taxonID	type	description	language	source
038E8D48617DA842FF73FC64F558F827.taxon	type_taxon	Type species: Physciolagria liturata Pic, 1930, monotypy.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617DA842FF73FC64F558F827.taxon	description	Physciolagria was originally erected to hold one African species, P. liturata Pic, 1930, on the basis of the following features (translated from Latin and annotated by the author for clarity in accordance with the present situation (as in the studied type specimen); author’s additions in square brackets): “ Body elongated; head short, eyes strong, strongly convex, not emarginate, setose; terminal [maxillary] palpomere elongate, truncate at apex; antennae short, strongly widened, second antennomere short, 3 rd antennomere triangular, distally widened, antennomeres 4 – 10 short and strongly transverse, terminal antennomere elongate, rather wide and depressed [possible meant flattened dorso-ventrally], subtruncate at apex; pronotum laterally emarginated, [pronotal hypomeron] impressed at base, [pronotal] base beaded and [slightly] impressed, anterolateral angles obsolete, posterolateral angles nearly rightangled; elytra wide, elongated, epipleuron narrow, posteriorly indistinct; legs slender, tarsi elongated, penultimate tarsomeres lobate, tarsal claws [basally] subdentate; pro- and mesocoxa conical, contiguous ” (Pic 1930: 7). The new genus was compared with Physcius Champion, 1889 (now Mycteridae: Eurypinae, at the time of description in Lacconotinae) but Pic (1930) informally placed Physciolagria in Lagriinae of Tenebrionidae (Lagriidae at the time of description) (translated from French): “ but seems to be that it needs to be placed inside the Lagriidae, rather than the Lacconotinae ” (Pic 1930: 7). Borchmann (1936) listed Physciolagria along other Lagriinae (Lagriidae at the time of the publication) genera, but did not include it in his key to Lagriinae genera, stating that (translated from German, my additions in square brackets) “ inadequate description prevented [from] including [it] in the key if [it] belongs in the family at all ” (Borchmann 1936: 23). Physciolagria had not been rediscovered and redescribed since that, so Bouchard et al. (2021) listed it among incertae sedis genera of Tenebrionidae.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA841FF73FF2FF258FBA2.taxon	materials_examined	Type species: Lacconotopedilus singularicornis Pic, 1935, monotypy. Type material. holotype ♀ L. singularicornis [MNHN]: Afrique Limawe [handwritten; it also could read “ Limowe ” or “ Limave ” or “ Limare ”] // type [handwritten] // Lacconotopedilus n g. singularicornis n sp [handwritten] // type [handwritten] // TYPE [printed, label red] // Museum Paris Coll. M. Pic [printed, label bluish] // HOLOTYPE [printed] Lacconotopedilus singularicornis Pic [handwritten] Examined by D. A. Pollock 19 [printed] 97 [handwritten] [label red, black frame] // MNHN, Paris EC 17269 [printed, provided with a QR code]. The type has extensively damaged elytra and metathoracic legs except for a basal fragment of the left metafemur are missing. Lacconotopedilus was established by Pic (1935) for one species, L. singularicornis, and was initially attributed to Lacconotinae (now a junior synonym of Eurypinae) of Mycteridae, but without a formal family placement. Pollock (1997) redescribed and figured Lacconotopedilus and the two species known at that time, and confirmed the placement of this genus within Mycteridae. Three species (Pic 1935, 1936; Telnov 2019) are currently attributed to this “ one of the most distinctive genera ” of Eurypinae as stated by Pollock (1997: 51). The most recent key to, and an annotated checklist of Lacconotopedilus were presented by Telnov (2019). Study of the type species of both Physciolagria and Lacconotopedilus suggests both genera are congeneric. Physciolagria liturata was described from a single female which is holotype by monotypy (Pic 1930: 7, footnote 1); L. singularicornis was also described from a single female, the holotype by monotypy (Pic 1935: 35). Physciolagria Pic, 1930 is therefore transferred to Eurypinae (Mycteridae) and a new synonym is proposed. Moreover, based on the study of the type material, P liturata was found to be conspecific with L. elongatus, and a new synonym is therefore proposed. Since one of the three species hitherto attributed to Lacconotopedilus appear invalid (junior synonym of P. liturata, see below), two new combinations are proposed: Physciolagria pollocki (Telnov, 2019) comb. nov., Physciolagria singularicornis (Pic, 1930) comb. nov. (both from Lacconotopedilus).	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA841FF73FBCBF539FA8A.taxon	description	(Figs 1 – 11, 20 – 21, 28 – 29, 36 – 38) Holotype ♀ BMNH, data as for the genus, provided above.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA846FF73FAE2F4E6F8F8.taxon	materials_examined	Type material. L. elongatus, holotype ♀ [MNHN] (Figs 5 – 8): U. 4 / IV. Co 2280 [handwritten] // Tanganyika (Conrads) [handwritten] // elongatus n sp [handwritten] // type [handwritten] // Museum Paris Coll. M. Pic [printed, label bluish] // TYPE [printed, label red] // HOLOTYPE [printed] Lacconotopedilus elongatus Pic [handwritten] Examined by D. A. Pollock 19 [printed] 97 [handwritten] [label red] // MNHN, Paris EC 15939 [printed, provided with QR code]. The type is missing both terminal antennomeres and both forelegs. Pollock (1997: 55 – 56) redescribed Lacconotopedilus from the type of L. singularicornis and additional material from Botswana, south-eastern DR Congo, and South Africa. Similar as in my case, Pollock was also unable to read in full the locality label of the holotype L. singularicornis: “ Afrique [with other illegible name, something like Lrimave] ” (Pollock 1995: 55) and therefore suggested possible conspecifity of L. elongatus and L. singularicornis but did not propose the formal synonymy due to lack of comparative material (Pollock 1997: 53). The holotype of L. singularicornis was obtained from unknown locality, since I was not able to attribute the type locality (spelled as Limawe, Limowe, Limare or Limare) to any existing place in Africa. The oldest label of the holotype of L. elongatus is handwritten by an unknown person, while the next one, claiming the specimen is from Conrads material from “ Tanganyika ”, was undoubtedly added later by Pic (judging from handwriting on both labels). Had the type specimen really been collected by P. Conrads, the letter “ U. ” on the initial label likely refers to Ukerewe Island, Lake Victoria, Tanzania, where Conrads extensively collected 1912 – 1931 (Lanjouw & Stafleu 1954). In support of this hypothesis, there are nine syntypes of Nanophyes conradsi Marshall, 1939 (Brentidae Billberg, 1820) present at BMNH, all described from Conrads collected material from Ukerewe Island, each bearing handwritten collecting locality code label “ 2215 ” which is close to the “ 2280 ” code as on the initial label of the L. elongatus holotype.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA846FF73FAE2F4E6F8F8.taxon	description	Measurements. Holotype ♀ (Figs 1 – 4), total body length 8.3 mm; head length 1.3 mm, head width across eyes 1.4 mm, pronotal length 1.2 mm, maximum pronotal width 1.5 mm, minimum pronotal 1.2 mm, elytral length 5.8 mm, combined maximum elytral width in apical third 3 mm. Selected comparative ♂♂ from Tanzania 5.7 – 5.8 mm long (see Checklist for locality data). Redescription, ♂ from Ukiriguru, Tanzania (Figs 9 – 11; consider Checklist below for specimen label data): Dorsal forebody generally dark brown, pronotal basal area narrowly paler, clypeus and labrum dirty yellow to yellowish brown. Elytra ochre with black, strongly elongate sutural spot in apical third leaving suture and apex of elytra narrowly yellow. Antennae black. Maxillary palps pale brown. Legs black-brown to brown, tarsi yellowish brown, trochanters yellowish brown. Venter generally brown, ventral and posterolateral parts of prothoracic hypomeron yellow to yellowish brown. Head (Figs 9 – 10) transverse, moderately glossy dorsally, flattened, with extremely large and globose hemispherical compound eyes occupying most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, rather long. Frontoclypeal suture or impression not indicated. Labrum broadly emarginate at anterior margin. Frons extremely narrow between compound eyes (at its narrowest point frons is narrower than length of basal antennomere), strongly widens posteriad. Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures dense and rather coarse, irregular (especially anteriad eyes). Intervening spaces moderately glossy on frons. Head dorsal setation yellowish, inconspicuous, suberect. Antenna (Fig. 11) extending beyond mesocoxa when directed backwards. Basal antennomere short, about 1.2 × as long as wide. Antennomere three about 1.2 – 1.3 × as long as antennomere two. Antennomeres 4 – 8 asymmetrical, 4 – 10 strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, rounded apically, with an inconspicuous, obtuse angulation about midlength of anterior margin, significantly longer than combined length of antennomeres 1 – 10. Terminal maxillary palpomere slightly fusiform, narrow. Pronotum (Figs 9 – 10) moderately glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate at anterior margin. Lateral margins subparallel. Anterolateral angles obsolete, posterolateral angles broadly obtuse to rounded. Pronotal base broadly rounded, with narrow bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, narrower than to twice as wide as punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, suberect. Scutellar shield triangular, rounded apically. Elytra (Fig. 9) elongate, nearly parallel-sided, slightly glossy, flattened dorsally, somewhat ‘ malacoderm-like’ thin and flimsy. Humerus broadly rounded. Punctures on elytral disc coarser and denser than those on pronotum, with irregular intervening spaces which are generally narrower than punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, suberect, moderately long, directed posteriad. Sutural striae not indicated. Epipleuron short, moderately broad, present from humerus towards about posterior margin of mesocoxal cavity. Metathoracic wings fully developed. All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Pro- and mesothoracic tibia about as long as corresponding femur, metathoracic tibia longer than metathoracic femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of all remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII subtruncate at posterior margin (Fig. 20), morphological sternite VII short and broad, subtruncate at posterior margin (Fig. 21). Tergite VIII and morphological sternite VIII short and broad, subtruncate at posterior margin. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally in dorsal and lateral view, strongly narrowed towards rounded apex (Figs 36 – 38). Gonopore armature as in Fig. 38. Apical portion with several pores, apical margin with 5 – 6 radial channels (Figs 36 – 37). Tegmen stout, with parameres externally evenly, broadly curved; paramere apex rounded, with several moderately long setae raising from pores (Figs 28 – 29). Outer margin of paramere narrow (Fig. 28). Inner margin of paramere nearly straight except at apex where it strongly curves inwards (Fig. 28 – 29). Ventral side of paramere with sparse, moderately long setae (Fig. 29). Redescription, ♀ holotype. Female (Figs 1 – 3) generally longer than male (see measurements, above). Frons comparatively wider between compound eyes. Antenna (Figs 1, 3) comparatively shorter than that of male. Antennomere three triangular, twice as long as antennomere two. Antennomeres 4 – 9 slightly asymmetrical, 5 – 10 strongly transverse, 9 – 10 distinctly broadly emarginate at posterior margin. Terminal antennomere asymmetrically ovoid, truncate apically, about as long as 4 – 4.5 preceding antennomeres. Terminal maxillary palpomere subtriangular, slightly widened apically. Basal metathoracic tarsomere longer than combined length of all remaining metathoracic tarsomeres. Tergite VII and morphological sternite VII subtruncate at posterior margin. Intraspecific variability. Specimens significantly vary in body length (see Measurements). Colour of pronotum varies from entirely black to brown. In ♀ holotype of P. liturata the dark apical elytral spot is not touching sutural or apical margin leaving them narrowly yellow. Elytron with or without indistinct tracks of four vaguely raised longitudinal costae.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA846FF73FAE2F4E6F8F8.taxon	biology_ecology	Ecology. Some specimens were attracted to light.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617AA844FF73FA4EF7AFF9FC.taxon	description	(Figs 12 – 14, 22 – 23, 26, 30 – 32, 39 – 41, 45 – 48)	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617AA844FF73FA4EF7AFF9FC.taxon	materials_examined	Type material: see above. This redescription is made from a male specimen collected in Botswana in the BMNH (see Telnov (2019: 783) for details).	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617AA844FF73FA4EF7AFF9FC.taxon	description	Measurements. Holotype ♂ (Figs 12 – 14), total body length 4.4 mm; head length 0.8 mm, head width across eyes 0.9 mm, pronotal length 0.8 mm, maximum pronotal width 1.0 mm, minimum pronotal width 0.8 mm, elytral length 2.8 mm, combined maximum elytral width in apical third 1.4 mm. Redescription, ♂: Head pale brown. Pronotum and elytra yellow, the latter with black, strongly elongate sutural spot in apical third touching suture but leaving apex of elytra narrowly yellow. Antennae dark brown, terminal antennomeres black. Maxillary palps pale brown. Legs brown, terminal tarsomeres yellow, trochanters brown. Venter uniformly brown. Head (Figs 12 – 14) slightly transverse (nearly circular), glossy dorsally, slightly convex in dorsal aspect, with extremely large and globose hemispherical compound eyes which occupy most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, moderately long. Frontoclypeal suture or impression not indicated. Labrum appears subtruncate at anterior margin. Frons extremely narrow between compound eyes (at its narrowest point frons is as long as to insignificantly narrower than length of basal antennomere), strongly widens posteriad. Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures large, dense and rather coarse (especially on anterior area), irregular. Intervening spaces glossy on occiput, subopaque on anterior area of head. Head dorsal setation yellowish, inconspicuous, suberect. Antenna (Figs 12 – 14) extending towards mesocoxa when directed backwards. Basal antennomere about 1.4 × as long as wide. Antennomere three barely shorter than antennomere two. Antennomeres 6 – 8 asymmetrical, 5 – 10 strongly shortened and strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, truncate apically, at anterior margin slightly narrowed apicad from about midlength where there is a minute, obtuse angulation, about 1.6 – 1.7 × as long as combined length of antennomeres 1 – 10. Terminal maxillary palpomere cylindrical, narrow. Pronotum (Figs 12 – 14) glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate at anterior margin. Lateral margins subparallel. Anterolateral angles rounded, posterolateral angles broadly obtuse. Pronotal base broadly rounded, with moderately wide bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, generally narrower than punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, suberect; setae along lateral margins longer and denser, somewhat curved apically. Scutellar shield triangular, rounded apically. Elytra (Figs 12 – 13) elongate, nearly parallel-sided, moderately glossy, flattened dorsally, somewhat ‘ malacoderm-like’ thin and flimsy. Humerus broadly rounded. Punctures on elytral disc dense, with glossy intervening spaces which are generally narrower than to as wide as punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, subdecumbent, moderately long, directed posteriad. Sutural striae not indicated. Metathoracic wings fully developed. All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Tibia about as long as corresponding femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII subtruncate at posterior margin (Fig. 22), morphological sternite VII short and broad, subtruncate at posterior margin (Fig. 23). Tergite VIII and morphological sternite VIII short and broad, subtruncate at posterior margin. Spiculum gastrale as in Fig. 26. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally, strongly narrowed in apical half, narrowly rounded apically, provided with irregular spine-like setae on its narrow part, with 7 – 8 radial channels at apical margin (Figs 39 – 41). Gonopore armature of numerous long, parallel baculi and one thicker, median rod (Fig. 41). Tegmen stout, with paramere strongly curved at apex only; paramere apex hook-like, rounded (Figs 30 – 32). Ventral surface and external margin of paramere with several long setae most dense on apical area (Figs 31 – 32). Outer and inner margin of paramere broad (Fig. 30). Sexual dimorphism. ♀ (holotype) is over 5 mm long (precise measurement impossible due to the condition of the specimen), terminal antennomere distinctly shorter than that of male with prominent basal denticle at inner margin (Figs 45, 47), pro- and mesothoracic legs rufous, interocular distance comparatively larger than that in male.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617AA844FF73FA4EF7AFF9FC.taxon	biology_ecology	Ecology. Some specimens attracted to light.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617AA844FF73FA4EF7AFF9FC.taxon	distribution	Distribution. The country is unknown for the type locality (see above). Distribution as given by Pollock (1997: 55 – 56) and Telnov (2019: 783) includes Botswana, southern DR Congo (Katanga Province), Namibia, and the Republic of South Africa.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	description	(Figs 15 – 19, 24 – 25, 27, 33 – 35, 42 – 44) https: // zoobank. org / 68 B 79 D 0 D- 502 D- 4536 - 9 B 65 - CD 90 D 7 C 41031	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	type_taxon	Holotype ♂ BMNH: SENEGAL 200 m Dindéfelo Camp 12 ° 22 ′ 43 ′′ N, 12 ° 19 ′ 27 ′′ W 27. v- 2. vi. 2019 Actinic Light [printed] // Aristophanous, M., Moretto, P., Mulvaney, L., leg. ANHRT: 2019.14, BMNH {E} 2022 - 15 [printed] // NHMUK 014663759 [printed, provided with QR code]. Paratypes 18 specimens, all BMNH: 1 ♂ — same labels as holotype but NHMUK 014663758; 1 ♂ — same labels but NHMUK 014663760; 1 ♂ — same labels but NHMUK 014663761; 1 ♂ — same labels but NHMUK 014663762; 1 ♀ — same labels but NHMUK 015018299; 1 ♂ — SENEGAL 30 m Mare de Simenti, Niokolo-Koba NP 13 ° 1 ′ 33 ′′ N, 13 ° 17 ′ 4 ′′ W 3 - 16. vi. 2019 Actinic Light [printed] // Aristophanous, M., Moretto, P., Mulvaney, L., leg. ANHRT: 2019.14, BMNH {E} 2020 - 19 [printed] // NHMUK 015013409 [printed, provided with QR code]; 1 ♂ — same labels but NHMUK 015013394; 1 ♂ — same labels but NHMUK 015013395; 1 ♂ — same labels but NHMUK 015013396; 1 ♂ — SENEGAL 30 m Simenti, Niokolo-Koba NP 13 ° 1 ′ 33 ′′ N, 13 ° 17 ′ 4 ′′ W 3 - 16. vi. 2019 MV LightTrap [printed] // Aristo phanous, M., Moretto, P., Mulvaney, L., leg. ANHRT: 2019.14, BMNH {E} 2020 - 19 [printed] // NHMUK 015013603; 1 ♀ — same labels but NHMUK 015013604; 1 ♂ — same labels but NHMUK 015013605; 1 ♂ — same labels but NHMUK 015018300; 1 ♀ — SENEGAL 30 m Simenti, Niokolo-Koba NP 13 ° 1 ′ 33 ′′ N, 13 ° 17 ′ 4 ′′ W 3 - 16. vi. 2019 LepiLED [printed] // Aristophanous, M., Moretto, P., Mulvaney, L., leg. ANHRT: 2019.14, BMNH {E} 2020 - 19 [printed] // NHMUK 015010042 [printed, provided with QR code]; 1 ♂ — same labels but NHMUK 015010041; 1 ♀ — same labels but NHMUK 015010043; 1 ♂ — same labels but NHMUK 015010044; 1 ♂ — same labels but NHMUK 015010045.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	etymology	Derivatio nominis. Patronymic. This species is named for Richard Smith (The African Natural History Research Trust, Kingsland, United Kingdom) to commemorate his investment in the study of African insects.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	description	Measurements. Holotype ♂ (Figs 15 – 17), total body length 3.7 mm; head length 0.6 mm, head width across eyes 0.9 mm, pronotal length 0.6 mm, maximum pronotal width 0.8 mm, minimum pronotal width 0.7 mm, elytral length 2.5 mm, combined maximum elytral width in apical third 1.3 mm. Selected ♂♂ paratypes 3.5 – 4.3 mm, selected ♀♀ paratypes 4.9 – 5.2 mm long. Description, ♂: Dorsal forebody black, pronotal basal area narrowly paler, clypeus and labrum dirty yellow to yellowish brown. Elytra ochre with black, strongly elongate sutural spot in apical third leaving suture and apex of elytra narrowly yellow. Antennae black. Maxillary palps pale brown. Legs black-brown to brown, tarsi yellowish brown, trochanters yellowish brown. Venter generally brown, ventral and posterolateral parts of prothoracic hypomeron yellow to yellowish brown. Head (Figs 15 – 17) transverse, glossy dorsally, flattened, with extremely large and globose hemispherical compound eyes which occupy most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, rather long. Frontoclypeal suture or impression not indicated. Labrum broadly emarginate at anterior margin. Frons extremely narrow between compound eyes (at its narrowest point frons is narrower than length of basal antennomere), widens posteriad. Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures large, dense and rather coarse, irregular. Intervening spaces glossy on frons. Occiput more regularly punctate, punctures circular, intervening spaces glossy. Head dorsal setation yellowish, inconspicuous, suberect. Antenna (Figs 15, 17) extending slightly beyond mesocoxa when directed backwards. Basal antennomere short, about 1.2 – 1.3 × as long as wide. Antennomere three about 1.5 – 1.7 × as long as antennomere two. Antennomeres 5 – 8 distinctly asymmetrical, 4 – 10 strongly shortened, of them 5 – 10 strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, rounded apically, at anterior margin slightly narrowed apicad from about midlength where there is an obtuse angulation, about 1.4 – 1.5 × as long as combined length of antennomeres 1 – 10. Terminal maxillary palpomere slightly fusiform, narrow. Pronotum (Figs 15 – 17) glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate to broadly rounded at anterior margin. Lateral margins subparallel. Anterolateral angles obsolete, posterolateral angles broadly obtuse to rounded. Pronotal base broadly rounded, with narrow bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, as wide as to 3 × as wide as punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, erect to suberect. Scutellar shield triangular, rounded apically. Elytra (Figs 15 – 16) elongate, nearly parallel-sided, moderately glossy, flattened dorsally, somewhat ‘ malacoderm-like’ thin and flimsy. Humerus broadly rounded. Punctures on elytral disc coarser and denser than those on pronotum, with irregular, glossy intervening spaces which are generally narrower than punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, suberect, moderately long, directed posteriad. Sutural striae not indicated. Epipleuron short, moderately broad, present from humerus towards about posterior margin of mesocoxal cavity. Metathoracic wings fully developed. All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Pro- and mesothoracic tibia about as long as corresponding femur, metathoracic tibia distinctly longer than metathoracic femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII truncate at posterior margin (Fig. 24), morphological sternite VII short and broad, subtruncate at posterior margin (Fig. 25). Tergite VIII short and broad, subtruncate at posterior margin, morphological sternite VIII of two sclerites interconnected by narrow, transparent membrane, subtruncate at posterior margin, short and broad. Spiculum gastrale as in Fig. 27. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally, gradually narrowed in apical half, broadly rounded apically, provided with irregular spines on its narrow part (Figs 42 – 44). Gonopore armature of numerous long, parallel baculi and one thicker, median rod (Fig. 44). Tegmen stout, with paramere externally curved in apical third only; paramere apex hook-like, rounded (Figs 33 – 35). Ventral surface and external margin of paramere with several long setae most dense on apical area (Fig. 35). Outer and inner margin of paramere broad (Fig. 33). Description, ♀. Female (Figs 18 – 19) generally longer and slenderer than male (consider Measurements above). Frons comparatively wider between compound eyes. Antenna (Figs 18 – 19) comparatively shorter than that of male. Antennomere three about as long as antennomere two. Antennomeres 4 – 8 distinctly asymmetrical, 5 – 10 strongly transverse, 9 – 10 distinctly broadly emarginate at posterior margin. Terminal antennomere of similar shape as of male but subtruncate apically, about as long as 4 – 4.5 preceding antennomeres. Terminal maxillary palpomere subtriangular, slightly widened apically. Basal metathoracic tarsomere longer than combined length of remaining metathoracic tarsomeres. Tergite VII and morphological sternite VII subtruncate at posterior margin. Intraspecific variability. Colour of pronotum varies from entirely black to yellowish-brown. Colour of venter vary from yellowish brown to dark brown. Pronotal disc shallowly concave dorsally in one ♂ paratype.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	diagnosis	Differential diagnosis. The new species is readily distinguished from other congeners primarily in the shape of the male genital organs (see Key below).	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	biology_ecology	Ecology. Savannah with patches or gully forest. All available specimens were attracted to various light sources (actinic, LED, MV) at ~ 30 – 300 m elevation. The type locality is within the Western African Mesic Woodland & Grassland vegetation type (Sayre et al. 2013).	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.taxon	distribution	Distribution. Only known from type locality — Niokolo-Koba National Park in SE Senegal.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D486173A84CFF73F92BF48BF85C.taxon	type_taxon	Type species: Physciolagria liturata Pic, 1930, monotypy.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D486173A84CFF73F92BF48BF85C.taxon	type_taxon	Type species: Lacconotopedilus singularicornis Pic, 1935, monotypy.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D486173A84CFF73F92BF48BF85C.taxon	synonymic_list	Physciolagria — Pic (1930: 7) original description, informally in Lagriinae; Borchmann (1936: 11, 23, 232) tentative in Lagriinae; Bouchard et al. (2021: 86, 300) incertae sedis in Tenebrionidae. Lacconotopedilus — Pic (1935: 35) original description, informally in Lacconotinae; Pollock (1997: 51); Pollock (2005: 250); Pollock (2010: 693); Telnov (2019: 779, 781).	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FF20F561FD1D.taxon	synonymic_list	= Lacconotopedilus elongatus Pic, 1936	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FF20F561FD1D.taxon	type_taxon	Type depository. BMNH (P. liturata), MNHN (L. elongatus).	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FF20F561FD1D.taxon	materials_examined	New material. 1 ♂ BMNH: [Kenya] Homa South Kavirondo Nov. - Dec. 1934 H. J. Allen Turner [printed] // Pres. by Comm Inst Ent B. M. 1981 – 315 [printed]; 1 ♂ BMNH: TANGANYIKA UKIRIGURU [printed] 29. 1. [handwritten] 196 [printed] 0 [handwritten] I. A. D. ROBERTSON [printed] L TRAP 22646 [handwritten] // 893 [handwritten] //? CLERIDAE Gen. + sp. incog. [handwritten] R. Madge det. 1965 [printed] // Pres. by Comn. Inst. Ent. BMNH 1981 – 315 [printed].	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FF20F561FD1D.taxon	distribution	Distribution. Kenya (first country record), Tanzania, Uganda.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FD52F41DFC27.taxon	description	Telnov (2019: 780, 782, 783) original description as Lacconotopedilus, checklist.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FD52F41DFC27.taxon	type_taxon	Type depository. BMNH.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FD52F41DFC27.taxon	distribution	Distribution. Eastern Gabon.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FBB7F431FA85.taxon	materials_examined	Type depository. MNHN. Distribution (follows Pollock (1997) and Telnov (2019 )). Botswana, Southern DR Congo (Katanga Province), Namibia, Republic of South Africa.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FAEAF449F9AF.taxon	materials_examined	Type depository. BMNH. Distribution. South-eastern Senegal.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA852FF73F9CEF38BFEF3.taxon	description	Confident identification is yet possible only for males. Female remains unknown for P. pollocki. Since pronotal colour varies in Physciolagria, this feature should be used carefully for treating species.	en	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
