taxonID	type	description	language	source
1548A2618568FFF6E7C0FA40FBF8861C.taxon	materials_examined	Type species: Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge & Yang, 2013	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618568FFF4E7C0F932FAB387A3.taxon	description	Figs 3, 8 A	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618568FFF4E7C0F932FAB387A3.taxon	materials_examined	Type locality: China: Da-laen-saen (type locality identity is discussed below). Type material examined. HOLOTYPE (male, BMNH): (1) Holotype, (2) G. C. Champion Coll. B. M. 1927 - 409, (3) Da-laen-saen, China. J. J. Walker, (4) HOLOTYPE; (5) Taizonia castanea Gruev sp. n.; ◉ PARATYPES (1 male; 4 female, BMNH): (1) Da-laen-saen, China. J. J. Walker, (2) PARATYPE. Additional material examined. 6 spec. (4 male, 2 female, BMNH): (1) G. C. Champion Coll. B. M. 1927 - 409; (2) Da-laen-saen, China. J. J. Walker; (3) Taizonia castanea Gruev det. A. S. Konstantinov, 2003; 9 spec. (9 male, BMNH): (1) Nyew-tew I., China, J. J. W [alker]; (2) ◉ 4 spec. (3 male, 1 female, NMPC, ADPC): (1) CHINA — Zhejiang prov., Lishui pref., Lu’aocun (炉岙村); Monkey Valley (Ñš ẽ); 27 ° 54 ′ 55.2 ″ N 119 ° 11 ′ 55.4 ″ E; 1100 m; 11. v. 2019; A. F. Damaška lgt.; (2) VOUCHER SPECIMEN DNA collection No. AFD- 235; 236; 282 283; 284 Albert F. Damaška coll.; (3) Cangshanaltica cf. castanea (spec. complex), A. F. Damaška det. 2022 ◉ 2 spec. (2 male, SZPT): (1) Hangzhou Wuyun Mt. (杭州五云山), on moss, under tree root, 2021 - III- 25, leg. Wei Wang (Ɨ ‼).	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618568FFF4E7C0F932FAB387A3.taxon	description	Redescription. Habitus. Body oval-rounded, convex, holotype 1.6 mm long, 1.2 mm wide, 0.9 mm high. Dorsal surfaces chestnut brown without metallic luster. Ventral surfaces light brown. Head. Nearly hypognathous, triangular, chestnut brown. Frontal calli flat, feebly developed, distinctly delimited anteriorly, feebly delimited posteriorly. Frontal ridge wide, flat, not projecting; interantennal space as wide as diameter of eye. Supraantennal, suprafrontal, orbital and supraorbital sulci shallow but developed. Orbit narrow. Frontal ridge, orbit and frons impunctate. Antennae with 11 antennomeres, short, reaching anterior ¼ of elytra. Antennomeres I and II large, wide, light yellowish brown; antennomere III slightly elongated, antennomeres IV – VI similar in length, slender, antennomere VI wider than V. Antennomere VII short, with a distinct distal protrusion; antennomeres VIII – XI gradually widening, forming a long and slender antennal club. Thorax. Pronotum brown, wide, convex, feebly punctate by scattered shallow punctures. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Posterior edges of pronotum sharp. Elytra convex, wide, feebly irregularly punctured. Maximal width of elytra 1 / 3 wider than maximal width of pronotum. Procoxal cavity widely open posteriorly. Anterior metaventral process completely covering mesoventrite between mesocoxae, developing a pentagonal “ horseshoe-like ” structure. Legs light brown. Metatibiae nearly straight, not visibly curved laterally. Metatarsomere I elongated. Abdomen and genitalia. Abdominal ventrite I as long as ventrites II and III combined. Median lobe of aedeagus variable in width, and in sharpness of the apex (Fig. 3 G, H). Apex pointed, arrow-like. Median lobe widened from apex to basal part. Basal orifice pyriform, reaching ¼ of the median lobe of aedeagus length. Spermatheca slender, duct without coils. Vaginal palpi widely separated apically. Generic placement. Cangshanaltica castanea had been described in Taizonia by Gruev (1985), a genus subsequently synonymized with Ivalia by Duckett et al. (2006). Due to homonymy with Ivalia castanea (Samuelson, 1966), a new name Ivalia gruevi was proposed for the species by Nadein (2013). Our placement of Gruev’s species in Cangshanaltica makes Nadein’s replacement name redundant (ICZN Art 59.4). We move the species to Cangshanaltica based on the following characters: (1) antennomere VII bearing a slight distal protrusion; (2) metaventrite bearing an anterior, horseshoe-like process reaching mesocoxae and covering the mesoventrite; (3) anterolateral pronotal setiferous pore placed in the middle of the pronotal margin; and (4) round, ovate, and convex body shape. Cangshanaltica castanea is similar to other Chinese Cangshanaltica species distributed in lowlands — C. sprynari, and C. fuanensis. The species can be distinguished using the key below.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618568FFF4E7C0F932FAB387A3.taxon	materials_examined	Type locality identity and distribution. The type locality “ China: Da-laen-saen ” named by Walker has been discussed by authors who have revised J. Walker’s Chinese material (Schawaller & Aston 2017; Smetana 2009). According to their findings, the most likely type locality of C. castanea is China, Zhejiang prov., Ningbo pref., Dalei Shan (kDZ 山), cca 29.5928 N, 121.2930 E. The second Walker locality in the BMNH material is “ Nyew-tew I. ” This locality refers most likely to the Nantian Dao Island (Éffl 岛) in Zhejiang province, named “ Nyew-tew Island ” in 19 th century Western maps, likely after the island’s peak Niutou Shan (牛Ë山). The second type locality should therefore be: China, Zhejiang prov., Ningbo pref., Nantian Dao (Éffl 岛); cca 29.1361 N, 121.9388 E. Additional specimens assigned to the species complex of C. castanea were collected recently in Lu’aocun (炉岙村) in the Lishui prefecture, and in Hangzhou, both in Zhejiang province, China. Species identity and variability. Cangshanaltica castanea was described based on a type series collected by J. J. Walker. Besides the type series, we examined additional Walker material from the type locality (Da Laen Saen) and its surroundings (Nyew-tew I.). We also collected fresh DNA-grade specimens in southern Zhejiang — they strongly resemble several specimens from the C. castanea type series. All the specimens have similar morphology of external characters and female genitalia, however, the morphology of median lobe of aedeagus is quite variable among all specimens, even within the type material (fig. 3 G – H). Specimens also differ in elytral punctuation (in the holotype and most of Walker’s specimens, elytra have very feeble punctures, in several of Walker’s specimens and all the fresh specimens from Lishui, punctures are shallow but distinctly visible, forming a pitted elytral relief, similar to the surface of a golf ball), and size (males are up to 1.9 mm long). A similar species in external morphology is C. fuanensis Ruan, Konstantinov & Damaška, 2022, which differs in having a more slender median lobe of aedeagus, but is strongly divergent from the Lishui specimens based on DNA phylogenetic analyses (Damaška et al. 2021). This strong difference in both mitochondrial and nuclear DNA markers among the two morphologically similar species suggests that the C. castanea species complex can consist of more species which are similar in morphology but phylogenetically isolated from each other. Due to the lack of any DNA-grade material from the two historical collecting sites in Zhejiang, we decided not to treat any of the specimens with different genitalia morphology as distinct species. However, the variability of median lobe of aedeagus in C. castanea in our understanding is apparently larger than in other Cangshanaltica species. The species definition of C. castanea may therefore represent a species complex. A careful analysis of DNA and morphology based on fresh material from more localities in Zhejiang and around (including Walker’s sites) is the only way to yield enough information to resolve the C. castanea species complex.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF2E7C0FF38FAB7820C.taxon	description	Fig. 8 B	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF2E7C0FF38FAB7820C.taxon	materials_examined	Type locality: China, Fujian, Fuan. Type series: Holotype — IZAS; paratypes: IZAS, SZPT, ADPC.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF2E7C0FF38FAB7820C.taxon	discussion	Remarks: The species was described originally by Ruan et al. (2020), however, the description did not comply with the ICZN, and the name has been considered as nomen nudum. Therefore, it has been described correctly by Ruan et al. (2022). The species is known only from the type series (30 specimens). It is very similar to the C. castanea species complex (from which it can be separated by using the key below) in having brown, ellipticalrounded body. However, Damaška et al. (2021) revealed that C. fuanensis is more closely related to the Taiwanese Cangshanaltica species complex, rather than to the C. castanea s. lat. from Zhejiang to which it is most similar.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF1E7C0F891FEFB8561.taxon	description	Figs 4, 8 C	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF1E7C0F891FEFB8561.taxon	materials_examined	Type locality: China, Guanxi, Panxie Gou. Type material. HOLOTYPE (male, NMPC): (1) CHINA: Guangxi, Pangxie Gou, 1.3 km S of Longsheng Hot Spring, 25 ° 53.0 ′ N 110 ° 12.6 ′ E, 385 m 13. iv. 2013, Fikáček, Hájek & Růžička lgt. (2) sifting of leaf litter and leaf accumulations along stony stream in evergreen lowland subtropical forest [sifting sample 13]; (3) VOUCHER SPECIMEN DNA collection No. AFD- 125 Albert F. Damaška coll.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF1E7C0F891FEFB8561.taxon	diagnosis	Differential diagnosis. The species can be distinguished from all other known Cangshanaltica species by its coloration — elytra ochraceous with one darkened spot in the center of each elytron. DNA sequences. 18 S: MZ 312403.1; 16 S: MZ 312387.1; Cox 1: MZ 303694.1, MZ 220536.1 (barcode). All sequences were published by Damaška et al., 2021.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF1E7C0F891FEFB8561.taxon	description	Description. Habitus. Body round, convex, 2 mm long, 1.7 mm wide, 0.9 mm high. Dorsal and ventral surfaces ochrebrown, each elytron with a darkened spot in the middle. Head. Nearly hypognathous, triangular. Labrum and clypeus blackish brown, vertex brown. Frontal ridge absent; frons shiny, impunctate, with bunches of setae below antennae. Frontal calli nearly absent. Suprafrontal sulcus shallow, orbital and supraorbital sulci developed. Eyes round. Antennae reaching basal 1 / 3 of elytra. Antennomeres I – VI and XI light brown, antennomeres V – X darkened. Antennomeres I and II bulbose, wide; antennomeres VII – XI elongate, as long or longer as antennomeres III and IV combined. Antennomere VII bearing a slight distal protrusion. Thorax. Pronotum brownish ochraceous, wide, convex, impunctate. Anterior margins of pronotum lobate, projecting, forming distinct “ flap-like ” structures towards head; posterior pronotal edges dull. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra convex, brownish ochraceous with a darkened spot in the central part of each elytron, irregularly punctured. Epipleura distinctly widened in basal part. Anterior metaventral process completely covering the mesoventrite between mesocoxae, as a sharply pentagonal, deeply excavated “ horseshoe-like ” structure. Legs brownish ochraceous. Metatibiae slightly curved, apical metatibial spine short. Metatarsomere I slightly elongated. Abdomen. Abdominal ventrite I longer than ventrite II, but not as long as ventrites II and III combined. Anterior process of abdominal ventrite I reaching space between metacoxae, bulbose, forming a sharp, ventrally projecting elevated ridge. Genitalia. Median lobe of aedeagus long but wide, parallel-sided in dorsal view, slightly curved in lateral view. Apex dull, slightly widening posteriorly. Female unknown.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF1E7C0F891FEFB8561.taxon	discussion	Remarks. The only known specimen is apparently not fully sclerotized — the coloration of a mature adult is probably somewhat darker. Despite that, we are describing the species, as all morphological characters are welldeveloped, and available DNA markers clearly indicate its isolated phylogenetic position among other studied Cangshanaltica species (Damaška et al. 2021).	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856CFFF1E7C0F891FEFB8561.taxon	etymology	Etymology. The species name is derived from Guanxi, the name of the Chinese province where the holotype was collected.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856FFFF0E7C0FAD8FB2B873C.taxon	description	Fig. s 5, 8 D	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856FFFF0E7C0FAD8FB2B873C.taxon	materials_examined	Type locality: Indonesia, Java Barat, Gunung Tangkuban Perahu. Type material. HOLOTYPE (male, MHNG): (1) INDONESIE — East Java 30 km N. Bandung, volcan Tangkuban Prahu, J. Robert 10. VIII. 85. ◉ Paratypes (1 male MHNG, 1 male NMPC) same labels as holotype; (2 female MHNG, 1 female NMPC): JAVA Volcan Tangkuban Prahu env 30 km N. Bandung J. Robert 7. VIII. 84.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856FFFF0E7C0FAD8FB2B873C.taxon	diagnosis	Differential diagnosis. The species can be easily distinguished from all known species by its genitalic morphology (median lobe of aedeagus pointed with sharp parallel margins on dorsal side, spermathecal duct long, U shaped). Additionally, the species can be distinguished by the key below.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856FFFF0E7C0FAD8FB2B873C.taxon	description	Description. Habitus. Body ovate, convex, 2.4 mm long, 1.7 mm wide, 1.1 mm high. Dorsal surfaces dark blackish-brown without metallic luster. Ventral side brown. Head. Nearly hypognathous. Labrum light brown, deeply incised; clypeus black. Labial and maxillary palpi light brown. Frons wide, brown, flat, frontal ridge absent. Suprafrontal sulcus well-developed, frontal calli feebly projecting, flat, feebly delimited posteriorly. Orbit narrow, orbital and supraorbital sulci deep. Vertex very finely punctured with shallow scattered punctures. Antennae reaching basal ¼ of elytra. Antennomeres I – IV and XI light yellowish-brown, antennomeres V – X brown. Antennomeres V – XI gradually widening, flat. Thorax. Pronotum wide, convex, punctured with small scattered punctures. Anterior pronotal margins feebly projecting anteriorly, but forming lobes. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra convex, about ¼ wider than maximal width of pronotum, irregularly punctured with distinct punctures twice as large as those on pronotum. Epipleura extremely widened in basal parts of elytra, nearly as wide as half of pronotal width. Procoxal cavity widely open posteriorly. Anterior metaventral process completely covering the mesoventrite between mesocoxae, as a deeply excavated “ horseshoe-like ” structure. Legs light brown, metafemora darkened. Metatibiae slightly curved laterally, with a long apical spine. Metatarsomere I elongated. Abdomen. Abdominal ventrite I as long as ventrites II and III combined. Anterior process of abdominal ventrite I reaching space between metacoxae, bulbose, forming a ventrally projecting elevated ridge. Genitalia. Median lobe of aedeagus long, slender, pointed apically, slightly curved in lateral view. Dorsal side of median lobe of aedeagus with sharp parallel margins narrowing in the middle, ventral side not narrowed. Spermathecal pump as long as receptacle, with digitiform apical part; receptacle twice as wide as pump, almost parallel-sided. Spermathecal duct long, U-shaped, reaching the apical 1 / 3 of the receptacle.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A261856FFFF0E7C0FAD8FB2B873C.taxon	etymology	Etymology. The species is named after the island of Java, where the type series was collected.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFFE7C0FF38FC9B81F8.taxon	description	Fig. 8 E	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFFE7C0FF38FC9B81F8.taxon	materials_examined	Type locality: Philippines, Luzon, Sagada. Type series: Holotype — MHNG, paratypes: MHNG, NMPC.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFFE7C0FF38FC9B81F8.taxon	discussion	Remarks. The species is known only from the type series.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFEE7C0F8ADFDEE84BA.taxon	description	Figs 6, 8 F	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFEE7C0F8ADFDEE84BA.taxon	materials_examined	Type locality: China, Yunnan, 24 ° 29 ′ 31 ″ N 98 ° 52 ′ 57 ″ E Type material. HOLOTYPE (male, DEIM): (1) CHINA (Yunnan), Dehong Dai Aut. Pref., mount. range 31 km E Luxi, 2280 m, 24 ° 29 ′ 31 ″ N / 98 ° 52 ′ 58 ″ E (grassland / pasture, under shrubs, moss / litter, sifted) 3. VI. 2007 D. W. Wrase [19 A]; (2) HOLOTYPE Cangshanaltica marginata sp. nov., Damaška, Ruan, Fikáček des. 2020. ◉ Paratypes (1 male, 2 female UABG, 1 male, 1 female NMPC, 1 female ADPC) have same labels as holotype.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFEE7C0F8ADFDEE84BA.taxon	diagnosis	Differential diagnosis. The species can be distinguished from all other known Cangshanaltica species by its coloration — elytra black with fulvous margin. Additionally, it can be distinguished from all other known species by the shape of the aedeagal apex (widened with a narrow tip).	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFEE7C0F8ADFDEE84BA.taxon	description	Description. Habitus. Body oblong-ovate, convex, holotype 1.8 mm long, 1.2 mm wide, 1 mm high. Head and pronotum brown, elytra black with elytral suture brown, basal and humeral margins of elytra with wide yellow spots, lateral elytral margins and apices of elytra yellow. Ventral side of head and prothorax light yellowish brown, ventral side of mesothorax, metathorax and abdomen brown. Head. Nearly hypognathous. Labrum pitchy black, widely, but not deeply incised. Maxillary and labial palpi brown. Clypeus and frons light brown, feebly pilose, frontal ridge absent. Suprafrontal sulcus shallow, frontal calli almost indistinct. Orbital and supraorbital sulci developed but shallow. Orbit and space around eyes slightly darker than vertex and frons in some specimens. Vertex shiny, almost impunctate. Eyes round. Antennae reaching basal ¼ of elytra. Antennomeres I – V and XI light brown, antennomeres VI – X dark brown. Antennomere VII slightly protruded. Thorax. Pronotum wide, convex, impunctate. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra slightly, irregularly punctured. Humeral part of lateral elytral margin and basal elytral margin with two wide light brownish yellow spots; lateral elytral margin with a wide yellowish-brown stripe; pleura brown. Elytral apices yellow. Procoxal cavity widely open posteriorly; prosternal intercoxal process elevated ventrally. Anterior metaventral process completely covered by mesoventrite between mesocoxae, with a deeply excavated, upturned U-shaped “ horseshoe-like ” structure. Legs brown. Tibiae feebly but distinctly pilose. Metatibiae slightly curved laterally, with a long apical spine. Metatarsomere I not strongly elongated. Abdomen. Ventrites slightly pilose. Abdominal ventrite I with a bulbose anterior process reaching the space between metaxoxae, with a slightly elevated ridge. Abdominal ventrite V slightly paler than ventrites I – IV. Genitalia. Median lobe of aedeagus short, wide and slightly curved in lateral view; in ventral view slightly widened towards apex in basal 2 / 3, apical 1 / 3 of median lobe of aedeagus narrowing, triangular, apex narrowly rounded. Spermatheca slender; spermathecal pump short, receptacle elongate-oval, spermathecal duct shortly Ushaped, without coils, ending in ½ of the receptacle. Vaginal palpi short, joined basally, V-shaped, slightly separated apically. Tignum slender.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618561FFFEE7C0F8ADFDEE84BA.taxon	etymology	Etymology. The species is named after its specific elytral color pattern — black with yellow lateral margin (marginata — with notable margin).	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618560FFFEE7C0FA94FD4086E8.taxon	description	Fig. 8 G	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618560FFFEE7C0FA94FD4086E8.taxon	materials_examined	Type locality: Philippines, Mindanao, Mt. Hamiguitan. Type series: Holotype — NMPC; paratypes: NMPC, ADPC, USNM.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618560FFFEE7C0FA94FD4086E8.taxon	discussion	Remarks. The species is described from moss-sifting samples in Mt. Hamiguitan, Mindanao, Philippines. A probable conspecific specimen is known from Mt. Musuan, Mindanao. In Damaška et al. (2021) the species was sister to all other Cangshanaltica species were included in their analyses (C. siamensis, C. fuanensis, C. guanxiensis, C. castanea, and C. nr. ogatai from Taiwan).	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618560FFFDE7C0F8A5FED68135.taxon	description	Figs 2, 8 H	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618560FFFDE7C0F8A5FED68135.taxon	materials_examined	Type locality: China, Yunnan prov., Dali, Cangshan (Ẽ 山). Type series: Holotype — IZAS, paratypes: IZAS, USNM. Type material examined. PARATYPE (1 male, USNM): China: Yunnan, env. Dali, Cangshan Mountains, 3478 m, 19 July 2011, 25 ° 41.100 ′ N, 100 ° 06.228 ′ E, moss, Konstantinov, Chamorro and Volkovitsh; (2) Paratype Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge, Yang, sp. nov., 2011. Additional material examined. 1 female (UABG): CHINA (Yunnan) Dali Bai Aut. Pref., Jizu Shan, summit plateau, 37 km NE Dali 3150 m 25 ° 58 ′ 30 ″ N, 100 ° 21 ′ 36 ″ E (mixed forest, sifted from litter, moss) 5. IX. 2009 D. W. Wrase [28].	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618563FFFDE7C0F90EFD3187AA.taxon	materials_examined	Type locality: Japan, Okinawa, Iriomote isl. Type series: Holotype — SEHU; paratypes: HSPC, TTPC.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618563FFFDE7C0F90EFD3187AA.taxon	discussion	Remarks. Cangshanaltica ogatai is the only species described, so far, which has an elytral pattern with multiple spots. This coloration is uncommon among Cangshanaltica species distributed in continental Asia. Species with a similar color pattern are distributed also in Taiwan, however, we hesitate to either assign them to C. ogatai or describe them as a new species (see comments below).	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618562FFFCE7C0FF38FCD282ED.taxon	description	Fig. 8 I	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618562FFFCE7C0FF38FCD282ED.taxon	materials_examined	Type locality: Thailand, Chiang Mai, Doi Inthanon. Type series: Holotype — NMPC; paratypes: NMPC, USNM, ADPC. Material examined. 1 spec., UABG: (1) THAILAND — Chiang Mai, Doi Inthanon 2500 m, VI. 87., Barber F., P. Schwedinger lgt.; (2) Taizonia martensi Medvedev, det. Döberl 1999; (3) coll. Manfred Döberl, Abensberg, 2016 coll. Ulf Arnold, Berlin. ◉ 1 male, 1 female, UABG: (1) THAILAND 18. IV / 23. V. 87 prov. Chiang Mai, Doi Inthanon 2500 m BF, P. Schwedinger; (2) Taizonia fulva Medvedev, det. Döberl 1999; (3) Cangshanaltica siamensis Damaška & Konstantinov, 2016 A. F. Damaška det. 2018. ◉ 2 spec., UABG: (1) THAILAND — Chiang Mai, Doi Inthanon 2530 m, VIII. 87., P. Schwedinger lgt.; (2) Taizonia martensi Medvedev, det. Döberl 1999; (3) Cangshanaltica siamensis Damaška & Konstantinov, 2016 A. F. Damaška det. 2018.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618562FFFCE7C0FF38FCD282ED.taxon	discussion	Remarks. The species is only known from Doi Inthanon, Thailand. The newly reported specimens were found in the collections misidentified as different Ivalia species.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618562FFFCE7C0F97AFC9B878B.taxon	description	Fig. 8 J	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618562FFFCE7C0F97AFC9B878B.taxon	materials_examined	Type locality: China, Hong Kong, Lantau. Type series: Holotype: NMPC; paratypes: NMPC, ADPC, AFCD, USNM, SYSU.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618562FFFCE7C0F97AFC9B878B.taxon	discussion	Remarks. The species is only known from the type series.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618565FFFBE7C0FAE6FAC285A4.taxon	discussion	Comments. We were unable to study the type specimen from Erland Shan Pass, Sichuan deposited in Medvedev’s private collection. Instead, we examined a photograph made by A. Moseyko of the box containing the type series of Ivalia sichuana. Based on this photograph, it seems likely that I. sichuana belongs to Cangshanaltica.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618565FFFAE7C0F9EAFDC983F6.taxon	description	Figs 7, 8 K	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618565FFFAE7C0F9EAFDC983F6.taxon	materials_examined	Material examined: 1 female, NMPC: (1) TAIWAN: New Taipei 2.5 km SWW of Wulai City, Meilushan Mt. above Baoqing Temple; 24.85445 ° N 121.53 ° E; 760 m; 6. v. 2018; Fikáček, Chang & Zelený lgt. (2) leaf litter accumulations in primary moist forest with sparse understory: sifting (3) VOUCHER SPECIMEN DNA collection No. AFD- 150 Albert F. Damaška coll. ◉ 1 female, NMPC: (1) TAIWAN: New Taipei 0.8 km SSE of Wulai City, Laka Trail; 24.85652 ° N 121.5536 ° E 5. v. 2018; 320 m; Fikáček, Leong & Chang lgt.; 2018 - TW 11; (2) sifting of small leaf litter accummulations in the secondary lowland moist forest on a steep slope, with sparse understory; (3) VOUCHER SPECIMEN DNA collection No. AFD- 184 Albert F. Damaška coll. ◉ 1 male, NMPC: (1) TAIWAN: New Taipei 2.5 km SWW of Wulai City, Meilushan Mt. above Baoqing Temple; 24.85445 ° N 121.53 ° E; 760 m; 6. v. 2018; Fikáček, Chang & Zelený lgt. 2018 - TW 12; (2) leaf litter accummulations in primary moist forest with sparse understory: sifting; (3) VOUCHER SPECIMEN DNA collection No. AFD- 035 Albert F. Damaška coll. ◉ 1 male, ADPC. (1) TAIWAN: Nantou county, Huisun forest res., Xiaochushan Mt., 24 ° 05 ′ 10.2 ″ N 121 ° 01 ′ 41.3 ″ E; sifting moss along a trail; 4. v. 2019; A. F. Damaška lgt. (2) VOUCHER SPECIMEN DNA collection No. AFD- 246 Albert F. Damaška coll. ◉ 1 female, ADPC: (1) TAIWAN: Nantou county, Huisun forest res., 24 ° 05 ′ 21.2 ″ N 121 ° 01 ′ 47.2 ″ E; FIT; 5. v. 2019; Hu Fang-Shuo lgt. ◉ (2) VOUCHER SPECIMEN DNA collection No. AFD- 287 Albert F. Damaška coll. ◉ 1 spec., TARI: (1) Taiwan, Taipei (# 22351), Neishuanghsi (內 ḎẔ); 17. IV. 2014. leg. H. - J. Chen ◉ 3 spec, TARI: (1) Taiwan, Nantou, Tatachia (ḆḆDZ), 16 – 23. VI. 2007, leg. C. - S. Tung	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
1548A2618565FFFAE7C0F9EAFDC983F6.taxon	discussion	Comments. In Taiwan, Cangshanaltica is a common genus, with at least two species well-delimited based on molecular analysis (Damaška et al. 2021). All examined species from Taiwan have spotted elytra and are generally similar to the Japanese C. ogatai Takemoto & Suenaga, 2021 in all external characters, except for the coloration: Taiwanese specimens have 4 elytral spots, C. ogatai has 6 elytral spots. The dorsal body coloration among different Taiwanese specimens is variable (for example, pronotum can be yellow or black, elytral suture yellow or black) and body size also varies greatly. However, the coloration variability does correspond to the molecular data provided by Damaška et al. (2021). We examined numerous specimens of Cangshanaltica from Taiwan, however, only a few provided molecular sequences, which made a broad molecular analysis impossible, similar to the case of C. castanea. The type series of C. ogatai also does not contain any DNA-grade specimens (H. Suenaga, pers. comm., 2021). An analysis containing both molecular and morphological characters would be, however, necessary to delimit the species. The fauna of the Iriomote island is very similar to Taiwan, both islands even share two species of the edaphic flea beetle genus Clavicornaltica (Suenaga & Yoshida 2016). It is therefore likely that C. ogatai may be distributed in Taiwan. Taiwanese Cangshanaltica diversity is clearly complicated and needs more data to be properly taxonomically resolved. As in the case of C. castanea, we therefore hesitate to describe any new species of Cangshanaltica from Taiwan.	en	Damaška, Albert František, Ruan, Yongying, Fikáček, Martin (2022): The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes. Zootaxa 5219 (1): 49-64, DOI: https://doi.org/10.11646/zootaxa.5219.1.2
