI darker (Fig. 31 A – C). Pronotum and hypomeron reddish brown (Fig. 30 A – B). Scutellum ochre or black (Fig. 32 D). Meso- and metathorax reddish-brown, each sclerite lighter laterally (Fig. 30 B). Elytra black; epipleuron dark ochre (Fig. 30 A). Coxae and trochanters dark ochre; femora dark ochre, apex lighter; tibiae dark ochre; lighter anteriorly and posteriorly; tarsi yellow (Fig. 30 B – C). Tergite VIII yellow. Ventrite 1 dark ochre, 2 – 4 ochre, 5 and 6 yellow (Fig. 35 C). Variation: few paratypes entirely dark brown, with lighter tarsi; others with reddish brown area less vibrant. HEAD. Punctation dense, fine (Fig. 31 A). Eyes globose in frontal view (Fig. 31 A). Labrum rectangular, lateral margins rounded, well delimited apically; central margin straight; sclerotized portion inwardly curved; lateral setae extending well beyond margins of labrum; little porose centrally (Fig. 31 D). Mandibles slightly curved; subapical serrations on left mandible conspicuous (Fig. 31 E – F). Maxillary palps elongated; galea and lacinia moderately pubescent, lacinia elongated (Fig. 31 G). Mentum slightly curved apically (Fig. 32 A – B). Setae of labial palpomere II extending beyond palpomere III; palpomere III longish with short apical setae (Fig. 32 A – B). Hypopharynx with wide and rounded sclerotized plate (Fig. 32 A – B). Post gena microsculptured with transversal lines; few gular pores (Fig. 32 C); gula narrow. Antennal club well distinct; antennomere XI longish, hexagonal (apex rounded in some specimens); antennal club different between sexes (Fig. 31 B – C). PROTHORAX. Pronotum smooth, shining; punctuation dense, fine; pubescence short, fine (Fig. 32 D – E); transverse, sub-straight laterally, forming an obtuse angle at lateral areas of posterior margin (Fig. 32 E). Hypomeron with strigulate microsculpture. Notosternal suture straight, inward directed (Fig. 32 F). Profurca thin and elongated, slightly extending beyond half of foramen (Fig. 32 G). Prosternal process acuminated (Fig. 33 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl 2005) strongly wavy (Fig. 33 B). Scutellum rounded posteriorly (Fig. 33 B). Anterior phragma large and straight (Fig. 33 C). Mesanepisternum with strigulate microsculpture. Procoxal rests wide; slightly curved posteriorly (Fig. 33 D). Mesoventral lines wavy and obtuse; median lines moderately wavy; area between median and mesoventral lines not specially enlarged (Fig. 33 D). Process of metaventrite short, slightly curved at top and base, forming a small ridge (Fig. 33 E). METATHORAX. Metanotum with alacrista enlarged anteriorly and turned to the sides; scutoscutellar suture strongly curved; median membranous area wide and short (Fig. 33 F). Metaventrite smooth; punctuation sparse, fine; coarsely punctate above intercoxal plates (Fig. 33 D). Mesocoxal line not forming an angle between coxal cavities, just a simple triangle, and finely punctate under coxal cavities (Fig. 33 D). Metanepisternum smooth; metepimeron with imbricate microsculpture. Intercoxal plates smooth. Metendosternite with arms almost straight; ‘ stalk ridge’ not exceeding half of stalk (Fig. 34 A); ventral longitudinal flange small, curved in lateral view (Fig. 34 B). WINGS. Elytra slightly wider than longer; partially covering tergite VI (Fig. 30 A); basal and lateral lines punctate; sutural lines dashed (Fig. 32 D); adsutural area with a row of setae; six rows of coarse punctures (not including sutural line) (Figs 30 A, 34 C); apical coarse punctation sparse; apical serrations almost inconspicuous (Fig. 34 D); pubescence short and fine. Epipleuron with diffuse and coarse punctures. Hind wings fully developed (Fig. 34 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Femora longish, somewhat fusiform (Fig. 34 F – H). Pro- and mesofemora sparsely and coarsely punctate; metafemora with shallow and sparse punctuation. Mesotibiae densely spinose, spines fine (Fig. 34 G); metatibiae sparsely spinose, spines fine (Fig. 34 H). ABDOMEN. Tergite VI – VIII with imbricate microsculpture (Fig. 35 A). Tergite VII trapezoidal (Fig 35 A) – triangular in some paratypes with tergite VIII not exposed; punctation sparse, fine; pubescence sparse, fine. Ventrites 1 – 5 (Fig. 35 B) sparsely and finely punctate; pubescence sparse, fine; with strigulate microsculpture (Fig. 35 C). Metacoxal lines finely punctate. Males MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.13 (0.06), 0.10 (0.04), 0.08 (0.03), 0.05 (0.03), 0.06 (0.04), 0.05 (0.05), 0.08 (0.09), 0.06 (0.09), 0.07 (0.10), 0.06 (0.11), 0.13 (0.11); (n = 7, including holotype; in mm): TL 2.00 – 2.22 (mean = 2.07, standard deviation ± 0.08), PL 0.70 – 0.80 (0.74 ± 0.03), PA 0.60 – 0.68 (0.63 ± 0.02), PB 1.28 – 1.40 (1.33 ± 0.04), SL 0.09 – 0.12 (0.10 ± 0.01), SW 0.10 – 0.12 (0.10 ± 0.01), EI 1.20 – 1.32 (1.25 ± 0.05), EL 1.40 – 1.54 (1.46 ± 0.05), EW 0.66 – 0.86 (0.77 ± 0.06), EH 0.50 – 0.62 (0.58 ± 0.04), HW 0.52 – 0.62 (0.57 ± 0.03), IS 0.17 – 0.18 (0.17 ± 0.00), WA 0.10 – 0.14 (0.13 ± 0.01), MC 0.52 – 0.61 (0.57 ± 0.02), MB 0.25 – 0.28 (0.27 ± 0.01), VL 0.42 – 0.56 (0.48 ± 0.04). Antennal club less distinct than in females (Fig. 31 B). Pro- and mesotarsomeres I – III enlarged, with tenet setae (Fig. 34 F – G). Tergite VIII hexagonal, straight posteriorly; punctuation inconspicuous; subglabrous (Fig. 35 D). Tergite IX with constricted ventral struts (Fig. 35 E – F). Sternite VIII sub-rectangular, with projection (Fig. 35 G). Sternite IX curved (Fig. 35 F). Aedeagus sclerotized, apex of median lobe short (Fig. 36 A, B); openings in dorsal view narrow, forming an obtuse angle (Fig. 36 C); internal sac with weak irregular sclerites with two hooks and a central plate (Fig. 36 D – E); parameres long, thin (Fig. 36 B). Females MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.13 (0.05), 0.10 (0.05), 0.09 (0.03), 0.05 (0.04), 0.06 (0.04), 0.05 (0.05), 0.08 (0.10), 0.09 (0.11), 0.09 (0.13), 0.09 (0.14), 0.13 (0.12); (n = 7, paratypes; in mm): TL 2.00 – 2.30 (mean = 2.19, standard deviation ± 0.12), PL 0.76 – 0.84 (0.78 ± 0.03), PA 0.6 – 0.7 (0.64 ± 0.04), PB 1.30 – 1.48 (1.41 ± 0.07), SL 0.11 – 0.15 (0.12 ± 0.01), SW 0.11 – 0.14 (0.12 ± 0.01), EI 1.22 – 1.44 (1.34 ± 0.07), EL 1.44 – 1.64 (1.58 ± 0.06), EW 0.80 – 0.88 (0.83 ± 0.02), EH 0.57 – 1.07 (0.65 ± 0.18), HW 0.31 – 0.62 (0.55 ± 0.11), IS 0.16 – 0.18 (0.17 ± 0.01), WA 0.14 – 0.17 (0.15 ± 0.01), MC 0.56 – 0.65 (0.61 ± 0.03), MB 0.27 – 0.32 (0.30 ± 0.01), VL 0.45 – 0.52 (0.50 ± 0.03). Antennal club more distinct than in males (Fig. 31 B – C). Tergite VIII hexagonal; punctuation inconspicuous; subglabrous (Fig. 37 A). Sternite VIII rectangular with a smooth projection (Fig. 37 B). Vagina and bursa copulatrix membranous without sclerites (Fig. 37 C). Vaginal plate with an apical sclerite (Fig. 37 D). Spermatheca not detected. Distal gonocoxites straight and thin (Fig. 37 C, E); gonostyli long, slender, larger at base (Fig. 37 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	biology_ecology	Host fungi Adults were collected from Psathyrella candolleana (1 record, 9 individuals), Psathyrella sp. (1, 6), Agaricus sp. (1, 4; von Groll et al. 2021: figs 1 – 4), A. dulcidulus (1, 4), A. sylvaticus (1, 5) and Entoloma (Inocephalus) sp. (1, 2).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	discussion	Remarks Extremely similar to C. lescheni sp. nov. (for comparison, see remarks under C. lescheni above and comparison of morphology below).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF437FFE79859FB78FC95F960.taxon	distribution	Distribution Known only from Mata da Biologia and Mata do Paraíso, campus of the Universidade Federal de Viçosa, Viçosa, state of Minas Gerais, Southeast Brazil (Fig. 46).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	description	urn: lsid: zoobank. org: act: 7 BE 94 FA 9 - A 727 - 4779 - B 8 FE- 975 D 5 C 178 ABC Figs 5, 38 – 46; Supp. file 1 E	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	diagnosis	Diagnosis TL: 3.35 – 4.20 mm in males and 4.20 – 4.35 mm in females. Black. Antennae entirely yellow; club lighter (Fig. 38 A). Anterior region of elytra reddish brown (Fig. 38 A). Hypomeron and mesanepisternum with close strigulate microsculpture. Metaventrite smooth; coarsely punctate above intercoxal plates (Fig. 38 B, E). Ventrites 1 – 5 densely and coarsely punctate (Fig. 43 C). Aedeagus strongly sclerotized, apex short; parameres short (Fig. 44 A – B). Sclerites of internal sac strong (Fig. 44 D – E). Distal gonocoxites short, straight and thick (Fig. 45 E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	etymology	Etymology In homage to Maurice Pic (1866 – 1957), who was responsible for gathering many specimens of scaphidiines deposited in museums, especially in the MNHN.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	materials_examined	Material examined Holotype BRAZIL • ♂; Mato Grosso, Cotriguaçu, Faz. São Nicolau, matinha do Fernando [Fernando’s woods]; 09 ° 50 ′ 19 ″ S, 58 ° 15 ′ 15 ″ W; FIT; 3 Nov. 2017; Vaz-de-Mello et al. leg.; “ \\ Cyparium pici von Groll & Lopes-Andrade HOLOTYPUS ” [red paper]; CEMT (Supp. file 1 E). Paratypes BRAZIL • 8 ♂♂, 5 ♀♀ (1 ♂ entirely dissected, preserved in glycerin); same collection data as for holotype; CEMT • 3 ♂♂, 2 ♀♀ (2 ♂♂, 1 ♀, entirely dissected, preserved in glycerin; 1 ♀, abdomen dissected, preserved in glycerin); same collection data as for holotype; CELC. All paratypes additionally labelled “ Cyparium pici von Groll & Lopes-Andrade PARATYPUS [yellow paper] ”.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	description	Description MEASUREMENTS (in mm). TL 3.45, PL 1.30, PA 1.12, PB 2.15, EW 1.42, EL 2.71, IS 0.28, HW 0.93. COLORATION. Black (Fig. 38 A – C). Frons dark brown; clypeus light brown; mouth parts and antennae yellow, club lighter (Fig. 39 A). Anterior region of elytra reddish brown (Fig. 38 A); epipleuron dark ochre. Femora dark brown; tibiae lighter; tarsi yellow (Fig. 38 B). Tergite VIII and ventrite 6 yellowish (Fig. 38 B). Variation: (1) entirely light brown (Fig. 38 D); (2) entirely dark brown with antennae, tarsi and tergite VIII yellow (Fig. 38 E – F). HEAD. Punctation dense, coarse (Fig. 39 A). Eyes slightly wider than head, rounded (Fig. 39 A). Labrum rectangular, lateral margins sub-straight, and well distinct from apical margin; central margin slightly curved centrally; sclerotized portion reaching apex; lateral setae slightly extending beyond margins; porose centrally (Fig. 39 D). Left mandible slightly curved and right mandible more curved; subapical serrations on left mandible conspicuous (Fig. 39 E – F). Maxillary palps elongated, palpomeres slender anteriorly; lacinia strongly robust, short and densely pubescent (Fig. 39 G). Mentum with lateral areas strongly rounded and apex well delimited (Fig. 40 A). Glossa heart-shaped (Fig. 40 A). Setae of labial palpomere II slightly exceeding palpomere III; palpomere III longish, with short apical setae (Fig. 40 A). Hypopharynx with wide and triangular sclerotized plate (Fig. 40 B). Post gena microsculptured with very close transversal lines; densely porose, except at region of gula; gula triangular and narrow (Fig. 40 C). Antennal club distinct; antennomere XI hexagonal, apex more or less acuminated, regardless of sex (Fig. 39 B – C). PROTHORAX. Pronotum smooth; punctuation dense, fine; pubescence short, fine (Fig. 40 D); transverse, rounded laterally, forming a small obtuse angle at lateral areas of posterior margin (Fig. 40 E). Hypomeron with close strigulate microsculpture. Notosternal suture straight (Fig. 40 F). Profurca elongated, only reaching half length of foramen (Fig. 40 G). Prosternal process short and round (Fig. 41 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl 2005) wavy (Fig. 41 B). Scutellum rounded posteriorly (Fig. 41 B). Anterior phragma straight (Fig. 41 C). Mesanepisternum with strigulate microsculpture. Procoxal rests triangular; slightly wavy posteriorly (Fig. 41 D). Mesoventral and median lines strongly wavy; area between median and mesoventral lines enlarged (Fig. 41 D). Process of metaventrite moderately long; apex more prominent, forming a ridge (Fig. 41 E). METATHORAX. Metanotum with alacrista triangular, small anteriorly and turned to posterior end; scutoscutellar suture not strongly wavy; median membranous area narrow and long (Fig. 41 F). Metaventrite smooth; punctuation sparse, fine; coarsely punctate above intercoxal plates (Figs 38 B, E, 41 D). Mesocoxal line forming a smooth angle between coxal cavities; finely punctate under coxal cavities (Fig. 41 D). Metanepisternum and metepimeron with imbricate microsculpture. Intercoxal plates smooth. Metendosternite with arms slightly curved; ‘ stalk ridge’ reaching half length of stalk (Fig. 42 A); ventral longitudinal flange curved in lateral view (Fig. 42 B). WINGS. Elytra slightly wider than longer; covering just until tergite V (Fig. 38 A, D); each elytron subrectangular (Fig. 42 C); basal (Fig. 40 D) and sutural lines dashed; adsutural area with a row of setae; six rows of coarse punctures (not including sutural line) (Figs 38 A, D, 42 C); lateral line punctate; apical coarse punctation moderately sparse; apical serrations almost inconspicuous (Fig. 42 D); pubescence short and fine. Epipleuron with a row of sparse and coarse punctures. Hind wings fully developed (Fig. 42 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Pro- and mesofemora fusiform (Fig. 42 F – G); punctuation sparse, coarse; metafemora longish, punctuation sparse, shallow (Fig. 42 H). Mesotibiae densely spinose, spines thick (Fig. 42 G); metatibiae sparsely spinose, spines fine (Fig. 42 H). ABDOMEN. Tergites VI – VIII with narrow imbricate microsculpture; punctures dense, coarse; pubescence sparse, coarse (Fig. 43 A). Ventrites 1 – 5 densely and coarsely punctate; pubescence moderately sparse, fine; close strigulate microsculpture (Fig. 43 B – C). Metacoxal lines finely punctate. Males MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.24 (0.08), 0.14 (0.07), 0.14 (0.06), 0.11 (0.06), 0.11 (0.07), 0.08 (0.09), 0.13 (0.16), 0.12 (0.18), 0.11 (0.19), 0.11 (0.20), 0.20 (0.22); (n = 10, including the holotype, unless otherwise specified; in mm): TL 3.35 – 4.20 (mean = 3.75, standard deviation ± 0.32), PL 1.30 – 1.64 (1.45 ± 0.13), PA 1.07 – 1.32 (1.16 ± 0.07), PB 2.15 – 2.62 (2.36 ± 0.17), SL (n = 8) 0.13 – 0.23 (0.17 ± 0.03), SW (n = 8) 0.13 – 0.23 (0.18 ± 0.03), EI 1.85 – 2.28 (2.06 ± 0.15), EL (n = 9) 2.37 – 2.75 (2.60 ± 0.13), EW 1.37 – 1.65 (1.49 ± 0.08), EH 0.75 – 1.15 (0.96 ± 0.11), HW (n = 9) 0.91 – 1.08 (0.99 ± 0.06), IS (n = 9) 0.26 – 0.31 (0.28 ± 0.02), WA (n = 9) 0.21 – 0.30 (0.24 ± 0.03), MC 0.93 – 1.10 (1.03 ± 0.05), MB 0.37 – 0.50 (0.45 ± 0.04), VL 0.62 – 0.78 (0.70 ± 0.04). Pro- and mesotarsomeres I – III enlarged, with tenet setae (Fig. 42 F – G). Tergite VIII trapezoidal, straight posteriorly; triangular in some dry paratypes (Fig. 43 D). Tergite IX with laterally curved ventral struts (Fig. 43 E – F). Sternite VIII sub-rectangular, with a smooth projection (Fig. 43 G). Sternite IX thick (Fig. 43 F). Aedeagus strongly sclerotized, apex of median lobe short; openings in dorsal view moderately narrow, forming an acute angle, almost right (Fig. 44 A – C); internal sac with strong and robust irregular sclerites (Fig. 44 D – E), parameres short (Fig. 44 A). Females MEASUREMENTS (n = 1, paratype; in mm). Antennomeres (length (width )): 0.25 (0.09), 0.15 (0.07), 0.15 (0.06), 0.11 (0.07), 0.10 (0.08), 0.08 (0.09), 0.13 (0.15), 0.13 (0.20), 0.11 (0.22), 0.11 (0.24), 0.19 (0.26); (n = 6, paratypes; in mm): TL 4.20 – 4.35 (mean = 4.26, standard deviation ± 0.06), PL 1.38 – 1.70 (1.62 ± 0.12), PA 1.22 – 1.35 (1.28 ± 0.05), PB 2.56 – 2.68 (2.62 ± 0.05), SL 0.15 – 0.25 (0.21 ± 0.03), SW 0.19 – 0.25 (0.22 ± 0.02), EI 2.25 – 2.47 (2.34 ± 0.10), EL 2.71 – 2.96 (2.79 ± 0.09), EW 1.62 – 1.80 (1.72 ± 0.06), EH 1.00 – 1.20 (1.10 ± 0.07), HW 1.05 – 1.15 (1.08 ± 0.03), IS 0.28 – 0.33 (0.31 ± 0.02), WA 0.22 – 0.28 (0.25 ± 0.02), MC 1.18 – 1.34 (1.26 ± 0.05), MB 0.56 – 0.62 (0.59 ± 0.03), VL 0.75 – 0.93 (0.85 ± 0.06). Tergite VIII hexagonal (Fig. 45 A), trapezoidal in dry specimens. Sternite VIII rectangular with a projection (Fig. 45 B). Vagina and bursa copulatrix membranous; vagina without sclerites; bursa copulatrix with irregular sclerites (Fig. 45 C). Vaginal plate with an apical M-shaped sclerite (Fig. 45 D). Spermatheca not detected. Distal gonocoxites short, straight and thick; gonostyli somewhat thick, parallel (Fig. 45 C – E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	discussion	Remarks Similar in body length to Cyparium rufohumerale, but differs in the yellowish antennae (bicolored in C. rufohumerale) and in the comparatively larger reddish brown mark on the elytra (only at the humeral region in C. rufohumerale). Also similar to C. humerale in body length and the entirely yellow antennae, but differs in the non-triangular mark on the elytra and in the shorter elytra.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF42DFF9298B3F902FE23FC27.taxon	distribution	Distribution Known only from Cotriguaçu, Faz. São Nicolau (09 ° 50 ′ 19 ″ S, 58 ° 15 ′ 15 ″ W), state of Mato Grosso, Midwest, Brazil (Fig. 46).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	description	Figs 4, 47 – 55; Supp. file 2 B, Supp. file 3 (Figs 1 A – 4 H)	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	materials_examined	Material examined BRAZIL – Sergipe • 3 ♂♂; Sta Luzia do Itanhi, Faz. Crasto; 9 – 12 Sep. 1999; A. Bonaldo leg.; MCN 166400 (abdomen dissected), 166404, 166405 • 3 ♀♀; same collection data as for preceding; MCN 166398, 166401, 167484 • 2 ♂♂; Areia Branca, Est. Ecol. Serra de Itabaiana; 14 – 20 Sep. 1999; A. Bonaldo leg.; MCN 166402, 166403 (abdomen dissected, preserved in glycerin) • 1 ♀; same collection data as for preceding; MCN 166406. – Minas Gerais • 1 ♂; Viçosa, EPTEA Mata do Paraíso; 8 Dec. 2014; I. PecciMaddalena leg.; CELC • 3 ♂♂, 5 ♀♀ (1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; 6 Dec. 2018; LabCol leg; CELC • 7 ♂♂, 3 ♀♀ (1 ♀ entirely dissected, preserved in glycerin; 1 ♂, 1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; 5 Nov. 2019; LabCol leg.; “ Fungo 02 \\ Em Pleurotus pulmonarius ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 03 \\ Em Mycena sp. ”; CELC • 14 ♂♂, 4 ♀♀ (1 ♂, 1 ♀ entirely dissected, preserved in glycerin; 1 ♂, 1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; “ Fungo 01 \\ Em Xerocomus aff. brasiliensis ”; CELC • 9 ♂♂, 3 ♀♀ (3 ♂♂ entirely dissected, preserved in glycerin); same collection data as for preceding; 7 Nov. 2019; LabCol leg.; “ Fungo 17 ”; CELC • 1 ♂; same collection data as for preceding; “ Fungo 14 \\ Em Leucoagaricus rubrotinctus ”; CELC • 1 ♂, 1 ♀; same collection data as for preceding; “ Fungo sem id ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo A 21 \\ Fotos: 712 - 713; 728 - 729, Em Heimiomyces neovelutipes ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo Que ”; CELC • 1 ♂, 2 ♀♀; same collection data as for preceding; 12 Nov. 2019; LabCol leg.; “ Fungo 09 ”; CELC • 2 ♀♀ (1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; 13 Nov. 2019; LabCol leg.; “ Fungo 18 \\ Em Macrolepiota colombiana ”; CELC • 1 ♂; same collection data as for preceding; 14 Nov. 2019; “ Fungo 22 \\ Em Marasmiellus sp. ”; CELC • 1 ♂; same collection data as for preceding; 19 Nov. 2019; “ Fungo 18; Em Marasmiellus cubensis ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 16 \\ Em Marasmiellus sp. ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 22 \\ Em Leucoprinus ianthinus ”; CELC • 1 ♂, 2 ♀♀; same collection data as for preceding; “ Fungo 13 \\ Em Marasmiellus cubensis ”; CELC • 2 ♀♀; same collection data as for preceding; “ Fungo 05 \\ Em Marasmius sp. ”; CELC • 2 ♀♀; same collection data as for preceding; “ Fungo 24 \\ Em Marasmiellus cubensis ”; CELC • 1 ♀; same collection data as for preceding; 21 Nov. 2019; LabCol leg.; “ Fungo 38 \\ Em Marasmiellus sp. ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 06 \\ Em Agaricus dulcidulus e Leucocoprinus brebissoni ”; CELC • 5 ♂♂, 2 ♀♀ (1 ♂, abdomen dissected, preserved in glycerin); same collection data as for preceding; “ Fungo 26 \\ Em Lulesia lignicola ”; CELC • 2 ♀♀; same collection data as for preceding; “ Fungo 16 \\ Em Marasmius araucariae ”; CELC • 2 ♂♂, 3 ♀♀ (1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; “ Fungo 12 \\ Em Marasmiellus aff. ramealis ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 52 \\ Em Marasmiellus sp. ”; CELC • 2 ♂♂, 1 ♀; same collection data as for preceding; “ Fungo 41 \\ Em Lepiota sp. ”; CELC • 2 ♂♂; same collection data as for preceding; 26 Nov. 2019; LabCol leg.; “ Fungo 23 \\ Em Marasmiellus cubensis ”; CELC • 1 ♂; same collection data as for preceding; “ Fungo 18 \\ Em Entoloma (Inocephalus) sp. ”; CELC • 2 ♀♀ (1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; “ Fungo 46 \\ Em Volvariella sp. ”; CELC • 1 ♂, 1 ♀; same collection data as for preceding; “ Fungo 24 \\ Em Marasmius sp. ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 19 \\ Pleteus sp. ”; CELC • 1 ♂; same collection data as for preceding; “ Fungo 37 \\ Conocybe sp. ”; CELC • 1 ♂; same collection data as for preceding; “ Fungo 26 \\ Em Leucocoprinus sp. ”; CELC • 3 ♂♂; same collection data as for preceding; “ Fungo 17 ”; CELC • 1 ♀; same collection data as for preceding; “ Fungo 39 \\ Em Marasmius hematocephalus ”; CELC • 2 ♂, 1 ♀ (1 ♂ entirely dissected, preserved in glycerin); same collection data as for preceding; 28 Nov. 2019; LabCol leg.; “ fotos 1103 - 05 \\ Em Marasmius sp. ”; CELC • 1 ♂; same collection data as for preceding, “ Trilha dos Gigantes ”; 3 Feb. 2019; LabCol leg.; CELC • 1 ♀; Viçosa, Mata da Biologia; 3 May 2014; S. Aloquio leg.; “ \\ ex. Lepiota sp. ”; CELC • 2 ♂♂, 2 ♀♀; Viçosa, Recanto das Cigarras, Mata da Biol.; 13 Nov. 2019; LabCol leg; “ Fungo 11 \\ Em Coprinellus disseminatus ”; CELC • 1 ♂; Viçosa, Mata da Biologia; 15 Oct. 2021; E. von Groll and A. Orsetti leg.; “ Fungo 06 ”; CELC • 1 ♂, 1 ♀; same collection data as for preceding; “ Fungo 26 \\ Em Leucocoprinus cepistipes ”; CELC • 1 ♂, 1 ♀; same collection data as for preceding; 23 Nov. 2021; E. von Groll and G. L. N. Martins leg.; “ \\ Em Favolus tenuiculus ”; CELC • 8 ♂♂, 1 ♀; same collection data as for preceding; 26 Nov. 2021; “ \\ Em Favolus tenuiculus ”; CELC.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	diagnosis	Diagnosis TL: 2.60 – 3.40 mm in males and 2.56 – 3.52 mm in females. Pronotum reddish brown, elytra black (Fig. 47 A). Eyes remarkably wider than head, rounded (Fig. 48 A). Antennomere XI hexagonal, longish (Fig. 48 B – C). Hypomeron only laterally with imbricate microsculpture. Metaventrite smooth, but laterally with imbricate microsculpture. Tergite VIII of males coarsely punctate (Fig. 52 D). Rounded ventral struts (Fig. 52 F). Aedeagus with short apex; openings in dorsal view thin, forming a wide acute angle; parameres short (Fig. 53 A – C). Distal gonocoxites curved and thick (Fig. 54 E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	description	Redescription COLORATION. Iridescent. Frons ochre, reddish brown; clypeus yellowish to ochre; mouthparts yellowbrown (Fig. 48 A, Supp. file 3 (Figs 1 D, 3 D )); antennomeres I – VI and apex of XI yellow-brown; VII – X and basal part of XI darker (Fig. 48 B). Pronotum and hypomeron reddish brown (Fig. 47 A – B, Supp. file 3 (Figs 1 E, 3 E )). Scutellum reddish brown or black (Fig. 49 D). Elytra black (Fig. 47 A); epipleuron dark ochre. Mesanepisternum, metanepisternum and metepimeron dark ochre. Metaventrite dark brown, lighter laterally (Fig. 47 B). Coxae and trochanters dark ochre; femora yellowish brown, apex yellow; tibiae and tarsi ochre (Fig. 47 B – C). Tergite VI and VII black; tergite VIII yellow. Ventrites 1 – 4 dark brown; 5 and 6 yellow. Teneral specimens entirely light brown (Fig. 47 D – E) or with pronotum lighter (Fig. 47 F). HEAD. Punctation dense, fine (Fig. 48 A). Eyes remarkably wider than head, rounded (Fig. 48 A). Labrum transverse, lateral areas rounded, not well marked at apex; central margin slightly curved; sclerotized portion rounded; lateral setae slightly exceeding margins of labrum; almost without pores centrally (Fig. 48 D). Mandibles smoothly curved, subapical serrations on left mandible conspicuous (Fig. 48 E – F). Maxillae with palpomere II long and thin; palpomere III somewhat globose; galea densely pubescent and lacinia moderately pubescent (Fig. 48 G). Mentum laterally rounded, curved apically (Fig. 49 A). Setae of labial palpomere II not exceeding palpomere III; palpomere III longish, with short apical setae (Fig. 49 A – B). Post gena microsculptured with close transversal lines; gular pores limited to region next to eyes and submentum; gula triangular, small (Fig. 49 C). Antennae not reaching or surpassing mesanepisternum, regardless of sex; antennal club smoothly delimited; antennomere XI hexagonal, longish (Fig. 48 B – C). PROTHORAX. Pronotum smooth, shining; punctation dense, fine; pubescence short, fine (Fig. 49 D); transverse, sub-straight laterally, almost forming a right angle at posterior margin (Fig. 49 E). Hypomeron laterally with imbricate microsculpture. Notosternal suture with an angle, outward directed (Fig. 49 F). Profurca thin, short, not exceeding half of foramen (Fig. 49 G). Prosternal process apically wavy (Fig. 50 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl 2005) wavy (Fig. 50 B). Scutellum rounded posteriorly (Fig. 50 B). Anterior phragma thin and somewhat straight (Fig. 50 C). Mesanepistenum with shallow imbricate microsculpture, inconspicuous in some. Procoxal rests triangular, straight posteriorly (Fig. 50 D). Mesoventral and median lines strongly wavy; area between median and mesocoxal lines large (Fig. 50 D). Mesoventral process curved, forming a wellmarked ridge (Fig. 50 E). METATHORAX. Metanotum with triangular alacrista, turned to the sides; scutoscutellar suture inconspicuous; median membranous area narrow and long (Fig. 50 F). Metaventrite smooth, but with imbricate microsculpture laterally; punctation sparse, fine. Mesocoxal line forming an angle between coxal cavities; finely punctate under coxal cavities (Fig. 50 D). Metanepisternum and metepimeron with imbricate microsculpture. Intercoxal plates smooth. Metendosternite with curved arms; ‘ stalk ridge’ exceeding half length of stalk (Fig. 51 A); ventral longitudinal flange wide and short in lateral view (Fig. 51 B). WINGS. Elytra slightly wider than long, partially covering tergite VI (Fig. 47 A); basal (Fig. 49 D) and sutural lines dashed and punctate; adsutural area with a row of setae, six rows of coarse punctures (not including sutural line) (Figs 47 A, 51 C); lateral lines punctate; apical coarse punctation dense; apical serrations moderately large, well visible (Fig. 51 D); pubescence short, fine. Epipleuron with diffuse coarse punctures. Hind wings fully developed (Fig. 51 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Femora somewhat straight (Fig. 51 F – H). Pro- and mesofemora sparsely and coarsely punctate; metafemora sparsely and finely punctate. Mesotibiae densely spinose, spines fine (Fig. 51 G). Metatibiae sparsely spinose, spines fine (Fig. 51 H). ABDOMEN. Tergites VI – VIII with imbricate microsculpture; tergite VII trapezoidal; punctation sparse, fine; pubescence dense, fine (Fig. 52 A). Ventrites 1 – 5 (Figs 52 B, S 5 F) sparsely and coarsely punctate; pubescence sparse, fine; imbricate microsculpture anteriorly, and with strigulate microsculpture posteriorly (Fig. 52 C). Metacoxal lines finely punctate. Males MEASUREMENTS (n = 1; in mm). Antennomeres (length (width): 0.15 (0.06), 0.13 (0.05), 0.14 (0.04), 0.09 (0.04), 0.11 (0.05), 0.08 (0.06), 0.07 (0.08), 0.07 (0.11), 0.08 (0.12), 0.08 (0.13), 0.18 (0.14); (n = 15, unless otherwise specified; in mm): TL 2.60 – 3.72 (mean = 3.16, standard deviation ± 0.34), PL 0.90 – 1.35 (1.12 ± 0.12), PA 0.75 – 1.00 (0.88 ± 0.07), PB 1.50 – 2.25 (1.90 ± 0.23), SL (n = 14) 0.13 – 0.21 (0.16 ± 0.02), SW (n = 14) 0.12 – 0.20 (0.16 ± 0.02), EI 1.50 – 2.15 (1.84 ± 0.19), EL 1.77 – 2.55 (2.18 ± 0.23), EW 0.85 – 1.42 (1.12 ± 0.15), EH 0.30 – 0.95 (0.74 ± 0.20), HW 0.68 – 0.89 (0.79 ± 0.06), IS 0.15 – 0.25 (0.21 ± 0.02), WA 0.15 – 0.20 (0.18 ± 0.01), MC 0.70 – 1.15 (0.91 ± 0.12), MB 0.33 – 0.56 (0.44 ± 0.07), VL 0.46 – 0.75 (0.62 ± 0.08). Pro- and mesotarsomeres I – III enlarged, with tenet setae (Fig. 51 F – G). Tergite VIII triangular, acuminate posteriorly; punctation dense, coarse; subglabrous (Fig. 52 A, D, Supp. file 3 (Figs 1 F, 3 G )). Tergite IX with rounded ventral struts (Fig. 52 E – F, Supp. file 3 (Figs 1 G, 4 A – B )). Sternite VIII rectangular, short (Fig. 52 G, Supp. file 3 (Figs 1 H, 4 C )). Sternite IX more or less spoon-like (Fig. 52 F, Supp. file 3 (Figs 1 G, 4 B )). Aedeagus sclerotized enlarged at base, apex of median lobe short; openings in dorsal view thin, forming a very acute angle (Fig. 53 A – C, Supp. file 3 (Figs 2 A – C, 4 D – F )); internal sac with irregular sclerites, crown-like (Fig. 53 D – E, Supp. file 3 (Fig. 4 G – H )); parameres short, curved and with denticular structures at base (Fig. 53 A – B, Supp. file 3 (Figs 2 A – B, 4 D – E )). Females MEASUREMENTS (n = 1; in mm). Antennomeres (length (width): 0.16 (0.06), 0.12 (0.06), 0.13 (0.04), 0.08 (0.04), 0.10 (0.05), 0.06 (0.06), 0.06 (0.08), 0.08 (0.10), 0.07 (0.12), 0.08 (0.14), 018 (0.15); (n = 14, unless otherwise specified; in mm): TL 2.56 – 3.64 (mean = 3.15, standard deviation ± 0.30), PL 0.92 – 1.37 (1.14 ± 0.11), PA 0.77 – 1.02 (0.90 ± 0.06), PB 1.50 – 2.20 (1.92 ± 0.18), SL (n = 12) 0.12 – 0.20 (0.16 ± 0.02), SW (n = 12) 0.14 – 0.19 (0.16 ± 0.01), EI (n = 12) 1.52 – 2.12 (1.87 ± 0.16), EL (n = 12) 1.85 – 2.52 (2.12 ± 0.19), EW 0.90 – 1.30 (1.14 ± 0.10), EH (n = 12) 0.35 – 0.97 (0.75 ± 0.23), HW 0.65 – 0.88 (0.79 ± 0.02), IS 0.20 – 0.26 (0.22 ± 0.02), WA 0.15 – 0.22 (0.18 ± 0.02), MC (n = 13) 0.72 – 1.06 (0.93 ± 0.09), MB 0.35 – 0.95 (0.49 ± 0.14), VL 0.52 – 0.77 (0.65 ± 0.07). Tergite VIII triangular, rounded posteriorly; punctation dense, coarse; subglabrous (Fig. 54 A). Sternite VIII rectangular with a sooth projection; with strigulate microsculpture (Fig. 54 B). Vagina and bursa copulatrix membranous without sclerites; vaginal plate with an apical sclerite, more or less short; oviduct bilobed, each lobe bearing a filiform spermatheca (Fig. 54 C – D, Supp. file 2 B). Distal gonocoxites curved and thick; gonostyli long, slender (Fig. 54 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	biology_ecology	Host fungi Adults were collected from Marasmiellus cubensis (Berk. & M. A. Curtis) Singer (4 records, 8 individuals), Marasmiellus spp. (5, 6), Marasmiellus aff. ramealis (1, 5), cf. Marasmiellus sp. (yellow) (2, 5), cf. Pleteus sp. (1, 1), Marasmius araucariae Singer (1, 2), Marasmius haematocephalus (1, 1), Leucocoprinus cepistipes (1, 2), Leucocoprinus ianthinus (1, 1), Leucoprinus sp. (1, 1), Inocybe sp. (Agaricales, Inocybaceae) (1, 3), cf. Valvariela sp. (1, 2), Leucoagaricus rubrotinctus (Peck) Singer (1, 1), Lepiota sp. (Agaricales, Agaricaceae) (1, 3), Macrolepiota colombiana Franco-Mol. (Agaricales, Agaricaceae) (1, 2), Heimiomyces neovelutipes (Hongo) E. Horak (Agaricales, Mycenaceae) (1, 1), Lulesia lignicola B. E. Lechner & J. E. Wright (Agaricales) (1, 7), Pleurotus pulmonarius (Fr.) Quél. (Agaricales, Pleurotaceae) (2, 28), Volvariella sp. (Agaricales, Pleurotaceae) (1, 2), Pluteus sp. (Agaricales, Pleurotaceae) (1, 1), Mycena sp. (Agaricales, Mycenaceae) (1, 1), Entoloma (Inocephalus) sp. (1, 1), Conocybe sp. (Agaricales, Bolbitiaceae) (1, 1), Coprinellus disseminatus (Pers.) J. E. Lange (Agaricales, Psathyrellaceae) (1, 4), unidentified mushrooms (1, 18) and Favolus tenuiculus P. Beauv. (Polyporaceae, Polyporales).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	discussion	Remarks Images of the type series were requested from the MNHN, where these specimens were supposedly deposited (Löbl 2018 a), but we were informed that they could not find any specimens. Nonetheless, we decided to treat the specimens that we examined as C. collare because their morphology and localities fit the original description and the known distribution of the species. Fourteen individuals of different body length and collected from different fungi at Viçosa, one from Areia Branca (SE) (Supp. file 3 (Figs 1 – 2 )), and another one from Sta Luzia do Itanhi (SE) (Supp. file 3 (Figs 3 – 4 )) were dissected. No differences were found in the terminalia, including the flagellum. Furthermore, no external differences were observed. The total body length is quite variable (2.60 – 3.72 mm). The three individuals from Areia Branca (SE) are the largest (3.64, 3.68 and 3.72 mm), while individuals collected at Viçosa (MG) and Sta Luzia do Itanhi (SE) are in the same size range (2.56 – 3.52 mm).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF458FF9E9855FC08FC11FE09.taxon	distribution	Distribution Known from Brazil [“ Matu Sinhos, Minas ”; “ Matto Grosso ”; “ Pery-Pery, Pernambuco ”; “ right bank of Parahyba ”]. New records from Areia Branca and Santa Luzia do Itanhi, state of Sergipe, Northeast, Brazil, and from “ Mata da Biologia ” and “ EPTEA Mata do Paraíso ”, campus of the Universidade Federal de Viçosa, Viçosa, Minas Gerais, Southeast Brazil (Fig. 55).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	description	Figs 5, 56 – 64	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	materials_examined	Material examined BRAZIL – Minas Gerais • 1 ♀; Viçosa, Mata da Biologia; 26 Oct. 2016; A. Orsetti and I. PecciMaddalena leg.; CELC • 7 ♂♂, 3 ♀♀ (2 ♂♂, entirely dissected, preserved in glycerin; 2 ♂♂, 1 ♀, abdomen dissected, preserved in glycerin); same collection data as for preceding; 26 Jun. 2019; von Groll and A. Orsetti leg.; CELC • 1 ♂, 2 ♀♀; Viçosa, EPTEA Mata do Paraíso; 9 Nov. 2016; I. PecciMaddalena and C. Lopes-Andrade leg.; “ \\ ex Psathyrella candolleana ”; CELC • 1 ♂; same collection data as for preceding; 12 Nov. 2019; LabCol leg; CELC.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	diagnosis	Diagnosis TL: 4.43 – 5.12 mm in males and 4.68 – 5.25 mm in females. Black (Figs 56 A – C, 64 A). Hypomeron with close strigulate microsculpture. Scutellum tapered posteriorly (Fig. 59 B). Mesanepisternum long with imbricate microsculpture. Metaventrite smooth (Fig. 56 B). Intercoxal plates with long imbricate microsculpture. Aedeagus poorly sclerotized, almost membranous, apex very long; parameres L-shaped, very long (Fig. 62 A – D). Distal gonocoxites curved and tapered (Fig. 63 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	description	Redescription COLORATION. Black, iridescent (Figs 56 A – C, 64 A). Frons dark brown; clypeus light brown (Fig. 57 A); mouth parts and antennomeres I – VI and apex of XI testaceous; antennomeres VII – X and base of XI dark (Fig. 57 B). Coxae and femora dark-brown reddish; tibiae lighter; tarsi and tergite VIII yellow; ventrites 5 and 6 brown. Variation: specimens entirely light brown to brown (Fig. 56 D – F). HEAD. Punctation dense, fine (Fig. 57 A); vertex with strigulate microsculpture. Eyes slightly wider than head, rounded apically (Fig. 57 A). Labrum rectangular, lateral margins rounded, and well distinct from apical margin; posterior margin slightly curved centrally; sclerotized portion curved; lateral setae slightly exceeding margins; porose centrally (Fig. 57 D). Mandibles curved, subapical serrations on left mandible conspicuous (Fig. 57 E – F). Maxillary palps elongated, palpomere III tapering smoothly; galea and lacinia densely pubescent, lacinia robust (Fig. 57 G). Mentum with lateral areas rounded and apex not well delimited; glossa concave (Fig. 58 A). Setae of labial palpomere II exceeding palpomere III; palpomere III longish, with long apical setae (Fig. 58 A). Hypopharynx with wide and rounded sclerotized plate (Fig. 58 A – B). Post gena microsculptured with very close transversal lines; densely porose all over; gula triangular with concave lateral areas (Fig. 58 C). Base of antennae long, antennal club distinct; antennomere XI hexagonal (Fig. 57 B – C), no difference between males and females. PROTHORAX. Pronotum smooth, densely and coarsely punctate; pubescence short, fine (Fig. 58 D – E); transverse and slightly curved laterally, forming an obtuse angle at lateral areas of posterior margin (Fig. 58 E). Hypomeron with close strigulate microsculpture. Notosternal suture straight and slightly turned to lateral areas (Fig. 58 F). Profurca elongated, extending half length of foramen (Fig. 58 G). Prosternal process short and curved (Fig. 59 A). MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines, Leschen & Löbl 2005) wavy (Fig. 59 B). Scutellum tapered posteriorly (Fig. 59 B). Anterior phragma large and straight (Fig. 59 C). Mesanepisternum with long imbricate microsculpture. Procoxal rests sub-squared, somewhat rounded posteriorly (Fig. 59 D). Mesoventral and median lines somewhat wavy; area between median and mesoventral lines somewhat enlarged (Fig. 59 D). Mesoventral process moderately short, with apex more prominent, forming a ridge (Fig. 59 E). METATHORAX. Metanotum with alacrista triangular, posterior portion longish; turned just slightly to posterior end; scutoscutellar suture long and wavy (Fig. 59 F). Metaventrite smooth; punctuation sparse, fine (Figs 56 B, E, 59 D). Mesocoxal line not forming an angle between coxal cavities, just a simple curve and finely punctate under coxal cavities (Fig. 59 D). Metanepisternum and metepimeron with imbricate microsculpture. Intercoxal plates with long imbricate microsculpture. Metendosternite with arms close and somewhat straight; ‘ stalk ridge’ not exceeding half length of stalk (Fig. 60 A); ventral longitudinal flange longish and rounded in lateral view (Fig. 60 B). WINGS. Elytra large, slightly wider than longer; covering tergite VI (Fig. 56 A); basal (Fig. 58 D) and sutural lines dashed; adsutural area with a row of setae; six rows of coarse punctures (not including sutural line) (Figs 56 A, 60 C); lateral line coarse punctate; apical coarse punctation dense; apical serrations small (Fig. 60 D); pubescence short and fine. Epipleuron with a row of sparse and coarse punctures. Hind wings fully developed (Fig. 60 E). LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Profemora fusiform; punctuation sparse, fine (Fig. 60 F). Mesofemora longish; punctures sparse, coarse (Fig. 60 G). Metafemora longish; punctuation sparse, fine (Fig. 60 H). Mesotibiae densely spinose; spines thick (Fig. 60 G); metatibia sparsely spinose, spines fine (Fig. 60 H). ABDOMEN. Tergites VI – VIII with narrow imbricate microsculpture (Fig. 61 A). Tergite VII trapezoidal when tergite VIII not exposed and triangular when exposed; punctation inconspicuous; pubescence sparse, fine. Ventrites 1 – 5 with strigulate microsculpture (Fig. 61 B). Ventrite 1 densely and coarsely punctate; pubescence sparse, fine (Fig. 61 C). Ventrites 2 – 5 densely and finely punctate; pubescence moderately sparse, fine. Metacoxal lines finely punctate. Males MEASUREMENTS (n = 1, in mm). Antennomeres (length (width )): 0.33 (0.10), 0.16 (0.08), 0.21 (0.08), 0.18 (0.08), 0.19 (0.09), 0.13 (0.11), 0.15 (0.17), 0.11 (0.18), 0.12 (0.21), 0.12 (0.23), 0.23 (0.25); (n = 8, unless otherwise specified; in mm): TL 4.43 – 5.12 (mean = 4.91, standard deviation ± 0.21), PL 1.68 – 2.00 (1.88 ± 0.09), PA 1.2 – 1.4 (1.34 ± 0.06), PB 2.72 – 3.16 (3.04 ± 0.14), SL 0.20 – 0.27 (0.24 ± 0.02), SW 0.21 – 0.28 (0.23 ± 0.02), EI 2.56 – 3.00 (2.85 ± 0.14), EL 2.96 – 3.56 (3.39 ± 0.19), EW 1.72 – 2.00 (1.90 ± 0.09), EH 1.00 – 1.25 (1.16 ± 0.08), HW 1.01 – 1.17 (1.13 ± 0.05), IS 0.30 – 0.37 (0.32 ± 0.02), WA 0.22 – 0.30 (0.26 ± 0.03), MC (n = 5) 1.36 – 1.44 (1.41 ± 0.03), MB (n = 6) 0.64 – 0.72 (0.68 ± 0.03), VL (n = 6) 0.80 – 0.96 (0.84 ± 0.06). Pro- and mesotarsomeres I – III enlarged, with tenet setae (Fig. 60 F – G). Tergite VIII pentagonal, slightly acuminate posteriorly; punctation sparse, fine; subglabrous (Fig. 61 D). Tergite IX with rectangular ventral struts (Fig. 61 E – F). Sternite VIII sub-rectangular, with a smooth projection (Fig. 61 G). Sternite IX thick, centrally constricted (Fig. 61 F). Aedeagus poorly sclerotized, almost membranous, apex of median lobe very long (Fig. 62 A – C); internal sac with irregular sclerites, with a large hook (Fig. 62 D – E); parameres L-shaped, very long (Fig. 62 A). Females MEASUREMENTS (n = 1, in mm). Antennomeres (length (width )): 0.28 (0.10), 0.15 (0.08), 0.20 (0.07), 0.17 (0.07), 0.17 (0.08), 0.12 (0.10), 0.14 (0.16), 0.13 (0.18), 0.11 (0.20), 0.11 (0.23), 0.22 (0.23); (n = 6, unless otherwise specified; in mm): TL 4.68 – 5.25 (mean = 4.91, standard deviation ± 0.19), PL 1.76 – 2.04 (1.86 ± 0.09), PA 1.28 – 1.40 (1.34 ± 0.04), PB 2.84 – 3.28 (3.04 ± 0.15), SL 0.24 – 0.27 (0.25 ± 0.01), SW 0.21 – 0.30 (0.25 ± 0.03), EI 2.64 – 3.04 (2.82 ± 0.13), EL 3.16 – 3.60 (3.34 ± 0.15), EW 1.72 – 2.00 (1.90 ± 0.10), EH 1.15 – 1.40 (1.23 ± 0.08), HW 1.06 – 1.15 (1.11 ± 0.03), IS 0.30 – 0.35 (0.32 ± 0.01), WA 0.25 – 0.30 (0.26 ± 0.02), MC (n = 5) 1.32 – 1.52 (1.40 ± 0.07), MB (n = 5) 0.64 – 0.80 (0.74 ± 0.06), VL (n = 5) 0.88 – 1.04 (0.94 ± 0.06). Tergite VIII triangular; punctation dense, fine; pubescence dense (Fig. 63 A). Sternite VIII sub-rectangular with a triangular projection (Fig. 63 B). Vagina and bursa copulatrix membranous, without sclerites (Fig. 63 C). Vaginal plate with an apical rectangular sclerite (Fig. 63 D) Spermatheca not detected. Distal gonocoxites curved and enlarged at base; gonostyli short, enlarged at base (Fig. 63 C, E).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	biology_ecology	Host fungi Adults were collected from Psathyrella candolleana (1 record, 3 individuals), unidentified mushroom (“ Lactarius? ”) (1, 10).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	discussion	Remarks The dorsal photo of one specimen belonging to the type series (Fig. 64 A) and the original description of C. oberthueri are consistent with the data of the specimens studied here. Furthermore, the itinerary of Philibert Germain (P. Germain) (Fig. 64 C) was evaluated: in 1885 he was in Minas Gerais and in 1886 at “ Province Matto Grosso ” where the type series was collected (Fig. 64 C) (Papavero 1971). The “ Province Matto Grosso ” at that time was a broad province that comprised what nowadays are the states of Mato Grosso, Mato Grosso do Sul and Rondônia. In 1887 he arrived in Cárceres (currently in the state of Mato Grosso) departing from Corumbá (currently Mato Grosso do Sul), and in 1889 he was in Cochabamba (Bolivia) (Fig. 64 C) (Papavero 1971). It is likely that “ Province Matto Grosso ” refers to somewhere between what nowadays is the northern part of Mato Grosso do Sul, and the southern part of Mato Grosso. We do not know whether the type series was collected in the Cerrado biome or in riparian forest similar, or even linked, to areas of the Atlantic Forest biome. The specimens we examined are all from two Atlantic Forest remnants of Southeast Brazil, far away from the type locality, but they fit the original description and are morphologically similar to the available image of a syntype (Fig. 64 A). At this moment, we prefer to call the specimens we have in hand as C. oberthueri, in the absence of any evidence to describe them as a new species.	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
A5415E5FF454FF88987FFE18FD7FFDB2.taxon	distribution	Distribution Known from “ Prov. Matto Grosso ”, Midwest, Brazil (1886, P. Germain). New records from Viçosa, Minas Gerais, Southeast Brazil (Fig. 64 C).	en	Groll, Elisa Von, Lopes-Andrade, Cristiano (2022): Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species. European Journal of Taxonomy 835 (1): 1-97, DOI: https://doi.org/10.5852/ejt.2022.835.1909, URL: http://dx.doi.org/10.5852/ejt.2022.835.1909
