occurrenceID	taxonID	catalogNumber	collectionCode	institutionCode	typeStatus	verbatimLabel	sex	individualCount	eventDate	recordedBy	recordNumber	decimalLatitude	decimalLongitude	minimumElevationInMeters	maximumElevationInMeters	minimumDepthInMeters	maximumDepthInMeters	country	stateProvince	municipality	locality	references	basisOfRecord
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3F30FF687653FF1A	03D6B32E820CD4203ECAFCEC7764FA22.taxon					Mexico: Jalisco, Michoacán, Puebla, Veracruz		1										Mexico	Jalisco		Jalisco	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3FA2FE6C7744FDDE	03D6B32E820CD4203ECAFCEC7764FA22.taxon				holotype	HOLOTYPE ♂ (by original designation). MEXICO: ESTADO DE MÉXICO: [Mpio. Tejupilco], Tejupilco, Temescaltepec [District of Temascaltepec], 19 - VI- 1933 / H. E. Hinton & R. L. Usinger collectors / Ambrysus cosmius La Rivers	male	1	1933-06-19	H. E. Hinton & R. L. Usinger								Mexico	Mexico	Tejupilco	Temescaltepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3C35FD9477DAFD82	03D6B32E820CD4203ECAFCEC7764FA22.taxon				holotype	HOLOTYPE / Ambrysus cosmius La Rivers determined by Ira La Rivers ' 50 / California Academy of Sciences Type No. 10056		1										Mexico	Mexico		California Academy of Sciences Type No.	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3D55FDF0700FFD82	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS	paratype	PARATYPES: same data as holotype (2 ♀ CAS)	female	2										Mexico	Mexico		California Academy of Sciences Type No.	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3CDAFDDC763CFD66	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS	allotype	ALLOTYPE		1										Mexico	Guerrero			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3DFBFDDD7688FD4A	03D6B32E820CD4203ECAFCEC7764FA22.taxon				allotype	GUERRERO: [Mpio. Chilapa de Álvarez], Chilapa, 29 - X- 1930, Prof. L Schultze / Ambrysus cosmius La Rivers		1	1930-10-29	Prof. L Schultze								Mexico	Guerrero		Chilapa	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3DEBFD387154FD2E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC, USNM	paratype	Paratype (1 ♀ EMEC; 2 ♂ USNM); [Mpio. Tecpan de Galeana], Tecpan, 02 - IV- 1930, Prof. L Schultze / Ambrysus cosmius La Rivers	female	3	1930-04-02	Prof. L Schultze								Mexico	Guerrero		Tecpan	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3A2BFD0470BCFD2E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			SEMC	paratype	Paratype (2 ♂, 3 ♀ SEMC)	male	5										Mexico	Guerrero			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3ECAFD6070BDFD12	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS, CNIN	paratype	same but / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♂ CAS; 1 ♀ CNIN)	male	2										Mexico	Guerrero	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3ECAFD4D76FFFCF6	03D6B32E820CD4203ECAFCEC7764FA22.taxon				paratype	MORELOS: 14 - VII- 1936, H. D. Thomas / Ambrysus cosmius La Rivers		1	1936-07-14	H. D. Thomas								Mexico	Morelos		MORELOS	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3DB8FD4C7031FCF6	03D6B32E820CD4203ECAFCEC7764FA22.taxon			SEMC	paratype	Paratype (1 ♂, 1 ♀ SEMC)	male	2										Mexico	Morelos			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3B7EFD4C71ECFCDA	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS	paratype	same but / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♀ CAS)	female	1										Mexico	Morelos	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3A88FCA87016FCBE	03D6B32E820CD4203ECAFCEC7764FA22.taxon				paratype	[Mpio. Temixco], Acatlipa, km 88 on Hwy Mexico City-Acapulco / 06 - V- 1945, J. G. Shaw / Ambrysus cosmius La Rivers		1	1945-05-06	J. G. Shaw								Mexico	Morelos	Hwy	City-Acapulco	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3B67FCF47403FC62	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC	paratype	Paratype (1 ♀ EMEC)	female	1										Mexico	Morelos			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3F69FCD0769FFC46	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS	paratype	same but / Ambrysus cosmius La Rivers determined by Ira La Rivers ' 50 / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♀ CAS)	female	1										Mexico	Morelos	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3D97FC3D776FFC2A	03D6B32E820CD4203ECAFCEC7764FA22.taxon				paratype	OAXACA: Oaxaca, 25 - VIII- 1937, H. D. Thomas / Ambrysus cosmius La Rivers		1	1937-08-25	H. D. Thomas								Mexico	Oaxaca		Oaxaca	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3C0FFC18766CFC2A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			SEMC	paratype	Paratype (1 ♀ SEMC)	female	1										Mexico	Oaxaca			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3E9AFC65762DFBF2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		Additional material examined. MEXICO: ESTADO DE MÉXICO: Mpio. Amatepec, Río San Felipe at Las Adjuntas, 20 - IV- 2004, R. Novelo coll. (1 ♂, 3 ♀ IEXA)	male	4	2004-04-20	R. Novelo								Mexico	Mexico	Rio San Felipe	Las Adjuntas	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3D4BFC407435FBD6	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		same but 05 - V- 2008, R. Novelo & K. Cuevas colls. (2 ♂, 5 ♀ IEXA)	male	7	2008-05-05	Novelo & Cuevas								Mexico	Mexico			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3F41FBAC773FFBBA	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		Mpio. Amatepec, Río San Felipe at San Felipe de Jesús, [approx. 18.799 2, - 100.1639], 1,000 m, 20 - IV- 2004, R. Novelo coll. (1 ♀ IEXA)	female	1	2004-04-20	R. Novelo				1000				Mexico	Mexico	Rio San Felipe	San Felipe de Jesus	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3C75FB887401FB9E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		[Mpio. Amatepec], Rio San Felipe at San Felipe de Jesús, 20 - IV- 2004, 3,280 ' (1 ♂ EMEC)	male	1	2004-04-20									Mexico	Mexico	Rio San Felipe	San Felipe de Jesus	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3F72FBD47669FB9E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		same but / William D. Shepard, leg. (2 ♂ EMEC)	male	2		William D. Shepard								Mexico	Mexico			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3D0AFBD471E3FB42	03D6B32E820CD4203ECAFCEC7764FA22.taxon					Mpio. Santo Tomás, [Río Tilastoc at] Santa Bárbara, 19 º 10 ' 29.4 '' N, 100 º 17 ' 20.4 '' W, 1,051 m, 20 - XII- 2012, L- 1463, Reynoso-Velasco coll. (1 ♂ UMC)	male	1	2012-12-20	Reynoso-Velasco		19.174833	-100.289	1051				Mexico	Mexico	Mpio. Santo Tomas	Santa Barbara	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3A8CFB3075D6FB0A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		[Mpio. Tejupilco], Tejupilco, Temescaltepec [District of Temascaltepec], 15 - VI- 1933 / H. E. Hinton & R. L. Usinger collectors (1 ♂, 3 ♀ EMEC)	male	4	1933-06-15	H. E. Hinton & R. L. Usinger								Mexico	Mexico	Tejupilco	Temescaltepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3EA3FB7870BCFB0A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS		same but Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♂ CAS)	male	1										Mexico	Mexico	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3ECAFB4474B1FAD2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		[Mpio. Tejupilco], Tejupilco, Temescaltepec [District of Temascaltepec], 19 - VI- 1933 / H. E. Hinton & R. L. Usinger collectors (2 ♂ EMEC)	male	2	1933-06-19	H. E. Hinton & R. L. Usinger								Mexico	Mexico	Tejupilco	Temescaltepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3FC3FAA0768EFAD2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		same but / Ambrysus signoreti Stål (1 ♀ EMEC)	female	1										Mexico	Mexico			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3DEEFAA0719DFAB6	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		[Mpio. Tejupilco], Tejupilco, Temescaltepec [District of Temascaltepec], 21 - VI- 1933 / H. E. Hinton & R. L. Usinger collectors (1 ♀ EMEC)	female	1	1933-06-21	H. E. Hinton & R. L. Usinger								Mexico	Mexico	Tejupilco	Temescaltepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3A99FA8C719AFA9A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			USNM		same but / exch. @ RLU 1968 / Ambrysus signoreti Stål det. R. L. Usinger / C. J. Drake Accession (1 ♀ USNM)	female	1		Drake Accession								Mexico	Mexico			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3A91FAE87403FA22	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		[Mpio. Tejupilco], Tejupilco, Temescaltepec [District of Temascaltepec], 13 - VII- 1932 [1933] / H. E. Hinton & R. L. Usinger collectors (1 ♀ EMEC)	female	1	1932-07-13	H. E. Hinton & R. L. Usinger								Mexico	Mexico	Tejupilco	Temescaltepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3F69FA1075FEFA06	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS		same but / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♀ CAS)	female	1										Mexico	Mexico	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3EB5FA7C75D6F9EA	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC		[Mpio. Tejupilco], Tejupilco, Temescaltepec [District of Temascaltepec], VII- 1934 / H. E. Hinton (2 ♂, 2 ♀ EMEC)	male	4	1934-07	H. E. Hinton								Mexico	Mexico	Tejupilco	Temescaltepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3EA1FA58766BF9CE	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS		Temascaltepec, 11 - VII- 1933, Robert L. Usinger, Cal. Acad. Sci. Coll. / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♂, 1 ♀ CAS)	male	2	1933-07-11	Robert L. Usinger & Cal. Acad. Sci. Coll.								Mexico	Mexico	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3D01F9A47467F9B2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS		same but no date, # 880, R. L. Usinger, H. E. Hinton (1 ♂, 3 ♀ CAS)	male	4		R. L. Usinger & Hinton								Mexico	Mexico			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3F12F98175C3F97A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		GUERRERO: Mpio. Teloloapan, [Arroyo El Naranjo] at km 20 on carr. Teloloapan-Apaxtla de Castrejón, nr. Tepozanalquillo, 18 º 14 ' 26.8 '' N, 99 º 54 ' 51 '' W, 1083 m, 29 - XII- 2012, L- 1497, Reynoso-Velasco coll. (1 ♂ IEXA)	male	1	2012-12-29	Reynoso-Velasco		18.240778	-99.91417	1083				Mexico	Guerrero	Teloloapan-Apaxtla de Castrejon	Tepozanalquillo	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3EACF9C8776BF95E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		Mpio. Teloloapan, Río Oxtotitlán at Oxtotitlán, 18 º 11 ' 35.7 '' N, 99 º 56 ' 24.3 '' W, 1009 m, 29 - XII- 2012, L- 1498, Reynoso-Velasco coll. (2 ♂, 1 ♀ IEXA)	male	3	2012-12-29	Reynoso-Velasco		18.193249	-99.94009	1009				Mexico	Guerrero	Rio Oxtotitlan	Oxtotitlan	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3C06F914779FF902	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC		Mpio. Tepecoacuilco, [Río Atoyac at San Agustín] Oapan, 17 º 57 ' 10.66 '' N, 99 º 27 ' 38.8 '' W, 496 m, J. Amith coll. (1 ♀ UMC)	female	1		J. Amith		17.952961	-99.46078	496				Mexico	Guerrero	Mpio. Tepecoacuilco	Oapan	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3C9AF9717650F8E6	03D6B32E820CD4203ECAFCEC7764FA22.taxon			USNM		JALISCO: [Mpio Tuxpan], nr Atentique [Atenquique], CL 740, 12 - VI- 1975, J. T. Polhemus / J. C. Drake Accession (1 ♀ USNM)	female	1	1975-06-12	J. T. Polhemus & J. C. Drake Accession								Mexico	Jalisco		Atentique	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3DDDF95B7155F8CA	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC		MICHOACÁN: [Mpio Aguililla, Río Aguililla at] Aguililla, [approx. 18.728 5, - 102.7934], 03 - IV- 2005, R. Novelo coll. (1 ♂, 1 ♀ UMC)	male	2	2005-04-03	R. Novelo								Mexico	Michoacan de Ocampo		Aguililla	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3A21F8B8748EF8AE	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		same but 02 - X- 2005, J. A. Gómez coll. (2 ♀ IEXA)	female	2	2005-10-02	J. A. Gomez								Mexico	Michoacan de Ocampo			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3FE8F8847729F892	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC		[Mpio. Chinicuila, Río La Tortuga at] Villa Victoria, [approx. 18.751 0, - 103.3600], 17 - V- 2002, R. Novelo coll. (1 ♀ UMC)	female	1	2002-05-17	R. Novelo								Haiti	Michoacan de Ocampo		Villa Victoria	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD42F3C45F8E077E4F876	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA		Mpio. Coalcomán, La Chichihua, [approx. 18 º 44 ' 36 '' N, 103 º 13 ' 1.39 '' W], 10 - VII- 2005, R. Novelo & J. A. Gómez colls. (1 ♀ IEXA)	female	1	2005-07-10	R. Novelo & J. A. Gomez		18.743334	-103.21706					Mexico	Michoacan de Ocampo	Mpio. Coalcoman	La Chichihua	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B170865820FD4213C97F8CC75EFFF1A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS		[Mpio. Tuxpan], 5 mi N of Tuxpan, 16 - VII- 1953, 5,500 ft / Univ. Kans. Mex. Expedition / Slesnick Field N 0. 48 / Ambrysus cosmius La Rivers 1953 (Ira La Rivers) / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (1 ♀ CAS); Mpio. Tuzantla, Río Tuzantla at Tuzantla, 19 º 13 ' 18.1 '' N, 100 º 34 ' 05.5 '' W, 593 m, 21 - XII- 2012, L- 1470, Reynoso-Velasco coll. (1 ♂ IEXA); [Mpio. Tzitzio], near Chinapa, 05 - IX- 1938 / Ambrysus cosmius Paratype (2 ♀ SEMC) [unpublished paratypes]. MORELOS: Mpio. Amacuzac, Río Amacuzac 1.6 km S (on dirt road) of km 50 carr. MEX 95 Cuernavaca-Taxco, 18 º 35 ' 48.05 '' N, 99 º 23 ' 57.88 '' W, 917 m, 04 - I- 2017, L- 1990, Reynoso-Velasco coll. (2 ♀ IEXA); Mpio. Jojutla, Río Amacuzac at Tehuiztla, 18 º 33 ' 45.2 '' N, 99 º 16 ' 51.3 '' W, 853 m, 28 - IV- 2011, L- 1308 Reynoso-Velasco coll. (4 ♂, 1 ♀ UMC); same but 18 º 32 ' 47.9 '' N, 99 º 16 ' 03.6 '' W, 854 m, L- 1309 (1 ♂, 3 ♀ UMC); Mpio. Xochitepec, [Río Cuentepec at] Balneario Palo Bolero, [approx. 18.766 8, - 99.2393], 07 - VII- 1987, R. Novelo coll. (1 ♂, 2 ♀ IEXA); Mpio. Xochitepec, Río Sabinos (Cuentepec) at Jardines de Xochitepec, [approx. 18.778 9, - 99.2439], 24 - IX- 1998, R. Novelo coll. (1 ♂, 2 ♀ IEXA); same but 05 - IV- 2002, (3 ♀ IEXA); same but 24 - IV- 2003 (1 ♂, 2 ♀ IEXA). OAXACA: Mpio. Putla Villa de Guerrero, Putla, [Río Cuchara] at Pte. La Cuchara, 17 º 01 ' 07 '' N, 97 º 55 ' 11.8 '' W, 711 m, 30 - V- 2012, L- 1396, ReynosoVelasco coll. (2 ♂, 1 ♀ UMC); same but [Río Cuchara at] 17 º 02 ' 21.5 '' N, 97 º 54 ' 42.9 '' W, 767 m, L- 1397 (1 ♂ UMC); Mpio. San Antonio Nanahuatípam, Río Salado at San Antonio Nanahuatípam, 18 ° 07 ' 59.4 '' N, 97 ° 07 ' 42.6 '' W, 790 m, 24 - III- 2015, L- 1877, Reynoso-Velasco, Sites, Shepard, & Barr colls. (4 ♂, 10 ♀ IEXA); [Mpio. San Antonio Nanahuatípam], Río Saldo at [San Gabriel] Casa Blanca, [approx. 18.155 1, - 97.1371], 01 - III- 1976, R. R. & F. H. Miller, M 76 - 41 (1 ♀ UMMZ); Mpio. San Felipe Jalapa de Díaz, Río Sto. Domingo at Sto. Domingo, 18 ° 02 ' 15.9 '' N, 96 ° 32 ' 57.2 '' W, 100 m, 25 - IV- 2015, L- 1881, Reynoso-Velasco, Sites, Shepard & Barr colls. (5 ♂, 4 ♀ IEXA); [Mpio. San Juan Bautista Cuicatlán], Río Las Huertas [Las Vueltas] at Santiago Dominguillo, 12 - VII- 1996, R. Barba & A. Rojas colls. / Colección del Instituto de Biología, UNAM. México D. F. (2 ♂, 3 ♀ UMC); Mpio. San Juan Bautista Cuicatlán, [Río Las Vueltas at] Santiago Dominguillo, 17 º 38 ' 46 '' N, 96 º 54 ' 36.7 '' W, 726 m, 01 - VI- 2012, L- 1405, Reynoso-Velasco coll. (1 ♂, 2 ♀ CNIN; 4 ♂, 9 ♀ IEXA; 18 ♂, 16 ♀ UMC); [Mpio. San Pedro Totolápam], Rio Grande [at] Totolapan [Totolápam], CL 1063, 29 - IV- 1964, J. T. & M. S. Polhemus / Ambrysus cosmius ' 67 La Riv. Det. J. T. Polhemus (1 ♂ EMEC); same but / C. J. Drake Accession (4 ♂, 3 ♀ USNM); [Mpio. Santa María Tecomavaca, Arroyo Los Cués at] San Juan de Los Cués, 14 - VI- 1984, Mario García coll. / Colección del Instituto de Biología, UNAM. México D. F. / C. J. Drake Accession (1 ♀ USNM); same but / J. T. Polhemus Collection 2014 (1 ♀ USNM); [Mpio. Santa María Tecomavaca], Río Salado [at Pte. El Saldo], 16 km S of San Juan de Los Cués on carr. MEX 135 Tehuacán-San Francisco Telixtlahuaca, 17 º 55 ' 50.5 '' N, 97 º 00 ' 04.2 '' W, 544 m, 01 - VI- 2014, L- 1407, Reynoso-Velasco coll. (2 ♂, 2 ♀ CNIN; 7 ♂, 15 ♀ IEXA; 2 ♀ UMC); Mpio. Valerio Trujano, Río Grande at Pte. Río Grande, km 124 carr. MEX 135 Tehuacán-San Francisco Telixtlahuaca on jct. to Tomellín, 17 º 45 ' 25 '' N, 96 º 56 ' 59.1 '' W, 607 m, 01 - VI- 2012, L- 1406, Reynoso-Velasco coll. (3 ♂, 2 ♀ CNIN; 21 ♂, 13 ♀ IEXA; 2 ♂, 2 ♀ NHMW; 12 ♂, 13 ♀ UMC); San Cristobal, 17 - VI- 84, H. Velazco coll. / Colección del Instituto de Biología, UNAM. México D. F. (1 ♀ UMC); 18 mi NW El Camarón, 20 - VIII- 1959, A. E. Menke coll. / Ambrysus cosmius La Rivers 1953 (Ira La Rivers) / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (2 ♂, 2 ♀ CAS; 1 ♀ CNIN); 25 mi S of Mitla, 04 - I- 1956, J. C. Schaffner / Ambrysus cosmius ' 67 LaR det. J. T. Polhemus / C. J. Drake Accession (1 ♀ USNM). PUEBLA: Mpio. Coxcatlán, [Río Tepazalco at] La Estación, 5 km SW of Coxcatlán, 18 ° 15 ' 07.3 '' N, 97 ° 11 ' 24.5 '' W, 950 m, 24 - III- 2015, L- 1876, Reynoso-Velasco, Sites, Shepard, & Barr colls. (2 ♀ IEXA). VERACRUZ: [Mpio. Fortín], Río Metalac [Metlac] 3 mi N of El Fortín [Fortín de Las Flores, approx. 18.931 2, - 97.0211], 2 - VII- 1955, R. B. & J. M. Selander (1 ♂, 2 ♀ BYU). Ambrysus morelosensis Reynoso & Sites NEW SPECIES (Figs. 5 – 7) Description. Macropterous female. HOLOTYPE, length 12.87; maximum width 8.12. Paratypes (n = 1), length 11.75; maximum width 7.63. General shape elliptical; widest across embolia. Overall dorsal coloration of hemelytra dark-brown with light-brown marks; head light-brown; pronotum brown with dark-brown marks; legs brown (Fig. 5 A). Dorsal surface coarsely punctate. Ventral coloration of head and thorax light-brown, abdomen with yellowishbrown pubescence. Head. Head length 1.73; maximum width 3.41. Mostly light-brown with dark-brown marks along midline and posteriorly; coarsely punctate. Eyes convergent anteriorly, synthlipsis 1.73; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes convex, extending anteriorly in front of eyes 10.71 % of head length; posterior margin between eyes strongly convex, extending posteriorly 28.57 % of head length. Labrum width 2.00 × length, evenly rounded. Labium with three visible light-brown segments, darkening distally, extending 0.76 beyond labrum not including extruded stylets. Antennal proportions 5: 14: 19: 9, length 0.94, not reaching lateral margin of eye, elongate hairs on segment 4 and distal half of 3. Thorax. Pronotum coarsely punctate, ground color brown, dark-brown marking medially, dark-brown line behind eye; transverse sulcus marking anterior border of transverse band in posterior 1 / 3; transverse band brown; lateral margins light-brown, convergent, evenly convex, explanate; posterior margin almost straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.86 × length; length at midline 2.35; maximum width at posterolateral corners 6.75. Prothorax ventrally pruinose throughout, except laterally; apices of propleura meeting at midline, not appressed to prosternellum; propleuron light-brown, medial 2 / 3 of posterior margin with golden setae. Probasisternum with sharp median carina and row of setae lateral to carina. Prosternellum not covered by apices of propleura. Scutellum coarsely punctate, triangular, brown, two reniform to ovate dark-brown marks medially, width 2.02 × length, width 4.52, length 2.23. Hemelytra densely punctate, mostly dark-brown, length 9.20 (chord measurement); corium dark-brown, lightbrown irregularly shaped and checkmark-shaped markings at lateral and posterior margins, respectively. Clavus dark-brown, light-brown at mid-length and apically, length 3.80 × width, length 4.71, width 1.24; claval commissure black, length 1.24. Embolium length 3.53, greatest width 1.30; lateral margin convex, yellowish-brown in anterior 2 / 3, dark-brown posteriorly. Membrane dark-brown (Fig. 5 A). Hindwings extending to posterior margin of tergum VI. Mesobasisternum midventral tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina. Legs. All legs segments brown. Profemur posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with occlusal inner surface flattened; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibiae with ventrolateral and dorsolateral rows of stout reddishbrown spines; mesolateral rows intermixed with combs of spines. Metatibia with ventrolateral and dorsolateral rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apex of dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin. Meso- and metatibiae and metatarsus with long, yellow to brown swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 3.10, tibia 2.41, tarsus 0.55; middle leg, femur 2.97, tibia 2.79, tarsomeres 1 – 3, 0.18, 0.54, 0.66; hind leg, femur 3.90, tibia 4.65, tarsomeres 1 – 3, 0.31, 1.00, 0.90. Abdomen. Dorsally with lateral margins of III – VIII exposed, IV – VI each dark-brown in anterior 1 / 3 and yellowish-brown posteriorly; lateral margins of II to basal half of III smooth, distal half of III – VI finely serrate, marginal row of profuse and short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corner of II (visible ventrally) right angled, III – VI acute. Ventrally yellowish-brown to brown, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches clearly visible around spiracles on laterosternites II – VI. Mediosternite VII (subgenital plate) asymmetrical, width 1.36 × length; length at midline 1.26; maximum width 1.72; left margin with broadly rounded lobe occupying its entire length; right margin with small apical lobe; posterolateral corners short, pointed, subtriangular, extending posteriorly further than the central lobe; central lobe poorly developed, inconspicuous, shallowly concave medially (Fig. 5 B). Macropterous male. Paratypes (n = 1), length 11.88; maximum width 7.75. Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for holotype. Otherwise hindwings extending to posterior margin of tergum V. Accessory genitalic process of tergum VI gently curved to right close to base, distal 1 / 3 angled to right to continue perpendicular to body axis (Fig. 5 C). Medial lobes of tergum VIII (pseudoparameres) symmetrical, posterolateral corners narrowly rounded, posteromesal corners broadly rounded, posterior margin concave (Fig. 5 D). Phallosoma elongate, linear, slender, broadly rounded dorsoapically; left endosomal sclerite reduced, with few small and inconspicuous denticles; right endosomal sclerite vestigial, rounded, without denticles (Fig. 5 E). Parameres symmetrical, slightly longer than maximum width, mesal margin rounded, setae emanating from distal half of dorsal surface. Proctiger almost as long as width at base. Pygophore with anterior margin obtuse to subobtuse medially, elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 5 F). FIGURE 5. Structures of Ambrysus morelosensis Reynoso & Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6 th and 7 th abdominal terga of male, (D) 8 th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite. Diagnosis. This species can be distinguished from the other three in this section by the female SGP with a large, plate-like, rounded lobe on the left margin and symmetrical subtriangular posterolateral corners (Fig. 5 B). Additionally, the male AGP tapers distally to a pointed and laterally directed apex (Fig. 5 C) and the pseudoparamere posterolateral corners are narrowly rounded (Fig. 5 D). Comparative notes. Ambrysus morelosensis presents a unique female SGP; none of the other three species in this group exhibits a similar large lobe on the left margin. The only other two species in the complex with a " comparable " feature are A. scalenus La Rivers and A. sitesi Reynoso; nonetheless, the lobe is truncate in A. scalenus and in both species it is located at mid-length, not occupying the entire margin as in the new species. Additionally, these two species present the central lobe almost at the same level as the posterolateral corners, whereas in A. morelosensis n. sp. the central lobe is poorly expressed, almost flat, and clearly below the level of the posterolateral corners. This new species is superficially similar to A. sitesi; this similarity together with the " comparable " female SGP and the similar male pseudoparameres suggest a close relationship. Variation. The single female paratype exhibits the posterior margin of the SGP evenly concave between the blunt posterolateral corners and the small apical lobe on the right margin is absent. Habitat description. The type locality, Río Amacuzac at Huajintlán, is in the southwestern area of the Mexican state of Morelos at an elevation of 931 meters. The first author visited the type locality (L- 1989) three decades after the type specimens were collected; unfortunately, no more specimens of A. morelosensis were found, only numerous specimens of A. hydor La Rivers. At the time, the river at the type locality was approximately 40 meters wide and shallow, exposing boulders and cobble (Fig. 6). Distribution. Ambrysus morelosensis is known from only two localities, one in the state of Michoacán and the other in Morelos, in central-western and central Mexico, respectively (Fig. 7). Both localities are at elevations below 1,000 meters and within the limits of the Balsas Basin biogeographical province; more specifically, in the middle (Morelos) and lower (Michoacán) sections of the basin. Etymology. The specific epithet " morelosensis " is in reference to the central Mexican state of Morelos, where the type locality is located. The species name means The Ambrysus from Morelos. Repository. The holotype and paratypes will be deposited in the Colección Entomológica del Instituto de Ecología A. C. (IEXA). FIGURE 6. Type locality of Ambrysus morelosensis Reynoso & Sites n. sp. at Río Amacuzac, in the Mexican central state of Morelos Type material examined. HOLOTYPE ♀. MEXICO: MORELOS: [Mpio. Amacuzac, Río Amacuzac at Huajintlán], km 50 [on carr. 95] Cuernavaca-Taxco, [18 º 36 ' 35.56 '' N, 99 º 25 ' 40.38 '' W, 931 m], 09 - III- 1985, M. Albores coll. PARATYPES: same data as holotype (1 ♂ IEXA). MICHOACÁN: Mpio. Buenavista, Buenavista River at Buenavista Tomatlán, below dam, 19 º 12 ' 57.8 '' N, 102 º 34 ' 30.2 '' W, 440 m, 24 - XII- 2012, L- 1481, ReynosoVelasco, D. coll. (1 ♀ IEXA).	female	1	1953-07-16	Univ. Kans. Mex. Expedition & Slesnick Field				1676				Mexico	Michoacan de Ocampo	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203B12FF2077DDFF37	03D6B32E820CD4203ECAFCEC7764FA22.taxon					[Mpio. Tzitzio], near Chinapa, 05 - IX- 1938 / Ambrysus cosmius		1	1938-09-05									Mexico	Michoacan de Ocampo		Chinapa	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203CA0FF0C76CEFF36	03D6B32E820CD4203ECAFCEC7764FA22.taxon			SEMC	paratype	Paratype (2 ♀ SEMC)	female	2										Mexico	Morelos			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203DB3FF0C713AFEFE	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	[unpublished paratypes]. MORELOS: Mpio. Amacuzac, Río Amacuzac 1.6 km S (on dirt road) of km 50 carr. MEX 95 Cuernavaca-Taxco, 18 º 35 ' 48.05 '' N, 99 º 23 ' 57.88 '' W, 917 m, 04 - I- 2017, L- 1990, Reynoso-Velasco coll. (2 ♀ IEXA)	female	2	2017-01-04	Reynoso-Velasco		18.596682	-99.39941	917				Mexico	Morelos	Rio Amacuzac	Cuernavaca-Taxco	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203A71FEB47072FEA2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC	paratype	Mpio. Jojutla, Río Amacuzac at Tehuiztla, 18 º 33 ' 45.2 '' N, 99 º 16 ' 51.3 '' W, 853 m, 28 - IV- 2011, L- 1308 Reynoso-Velasco coll. (4 ♂, 1 ♀ UMC)	male	5	2011-04-28	Reynoso-Velasco		18.562555	-99.28092	853				Mexico	Morelos	Rio Amacuzac	Tehuiztla	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203B3EFE90764BFE86	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC	paratype	same but 18 º 32 ' 47.9 '' N, 99 º 16 ' 03.6 '' W, 854 m, L- 1309 (1 ♂, 3 ♀ UMC)	male	4				18.546638	-99.26767	854				Mexico	Morelos			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203D27FEFC71AFFE6A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	Mpio. Xochitepec, [Río Cuentepec at] Balneario Palo Bolero, [approx. 18.766 8, - 99.2393], 07 - VII- 1987, R. Novelo coll. (1 ♂, 2 ♀ IEXA)	male	3	1987-07-07	R. Novelo								Mexico	Morelos	Mpio. Xochitepec	Balneario Palo Bolero	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203AC5FED875C3FE32	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	Mpio. Xochitepec, Río Sabinos (Cuentepec) at Jardines de Xochitepec, [approx. 18.778 9, - 99.2439], 24 - IX- 1998, R. Novelo coll. (1 ♂, 2 ♀ IEXA)	male	3	1998-09-24	R. Novelo								Mexico	Morelos	Rio Sabinos	Cuentepec	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203EAFFE007754FE32	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	same but 05 - IV- 2002, (3 ♀ IEXA)	female	3	2002-04-05									Mexico	Morelos			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203C21FE007100FE32	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	same but 24 - IV- 2003 (1 ♂, 2 ♀ IEXA)	male	3	2003-04-24									Mexico	Morelos			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203A73FE0174C5FDFA	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC	paratype	OAXACA: Mpio. Putla Villa de Guerrero, Putla, [Río Cuchara] at Pte. La Cuchara, 17 º 01 ' 07 '' N, 97 º 55 ' 11.8 '' W, 711 m, 30 - V- 2012, L- 1396, ReynosoVelasco coll. (2 ♂, 1 ♀ UMC)	male	3	2012-05-30	ReynosoVelasco		17.01861	-97.919945	711				Mexico	Oaxaca	Putla	Pte. La Cuchara	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203FB0FE4870BCFDFA	03D6B32E820CD4203ECAFCEC7764FA22.taxon				paratype	same but [Río Cuchara at] 17 º 02 ' 21.5 '' N, 97 º 54 ' 42.9 '' W, 767 m, L- 1397 (1 ♂ UMC)	male	1				17.039305	-97.91192	767				Mexico	Oaxaca		Rio Cuchara	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203ECAFD94718CFD82	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	Mpio. San Antonio Nanahuatípam, Río Salado at San Antonio Nanahuatípam, 18 ° 07 ' 59.4 '' N, 97 ° 07 ' 42.6 '' W, 790 m, 24 - III- 2015, L- 1877, Reynoso-Velasco, Sites, Shepard, & Barr colls. (4 ♂, 10 ♀ IEXA)	male	14	2015-03-24	Reynoso-Velasco & Sites & Shepard & Barr		18.133167	-97.1285	790				Mexico	Oaxaca	Rio Salado	San	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203AEAFDF074C6FD4A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMMZ	paratype	[Mpio. San Antonio Nanahuatípam], Río Saldo at [San Gabriel] Casa Blanca, [approx. 18.155 1, - 97.1371], 01 - III- 1976, R. R. & F. H. Miller, M 76 - 41 (1 ♀ UMMZ)	female	1	1976-03-01	R. R. & F. H. Miller								Mexico	Oaxaca	Rio Saldo	Casa Blanca	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203FB0FD387060FD2E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	Mpio. San Felipe Jalapa de Díaz, Río Sto. Domingo at Sto. Domingo, 18 ° 02 ' 15.9 '' N, 96 ° 32 ' 57.2 '' W, 100 m, 25 - IV- 2015, L- 1881, Reynoso-Velasco, Sites, Shepard & Barr colls. (5 ♂, 4 ♀ IEXA)	male	9	2015-04-25	Reynoso-Velasco & Sites & Shepard & Barr		18.03775	-96.549225	100				Mexico	Oaxaca	Rio Sto. Domingo	Sto. Domingo	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203B12FD04715FFCF6	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC	paratype	[Mpio. San Juan Bautista Cuicatlán], Río Las Huertas [Las Vueltas] at Santiago Dominguillo, 12 - VII- 1996, R. Barba & A. Rojas colls. / Colección del Instituto de Biología, UNAM. México D. F. (2 ♂, 3 ♀ UMC)	male	5	1996-07-12	R. Barba & A. Rojas								Mexico	Oaxaca	Santiago Dominguillo	Coleccion del Instituto de Biologia	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203ADAFD4C769CFCBE	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CNIN	paratype	Mpio. San Juan Bautista Cuicatlán, [Río Las Vueltas at] Santiago Dominguillo, 17 º 38 ' 46 '' N, 96 º 54 ' 36.7 '' W, 726 m, 01 - VI- 2012, L- 1405, Reynoso-Velasco coll. (1 ♂, 2 ♀ CNIN; 4 ♂, 9 ♀ IEXA; 18 ♂, 16 ♀ UMC)	male	50	2012-06-01	Reynoso-Velasco		17.64611	-96.910194	726				Mexico	Oaxaca	San Juan	Santiago Dominguillo	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203D9EFCF474EAFC46	03D6B32E820CD4203ECAFCEC7764FA22.taxon			EMEC	paratype	[Mpio. San Pedro Totolápam], Rio Grande [at] Totolapan [Totolápam], CL 1063, 29 - IV- 1964, J. T. & M. S. Polhemus / Ambrysus cosmius ' 67 La Riv. Det. J. T. Polhemus (1 ♂ EMEC)	male	1	1964-04-29	J. T. & M. S. Polhemus								Mexico	Oaxaca	Rio Grande	Totolapan	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203F87FC3C7133FC46	03D6B32E820CD4203ECAFCEC7764FA22.taxon			USNM	paratype	same but / C. J. Drake Accession (4 ♂, 3 ♀ USNM)	male	7		Drake Accession								Mexico	Oaxaca			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203A7EFC3C7600FC0E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			USNM	paratype	[Mpio. Santa María Tecomavaca, Arroyo Los Cués at] San Juan de Los Cués, 14 - VI- 1984, Mario García coll. / Colección del Instituto de Biología, UNAM. México D. F. / C. J. Drake Accession (1 ♀ USNM)	female	1	1984-06-14	Mario Garcia & C. J. Drake Accession								Mexico	Oaxaca	San Juan de Los Cues	Coleccion del Instituto de Biologia	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203D72FC6470BCFC0E	03D6B32E820CD4203ECAFCEC7764FA22.taxon			USNM	paratype	same but / J. T. Polhemus Collection 2014 (1 ♀ USNM)	female	1		Polhemus Collection								Mexico	Oaxaca			https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203ECAFC4077E6FBBA	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CNIN, IEXA	paratype	[Mpio. Santa María Tecomavaca], Río Salado [at Pte. El Saldo], 16 km S of San Juan de Los Cués on carr. MEX 135 Tehuacán-San Francisco Telixtlahuaca, 17 º 55 ' 50.5 '' N, 97 º 00 ' 04.2 '' W, 544 m, 01 - VI- 2014, L- 1407, Reynoso-Velasco coll. (2 ♂, 2 ♀ CNIN; 7 ♂, 15 ♀ IEXA; 2 ♀ UMC)	male	28	2014-06-01	Reynoso-Velasco		17.930695	-97.00117	544				Mexico	Oaxaca	Rio Salado	Tehuacan-San Francisco Telixtlahuaca	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203C93FB887041FB42	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CNIN	paratype	Mpio. Valerio Trujano, Río Grande at Pte. Río Grande, km 124 carr. MEX 135 Tehuacán-San Francisco Telixtlahuaca on jct. to Tomellín, 17 º 45 ' 25 '' N, 96 º 56 ' 59.1 '' W, 607 m, 01 - VI- 2012, L- 1406, Reynoso-Velasco coll. (3 ♂, 2 ♀ CNIN; 21 ♂, 13 ♀ IEXA; 2 ♂, 2 ♀ NHMW; 12 ♂, 13 ♀ UMC)	male	68	2012-06-01	Reynoso-Velasco		17.756945	-96.94975	607				Mexico	Oaxaca	Rio Grande	Rio Grande	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203B2FFB307075FB26	03D6B32E820CD4203ECAFCEC7764FA22.taxon			UMC	paratype	San Cristobal, 17 - VI- 84, H. Velazco coll. / Colección del Instituto de Biología, UNAM. México D. F. (1 ♀ UMC)	female	1		H. Velazco								Mexico	Oaxaca	San	Coleccion del Instituto de Biologia	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203B01FB1C71EAFAEE	03D6B32E820CD4203ECAFCEC7764FA22.taxon			CAS, CNIN	paratype	18 mi NW El Camarón, 20 - VIII- 1959, A. E. Menke coll. / Ambrysus cosmius La Rivers 1953 (Ira La Rivers) / Ira La Rivers Collection bequeathed to the California Academy of Sciences - 1978 (2 ♂, 2 ♀ CAS; 1 ♀ CNIN)	male	5	1959-08-20	A. E. Menke								Mexico	Oaxaca	Ira La Rivers Collection	California Academy of Sciences	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203A84FB4570BDFAD2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			USNM	paratype	25 mi S of Mitla, 04 - I- 1956, J. C. Schaffner / Ambrysus cosmius ' 67 LaR det. J. T. Polhemus / C. J. Drake Accession (1 ♀ USNM)	female	1	1956-01-04	Schaffner & Drake Accession								Mexico	Oaxaca		25 mi S of Mitla	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203ECAFA8D71B5FA9A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	PUEBLA: Mpio. Coxcatlán, [Río Tepazalco at] La Estación, 5 km SW of Coxcatlán, 18 ° 15 ' 07.3 '' N, 97 ° 11 ' 24.5 '' W, 950 m, 24 - III- 2015, L- 1876, Reynoso-Velasco, Sites, Shepard, & Barr colls. (2 ♀ IEXA)	female	2	2015-03-24	Reynoso-Velasco & Sites & Shepard & Barr		18.252028	-97.19014	950				Mexico	Puebla	Coxcatlan	La Estacion	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658200D4203AC3FAE97718FA22	03D6B32E820CD4203ECAFCEC7764FA22.taxon			BYU	paratype	VERACRUZ: [Mpio. Fortín], Río Metalac [Metlac] 3 mi N of El Fortín [Fortín de Las Flores, approx. 18.931 2, - 97.0211], 2 - VII- 1955, R. B. & J. M. Selander (1 ♂, 2 ♀ BYU)	male	3	1955-07-02	R. B. & J. M. Selander								Mexico	Veracruz de Ignacio de la Llave		Rio Metalac	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658204D4243C95FF21740FFF1A	03D6B32E820CD4203ECAFCEC7764FA22.taxon				holotype	MEXICO: MORELOS: [Mpio. Amacuzac, Río Amacuzac at Huajintlán], km 50 [on carr. 95] Cuernavaca-Taxco, [18 º 36 ' 35.56 '' N, 99 º 25 ' 40.38 '' W, 931 m], 09 - III- 1985, M. Albores coll.		1	1985-03-09	M. Albores		18.609879	-99.42788	931				Mexico	Morelos		Cuernavaca-Taxco	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658204D4243F69FF68764BFF1A	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	PARATYPES: same data as holotype (1 ♂ IEXA)	male	1	1985-03-09	M. Albores		18.609879	-99.42788	931				Mexico	Morelos		Cuernavaca-Taxco	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E820CD4203ECAFCEC7764FA22.mc.3B1708658204D4243D21FF5774FEFEA2	03D6B32E820CD4203ECAFCEC7764FA22.taxon			IEXA	paratype	MICHOACÁN: Mpio. Buenavista, Buenavista River at Buenavista Tomatlán, below dam, 19 º 12 ' 57.8 '' N, 102 º 34 ' 30.2 '' W, 440 m, 24 - XII- 2012, L- 1481, ReynosoVelasco, D. coll. (1 ♀ IEXA)	female	1	2012-12-24	ReynosoVelasco, D.		19.216057	-102.57506	440				Mexico	Michoacan de Ocampo	Buenavista River	Buenavista Tomatlan	https://treatment.plazi.org/id/03D6B32E820CD4203ECAFCEC7764FA22	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263FA8FEB47622FE86	03D6B32E8204D4263ECAFB6370BEFD12.taxon				holotype	HOLOTYPE ♀ (by original designation). MEXICO: OAXACA: Mpio. Sta. Maria Zacatepec, km 179 carr. Sta. Maria Zacatepec-Putla, El Rosario, Pte. El Rosario, 29 - May- 2012, L- 1394 / 361 m, 16 º 51 ' 34.6 '' N, 97 º 58 ' 44.7 '' W, Reynoso-Velasco coll.	female	1	2012-05-29			16.859612	-97.97908	361				Mexico	Oaxaca		El Rosario	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263D52FEFC7657FE4E	03D6B32E8204D4263ECAFB6370BEFD12.taxon			IEXA, UMC, CNIN, USNM	paratype	PARATYPES: same data as holotype (3 ♂ IEXA; 6 ♂, 6 ♀ UMC); Mpio. Mesones Hidalgo, km 173 carr. Sta. Maria Zacatepec-Putla, Pte. La Junta, 29 - May- 2012, L- 1395 / 396 m, 16 º 53 ' 59.7 '' N 97 º 58 ' 02 '' W, Reynoso-Velasco coll. (1 ♂ UMC);	male	34	2012-05-27	Reynoso-Velasco		16.705252	-98.014694	396				Mexico	Hidalgo	San	Rio Rana	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263D21FE247772FE16	03D6B32E8204D4263ECAFB6370BEFD12.taxon			IEXA, UMC, CNIN, USNM	paratype	Mpio. San Gabriel Mixtepec, Río Rana, km 194 carr. Puerto Escondido-Oaxaca, San Gabriel Mixtepec, L- 1380 / 27 - May- 2012, 712 m, 16 º 06 ' 06.3 '' N 97 º 03 ' 52.9 '' W, Reynoso-Velasco coll. (1 ♂, 1 ♀ UMC);	male	34	2012-05-27	Reynoso-Velasco		16.705252	-98.014694	396				Mexico	Hidalgo	San	Rio Rana	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263C3DFE6C762AFDFA	03D6B32E8204D4263ECAFB6370BEFD12.taxon			IEXA, UMC, CNIN, USNM	paratype	same but 3 - Apr- 2015, L- 1906, Reynoso-Velasco, Sites, Shepard, & Barr colls., rocks & gravel (5 ♂, 1 ♀ IEXA; 3 ♂, 1 ♀ UMC); [	male	34	2012-05-27	Reynoso-Velasco & Sites & Shepard & Barr		16.705252	-98.014694	396				Mexico	Hidalgo	San	Rio Rana	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263D53FE487074FDDE	03D6B32E8204D4263ECAFB6370BEFD12.taxon			IEXA, UMC, CNIN, USNM	paratype	Mpio. San Gabriel Mixtepec], San Gabriel Mixtepec, 20 - V- 84, M. García / Colección del Instituto de Biología, UNAM. México D. F. (1 ♂ CNIN; 1 ♂, 1 ♀ USNM);	male	34	2012-05-27	M. García		16.705252	-98.014694	396				Mexico	Hidalgo	San	Rio Rana	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263B04FD947685FD66	03D6B32E8204D4263ECAFB6370BEFD12.taxon			IEXA, UMC, CNIN, USNM	paratype	Mpio. Sta. Maria Zacatepec, km 189 carr. Sta. Maria Zacatepec-Putla, S of El Tapanco, L- 1393 / 29 - May- 2012, 341 m, 16 º 47 ' 28.3 '' N 97 º 59 ' 20.4 '' W, Reynoso-Velasco coll. (2 ♂ UMC);	male	34	2012-05-27	Reynoso-Velasco		16.705252	-98.014694	396				Mexico	Hidalgo	San	Rio Rana	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263DF0FDDC765BFD2E	03D6B32E8204D4263ECAFB6370BEFD12.taxon			IEXA, UMC, CNIN, USNM	paratype	Mpio. Sta Maria Zacatepec, Km 201 carr. Mártires de Tacubaya-Santa Maria Zacatepec ca. 13 km NE of San Pedro Amuzgos, L- 1391 / 29 - May- 2012, 276 m, 16 º 42 ' 18.9 '' N 98 º 00 ' 52.9 '' W, Reynoso-Velasco coll. (1 ♂ UMC)	male	34	2012-05-27	Reynoso-Velasco		16.705252	-98.014694	396				Mexico	Hidalgo	San	Rio Rana	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8204D4263ECAFB6370BEFD12.mc.3B1708658206D4263DD4FD0570B2FD12	03D6B32E8204D4263ECAFB6370BEFD12.taxon			UMMZ	paratype	GUERRERO: stream tributary to Laguna Tres Palos, about 22 miles from Acapulco on highway to Mexico City, III: 22: 1951 R. R. & M. Miller (2 ♂, 1 ♀ UMMZ)	male	3		R. R. & M. Miller								Mexico	Guerrero		GUERRERO	https://treatment.plazi.org/id/03D6B32E8204D4263ECAFB6370BEFD12	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393C67FABB71CAFAA8	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		COSTA RICA: Río Virilla, 26 - XII- 1931, Heinrich Schmidt / purchased from collector / Ambrysus variegatus Usinger determined by Ira La Rivers 1948 (1 ♀ SEMC; 1 ♀ UMC).	female	10	1931-07			10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393AA9FA8776F4FA8C	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		[CARTAGO]: Turrialba, Río Tuis (800 msnm), 08 - IV- 1993, R. Novelo coll. (1 ♀ UMC).	female	10	1931-07	R. Novelo & A. E. Z. Short & de Monteverde & R. W. Sites & San Jose & Crandall Reserve, Instituto de Monteverde & Mus. Expd. S. Meek & Field Museum Coll. & San Jose, VI & Heinrich Schmidt		10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393D8BFAE371EBFA70	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		HEREDIA: Río Sarapiqui (margin) just E of Hwy 4 [126], 10 º 23.22 ' N, 84 º 08.20 ' W, alt. 620 ', 25 - VI- 2003, A. E. Z. Short coll. (1 ♂ UMC).	male	10	1931-07	A. E. Z. Short		10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393A8BFACF7620FA39	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		PUNTARENAS: Qda. Máquina at Crandall Reserve, Instituto de Monteverde, 10 º 18 ' 29.80 '' N, 84 º 48 ' 29.56 '' W, elev. 1400 m, 17 - IV 2021, R. W. Sites coll., rootmats in current (1 ♂ UMC). [SAN JOSÉ]: San José, 18 - IV- [19] 12, C. R. / Mus. Expd. S. Meek coll. / Field Museum Coll. (1 ♂ FMNH); San José, VI & VII- 1931, Heinrich Schmidt / bought from collector (2 ♂ SEMC);	male	10	1931-07	R W Sites		10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393D59FA77740CF9C0	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		SAN JOSÉ]: San José, 18 - IV- [19] 12, C. R. / Mus. Expd. S. Meek coll. / Field Museum Coll. (1 ♂ FMNH); San José, VI & VII- 1931, Heinrich Schmidt / bought from collector (2 ♂ SEMC);	male	10	1931-07	Field Museum		10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393F72F9BE707FF9C0	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		same but / bought from Heinrich Schmidt / Ambrysus variegatus det. R. L. Usinger (1 ♂ UMC);	male	10	1931-07			10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
03D6B32E8206D4393ECAF9257609F9A4.mc.3B1708658219D4393B3EF9BE760DF9A4	03D6B32E8206D4393ECAF9257609F9A4.taxon			SEMC, UMC, FMNH		same but Ambrysus variegatus det. H. E. Hungerford (1 ♂ UMC)	male	10	1931-07			10.308278	-84.80821	1400				Costa Rica	Cartago	Heinrich Schmidt	Ira La Rivers	https://treatment.plazi.org/id/03D6B32E8206D4393ECAF9257609F9A4	MaterialCitation
