identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038E8D48617DA842FF73FC64F558F827.text	038E8D48617DA842FF73FC64F558F827.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria Pic 1930	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria Pic, 1930</p>
            <p>Pic (1930: 7)</p>
            <p> Type species:  Physciolagria liturata Pic, 1930 , monotypy. </p>
            <p> Type material (Figs 1–4),   holotype ♀ [BMNH]: Type [printed, label circular, red frame] // W. Nile IX-1928. G.D.H.C. handwritten] // Pres. by Imp.Inst.Ent. Brit.Mus. 1930–298. [printed] // Très désiré [handwritten, underlined] //  Physciolagria n gen. liturata  n sp. [handwritten] // NHMUK014663499 [printed, provided with QR code]. The type is in good condition  . </p>
            <p> Physciolagria was originally erected to hold one African species,  P. liturata Pic, 1930 , on the basis of the following features (translated from Latin and annotated by the author for clarity in accordance with the present situation (as in the studied type specimen); author’s additions in square brackets): “Body elongated; head short, eyes strong, strongly convex, not emarginate, setose; terminal [maxillary] palpomere elongate, truncate at apex; antennae short, strongly widened, second antennomere short, 3 rd antennomere triangular, distally widened, antennomeres 4–10 short and strongly transverse, terminal antennomere elongate, rather wide and depressed [possible meant flattened dorso-ventrally], subtruncate at apex; pronotum laterally emarginated, [pronotal hypomeron] impressed at base, [pronotal] base beaded and [slightly] impressed, anterolateral angles obsolete, posterolateral angles nearly rightangled; elytra wide, elongated, epipleuron narrow, posteriorly indistinct; legs slender, tarsi elongated, penultimate tarsomeres lobate, tarsal claws [basally] subdentate; pro- and mesocoxa conical, contiguous” (Pic 1930: 7). </p>
            <p> The new genus was compared with  Physcius Champion, 1889 (now  Mycteridae :  Eurypinae , at the time of description in  Lacconotinae ) but Pic (1930) informally placed  Physciolagria in  Lagriinae of  Tenebrionidae (  Lagriidae at the time of description) (translated from French): “but seems to be that it needs to be placed inside the  Lagriidae , rather than the  Lacconotinae ” (Pic 1930: 7). </p>
            <p> Borchmann (1936) listed  Physciolagria along other  Lagriinae (  Lagriidae at the time of the publication) genera, but did not include it in his key to  Lagriinae genera, stating that (translated from German, my additions in square brackets) “inadequate description prevented [from] including [it] in the key if [it] belongs in the family at all” (Borchmann 1936: 23).  Physciolagria had not been rediscovered and redescribed since that, so Bouchard et al. (2021) listed it among incertae sedis genera of  Tenebrionidae . </p>
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	https://treatment.plazi.org/id/038E8D48617DA842FF73FC64F558F827	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA841FF73FF2FF258FBA2.text	038E8D48617EA841FF73FF2FF258FBA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lacconotopedilus , Afrotropical 1935	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> =  Lacconotopedilus Pic, 1935 syn. nov. </p>
            <p>Pic (1935: 35)</p>
            <p> Type species:  Lacconotopedilus singularicornis Pic, 1935 , monotypy. </p>
            <p> Type material. holotype ♀  L. singularicornis [MNHN]:Afrique Limawe [handwritten; it also could read “Limowe” or “Limave” or “Limare”] // type [handwritten] //  Lacconotopedilus n g. singularicornis n sp [handwritten] // type [handwritten] // TYPE [printed, label red] //Museum Paris Coll.M. Pic [printed, label bluish] // HOLOTYPE [printed]  Lacconotopedilus singularicornis Pic [handwritten] Examined by D.A. Pollock 19[printed] 97 [handwritten] [label red, black frame] // MNHN, Paris EC17269 [printed, provided with a QR code]. The type has extensively damaged elytra and metathoracic legs except for a basal fragment of the left metafemur are missing. </p>
            <p> Lacconotopedilus was established by Pic (1935) for one species,  L. singularicornis , and was initially attributed to  Lacconotinae (now a junior synonym of  Eurypinae ) of  Mycteridae , but without a formal family placement. Pollock (1997) redescribed and figured  Lacconotopedilus and the two species known at that time, and confirmed the placement of this genus within  Mycteridae . Three species (Pic 1935, 1936; Telnov 2019) are currently attributed to this “one of the most distinctive genera” of  Eurypinae as stated by Pollock (1997: 51). The most recent key to, and an annotated checklist of  Lacconotopedilus were presented by Telnov (2019). </p>
            <p> Study of the type species of both  Physciolagria and  Lacconotopedilus suggests both genera are congeneric.  Physciolagria liturata was described from a single female which is holotype by monotypy (Pic 1930: 7, footnote 1);  L. singularicornis was also described from a single female, the holotype by monotypy (Pic 1935: 35).  Physciolagria Pic, 1930 is therefore transferred to  Eurypinae (Mycteridae) and a new synonym is proposed. </p>
            <p> Moreover, based on the study of the type material,  P liturata was found to be conspecific with  L. elongatus , and a new synonym is therefore proposed. </p>
            <p> Since one of the three species hitherto attributed to  Lacconotopedilus appear invalid (junior synonym of  P. liturata , see below), two new combinations are proposed:  Physciolagria pollocki (Telnov, 2019) comb. nov. ,  Physciolagria singularicornis (Pic, 1930) comb. nov. (both from  Lacconotopedilus ). </p>
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	https://treatment.plazi.org/id/038E8D48617EA841FF73FF2FF258FBA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA841FF73FBCBF539FA8A.text	038E8D48617EA841FF73FBCBF539FA8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria liturata Pic 1930	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria liturata Pic, 1930</p>
            <p>(Figs 1–11, 20–21, 28–29, 36–38)</p>
            <p> Holotype ♀ BMNH, data as for the genus, provided above.</p>
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	https://treatment.plazi.org/id/038E8D48617EA841FF73FBCBF539FA8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617EA846FF73FAE2F4E6F8F8.text	038E8D48617EA846FF73FAE2F4E6F8F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lacconotopedilus elongatus Pic 1936	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> =  Lacconotopedilus elongatus Pic, 1936 syn. nov.</p>
            <p> Type material.  L. elongatus ,   holotype ♀ [MNHN] (Figs 5–8): U. 4/IV. Co 2280 [handwritten] // Tanganyika (Conrads) [handwritten] // elongatus n sp [handwritten] // type [handwritten] // Museum Paris Coll. M. Pic [printed, label bluish] // TYPE [printed, label red] // HOLOTYPE [printed]  Lacconotopedilus elongatus Pic [handwritten] Examined by D.A. Pollock 19 [printed] 97 [handwritten] [label red] // MNHN, Paris EC 15939 [printed, provided with QR code]. The type is missing both terminal antennomeres and both forelegs. </p>
            <p> Pollock (1997: 55–56) redescribed  Lacconotopedilus from the type of  L. singularicornis and additional material from Botswana, south-eastern DR Congo, and South Africa. Similar as in my case, Pollock was also unable to read in full the locality label of the holotype  L. singularicornis : “Afrique [with other illegible name, something like Lrimave]” (Pollock 1995: 55) and therefore suggested possible conspecifity of  L. elongatus and  L. singularicornis but did not propose the formal synonymy due to lack of comparative material (Pollock 1997: 53). </p>
            <p> The holotype of  L. singularicornis was obtained from unknown locality, since I was not able to attribute the type locality (spelled as Limawe, Limowe, Limare or Limare) to any existing place in Africa. </p>
            <p> The oldest label of the holotype of  L. elongatus is handwritten by an unknown person, while the next one, claiming the specimen is from Conrads material from “Tanganyika”, was undoubtedly added later by Pic (judging from handwriting on both labels). Had the type specimen really been collected by P. Conrads, the letter “U.” on the initial label likely refers to Ukerewe Island, Lake Victoria, Tanzania, where Conrads extensively collected 1912–1931 (Lanjouw &amp; Stafleu 1954). In support of this hypothesis, there are nine syntypes of  Nanophyes conradsi Marshall, 1939 (  Brentidae Billberg, 1820 ) present at BMNH, all described from Conrads collected material from Ukerewe Island, each bearing handwritten collecting locality code label “2215” which is close to the “2280” code as on the initial label of the  L. elongatus holotype. </p>
            <p>Measurements. Holotype ♀ (Figs 1–4), total body length 8.3 mm; head length 1.3 mm, head width across eyes 1.4 mm, pronotal length 1.2 mm, maximum pronotal width 1.5 mm, minimum pronotal 1.2 mm, elytral length 5.8 mm, combined maximum elytral width in apical third 3 mm. Selected comparative ♂♂ from Tanzania 5.7–5.8 mm long (see Checklist for locality data).</p>
            <p>Redescription, ♂ from Ukiriguru, Tanzania (Figs 9–11; consider Checklist below for specimen label data): Dorsal forebody generally dark brown, pronotal basal area narrowly paler, clypeus and labrum dirty yellow to yellowish brown. Elytra ochre with black, strongly elongate sutural spot in apical third leaving suture and apex of elytra narrowly yellow. Antennae black. Maxillary palps pale brown. Legs black-brown to brown, tarsi yellowish brown, trochanters yellowish brown. Venter generally brown, ventral and posterolateral parts of prothoracic hypomeron yellow to yellowish brown. Head (Figs 9–10) transverse, moderately glossy dorsally, flattened, with extremely large and globose hemispherical compound eyes occupying most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, rather long. Frontoclypeal suture or impression not indicated. Labrum broadly emarginate at anterior margin. Frons extremely narrow between compound eyes (at its narrowest point frons is narrower than length of basal antennomere), strongly widens posteriad. Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures dense and rather coarse, irregular (especially anteriad eyes). Intervening spaces moderately glossy on frons. Head dorsal setation yellowish, inconspicuous, suberect. Antenna (Fig. 11) extending beyond mesocoxa when directed backwards. Basal antennomere short, about 1.2× as long as wide. Antennomere three about 1.2–1.3× as long as antennomere two. Antennomeres 4–8 asymmetrical, 4–10 strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, rounded apically, with an inconspicuous, obtuse angulation about midlength of anterior margin, significantly longer than combined length of antennomeres 1–10. Terminal maxillary palpomere slightly fusiform, narrow. Pronotum (Figs 9–10) moderately glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate at anterior margin. Lateral margins subparallel. Anterolateral angles obsolete, posterolateral angles broadly obtuse to rounded. Pronotal base broadly rounded, with narrow bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, narrower than to twice as wide as punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, suberect. Scutellar shield triangular, rounded apically. Elytra (Fig. 9) elongate, nearly parallel-sided, slightly glossy, flattened dorsally, somewhat ‘malacoderm-like’ thin and flimsy. Humerus broadly rounded. Punctures on elytral disc coarser and denser than those on pronotum, with irregular intervening spaces which are generally narrower than punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, suberect, moderately long, directed posteriad. Sutural striae not indicated. Epipleuron short, moderately broad, present from humerus towards about posterior margin of mesocoxal cavity. Metathoracic wings fully developed. All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Pro- and mesothoracic tibia about as long as corresponding femur, metathoracic tibia longer than metathoracic femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of all remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII subtruncate at posterior margin (Fig. 20), morphological sternite VII short and broad, subtruncate at posterior margin (Fig. 21). Tergite VIII and morphological sternite VIII short and broad, subtruncate at posterior margin. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally in dorsal and lateral view, strongly narrowed towards rounded apex (Figs 36–38). Gonopore armature as in Fig. 38. Apical portion with several pores, apical margin with 5–6 radial channels (Figs 36–37). Tegmen stout, with parameres externally evenly, broadly curved; paramere apex rounded, with several moderately long setae raising from pores (Figs 28–29). Outer margin of paramere narrow (Fig. 28). Inner margin of paramere nearly straight except at apex where it strongly curves inwards (Fig. 28–29). Ventral side of paramere with sparse, moderately long setae (Fig. 29).</p>
            <p>Redescription, ♀ holotype. Female (Figs 1–3) generally longer than male (see measurements, above). Frons comparatively wider between compound eyes. Antenna (Figs 1, 3) comparatively shorter than that of male. Antennomere three triangular, twice as long as antennomere two. Antennomeres 4–9 slightly asymmetrical, 5–10 strongly transverse, 9–10 distinctly broadly emarginate at posterior margin. Terminal antennomere asymmetrically ovoid, truncate apically, about as long as 4–4.5 preceding antennomeres.Terminal maxillary palpomere subtriangular, slightly widened apically. Basal metathoracic tarsomere longer than combined length of all remaining metathoracic tarsomeres. Tergite VII and morphological sternite VII subtruncate at posterior margin.</p>
            <p> Intraspecific variability. Specimens significantly vary in body length (see Measurements). Colour of pronotum varies from entirely black to brown. In ♀ holotype of  P. liturata the dark apical elytral spot is not touching sutural or apical margin leaving them narrowly yellow. Elytron with or without indistinct tracks of four vaguely raised longitudinal costae. </p>
            <p>Ecology. Some specimens were attracted to light.</p>
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	https://treatment.plazi.org/id/038E8D48617EA846FF73FAE2F4E6F8F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617AA844FF73FA4EF7AFF9FC.text	038E8D48617AA844FF73FA4EF7AFF9FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria singularicornis (Pic 1935) Telnov 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria singularicornis (Pic, 1935) comb. nov.</p>
            <p>(Figs 12–14, 22–23, 26, 30–32, 39–41, 45–48)</p>
            <p>Type material: see above.</p>
            <p>  This redescription is made from a male specimen collected in Botswana in the BMNH (see Telnov (2019: 783) for details)  . </p>
            <p>Measurements. Holotype ♂ (Figs 12–14), total body length 4.4 mm; head length 0.8 mm, head width across eyes 0.9 mm, pronotal length 0.8 mm, maximum pronotal width 1.0 mm, minimum pronotal width 0.8 mm, elytral length 2.8 mm, combined maximum elytral width in apical third 1.4 mm.</p>
            <p>Redescription, ♂: Head pale brown. Pronotum and elytra yellow, the latter with black, strongly elongate sutural spot in apical third touching suture but leaving apex of elytra narrowly yellow. Antennae dark brown, terminal antennomeres black. Maxillary palps pale brown. Legs brown, terminal tarsomeres yellow, trochanters brown. Venter uniformly brown. Head (Figs 12–14) slightly transverse (nearly circular), glossy dorsally, slightly convex in dorsal aspect, with extremely large and globose hemispherical compound eyes which occupy most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, moderately long. Frontoclypeal suture or impression not indicated. Labrum appears subtruncate at anterior margin. Frons extremely narrow between compound eyes (at its narrowest point frons is as long as to insignificantly narrower than length of basal antennomere), strongly widens posteriad. Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures large, dense and rather coarse (especially on anterior area), irregular. Intervening spaces glossy on occiput, subopaque on anterior area of head. Head dorsal setation yellowish, inconspicuous, suberect. Antenna (Figs 12–14) extending towards mesocoxa when directed backwards. Basal antennomere about 1.4× as long as wide. Antennomere three barely shorter than antennomere two. Antennomeres 6–8 asymmetrical, 5–10 strongly shortened and strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, truncate apically, at anterior margin slightly narrowed apicad from about midlength where there is a minute, obtuse angulation, about 1.6–1.7× as long as combined length of antennomeres 1–10. Terminal maxillary palpomere cylindrical, narrow. Pronotum (Figs 12–14) glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate at anterior margin. Lateral margins subparallel. Anterolateral angles rounded, posterolateral angles broadly obtuse. Pronotal base broadly rounded, with moderately wide bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, generally narrower than punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, suberect; setae along lateral margins longer and denser, somewhat curved apically. Scutellar shield triangular, rounded apically.Elytra (Figs 12–13) elongate, nearly parallel-sided, moderately glossy, flattened dorsally, somewhat ‘malacoderm-like’ thin and flimsy. Humerus broadly rounded. Punctures on elytral disc dense, with glossy intervening spaces which are generally narrower than to as wide as punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, subdecumbent, moderately long, directed posteriad. Sutural striae not indicated. Metathoracic wings fully developed. All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Tibia about as long as corresponding femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII subtruncate at posterior margin (Fig. 22), morphological sternite VII short and broad, subtruncate at posterior margin (Fig. 23). Tergite VIII and morphological sternite VIII short and broad, subtruncate at posterior margin. Spiculum gastrale as in Fig. 26. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally, strongly narrowed in apical half, narrowly rounded apically, provided with irregular spine-like setae on its narrow part, with 7–8 radial channels at apical margin (Figs 39–41). Gonopore armature of numerous long, parallel baculi and one thicker, median rod (Fig. 41). Tegmen stout, with paramere strongly curved at apex only; paramere apex hook-like, rounded (Figs 30–32). Ventral surface and external margin of paramere with several long setae most dense on apical area (Figs 31–32). Outer and inner margin of paramere broad (Fig.30).</p>
            <p>Sexual dimorphism. ♀ (holotype) is over 5 mm long (precise measurement impossible due to the condition of the specimen), terminal antennomere distinctly shorter than that of male with prominent basal denticle at inner margin (Figs 45, 47), pro- and mesothoracic legs rufous, interocular distance comparatively larger than that in male.</p>
            <p>Ecology. Some specimens attracted to light.</p>
            <p>Distribution. The country is unknown for the type locality (see above). Distribution as given by Pollock (1997: 55–56) and Telnov (2019: 783) includes Botswana, southern DR Congo (Katanga Province), Namibia, and the Republic of South Africa.</p>
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	https://treatment.plazi.org/id/038E8D48617AA844FF73FA4EF7AFF9FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48617BA84CFF73F971F2D9F98B.text	038E8D48617BA84CFF73F971F2D9F98B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria smithi Telnov 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria smithi sp. nov.</p>
            <p>(Figs 15–19, 24–25, 27, 33–35, 42–44)</p>
            <p>https://zoobank.org/ 68B79D0D-502D-4536-9B65-CD90D7C41031</p>
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                  Holotype ♂ BMNH: SENEGAL 200m  
                <a title="Search Plazi for locations around (long -12.324166/lat 12.378612)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-12.324166&amp;materialsCitation.latitude=12.378612">Dindéfelo Camp</a>
                 12°22′43′′N, 12°19′27′′W 27.v-2.vi.2019 Actinic Light [printed] // Aristophanous,M., Moretto,P., Mulvaney, L., leg. ANHRT:2019.14, BMNH {E} 2022-15 [printed] // NHMUK014663759 [printed, provided with QR code]. 
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                 Paratypes 18 specimens, all BMNH :  1♂ —same labels as holotype but NHMUK014663758 ;  1♂ —same labels but NHMUK014663760 ;  1♂ —same labels but NHMUK014663761 ;  1♂ —same labels but NHMUK014663762 ;  1♀ —same labels but NHMUK015018299 ;   1♂ — SENEGAL 30m  
                <a title="Search Plazi for locations around (long -13.284445/lat 13.025833)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-13.284445&amp;materialsCitation.latitude=13.025833">Mare de Simenti</a>
                 , Niokolo-Koba NP 13°1′33′′N, 13°17′4′′W 3-16.vi.2019 Actinic Light [printed] // Aristophanous,M., Moretto,P., Mulvaney, L., leg. ANHRT  :2019.14, BMNH{  E} 2020-19 [printed] // NHMUK015013409 [printed, provided with QR code];  1♂ —same labels but NHMUK015013394 ;  1♂ —same labels but NHMUK015013395 ;  1♂ —same labels but NHMUK015013396 ;   1♂ — SENEGAL 30m  
                <a title="Search Plazi for locations around (long -13.284445/lat 13.025833)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-13.284445&amp;materialsCitation.latitude=13.025833">Simenti</a>
                 , Niokolo-Koba NP 13°1′33′′N, 13°17′4′′W 3-16.vi.2019 MV   LightTrap [printed] // Aristo phanous,M., Moretto,P., Mulvaney, L., leg. ANHRT  :2019.14, BMNH{  E} 2020-19 [printed] // NHMUK015013603 ;  1♀ —same labels but NHMUK015013604 ;  1♂ —same labels but NHMUK015013605 ;  1♂ —same labels but NHMUK015018300 ;   1♀ — SENEGAL 30m  
                <a title="Search Plazi for locations around (long -13.284445/lat 13.025833)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-13.284445&amp;materialsCitation.latitude=13.025833">Simenti</a>
                 , Niokolo-Koba NP 13°1′33′′N, 13°17′4′′W 3-16.vi.2019 LepiLED [printed] // Aristophanous,M., Moretto,P., Mulvaney, L., leg. ANHRT  :2019.14, BMNH{  E} 2020-19 [printed] // NHMUK015010042 [printed, provided with QR code];  1♂ —same labels but NHMUK015010041 ;  1♀ —same labels but NHMUK015010043 ;  1♂ —same labels but NHMUK015010044 ;  1♂ —same labels but NHMUK015010045 . 
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            <p>Derivatio nominis. Patronymic.This species is named for Richard Smith (The African Natural History Research Trust, Kingsland, United Kingdom) to commemorate his investment in the study of African insects.</p>
            <p>Measurements. Holotype ♂ (Figs 15–17), total body length 3.7 mm; head length 0.6 mm, head width across eyes 0.9 mm, pronotal length 0.6 mm, maximum pronotal width 0.8 mm, minimum pronotal width 0.7 mm, elytral length 2.5 mm, combined maximum elytral width in apical third 1.3 mm. Selected ♂♂ paratypes 3.5–4.3 mm, selected ♀♀ paratypes 4.9–5.2 mm long.</p>
            <p>Description, ♂: Dorsal forebody black, pronotal basal area narrowly paler, clypeus and labrum dirty yellow to yellowish brown. Elytra ochre with black, strongly elongate sutural spot in apical third leaving suture and apex of elytra narrowly yellow. Antennae black. Maxillary palps pale brown. Legs black-brown to brown, tarsi yellowish brown, trochanters yellowish brown. Venter generally brown, ventral and posterolateral parts of prothoracic hypomeron yellow to yellowish brown. Head (Figs 15–17) transverse, glossy dorsally, flattened, with extremely large and globose hemispherical compound eyes which occupy most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, rather long. Frontoclypeal suture or impression not indicated. Labrum broadly emarginate at anterior margin.Frons extremely narrow between compound eyes (at its narrowest point frons is narrower than length of basal antennomere), widens posteriad.Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures large, dense and rather coarse, irregular. Intervening spaces glossy on frons. Occiput more regularly punctate, punctures circular, intervening spaces glossy. Head dorsal setation yellowish, inconspicuous, suberect.Antenna (Figs 15, 17) extending slightly beyond mesocoxa when directed backwards. Basal antennomere short, about 1.2–1.3× as long as wide. Antennomere three about 1.5–1.7× as long as antennomere two. Antennomeres 5–8 distinctly asymmetrical, 4–10 strongly shortened, of them 5–10 strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, rounded apically, at anterior margin slightly narrowed apicad from about midlength where there is an obtuse angulation, about 1.4–1.5× as long as combined length of antennomeres 1–10. Terminal maxillary palpomere slightly fusiform, narrow. Pronotum (Figs 15–17) glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate to broadly rounded at anterior margin. Lateral margins subparallel. Anterolateral angles obsolete, posterolateral angles broadly obtuse to rounded. Pronotal base broadly rounded, with narrow bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, as wide as to 3× as wide as punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, erect to suberect. Scutellar shield triangular, rounded apically. Elytra (Figs 15–16) elongate, nearly parallel-sided, moderately glossy, flattened dorsally, somewhat ‘malacoderm-like’ thin and flimsy. Humerus broadly rounded. Punctures on elytral disc coarser and denser than those on pronotum, with irregular, glossy intervening spaces which are generally narrower than punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, suberect, moderately long, directed posteriad. Sutural striae not indicated. Epipleuron short, moderately broad, present from humerus towards about posterior margin of mesocoxal cavity. Metathoracic wings fully developed.All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Pro- and mesothoracic tibia about as long as corresponding femur, metathoracic tibia distinctly longer than metathoracic femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII truncate at posterior margin (Fig. 24), morphological sternite VII short and broad, subtruncate at posterior margin (Fig. 25). Tergite VIII short and broad, subtruncate at posterior margin, morphological sternite VIII of two sclerites interconnected by narrow, transparent membrane, subtruncate at posterior margin, short and broad. Spiculum gastrale as in Fig. 27. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally, gradually narrowed in apical half, broadly rounded apically, provided with irregular spines on its narrow part (Figs 42–44). Gonopore armature of numerous long, parallel baculi and one thicker, median rod (Fig. 44). Tegmen stout, with paramere externally curved in apical third only; paramere apex hook-like, rounded (Figs 33–35). Ventral surface and external margin of paramere with several long setae most dense on apical area (Fig. 35). Outer and inner margin of paramere broad (Fig. 33).</p>
            <p>Description, ♀. Female (Figs 18–19) generally longer and slenderer than male (consider Measurements above). Frons comparatively wider between compound eyes. Antenna (Figs 18–19) comparatively shorter than that of male. Antennomere three about as long as antennomere two. Antennomeres 4–8 distinctly asymmetrical, 5–10 strongly transverse, 9–10 distinctly broadly emarginate at posterior margin. Terminal antennomere of similar shape as of male but subtruncate apically, about as long as 4–4.5 preceding antennomeres. Terminal maxillary palpomere subtriangular, slightly widened apically. Basal metathoracic tarsomere longer than combined length of remaining metathoracic tarsomeres. Tergite VII and morphological sternite VII subtruncate at posterior margin.</p>
            <p>Intraspecific variability. Colour of pronotum varies from entirely black to yellowish-brown. Colour of venter vary from yellowish brown to dark brown. Pronotal disc shallowly concave dorsally in one ♂ paratype.</p>
            <p>Differential diagnosis. The new species is readily distinguished from other congeners primarily in the shape of the male genital organs (see Key below).</p>
            <p> Ecology.   Savannah with patches or gully forest. All available specimens were attracted to various light sources (actinic, LED, MV) at ~ 30–300 m elevation. The type locality is within the  Western African Mesic Woodland &amp; Grassland vegetation type (Sayre et al. 2013)  . </p>
            <p>Distribution. Only known from type locality—Niokolo-Koba National Park in SE Senegal.</p>
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	https://treatment.plazi.org/id/038E8D48617BA84CFF73F971F2D9F98B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D486173A84CFF73F92BF48BF85C.text	038E8D486173A84CFF73F92BF48BF85C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria Pic 1930	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria Pic, 1930</p>
            <p> Type species:  Physciolagria liturata Pic, 1930 , monotypy. </p>
            <p> =  Lacconotopedilus Pic, 1935</p>
            <p> Type species:  Lacconotopedilus singularicornis Pic, 1935 , monotypy. </p>
            <p> Physciolagria — Pic (1930: 7) original description, informally in  Lagriinae ; Borchmann (1936: 11, 23, 232) tentative in  Lagriinae ; Bouchard et al. (2021: 86, 300) incertae sedis in  Tenebrionidae . </p>
            <p> Lacconotopedilus — Pic (1935: 35) original description, informally in  Lacconotinae ; Pollock (1997: 51); Pollock (2005: 250); Pollock (2010: 693); Telnov (2019: 779, 781). </p>
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	https://treatment.plazi.org/id/038E8D486173A84CFF73F92BF48BF85C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FF20F561FD1D.text	038E8D48616CA853FF73FF20F561FD1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria liturata Pic 1930	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria liturata Pic, 1930</p>
            <p> =  Lacconotopedilus elongatus Pic, 1936</p>
            <p> Physciolagria liturata — Pic (1930: 7) original description; Borchmann (1936: 232) German translation of the original description; Bouchard et al. (2021: 300). </p>
            <p> Lacconotopedilus elongatus — Pic (1936: 19) original description; Pollock (1997: 56) redescription; Telnov (2019: 779, 781) checklist. </p>
            <p> Type depository. BMNH (  P. liturata ), MNHN (  L. elongatus ). </p>
            <p>  New material. 1♂ BMNH: [Kenya] Homa South Kavirondo Nov.-Dec. 1934 H.J.Allen Turner [printed] // Pres. by Comm Inst Ent B.M. 1981–315 [printed]; 1♂ BMNH: TANGANYIKA UKIRIGURU [printed] 29. 1. [handwritten] 196 [printed] 0 [handwritten] I. A.D. ROBERTSON [printed] L TRAP 22646 [handwritten] // 893 [handwritten] //?  CLERIDAE Gen. +sp. incog. [handwritten] R. Madge det.1965 [printed] // Pres. by Comn. Inst. Ent. BMNH 1981–315 [printed]  . </p>
            <p>Distribution. Kenya (first country record), Tanzania, Uganda.</p>
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	https://treatment.plazi.org/id/038E8D48616CA853FF73FF20F561FD1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FD52F41DFC27.text	038E8D48616CA853FF73FD52F41DFC27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria pollocki (Telnov 2019) Telnov 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria pollocki (Telnov, 2019)</p>
            <p> Telnov (2019: 780, 782, 783) original description as  Lacconotopedilus , checklist. </p>
            <p> Type depository. BMNH.</p>
            <p>Distribution. Eastern Gabon.</p>
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	https://treatment.plazi.org/id/038E8D48616CA853FF73FD52F41DFC27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FBB7F431FA85.text	038E8D48616CA853FF73FBB7F431FA85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria singularicornis (Pic 1935) Telnov 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria singularicornis (Pic, 1935)</p>
            <p> Lacconotopedilus singularicornis — Pic (1935: 35) original description; Pic (1936: 19) comparative diagnosis; Pollock (1997: 53) redescription, new records; Telnov (2019: 779, 783) checklist, taxonomy, new record. </p>
            <p> Type depository. MNHN . </p>
            <p>Distribution (follows Pollock (1997) and Telnov (2019)). Botswana, Southern DR Congo (Katanga Province), Namibia, Republic of South Africa.</p>
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	https://treatment.plazi.org/id/038E8D48616CA853FF73FBB7F431FA85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA853FF73FAEAF449F9AF.text	038E8D48616CA853FF73FAEAF449F9AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria smithi Telnov 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Physciolagria smithi sp. nov.</p>
            <p>Described above.</p>
            <p> Type depository. BMNH.</p>
            <p>Distribution. South-eastern Senegal.</p>
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	https://treatment.plazi.org/id/038E8D48616CA853FF73FAEAF449F9AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
038E8D48616CA852FF73F9CEF38BFEF3.text	038E8D48616CA852FF73F9CEF38BFEF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Physciolagria Pic 1930	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to species of  Physciolagria</p>
            <p> Confident identification is yet possible only for males. Female remains unknown for  P. pollocki . Since pronotal colour varies in  Physciolagria , this feature should be used carefully for treating species. </p>
            <p> 1 Elytral pubescence double, with distinct, short undersetae and longer ordinary setae; parameres of male aedeagus apically with distinct, curved spine apically; pronotum pale, disc with rufobrunneous, median, longitudinal spot.........................................................................................................  P. pollocki (Telnov)</p>
            <p>- Elytral pubescence simple (without undersetae); parameres of male aedeagus not spinose apically; pronotum nearly uniformly dark or pale, without contrasting median dark spot.......................................................... 2</p>
            <p> 2 Tegmen of aedeagus as in Figs 28–29; median lobe as in Figs 36–38; female terminal antennomere as in Fig. 1, comparatively long, with an inconspicuous basal denticle at inner margin..........................................  P. liturata Pic</p>
            <p>- Tegmen and median lobe of aedeagus different; female terminal antennomere comparatively long, with an inconspicuous basal denticle at inner margin (Figs 18–19) or distinctly shorter, with conspicuous basal denticle at inner margin (Figs 45, 47)... 3</p>
            <p> 3 Tegmen of aedeagus as in Figs 30–32; median lobe as in Figs 39–41; female terminal antennomere rather short, with conspicuous basal denticle at anterior margin (Figs 45, 47); pronotum pale, about same colour as elytra; pronotal dorsal setae about same colour as pronotum.................................................................  P. singularicornis (Pic)</p>
            <p> - Tegmen of aedeagus as in Figs 33–35; median lobe as in Figs 42–44; female terminal antennomere comparatively longer, with inconspicuous median denticle at anterior margin (Figs 18–19); pronotum distinctly darker than elytra; pronotal dorsal setae distinctly paler than pronotum..............................................................  P. smithi sp. nov.</p>
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	https://treatment.plazi.org/id/038E8D48616CA852FF73F9CEF38BFEF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Telnov, Dmitry	Telnov, Dmitry (2022): Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key. Zootaxa 5219 (2): 121-138, DOI: 10.11646/zootaxa.5219.2.2
